identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
4D4087B5FFACCA06D9A6AF09FD11FE03.text	4D4087B5FFACCA06D9A6AF09FD11FE03.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amolops nidorbellus Biju & Mahony & Kamei 2010	<div><p>Amolops nidorbellus sp. nov.</p> <p>(Figs. 1–3, 6; Table 1)</p> <p>Holotype. ZSI A 10965, adult male, collected by SDB and RGK on 27 June 2007 from <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=94.05862&amp;materialsCitation.latitude=25.639723" title="Search Plazi for locations around (long 94.05862/lat 25.639723)">Lorü</a>, 25°38’23’’N 94°03’31”E, 1575 m asl., Jotsoma village, Kohima district, Nagaland, northeast India (Fig. 1).</p> <p>Paratypes. ZSI A 10968, adult female, collected along with holotype; ZSI A 10969–10970, two adult females, and BNHS 5276, adult male, collected by RGK on 4 June 2009 from the holotype locality.</p> <p>Referred specimen. SDB 2009.335, adult male, collected along with paratype BNHS 5276.</p> <p>Diagnosis. Amolops nidorbellus sp. nov. is here placed in the genus Amolops based on its overall morphological similarity to A. viridimaculatus. Recently the systematic status of the latter species was confirmed phylogenetically (Cai et al. 2007). Amolops nidorbellus sp. nov. can be distinguished from known congeners by the following combination of characters: (1) large adult size (SVL male 76.4–82.3 mm; SVL female 85.4–98 mm); (2) head wider than long, HW:HL 102–107%; (3) tympanum distinct, TYD:EL 27– 34%; (4) digit I of hand without circummarginal groove on disk; (5) relative length of fingers I&lt;II&lt;IV&lt;III; (6) feet fully webbed; (7) externally visible vocal sacs absent; (8) dorsal and ventral spinules and asperities absent in both sexes; (9) dorsally brown with small irregularly arranged cobalt green spots; (10) ventrally pale brown with small white spots on throat, chest and anterior and lateral abdomen.</p> <p>Comparisons. Amolops nidorbellus sp. nov. differs from A. archotaphus, A. assamensis, A. chunganensis, A. compotrix, A. cucae, A. iriodes, A. larutensis, A. marmoratus, A. minutus, A. panhai, A. spinaspectoralis and A. vitreus by combination of its large adult male size, SVL&gt; 65 mm with absence of externally visible vocal sacs (vs. males SVL &lt;65 mm with externally visible vocal sacs); differs from A. daiyunensis, A. formosus, A. hongkongensis, A. ricketti, A. torrentis, and A. wuyiensis by the combination of its large adult male size, SVL&gt; 65 mm and absence of circummarginal groove on the first finger (vs. males SVL &lt;65 mm and presence of circummarginal groove on first finger); differs from A. aniqiaoensis, A. bellulus, A. chakratensis, A. cremnobatus, A. daorum, A. gerbilis, A. granulosus, A. jaunsari, A. jinjiangensis, A. kangtingensis, A. liangshanensis, A. longimanus, A. mengyangensis and A. monticola by its absence of dorsolateral folds or linear dorsolateral series of tubercles/pustules (vs. presence); differs from A. hainanensis, A. lifanensis, A. mantzorum and A. tuberodepressus by absence of tubercles on the dorsum or flanks (vs. presence); differs from A. loloensis by combination of its large adult male size, SVL&gt; 65 mm and absence of spinules (vs. adult male size, SVL &lt;65 mm and patches of spinules on the flanks, sides of head and posterior thigh); differs from A. himalayanus by absence of microspinules and externally visible vocal sacs (vs. microspinules arranged in patches at the rear of the mandible and often a large patch surrounding the vent region, externally visible vocal sacs present); from A. medogensis by its considerably smaller adult male size, SVL 76.5–82.3 mm (vs. 95 mm in A. medogensis).