identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
4D54A843FFBC1A650694FB85EDEDFA7C.text	4D54A843FFBC1A650694FB85EDEDFA7C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acylophorus orientalis	<div><p>A. orientalis group</p><p>In this group only one terminal segment in the maxillary palpi is pubescent. The terminal segment is narrow and elongate and either symmetric or only slightly asymmetric (except A. makhoreae). The penultimate segment is widened toward the apex, but it is more or less parallel-sided at the apex. The head has two pairs of interocular setae. The micro-punctures on the head and pronotum are absent or sparse and localised. There is either no or only a narrow pigmented strip in front of the antennal insertion on the head, so that it looks as though the antennae are inserted right on the front margin (except A. makhoreae). The paramere is bilobed or, in one case ( A. tshuapensis), the lobes have fused to form one lobe. The apices of the styles of tergite IX are thickly covered in bristles so that their outline is invisible (Fig. 30).</p></div>	https://treatment.plazi.org/id/4D54A843FFBC1A650694FB85EDEDFA7C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lott, Derek A.	Lott, Derek A. (2010): The species of Acylophorus Nordmann (Coleoptera: Staphylinidae: Staphylininae) in continental sub-Saharan Africa. Zootaxa 2402: 1-51, DOI: 10.5281/zenodo.275907
4D54A843FFBC1A690694FA48ED90FF12.text	4D54A843FFBC1A690694FA48ED90FF12.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acylophorus orientalis Fauvel	<div><p>Acylophorus orientalis Fauvel</p><p>(Figs 1, 28–33, 51, 77, 109)</p><p>Acylophorus orientalis Fauvel, 1907: 48; Eichelbaum, 1913: 129; Bernhauer and Schubert, 1916: 439; Bernhauer and Scott, 1931: 588; Scheerpeltz, 1974: 25; Herman, 2001: 3034.</p><p>Acylophorus picipennis Bernhauer, 1929: 124; Scheerpeltz, 1933: 1467; Herman, 2001: 3035 (new synonym). Acylophorus marginalis Cameron, 1948: 42; Tottenham, 1961: 205; Herman, 2001: 3033 (new synonym).</p><p>Redescription. Length 6.5–7.5 mm. Body black or pitchy, often with iridescent abdomen. Pronotum, elytra and hind margins of abdominal tergites sometimes paler, the elytra in particular sometimes being partly red. Abdominal tergite and sternite VIII both with dark median band between pale base and apex. Legs, antennae and palpi largely pale, but middle segments of antennae sometimes infuscated. The terminal segment of the maxillary palpi is also noticeably darker in most specimens.</p><p>Head of average size for the genus (pronotum 1.75x wider than head), nearly as long as wide (&lt;1.1x wider than long) with temples rounded and not very evident (Fig. 1). Mostly shining with sparse micro-punctures more or less confined to front of head and inside eyes, but with dense short pubescence behind eyes. Two pairs of interocular setae arising from foveate punctures much closer to eyes than each other. A line of five postocular setae on each side plus an additional rather short seta out of line on the hind margin of the eye. Underside of head with long, but sparse pubescence, strongly depressed at base. Gular sutures continued to base of head, separate throughout their length, though proximate where they enter the depression at the base of head (Figs 28, 29). Mandibles with one medial tooth on right and two on left (Fig. 33). Maxillary palpi with terminal segment elongate and densely pubescent, slightly more rounded on outer margin than inner margin, but almost symmetric, longer than glabrous penultimate segment which is slightly elongate (Fig. 51). Antennae inserted right on front margin of head with no pigmented area in front of the insertion. First segment of antenna longer than next three. Segments I to VII elongate, IX to XI transverse (Fig. 77).</p><p>Pronotum weakly transverse (1.2x wider than long) with rounded sides, widest in basal half. Shining with no micro-punctures. One pair of dorsal setae. One pair of lateral setae. Marginal setae long. Elytra strongly transverse (1.6x wider than long) with long, close pubescence and dense asperate punctures. Hind margins supporting a fringe of bristles that are thicker, but not longer than the remaining pubescence. Abdominal tergites also with long pubescence and strong asperate punctures, but not quite as dense as on elytra and becoming sparser toward the hind margin of each tergite. Punctures on tergite VII as strong as on tergite III except for apical band which is free of punctures. Dorsal aspect of the genital segment as in fig. 30. Apex of male sternite IX slightly incised (Fig. 31). Female gonocoxites as in fig. 32.