taxonID	type	description	language	source
645A64892988521F87ABCADEBC6BAFDC.taxon	description	Figs 1 A-C, 2 A, B	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
645A64892988521F87ABCADEBC6BAFDC.taxon	etymology	Etymology. The specific epithet is a noun in apposition, referring to the apical ancestor of the Achaemenid dynasty of rulers of Persia.	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
645A64892988521F87ABCADEBC6BAFDC.taxon	diagnosis	Diagnosis. The new species differs from its congeners occurring in the region by the very long receptacle (Re), longer than the posterior margin of the dorsal arch (Da) (vs. shorter).	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
645A64892988521F87ABCADEBC6BAFDC.taxon	description	Description. Female. Habitus as in Fig. 1 A-C. Total length 8.13. Carapace 3.28 long, 2.58 wide. Eye diameters: AME 0.12, PME 0.12, PLE 0.14. Carapace, sternum, chelicerae, labium, and maxillae reddish. Legs pale orange. Abdomen pale cream-coloured, without any pattern. Spinnerets uniformly pale cream-coloured. Measurements of legs: I: 8.77 (2.64, 1.56, 2.21, 1.78, 0.58), II: 8.49 (2.42, 1.44, 2.02, 2.08, 0.53), III: 6.54 (1.87, 0.99, 1.30, 1.92, 0.46), IV: 8.99 (2.51, 1.32, 2.07, 2.47, 0.62). Spination: I: Fe: 3 pl. II: Fe: 1 pl. III: Fe: 1 pl; Ti: 4 pl, 1 rl, 6 v; Mt: 3 pl, 2 rl, 6 v. IV: Fe: 9 d, 1 rl; Ti: 4 pl, 6 v; Mt: 3 pl, 2 rl, 5 v. Endogyne as in Fig. 2 A, B; length / width ratio ca. 2.6; receptacle slightly arched, 5 x longer than wide, anterior angles (Aa) rounded, not long; dorsal arch (Da) trapezoidal; transverse bar (Tb) as wide as receptacle, lateral edges (Le) directed posterolaterally; posterior diverticulum (Pd) hexagonal. Male. Unknown.	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
645A64892988521F87ABCADEBC6BAFDC.taxon	distribution	Distribution. Known only from the type locality in Fars Province, southern Iran (Fig. 35).	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
D067C2D80FD45FE7AE4798F1868E023C.taxon	description	Figs 3, 4 A-D	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
D067C2D80FD45FE7AE4798F1868E023C.taxon	etymology	Etymology. The specific epithet is a noun in apposition, referring to an Iranian tribe primarily inhabiting Chaharmahal & Bakhtiari, Khuzestan, Lorestan, Bushehr, and Isfahan provinces.	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
D067C2D80FD45FE7AE4798F1868E023C.taxon	diagnosis	Diagnosis. This species can be distinguished from other species of the Dysdera aculeata group occurring in the region by having a wider psembolus (i. e., 1.5 x wider than the tegulum).	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
D067C2D80FD45FE7AE4798F1868E023C.taxon	description	Description. Male. Habitus as in Fig. 3. Total length 5.97. Carapace 2.55 long, 1.97 wide. Eye diameters: AME 0.10, PME 0.08, PLE 0.11. Carapace, sternum, chelicerae, labium, and maxillae reddish brown. Legs orange. Abdomen cream-coloured, without any pattern. Spinnerets uniformly cream-coloured. Measurements of legs: I: 7.40 (2.07, 1.26, 1.80, 1.82, 0.45), II: 6.68 (1.83, 1.17, 1.61, 1.61, 0.46), III: 5.02 (1.42, 0.81, 0.98, 1.40, 0.41), IV: 6.51 (1.87, 0.97, 1.43, 1.77, 0.47). Spination: I: Fe: 4 pl. II: Fe: 3 pl. III: Fe: 3 pl, 1 rl; Ti: 4 pl, 2 rl, 5 v; Mt: 6 pl, 2 rl, 2 v. IV: Fe: 7 d, 1 pl; Ti: 5 pl, 2 rl, 5 v; Mt: 5 pl, 2 rl, 5 v. Palp as in Fig. 4 A-D; bulb ca. 2 x longer than wide; tegulum bell-shaped, 1.2 x longer than wide; psembolus 1.5 x longer than tegulum; median crest (Mc) rounded, ca. 2.2 x shorter than length of psembolus, ca. 4 x wider than high; posterior apophysis (Ap) very large, rounded; incision between tegulum and psembolus absent; retrolateral crest (Rc) gradually rounded. Female. Unknown.	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
D067C2D80FD45FE7AE4798F1868E023C.taxon	distribution	Distribution. Known only from the type locality in Chaharmahal & Bakhtiari Province, southwestern Iran (Fig. 35).	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
45DFBDC3F6665D64AAF6CC8B4DE06615.taxon	description	Figs 25 A-C, 26 A-D	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
45DFBDC3F6665D64AAF6CC8B4DE06615.taxon	etymology	Etymology. The specific epithet is an adjective, referring to the type locality of the species.	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
45DFBDC3F6665D64AAF6CC8B4DE06615.taxon	diagnosis	Diagnosis. The male of the new species is most similar to that of D. concinna L. Koch, 1878 from Azerbaijan, but differs by longer bulb (i. e., bulb length / tegulum width = 3.1, vs. 2.7), relatively shorter median crest, and longer stylus (cf. Fig. 26 A and Dunin 1982: fig. B). Dysdera damavandica sp. nov. is also similar to D. tapuria sp. nov. but differs by the median crest higher than wide (vs. wider than high), relatively longer stylus (cf. Fig. 26 A and Fig. 30 A) and posterior apophysis located at distal half of the bulb (vs. located at mid-part).	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
