taxonID	type	description	language	source
480C87994001761CDE62D0D8FDDCFA64.taxon	materials_examined	Type species: Mygalea antiqua (Pomel, 1848).	en	Ziegler, Reinhard (2003): Moles (Talpidae) from the late Middle Miocene of South Germany. Acta Palaeontologica Polonica 48 (4): 617-648, DOI: 10.5281/zenodo.13396039
480C87994001761DDE62D08BFF28FBCB.taxon	description	Fig. 1. Material and measurements. — Petersbuch 6: NHMA P 6 − 1065 / 1, left dentary fragment with m 1 – m 3; Lm 1 – m 3 (6.45), h of dentary below m 1 (2.80), m 1 (2.38 × 1.17 × 1.36), m 2 (2.32 × 1.25 × 1.35), m 3 (1.97 × 1.09 × 1.02); NHMA P 6 − 1065 / 2, left dentary fragment with m 3 and complete ascending ramus; h of dentary below m 1 (2.70), h 1 of coronoid (9.15), h 2 of coronoid (7.80), m 3 (1.86 × 1.04 × 0.95). Description Dentary. — The coronoid process, which is slightly bent posteriorly, forms a nearly right angle with the horizontal ramus. The masseteric fossa is moderately deep. The internal temporal fossa is deeply excavated. The mandibular foramen opens directly below the mylohyoid ridge, slightly posterior to the centre of the ascending ramus. The angular process is shovel−shaped with an internal concavity and a crest on the external side. The condylar process lies high above the level of the tooth−row. There are two mental foramina: one between the roots of m 1, another beneath p 1 or the posterior root of p 2. There are 10 alveoles anterior to m 1: two for p 2 – p 4 each and one for i 1, i 2, c and p 1 each. Consequently, the mandibular dental formula is 2 − 1 − 4 − 3. According to the alveoles the lower premolars decrease in size anteriorly, the i 1 is procumbent. Lower molars. — The size relation of the lower molars is m 1> m 2> m 3. In the m 1 the trigonid is distinctly narrower than the talonid, in the m 2 only slightly, in the m 3 it is somewhat wider. The protoconid is slightly higher than the hypoconid in the m 1, but distinctly higher in m 2 and m 3. Most conspicuous in the lower molars is the strong cingulid, which extends from below the paraconid to the hypoconid. Postcingulids are somewhat weaker on m 1 and m 2 and rudimentary on m 3. Lingual cingulids are absent. A short entostylid is developed in m 1 and m 2. The oblique cristid terminates at the posterior wall of the trigonid below the protocristid, slightly lingual to the protocristid notch. In the m 2 a faint metacristid is developed. Discussion There are three Mygalea species in the Miocene of Europe. Mygaleamagna Ziegler, 1990, the oldest and largest species, is only known from the type locality Budenheim or Hessler from the Calcareous Tertiary in the Mainz Basin. It is correlated with the Lower Miocene (Middle Agenian, MN 2 a). This species has distinctly larger, primarily wider, teeth than the Petersbuch specimens and has the i 3 retained.	en	Ziegler, Reinhard (2003): Moles (Talpidae) from the late Middle Miocene of South Germany. Acta Palaeontologica Polonica 48 (4): 617-648, DOI: 10.5281/zenodo.13396039
480C87994001761DDE62D08BFF28FBCB.taxon	description	Mygalea is allocated to the Desmaninae by most students except Rümke (1985: 16). She considers the subfamilial allocation not justified, unless the intermediate position of Mygalea can be demonstrated. I think this is not necessary. The humeri of all three Mygalea species are known and show clear desmanine affinities. In the most advanced species, M. antiqua, the i 3 is absent. Hence it cannot be ancestral to any extant desmanine. The genus may be a desmanine side branch that became extinct in the early Late Miocene.	en	Ziegler, Reinhard (2003): Moles (Talpidae) from the late Middle Miocene of South Germany. Acta Palaeontologica Polonica 48 (4): 617-648, DOI: 10.5281/zenodo.13396039
480C87994000761DDE62D18FFF1FF843.taxon	materials_examined	Type species: Leptoscaptor bavaricum gen. et sp. nov. Etymology: From Greek leptos, slender, asthenic; skaptein, to dig, to plug. ComparedtootherScalopini, Leptoscaptor hasaslenderhumerus. Included species: Leptoscaptor robustior gen. et sp. nov. Diagnosis. — Medium−sized scalopine mole. Tentative dental formula I? 1 / 2, C 1 / 1, P 4 / 3, M 3 / 3. i 2> i 1> c. Lower canine incisor−shaped, single−rooted. i 2 more procumbent than c. Lower premolars double−rooted, increasing in size posteriorly. m 1 <m 2> m 3. Oblique cristid joins metacristid in m 2 and m 3, no metastylid developed. Talonid lingually open in m 1. Upper incisor enlarged. Upper canine single− or double−rooted. P 1 – P 3 double−rooted, P 4 with tiny parastyle, protocone in most P 4 a small but distinct cusp. Mesostyles on upper molars divided, no lingual conules on M 1 and M 3, on M 2 weakly developed para− and metaconules. Humerus more or less slender, head elliptical, directed parallel to shaft, brachialis fossa large but shallow, deltoid process short, teres tubercle long, crest−shaped, pectoral tubercle in midshaft position, scalopine ridge prominent, running diagonally from the head to the medial side of the lesser tuberosity, the trochlea is wide, separated by a narrow notch from the fossa for the m. flexor digitorum profundus ligament, olecranon fossa large.	en	Ziegler, Reinhard (2003): Moles (Talpidae) from the late Middle Miocene of South Germany. Acta Palaeontologica Polonica 48 (4): 617-648, DOI: 10.5281/zenodo.13396039
480C879940007616DD28D5BCFC5CFDC8.taxon	description	Fig. 2. Etymology: From Latin bavaricum, Bavarian. The species is recorded from the state of Bavaria, Germany. Holotype: Left dentary fragment with c, p 2 – m 1 and the alveoles of i 1 – i 2, P 10 − 608 / 6, fig. 2 A. Measurementsoftheholotype: lc – p 4 (3.35), lp 2 – p 4 (2.52), c (0.64 × 0.38), p 2 (0.67 × 0.40), p 3 (0.80 × 0.45), p 4 (1.03 × 0.65), m 1 (1.54 × 0.90 × 1.16); h of the corpus below the lingual side of m 1 (1.80). Type locality: Petersbuch 10 (details see p. 618). Age: Uppermost part of the Middle Miocene (MN 8 according to Rummel 2000, means MN 7 + 8). Paratypes (measurements see Tables 1, 2). — Petersbuch 10: NHMA P 10 − 608 / 36, 37, 2 left dentary fragments with teeth; NHMA P 10 − 624 / 1 + 3, 2 left maxilla fragments wilth teeth; NHMA P 10 − 625 D 1 + H 4, 2 upper molars; NHMA P 10 − 610.2 + 612, 2 left humerus fragments; CRW P 10 − 608 – 611, 613, 624, 625, 102 dentary fragments with teeth, 27 maxilla fragments with teeth, 44 isolated teeth, 11 humerus fragments, 5 ulna fragments. Referred material, L. bavaricum vel robustior (measurements see Tables 1, 2). — Petersbuch 6: CRW P 6 − 1063, 5 dentary fragments with teeth; Petersbuch 18: CRW P 18 − 752, 756, 5 dentary fragments with teeth, 2 maxilla fragments with teeth, 5 isolated teeth. Diagnosis. — Leptoscaptor species with slender humerus and a single mental foramen in the vast majority of the dentaries. Description of the holotype Only the horizontal ramus of the dentary anterior to m 2 with c and p 2 – m 1 insitu is preserved. The mental foramen is situated beneath the anterior alveolus of p 3; the symphysis extends posteriorly to c / p 2. The dental formula is reduced, two antemolar teeth being lost: probably the i 3 and the p 1. The lost incisor is interpreted as i 3, because it is the smallest one in those scalopines where it is still present, for example, in Proscapanus and Scalopoides. In the designation of the missing tooth between i 2 and p 2 as the p 1 (Hutchison 1968: 63) is followed. Based on the alveoles, the i 2 was distinctly larger than the i 1, both being procumbent. All teeth are heavily worn. The canine is single−rooted, chisel−shaped and slightly inclined anteriorly. The p 2 – p 4 are double−rooted, increasing in size posteriorly. The protoconid of p 2 and p 3 is centred over the anterior root, a small heel over the posterior one. The p 4 is more inflated, the heel a veritable talonid with posterior cuspule. In m 1 the talonid is distinctly wider than the trigonid. The oblique cristid extends lingually, but does not join the postero−lingual face of the metaconid. The only cingulid is a short ectocingulid below the hypoflexid. The entostylid is small. Description of paratypes and referred material Dentary. — Some other specimens show the alveoli of two incisors, the canine, three premolars, and three molars. The mandibular dental formula is 2 i, 1 c, 3 p, 3 m. There is some variability in the position of the mental foramen. In the Petersbuch 6 sample there are four dentaries with the mental foramen preserved. In one specimen there are two foramina, one below the posterior root of p 2, the other beneath the posterior root of p 4. In three further dentaries a single foramen is situated either beneath the anterior root of p 3, or between the roots or under the posterior root of p 3. In the Petersbuch 10 sample there is consistently one mental foramen: twice between the roots of p 2 and p 3, 24 times below the anterior root of p 3, thrice between the roots of p 3 and five times beneath the posterior root of p 3. In the Petersbuch 18 sample there are four dentaries with the mental foramina preserved: one with one foramen between the roots of p 4 and another between the roots of p 2 and p 3, one with the foramina under the anterior root of m 1 and beneath p 3 / p 4, respectively, and two dentaries with a single foramen below the posterior root of p 4. Lower dentition. — The i 2 has a strong root, a laterally compressed crown with a mesial crest and no cingulids. It is distinctly larger than the canine. The p 2 is similar to p 3, but smaller. The size relation of the lower molars is consistently m 2> m 1> m 3. In the m 1 the trigonid is longer than in the m 2, the talonid is distinctly wider than the trigonid. The oblique cristid terminates at the posterior wall of the trigonid well below the trigonid notch. As there is no metacristid the talonid is lingually open. The m 2 has a very narrow trigonid, as it is known from Proscapanus. The trigonid is wider than the talonid. The oblique cristid runs lingually to join the marked metacristid; the talonid is lingually closed. There is a prominent precingulid and a weak ectocingulid below the hypoflexid. The entostylid is small. The m 3 is distinctly smaller than the m 2, has a talonid reduced in width and no entostylid. Maxilla. — Only two anterior fragments and some with one or two teeth are preserved. The two anterior fragments carry the double−rooted P 1 – P 3 and the alveolus of the canine. In one specimen the canine is single−rooted but in the other double−rooted. Lingually to the canine alveolus of each fragment a