taxonID	type	description	language	source
4F16166CFFF2FF954DFFA5DE3BF5FCD4.taxon	materials_examined	Type species: Harnischia (Cladopelma) tylus Townes, 1945 Other included species: S. digitata Yan et al., 2011, S. glabra Yan et al., 2011, S. hirta Saether, 1983, S. reissi Jackson, 1977, S. separate Yan et al., 2011, S. tamanipparai (Sasa, 1983). Emended diagnosis. Male: as in Cranston et al. (1989: 410) with the following emendations: wing length 2.0 – 2.4 mm; AR 1.81 – 2.10; antepronotals 1 – 5, acrostichals 6 – 11, dorsocentrals 6 – 13, supraalar 1 – 2, prealars 2 – 5; squama with 2 – 10 setae; superior volsella consist of bare dorsal lobe with a finger-shaped sclerotized projection and ventral lobe covered with microtrichia and bearing 2 setae. Pupa: as in Pinder and Reiss (1986: 350) with the following emendations: sternite IV with weak median shagreen; segment I with 1 L seta. Fourth instar larva: as in Epler et al. (2013: 434) with the following emendations: SIII hair-like.	en	Orel, Oksana V. (2014): To systematics of the genus Saetheria Jackson (Diptera, Chironomidae) from the Russian Far East. Zootaxa 3802 (1): 65-74, DOI: 10.11646/zootaxa.3802.1.5
4F16166CFFF2FF904DFFA7F73C56F8EA.taxon	materials_examined	Material examined. PRIMORYE TERRITORY: male, Nadezhdenky Region, Razdol’naya River, 31.05.1990, Vshivkova T. S.; 1 male, same data, 0 2.06.1990, Vshivkova T. S.; male, Khankaisky Region, Khanka Lake basin, Kommisarovka River, 17.06.1999, Makarchenko E. A.; 2 males, 2 larvae, Khasansky Region, Barabashevka River, 0 1.06.2002, Teslenko V. A.; P-male, same data, 0 4.06.2002, Teslenko V. A.; male, larva, same data, 0 1.06.2003, Makarchenko E. A.; larva, same data, 12.05.2004, Makarcheko E. A.; male, Khasansky Region, Izvestkovaya River, 31.05.2003, Lubaretz V.; larva, Partizansraya River about Peretino Vallage, 29.05.2004, Vshivkova T. S.; male, Terneisky Region, Samarga River, 0 7.08.2006, Zorina O. V.; KHABAROVSK TERRITORY: Р-male, Chegdomynsky Region, Chegdomyn River, 13.07.2003, Tiunova T. M.; larva, Urgal River, 18.07.2003, Tiunova T. M. AMUR TERRITORY: P-male, 2 larvae, Zeya River basin, Ulunga River near Aprel’sky Village, 20.07.2008, Kotzuk D. B. SAKHALIN REGION: male, Otchepukha River about Lesnoe Village, 0 3.07.1985, Makarchenko E. A.; MAGADAN TERRITORY: P-male, pupal exuviae; Tauy River; 17.07.2001, Kocharina C.; SOKHA REPUBLIC (YAKUTIA): P-male with larval exuviae, Aldan River basin, Chul’man River, 25.07.2006, Makarchenko E. A. Diagnostic characters. The male of S. reissi Jackson is distinguished by the following combination of features: wing length 2.1 – 2.4 mm; AR 1.81 – 2.10; small frontal tubercles always present; anal point spoon-shaped; superior volsella consist of bare dorsal lobe with a finger-shaped sclerotized projection and triangular ventral lobe covered with microtrichia and bearing 1 strong apical seta and 1 weaker median seta; gonostylus narrowed in middle. The pupa with 1 L seta on segment I, with 2 – 3 L setae on segments II – IV, with weak median shagreen on sternite IV, with median shagreen on tergite V. The larva with setae SI seta-like, SII broadly blade-like, SIII hairlike; AR 1.13 – 1.26; segment 2 of antenna 5 – 7 µm long, apically with the blade reaches the apex of the segment 5; premandible with 3 large distal teeth and 1 small proximal tooth; seta subdentalis a wide blade-like; mentum with broad triangular or round median tooth and 6 pair of lateral teeth; third pair lateral teeth below the remaining teeth.	en	Orel, Oksana V. (2014): To systematics of the genus Saetheria Jackson (Diptera, Chironomidae) from the Russian Far East. Zootaxa 3802 (1): 65-74, DOI: 10.11646/zootaxa.3802.1.5