45DFBDC3F6665D64AAF6CC8B4DE06615.taxon	description	Description. Male. Habitus as in Fig. 25 A-C. Total length 10.40. Carapace 5.53 long, 4.12 wide. Eye diameters: AME 0.18, PME 0.19, PLE 0.23. Carapace, sternum, chelicerae, labium, and maxillae reddish brown. Legs dark orange. Abdomen greyish, without any pattern. Spinnerets uniformly greyish. Measurements of legs: I: 15.64 (4.51, 2.54, 4.12, 3.56, 0.91), II: 15.12 (4.16, 2.67, 3.73, 3.69, 0.87), III: 10.91 (3.21, 1.73, 2.22, 3.01, 0.74), IV: 13.75 (3.82, 2.02, 3.13, 3.94, 0.84). Spination: I: Fe: 2 pl. II: Fe: 2 pl. III: Fe: 1 pl; Ti: 4 pl, 2 rl, 5 v; Mt: 3 pl, 2 rl, 3 v. IV: Fe: 7 d; Ti: 2 pl, 3 rl, 5 v; Mt: 4 pl, 3 rl, 5 v. Palp as in Fig. 26 A-D; bulb ca. 3.1 x longer than wide; tegulum bell-shaped, almost as long as wide; psembolus 2.77 x longer than tegulum; median crest (Mc) triangular, ca. 6.65 x shorter than length of psembolus, higher than wide; posterior apophysis (Ap) rounded; incision between tegulum and psembolus present; retrolateral crest almost straight; stylus (St) straight, longer than median crest. Female. Unknown.	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
45DFBDC3F6665D64AAF6CC8B4DE06615.taxon	distribution	Distribution. Known only from the type locality in Mazandaran Province, northern Iran (Fig. 35).	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
DF2FB01EBB6C53BD92ABA606E75F0616.taxon	description	Figs 32 A-F, 33 A-F, 34 A, B	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
DF2FB01EBB6C53BD92ABA606E75F0616.taxon	etymology	Etymology. The specific epithet is an adjective, referring to the type locality of the species.	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
DF2FB01EBB6C53BD92ABA606E75F0616.taxon	diagnosis	Diagnosis. The male of the new species differs from those of its congeners by having weakly sclerotized bent stylus (St) (Fig. 32 C, F) (vs. stylus, if present, not bent). The female of this species is very similar to that of D. iranica sp. nov. by having a wide receptacle (i. e.,> 2 x wider than transverse bar), but differs by the relatively wider receptacle lacking an anterior triangular projection, and the presence of a median concavity on the transverse bar (vs. receptacle with triangular projection, transverse bar without concavity).	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
DF2FB01EBB6C53BD92ABA606E75F0616.taxon	description	Description. Male (Holotype). Habitus as in Fig. 32 A-C. Total length 5.49. Carapace 2.42 long, 1.88 wide. Eye diameters: AME 0.15, PME 0.13, PLE 0.11. Carapace, sternum, chelicerae, labium, and maxillae pale reddish. Legs orange. Abdomen cream-coloured, without any pattern. Spinnerets uniformly cream-coloured. Measurements of legs: I: 11.78 (3.23, 1.90, 2.87, 3.02, 0.76), II: 10.21 (2.91, 1.60, 2.52, 2.45, 0.73), III: 7.98 (2.31, 1.15, 1.65, 2.29, 0.58), IV: 10.30 (2.89, 1.44, 2.28, 2.96, 0.73). Spination: I: Fe: 2 pl. II: Fe: 1 pl. III: Fe: 1 pl; Ti: 3 pl, 2 v; Mt: 3 pl, 4 rl, 3 v. IV: Fe: 4 d; Pa: 1 pl; Ti: 3 pl, 1 rl, 4 v; Mt: 1 pl, 1 rl, 3 v. Palp as in Fig. 33 A-F; bulb ca. 2.3 x longer than wide; tegulum bell-shaped, almost as long as wide; psembolus as long as tegulum, length / width ratio ca. 1.25; median crest (Mc) triangular; apex with two triangular laminae (La); posterior apophysis (Ap) claw-shaped, accompanied by a triangular outgrowth (To) anteriorly; incision between tegulum and psembolus absent; stylus (St) bent, long, and weakly sclerotized. Female. Habitus as in Fig. 32 D-F. Total length 6.52. Carapace 2.06 long, 1.59 wide. Eye diameters: AME 0.11, PME 0.11, PLE 0.10. Colouration as in male. Measurements of legs: I: 7.71 (1.93, 1.37, 2.01, 1.88, 0.52), II: 7.31 (1.99, 1.26, 1.77, 1.74, 0.55), III: 5.84 (1.58, 0.92, 1.19, 1.65, 0.50), IV: 7.77 (2.19, 1.21, 1.79, 2.04, 0.54). Spination: I: Fe: 2 pl. II: Fe: 2 pl. III: Fe: 1 pl; Ti: 2 pl, 1 rl, 5 v; Mt: 5 pl, 2 rl. IV: Fe: 5 d, 1 rl; Pa: 1 rl, 1 v; Ti: 5 pl, 1 rl, 5 v; Mt: 4 pl, 2 rl, 3 v. Endogyne as in Fig. 34 A, B; length / width ratio ca. 2; receptacle (Re) inverted trapezoidal, anterior margin 1.4 x longer than posterior; dorsal arch (Da) trapezoidal, posterior margin 1.6 x longer than anterior margin, 1.25 x longer than receptacle; anterior margin of transverse bar (Tb) with median concavity, posterior margin arched; lateral edges (Le) small, approximately as long as wide; posterior diverticulum (Pd) bilobed.	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
DF2FB01EBB6C53BD92ABA606E75F0616.taxon	distribution	Distribution. Known only from the type locality in Hormozgan Province, southern Iran (Fig. 35).	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
245D23FCCEC85241BC6C79DB4629F9C8.taxon	description	Figs 5 A-C, 6 A, B	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
245D23FCCEC85241BC6C79DB4629F9C8.taxon	etymology	Etymology. The specific epithet is an adjective referring to Hormuz Island, from where the holotype was collected.	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