foramen pierces the palate, probably the fissura palatina. In front of the canine alveolus there is one large incisor alveolus. The anteriormost part is broken. But on this small flake of bone broken off probably have been no incisor alveoli. Thus two incisors are eliminated, which ones cannot be determined. The maxillary dental formula is? 1 − 1 − 4 − 3. In one specimen the origin of the zygomatic arch above the metastyle of M 2 is preserved; some other fragments show the lacrimal foramen above the anterior root of M 1. Upperdentition. — The antemolar size relation is I >> C ~ P 3> P 1> P 2. P 1 – P 3 are double−rooted and monocuspulate. The P 3 has a small posterior cingulum. The only P 4 of the Petersbuch 18 sample has a small protocone and a tiny parastyle, which is rather a small protuberance of the anterior cingulum. In the Petersbuch 10 sample in 15 P 4 the parastyle is tiny with some transitions to a projecting parastyle in three specimens. The protocone is conical in 13 P 4 and more or less fused with the lingual cingulum in 5 P 4. The mesostyle is clearly divided in all molars. Para− and metaconule are hardly individualised in the M 1. The preprotocrista is continuous with paracingulum, which joins a more or less projecting parastyle. Postproto− and postmetacrista run parallel to one another, the premetacrista parallel to the anterior margin. The four roots are situated above protocone, paracone, metastyle, and a very small one above and slightly labial to the centre. In the M 2 para− and metaconule are somewhat better developed. The parastyle is completely fused with the preparacrista. Neither a para− nor a metacingulum is developed. On the M 3 there are no lingual conules and no paracingulum. There are no labial and lingual cingula in the upper molars. Humerus. — No complete humerus is preserved, but 12 distal fragments and one proximal from Petersbuch 10. The overall morphology and slenderness indicates a moderate stage of fossorial adaptation. The proximal epiphysis is wider than the distal one. The head is directed parallel to the shaft. The brachialis fossa is large but not very deep. The teres tubercle is moderately long and situated close to the pectoral crest. The pectoral tubercle is situated in mid−shaft position. The deltoid process is short. A prominent scalopine ridge runs from the head to the medial side of the lesser tubercle and separates two areas in different planes. The area delimited by pectoral crest, pectoral ridge and greater tubercle is slightly concave. The notch between head and lesser tuberosity is well defined. On the distal epiphysis there is a large olecranon fossa and a somewhat smaller supratrochlear fossa. The trochlea is broad, thus leaving only a narrow notch between trochlea and the fossa for the m. flexor digitorum profundus ligament. Ulna. — Only the proximal part is preserved. The abductor fossa is deeply excavated. The proximal crest forms a large blade widely separated from the semilunar notch. A prominent processus anconaeus and smaller but distinct coronoid process delimit the well−defined semilunar notch. Comparisons Leptoscaptor shows clear scalopine affinities as defined by Hutchison (1968: 58): the enlarged i 2, not enlarged p 1 and upper canine, moderately to very broad humerus with a moderately deep brachialis fossa. The allocation with any other talpine tribe can be excluded with certainty. Consequently, with few exceptions, we can restrict our comparison to scalopine genera. The only Recent Old World member of the Scalopini as defined by Hutchison (1968) is Scapanulus oweni Thomas, 1912, the Kansu mole, which lives in parts of China. It corresponds to Leptoscaptor in dental formula, the number of roots in the P 1 – P 3 and in the divided mesostyles of the upper molars. However, Leptoscaptor differs from this species in: – the absence of a metastylid on m 2 and m 3, – the trigonid of the m 1, which is not compressed antero−posteriorly, – the double−rooted p 2, – the small but present parastyle on P 4, – the pectoral tubercle situated more in the midline of the shaft, – the greater tubercle and head of the humerus not being twisted medially. No specimen has been seen. The Scapanulus oweni criteria have been concluded from Storch and Qiu (1983: 119) and from Hutchison (1968: figs. 10 D, 11).	en	Ziegler, Reinhard (2003): Moles (Talpidae) from the late Middle Miocene of South Germany. Acta Palaeontologica Polonica 48 (4): 617-648, DOI: 10.5281/zenodo.13396039
480C879940007616DD28D5BCFC5CFDC8.taxon	description	– the well−developed metastylids on m 2 and m 3, – the more lingual termination of the oblique cristid on m 1, – the better−developed cingulids, – the more robust humerus, which indicates a better fossorial adaptation.	en	Ziegler, Reinhard (2003): Moles (Talpidae) from the late Middle Miocene of South Germany. Acta Palaeontologica Polonica 48 (4): 617-648, DOI: 10.5281/zenodo.13396039
480C879940007616DD28D5BCFC5CFDC8.taxon	description	– the absence of a metacristid on m 2, – the undivided mesostyle on M 2 and M 3, – the prominent para− and metaconule on M 2.	en	Ziegler, Reinhard (2003): Moles (Talpidae) from the late Middle Miocene of South Germany. Acta Palaeontologica Polonica 48 (4): 617-648, DOI: 10.5281/zenodo.13396039
480C879940007616DD28D5BCFC5CFDC8.taxon	description	– the well−developed metastylid on m 2 and m 3, – the weakly divided mesostyle on the upper molars, – the better−developed metaconule and metacingulum on M 1.	en	Ziegler, Reinhard (2003): Moles (Talpidae) from the late Middle Miocene of South Germany. Acta Palaeontologica Polonica 48 (4): 617-648, DOI: 10.5281/zenodo.13396039
480C879940007616DD28D5BCFC5CFDC8.taxon	description	– a not hypertrophied i 2, – lower molars with antero−posteriorly more compressed trigonids.	en	Ziegler, Reinhard (2003): Moles (Talpidae) from the late Middle Miocene of South Germany. Acta Palaeontologica Polonica 48 (4): 617-648, DOI: 10.5281/zenodo.13396039
480C879940007616DD28D5BCFC5CFDC8.taxon	description	– in the presence of three upper and lower incisors, – in the single−rooted p 2, – the presence of a small metaconid on p 4, – the better developed cingula on all teeth, – the oblique cristid terminating more buccally on m 2 and m 3, – the undivided mesostyle on the upper molars.	en	Ziegler, Reinhard (2003): Moles (Talpidae) from the late Middle Miocene of South Germany. Acta Palaeontologica Polonica 48 (4): 617-648, DOI: 10.5281/zenodo.13396039
480C879940007616DD28D5BCFC5CFDC8.taxon	description	– the complete set of three lower incisors with an enlarged i 1, – the single−rooted p 2, – the higher−crowned lower molars, – the undivided mesostyles on the upper molars, – the head of the humerus, which is directed medio−distally.	en	Ziegler, Reinhard (2003): Moles (Talpidae) from the late Middle Miocene of South Germany. Acta Palaeontologica Polonica 48 (4): 617-648, DOI: 10.5281/zenodo.13396039
480C879940007616DD28D5BCFC5CFDC8.taxon	description	– the oblique cristid joining the metacristid. Van den Hoek Ostende (2001) described the new talpid subfamily Suleimaninae with the only species Suleimania ruemkae Van den Hoek Ostende, 2001 from the Lower Miocene localities Harami, Kilçak, and Keseköy in Anatolia. This species is mainly known from isolated teeth. This large−sized species is distinctive by the loss of the M 3 and the loss of the talonid in the m 3, a character known from the erinaceines and the dimylid Exoedaenodus. Furthermore, this species differs from Leptoscaptor in: – the sharp cutting edges of the premolars, – the inflated cusps of the m 1 and M 1, – the presence of a well−developed hypocone in the upper molars. Hugueneya Van den Hoek Ostende, 1989 is a monospecific species from the Early Miocene of South Germany. The only species H. primitiva (Hutchison, 1974), in spite of being also a scalopine, cannot be confused with Leptoscaptor. Hugueneya differs from Leptoscaptor in (cf. Hutchison 1974: fig. 21, pl. 39): – the presence of four lower premolars, – the more inflated teeth, – the more prominent metaconule and the more spaced mesostyles of the upper molars, – the distinctly more robust humerus. Discussion Along with Talpa minuta, Leptoscaptor bavaricum represents the most common talpid in the samples of Petersbuch 6, 10, and 18. Petersbuch 10 yielded the most numerous sample of this species and the only one with postcranial bones. Therefore it was chosen as type locality though the dentaries are better preserved in Petersbuch 6 and 18. The association of lower and upper dentition and of the humerus fragments to the dental remains is without alternative and is certainly correct. The ulna fragments match the humerus fragments in size. The only noticeable difference between the three samples is the position of the mental foramen. It is more variable in the smaller samples than in Petersbuch 10. In the size of the teeth there are no significant differences between the three samples even though some specimens of Petersbuch 18 are slightly smaller (P 3) or larger (p 2) than the corresponding teeth of the Petersbuch 10 sample. In the talpid samples of Petersbuch 6 and 18 there are no postcranial remains left that can be associated with the teeth of Leptoscaptor. There is another species of Leptoscaptor with a somewhat more robust humerus, L. robustior. This species can unambiguously be identified in its type locality Petersbuch 35. As the fissures from Petersbuch 6, 10, and 18 are directly adjacent to one another, they probably all belong to one fissure system and all three fissure fills may result from the same filling process. Consequently, the three samples possibly represent only one population instead of three different ones. This spatial viewpoint argues in favour of an affiliation between the Petersbuch 6 and 18 samples with L. bavaricum. However, the position of the mental foramen is more variable in Petersbuch 6 and especially in Petersbuch 18, as it is characteristic of the Petersbuch 35 sample, which undoubtedly represents L. robustior. Therefore, we cannot exclude with certainty that either Petersbuch 6 or Petersbuch 18 or even both samples represent L. robustior. The fact that two species only can unambiguously be identified by their humeri is not unique to the genus Leptoscaptor. Regarding Paratalpa, an Oligocene to Agenian genus, and Desmanodon, which appeared in Europe in the Orleanian, there are even two different genera that are only distinguishable by their humeri (see discussion in van den Hoek Ostende 1989, Ziegler 1990). However, in talpids the humeri usually allow discrimination to the level of the tribe and the dentition is more distinctive. Regarding the loss of two lower antemolars, Leptoscaptor is more advanced than the majority of the scalopine genera. Only the living Scalopus from North America has more reduced dentition with single−rooted premolars. For Scapanulus the data are somewhat contradictory. Storch and Qiu (1983: 119) mention the loss of two upper and lower antemolars of questionable homologies. Consequently, the dental formula could be as in Leptoscaptor. Hutchison (1968: 74), in contrast, mentions a complete lower dentition. Gerhard Storch told me that his antemolar count is correct (personal communication, 18 th December 2002). Ziegler (1971: 59) gives the same conclusion as Storch and Qiu, referring to the original description by Thomas (1912: 397). Obviously, Hutchison's antemolar count is erroneous. Concerning the number of roots in the lower premolars, Leptoscaptor is less advanced than all living scalopines. As Scalopus is too specialised in other characteristics and as the other extant species have more complete dentitions, Leptoscaptor cannot be ancestral to any extant genus. Obviously it is a Miocene offshoot that became extinct somewhat later.	en	Ziegler, Reinhard (2003): Moles (Talpidae) from the late Middle Miocene of South Germany. Acta Palaeontologica Polonica 48 (4): 617-648, DOI: 10.5281/zenodo.13396039