</p><p>Aedeagus narrow with the paramere divided into two closely aligned, but separate parallel lobes, each lobe being straight, narrow and strongly keeled on the dorsal surface so that they are laterally compressed (Fig. 109). The internal pegs are confusedly distributed along the margins of the apical half of each lobe, sometimes more densely packed at the mid-point and the apex giving the appearance of a gap. The median lobe is generally slightly expanded at the apex and normally only just surpasses the paramere.</p><p>Type material. Fauvel described the species from specimens collected by Alluaud between June 1903 and May 1904 in Kenya at Voï, Kisoumou, “baie de Kavirondo” and “steppe Masai près Nairobi”. Two specimens in the Fauvel collection can be recognised as belonging to the type series. One specimen carries conflicting labels referring to two separate localities, both conforming to data for the type locality given by Fauvel. The other carries a date label that conflicts with data given by Fauvel, possibly the result of a transcription error. They are conspecific, but one specimen is abnormally pale to the extent that it does not match the original description, so the darker specimen is here designated as the lectotype in order to fix the identity of the species. Lectotype Ƥ: “AFRIQUE OR le ANGLAISE; BAIE DE KAVIRONDO; (VICTORIA-NYANZA N.- E.); CH. ALLUAUD IX–X. 1903 / Voï 1–4; afr. or. angl. / orientalis Fvl. / R.I.Sc.N.B. 17.479; Coll. et det. A. Fauvel / Syntype / LECTOTYPE Acylophorus orientalis Fauvel Ƥ det. DA Lott, 2009” (IRSNB). Paralectotype Ƥ: “MUSEUM PARIS; AFRIQUE ORIENT. ANGL; KISOUMOU; (VICTORIA-NYANZA); CH ALLUAUD 1904 / Oct. / Coll. et det. A. Fauvel; Acylophorus orientalis Fauv.; R.I.Sc.N.B. 17.479 / Syntype / PARALECTOTYPE Acylophorus orientalis Fauvel Ƥ det. DA Lott, 2009” (IRSNB).</p><p>Acylophorus picipennis Bernhauer. The species was described from material collected by Schouteden at Luebo on 16th September 1921. Only one specimen conforming to these details has been located, here designated as the lectotype in order to fix the identity of the species. Lectotype 3: “ MUSÉE DU CONGO / Luebo; - IX – 1921; Schouteden / picipennis Brh. Typ / Acylophorus picipennis Brh. Typus. / Chicago NHMus M. Bernhauer Collection / LECTOTYPE Acylophorus picipennis Bernhauer 3 det. DA Lott, 2009 / Acylophorus orientalis Fauvel 3 det. DA Lott, 2009” (FMNH). Bernhauer separated A. picipennis from A. orientalis on colour, head shape, pronotum shape, punctation on the elytra and iridescence on the abdomen. In fact, all these characters in the type fall within the range of variation exhibited by A. orientalis . Consequently, A. picipennis is here synonymised with A. orientalis .</p><p>Acylophorus marginalis Cameron. The species was described from specimens collected in Chad. In his description Cameron indicated that the type was collected from the Rivière Gribingui in the Bassin du Chari, but he also referred to the locality Fort Archambault in the Moyen Chari. I have had the opportunity to examine a specimen from the second locality. Under article 72.4.6 of the nomenclatural code (ICZN, 1999), this specimen cannot be considered to be part of the type series, but it was part of the material studied by Cameron when defining the species and is likely to be conspecific with the type. Material studied 13: “ Paratype / MUSEUM PARIS; MOYEN CHARI; FORT ARCHAMBAULT; BOUNGOUL (BA-KARÉ); MISSION CHARI-TCHAD; DR J. LECORSE 1904 / JANVIER / M. Cameron Bequest B.M. 1955-147 / Acylophorus marginalis Cam. Paratype / Acylophorus orientalis Fauvel 3 det. DA Lott, 2009” (BMNH). The specimen is a typical A. orientalis . There is no reason to doubt that this identity applies to the type. Consequently, A. marginalis is here synonymised with A. orientalis .</p><p>Further material examined. ANGOLA: Cunene: R. Cunene, Rocadas, 19–22. ii.1972, 29 (BMNH). BURKINA FASO: Bam: L. Bam, 13 O 18’N 1 O 31’W, DA Lott, 28.x.2004, 433Ƥ (cLott); Tikare: Bge de Soukoundougou, 13 O 31’N 1 O 40’W, DA Lott, 24.x.2003, 433Ƥ (cLott); Bge de Bagara, 13 O 19’N 1 O 43’W, DA Lott, 28. x.2004, 14 (cLott); Bge de Tikare, 13 O 17’N 1 O 44’W, DA Lott, 29. x.2004, 1 Ƥ (cLott); Comoe: Bge de Bounouna, 10 O 39’N 4 O 44’W, DA Lott, 20.x.2004, 43 3Ƥ (cLott); Lerikoseni, 10 O 40’N 4 O 55’W, DA Lott, 20. x.2004, 23 2Ƥ (cLott); Karfiguela, 10 O 43’N 4 O 49’W, DA Lott, 22.x.2004, 33 1Ƥ (cLott); Siniéna, 10 O 33’N 4 O 46’W, DA Lott, 22. x.2004, 2 Ƥ (cLott); R. Comoe, 10 O 28’N 4 O 47’W, DA Lott, 22. x.2004, 13 (cLott); Ioba: clay pit by R. Mouhoun, 11 O 1’N 2 O 49’W, DA Lott, 15. x.2004, 5 (cLott); Djipologo, 10 O 56’N 3 O 6’W, DA Lott, 15. x.2004, 1 Ƥ (cLott); Gnagna: Botou, C Alluaud &amp; PA Chapuis, xii.1930 – iv.1931, 1 (FMNH); Naouri: Nazinga Park: Bge de Barka, 11 O 8’N 1 O 37’W, DA Lott, 29.x.2003 &amp; 11. x.2004, 22 (cLott); Bge de Bouzounga, 11 O 8’N 1 O 36’W, DA Lott, 30. x.2003, 1 Ƥ, (cLott); buffalo wallow, 11 O 9’N 1 O 34’W, DA Lott, 11.x.2004, 63 2Ƥ (cLott); Bge de Naguio, 11 O 8’N 1 O 35’W, DA Lott, 12. x.2004, 8 (cLott); Mare Nagale, 11 O 9’N 1 O 38’W, DA Lott, 