</p> <p>In general appearance and morphology, Amolops nidorbellus sp. nov. is most similar to A. caelumnoctis / A. splendissimus, A. kaulbacki and A. viridimaculatus. From A. caelumnoctis / A. splendissimus, this species differs by possessing oblique loreal region (vs. vertical for A. caelumnoctis and A. splendissimus), nostril closer to the eye than snout (vs. nostril located midway between tip of snout and eye for A. caelumnoctis and unknown for A. splendissimus), tympanic region and posterior angle of the jaw with dense tubercles (vs. smooth on A. caelumnoctis and A. splendissimus), and dorsal colouration is brown with clusters of small green irregularly shaped spots, many of which join to form small blotches (vs. purple/black with small circular yellow spots for A. caelumnoctis and brown/black with small irregular shaped yellow spots for A. splendissimus), throat, chest, anterior and lateral abdomen brown with distinct small white spots (vs. throat, chest and abdomen grey without distinct spots for A. caelumnoctis and A. splendissimus); it differs from A. kaulbacki by snout subelliptical dorsally (vs. rounded), smaller tympanum, TYD:EL 27.0–33.8% (vs. 40.0% on holotype), and fore and hind limbs not banded, small white spots on ventral aspect of body (vs. limbs distinctly banded, ventrum plain); from A. viridimaculatus by HW&gt;HL (vs. HW&lt;HL), fore and hind limbs not banded (vs. distinctly banded), dorsum brown with clusters of small green irregularly shaped spots, many of which join to form small blotches (vs. maroon red with large green blotches).</p> <p>Description of holotype. (all measurements in mm): adult male (SVL 76.4), body robust (Figs. 2A–B, 3A); head dorsally subelliptical (Figs. 2D–E), wider than long (HW 26.9, HL 25.8, HW:HL 104%), flat above; snout rounded and slightly protruding in profile (Fig. 2C), its length (SL 10.8) longer than the horizontal diameter of the eye (EL 7.4); canthus rostralis rounded, loreal region strongly concave, obtuse (Fig. 2C); interorbital space flat, interorbital distance (IUE 6.8) greater than width of the eye lids (UEW 6.5), and narrower than the internarial distance (IN 9.1); nostrils laterally positioned, vertically ovular with slightly raised rim anteriorly, slightly closer to the eye (EN 5.1) than to the snout (SN 5.4); pupil horizontal; tympanum fully exposed (TYD 2.0), circular, tympanum–eye distance (TYE 4.6); pineal ocellus barely visible; vomerine ridge distinct, obtuse, bearing 6 long teeth each, slightly closer to each other than choanae, positioned level to choanae which are oval and transverse.</p> <p>Arms moderately long, thick, forearm enlarged, length (FAL 17.8) shorter than the hand (HAL 26.0); relative length of fingers I&lt;II&lt;IV&lt;III (FIL 9.4, FIIL 12.5, FIIIL 19.8, FIVL 14.1); finger tips on II–IV dilated with wide oval disks (Fig. 2F), largest on digit III (FDIII 5.7, FWIII 1.8), finger I with distinct disk but only slightly dilated in relation to adjoining finger width, relative width of finger disks I&lt;II&lt;IV&lt;III, circummarginal grooves present on digits II–IV only; terminal phalange shape unknown; fingers without distinct lateral fringes, webbing on fingers absent; subarticular tubercles prominently domed, circular; prepollex and outer metacarpal tubercle oval, distinct and flat, inner metacarpal tubercle absent; supernumerary tubercle on the base of finger IV barely distinguishable, others indistinct (Fig. 2F).</p> <p>Hind limbs long, shank (SHL 45.8) longer than thigh (TL 40.7) and foot (FOL 40.6); toes long and thin, relative lengths I&lt;II&lt;III&lt;V&lt;IV; tips of all toes expanded with transversely oval disks, smaller than those of the fingers (TDIII 4.4), relative width of disks 1&lt;5&lt;4&lt;2&lt;3, all with circummarginal grooves; toes completely webbed (Fig. 2G); post axial groove on toe V extends from tip of toe to the basal subarticular tubercle; subarticular tubercles all present, prominently domed and circular, inner metatarsal tubercle prominent, oval and relatively long (IMT 4.4); tarsal glandular ridge, outer metatarsal and supernumerary tubercles all absent (Fig. 2G).</p> <p>Skin on dorsal surface of head, flanks, dorsal and ventral aspect of the body, throat and limbs smooth; area surrounding vent and adjoining posterior and ventral surface of the thighs granular, dorsal tubercles or pustular warts absent with exception of the temporal region posterior to the eye below the supratympanic fold, which is covered with enlarged glandular granules and warts; dorsolateral fold absent; supratympanic fold present but weakly developed; co-ossified skin absent; rictal and humeral glands absent (Fig. 2C); dorsal and ventral asperities and/or horny spinules absent.