245D23FCCEC85241BC6C79DB4629F9C8.taxon	diagnosis	Diagnosis. The new species differs from all Dysdera species occurring in the region by the receptacle divided into two chambers (vs. undivided), and the indistinct dorsal arch (vs. distinct).	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
245D23FCCEC85241BC6C79DB4629F9C8.taxon	description	Description. Female. Habitus as in Fig. 5 A-C. Total length 8.11. Carapace 3.26 long, 2.56 wide. Eye diameters: AME 0.13, PME 0.13, PLE 0.14. Carapace, sternum, chelicerae, labium, and maxillae orangish. Legs pale orange. Abdomen pale cream-coloured, without any pattern. Spinnerets uniformly pale cream-coloured. Measurements of legs: I: 7.87 (2.19, 1.29, 2.01, 1.85, 0.53), II: 7.31 (1.99, 1.19, 1.77, 1.83, 0.53), III: 5.83 (1.61, 0.88, 1.13, 1.74, 0.47), IV: 7.27 (1.83, 1.05, 1.56, 2.22, 0.61). Spination: I: Fe: 2 pl. II: Fe: 3 pl. III: Fe: 3 d; Pa: 1 d, 1 pl; Ti: 5 pl, 2 rl, 6 v; Mt: 3 d, 6 pl, 2 rl. IV: Fe: 8 d; Pa: 1 pl; Ti: 5 pl, 3 rl, 6 v; Mt: 3 pl, 4 rl, 3 v. Endogyne as in Fig. 6 A, B; length / width ratio ca. 2.6; receptacle 2 x longer than wide, divided in two chambers, with an anterior median concavity; anterior angles located almost at mid-part of each chamber and directed anteriorly, approximately as long as wide; dorsal arch indistinct; transverse bar ca. 4.6 x longer than wide, mid-part similar to an inverted trapezoid, anterior part 1.3 x longer than posterior part; lateral edges directed postero-laterally, clearly separated from transverse bar by an incision; posterior diverticulum elongated horizontally. Male. Unknown.	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
245D23FCCEC85241BC6C79DB4629F9C8.taxon	distribution	Distribution. Known only from the type locality in Hormuz Island, the Persian Gulf (Fig. 35).	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
D1F011A443AA549C96658F977D9D0D91.taxon	description	Figs 7 A-F, 8 A-D, 9 A, B	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
D1F011A443AA549C96658F977D9D0D91.taxon	etymology	Etymology. The specific epithet is an adjective and refers to the country from where the specimens of the new species were collected.	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
D1F011A443AA549C96658F977D9D0D91.taxon	diagnosis	Diagnosis. The male of the new species is somewhat similar to that of D. arabica Deeleman-Reinhold, 1988 from Oman (cf. Fig. 8 A-D and Deeleman-Reinhold and Deeleman 1988: figs 309 - 310), but differs by the small, claw-like posterior apophysis (vs. broad and rounded) and keel-like median crest (vs. rounded). The male of D. iranica sp. nov. differs from those of its congeners occurring in Iran by the elongate keel-like median crest (vs. rounded or triangular). The female of D. iranica sp. nov. is most similar to that of D. tapuria sp. nov. by having a very wide receptacle (i. e.,> 2 x wider than the transverse bar), but differs by having a triangular extension (Te) in the anterior margin of the receptacle (vs. absent).	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
D1F011A443AA549C96658F977D9D0D91.taxon	description	Description. Male (Holotype). Habitus as in Fig. 7 D-F. Total length 7.32. Carapace 3.35 long, 2.63 wide. Eye diameters: AME 0.15, PME 0.15, PLE 0.15. Carapace, sternum, chelicerae, labium, and maxillae reddish. Legs yellowish orange. Abdomen cream-coloured, without any pattern. Spinnerets uniformly dark yellowish. Measurements of legs: I: 11.4 (3.14, 1.92, 2.86, 2.84, 0.63), II: 10.26 (2.72, 1.78, 2.46, 2.68, 0.62), III: 7.61 (2.14, 1.17, 1.54, 2.13, 0.63), IV: 9.75 (2.69, 1.53, 2.12, 2.74, 0.67). Spination: I: Fe: 3 pl. II: Fe: 3 pl. III: Fe: 3 pl, 1 rl; Pa: 1 pl; Ti: 5 pl, 3 rl, 5 v; Mt: 6 pl, 2 rl, 2 v. IV: Fe: 7 d, 2 pl, 1 rl; Pa: 1 pl; Ti: 6 pl, 2 rl, 6 v; Mt: 3 pl, 2 rl, 5 v. Palp as in Fig. 8 A-D; bulb ca. 2.5 x longer than wide; tegulum bell-shaped, almost as long as wide; psembolus 1.4 x longer than tegulum; median crest rounded, ca. 2.5 x shorter than length of psembolus, ca. 2.5 x wider than high; posterior apophysis claw-shaped, 1.5 x longer than wide; incision between tegulum and psembolus absent; retrolateral crest gradually rounded. Female. Habitus as in Fig. 7 A-C. Total length 8.20. Carapace 3.03 long, 2.45 wide. Eye diameters: AME 0.14, PME 0.13, PLE 0.14. Colouration as in male. Measurements of legs: I: 8.45 (2.48, 1.44, 1.94, 2.01, 0.58), II: 7.79 (2.17, 1.35, 1.88, 1.91, 0.48), III: 6.25 (1.77, 1.05, 1.16, 1.71, 0.56), IV: 7.88 (2.28, 1.29, 1.72, 2.01, 0.58). Spination: I: Fe: 3 pl. II: Fe: 2 pl. III: Fe: 2 d, 3 pl, 3 rl; Ti: 4 pl, 2 rl, 5 v; Mt: 5 pl, 2 rl, 3 v. IV: Fe: 7 d, 1 rl; Ti: 3 pl, 2 rl, 6 v; Mt: 3 pl, 2 rl, 3 v. Endogyne as in Fig. 9 A, B; length / width ratio ca. 2.1; receptacle 4 x longer than wide, almost inverted trapezoidal; anterior angle large and triangular, with its base as wide as receptacle, directed antero-laterally; receptacle with median triangular extension (Te); dorsal arch trapezoidal, anterior corners rounded, posterior margin 1.6 x longer than anterior, anterior margin ca. 1.25 x longer than width of dorsal arch; transverse bar slightly arched, 1.5 x longer than receptacle; lateral edges with horizontal anterior margins; posterior diverticulum trapezoidal.	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