4F16166CFFF2FF904DFFA7F73C56F8EA.taxon	description	Male (n = 7). Total length 3.2 – 3.8 mm. Wing length 2.1 – 2.4 mm. Total length / wing length 1.52 – 1.76. Colouration. Pedicel of antenna, mesonotal stripes and postnotum yellowish brown, ground colour and scutellum pale yellowish. Abdomen yellowish or yellowish-brown. Fore legs yellowish with brown distal end of femora, proximal third and distal end of tibia, ta 1 – 5 gradually darkened toward ends, tibiae sometimes brown. Mid and hind legs yellowish, ta 1 – 5 gradually darkened toward ends. Head. Frontal tubercles small (3.4 – 10.2 µm long, 3.4 – 6.8 µm wide). Antenna 944 – 1080 µm long; ultimate flagellomere 608 – 720 µm long; AR 1.81 – 2.10. Verticals 12 – 19. Clypeus with 8 – 10 setae. Maxillary palp 435 – 510 µm long; length of palpomeres 2 – 5 (in µm): 34 – 44, 109 – 126, 116 – 139, 170 – 204. Head width / palp length 0.42 – 0.43; antenna length / palp length 2.12 – 2.28. Thorax chaetotaxy. Antepronotals 1 – 3, acrostichals 6 – 11, dorsocentrals 6 – 13, supraalar 1 – 2, prealars 2 – 3. Scutellum with 5 – 9 setae. Wing 0.51 – 0.60 mm wide. Veins R with 0 – 7 setae, R 4 + 5 with 1 – 2 setae. Squama with 2 – 7 setae; brachiolum with 1 – 2 setae. VR 1.19 – 1.25. Legs. Spurs of middle tibia 24 – 27 µm, of hind tibia 27 – 31 and 34 µm long. Lengths and proportions of legs as in Table 1. Hypopygium (Figs 1 – 2). Laterosternite IX with 1 – 4 strong setae. Anal point spoon-shaped (length 54 – 61 µm, width 14 – 17 µm) at the base with 2 – 4 strong medial and 4 – 7 weak lateral setae. Gonocoxite 112 – 126 µm long, with 3 – 4 setae on inner margin. Transverse sternapodeme 68 – 82 µm wide. Phallapodeme 85 – 92 µm long. Superior volsella triangular (54 – 68 µm long and 14 – 17 µm wide in the base) composed of dorsal and ventral lobes. Dorsal lobe bare with a finger-shaped sclerotized projection (14 – 20 µm long), ventral lobe covered with microtrichia and bearing 1 strong apical seta (34 – 37 µm long) and 1 weaker seta (17 – 20 µm long) on middle. Inferior volsella with rounded apical part and covered with microtrichia. Gonostylus (147 – 170 µm long, 24 – 31 µm wide) narrowed in middle, apically round. HR 0.66 – 0.80. Pupa (n = 5, males). Cephalothorax (Figs 3 – 4). Cephalic tubercle conical (20 – 31 µm long, 34 – 51 µm wide) with apical spine; frontal setae 68 – 109 µm long (Fig. 3). Base of thoracic horn oval form (34 µm long, 17 µm wide). Lamelliform precorneals 2 (1 st 170 µm long, 2 nd 150 – 160 µm long), lamelliform antepronotals 2 (1 median 119 – 122 µm long and 1 lateral 290 µm long), dorsocentrals 4 (Dc 1 34 – 54 µm long, Dc 2 85 – 202 µm long, Dc 3 41 – 61 µm long, Dc 4 41 – 51 µm long); distance between Dc 1 and Dc 2 41 – 61 µm, Dc 2 and Dc 3 150 – 180 µm, Dc 3 and Dc 4 10 – 14 µm (Fig. 4). Abdomen (Figs 5 – 7) 2.2 – 2.4 mm long. Tergites I – III, VIII without shagreen. Hook row 153 – 204 µm long, with 44 – 50 spines. Tergites III – V on posterior margin with transverse band of strong dark spines distributed in 2 – 4 rows (tergite III with 47 – 62 spines, IV with 50 – 70 spines, V with 51 – 75 spines), tergite VI with weaker spines. Tergite IV with pair of anterolateral patches of shagreen (Fig. 5), sometimes present weak median shagreen (Fig. 6). Tergites V – VII with shagreen in a T-shaped (Fig. 7), sometimes tergite V with pair anterolateral