</p> <p>Colouration of holotype. In preservation (Fig. 2A): dorsal surface of head, body, limbs and upper flanks brown; green spots are faded to greyish blue; all ventral surface are faded to pale brown and small round light spots with dark rim only visible on throat and chest; webbing cream with fine grey speckling. In life (Fig. 3A): entire dorsal portion of the head, back, limbs and digits uniform brown with irregularly arranged clusters of small green spots; webbing yellow marbled with greyish brown; throat, chest and ventral surface of forelimb and thighs pale brown; abdomen, lower flanks, ventral surface of shank and tarsus yellowish brown, small white spots on the throat, chest, anterior and lateral abdomen; ventral surface of the hands and all digital disks, subarticular and inner metatarsal tubercles of the feet bluish grey; iris brownish black with dense yellowish green mottling.</p> <p>Condition of type series. All type specimens are in good condition and fully intact with the exception of ZSI A 10965 (holotype), BNHS 5276 and ZSI A 10969 (paratypes) which have a portion of the ventral thigh muscle removed for molecular analysis.</p> <p>Secondary sexual characters. Male: nuptial pad present, large, smooth (Fig. 2E), mostly on basal phalange, extending slightly onto the distal phalange, dorsally light, posteriorly and ventrally dark blue/grey (Fig. 3D); forearms considerably enlarged when compared to females; external vocal pouch absent. Female: posses a patch of small light tipped granules posterior to the supratympanic fold.</p> <p>Variation. Mensural differences are provided in Table 1, otherwise the paratypes and referred specimen agree in all general morphology and colouration (Fig. 3B) with the exception of the following details: weak supernumerary tubercles are variously distinct or indistinct on the base of all digits on hands; vomerine teeth range from 5–7 and vomerine ridges are subequally distant from each other and the choanae on ZSI A 10968 and SDB 2009.335; supratympanic fold is well developed on all paratypes and SDB 2009.335; relative toe length varies from III&lt;V to V &lt;III; pineal ocellus is not visible on ZSI A 10969 and SDB 2009.335, and only weakly visible on remaining paratypes. The extent of dark pigment on webbing varies; on all recently collected specimens (ZSI A 10969, 10970, BNHS 5276 and SDB 2009.335)––ventral brown with pale green spots on throat, chest and anterior and lateral abdomen, few smaller spots are scattered on the ventral thigh and tarsus (Figs. 3C–F).</p> <p>Natural history. This species is currently known only from the type locality, Lorü, Jotsoma village, about 5 km west of Kohima town (Fig. 1). All specimens were collected after heavy showers between 19.00–23.00 h, from rocks in a small cascading stream. This species is found sympatrically with Amolops kohimaensis sp. nov. The stream is bordered on both sides by steep vegetated banks within a patch of degraded secondary broadleaf semi-evergreen forest containing sparse undergrowth. Holotype and two paratypes were found sitting on vertical rock surfaces below a section of a multileveled cascade. Another paratype was found clinging to a moss covered rock 2 m above ground level on the stream bank. The otherwise conspicuous markings of this species provide surprisingly good camouflage with the moss-covered rocks on which it was found, indicating that this species may occupy only a very specialized habitat niche. All individuals were collected within an approximately one metre stretch of the stream. No further individuals were observed neither up nor downstream of this location along the ca. 100 meters of stream bank surveyed. No behaviour associated with reproduction was observed, nor were tadpoles apparent in the stream during the survey period. When handled these frogs immediately produce excessive secretions from the skin which is extremely pungent and causes considerable irritation when in contact with broken skin on the collector’s hands.</p> <p>Etymology. Derived from two Latin words–– nidor meaning ‘odor’ and bellus meaning ‘beauty’, referring to the beautiful appearance with strong odor of this species.</p> <p>Suggested common name. Spotted Stinky Torrent Frog.</p> <p>mm. H. holotype, P. paratype, RS. referred specimen.</p></div> 	https://treatment.plazi.org/id/4D4087B5FFACCA06D9A6AF09FD11FE03	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Biju, S. D.;Mahony, Stephen;Kamei, Rachunliu G.	Biju, S. D., Mahony, Stephen, Kamei, Rachunliu G. (2010): Description of two new species of torrent frog, Amolops Cope (Anura: Ranidae) from a degrading forest in the northeast Indian state of Nagaland. Zootaxa 2408 (1): 31-46, DOI: 10.11646/zootaxa.2408.1.2, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2408.1.2