12. x.2004, 13 2Ƥ (cLott); Bge de Kalieboulou, 11 O 12’N 1 O 30’W, DA Lott, 13. x.2004, 1 Ƥ (cLott); R. Akwazena, 11 O 9’N 1 O 36’W, DA Lott, 13. x.2004, 13 1Ƥ (cLott); Poni: near Tioyo, 10 O 37’N 3 O 14’W, DA Lott, 16. x.2004, 5 (cLott); near Houli, 10 O 16’N 3 O 14’W, DA Lott, 16. x.2004, 13 3Ƥ (cLott); 43 1Ƥ, Sissili: Sissili, 11 O 11’N 2 O 1’W, DA Lott, 14.x.2004, 43 1Ƥ (cBord, cJanak &amp; cLott); Bourbye, 11 O 1’N 2 O 29’W, DA Lott, 15. x.2004, 15 (cLott); R. Yali, 11 O 13’N 1 O 58’W, DA Lott, 14. x.2004, 7 (cJanak &amp; cLott). R.D. CONGO: Haut-Uelé: Moto, L. Burgeon, 1920, 1Ƥ (MRAC); Ituri: Mahagi, H Schouteden, 5. v.1925, 13 (MRAC); Kasai: Luebo, H Schouteden, 17. viii.1921, 13 (MRAC); Katanga: Katompe, P. Gérard, 1 – 15/6/1930, 1 (FMNH). CÔTE D’IVOIRE: Riv. Leraba, C Alluaud &amp; PA Chapuis, xii.1930 – iv.1931, 131Ƥ (FMNH). GHANA: near Addah, W Rossi, 22. iv.1984, 13 (teneral) (cBord). NIGERIA: Kano: Kano, ROS Clark, iii.1965, 13 (BMNH). SIERRA LEONE: Northern Province: Binkolo / Kamabesi, W Rossi, 13. i.1997, 13 (cBord). TANZANIA: Iringa: Parvagga, xii.1912, 2 (ZMHB); Lower Kihansi Valley, 24. xi.1912, 2 (ZMHB); Moero: Kiambi, P Gérard, vi–vii.1930, 2 Ƥ (MRAC); Ngerengere, Methner, xi.1912, 2 (FMNH); Ussure, Methner, 2. v.1911, 2 (FMNH); Wembare, Methner, 29. vi.1911, 1 (FMNH). ZAMBIA: Mwengwa, 13 O S 27 O 40’E, HC Dollman, 2.vi.1913 &amp; 16. vii.1914, 2 Ƥ (BMNH); Lukanga, HC Dollman, iv.1915, 13 (FMNH); Kashitu N. of Broken Hill, HC Dollman, 20. iii.1915, 1 Ƥ (FMNH).</p><p>Distribution and bionomics. A. orientalis is a widespread, relatively frequently collected species in continental Africa. Specimens have been seen from West Africa, Chad, East Africa north to Kenya, Congo, Zambia and Angola (Fig. 140). It has also been recorded in the literature from Sudan (Scheerpeltz, 1974) and Ethiopia (Bernhauer &amp; Scott, 1931). In Burkina Faso, it was found by standing water or slow-flowing rivers on wet mud and clay under litter or at least some vegetative cover, however sparse. It was never found on floating rafts of vegetation. The sparse data on specimen labels from elsewhere suggests a similar microhabitat in other parts of its range. In Burkina Faso it was found commonly in secondary habitats, especially irrigation reservoirs.</p><p>Discussion. The variability of this species has led to several manuscript names being applied to various specimens in museum collections and to two published synonyms. Except for one or two areas, the amount of material available from different regions is too sparse to allow a proper evaluation of local races and varieties. However, it appears that some colour varieties may be prominent in certain areas. Specimens from Angola and Zambia were larger than specimens from further north and had correspondingly larger aedeagi. It may be that southern populations deserve subspecific or even specific status, but the absence of specimens from the vast tracts of territory connecting to more northern populations prevents an assessment of any cline in characters, so it would be premature to erect a subspecies at this stage.</p></div>	https://treatment.plazi.org/id/4D54A843FFBC1A690694FA48ED90FF12	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lott, Derek A.	Lott, Derek A. (2010): The species of Acylophorus Nordmann (Coleoptera: Staphylinidae: Staphylininae) in continental sub-Saharan Africa. Zootaxa 2402: 1-51, DOI: 10.5281/zenodo.275907
4D54A843FFB01A690694FEFDEFD5F901.text	4D54A843FFB01A690694FEFDEFD5F901.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acylophorus delphinus Fauvel	<div><p>Acylophorus delphinus Fauvel</p><p>(Figs 104–107)</p><p>Acylophorus delphinus Fauvel, 1905: 181; Bernhauer and Schubert, 1916: 438; Jarrige, 1965: 141; Herman, 2001: 3028.</p><p>Redescription. Length 7 mm. Body pitchy with abdomen shiny, but not iridescent, pronotum, paler. Abdominal tergite and sternite VIII both with dark medial band between pale base and apex. Legs, palpi and antennal segment I pale, remaining antennal segments infuscated.</p><p>Head of average size (pronotum 1.75x wider than head), slightly transverse (1.1x wider than long) with temples very slightly angled (Fig. 104). Shining with micro-punctures extremely sparse, but with dense short pubescence behind eyes. Two pairs of interocular setae arising from foveate punctures much closer to eyes than each other. A line of five postocular setae on each side plus an additional long seta out of line near the hind border of the eye. Maxillary palpi with terminal segment pubescent, slightly asymmetric, elongate, but less so than A. orientalis, longer than glabrous penultimate segment which is not elongate (Fig. 105). Antennae inserted right on front margin of head with no pigmented area in front of the insertion. First segment of antenna longer than next three. Segments I to VIII elongate, X to XI transverse (Fig. 106).