taxonID	type	description	language	source
4F248367DB5D5C319ADBFEA636B98710.taxon	diagnosis	Diagnosis. Dorsal shield bearing 29 pairs of long and pointed setae, z 1 absent; anterior margin of ventri-anal shield conspicuously concave, with three pairs of pre-anal setae, post-anal seta longer than other ventri-anal setae; anterolateral margins of ventrianal shield closely adjacent to relatively long and narrow metapodal shield; presternal shields absent; peritrematal shields fused with exopodal shields, their posterior ends reaching beyond coxae IV; soft integument between peritrematal shields and anterolateral margins of ventri-anal shield with one pair of small oval platelets; epigynal shield reduced in size, relatively small, posterior margin with a transverse row of four elongate platelets and a pair of small oval platelets; lateral and opisthogastric soft integument with 11 pairs of setae. Specimens examined. Holotype female, Trabzon Province, Demirkapı Village, 40 ° 34 ' 23 '' N, 40 ° 24 ' 06 '' E, alt. 1740 m, 17 June 2014, litter under Pinus sp. Paratypes: 16 females, same data as holotype.	en	Özbek, Hasan Hüseyin, Halliday, Bruce (2015): Two new species of Pachyseius Berlese (Acari: Pachylaelapidae) from Turkey, with a key to the world species. Zootaxa 3957 (1): 98-108, DOI: 10.11646/zootaxa.3957.1.8
4F248367DB5D5C319ADBFEA636B98710.taxon	description	Female. Dorsum (Fig. 1). Dorsal shield 485 – 535 long, 275 – 300 wide at widest point, oblong (length / width, 1.70 – 1.78), ornamented with net-like pattern, and bearing 29 pairs of mostly long and pointed needle-like setae, setae z 1 absent (Fig. 19); setae j 1 shortest, 10 – 12; j 2 34 – 50; j 3, s 4, s 5, and Z 1 70 – 75; z 6 60 – 65; J 5 22 – 28, other dorsal setae 35 – 60; a pair of lyrifissures near bases of j 2; slit-like gland pores gdj 3 present between seta j 2 and s 2; anterior ends of peritremes extending forward beyond level of gdj 3. Venter (Fig. 2). Pre-sternal shields absent. Sternal shield 135 – 145 long, 65 – 75 wide at level of coxae II; central region ornamented with punctate polygonal lines; carrying three pairs of pointed needle-like setae and two pairs of lyrifissures, edges of shield strongly sclerotised. Length of sternal setae st 1 - 3 20 – 28. Metasternal shields small, each bearing a metasternal seta (18 – 22) and a poroid. Epigynal shield 58 – 65 long, 70 – 85 wide at widest point, with punctate ornamentation, and a row of four elongate platelets and a pair of small oval platelets along its posterior margin; genital pores situated outside postero-lateral margins of shield, genital setae length 18 – 22. Ventri-anal shield with a concave anterior margin (Fig. 20) 182 – 210 long, 182 – 200 wide at widest point (length / width, 0.95 – 1.07), with polygonal ornamentation, bearing three pairs of pointed pre-anal setae (Jv 1 and Jv 2, 20 – 25; Jv 3 shorter, 14 – 16), a pair of longer para-anal setae (28 – 30), and a long postanal seta (30 – 38). Peritrematal shields fused with exopodal shields, with four pairs of small post-stigmatal pores, their posterior ends rounded, enclosing posterior margins of coxae IV and extending medially almost to genital setae. Opisthogastric soft integument between peritrematal shields and anterolateral margins of ventri-anal shield with one pair of oval platelets. Metapodal shields elongated (60 – 62 long) and very close to anterolateral margins of ventri-anal shield. Lateral and opisthogastric soft integument bearing 11 pairs of setae (15 – 22). Gnathosoma (Fig. 3). Corniculi short and horn-like; hypostomal seta h 3 longest (45 – 52), h 2 shortest (10 – 14), and h 1 23 – 27 long; pc 15 – 19 long; deutosternal groove with five rows of denticles and two smooth transverse lines (Fig. 21). Palp with 3 - tined apotele. Anterior and lateral margins of epistome denticulate (Fig. 4). Cheliceral digits slender, movable digit about 45 long, with two subdistal teeth and an terminal hook, fixed digit about 40 long (to dorsal seta), medially edentate, terminally with bidentate hook; pilus dentilis minute; arthrodial brush short (Fig. 5). Spermathecal apparatus. Spermathecal tubes not visible in the available specimens. Legs. Chaetotaxy: Leg I. Coxa 0 0 / 2 0, trochanter 1 1 / 3 1, femur 2 5 / 4 2, genu 2 5 / 3 2, tibia 2 5 / 3 2. Leg II. Coxa 0 0 / 2 0, trochanter 1 0 / 3 1, femur 2 5 / 3 1, genu 2 5 / 2 2, tibia 2 4 / 2 2, tarsus 3 7 / 5 3. Leg III. Coxa 0 0 / 2 0, trochanter 1 0 / 3 1, femur 1 3 / 1 1, genu 2 4 / 1 1, tibia 1 3 / 2 1, tarsus 3 7 / 5 3. Leg IV. Coxa 0 0 / 1 0, trochanter 1 0 / 3 1, femur 1 3 / 1 1, genu 2 4 / 1 1, tibia 1 3 / 2 1, tarsus 3 7 / 5 3. Tarsus II with one large spur-like seta (Fig. 6).	en	Özbek, Hasan Hüseyin, Halliday, Bruce (2015): Two new species of Pachyseius Berlese (Acari: Pachylaelapidae) from Turkey, with a key to the world species. Zootaxa 3957 (1): 98-108, DOI: 10.11646/zootaxa.3957.1.8