480C8799400B7617DD28D7DCFB44FA1D.taxon	description	Fig. 3. Etymology: From Latin robustior, more robust. The humerus is more robust than in the type species L. bavaricum. Holotype: Right humerus, NHMA P 35 − 58 / 6, fig. 3 A. Measurementsoftheholotype: GL (10.5), Bp (6.35), BpwT (5.70); SD (2.56), BdwE (5.14), Bp * 100 / GL (60.5). Type locality: Petersbuch 35 (details see p. 618). Age: Uppermost part of the Middle Miocene (MN 8 according to Rummel 2000, means MN 7 + 8.) Paratypes (measurements see Tables 1, 2). — Petersbuch 35: NHMA P 35 − 57 A 1, right dentary fragment with teeth; CRW P 35 − 57 + 58, 4 dentary fragments with teeth, left maxilla fragment with p 4, 7 isolated teeth, 11 humerus fragments, right ulna fragments. Referred material (measurements see Tables 1, 2). — Petersbuch 48: CRW P 48 − 93 – 94, left dentary fragment with p 3 – p 4, right maxilla fragment with M 2, 3 isolated teeth, 3 humerus fragments. Diagnosis. — Medium−sized species of Leptoscaptor with postcranial elements more robust than in the type species and with two mental foramina on the dentary. Description of the holotype Pectoral crest, deltoid process and the epicondylar spines are broken. The long axis of the elliptical head is directed parallel to the shaft. The marked scalopine ridged separates two areas in different planes. The brachialis fossa is moderately deep. The anterior aspect shows the pectoral process in midshaft position and a concave area delimited by pectoral crest, pectoral ridge, and greater tubercle. Above the distal epiphysis there is a wide olecranon fossa and a small supratrochlear fossa. The broad trochlea only leaves a narrow notch, separating trochlea and the fossa for the m. flexor digitorum profundus ligament. Description of paratypes and referred material Dentary. — There are five fragments of the horizontal ramus from Petersbuch 35, and one from Petersbuch 48. The consistent presence of two mental foramina is characteristic: below the anterior roots of p 3 and p 4 (once), beneath the posterior root of p 2 and between the roots of p 4 (twice), below the anterior root of p 3 and under the posterior root of p 4 (once) in the Petersbuch 35 sample. In the Petersbuch 48 specimen one mental foramen is situated beneath the anterior root of p 3, another below the anterior root of p 4. In two specimens the alveoles of the double−rooted p 2 – p 3 and the single−rooted canine are preserved. The Petersbuch 48 dentary also preserves the canine alveolus and the two incisor alveoli, indicating that i 2 was larger than i 1. Teeth. — In the morphology of the preserved teeth there is no difference to those of L. bavaricum. Postcranialbones. — Aside from the type specimen there are 11 additional humerus fragments from Petersbuch 35 and three from Petersbuch 48, which correspond in robustness. The ulna fragment from Petersbuch 35 was referred because of matching size. It has a deep abductor fossa. Comparisons L. robustior is quite similar to the type species of the genus L. bavaricum. Only this species can be confused and needs to be differentiated. L. robustior differs in: – the consistent presence of two mental foramina, which are situated slightly more posterior, – a distinctly more robust humerus. Discussion In the small talpid sample from Petersbuch 35 there are only two species: Proscapanus sansaniensis and Leptoscaptor robustior. As both species differ markedly in size and robustness of the humerus they are easily distinguishable. Assuming that there are not two different species, one being represented by teeth only and the other exclusively by postcranial elements, the association of postcranial elements and dental remains in Leptoscaptor robustior is without alternative. In the Petersbuch 48 talpid fauna there are only three species, which also can be readily distinguished: P. sansaniensis, Talpa minuta and L. robustior. The difference in robustness of the nearly complete humeri from Petersbuch 35 and 48 lies well within the range of a population. The more robust humerus of L. robustior indicates a better fossorial adaptation than in L. bavaricum. Without humeri the Leptoscaptor species are hardly distinguishable. Nevertheless, I am convinced that they represent different biological adaptations (see also chapter discussion of L. bavaricus).	en	Ziegler, Reinhard (2003): Moles (Talpidae) from the late Middle Miocene of South Germany. Acta Palaeontologica Polonica 48 (4): 617-648, DOI: 10.5281/zenodo.13396039
480C8799400A7617DD28D0B0FAB9F98C.taxon	materials_examined	Type species: Proscapanus sansaniensis (Lartet, 1851).	en	Ziegler, Reinhard (2003): Moles (Talpidae) from the late Middle Miocene of South Germany. Acta Palaeontologica Polonica 48 (4): 617-648, DOI: 10.5281/zenodo.13396039
480C8799400A7612DD28D36FFC4EF843.taxon	description	Fig. 4. Material (measurements see Tables 3, 4). — Petersbuch 10: CRW P 10 − 620 – 621, left dentary fragment with p 1 – p 3, right humerus. Petersbuch 31: NHMA P 31 − 163 A 1, 163 C 1, 163 E 3, 162 B 2, 164 A 1, 2 dentary fragments with teeth, 2 upper teeth, left humerus; CRW P 31 − 163, 164, 4 dentary fragments with teeth, 2 maxilla fragments with teeth, 8 isolated teeth, 6 humeri, 5 ulnae, 3 radii. Petersbuch 35: CRW P 35 − 55, 56, 5 isolated teeth, left humerus, left ulna fragment, right radius. Petersbuch 48: NHMA P 48 − 89 A 1 + B 2, right dentary fragment with teeth, left M 1; CRW P 48 − 89, 90, right maxilla fragment with P 4, 4 isolated teeth, 4 humeri. Description Dentary — Only some more or less complete fragments of the horizontal ramus are preserved. The jaw slightly tapers anteriorly. The dentary fragment from Petersbuch 10 shows mental foramina below p 4 / m 1 and beneath p 2. In two specimens from Petersbuch 31 the anterior mental foramen is situated between the roots of p 2 and p 3, the posterior one under the trigonid of m 1. In the dentary fragment from Petersbuch 48 there are three mental foramina, one situated between the roots of p 1 and p 2, and one below the anterior and posterior root of p 4 each. All teeth anterior to p 4 are single−rooted. According to their alveoles the three incisors are increasingly inclined anteriorly, i 2 being larger than i 1 and i 3. The canine is only slightly inclined anteriorly. Lower dentition. — p 1 to p 3 are slightly inflated and increase in size. There is an incipient anterior crest extending toward the apex of the protoconid. A faint posterior basal cuspule is developed. The p 4 is rectangular in occlusal outline and has an antero−buccal and posterior cingulid, the latter culminating in a postero−lingual basal cuspule. The sizes of the molars are ranked in the following order, m 2> m 1> m 3. The oblique cristid extends lingually to join the metacristid. In the m 1 the paralophid is curved and leaves a long trigonid, whereas it is very short in m 2 and m 3. There is a marked precingulid in m 2 and m 3. In the Petersbuch 31 sample the pre− and ectocingulid of m 1 are indistinct, whereas they are better developed in the Petersbuch 48 dentary. In the m 1 the talonid is wider than the trigonid, in the m 2 the trigonid is somewhat wider and in the m 3 distinctly wider than the talonid. Maxilla. — Two fragments from the Petersbuch 31 sample show the infraorbital foramen above the mesostyle of M 1, the anterior opening of the infraorbital canal above the posterior root of M 2 and the origin of the zygomatic arch above M 3. Upperdentition. — Only P 4 to M 3 are preserved. The P 4 has an indistinct, hardly projecting parastyle. On the lingual talon there is no vestige of a protocone, but a lingual cingulum. The molars have four roots, the posterior one being the strongest and the central one the weakest. The M 1 has a deeply divided mesostyle, a slightly projecting parastyle and indistinct para− and metaconule. The marked paracingulum joins the parastyle; the metacingulum tapers but extends to the postero−labial corner. The M 2 also has a divided mesostyle. In some specimens the lingual conules are somewhat better developed than in the M 1. Para− and metacingulum are either very thin or even absent. In the M 3 the mesostyle is only superficially divided. There are only three M 3 from the Petersbuch 31 sample, which are assumed to belong with Proscapanus sansaniensis because of their size. One has a marked paracingulum, in two M 3 it is absent. Postcranial bones — The humerus strongly resembles the specimens from Sansan in all morphological details and in gracility. The brachialis fossa is deeply excavated, the teres tubercle long, the supratrochlear fossa small, and the scalopine ridge marked and shelf−like. In the ulna the most conspicuous character is the deep abductor fossa, which extends on the proximal moiety of the lateral side. The proximal crest delimits the large area of insertion for the triceps. The radius has a capitular process projecting proximally. The distal joint is characterised by the large scaphoid articular facet. Discussion	en	Ziegler, Reinhard (2003): Moles (Talpidae) from the late Middle Miocene of South Germany. Acta Palaeontologica Polonica 48 (4): 617-648, DOI: 10.5281/zenodo.13396039