</p><p>Pronotum slightly transverse (1.1x wider than long) with rounded sides, widest in basal half. Shining with no micro-punctures. One pair of dorsal setae. One pair of lateral setae. Marginal setae largely abraded on the material available for study. Elytra transverse (1.6x wider than long) with long, close pubescence and dense asperate punctures. Fringe of bristles on hind margins thicker, but not longer than the general pubescence. Abdominal tergites also with long pubescence and strong asperate punctures, but not quite as dense as on elytra and becoming sparser toward the hind margin of each tergite. Punctures on tergite VII as strong as on tergite III.</p><p>Aedeagus narrow with the paramere divided into two parallel lobes, each lobe being straight, narrow and somewhat ridged dorsally, though much less so than in A. orientalis (Fig. 107). The internal pegs are confusedly distributed toward the apex. The median lobe is appreciably longer than the paramere.</p><p>Type material. Fauvel described the species from a single specimen collected at Abovombé in Madagascar. This specimen is the holotype by monotypy. Holotype 3: “Ambovombé Madag. / Acylophorus Nordmann / delphinus Fvl. / R.I.Sc.N.B. 17.479 Coll. et det. A. Fauvel / TYPE / HOLOTYPE Acylophorus delphinus Fauvel 3 det. DA Lott, 2009” (IRSNB). Jarrige (1965) wrongly supposed this specimen to be a female. When the type was examined in 2009, the genital segment was found to be pushed back into the abdomen with the dorsal sclerite and parameral surface of the aedeagus broken off, as though an attempt at dissecting the genitalia had already been made and aborted. Fortunately, the more significant features of the aedeagus were not damaged.</p><p>Distribution and bionomics. A. delphinus is only known from the type specimen taken in Madagascar. I have not seen it from continental Africa.</p><p>Comparative notes. This species is similar to A. orientalis, but can be distinguished by the pale terminal segment of the maxillary palpi, the longer additional postocular seta on the hind margin of the eye and the form of the aedeagus.</p></div>	https://treatment.plazi.org/id/4D54A843FFB01A690694FEFDEFD5F901	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lott, Derek A.	Lott, Derek A. (2010): The species of Acylophorus Nordmann (Coleoptera: Staphylinidae: Staphylininae) in continental sub-Saharan Africa. Zootaxa 2402: 1-51, DOI: 10.5281/zenodo.275907
4D54A843FFB01A680694F8EAEDD7F935.text	4D54A843FFB01A680694F8EAEDD7F935.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acylophorus nitens Lott	<div><p>Acylophorus nitens Lott, new species</p><p>(Figs 2, 34, 52, 78, 110)</p><p>Description. Length 7–8mm. Variable in colour. Head black. Pronotum orange-brown with disc sometimes diffusely infuscated, rarely all black. Elytra dark brown to black. In all the Zambian specimens seen, the pronotum is bright orange contrasting strongly with the black head and elytra. Abdomen strongly iridescent. Legs pale. Antennae dark with base of segment I and sometimes segment XI pale. The terminal segment of the maxillary palpi is darker than the penultimate segment.</p><p>Head of average size (pronotum 1.7x wider than head), more or less as long as wide with rounded temples and antennal insertion right on front margin (Fig. 2). Micro-punctures very sparse and confined to front of head and inside eyes. Dense short pubescence behind eyes. Two pairs of interocular setae arising from foveate punctures much closer to eyes than each other. A line of five postocular setae plus an additional short seta on hind margin of eye. Underside of head sparsely pubescent, strongly depressed at base with gular sutures separate, but very approximate toward base. Right mandible with three medial teeth; left mandible with two medial teeth (Fig. 34). Maxillary palpi with terminal segment elongate and densely pubescent, more or less symmetric, longer than glabrous penultimate segment which is not elongate (Fig. 52). First segment of antenna as long as next four. Segments I to VIII elongate, X transverse (Fig. 78).</p><p>Pronotum only slightly transverse (1.1x wider than long) with rounded sides and widest in basal half. Shining with no micro-punctures. One pair of dorsal setae. One pair of lateral setae. Marginal setae shorter than in A. orientalis . Elytra transverse (1.5x wider than long) with pubescence arising from relatively fine asperate punctures. Apical fringe of thick bristles about as long as the hairs on the rest of the elytra. Abdominal tergites with evenly spaced, relatively sparse asperate punctures and a marginal fringe of bristles similar to those on the elytra.</p><p>Paramere bilobed, lobes parallel and proximate, each lobe ridged, though not as strongly as in A. orientalis, pegs arranged along inner margins at apex (Fig. 110). Median lobe longer than paramere with rather narrow, spoon-shaped apex.