4D4087B5FFA9CA0DD9A6A96AFC9DFD27.text	4D4087B5FFA9CA0DD9A6A96AFC9DFD27.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amolops kohimaensis Biju & Mahony & Kamei 2010	<div><p>Amolops kohimaensis sp. nov.</p> <p>(Figs. 1, 4–6; Table 2)</p> <p>Holotype. ZSI A 10972, adult male, collected by SDB and RGK on 27 June 2007 from <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=94.05862&amp;materialsCitation.latitude=25.639723" title="Search Plazi for locations around (long 94.05862/lat 25.639723)">Lorü</a>, 25°38’23’’N 94°03’31”E, 1575 m asl., Jotsoma village, Kohima district, Nagaland, northeast India (Fig. 1).</p> <p>Paratypes. BNHS 5282, ZSI A10966 and ZSI A 10971, three adult males collected along with holotype.</p> <p>Referred specimen. SDB 2007.129, one adult male, collected along with holotype.</p> <p>Diagnosis. Amolops kohimaensis sp. nov. is here placed in the genus Amolops, based on its overall morphological similarity to A. granulosus. Recently the systematic status of the latter species has been confirmed phylogenetically (Cai et al. 2007). Amolops kohimaensis sp. nov. is characterized by a combination of the following attributes: (1) SVL of adult males 42.8–48.6 mm; (2) head longer than wide, HW:HL 91– 95%; (3) entire tympanum distinct, TYD:EL 40.4–45.5%; (4) circummarginal groove present on all digits of hands and feet; (5) outer metatarsal tubercle absent; (6) fourth toe webbed to the distal subarticular tubercle; (7) external vocal pouches, dorsal spinules and asperities present in males; (8) male with nuptial pad on digit I covered with microgranules; (9) dorsolateral folds present; (10) absence of glandular tarsal ridge; (11) microgranules in a crescentic patch on ventral surface of jaw and second broad patch covering the entire chest, extending onto posterior throat, to the level of vocal pouches.</p> <p>Comparisons. Amolops kohimaensis sp. nov. differs from A. bellulus, A. caelumnoctis, A. cremnobatus, A. daiyunensis, A. formosus, A. granulosus, A. hainanensis, A. hongkongensis, A. jinjiangensis, A. kangtingensis, A. kaulbacki, A. lifanensis, A. loloensis, A. mantzorum, A. medogensis, A. nidorbellus, A. ricketti, A. splendissimus, A. torrentis, A. tuberodepressus, A. viridimaculatus and A. wuyiensis, by presence of externally visible vocal sacs on males (vs. absence); from A. aniqiaoensis, A. assamensis, A. himalayanus, A. jaunsari, A. marmoratus, A. panhai and A. spinaspectoralis, by fourth digit of the feet webbed to the distal subarticular tubercle (vs. feet fully webbed to disks or beyond the distal subarticular tubercle); from A. archotaphus, A. compotrix, A. cucae, A. larutensis and A. vitreus, by absence of a distinct outer metatarsal tubercle (vs. presence); differs from A. chunganensis, A. gerbilis, A. iriodes, A. liangshanensis, A. mengyangensis and A. minutus by presence (vs. absence) of extensive dorsal spinules on males in breeding condition; from A. monticola by presence (vs. absence) of dorsal spinules on males in breeding condition, larger tympanum, TYD:EL 40.4–45.5% (vs. TYD:EL 33.3–36.6%) and circummarginal groove on first finger weakly developed (vs. strongly developed); from A. daorum by larger adult male size, SVL 42.8–48.6 (vs. male SVL 32–38), dorsum brown (vs. green) and larger male tympanum, TYD:EL 40.4–45.5% (vs. male TYD:EL 29%); from A. chakratensis known only by the female holotype, by HW&lt;HL (vs. HL&lt;HW), absence of elongated glands on the lateral aspect of belly and shoulder (vs. presence), and brown flanks (vs. green flanks); differs from A. longimanus by EN&lt;SN (vs. SN&lt;EN) and a considerably larger tympanum, TYD:EL 40.4–45.5% (vs. TYD:EL ~25%).