480C8799400D7610DD28D794FB12FCA8.taxon	materials_examined	Type species: Talpa europaea Linnaeus, 1758.	en	Ziegler, Reinhard (2003): Moles (Talpidae) from the late Middle Miocene of South Germany. Acta Palaeontologica Polonica 48 (4): 617-648, DOI: 10.5281/zenodo.13396039
480C8799400D760FDD28D641FBFFFC23.taxon	description	Fig. 7. Material (measurements see Tables 7, 8). — Petersbuch 6: NHMA P 6 − 1058 B 1, 1059 / 4, 6, 1060 / 1, 1062 / 2, 3, right dentary and right maxilla fragment with teeth, right P 4, left humerus, 2 left ulnae; CRW P 6 − 1058 – 1062, Petersbuch 6, 19 dentary fragments with teeth, 8 maxilla fragments with teeth, 18 humeri, 4 ulnae. Petersbuch 10: CRW P 10 − 0601 – 607, 622 − 62, 6 dentary fragments with teeth, 3 maxilla fragments with teeth, 5 isolated teeth, 26 humeri, 13 ulnae. Petersbuch 18: CRW P 18 − 0749 – 751, 6 dentary fragments with teeth, 3 maxilla fragments with teeth, 5 isolated teeth, 2 humeri. Petersbuch 31: CRW P 31 − 0159 – 162, 3 dentary fragments with teeth, 2 maxilla fragments with teeth, 33 isolated teeth, 43 humeri, 8 ulnae, 3 radii. Petersbuch 48: CRW P 48 − 0091, 92, 7 isolated teeth, 6 humeri. Description Dentary. — In general shape the dentary is similar to that of the extant Talpaeuropaea. The complete ascending ramus is only preserved in the Petersbuch 6 sample. It forms a nearly right angle with the horizontal ramus. The mandibular foramen is situated on the ventral margin of the mylohyoid ridge, roughly in the centre of the ascending ramus. The masseteric fossa is deeply excavated, the pterygoid fossa only moderately deep. The angular process is shovel−shaped with an internal concavity. The cylindrical condylus is situated high above the level of the tooth row. The coronoid process itself is rectangular. In the Petersbuch 6 sample the posterior mental foramen is located either under the anterior root of m 1 (twice) or between its roots (four times), the anterior mental foramen below the posterior root of p 2 (twice) or between p 2 and p 2 (twice) or under the anterior root of p 3 (once). The incisor alveoli are not preserved. Lower dentition. — The canine is incisor−shaped and slightly procumbent. All premolars are double−rooted, p 2 and p 3 overlap one another. p 1 is distinctly larger than p 2 and p 3, but all are similar in shape. The cusp is centred over the anterior root and origin of a marked posterior crest, joining a posterior cuspule, and a less well−developed anterior crest. The buccal face is convex, the lingual side flat. A faint cingulid is restricted to the posterior part of the crown base. The p 4 has a talonid and a trilateral crown with convex buccal and lingual faces and a concave posterior side. The postero−lingual crest joins the posterior cuspule. An anterior cuspule is also developed. The size relation between the molars is m 1 <or> m 3 <m 2. Buccal cingulids are rudimentary and confined to the hypoflexid. The trigonid is lingual open. In the m 1 the talonid is wider than the trigonid, the oblique cristid joins the middle of the protocristid or ends slightly labial to the middle. The most conspicuous feature is the faint ascending precingulid, which joins the paracristid. The protoconid of the m 2 is the highest cusp in the tooth row. The trigonid is somewhat wider than the talonid. There is a marked precingulid and a tiny cuspule in front of the entoconid. The m 3 is similar to a small m 2 without entostylid and with a narrower talonid. Maxilla. — Larger fragments of maxillae are preserved only in the Petersbuch 6 sample. The origin of the zygomatic arch lies above M 3. The infraorbital foramen opens high above the middle of M 1. The infraorbital canal runs in a deep groove. Its anterior and posterior opening are separated by a narrow bony bridge above the anterior root of M 3. The same configuration is found in extant Talpa. Upper dentition. — According to the alveoles there are three single−rooted incisors. Only two chisel−shaped I 2 and I 3 are preserved. The double−rooted canine has a mesio−lingual groove and a distal crest. P 1 – P 3 are double−rooted, P 2 is smaller than both the others. No P 1 is preserved. P 2 and P 3 have an oval outline in occlusal view. There is a distal crest. A faint disto−buccal cingulum may be developed. The P 4 has a more or less projecting parastyle and a conical protocone. A straight distal crest originates from the paracone. The upper molars have an undivided mesostyle and neither paranor metaconule. In the M 1 the marked paracingulum joins the projecting parastyle. The metacingulum is interrupted along the postmetacrista or reduced to a short spur above the metastyle. There are four roots, the strongest above the protocone, a mesio−distally compressed one above paracone and postmetacrista respectively, and a small round one on both the labial roots. M 2 and M 3 have three roots and neither para− nor metacingulum. Postcranial bones. — Humerus, ulna, and radius are smaller than in the extant Talpa, but do not show any morphological differences. Therefore we can abstain from a formal description. Even in the gracility index, defined as Bp * 100 / Gl, they correspond to T. europaea. This means that both have similar fossorial abilities. However, the extant Talpa europaea is distinctly larger. Discussion With the exception of Petersbuch 35, which yielded only two talpid species, Talpaminuta is represented in all samples and it is the most common talpid in all samples. There are no morphological differences among the specimens under study and Talpa minuta from the type locality Sansan. The position of the anterior mental foramen from this locality is as variable as in the samples under study. In the Sansan sample the posterior mental foramen is consistently situated under the anterior root of m 1 whereas it slightly varies in position in the Petersbuch samples. However, there is a distinct difference in size, the dentition and postcranial bones from Sansan being smaller than in Petersbuch. It is assumed that the size difference lies within the variability of a species and that the delineation of a new species or subspecies is not justified. Talpa minuta is one of the most common and most wide spread talpid species in the Miocene of Europe. It is recorded from faunas correlated with MN 3, e. g., Wintershof−West, to MN 9, for example, Nebelbergweg (Ziegler 1994; Kälin and Engesser 2001), from Slovakia in the East to Southeast France in the West.	en	Ziegler, Reinhard (2003): Moles (Talpidae) from the late Middle Miocene of South Germany. Acta Palaeontologica Polonica 48 (4): 617-648, DOI: 10.5281/zenodo.13396039
480C87994012760CDD28D687FC65FD08.taxon	materials_examined	Type species: Urotrichus talpoides Temminck, 1841.	en	Ziegler, Reinhard (2003): Moles (Talpidae) from the late Middle Miocene of South Germany. Acta Palaeontologica Polonica 48 (4): 617-648, DOI: 10.5281/zenodo.13396039
480C87994012760CDD28D687FC65FD08.taxon	materials_examined	Material and measurements. — Petersbuch 6: NHMA P 6 − 1066, left dentary fragment with p 4 – m 1 (trigonid), p 4 (1.05 × 0.67), m 1 Wa (1.03); NHMA P 6 − 1067, right humerus, GL (7.60), Bp (3.64), BpwT (3.00), DS (1.30), Bd (3.86), BdwE (3.32), Bpx 100 / GL (48.0). Description Dentary. — There is only a short fragment of the horizontal ramus with two small mental foramina beneath the trigonid of m 1 and a bigger one under the second alveolus anterior to p 4. The third alveolus anterior to p 4 is slightly inclined. This means that the fracture is near the real tip of the dentary and that the antemolar part is reduced. The cusp of the p 4 is cone−shaped with a more or less flat lingual face and a postero−lingual crest. Close to the base of this crest there is a small accessory cuspule. The p 4 is slightly heeled and has a precingulid and a postcingulid respectively. The m 1 protoconid is somewhat higher than the p 4. The paracristid is angular, the protocristid notched. The oblique cristid extends to below the protocristid notch and does not join the weak metacristid. There is a well−developed precingulid. The talonid is broken off.	en	Ziegler, Reinhard (2003): Moles (Talpidae) from the late Middle Miocene of South Germany. Acta Palaeontologica Polonica 48 (4): 617-648, DOI: 10.5281/zenodo.13396039
480C87994012760CDD28D687FC65FD08.taxon	description	Humerus. — The humerus is nearly complete. Only the epicondylar spines and the deltoid process are broken. It is slender in overall shape, which means only modest fossorial specialisation. Posteriorly the olecranon fossa is extensive, but shallow. The long axis of the head points latero−distally. The ridge running from the lesser tubercle to beneath the head is weak and curved. The brachialis fossa is moderately deep. Anteriorly the pectoral tubercle terminates far laterally about half way down the shaft. Teres tubercle and lesser tubercle are separated by a wide sulcus, head and greater tubercle by a deep and narrow notch. The supratrochlear fossa is wide and deeply pocketed. There is a deep notch between the trochlea and the fossa for the m. flexor digitorum profundus ligament. Discussion Dentary and humerus have been lumped together, because they match in overall size, both represent urotrichines and both are the only talpid leavings in the Petersbuch 6 talpid fauna. It is considered more probable that dentary and humerus represent only one species instead of two. The humerus shows undeniable urotrichine affinities and fits well with? Urotrichus dolichochir from La Grive in all morphological details and in gracility. The small size difference is not worth mentioning. The Petersbuch 6 specimen also resembles the humeri of? Urotrichus cf. dolichochir from Sandelzhausen (cf. Ziegler 2000: 81). The type humerus from La Grive was named Scaptonyx? dolichochir by Gaillard (1899: 30) even though he noted the resemblance to the extant genus Urotrichus. As this species is not referable to Scaptonyx and in the absence of associated dental material Hutchison (1974: 226) referred the species tentatively to Urotrichus on the basis of the humeral morphology. He also considered the lectotype dentary of “ Scaptonyx ” edwardsi from the same site a possible candidate for the association with? Urotrichus dolichochir. However, Hutchison (1974: 228) also wrote that the dentary may belong with the humerus of cf. Scalopoides sp. from La Grive. In fact, we do not know the dentition of? Urotrichus dolichochir. The dentary fragment under study differs from Scaptonyx edwardsi in the duplication of the posterior mental foramen, in the presence of a postero−lingual cuspule on p 4 and of a weak metacristid in m 1. In order to resolve the problem whether any dentary belongs to any humerus, much more material is needed. In the present study both humerus and dentary are reservedly referred to? Urotrichus dolichochir. They represent the second record of this species in Germany.	en	Ziegler, Reinhard (2003): Moles (Talpidae) from the late Middle Miocene of South Germany. Acta Palaeontologica Polonica 48 (4): 617-648, DOI: 10.5281/zenodo.13396039