</p><p>Type material. Holotype 3: “S. SUDAN: Bahr el Ghazal Prov. 8 O 30’N-28 O 30’E / C.E. Tottenham collection. B.B. 1974-587 / HOLOTYPE Acylophorus nitens sp. n. det. DA Lott, 2009” (BMNH); Paratypes 76: same data as holotype. It is almost certain that the type series was collected at Wunatong, near Wau, on 19.iii.1955, as these data were recorded on a label on a specimen in the same tray in the Tottenham collection.</p><p>This species was given the manuscript names, A. dollmani and A. nigeriae, by Bernhauer. Types were labelled accordingly, but he never published a description under either name.</p><p>Further material examined. R.D. CONGO: Nizi, Blukwa, A Collart, 25.i.1929, 131Ƥ (ISRNB). NIGERIA: Gombe: Matzoro Lakes, L Lloyd, i.1929, 13 (BMNH). SIERRA LEONE: Northern Province: Kambai / Binkolo, W Rossi, 16. ii.1993, 13 (cBord). SUDAN: Bahr el Ghazal: Wunatong, near Wau, 8 O 80’N 28 O 30’E 19. iii.1955, 10 (BMNH). ZAMBIA: Namwala, HC Dollman, iii.1913, 7 (BMNH &amp; FMNH). ZIMBABWE: Kutsaga near Harare Airport, W Rossi, 18. vi.1997, 2 Ƥ (cBord).</p><p>Distribution and bionomics. Widespread in continental Africa ranging from Zimbabwe to Sudan and Sierra Leone (Fig. 141). There are long series of specimens taken at two localities in the Sudd wetlands in the floodplain of the Bahr el Ghazal in southern Sudan. There are no ecological data.</p><p>Comparative notes. Similar to A. orientalis, but distinguished by the larger size, shorter marginal setae on the pronotum, the form of the mandibles and the distinctive aedeagus. In addition, the last segment of the maxillary palpi is more elongate as well as the medial segments of the antennae. See also notes under A. rossii .</p><p>Discussion. It is possible that particular geographic areas might be populated by distinctive colour forms. Specimens from Zambia were very distinctive by virtue of their contrasting orange pronota. This species and A. rossii appear to be closely related. The co-occurrence of the two species in a series from Sierra Leone, precludes treating the two taxa as subspecies or geographic races.</p></div>	https://treatment.plazi.org/id/4D54A843FFB01A680694F8EAEDD7F935	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lott, Derek A.	Lott, Derek A. (2010): The species of Acylophorus Nordmann (Coleoptera: Staphylinidae: Staphylininae) in continental sub-Saharan Africa. Zootaxa 2402: 1-51, DOI: 10.5281/zenodo.275907
4D54A843FFB11A6B0694F89FEC7AF93C.text	4D54A843FFB11A6B0694F89FEC7AF93C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acylophorus rossii Bordoni	<div><p>Acylophorus rossii Bordoni</p><p>(Figs 3, 35, 53, 79, 111)</p><p>Acylophorus rossii Bordoni, 1994: 309; Herman, 2001: 3036.</p><p>Diagnostic characters. Bordoni’s description contains illustrations of the aedeagus, head, maxillary palpi and posterior tarsi in addition to a textual description of morphological features. The following section contains additional diagnostic features as well as variations exhibited by further specimens studied.</p><p>Length 7–8mm (The figure in the original description of 8.5 mm for overall body length does not agree with my own measurements of type material). Variable in colour. Head black. Pronotum dark brown to black, often with diffuse reddish margins. Elytra dark brown to black. Abdomen weakly iridescent. Legs pale. Antennae dark with base of segment I and sometimes segment XI pale. The terminal segment of the maxillary palpi is darker than the penultimate segment.</p><p>Head of average size (pronotum 1.7x wider than head), slightly transverse (1.05x wider than long) with rounded temples bulging behind relatively small eyes and antennal insertion right on front margin (Fig. 3). Micro-punctures very sparse and confined to front of head and inside eyes. Dense short pubescence behind eyes. Two pairs of interocular setae. A line of five postocular setae plus an additional short seta on hind margin of eye. Right mandible with three medial teeth, but with middle tooth reduced; left mandible with two medial teeth (Fig. 35). Maxillary palpi with terminal segment elongate and densely pubescent, more or less symmetric, longer than glabrous penultimate segment which is also elongate (Fig. 53). First segment of antenna almost as long as next four. Segments I to VIII elongate, X transverse (Fig. 79).</p><p>Pronotum only slightly transverse (1.1x wider than long) with rounded sides and widest in basal half. Shining with no micro-punctures. One pair of dorsal setae. One pair of lateral setae. Marginal setae shorter than in A. orientalis . Elytra transverse (1.55x wider than long) with pubescence arising from relatively fine asperate punctures. Marginal fringe of thick bristles about as long as the hairs on the rest of the elytra. Abdominal tergites with evenly spaced, relatively sparse asperate punctures and a marginal fringe of bristles similar to those on the elytra.