patches of spines and median spines (Fig. 5) Sternite IV with median shagreen. Sternite V with or without posterolateral spines. Segment I with 1 L seta, segments II – IV with 2 – 3 L seta, V – VIII with 4 LS setae. Anal lobe with 28 – 40 lamelliform setae. Length of anal lobe 177 – 221 µm, width 187 – 279 µm. ALR 0.79 – 1.0. Fourth instar larva (n = 5). Green or greenish yellow (in formaldehyde). Head. Yellowish, 0.3 mm long, 0.2 mm wide Antenna 102 – 122 µm long, length of segments (in µm): 58 – 78, 5.1 – 6.8, 24 – 31, 5.1, 6.8, 5.1 (Fig. 8). AR 1.13 – 1.26. Basal segment with ring organ in proximal 1 / 4; distance from ring organ to base of antenna 17 µm; ROR 4.0 – 5.7. Blade 34 – 41 µm long, seated apically on segment 2, reaches the apex of the segment 5; accessory blade 20 – 27 µm. Lauterborn organ 3.4 µm long. Style 10 µm long. Seta SI seta-like 24 µm long, SII broadly blade-like 41 µm long, SIII hair-like 17 µm long, SIVa 3 - segmented 10 µm long and SIVb 10 µm long. Labral lamella absent. Premandible 54 µm long, with 3 large apical teeth and 1 small proximal tooth; premandibular brush well developed. Pecten epipharyngis triangular, 10 µm long (Fig. 9). Mandible 75 – 88 µm long, 41 µm wide, with apical (14 µm long) tooth and 2 brown inner teeth; seta subdentalis a wide blade-like 20 – 24 µm long; pecten mandibularis consist of 5 lamellae (Fig. 10). Maxillary palp 20 µm long, 12 – 14 µm wide; distance from ring organ to base of maxillary palp 7 – 10 µm; ROR 2 – 3 (Fig. 11). Mentum (61 – 68 µm wide) with brownish median tooth and 6 pair of lateral teeth. Median tooth (17 – 20 µm wide) triangular or round form; third lateral tooth below the remaining teeth (Figs 12 – 13). Ventromental plate 51 – 58 µm wide, 37 – 41 µm high; number of striae 21; distance between ventromental plates 31 – 34 µm. VmPR 1.25 – 1.55. VmPSR 1.5 – 1.9. Body. Posterior parapod with 17 yellow simple claws. Anal tubules conical, upper pair 85 – 102 µm long, bottom pair 102 – 119 µm long. Procercus 24 µm long, with 8 simple anal setae.	en	Orel, Oksana V. (2014): To systematics of the genus Saetheria Jackson (Diptera, Chironomidae) from the Russian Far East. Zootaxa 3802 (1): 65-74, DOI: 10.11646/zootaxa.3802.1.5
4F16166CFFF2FF904DFFA7F73C56F8EA.taxon	discussion	Remarks. The male of S. reissi Jackson is very similar to S. tylus (Townes) in the shape of the hypopygium, but can be separated by a larger body size (TL 3.2 – 3.8 mm), superior volsella roughly triangular form, gonostylus narrowed in middle. Whereas, the male of S. tylus has total length 2.30 – 3.18 mm, superior volsella an elongated foot-shaped, gonostylus narrowed in the proximal third (Townes 1945; Jackson 1977; Sasa 1989; Sasa & Tanaka 2001; Kobayashi 2007). The Russian far eastern specimens of S. reissi are different by larger body size from its found in the Irkutsk Territory (original description), Japan, and Korea (Jackson 1977; Sasa & Kondo 1993; Ree 2009). The pupa of S. reissi is also similar to S. tylus, but can be separated by the presence of 1 L seta on segment I, 2 – 3 L setae on segments II – IV, median shagreen on tergite V and on sternite IV. While, the pupa of S. tylus characterized by the absence of L setae on segment I, median shagreen on tergite V and sternite IV, and the presence of only 2 L setae on segments II – IV (Jackson 1977; Saether 1983; Pinder & Reiss 1986). According to Langton (1991) pupa of S. reissi has not shagreen on tergites IV – V and 1 L seta on segment I. Whereas pupae from our material are