480C87994011760CDD28D793FB07FCAD.taxon	materials_examined	Type species: Myxomygale antiqua Filhol, 1890.	en	Ziegler, Reinhard (2003): Moles (Talpidae) from the late Middle Miocene of South Germany. Acta Palaeontologica Polonica 48 (4): 617-648, DOI: 10.5281/zenodo.13396039
480C87994011760ADD28D641FE81F966.taxon	description	Fig. 9. Etymology: From Latin gracilis, slender, lank; becauseof thegracilehumerus. Holotype: Right humerus, P 10 − 615 / 2, fig. 9 D. Measurementsoftheholotype: GL (8.05), Bp (3.50), BpwT (3.15), DS (1.30), Bd (3.92), BdwE (3.46), Bp * 100 / GL (43.5). Type locality: Petersbuch 10 (details see p. 618). Age: UppermostpartoftheMiddleMiocene (MN 7 / 8, Rummel 2000). Paratypes and measurements. — Petersbuch 10: CRW P 10 − 614 / 1, left dentary fragment with m 1 – m 2; m 1 (1.47 × 0.89 × 1.07); m 2 (1.69 × 1.09 × –); NHMA P 10 − 614 / 2, right dentary fragment with m 2 – m 3; m 2 (1.64 × 1.15 × 1.04); m 3 (1.35 × 0.88 × 0.6); NHMA P 10 − 614 / 3, right dentary fragment with m 2 – m 3; m 2 (1.65 × 1.09 × 0.97), m 3 (1.37 × 0.84 × 0.60); NHMA P 10 − 614 / 4, left maxilla fragment with M 1, M 1 (2.10 × 1.84); CRW P 10 − 614 / 5, right M 1 (2.03 × 1.77); CRW P 10 − 615.1, left humerus fragment, DS (1.44), Bd (ca. 4.1), BdwE (3.40). Diagnosis. — Small−sized Myxomygale characterised a mental foramen situated under the protoconid of m 1, an oblique cristid, extending labially in m 1, joining a marked metacristid in m 2 and m 3. M 1 without parastyle and preparacrista, with deeply divided mesostyle. The humerus is urotrichine and slender with long ledge−like teres tubercle and a pectoral tubercle situated laterally. Description of the holotype Only the deltoid process and the epicondylar spines are broken off. In anterior view the humerus shows a long teres tubercle with a proximal end hidden by the marked pectoral ridge. The pointed pectoral tubercle extends further distally than the teres tubercle and is situated on the lateral margin of the shaft. It is visible in posterior view. The brachialis fossa is moderately deep. The sulcus between head and major tubercle is a narrow groove. There is a large, pocketed supratrochlear fossa, between trochlea and the fossa m. flexor digitorum profundus ligament a deep, concave notch. The posterior face shows the shallow olecranon fossa and the head with a long axis slightly directed disto−laterally. The ridge between head and lesser tubercle is short and tapers towards the lesser tubercle. Description of the paratypes Dentary. — Among the 4 short fragments of the horizontal ramus two show the posterior mental foramen beneath the trigonid of m 1. The specimens yield no information concerning the antemolar dentition. Lower molars. — The size relation is m 2> m 1> m 3. In the m 1 the oblique cristid runs rather bucally, extending to the posterior base of the protoconid. There is no metacristid. The precingulid does not extend to below the paraconid but is continuous with the ectocingulid. There is only a weak postcingulid hidden by the marked precingulid of the m 2. The entostylid is broken off. In the m 2 the protoconid is more elevated than in the m 1 and the trigonid is distinctly narrower. The oblique cristid joins the marked metacristid. The precingulid is more marked than in the m 1. The m 3 differs from the m 2 in the smaller size and in the reduced talonid without entostylid. Maxilla. — One fragment with M 1 shows the lacrimal foramen above the anterior root of M 1. M 1. — There are neither preparacrista nor parastyle. The mesostyle is deeply divided. Para− and metaconule are only moderately differentiated. The preparaconuluscrista is continuous with the precingulum, which extends to the buccal margin. The postmetaconuluscrista joins the metacingulum which itself tapers in its mid part and ends in a marked short crest above the metastyle. Humerus. — In addition to the type specimen there is one humerus with the proximal third and the point of the pectoral tubercle broken off. It yields no further information. Comparisons There are five species of Myxomygale known thus far most of which are only scarcely represented. They are listed in ascending order with respect to their stratigraphic range: the type species Myxomygale antiqua Filhol, 1890 from the Quercy (Oligocene), Myxomygale vauclusensis Crochet, 1995 from Saint−Martin−de−Castillon (Oligocene, MP 23, figures and measurements in Hugueney 1972: 53 f), Myxomygaleminor Ziegler, 1990 from Ulm−Westtangente (Lower Miocene, MN 2 a), Myxomygale hutchisoni (Ziegler, 1985) from Petersbuch 2 (Lower Miocene, MN 4), Myxomygale engesseri Doukas, 1986 from Aliveri in Greece (Lower Miocene, MN 4). The humerus is known, i. e. published, described and figured, only from M. hutchisoni (Ziegler 1985: fig. 8, tab. 3). Crochet (1995: 56) also mentions the humerus in the revised genus diagnosis of Myxomygale. However, it is not known to which species he refers. As much as is known from published evidence, the humerus is only preserved in M. hutchisoni. Perhaps Crochet knows unpublished humeri from M. antiqua. M. gracilis differs from M. antiqua in: – the distinctly smaller size, – the size relation between m 1 and m 2 (m 1 <m 2) and the weaker ectocingulids. M. gracilis differs from M. vauclusensis in: – the distinctly smaller size, – the more anterior position of the mental foramen, – the absence of a parastyle, the divided mesostyle and the less developed para− and metaconule of M 1. M. gracilis differs from M. minor in: – the distinctly wider M 1, – the absence of a parastyle and the divided mesostyle in M 1, M. gracilis differs from M. hutchisoni in: – the absence of a parastyle and the divided mesostyle in M 1, – the smaller and in particular more slender humerus, with a relatively longer teres tubercle and a less marked ridge running from the head to the lesser tubercle. M. gracilis differs from M. engesseri in: – the wider m 2, – the somewhat bigger M 1 without parastyle and with continuous preparaconuluscrista and paracingulum respectively and deeply divided mesostyle.? Urotrichusdolichochir (Gaillard, 1899) known from the type locality La Grive (Middle Miocene, MN 7 / 8) by the humerus only, and from Petersbuch 6, is rather similar, thus deserves to be mentioned. M. gracilis differs from? U. dolichochir in: – the somewhat bigger humerus, – the longer teres tubercle hidden proximally by the pectoral ridge. In the Petersbuch 10 fauna there is another small urotrichine, which is not determinable beyond the tribe. It is represented by a humerus and tentatively associated dentition. M. gracilis differs from Urotrichini gen. et sp. indet. I in the: – smaller overall size, – more posterior position of the mental foramen, – m 1 without metacristid and an oblique cristid running more buccally, – more gracile humerus with the longer teres tubercle, – lateral position of the pectoral tubercle, – less marked ridge running from the head and tapering towards the lesser tubercle. The Petersbuch 31 fauna yielded a small urotrichine, represented by humeri and dentition. M. gracilis differs from Urotrichini gen. et sp. indet. II in the: – less pronounced para− and metaconule, the absence of the parastyle and the divided mesostyle in M 1, – more gracile humerus with the longer teres tubercle and a weaker and discontinuous ridge between head and lesser tubercle. Discussion	en	Ziegler, Reinhard (2003): Moles (Talpidae) from the late Middle Miocene of South Germany. Acta Palaeontologica Polonica 48 (4): 617-648, DOI: 10.5281/zenodo.13396039
480C87994017760ADE62D3AFFC34FB24.taxon	materials_examined	Type species: Tenuibrachiatum storchi gen. et sp. nov. Etymology: From Latin tenuis, tenuous; brachium, arm; tenuibrachiatum, with tenuous arms; because of the slender humerus. Included species: Type species only. Diagnosis. — Small urotrichine mole. Tentative mandibular dental formula 2 i, 1 c, 4 p, 3 m. Incisors procumbent, i 1> i 2. All lower premolars double−rooted. m 1 <m 2> m 3. Oblique cristid terminates bucally in m 1, joins the metacristid in m 2 and m 3. P 4 with tiny parastyle and marked protocone. M 1 and M 2 with undivided mesostyle. Para− and metaconule weakly developed in M 1, well differentiated in M 2. Humerus typical urotrichine, slender, long teres tubercle, pectoral tubercle situated laterally, large, pocketed supratrochlear fossa. In anterior aspect the trochlea is broadening towards the capitulum.	en	Ziegler, Reinhard (2003): Moles (Talpidae) from the late Middle Miocene of South Germany. Acta Palaeontologica Polonica 48 (4): 617-648, DOI: 10.5281/zenodo.13396039
480C879940177609DD28D1E5FB5BFB88.taxon	description	Fig. 10. Etymology: In honour of Dr. Gerhard Storch, Frankfurt, thus appreciating his outstanding contributions to our knowledge of the Tertiary small mammals. Holotype: Left dentary fragment with p 1, p 4 – m 2 and the alveoles of i 1, i 2, c, p 2 – p 3; NHMA P 31 − 166 A 1, Fig. 10 A. Measurements of the holotype: p 1 (0.70 × 0.43), p 4 (0.97 × 0.58), m 1 (1.40 × 0.88 × 1.00), m 2 (1.54 × 1.00 × 0.95), lingual height of the dentary below m 1 (1.35). Type locality: Petersbuch 31 (details see p. 618).	en	Ziegler, Reinhard (2003): Moles (Talpidae) from the late Middle Miocene of South Germany. Acta Palaeontologica Polonica 48 (4): 617-648, DOI: 10.5281/zenodo.13396039