D1F011A443AA549C96658F977D9D0D91.taxon	distribution	Distribution. Known from the listed localities in Fars and Hormozgan provinces, south-central and southern Iran (Fig. 35).	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
CA37509DB3BD5DE4B503426AA9B08763.taxon	description	Figs 10 A-D, 11 A, B, 12 A-D, 14 A-D	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
CA37509DB3BD5DE4B503426AA9B08763.taxon	etymology	Etymology. The specific epithet is an adjective, referring to the type locality of the species.	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
CA37509DB3BD5DE4B503426AA9B08763.taxon	diagnosis	Diagnosis. The male of this species differs from those of the other species of the Dysdera aculeata group occurring in the region by the very long psembolus (i. e., length of psembolus / length of tegulum = 1.85 in the new species, vs. 1.6 or less in most other species), rounded arch-like ridge (Ar), presence of the notch of posterior apophysis (vs. absent), and median position of the posterior apophysis on the psembolus (vs. close to tegulum). Dysdera persica sp. nov. also bears a long psembolus (i. e., length of psembolus / length of tegulum = 2), but differs from D. isfahanica sp. nov. in the shape of the posterior apophysis. The female of this species can be recognized by its long anterolaterally stretched angles of the receptacle.	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
CA37509DB3BD5DE4B503426AA9B08763.taxon	description	Description. Male. Habitus as in Figs 10 A, C, 11 A, B. Total length 10.47. Carapace 4.15 long, 3.45 wide. Eye diameters: AME 0.17, PME 0.14, PLE 0.15. Carapace, sternum, chelicerae, labium, and maxillae orange. Legs yellowish. Abdomen pale beige, without any pattern. Spinnerets uniformly pale beige. Measurements of legs: I: 13.57 (3.77, 2.47, 3.23, 3.21, 0.89), II: 12.83 (3.47, 2.34, 2.96, 3.16, 0.90), III: 9.23 (2.60, 1.39, 1.95, 2.58, 0.71), IV: 12.28 (3.47, 1.91, 2.59, 3.43, 0.88). Spination: I: Fe: 1 - 2 pl. II: no spines. III: Ti: 4 pl, 2 rl, 4 v; Mt: 6 pl, 6 rl. IV: Fe: 6 d; Ti: 2 d, 2 pl, 2 rl, 2 v; Mt: 6 pl, 5 rl. Palp as in Fig. 12 A-D; bulb ca. 2.5 x longer than wide; tegulum bell-shaped, almost as long as wide; psembolus 1.8 x longer than tegulum; median crest rounded, ca. 3.4 x shorter than length of psembolus, ca. 2 x wider than high; posterior apophysis claw-like, 2.3 x longer than wide; incision between tegulum and psembolus absent; retrolateral crest forming right angle. Female. Habitus as in Fig. 10 B, D. Total length 9.45. Carapace 3.30 long, 2.55 wide. Eye diameters: AME 0.15, PME 0.14, PLE 0.12. Colouration as in male. Measurements of legs: I: 9.74 (2.76, 1.66, 2.31, 2.40, 0.61), II: 8.80 (2.52, 1.50, 2.08, 2.13, 0.57), III: 6.74 (1.94, 1.08, 1.28, 1.90, 0.54), IV: 8.68 (2.47, 1.31, 1.90, 2.46, 0.54). Spination: I: Fe: 3 pl. II: Fe: 4 pl. III: Fe: 5 d, 1 rl; Pa: 3 pl; Ti: 4 pl, 2 rl, 6 v; Mt: 3 pl, 2 rl, 6 v. IV: Fe: 8 - 11 d, 2 pl; Pa: 1 pl; Ti: 5 pl, 4 rl, 6 v; Mt: 2 pl, 3 rl, 6 v. Endogyne as in Fig. 14 A-D; length / width ratio ca. 2; receptacle ca. 5 x longer than wide, with long anterior angles (i. e., longer than width of their bases), anterior margin almost straight; dorsal arch trapezoidal; transverse bar straight, 1.6 x longer than receptacle; anterior margin of lateral edge inclined, lateral edge approximately as wide as receptacle; posterior diverticulum almost rectangular, its posterior edge rounded.	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
CA37509DB3BD5DE4B503426AA9B08763.taxon	distribution	Distribution. Known only from the type locality in Isfahan Province, central Iran (Fig. 35).	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
E880632E356B528F934A5208AE2CF592.taxon	diagnosis	Diagnosis. Dysdera can be diagnosed from other dysderine genera by the interdistance of PLE and PME less than half of their diameter, three or four cheliceral teeth in one series, punctiform (= highly reduced) fovea, and femur I at least twice as long as coxa I. The bulb is cylindrical, bearing a broad posterior apophysis and a distal psembolus. The endogyne is composed by an anterior diverticulum bearing a dorsal arch and a ventral arch, a transverse receptacle and a posterior diverticulum bearing a transverse bar (Deeleman-Reinhold and Deeleman 1988; Le Peru 2011).	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
83EF3B31E0E75DF794A2906ED9D5CE02.taxon	description	Figs 13 A, B, 14 E-G	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