4F248367DB5D5C319ADBFEA636B98710.taxon	etymology	Etymology. The name destitutus, lacking, refers to the loss of dorsal shield seta z 1 in this species. Notes. The new species may be easily distinguished from the other congeners by the dorsal shield with 29 pairs of setae, z 1 absent, and the peritrematal shields fused with all the exopodal shields. The only other species of Pachyseius that lacks seta z 1 is P. cavernicola (we have examined the holotype of P. cavernicola, and confirm that z 1 setae are absent), but these two species are clearly distinguished by the number of pre-anal setae (two pairs in P. cavernicola, three in P. destitutus), the different length and form of the dorsal shield setae (spatulate in P. cavernicola, pointed in P. destitutus), the shape of the epistome (broadly arched in P. cavernicola, narrower in P. destitutus), and the number of spur-like seta on tarsus II (two in P. cavernicola, one in P. destitutus). The collection of 17 females and no males, and the apparent absence of insemination structures in the females, suggests that P. destitutus may be thelytokous, but that is unconfirmed.	en	Özbek, Hasan Hüseyin, Halliday, Bruce (2015): Two new species of Pachyseius Berlese (Acari: Pachylaelapidae) from Turkey, with a key to the world species. Zootaxa 3957 (1): 98-108, DOI: 10.11646/zootaxa.3957.1.8
4F248367DB5F5C349ADBFBAD3788821D.taxon	diagnosis	Diagnosis. In female, peritrematal shields fused with exopodal shields; ventri-anal shield bearing four pairs of preanal setae; pre-sternal shields absent; area between peritrematal shields and anterolateral margins of ventri-anal shield with two pairs of oval platelets, lateral and opisthogastric soft integument with seven pairs of setae. In male, holoventral shield present, opisthogastric region of shield with four pairs of ventral setae; spermatodactyl 90 – 110 long, and distally narrowed. Tarsus II with two spur-like setae. Specimens examined. Holotype female, Gümüşhane Province, Çıkrıkdüzü Wold, 40 ° 39 ' 58 '' N, 38 ° 59 ' 50 '' E, alt. 1990 m, 12 October 2013, litter under Abies sp. Paratypes: 2 females, same data as holotype; 1 female and 6 males; Gümüşhane Province, Kürtün Town, Örümcek Forest, 40 ° 39 ' 48 '' N, 39 ° 01 ' 46 '' E, alt. 1455 m, 1 June 2009, litter under Picea sp.; 1 female, Gümüşhane Province, Kürtün Town, Kızılali Wold, 40 ° 48 ' 17 '' N, 39 ° 04 ' 04 '' E, alt. 1555 m, 13 October 2013, litter under Rubus sp.	en	Özbek, Hasan Hüseyin, Halliday, Bruce (2015): Two new species of Pachyseius Berlese (Acari: Pachylaelapidae) from Turkey, with a key to the world species. Zootaxa 3957 (1): 98-108, DOI: 10.11646/zootaxa.3957.1.8
4F248367DB5F5C349ADBFBAD3788821D.taxon	description	Female. Dorsum (Fig. 7). Dorsal shield 600 – 676 long, 310 – 368 wide at widest point, oblong (length / width, 1.71 – 1.94), ornamented with reticulate pattern, carrying 30 pairs of setae, all smooth and pointed; setae z 1 shortest (8 – 10); j 1 15 – 20; z 2, s 2, r 3 and J 5 distinctly longer (30 – 40), other dorsal setae 20 – 30 long; a pair of lyrifissures present between j 2 and z 1; a pair of slit-like gland pores gdj 3 present between seta z 1 and z 2. Anterior ends of peritremes reaching or extending beyond level of z 1. Venter (Fig. 8). Pre-sternal shields absent. Sternal shield 140 – 152 long, 80 – 90 wide at level of coxae II, central region ornamented with punctate polygonal lines, carrying three pairs of smooth pointed setae and two pairs of lyrifissures, edges of shield strongly sclerotised. Length of setae st 1 – 3 25 – 36. Metasternal shields very small, each bearing a metasternal seta (20 – 25) and a poroid. Epigynal shield 85 – 95 long, 100 – 110 wide at widest point, with punctate ornamentation; one pair of elongate platelets on its posterior margin; genital pores situated outside shield; genital setae length 23 – 27. Ventri-anal shield (Fig. 22) longer than wide, 248 – 270 long, 208 – 233 wide at widest point (length / width 1.10 – 1.21), reticulated with punctate lines, and