480C879940177609DD28D1E5FB5BFB88.taxon	diagnosis	Diagnosis. — As for the genus. Description of the holotype The horizontal ramus from the first incisor alveolus to the fracture behind m 2 is preserved. The dentary was broken between p 4 and m 1 and was glued. It tapers anteriorly. The anterior mental foramen is situated under the anterior root of p 2. The posterior foramen beneath the trigonid of m 1 is filled with not removable sediment, thus hardly visible. The four alveoli of p 2 and p 3 are slightly overlapping. There are three alveoli anterior to p 1: the first two nearly procumbent, the third slightly inclined anteriorly. They are interpreted as alveoli of i 1, i 2, and c, resulting in the tooth formula 2143. Alternatively they can be interpreted as i 2, i 3, and c, resulting in the same tooth formula, or as i 1, i 2, and i 3. In the latter case the tooth formula was 3043. According to their alveoli the size relation is i 1> i 2> c. The symphysis extends to c / p 1. The double−rooted p 1 is oval in occlusal outline. Its cusp is situated above the anterior root. There is a weak posterior and lingual cingulid, respectively. The p 4 has a postero−lingual and an anterior crest. It is surrounded by a weak cingulum. There is a tiny parastyle and a well−developed heel. In the m 1 the oblique cristid terminates far labially, whereas in m 2 it ascends and joins the marked metacristid. In the m 1 there is only a vestige of a short ectocingulid below the hypoflexid, in the m 2 a faint precingulid. Both have a well−developed entostylid. Description of the paratypes Dentary. — In two further fragments the posterior mental foramen is situated below the trigonid of m 1; in one specimen, like in the type, the p 3 is obliquely implanted. The slightly crowded p 3 and p 2 seem to be a consistent character of the species.	en	Ziegler, Reinhard (2003): Moles (Talpidae) from the late Middle Miocene of South Germany. Acta Palaeontologica Polonica 48 (4): 617-648, DOI: 10.5281/zenodo.13396039
480C879940177609DD28D1E5FB5BFB88.taxon	description	Lower dentition. — With respect to the lower dentition there is no additional information except for the m 3. In the last molar the precingulid is more marked, the oblique cristid joins the metacristid. The reduced talonid has no entostylid. In size and morphology all five m 3 fit well with one another. However, it cannot be excluded that one or more belong to Urotrichini gen. et sp. indet. II and / or to Desmanella sp., which is of roughly the same size. The well−developed precingulids of these m 3 would argue in favour of the association with Desmanella sp. whose m 1 and m 2 have well−developed precingulids. However, Desmanella sp. is represented only by six specimens. It is unlikely, that all five m 3 belong to this species. Upper dentition. — The P 4 has a tiny parastyle and a well−developed protocone. There is a weak precingulum and a more pronounced postcingulum. The M 1 has an undivided mesostyle, hardly differentiated para− and metaconule respectively, a marked paracingulum, which joins the projecting parastyle and a weak metacingulum, which tapers distolabially. In the M 2 there is a marked paraconule and a well−developed metaconule. Para− and metacingulum are absent. The mesostyle is undivided. Humerus. — The associated humeri are the smallest and most gracile ones among the Petersbuch 31 talpid humeri. There is a suite of urotrichine characters: large, pocketed supratrochlear fossa, deep, concave notch between trochlea and the fossa m. flexor digitorum profundus ligament, small and moderately deep brachialis fossa. In anterior view the trochlea is broadening towards the capitulum. The long axis of the elliptical head runs nearly parallel to the shaft. The marked ledge extending from the lesser tubercle to beneath the head is rounded. The olecranon fossa is shallow. The teres tubercle forms a long crest proximally not covered by the pectoral ridge. The pectoral tubercle is situated labially on the shaft. There is a deep groove between head and major tubercle. The passage of the biceps tendon between teres tubercle and pectoral crest is a bicipital notch. Comparisons As the material under study without any doubt represents a species of the tribe Urotrichini, the comparisons mainly refer to the fossil and extant species of this tribe. Storch and Qiu (1983: tabes 6 and 7) listed a suite of features characterising both the Recent genera Urotrichus Temminck, 1841 and Neurotrichus Günther, 1880. The Recent Urotrichustalpoides Temminck, 1841 and U. pilirostris (True, 1886) differ from Tenuibrachiatum storchi in the: – humerus with a long axis of the head directed disto−laterally, – shorter, proximally angled teres tubercle (see Storch and Qiu 1983: figs. 20, 21), – pectoral tubercle bent laterally, thus being visible in posterior view, – dentary with a reduced antemolar region not tapered anteriorly, – more posterior position of the mental foramina, – at large more compact teeth (see Hugueney 1972: figs. 16, 17).	en	Ziegler, Reinhard (2003): Moles (Talpidae) from the late Middle Miocene of South Germany. Acta Palaeontologica Polonica 48 (4): 617-648, DOI: 10.5281/zenodo.13396039
480C879940177609DD28D1E5FB5BFB88.taxon	description	– a reduced antemolar region, – a p 4 with a disto−lingual cuspule, – a m 1 with a better−developed precingulid.	en	Ziegler, Reinhard (2003): Moles (Talpidae) from the late Middle Miocene of South Germany. Acta Palaeontologica Polonica 48 (4): 617-648, DOI: 10.5281/zenodo.13396039
480C879940177609DD28D1E5FB5BFB88.taxon	description	M 1, – the presence of an metacingulum and the weaker paraconule of m 2. Neurotrichuspolonicus Skoczeń, 1980 from the Pliocene of Poland differs from Tenuibrachiatum storchi in: – being distinctly bigger (see Skoczeń 1980: tables 5, 7), – the morphological characters listed above for N. gibbsi.	en	Ziegler, Reinhard (2003): Moles (Talpidae) from the late Middle Miocene of South Germany. Acta Palaeontologica Polonica 48 (4): 617-648, DOI: 10.5281/zenodo.13396039
480C879940177609DD28D1E5FB5BFB88.taxon	description	– the complete set of lower antemolars, but single−rooted lower premolars, – lower molars with marked pre− and ectocingulids, – more robust humerus, which shows advanced fossorial adaptations. M. gracilis differs from Tenuibrachiatum in having: – lower molars with better developed pre− and ectocingulid respectively, – M 1 without projecting parastyle and with divided mesostyle, – a slightly bigger humerus with a pectoral tubercle situated more laterally and being visible in posterior view.	en	Ziegler, Reinhard (2003): Moles (Talpidae) from the late Middle Miocene of South Germany. Acta Palaeontologica Polonica 48 (4): 617-648, DOI: 10.5281/zenodo.13396039
480C879940177609DD28D1E5FB5BFB88.taxon	description	– less projecting parastyle of M 1, – hardly developed metaconule of M 2.	en	Ziegler, Reinhard (2003): Moles (Talpidae) from the late Middle Miocene of South Germany. Acta Palaeontologica Polonica 48 (4): 617-648, DOI: 10.5281/zenodo.13396039
480C879940177609DD28D1E5FB5BFB88.taxon	description	– projecting parastyle of P 4, – better developed para− and metacingulum of M 1 and M 2. Paratalpa Lavocat, 1951 is known from the Oligocene species P. micheli Lavocat, 1951, the Agenian P. micheli saulcetensis Hugueney, 1972, P. brachychir (von Meyer, 1846), and P. meyeri (Schlosser, 1887). They are all distinctly bigger, have a dentary with a more reduced antemolar region, lower molars with better−developed cingulids and further buccally terminating oblique cristids, upper molars with deeply divided and spaced mesostyles. The humerus, known from P. micheli, P. brachychir, and P. meyeri, is bigger, more robust and has a shorter teres tubercle.	en	Ziegler, Reinhard (2003): Moles (Talpidae) from the late Middle Miocene of South Germany. Acta Palaeontologica Polonica 48 (4): 617-648, DOI: 10.5281/zenodo.13396039
480C879940177609DD28D1E5FB5BFB88.taxon	discussion	Discussion Both dentition and associated humeri show a suite of urotrichine characters, which leaves the tribal assignation beyond any doubt. I think the association of humeri and dentition is correct. The smallest are expected to belong to the smallest dentition. Additionally, it is assumed that the humeri have to be associated with the dentition that roughly corresponds in the number of specimens. The Petersbuch 31 fauna also yielded four dentary fragments and two m 2 of Desmanella, which correspond in size to Tenuibrachiatum. The only Desmanella species with associated humeri is D. engesseri Ziegler, 1985 from the Early Miocene fissure fill Petersbuch 2 (Ziegler 1985: fig. 2). In this species the teres tubercle is distinctly shorter and forms no ledge. Consequently, we can be rather confident that the association of humeri – dentition for Tenuibrachiatum is correct. T. storchi cannot be ancestral to the Recent Neurotrichus nor to Quyania. Both have all three lower incisors, whereas one is lost in Tenuibrachiatum. In anterior view the humerus of the later genera shows a narrow trochlea, which is connected to the capitulum by a thin bridge of the articular facets (see Storch and Qiu 1983: figs. 17 – 19). In the humerus morphology Tenuibrachiatum shows more affinities to Urotrichus. In the extant species the antemolar region is more reduced. Tenuibrachiatum is a possible candidate for the ancestry of Urotrichus. The ancestor of Tenuibrachiatum is expected to have the full set of lower antemolars and double−rooted lower premolars, hence one tooth (? i 3) with one additional alveolus. Neither Myxomygale nor Paratalpa fulfil this qualification. In spite of the Oligocene to Early Miocene correlation of most species, both genera are more advanced with respect to the degree of reduction in the lower antemolar region. In dental morphology Tenuibrachiatum is somewhat closer to Myxomygale.	en	Ziegler, Reinhard (2003): Moles (Talpidae) from the late Middle Miocene of South Germany. Acta Palaeontologica Polonica 48 (4): 617-648, DOI: 10.5281/zenodo.13396039
480C879940147609DD28D11EFAE8FB23.taxon	materials_examined	Type species: Desmanella stehlini Engesser, 1972.	en	Ziegler, Reinhard (2003): Moles (Talpidae) from the late Middle Miocene of South Germany. Acta Palaeontologica Polonica 48 (4): 617-648, DOI: 10.5281/zenodo.13396039