83EF3B31E0E75DF794A2906ED9D5CE02.taxon	etymology	Etymology. The specific epithet is a noun in apposition, named after an Iranian language of the northwestern branch spoken by the Mazandarani people.	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
83EF3B31E0E75DF794A2906ED9D5CE02.taxon	diagnosis	Diagnosis. The new species is similar to D. isfahanica sp. nov., but differs by the arched anterior margin of receptacle (vs. almost straight), almost square-shaped dorsal arch (vs. distinctly trapezoidal), and shorter anterior angles (vs. longer, cf. Fig. 14 A-D, E-G).	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
83EF3B31E0E75DF794A2906ED9D5CE02.taxon	description	Description. Female (Holotype). Habitus as in Fig. 13 A, B. Total length 8.90. Carapace 4.25 long, 3.37 wide. Eye diameters: AME 0.21, PME 0.19, PLE 0.22. Carapace, sternum, chelicerae, labium, and maxillae orange. Legs yellowish. Abdomen pale beige, without any pattern. Spinnerets uniformly pale beige. Measurements of legs: I: 10.93 (3.48, 2.02, 2.50, 2.25, 0.68), II: 10.00 (2.95, 1.86, 2.31, 2.25, 0.63), III: 7.46 (2.38, 1.08, 1.44, 1.91, 0.65), IV: 9.92 (2.71, 1.50, 2.12, 2.88, 0.71). Spination: I, II: no spines. III: Fe: 1 d; Ti: 2 pl, 2 rl, numerous v spine-like setae; Mt: 2 pl, 4 rl, 5 v. IV: Fe: 2 d; Ti: 2 pl, 2 rl, numerous v spine-like setae; Mt: 3 pl, 2 rl, 6 v. Endogyne as in Fig. 14 E-G; length / width ratio ca. 2.2; receptacle with slightly arched anterior margin, ca. 5 x longer than wide, anterior angles slightly rounded; dorsal arch almost square-shaped, posterior margin 1.2 x longer than anterior margin; transverse bar straight, 1.7 x longer than receptacle; lateral edges broad, wider than receptacle; posterior diverticulum narrowing posteriorly. Male. Unknown.	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
83EF3B31E0E75DF794A2906ED9D5CE02.taxon	distribution	Distribution. Known only from the listed localities in Mazandaran Province, northern Iran (Fig. 35).	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
109EDCC1C8CE50918772C0B4B0A10F88.taxon	description	Figs 27 A-C, 28 A-D	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
109EDCC1C8CE50918772C0B4B0A10F88.taxon	etymology	Etymology. The specific epithet is a noun in apposition, referring to an ancient Iranian people who inhabited an area known as Media between western and northern Iran.	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
109EDCC1C8CE50918772C0B4B0A10F88.taxon	diagnosis	Diagnosis. The male of the new species is similar to that of D. granulata Kulczynski, 1897 from Italy and the Balkan Peninsula, but differs by the shape of the tegulum (i. e., almost as wide as long, vs. 1.5 x longer than wide), and by thinner psembolus (as wide as tegulum, vs. wider than tegulum). The male of D. medes sp. nov. differs from those of its congeners occurring in Iran by the very long median crest (i. e., longer than half of psembolus, vs. shorter), abrupt tip of psembolus in ventral and dorsal views (Fig. 28 C, D) (vs. not abrupt), and posterior apophysis with two teeth (vs. one).	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
109EDCC1C8CE50918772C0B4B0A10F88.taxon	description	Description. Male. Habitus as in Fig. 27 A-C. Total length 10.0. Carapace 4.19 long, 3.17 wide. Eye diameters: AME 0.14, PME 0.14, PLE 0.14. Carapace, sternum, chelicerae, labium, and maxillae reddish brown. Legs orange. Abdomen greyish, without any pattern. Spinnerets uniformly dark yellowish. Measurements of legs: I: 12.33 (3.54, 2.03, 2.89, 3.00, 0.87), II: 13.50 (3.80, 2.40, 3.12, 3.29, 0.89), III: 9.99 (2.95, 1.59, 1.86, 2.69, 0.90), IV: 12.31 (3.73, 1.80, 2.46, 3.30, 1.02). Spination: I, II: no spines. III: Ti: 4 pl, 2 rl, 5 v; Mt: 6 pl, 3 rl, 2 v. IV: Fe: 7 d; Ti: 7 pl, 4 rl, 8 v; Mt: 7 pl, 3 rl, 4 v. Palp as in Fig. 28 A-D; bulb ca. 2.8 x longer than wide; tegulum bell-shaped, almost as long as wide; psembolus 1.9 x longer than tegulum; median crest rounded, ca. 2.42 x shorter than length of psembolus, ca. 5.3 x wider than high; posterior apophysis claw-shaped, with 2 teeth (Fig. 28 B, D); incision between tegulum and psembolus absent; retrolateral crest almost straight. Female. Unknown.	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
109EDCC1C8CE50918772C0B4B0A10F88.taxon	distribution	Distribution. Known only from the type locality in Tehran Province, northern Iran (Fig. 35).	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
4E0DFB130C1E54019D88770C09E499B6.taxon	description	Figs 15 A-F, 16 A-D, 17 A, B	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
4E0DFB130C1E54019D88770C09E499B6.taxon	etymology	Etymology. The specific epithet is an adjective, referring to the historical region of the Middle East, located in eastern Mesopotamia, which is now Iran.	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