characterized by the presence of shagreen on tergites IV – V and 1 L seta on segment I. The larva also is very close to S. tylus, but can be easily separated by the presence of well developed premandibular brush, the blade of antenna reaches the apex of the segment 5. Whereas, the larva of S. tylus has not premandibular brush, the blade of antenna reaches of middle of the segment 5 (Jackson 1977). The larva of Chironominae genuinae № 9 described by Lipina (1926) from Oka River has 3 large teeth in the apical part and a large tooth in middle of premandible, and seta subdentalis a narrow blade-like. The larvae of S. reissi in our material with 3 large apical teeth and 1 small proximal tooth and seta subdentalis a wide blade-like.	en	Orel, Oksana V. (2014): To systematics of the genus Saetheria Jackson (Diptera, Chironomidae) from the Russian Far East. Zootaxa 3802 (1): 65-74, DOI: 10.11646/zootaxa.3802.1.5
4F16166CFFF2FF904DFFA7F73C56F8EA.taxon	distribution	Distribution. Saetheria reissi Jackson is quite widespread in the West Europen (Saether & Spies 2004). This species was so far reported from North Korea and Japan (Sasa & Kondo 1993; Ree 2009; Saether et al. 2000). In Russian S. reissi is known from Angara River near Irkutsk Townes by original description (Jackson 1977). This species is widespread in Russian Far East: Primorye, Khabarovsk and Amur Territories, Magadan and Sakhalin Regions, Sokha Republic (Yakutia) (Makarchenko et al. 2005).	en	Orel, Oksana V. (2014): To systematics of the genus Saetheria Jackson (Diptera, Chironomidae) from the Russian Far East. Zootaxa 3802 (1): 65-74, DOI: 10.11646/zootaxa.3802.1.5
4F16166CFFF6FF924DFFA4DB3D55F991.taxon	description	(Figs 14 – 15)	en	Orel, Oksana V. (2014): To systematics of the genus Saetheria Jackson (Diptera, Chironomidae) from the Russian Far East. Zootaxa 3802 (1): 65-74, DOI: 10.11646/zootaxa.3802.1.5
4F16166CFFF6FF924DFFA4DB3D55F991.taxon	materials_examined	Material examined. KHABAROVSK TERRITORY: 10 males, Jewish Autonomous Region, Bira River at about 4 km above of Zheltyi Yar Village, 12 – 15.08.2004, Tiunova T. M. Diagnostic characters. The male S. tamanipparai (Sasa) can be separated from other species of this genus by the following combination of characters: wing length 2.0 mm; AR 2.0; small frontal tubercles present; anal point drop-shaped; superior volsella consist of bare dorsal lobe with a finger-shaped sclerotized projection and footshaped ventral lobe covered with microtrichia and bearing 1 strong apical seta and 1 weaker seta in apical third; gonostylus widest in the proximal third and gradually narrowed toward apex.	en	Orel, Oksana V. (2014): To systematics of the genus Saetheria Jackson (Diptera, Chironomidae) from the Russian Far East. Zootaxa 3802 (1): 65-74, DOI: 10.11646/zootaxa.3802.1.5
4F16166CFFF6FF924DFFA4DB3D55F991.taxon	description	Male (n = 1). Total length 3.3 mm. Wing length 2.0 mm. Total length / wing length 1.65. Colouration. Pedicel of antenna, mesonotal stripes and postnotum yellowish brown, ground colour and scutellum pale yellowish. Abdomen yellowish gradually darkened toward end. Fore legs yellowish with brown distal end of femora, proximal third and distal end of tibia, ta 1 – 5 gradually darkened toward ends. Mid and hind legs yellowish, ta 1 – 5 gradually darkened toward ends. Head. Frontal tubercles small (6.8 µm long, 5.1 µm wide). Antenna 960 µm long; ultimate flagellomere 640 µm long; AR 2.0. Verticals 17. Clypeus with 15 setae. Maxillary palp 575 µm long; length of