480C879940147604DD28D1C9FE22FBF7.taxon	description	Fig. 11. Material and measurements. — Petersbuch 6: NHMA P 6 − 1064, left dentary fragment with m 1 – m 3, Lm 1 – m 3 (4.19), m 1 (1.53 × 1.00 × 1.03), m 2 (1.59 × 1.04 × 1.02), m 3 (1.26 × 0.80 × 0.62). Petersbuch 18: NHMA P 18 − 754, left dentary fragment with p 4 – m 3, Lm 1 – m 3 (4.15), m 1 (1.57 × 0.93 × 1.07), m 2 (1.61 × 0.99 × 1.00), m 3 (1.26 × 0.76 × 0.65). Petersbuch 31: NHMA P 31 − 0165 / 1, left dentary fragment with p 3 + m 2, p 3 (0.49 × 0.45), m 2 (> 1.5 × 0.97 × 0.96); CRW P 31 − 0165 / 2, right dentary fragment with p 3 – p 4, p 3 (0.42 × 0.40), p 4 (0.97 × 0.69); NHMA P 31 − 0165 / 3, right dentary fragment with p 4 – m 1, p 4 (0.92 × 0.67), m 1 (1.46 × 0.95 × 1.08); NHMA P 31 − 0165 / 4, left dentary fragment with m 1, m 1 (1.48 × 1.01 × 1.10). Description Dentary. — Only short fragments of the horizontal ramus are preserved. The specimen from Petersbuch 18 shows the posterior mental foramen between the roots of p 4 and m 1 and the anterior one under the third root anterior to p 4, the p 1 alveolus. In the small sample from Petersbuch 31 the anterior mental foramen is below p 2 (twice) or between the roots of p 1 and p 2 (once) and the posterior one beneath the trigonid of m 1. One specimen shows the complete set of antemolar alveoles. There is one alveolus each for i 2, i 3, c, p 1, p 2, and p 3. Consequently, the lower tooth formula is 2 − 1 − 4 − 3. Lower dentition. — The teeth anterior to p 3 are not preserved. Their alveoli show that all are single−rooted. The likewise single−rooted p 3 has a conical cusp with a posterior cingulid. The double−rooted p 4 is oval in occlusal outline. The crown has a concave posterior face, a convex mesiobuccal side, and a flat mesio−lingual one. The posterior cingulid encompasses a short talonid. The molars are slightly inflated and low−crowned. The size relation is m 2> m 1> m 3. In m 1 the oblique cristid joins the centre of the protocristid, in m 2 and m 3 it extends more lingually but does not join the weak metacristid. There is no well−developed metacristid in m 1, but a weak, descending entocristid. A moderately developed cingulid runs from below the paraconid to the hypoflexid. The weak postcingulid joins the entostylid. The m 2 and m 3 are characterised by their short talonid. In m 2 the protoconid is somewhat higher, the talonid narrower, and the oblique cristid joins the marked metacristid. The m 3 has neither postcingulid nor entostylid. Discussion Desmanellastehlini, the genotype, was described for the first time by Engesser (1972) on the basis of six isolated molars from the Anwil fauna. The specimens under study fit well in morphology and length with the molars from the type locality, but they are narrower. The dentaries and teeth from Petersbuch 6 + 18 correspond well in size with those from Petersbuch 31, but differ in the slightly more posterior position of the mental foramen and in the somewhat weaker preand ectocingulid of m 2. The position of the mental foramen is not known from D. stehlini from the type locality. For want of the upper dentition in our material the presence of important characters cannot be verified. Hence the determination is Desmanella cf. stehlini. To date, the record of D. stehlini is extremely sparse. Kälin (1993) reported on five isolated teeth of D. aff. stehlini from Le Locle sous le Stand, Switzerland, which is correlatable with MN 7 + 8. Kälin and Engesser (2001) designated two isolated teeth from Nebelbergweg, a MN 9 − fauna from Switzerland, Desmanella sp. Probably they also represent D. stehlini. The authors refrained from specific determination because of insufficient material. Crochet and Green (1982) referred 15 isolated teeth and a dentary fragment from Montredon, an Upper Miocene (MN 10) fauna from France, to Desmanella cf. stehlini. The genus Desmanella itself has a long stratigraphic range. The earliest records are from the Oligocene / Miocene transition in South Germany (Ziegler 1990), the latest is represented by Desmanella gardiolensis Crochet, 1986 from the Late Pliocene (MN 16) fauna Balaruc 2 in South France (Crochet 1986). The subfamilial allocation of the genus is a matter of continuous dispute. Desmanella was referred to the Desmaninae (Engesser 1972), the Talpinae (Storch 1978) and by most students to the Uropsilinae (e. g., Rümke 1974, 1976; Engesser 1980; Ziegler 1985; Crochet 1986; and van den Hoek Ostende 2001). The whole story is reviewed and comprehensively discussed in Dahlmann (2001) and van den Hoek Ostende (2001). My arguments for an allocation with the Uropsilinae have been the associated humeri of D. engesseri, which are characterised by the absence of a bicipital tunnel. According to Campbell (1939), except from the Uropsilinae in all mole humeri the walls of the bicipital groove are fused to form a tunnel. However, in the Recent Urotrichus talpoides the bicipital groove is not fully ossified but rather closed by cartilage (Dahlmann 2001: 47). Another particularly important character of the uropsiline humerus is the rounded caput, which is elliptical in all other talpids. The humerus of D. engesseri has an elliptical caput (see Ziegler 1985: fig. 2 b, 1994: pl. 1: 6, 7), hence they cannot belong to an uropsiline. I think, with respect to humerus morphology, Desmanella is better placed within the Urotrichini. Dental morphology and tooth formula is compatible with this allocation. The presence of a functional milk dentition, the main argument of the Uropsilinae advocates, is also known from some Urotrichini, for example Urotrichus Temminck, 1841 and Quyania chowi Storch and Qiu, 1983. Urotrichini gen. et sp. indet. I Fig. 12. Material and measurements. — Petersbuch 10: NHMA P 10 − 616 / 1, left dentary fragment with p 3, p 3 (0.93 × 0.53); NHMA P 10 − 616 / 2, right m 1 (1.78 × 0.87 × 1.01); NHMA P 10 − 617, right humerus GL (10.3), Bp (6.17), BpwT (4.55), DS (2.03), Bp * 100 / GL (59.9). Description Dentary. — There is one dentary fragment with the alveoli of i 1 – m 1, a double−rooted p 3, and the talonid of p 4. In this specimen a small posterior mental foramen is situated under the anterior root of p 4, the anterior one beneath p 2. There are five alveoli anterior to p 3 for the single−rooted i 1, i 2, i 3, p 1, and p 2. The incisors are increasingly inclined, i 1 being the largest. The canine is assumed to be eliminated. In the p 3 the crown is buccally convex and flat on the lingual side. It is surrounded by a cingulid which tapers lingually. From the p 4 only the posteriormost part with the marked postcingulid is preserved. m 1. — There is one isolated specimen with a notched paraand protocristid respectively. The oblique cristid extends far lingually but does not join the metacristid. The precingulid is short; the ectocingulid restricted to the hypoflexid, the postcingulid is extremely weak and short. Humerus. — The supratrochlear fossa, the fossa for the m. flexor digitorum profundus ligament with the medial epicondyle and the greater tubercle with the deltoid process are broken away. In spite of the fracture the deep notch between trochlea and the fossa, which characterises the urotrichines, is partly preserved. The pectoral tubercle extends halfway down the shaft and is situated in its mid, not laterally as in the other urotrichines. The brachialis fossa is only partly preserved; it was only moderately deep. Greater tubercle and head are separated by a deep groove. There is a notch between and lesser tubercle and teres tubercle. Discussion The humerus is the largest and most robust among the urotrichine humeri of all samples under study. It is associated with the biggest urotrichine dentary and the biggest urotrichine m 1 of the Petersbuch 10 talpid sample. There is no more probable alternative to this association, nonetheless it is considered tentative. Neither humerus nor dental remains fit well with any known species or genus. In view of the uncertainties concerning the association and as the material is too scarce the description of a new taxon is not possible. Urotrichini gen. et sp. indet. II Fig. 13. Material and measurements. — Petersbuch 31, P 31 − 171 / 1 – 6: NHMA P 31 − 171 / 1, leftm 2 (1.61 × 0.99 × 0.93); CRW P 31 − 171 / 2, right m 2 (1.77 × 1.05 × 1.08); NHMA P 31 − 171 / 3, right M 1 (2.06 × 1.47); NHMA P 31 − 171 / 4, left M 2 (1.90 × 1.87); CRW P 31 − 171 / 5, right M 2 (1.94 × 1.97); CRW P 31 − 171 / 6, right M 2 (1.57 × 1.85). Description Lower dentition. — In both m 2 the metacristid does reach the cusp of the metaconid. It instead ends in a small metastylid below the metaconid. The oblique cristid joins the metacristid. There is a marked precingulid, a short ectocingulid beneath the hypoflexid and a vestigial postcingulid close to the entostylid. Upper dentition. — The mesostyle of the M 1 is nearly confluent in the moderately worn tooth. A slight notch shows that the mesostyle was divided in the unworn tooth. Paraconule and metaconule are differentiated. The preparaconuluscrista is confluent with the strong paracingulum, which itself joins the projecting parastyle. Postmetaconuluscrista and metacingulum are also confluent and terminate in the metastyle. In all three M 2 the mesostyle is deeply divided. Paraconule and metaconule a more marked than in the M 1. The preparaconuluscrista terminates at the mesial basis of the paracone, the postmetaconuluscrista at the distal basis of the metacone. Discussion The teeth cannot be referred to any other species of the Petersbuch 31 talpid fauna. They differ from Tenuibrachiatum storchi in the bigger size, the better developed precingulids of m 2 and in the divided mesostyles of the M 1 and M 2. The marked lingual conules in the upper molars, especially in M 2, is a distinct desmanine and urotrichine character. As desman teeth are more massive and differ in a suite of other characters, an affiliation with this subfamily can be excluded. It is assumed that all teeth form a homogeneous sample, in spite of the small M 2 (no. 6). The teeth are too small for an association with the dentary of Scalopini gen. et sp. indet. Furthermore, the marked lingual conules of the upper molars better fit with the urotrichines. We cannot exclude that the teeth represent the same species as the indeterminable urotrichine from Petersbuch 10. As both samples have neither teeth nor postcranial elements in common this assumption cannot be corroborated. Hence, the determination is Urotrichini gen. et sp. indet. II. Talpidae incertae sedis	en	Ziegler, Reinhard (2003): Moles (Talpidae) from the late Middle Miocene of South Germany. Acta Palaeontologica Polonica 48 (4): 617-648, DOI: 10.5281/zenodo.13396039