4E0DFB130C1E54019D88770C09E499B6.taxon	diagnosis	Diagnosis. The male of this species differs from those of the other species of the Dysdera aculeata group occurring in Iran by the extremely long bulb (especially psembolus, i. e., twice longer than tegulum), and by the tegulum with posterior margin 1.3 x longer than anterior margin (vs. equal or shorter in length). The female of D. persica sp. nov. differs from those of its congeners by having the broadest dorsal arch, bearing almost angled anterior corners (vs. rounded).	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
4E0DFB130C1E54019D88770C09E499B6.taxon	description	Description. Male (Holotype). Habitus as in Fig. 15 D-F. Total length 9.06. Carapace 4.19 long, 3.20 wide. Eye diameters: AME 0.14, PME 0.16, PLE 0.17. Carapace, sternum, chelicerae, labium, and maxillae reddish brown. Legs orange. Abdomen greyish, without any pattern. Spinnerets uniformly dark yellowish. Measurements of legs: I: 13.98 (3.96, 2.36, 3.43, 3.52, 0.71), II: 12.87 (3.60, 2.11, 3.07, 3.27, 0.82), III: 9.38 (2.74, 1.51, 1.84, 2.70, 0.59), IV: 12.37 (3.54, 1.95, 2.66, 3.40, 0.82). Spination: I: Fe: 2 pl. II: Fe: 1 pl. III: Fe: 1 pl; Ti: 5 pl, 3 rl, 4 v; Mt: 5 d, 9 pl, 5 rl, 3 v. IV: Fe: 4 d; Ti: 4 pl, 6 rl, 1 v; Mt: 5 d, 6 pl, 7 rl, 6 v. Ti III-IV with a row of thin rigid ventral setae. Palp as in Fig. 16 A-D; bulb ca. 2.3 x longer than wide; tegulum bell-shaped, almost as long as wide; psembolus 2 x longer than tegulum; median crest triangular, ca. 2 x shorter than length of psembolus, ca. 2.4 x wider than high; posterior apophysis broad; incision between tegulum and psembolus present; retrolateral crest gradually rounded. Female. Habitus as in Fig. 15 A-C. Total length 11.0. Carapace 4.25 long, 3.16 wide. Eye diameters: AME 0.17, PME 0.18, PLE 0.15. Carapace and chelicerae dark reddish violet; sternum, labium, and maxillae reddish brown. Leg reddish brown. Abdomen greyish, without any pattern. Spinnerets uniformly greyish. Measurements of legs: I: 11.02 (3.15, 1.98, 2.59, 2.51, 0.79), II: 9.61 (2.72, 1.76, 2.32, 2.14, 0.67), III: 8.07 (2.27, 1.33, 1.62, 2.14, 0.71), IV: 10.54 (3.02, 1.66, 2.15, 2.89, 0.82). Spination: I: Fe: 3 pl. II: Fe: 1 pl. III: Fe: 1 d; Ti: 4 pl, 4 rl; Mt: 12 pl, 5 rl, 4 v. IV: Fe: 4 d; Ti: 5 pl, 4 rl, 1 v; Mt: 14 pl, 9 rl, 3 v. Ti III-IV with a row of thin rigid ventral setae. Endogyne as in Fig. 17 A, B; length / width ratio ca. 1.9; receptacle 3.5 x longer than wide, 1.2 x wider than transverse bar; anterior angles rounded; dorsal arch trapezoidal; transverse bar straight, ca. 1.8 x longer than receptacle; lateral edges with almost horizontal anterior margins, as long as width of receptacle; posterior diverticulum rectangular.	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
4E0DFB130C1E54019D88770C09E499B6.taxon	distribution	Distribution. Known only from listed localities in Golestan and Mazandaran provinces, northern Iran (Fig. 35).	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
64C95D094759543D97BA74A53233F151.taxon	distribution	Distribution. Iran, Turkmenistan.	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
9671F65BF7635D5DBC744D187D1FB975.taxon	description	Figs 18 A-F, 19 A-D, 20 A, B	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
9671F65BF7635D5DBC744D187D1FB975.taxon	etymology	Etymology. The specific epithet is a noun in apposition, referring to an ancient tribe dwelling in the Iranian plateau.	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
9671F65BF7635D5DBC744D187D1FB975.taxon	diagnosis	Diagnosis. The male of this species differs from those of the other species of the Dysdera aculeata group occurring in Iran by the strong dorsal incision between tegulum and psembolus, and the posterior apophysis bent on right angle (vs. no or small incision, and posterior apophysis not bent on right angle); the most similar species is D. mikhailovi Fomichev & Marusik, 2021 from Tajikistan, from which the new species differs by having a dorsal incision between tegulum and psembolus, the parallel dorsal sides of tegulum and psembolus (vs. dorsal margin of psembolus inclined), and smaller retrolateral crest angled at distal 1 / 3 of psembolus (vs. larger and angled at mid-part). The female of D. sagartia sp. nov. differs from those of its congeners occurring in Iran by having an arched anterior margin of receptacle in combination with a lack of anterior angles (vs. species with arched receptacle have anterior angles), and the almost semiround dorsal arch (vs. trapezoidal).	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