</p><p>Paramere bilobed, lobes parallel and proximate, each lobe ridged, though not as strongly as in A. orientalis, pegs confusedly arranged along inner margins at apex (Fig. 111). Median lobe longer than paramere with rather narrow, spoon-shaped apex.</p><p>Type material. Paratype 3: “S. Leone W. Area Regent Rossi lg 20.xii.92 / PARATYPUS Acylophorus rossii sp. n. Bordoni det. 1992” (cBord).</p><p>This species was given the manuscript name, A. natalensis, by Bernhauer. Types were labelled accordingly, but he never published a description.</p><p>Further material examined. SIERRA LEONE: Northern Province: near Bumbuna, W Rossi, 29. ii.1992, 1 (cBord); Kambai / Binkolo, W Rossi, 16.ii.1993, 132Ƥ (cBord &amp; cLott); Binkolo / Kamabesi, W Rossi, 13. i.1997, 5 (cBord); Southern Province: Tiwai Island, W Rossi, 8. x.1989, 13 (cBord); Western Area: Picket Hill, W Rossi, 1. xi.1995, 10 (cBord); Regent, W Rossi, 31.iii–11. iv.1996, 18 (cBord); York, W Rossi, 18. xii.1992, 13 (cBord). SOUTH AFRICA: Natal, no date, Sharp collection, 131Ƥ (BMNH &amp; FMNH).</p><p>Distribution and bionomics. Probably widespread in continental Africa having been found in South Africa and Sierra Leone, but so far unknown from in between and collected less frequently than A. orientalis and A. nitens (Fig. 141). There are long series of specimens taken from several localities in Sierra Leone, where it has been found in marshes and river margins (Bordoni, 1994).</p><p>Comparative notes. Ve ry similar to A. nitens but distinguished by the smaller eyes in front of more pronounced temples and the more slender and separated lobes of the paramere in the aedeagus. There are also subtle differences in the shape of the medial teeth of the mandibles, but it is not known how much these might be subject to intraspecific variation.</p></div>	https://treatment.plazi.org/id/4D54A843FFB11A6B0694F89FEC7AF93C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lott, Derek A.	Lott, Derek A. (2010): The species of Acylophorus Nordmann (Coleoptera: Staphylinidae: Staphylininae) in continental sub-Saharan Africa. Zootaxa 2402: 1-51, DOI: 10.5281/zenodo.275907
4D54A843FFB21A6A0694F8B8ED43FAB2.text	4D54A843FFB21A6A0694F8B8ED43FAB2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acylophorus bumbunae Bordoni	<div><p>Acylophorus bumbunae Bordoni</p><p>(Figs 4, 54, 80, 112)</p><p>Acylophorus bumbunae Bordoni, 1994: 311; Herman, 2001: 3027.</p><p>Diagnostic characters. Bordoni’s description contains illustrations of the aedeagus, head, maxillary palpi and posterior tarsi in addition to a textual description of morphological features. Note, however, that the illustration of the aedeagus in Bordoni (1994) is incorrectly labelled. Fig. 9 in that work shows the aedeagus of A. bumbunae, not fig. 8. The following section contains additional diagnostic features.</p><p>Length 7mm. Head black. Pronotum dark brown to black, often with diffuse reddish margins. Elytra dark brown to black. Abdomen weakly iridescent. Legs pale. Antennae dark with base of segment I and sometimes segment XI pale. The terminal segment of the maxillary palpi is darker than the penultimate segment.</p><p>Head of average size (pronotum 1.7x wider than head), more or less as long as wide with rounded temples bulging behind relatively small eyes and antennal insertion right on front margin (Fig. 4). Micro-punctures very sparse and confined to front of head and inside eyes. Dense short pubescence behind eyes. Two pairs of interocular setae. A line of five postocular setae plus an additional short seta on hind margin of eye. Maxillary palpi with terminal segment elongate and densely pubescent, more or less symmetric, longer than glabrous penultimate segment which is also elongate (Fig. 54). First segment of antenna longer than next three. Segments I to VIII elongate (Fig. 80).</p><p>Pronotum slightly transverse (1.15x wider than long) with rounded sides and widest in basal half. Shining with sparse, but evenly distributed micro-punctures. One pair of dorsal setae. One pair of lateral setae. Marginal setae shorter than in A. orientalis . Elytra strongly transverse (1.7x wider than long) with pubescence arising from relatively fine asperate punctures. Apical fringe of thick bristles about as long as the hairs on the rest of the elytra. Abdominal tergites with evenly spaced, relatively sparse asperate punctures and a marginal fringe of bristles similar to those on the elytra.</p><p>Paramere bilobed, lobes parallel and well separated, slightly converging toward apex, each lobe keeled as in A. orientalis, pegs confusedly arranged along inner margins at apex (Fig. 112). Median lobe not longer than paramere, barely expanded but strongly flattened at apex.