480C879940147604DD28D1C9FE22FBF7.taxon	description	Fig. 11. Material and measurements. — Petersbuch 6: NHMA P 6 − 1064, left dentary fragment with m 1 – m 3, Lm 1 – m 3 (4.19), m 1 (1.53 × 1.00 × 1.03), m 2 (1.59 × 1.04 × 1.02), m 3 (1.26 × 0.80 × 0.62). Petersbuch 18: NHMA P 18 − 754, left dentary fragment with p 4 – m 3, Lm 1 – m 3 (4.15), m 1 (1.57 × 0.93 × 1.07), m 2 (1.61 × 0.99 × 1.00), m 3 (1.26 × 0.76 × 0.65). Petersbuch 31: NHMA P 31 − 0165 / 1, left dentary fragment with p 3 + m 2, p 3 (0.49 × 0.45), m 2 (> 1.5 × 0.97 × 0.96); CRW P 31 − 0165 / 2, right dentary fragment with p 3 – p 4, p 3 (0.42 × 0.40), p 4 (0.97 × 0.69); NHMA P 31 − 0165 / 3, right dentary fragment with p 4 – m 1, p 4 (0.92 × 0.67), m 1 (1.46 × 0.95 × 1.08); NHMA P 31 − 0165 / 4, left dentary fragment with m 1, m 1 (1.48 × 1.01 × 1.10). Description Dentary. — Only short fragments of the horizontal ramus are preserved. The specimen from Petersbuch 18 shows the posterior mental foramen between the roots of p 4 and m 1 and the anterior one under the third root anterior to p 4, the p 1 alveolus. In the small sample from Petersbuch 31 the anterior mental foramen is below p 2 (twice) or between the roots of p 1 and p 2 (once) and the posterior one beneath the trigonid of m 1. One specimen shows the complete set of antemolar alveoles. There is one alveolus each for i 2, i 3, c, p 1, p 2, and p 3. Consequently, the lower tooth formula is 2 − 1 − 4 − 3. Lower dentition. — The teeth anterior to p 3 are not preserved. Their alveoli show that all are single−rooted. The likewise single−rooted p 3 has a conical cusp with a posterior cingulid. The double−rooted p 4 is oval in occlusal outline. The crown has a concave posterior face, a convex mesiobuccal side, and a flat mesio−lingual one. The posterior cingulid encompasses a short talonid. The molars are slightly inflated and low−crowned. The size relation is m 2> m 1> m 3. In m 1 the oblique cristid joins the centre of the protocristid, in m 2 and m 3 it extends more lingually but does not join the weak metacristid. There is no well−developed metacristid in m 1, but a weak, descending entocristid. A moderately developed cingulid runs from below the paraconid to the hypoflexid. The weak postcingulid joins the entostylid. The m 2 and m 3 are characterised by their short talonid. In m 2 the protoconid is somewhat higher, the talonid narrower, and the oblique cristid joins the marked metacristid. The m 3 has neither postcingulid nor entostylid. Discussion Desmanellastehlini, the genotype, was described for the first time by Engesser (1972) on the basis of six isolated molars from the Anwil fauna. The specimens under study fit well in morphology and length with the molars from the type locality, but they are narrower. The dentaries and teeth from Petersbuch 6 + 18 correspond well in size with those from Petersbuch 31, but differ in the slightly more posterior position of the mental foramen and in the somewhat weaker preand ectocingulid of m 2. The position of the mental foramen is not known from D. stehlini from the type locality. For want of the upper dentition in our material the presence of important characters cannot be verified. Hence the determination is Desmanella cf. stehlini. To date, the record of D. stehlini is extremely sparse. Kälin (1993) reported on five isolated teeth of D. aff. stehlini from Le Locle sous le Stand, Switzerland, which is correlatable with MN 7 + 8. Kälin and Engesser (2001) designated two isolated teeth from Nebelbergweg, a MN 9 − fauna from Switzerland, Desmanella sp. Probably they also represent D. stehlini. The authors refrained from specific determination because of insufficient material. Crochet and Green (1982) referred 15 isolated teeth and a dentary fragment from Montredon, an Upper Miocene (MN 10) fauna from France, to Desmanella cf. stehlini. The genus Desmanella itself has a long stratigraphic range. The earliest records are from the Oligocene / Miocene transition in South Germany (Ziegler 1990), the latest is represented by Desmanella gardiolensis Crochet, 1986 from the Late Pliocene (MN 16) fauna Balaruc 2 in South France (Crochet 1986). The subfamilial allocation of the genus is a matter of continuous dispute. Desmanella was referred to the Desmaninae (Engesser 1972), the Talpinae (Storch 1978) and by most students to the Uropsilinae (e. g., Rümke 1974, 1976; Engesser 1980; Ziegler 1985; Crochet 1986; and van den Hoek Ostende 2001). The whole story is reviewed and comprehensively discussed in Dahlmann (2001) and van den Hoek Ostende (2001). My arguments for an allocation with the Uropsilinae have been the associated humeri of D. engesseri, which are characterised by the absence of a bicipital tunnel. According to Campbell (1939), except from the Uropsilinae in all mole humeri the walls of the bicipital groove are fused to form a tunnel. However, in the Recent Urotrichus talpoides the bicipital groove is not fully ossified but rather closed by cartilage (Dahlmann 2001: 47). Another particularly important character of the uropsiline humerus is the rounded caput, which is elliptical in all other talpids. The humerus of D. engesseri has an elliptical caput (see Ziegler 1985: fig. 2 b, 1994: pl. 1: 6, 7), hence they cannot belong to an uropsiline. I think, with respect to humerus morphology, Desmanella is better placed within the Urotrichini. Dental morphology and tooth formula is compatible with this allocation. The presence of a functional milk dentition, the main argument of the Uropsilinae advocates, is also known from some Urotrichini, for example Urotrichus Temminck, 1841 and Quyania chowi Storch and Qiu, 1983. Urotrichini gen. et sp. indet. I Fig. 12. Material and measurements. — Petersbuch 10: NHMA P 10 − 616 / 1, left dentary fragment with p 3, p 3 (0.93 × 0.53); NHMA P 10 − 616 / 2, right m 1 (1.78 × 0.87 × 1.01); NHMA P 10 − 617, right humerus GL (10.3), Bp (6.17), BpwT (4.55), DS (2.03), Bp * 100 / GL (59.9). Description Dentary. — There is one dentary fragment with the alveoli of i 1 – m 1, a double−rooted p 3, and the talonid of p 4. In this specimen a small posterior mental foramen is situated under the anterior root of p 4, the anterior one beneath p 2. There are five alveoli anterior to p 3 for the single−rooted i 1, i 2, i 3, p 1, and p 2. The incisors are increasingly inclined, i 1 being the largest. The canine is assumed to be eliminated. In the p 3 the crown is buccally convex and flat on the lingual side. It is surrounded by a cingulid which tapers lingually. From the p 4 only the posteriormost part with the marked postcingulid is preserved. m 1. — There is one isolated specimen with a notched paraand protocristid respectively. The oblique cristid extends far lingually but does not join the metacristid. The precingulid is short; the ectocingulid restricted to the hypoflexid, the postcingulid is extremely weak and short. Humerus. — The supratrochlear fossa, the fossa for the m. flexor digitorum profundus ligament with the medial epicondyle and the greater tubercle with the deltoid process are broken away. In spite of the fracture the deep notch between trochlea and the fossa, which characterises the urotrichines, is partly preserved. The pectoral tubercle extends halfway down the shaft and is situated in its mid, not laterally as in the other urotrichines. The brachialis fossa is only partly preserved; it was only moderately deep. Greater tubercle and head are separated by a deep groove. There is a notch between and lesser tubercle and teres tubercle. Discussion The humerus is the largest and most robust among the urotrichine humeri of all samples under study. It is associated with the biggest urotrichine dentary and the biggest urotrichine m 1 of the Petersbuch 10 talpid sample. There is no more probable alternative to this association, nonetheless it is considered tentative. Neither humerus nor dental remains fit well with any known species or genus. In view of the uncertainties concerning the association and as the material is too scarce the description of a new taxon is not possible. Urotrichini gen. et sp. indet. II Fig. 13. Material and measurements. — Petersbuch 31, P 31 − 171 / 1 – 6: NHMA P 31 − 171 / 1, leftm 2 (1.61 × 0.99 × 0.93); CRW P 31 − 171 / 2, right m 2 (1.77 × 1.05 × 1.08); NHMA P 31 − 171 / 3, right M 1 (2.06 × 1.47); NHMA P 31 − 171 / 4, left M 2 (1.90 × 1.87); CRW P 31 − 171 / 5, right M 2 (1.94 × 1.97); CRW P 31 − 171 / 6, right M 2 (1.57 × 1.85). Description Lower dentition. — In both m 2 the metacristid does reach the cusp of the metaconid. It instead ends in a small metastylid below the metaconid. The oblique cristid joins the metacristid. There is a marked precingulid, a short ectocingulid beneath the hypoflexid and a vestigial postcingulid close to the entostylid. Upper dentition. — The mesostyle of the M 1 is nearly confluent in the moderately worn tooth. A slight notch shows that the mesostyle was divided in the unworn tooth. Paraconule and metaconule are differentiated. The preparaconuluscrista is confluent with the strong paracingulum, which itself joins the projecting parastyle. Postmetaconuluscrista and metacingulum are also confluent and terminate in the metastyle. In all three M 2 the mesostyle is deeply divided. Paraconule and metaconule a more marked than in the M 1. The preparaconuluscrista terminates at the mesial basis of the paracone, the postmetaconuluscrista at the distal basis of the metacone. Discussion The teeth cannot be referred to any other species of the Petersbuch 31 talpid fauna. They differ from Tenuibrachiatum storchi in the bigger size, the better developed precingulids of m 2 and in the divided mesostyles of the M 1 and M 2. The marked lingual conules in the upper molars, especially in M 2, is a distinct desmanine and urotrichine character. As desman teeth are more massive and differ in a suite of other characters, an affiliation with this subfamily can be excluded. It is assumed that all teeth form a homogeneous sample, in spite of the small M 2 (no. 6). The teeth are too small for an association with the dentary of Scalopini gen. et sp. indet. Furthermore, the marked lingual conules of the upper molars better fit with the urotrichines. We cannot exclude that the teeth represent the same species as the indeterminable urotrichine from Petersbuch 10. As both samples have neither teeth nor postcranial elements in common this assumption cannot be corroborated. Hence, the determination is Urotrichini gen. et sp. indet. II.	en	Ziegler, Reinhard (2003): Moles (Talpidae) from the late Middle Miocene of South Germany. Acta Palaeontologica Polonica 48 (4): 617-648, DOI: 10.5281/zenodo.13396039
480C879940197604DE62D1FCFDB0FB4E.taxon	materials_examined	Type species: Desmanodon major Engesser, 1980.	en	Ziegler, Reinhard (2003): Moles (Talpidae) from the late Middle Miocene of South Germany. Acta Palaeontologica Polonica 48 (4): 617-648, DOI: 10.5281/zenodo.13396039