9671F65BF7635D5DBC744D187D1FB975.taxon	description	Description. Male (Holotype). Habitus as in Fig. 18 D-F. Total length 8.38. Carapace 4.29 long, 3.17 wide. Eye diameters: AME 0.18, PME 0.15, PLE 0.15. Carapace, sternum, chelicerae, labium, and maxillae reddish. Legs yellowish orange. Abdomen cream-coloured, without any pattern. Spinnerets uniformly cream-coloured. Measurements of legs: I: 14.51 (3.99, 2.46, 3.61, 3.68, 0.77), II: 12.87 (3.41, 2.02, 3.19, 3.50, 0.75), III: 9.24 (2.75, 1.17, 2.04, 2.47, 0.81), IV: 11.84 (3.51, 1.62, 2.70, 3.13, 0.88). Spination: I: Fe: 3 pl. II: Fe: 3 pl. III: Fe: 1 d, 3 pl; Ti: 5 pl, 2 rl, 5 v; Mt: 6 pl, 4 rl, 2 v. IV: Fe: 8 d, 1 pl, 1 rl; Ti: 6 pl, 3 rl, 7 v; Mt: 6 pl, 2 rl, 5 v. Palp as in Fig. 19 A-D; bulb ca. 2.1 x longer than wide; tegulum bell-shaped, almost as long as wide; psembolus 1.44 x longer than tegulum; median crest rounded, ca. 2.3 x shorter than length of psembolus, ca. 3.1 x wider than high; posterior apophysis broad; incision between tegulum and psembolus present; retrolateral crest bent on obtuse, almost right, angle. Female. Habitus as in Fig. 18 A-C. Total length 11.7. Carapace 3.85 long, 2.91 wide. Eye diameters: AME 0.16, PME 0.14, PLE 0.12. Colouration as in male. Measurements of legs: I: 10.60 (2.87, 1.75, 2.62, 2.73, 0.63), II: 11.38 (3.24, 1.86, 3.04, 2.81, 0.43), III: 8.27 (2.46, 1.27, 1.60, 2.32, 0.62), IV: 10.86 (3.07, 1.50, 2.36, 3.16, 0.77). Spination: I: Fe: 3 pl. II: Fe: 3 pl. III: Fe: 1 d, 1 pl; Ti: 6 pl, 2 rl, 5 v; Mt: 2 d, 4 pl, 2 rl, 3 v. IV: Fe: 7 d; Ti: 2 pl, 2 rl, 8 v; Mt: 2 d, 4 pl, 5 v. Endogyne as in Fig. 20 A, B; length / width ratio ca. 2; receptacle slightly arched, ca. 4.7 x longer than wide, anterior angles indistinct; dorsal arch semi-oval, its posterior margin 1.5 x longer than anterior; transverse bar 1.5 x longer than receptacle and approximately as wide, slightly arched, lateral edges longer than wide; posterior diverticulum rounded.	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
9671F65BF7635D5DBC744D187D1FB975.taxon	distribution	Distribution. Known only from the listed localities in Tehran Province, northern Iran (Fig. 35).	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
5C3653107DE7552F82633A0D44AC1FB2.taxon	description	Figs 29 A-F, 30 A-D, 31 A-D	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
5C3653107DE7552F82633A0D44AC1FB2.taxon	etymology	Etymology. The specific epithet is a noun in apposition, referring to the term applied to a mountainous region located in the Caspian coast of northern Iran.	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
5C3653107DE7552F82633A0D44AC1FB2.taxon	diagnosis	Diagnosis. The male of the new species is most similar to that of D. concinna, but differs by longer bulb (i. e., bulb length / tegulum width = 3.1, vs. 2.7), median crest wider than high (vs. higher than wide), and shorter stylus (cf. Fig. 30 B and Dunin 1982: fig. B). The male of D. tapuria sp. nov. is also similar to that of D. damavandica sp. nov., but differs by the median crest wider than high (vs. higher than wide) and relatively shorter stylus (cf. Fig. 30 A and Fig. 26 A). The female of this species differs from those of its congeners occurring in the region by the very wide lateral edges of the receptacle (i. e., approximately half of the receptacle's width, vs. less than half).	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
5C3653107DE7552F82633A0D44AC1FB2.taxon	description	Description. Male (Holotype). Habitus as in Fig. 29 A-C. Total length 9.55. Carapace 3.76 long, 2.86 wide. Eye diameters: AME 0.14, PME 0.13, PLE 0.13. Carapace, sternum, chelicerae, labium, and maxillae reddish brown. Legs orange. Abdomen cream-coloured, without any pattern. Spinnerets uniformly dark yellowish. Measurements of legs: I: 14.04 (4.16, 2.14, 3.65, 3.24, 0.85), II: 12.72 (3.79, 2.08, 3.00, 3.11, 0.74), III: 10.01 (2.93, 1.44, 2.14, 2.77, 0.73), IV: 11.90 (3.72, 1.48, 2.73, 3.19, 0.78). Spination: I: Fe: 2 pl. II: Fe: 1 pl. III: Ti: 5 pl, 3 rl, 1 v; Mt: 5 pl, 2 rl, 3 v. IV: Fe: 8 d, 1 rl; Pa: 1 pl; Ti: 5 pl, 3 rl, 6 v; Mt: 6 pl, 2 rl, 5 v. Palp as in Fig. 30 A-D; bulb ca. 3.1 x longer than wide; tegulum bell-shaped, almost as long as wide; psembolus 1.8 x longer than tegulum; median crest triangular, ca. 7 x shorter than length of psembolus, ca. 2.5 x wider than high; posterior apophysis claw-shaped; incision between tegulum and psembolus present; retrolateral crest roundly bent in proximal part and almost straight distally; stylus straight, as long as median crest. Female. Habitus as in Fig. 29 D-F. Total length 18.1. Carapace 7.77 long, 6.01 wide. Eye diameters: AME 0.30, PME 0.30, PLE 0.34. Colouration as in male. Measurements of legs: I: 14.24 (3.99, 2.54, 3.62, 3.44, 0.65), II: 13.46 (4.31, 2.10, 3.22, 3.06, 0.77), III: 10.20 (2.96, 1.70, 1.97, 2.75, 0.82), IV: 13.08 (3.99, 1.72, 2.85, 3.62, 0.90). Spination: I: Fe: 2 pl. II: Fe: 2 pl. III: Fe: 1 pl; Ti: 2 pl, 2 rl, 4 v; Mt: 3 pl, 4 rl, 5 v. IV: Fe: 3 d; Ti: 2 pl, 2 rl, 6 v; Mt: 5 pl, 4 rl, 5 v. Endogyne as in Fig. 31 A-D; length / width ratio ca. 3; receptacle (Re) with shallow median concavity, ca. 4.5 x longer than wide, anterior angles (Aa) indistinct; dorsal arch (Da) trapezoidal, posterior margin ca. 1.6 x longer than anterior, anterior angles rounded; transverse bar (Tb) almost 2 x longer and slightly wider than receptacle; transverse bar's anterior margin straight, posterior margin arched; lateral edges (Le) very wide, directed latero-posteriorly; posterior diverticulum (Pd) rounded.	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