bearing four pairs of sub-equal pre-anal setae (20 – 30), a pair of longer para-anal setae (30 – 35) and a long post-anal seta (40 – 46). Peritrematal shields fused with exopodal shields, with two pairs of small post-stigmatal pores and truncated posterior margin, their posterior ends rounded, enclosing posterior margins of coxae IV and extending beyond level of genital setae. Soft integument between peritrematal shields and anterolateral margins of ventri-anal shield with two pairs of oval platelets. Metapodal shields elongate (60 – 65 long) and not well separated from anterolateral margins of ventri-anal shield. Lateral and opisthogastric soft integument bearing seven pairs of setae (15 – 25). Gnathosoma (Fig. 9). Subcapitulum with four pairs of setae, all smooth and needle-like, h 3 longest (48 – 52), h 2 shortest (12 – 15), setae pc 18 – 22 long, h 1 26 – 30 long. Deutosternum wide, with five long transverse rows of denticles. Corniculi horn-like. Palp with 3 - tined apotele. Anterior and lateral margins of tongue-shaped epistome denticulate (Fig. 10). Cheliceral digits slender, movable digit 54 – 58 long, with two subdistal teeth and a terminal hook; arthrodial brush short; fixed digit 40 – 45 long (to dorsal seta), with a single submedial tooth and bidentate terminal hook; pilus dentilis not seen (Fig. 11). Spermathecal apparatus. Spermathecal tubes long and fine, associated with coxae III (Fig. 12). Legs. Chaetotaxy: Leg I. Coxa 0 0 / 2 0, trochanter 1 1 / 3 1, femur 2 5 / 4 2, genu 2 5 / 3 2, tibia 2 5 / 3 2. Leg II. Coxa 0 0 / 2 0, trochanter 1 0 / 3 1, femur 2 5 / 3 1, genu 2 5 / 2 2, tibia 2 4 / 2 2, tarsus 3 7 / 5 3. Leg III. Coxa 0 0 / 2 0, trochanter 1 0 / 3 1, femur 1 3 / 1 1, genu 2 4 / 1 1, tibia 1 3 / 2 1, tarsus 3 7 / 5 3. Leg IV. Coxa 0 0 / 1 0, trochanter 1 0 / 3 1, femur 1 3 / 1 1, genu 2 4 / 1 1, tibia 1 3 / 2 1, tarsus 3 7 / 5 3. Tarsus II with two large spur like setae (Fig. 13). Male. Dorsum. Dorsal shield 510 – 558 long, 313 – 342 wide at widest point, oblong (length / width, 1.59 – 1.66), ornamented with reticulate pattern. Other features as in female. Venter (Fig. 14). Holoventral shield present, 423 – 464 long, 80 – 85 wide at level of coxae II; sternal region with punctate-reticulate ornamentation, and bearing five pairs of needle-like setae and three pairs of lyrifissures; opisthogastric portion of shield ornamented with punctate polygonal lines, with four pairs of setae (15 – 28); anal portion with a pair of para-anal setae (25 – 30) and longer post-anal seta (35 – 40). Peritrematal shields connected to holoventral shield. Other features as in female. Gnathosoma. Epistome triangular, with denticulate anterior margin (Fig. 15). Movable digit of chelicera 60 – 65 long, with a larger subterminal tooth and bidentate terminal hook; spermatodactyl directed posteriorly, 90 – 110 long, and distally narrowed; fixed digit about 46 long (to dorsal seta), slender, edentate and with a minute pilus dentilis; arthrodial brush short (Fig. 16). Legs. Femur II with well-developed ventral spur, genu II and tibia II each with smaller ventral spur (Fig. 17); tarsus II with two spur-like dorsal setae (Fig. 18).	en	Özbek, Hasan Hüseyin, Halliday, Bruce (2015): Two new species of Pachyseius Berlese (Acari: Pachylaelapidae) from Turkey, with a key to the world species. Zootaxa 3957 (1): 98-108, DOI: 10.11646/zootaxa.3957.1.8
4F248367DB5F5C349ADBFBAD3788821D.taxon	etymology	Etymology. The name quadrigeminus, four pairs, refers to the presence of four pairs of setae on the ventri-anal shield, in addition to the circumanal setae. Notes. The new species is easily distinguishable from the other congeners by the ventri-anal shield with four pairs of pre-anal setae, peritrematal shields fused with exopodal shields, and the male with a holoventral shield. The ventri-anal shield in Pachyseius usually has two or three pairs of pre-anal setae. Only P. sinicus has four pairs, but according to Özbek & Halliday (2014), two of these pairs are on the extreme lateral edges of the shield, where their positions may show intra-specific or artificial variation. All of our females of P. quadrigeminus sp. nov. carry four pairs of setae, and they are well inside the lateral edges of the shield.	en	Özbek, Hasan Hüseyin, Halliday, Bruce (2015): Two new species of Pachyseius Berlese (Acari: Pachylaelapidae) from Turkey, with a key to the world species. Zootaxa 3957 (1): 98-108, DOI: 10.11646/zootaxa.3957.1.8