palpomeres 2 – 5 (in µm): 48, 146, 143, 238. Antenna length / palp length 1.67. Thorax chaetotaxy. Antepronotals 3 – 5, acrostichals 11, dorsocentrals 10, supraalar 1, prealars 4 – 5. Scutellum with 18 setae. Wing 0.54 mm wide. Veins R with 9 – 10 setae, R 4 + 5 with 2 setae. Squama with 10 setae; brachiolum with 1 setae. VR 1.12. Legs. Spurs of middle tibia 30 µm, of hind tibia 27 and 34 µm long. Lengths and proportions of legs as in Table 3. Hypopygium (Figs 14 – 15). Laterosternite IX with 1 strong seta. Anal point spoon-shaped (length 68 µm, width 17 µm) in the base with 7 strong median and 5 – 6 weak lateral setae. Gonocoxite 129 µm long, with 3 setae on inner margin. Transverse sternapodeme 75 µm wide. Phallapodeme 102 µm long. Superior volsella foot-shaped (75 µm long and 20 µm wide in the base) composed of dorsal and ventral lobes. Dorsal lobe bare with a finger-shaped sclerotized projection (17 µm long), ventral lobe covered with microtrichia and bearing 1 strong apical seta (27 µm long) and 1 weaker seta (14 µm long) in the apical third. Inferior volsella with rounded apical part and covered with microtrichia. Gonostylus (170 µm long, 31 µm wide) widest in proximal third and gradually narrowed toward apex. HR 0.79. Pupa unknown. Larva unknown.	en	Orel, Oksana V. (2014): To systematics of the genus Saetheria Jackson (Diptera, Chironomidae) from the Russian Far East. Zootaxa 3802 (1): 65-74, DOI: 10.11646/zootaxa.3802.1.5
4F16166CFFF6FF924DFFA4DB3D55F991.taxon	discussion	Remarks. Far eastern male of S. tamanipparai differ from the Japanese smaller body size (TL 3.3 mm), AR (2.0) and colour of fore femora (brown on distal end). Whereas, Japanese male has total length 3.75 mm, AR 1.67, and femora yellow (Sasa 1983).	en	Orel, Oksana V. (2014): To systematics of the genus Saetheria Jackson (Diptera, Chironomidae) from the Russian Far East. Zootaxa 3802 (1): 65-74, DOI: 10.11646/zootaxa.3802.1.5
4F16166CFFF6FF924DFFA4DB3D55F991.taxon	distribution	Distribution. Saetheria tamanipparai (Sasa) is recorded from Japan and Russian Far East (Sasa 1983; Makarchenko et al. 2005; Zorina 2006).	en	Orel, Oksana V. (2014): To systematics of the genus Saetheria Jackson (Diptera, Chironomidae) from the Russian Far East. Zootaxa 3802 (1): 65-74, DOI: 10.11646/zootaxa.3802.1.5
4F16166CFFF5FF9C4DFFA2C53A8DFCC1.taxon	description	(Figs 16 – 17)	en	Orel, Oksana V. (2014): To systematics of the genus Saetheria Jackson (Diptera, Chironomidae) from the Russian Far East. Zootaxa 3802 (1): 65-74, DOI: 10.11646/zootaxa.3802.1.5
4F16166CFFF5FF9C4DFFA2C53A8DFCC1.taxon	materials_examined	Material examined. PRIMORYE TERRITORY: male, Ussuryiskiy Region, Razdol’naya River basin, Kraunovka River, 21.07.2004, Arephina T. I. Diagnostic characters. The male of S. tylus (Townes) is distinguished by the following combination of features: wing length 1.7 mm; AR 1.74; small frontal tubercles present; anal point spoon-shaped; superior volsella consist of bare dorsal lobe with a finger-shaped sclerotized projection and elongated foot-shaped ventral lobe covered with microtrichia and bearing 1 strong apical seta and 1 weaker seta in the apical third; gonostylus narrowed in the proximal third.	en	Orel, Oksana V. (2014): To systematics of the genus Saetheria Jackson (Diptera, Chironomidae) from the Russian Far East. Zootaxa 3802 (1): 65-74, DOI: 10.11646/zootaxa.3802.1.5