</p><p>Type material. Holotype 3: “ SIERRA LEONE Northern Province near Bumbuna 12.v. 91 W. Rossi / Laboulbeniales n. 1646 Walter Rossi / HOLOTYPUS / HOLOTYPUS Acylophorus bumbunae sp. n. Bordoni det. 1992” (cBord).</p><p>Distribution and bionomics. Only known from three specimens collected in dead leaves on the banks of the River Seli in Sierra Leone (Bordoni, 1994) (Fig. 142).</p><p>Comparative notes. Very similar to A. rossii . The external differences given by Bordoni (1994) include the larger eyes, the more transverse pronotum and elytra, but these characters can be difficult to appreciate, given that the differences are small and that comparison of these particular characters is difficult, unless specimens are set perfectly during their preparation. The significant difference between the two species occurs in the form of the aedeagus. The illustration accompanying the original description exaggerates certain features. Fig. 112 is redrawn from the holotype. The important features are the relatively short median lobe in comparison with the paramere and its flattened apex.</p></div>	https://treatment.plazi.org/id/4D54A843FFB21A6A0694F8B8ED43FAB2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lott, Derek A.	Lott, Derek A. (2010): The species of Acylophorus Nordmann (Coleoptera: Staphylinidae: Staphylininae) in continental sub-Saharan Africa. Zootaxa 2402: 1-51, DOI: 10.5281/zenodo.275907
4D54A843FFB31A6D0694FA0DEABDFB5E.text	4D54A843FFB31A6D0694FA0DEABDFB5E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acylophorus capensis Cameron	<div><p>Acylophorus capensis Cameron</p><p>(Figs 5, 36, 55, 82, 113)</p><p>Acylophorus capensis Cameron, 1945: 716; Herman, 2001: 3028.</p><p>Redescription. Length 7mm. Body black with iridescent abdomen. Abdominal tergite and sternite VIII black, occasionally pale at extreme base and apex. Antennae dark brown with segment XI and base of segment I occasionally red-brown. Legs pale with darkened meso- and meta-femora and tibiae. Palpi completely pale.</p><p>Head large (pronotum 1.4–1.6x wider than head), as long as wide with temples well developed (Fig. 5). Mostly shining with sparse micro-punctures confined to front of head, but with dense short pubescence behind eyes. Antennae inserted close to front margin of head, but separated from it by a small pigmented strip. Temples long and pronounced behind relatively small eyes. Two pairs of interocular setae arising from foveate punctures much closer to eyes than each other. Four postocular setae visible from above on each side with a further seta out of line on the hind-border of the eye. Mandibles with two medial teeth on each side (Fig. 36). Maxillary palpi with terminal segment elongate and sparsely pubescent, almost symmetric with attenuated apex, longer than glabrous penultimate segment which is slightly elongate (Fig. 55). First segment of antenna longer than next three. Segments I to VI elongate, IX to XI1 transverse (Fig. 81).</p><p>Pronotum weakly transverse (1.025–1.1x wider than long) with rounded sides, widest in basal half. One pair of dorsal setae (The type specimen has two proximate pairs of dorsal punctures, but this is probably an aberration.) One pair of lateral setae. Marginal setae long. Elytra transverse (1.5x wider than long) with long, close pubescence and asperate punctures which are sparser, but stronger than in A. orientalis . Fringe of bristles on hind margins longer than the general pubescence. Abdominal tergites also with long pubescence and strong asperate punctures, but not quite as dense as on elytra. Punctures sparser on apical half of each tergite than on basal half.</p><p>Aedeagus with the paramere divided into two slightly converging, proximate lobes, each lobe being ridged dorsally, though much less so than in A. orientalis (Fig. 113). The internal pegs are confusedly distributed toward the apex. The median lobe is appreciably longer than the paramere and markedly expanded at the apex.</p><p>Type material. Cameron described the species from a unique type in his collection. This specimen is the holotype by monotypy. Holotype Ƥ: “ Type / Marsh / Simon’s Town iv-vi-15 MC / A. capensis Cam. TYPE ” (BMNH).</p><p>The name capensis was also used for this species by Fauvel, but in manuscript form on a specimen label. He never published any description.</p><p>Further material examined. SOUTH AFRICA: Eastern Cape: Hogsback, 970–1300m, 32 O 35’S 26 O 56-57’E, J Janák, 5–7. xii.2006, 23 (cJanák); KwaZulu-Natal: Pietermaritzburg, Queen Elizabeth Park, 29 O 34.157 ’S 30 O 19.299 ’E, J Janák, 22.xi.2006, 233Ƥ (cJanák &amp; cLott); Western Cape: Cape Town, ex. Coll. Fauvel, 1Ƥ (IRSNB); Swellendam, R. Kmeco, 28.xi–1. xii.1997, 13 (cJanák); Harkerville S.F., Kranshoek Picnic Area, 145m, 34 O 04.7 ’S 23 O 13.6 ’E, 1. ii.2004, 13 (FMNH); Grootvadersbosch N.R., Bushbuck Trail (Bosbockrand), 360m, 33 O 58.9 ’S 20 O 49.1 ’E, 26. i.2004, 1 Ƥ (FMNH).