</p> <p>Description of holotype. (all measurements in mm): adult male (SVL 48.6), body habitus slender (Figs. 4A–B, 5); head dorsally subovoid, longer than wide (HW 15.2, HL 16.3, HW:HL 93%), flat above; snout rounded and strongly protruding in profile (Fig. 4C), its length (SL 6.9) longer than the horizontal diameter of the eye (EL 5.9); canthus rostralis distinct, rounded, loreal region concave, obtuse; interorbital space slightly convex, interorbital distance (IUE 4.1) slightly less than width of the eye lids (UEW 4.2) and narrower than the internarial distance (IN 5.0); nostrils laterally positioned, vertically ovular with raised rim anteriorly and posteriorly, slightly closer to the eye (EN 3.2) than to the snout (SN 4.0); pupil horizontal; tympanum fully exposed (TYD 2.6), circular with raised rim, tympanum–eye distance (TYE 1.3); pineal ocellus distinct; vomerine ridges moderately well developed, obtuse, bearing 5 short teeth each, slightly closer to each other than to choanae, positioned level to posterior border of choanae which are large, oval and slightly acute; tongue long, narrow and deeply emarginated.</p> <p>Arms moderately long, thick, forearm enlarged, length (FAL 12.5) shorter than the hand (HAL 14.7); relative length of fingers I&lt;II&lt;IV&lt;III (FIL 6.0, FIIL 7.0, FIIIL 10.7, FIVL 7.2); finger tips on II–IV dilated with wide oval disks (Fig. 4F), largest on digit III (FDIII 1.7 mm, FWIII 1.1), finger I with distinct disk but only slightly dilated in relation to adjoining finger width, relative width of finger disks I&lt;II=III=IV, circummarginal grooves present on all digits, but only weakly developed on digit I; terminal phalange shape unknown; fingers without distinct lateral fringes, webbing on fingers absent; subarticular tubercles prominently domed, circular, prepollex indistinct, outer and inner metacarpal tubercles oval, flat, barely visible; supernumerary tubercle on the base of all fingers, flat and barely visible.</p> <p>Hind limbs long, shank (SHL 30.2) longer than thigh (TL 26.1 mm) and foot (FOL 28.0); toes long and thin, relative lengths I&lt;II&lt;III&lt;V&lt;IV; tips of all toes expanded with transversely oval disks, subequal in width to those of the fingers (TDIII 1.6), relative width of toe disks 1&lt;5&lt;4&lt;2&lt;3, all with circummarginal grooves; toes moderately webbed (Fig. 4G); post axial groove on toe V extends from tip of toe to the basal subarticular tubercle, pre axial groove on digit I extends from the base of the disk to the inner metatarsal tubercle; subarticular tubercles all present, prominent, longitudinally ovoid, inner metatarsal tubercle prominent, oval and relatively long (IMT 2.1); tarsal glandular ridge, outer metatarsal and supernumerary tubercles all absent (Fig. 4G).</p> <p>Skin on almost entire dorsal and ventral surface of head, body, limbs and flanks smooth, with the following exceptions: two patches of microgranules with granule size subequal to those of the nuptial pad. One forms a crescent shaped patch following the curvature of the lower jaw, and the other covers the entire chest extending anteriorly to a rounded apex between the vocal pouches (Fig. 4E). A patch of dense tubercles on the proximal surface of the posterior thighs below the vent; dorsal and lateral tubercles or pustular warts absent; dorsolateral fold wide and continuous from posterior border of the eye to above the vent; supratympanic folds present, barely distinguishable; co-ossified skin absent; enlarged rictal glands absent replaced by a patch of small tubercles; humeral glands absent; conspicuous yellow/white horny spinules present (Fig. 4A, C, D), covering the dorsal aspect of the body from the vent anteriorly to the level of the insertion of the forelimbs, extending further anteriorly the length of dorsolateral ridges, a patch on the posterior upper eye lids, temporal area posterior to the supratympanic region, at the rear axis of the mandibles and between the tympanum and eye, densely covering granular area below the vent, extending in a narrow row along the posterior outer dorsal surface of the thigh, and entire outer surface of the tibia; those of the outer tarsus, lores and lips reduced to fine white asperities; asperities or spinules absent on the ventral aspects of the body.