5C3653107DE7552F82633A0D44AC1FB2.taxon	distribution	Distribution. Known only from the listed localities in Mazandaran Province, northern Iran (Fig. 35).	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
AFD3A047BC9C5FB69377ABA45FFAB955.taxon	description	Figs 21 A-C, 22 A-D	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
AFD3A047BC9C5FB69377ABA45FFAB955.taxon	etymology	Etymology. The specific epithet is a noun in apposition, referring to an Old Persian word for the Gorgan region, meaning " land of wolves ".	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
AFD3A047BC9C5FB69377ABA45FFAB955.taxon	diagnosis	Diagnosis. The male of the new species is most similar to that of D. sagartia sp. nov., but differs by the more rounded median crest, the posterior apophysis not bent on right angle (cf. Fig. 22 A and Fig. 19 A), and the relatively longer psembolus (i. e., length of psembolus / length of tegulum = 1.66 in D. verkana sp. nov., vs. 1.44 in D. sagartia sp. nov.).	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
AFD3A047BC9C5FB69377ABA45FFAB955.taxon	description	Description. Male. Habitus as in Fig. 21 A-C. Total length 10.8. Carapace 5.65 long, 4.26 wide. Eye diameters: AME 0.24, PME 0.23, PLE 0.21. Carapace, sternum, chelicerae, labium, and maxillae reddish brown. Legs orange. Abdomen cream-coloured, without any pattern. Spinnerets uniformly dark yellowish. Measurements of legs: I: 13.04 (3.90, 2.23, 3.10, 3.06, 0.75), II: 11.76 (3.39, 1.98, 2.91, 2.71, 0.77), III: 8.56 (2.64, 1.26, 1.69, 2.33, 0.64), IV: 11.46 (3.32, 1.76, 2.48, 3.20, 0.70). Spination: I: Fe: 2 pl. II: Fe: 1 pl. III: Fe: 1 d, 2 pl; Ti: 4 pl, 2 rl, 5 v; Mt: 3 pl, 2 rl, 6 v. IV: Fe: 6 d, 1 pl; Ti: 4 pl, 4 rl, 5 v; Mt: 4 pl, 2 rl, 6 v. Palp as in Fig. 22 A-D; bulb ca. 2.2 x longer than wide; tegulum bell-shaped, almost as long as wide; psembolus 1.66 x longer than tegulum; median crest rounded, ca. 2.3 x shorter than length of psembolus, ca. 2.7 x wider than high; posterior apophysis broad; incision between tegulum and psembolus present; retrolateral crest roundly bent, forming right angle. Female. Unknown.	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
AFD3A047BC9C5FB69377ABA45FFAB955.taxon	distribution	Distribution. Known only from the type locality in Golestan Province, northern Iran (Fig. 35).	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
7ADCDF8AAA3B50F88C89E45A9071A4D2.taxon	description	Figs 23 A-C, 24 A-D	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
7ADCDF8AAA3B50F88C89E45A9071A4D2.taxon	etymology	Etymology. The new species is named after Xerxes I, the fourth King of Kings of the Achaemenid Empire, ruling from 486 to 465 BC; adjective.	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
7ADCDF8AAA3B50F88C89E45A9071A4D2.taxon	diagnosis	Diagnosis. The new species differs from all of its congeners occurring in the region by having a stylus (St), rounded median crest (Mc) and wide posterior apophysis (Ap); none of the other species has a rounded median crest, and those with a stylus, have a small posterior apophysis.	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
7ADCDF8AAA3B50F88C89E45A9071A4D2.taxon	description	Description. Male. Habitus as in Fig. 23 A-C. Total length 4.02. Carapace 2.14 long, 1.60 wide. Eye diameters: AME 0.11, PME 0.09, PLE 0.08. Carapace, sternum, chelicerae, labium, and maxillae reddish brown. Legs missing. Abdomen cream-coloured, without any pattern. Spinnerets uniformly cream-coloured. Palp as in Fig. 24 A-D; bulb ca. 2.3 x longer than wide; tegulum bell-shaped, almost as long as wide; psembolus 1.48 x longer than tegulum; median crest (Mc) rounded, ca. 3.45 x shorter than length of psembolus, ca. 2.3 x wider than high; posterior apophysis (Ap) broad and hook-shaped; incision between tegulum and psembolus absent; retrolateral crest (Rc) gradually rounded; stylus (St) membranous and shorter than median crest. Female. Unknown.	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
7ADCDF8AAA3B50F88C89E45A9071A4D2.taxon	distribution	Distribution. Known only from the type locality in Bushehr Province, southern Iran (Fig. 35).	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
42DB282680915FE7B12EBB80F0F2CAE9.taxon	diagnosis	Diagnosis. This subfamily can be diagnosed from other dysderids by the edge of sternum-labium joint ca. 2.5 - 3 x longer than the edge of the maxilla-sternum joint, all tarsi bearing claw tufts, posterior metatarsi bearing scopulae, and the spineless anterior tibiae and metatarsi. Furthermore, the bulb of dysderines does not bear a free embolus (with the exception of Harpactocrates Simon, 1914), and the posterior diverticulum of endogyne is large and wide (Deeleman-Reinhold and Deeleman 1988; Le Peru 2011; Kunt et al. 2019).	en	Zamani, Alireza, Marusik, Yuri M., Szűts, Tamas (2023): A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys 1146: 43-86, DOI: http://dx.doi.org/10.3897/zookeys.1146.97517, URL: http://dx.doi.org/10.3897/zookeys.1146.97517