4F16166CFFF5FF9C4DFFA2C53A8DFCC1.taxon	description	Male (n = 1). Total length 3.0 mm. Wing length 1.7 mm. Total length / wing length 1.76. Colouration. Pedicel of antenna, mesonotal stripes and postnotum yellowish brown, ground colour and scutellum pale yellowish. Abdomen yellowish. Fore legs yellowish with brown distal end of femora, proximal third and distal end of tibia, ta 1 – 5 gradually darkened toward ends, tibiae sometimes brown. Mid and hind legs yellowish, ta 1 – 5 gradually darkened toward ends. Head. Frontal tubercles small (5.1 µm long, 3.4 µm wide). Antenna 856 µm long; ultimate flagellomere 544 µm long; AR 1.74. Verticals 12 – 14. Clypeus with 10 setae. Maxillary palp 462 µm long; length of palpomeres 2 – 5 (in µm): 37, 109, 122, 194. Antenna length / palp length 1.85. Thorax chaetotaxy. Antepronotals 2, acrostichals 4, dorsocentrals 8 – 9, supraalar 1, prealars 3. Scutellum with 6 setae. Wing 0.48 mm wide. Veins R with 6 – 7 setae, R 4 + 5 with 2 setae. Squama with 4 setae; brachiolum with 1 setae. VR 1.31. Legs. Spurs of middle tibia 24 µm, of hind tibia 24 and 31 µm long. Lengths and proportions of legs as in Table 3. Hypopygium (Figs 16 – 17). Laterosternite IX with 3 strong setae. Anal point drop-shaped (length 51 µm, width 17 µm) at the base with 3 strong medial and 5 – 6 weak lateral setae. Gonocoxite 102 µm long, with 3 setae on inner margin. Transverse sternapodeme 51 µm wide. Phallapodeme 75 µm long. Superior volsella elongated foot-shaped (58 µm long and 31 µm wide in the base) composed of dorsal and ventral lobes. Dorsal lobe bare with a fingershaped sclerotized projection (17 µm long), ventral lobe covered with microtrichia and bearing 1 strong apical seta (27 µm long) and 1 weaker seta (10 – 14 µm long) in the apical third. Inferior volsella with rounded apical part and covered with microtrichia. Gonostylus (136 µm long, 20 – 24 µm wide) narrowed in the proximal third and apically round. HR 0.75. Pupa is described by Jackson (1977) and redescribed by Saether (1983) and Pinder & Reiss (1986). Larva is described by Jackson (1977).	en	Orel, Oksana V. (2014): To systematics of the genus Saetheria Jackson (Diptera, Chironomidae) from the Russian Far East. Zootaxa 3802 (1): 65-74, DOI: 10.11646/zootaxa.3802.1.5
4F16166CFFF5FF9C4DFFA2C53A8DFCC1.taxon	discussion	Remarks. The adult male of S. tylus (Townes) from Primorye Territory is almost indistinguishable from the description of specimens of this species by other authors (Table 4). The species affiliation larvae of S. tylus described by Kobayashi (2007), is questionable, as the larvae have 5 - segmented antenna and it are not associated with the male. The larva of Chironominae genuinae № 9 described by Lipina (1926) from Oka River is somewhat different from larva of S. tylus by presence of a large tooth in middle of premandible and seta subdentalis a narrow blade-like. The larva of S. tylus has 3 large apical teeth and 1 small proximal tooth on premandible and seta subdentalis a wide blade-like (Jackson 1977).	en	Orel, Oksana V. (2014): To systematics of the genus Saetheria Jackson (Diptera, Chironomidae) from the Russian Far East. Zootaxa 3802 (1): 65-74, DOI: 10.11646/zootaxa.3802.1.5
4F16166CFFF5FF9C4DFFA2C53A8DFCC1.taxon	distribution	Distribution. Saetheria tylus (Townes) is distributed in the Holarctic Region (Saether & Spies 2004). This species is known from Japan (Kobayashi 2007). In Russian Far East it is recorded from Primorye Territory only.	en	Orel, Oksana V. (2014): To systematics of the genus Saetheria Jackson (Diptera, Chironomidae) from the Russian Far East. Zootaxa 3802 (1): 65-74, DOI: 10.11646/zootaxa.3802.1.5