</p> <p>Colour of holotype. In preservation (Fig. 4): As in life but dorsal surface has darkened to dull brown; yellow orange colouration on the groin has completely faded to cream white. Entire ventral surface of body, and limbs cream white with some faint grey mottling on the chest and along the lower margin of the jaw; ventral surface of the hands and feet mottled cream and grey; ventral nuptial pad grey; ventral tarsus solid dark brown, posterior thighs marbled dark brown and cream; crescentic faded yellow marking posterior to the microgranular patch on the throat. In life (Fig. 5): Dorsal surface of head, body, limbs and lores primarily medium brown. Dorsal head and body with randomly arranged patches of dark brown speckling; dark brown stripe from snout tip, through nostril to eye, tympanic region and upper flanks dark brown, fading to light greyish brown on the lower flanks; groin with hint of yellowish orange. Dorsal surface of thighs with 5 dark brown transverse stripes with poorly defined borders, four on shank and two on tarsus; outer forelimb with extensive dark brown mottling and dark patch covering the proximal anterior humeral region posterior to the vocal patches. Dorsal surface of hands, feet, and lips with dark mottling; webbing plain light brown, opaque; nuptial pads pale pink above. Upper third of iris pale yellow, lower two thirds dark reddish brown. Ventral surface greyish white with some brown mottling on the chest and a crescent shaped blotch on the throat.</p> <p>Condition of type series. All specimens are fully intact, with the exception of ZSI A 10972 (holotype), which has a portion of the ventral thigh muscle removed for molecular analysis.</p> <p>Secondary sexual characters. Male: large nuptial pad on the proximal dorsal surface of first finger, covered in microgranules; forearms are enlarged relative to upper arm; a pair of externally visible vocal pouches at the rear axis of the jaws.</p> <p>Variation. Mensural differences are provided in Table 2. Considerable variation exists in several characters: relative finger lengths of ZSI A 10966 is I&lt;IV&lt;II&lt;III, all others including holotype and referred specimen have I&lt;II&lt;IV&lt;III; pineal ocellus is not visible on BNHS 5282, but is weakly visible on the remaining type series and referred specimen. Considerable variation is also apparent with regards to vomerine ridges and teeth: SDB 2007.129 has moderately well developed ridges and teeth 5/3 (right/left); BNHS 5282 and ZSI A 10966 both have weakly developed ridges, the former without vomerine teeth, the latter with 6 short teeth on each ridge; ZSI A 10971 has one weak vomerine ridge on the left side only, without teeth. Markings vary considerably between specimens: the dark dorsal blotches may be well defined e.g., SDB 2007.129, or reduced to fine scattered speckling as seen in the holotype; ZSI A 10971 has a considerable amount of dark grey mottling on the throat, chest and anterior abdomen and some fine speckling on ventral hind limbs; ventral markings on SDB 2007.129 are restricted to the throat and ventral shanks only.</p> <p>Natural history. This species is also known only from the type locality, Lorü, Jotsoma village, about 5 km west of Kohima town (Fig. 6). All specimens were collected from a small side pool of the same stream described above for Amolops nidorbellus sp. nov. between 20:00–24:00 h. The pool was formed by a diversion of the stream made by locals for trapping fish. Males were found calling from stones lining this pool, rather than the adjoining, fast flowing portion of the stream. It appears that males congregate for breeding, however no females or tadpoles were observed at this locality. The 2009 visit to this same stream, three weeks earlier in the season than the previous visit in 2007, resulted in no observations of this species. No further individuals were observed elsewhere along this stream during either visit.</p> <p>Etymology. Named after Kohima district, Nagaland, where the type series was collected.</p> <p>Suggested common name. Kohima Spiny Torrent Frog.</p></div> 	https://treatment.plazi.org/id/4D4087B5FFA9CA0DD9A6A96AFC9DFD27	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Biju, S. D.;Mahony, Stephen;Kamei, Rachunliu G.	Biju, S. D., Mahony, Stephen, Kamei, Rachunliu G. (2010): Description of two new species of torrent frog, Amolops Cope (Anura: Ranidae) from a degrading forest in the northeast Indian state of Nagaland. Zootaxa 2408 (1): 31-46, DOI: 10.11646/zootaxa.2408.1.2, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2408.1.2