</p><p>Distribution and bionomics. This species is only known so far from South Africa (Fig. 141). All recent specimens have been taken from stream margins. In two localities these were in “afromontane forest”.</p><p>Comparative notes. Distinctive within the species group by virtue of its large head. Most s imilar to A. rossii but has even smaller eyes. The distinctive shape of the terminal segment of the maxillary palp, the arrangement of medial teeth on the mandibles, the short paramere in comparison to the median lobe of the aedeagus and the sparse, but strong punctures on the elytra are also useful diagnostic characters.</p></div>	https://treatment.plazi.org/id/4D54A843FFB31A6D0694FA0DEABDFB5E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lott, Derek A.	Lott, Derek A. (2010): The species of Acylophorus Nordmann (Coleoptera: Staphylinidae: Staphylininae) in continental sub-Saharan Africa. Zootaxa 2402: 1-51, DOI: 10.5281/zenodo.275907
4D54A843FFB41A6C0694FAA9EF75FC72.text	4D54A843FFB41A6C0694FAA9EF75FC72.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acylophorus dankalensis Bordoni	<div><p>Acylophorus dankalensis Bordoni</p><p>(Figs 6, 37, 56, 82, 114)</p><p>Acylophorus dankalensis Bordoni, 1994: 311; Herman, 2001: 3028.</p><p>Diagnostic characters. Bordoni’s description contains illustrations of the aedeagus, head, maxillary palpi and posterior tarsi in addition to a textual description of morphological features. Note, however, that the illustration of the aedeagus in Bordoni (1994) is incorrectly labelled. Fig. 8 in that work shows the aedeagus of A. dankalensis, not fig. 7. The following section contains additional diagnostic features as well as variations exhibited by further specimens studied.</p><p>Length 7–7.5mm. Head black. Pronotum brown to dark brown. Elytra brown. Abdomen brown and iridescent. Legs pale. Antennae pale with segments II to VIII infuscated. The maxillary palpi are uniformly pale.</p><p>Head large (pronotum 1.6x wider than head), as long as wide with rounded temples and antennal insertion right on front margin (Fig. 6). Eyes relatively small. Micro-punctures sparse and concentrated toward front of head. Dense short pubescence behind eyes. Two pairs of interocular setae. A line of five postocular setae plus an additional short seta on hind margin of eye. Underside of head sparsely pubescent, strongly depressed at base with gular sutures separate, but very proximate toward base. Mandibles with outer margin strongly curved. Right mandible with medial tooth and flange; left mandible with one sharp tooth and one blunt tooth (Fig. 37). Maxillary palpi with terminal segment elongate and densely pubescent, almost symmetric, longer than glabrous penultimate segment which is also elongate (Fig. 56). First segment of antenna longer than next three. Segments I to VIII elongate, X transverse (Fig. 82).</p><p>Pronotum only slightly transverse (1.2x wider than long) with rounded sides and widest in basal half. Shining with no micro-punctures. One pair of dorsal setae. One pair of lateral setae. Marginal setae long. Elytra transverse (1.4x wider than long) with pubescence arising from relatively fine asperate punctures. Apical fringe of thick bristles slightly longer than the hairs on the rest of the elytra. Abdominal tergites with long semi-erect pubescence arising from asperate punctures, longer than marginal fringe of bristles.</p><p>Paramere bilobed, lobes parallel, narrow and proximate, each lobe strongly ridged, though not as strongly as in A. orientalis, pegs arranged confusedly in apical half of each lobe (Fig. 114). Median lobe not longer than paramere with rounded and slightly expanded apex, flattened at tip.</p><p>Type material. Holotype 3: “ SIERRA LEONE MT i LOMA CASCATE DENKALE m 800 ca 25– 26.xi.84 Leg. W. ROSSI / HOLOTYPUS / HOLOTYPUS Acylophorus dankalensis sp. n. Bordoni det. 1992” (cBord).</p><p>Further material examined. R.D. CONGO: Kivu: Masisi, Mutakato, 800m, N. Leleup, ix.1953, 13 (ISRNB); Tshuapa: Lac Tumba, Mabali, 350m, N. Leleup, x.1955, 13 (ISRNB). NAMIBIA: Popa Falls, Snizek, i.1994, 13 (cJanak).</p><p>Distribution and bionomics. Collected from waterfalls in Sierra Leone and Namibia and from forest humus at two localities in the Democratic Republic of Congo (Fig. 142).</p><p>Comparative notes. Distinguished within the species group by a combination of the small eyes, the elongate antennal segments, the curved mandibles and the arrangement of medial teeth, as well as the form of the aedeagus.</p></div>	https://treatment.plazi.org/id/4D54A843FFB41A6C0694FAA9EF75FC72	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lott, Derek A.	Lott, Derek A. (2010): The species of Acylophorus Nordmann (Coleoptera: Staphylinidae: Staphylininae) in continental sub-Saharan Africa. Zootaxa 2402: 1-51, DOI: 10.5281/zenodo.275907
4D54A843FFB51A6F0694FC5DEB5EFEA9.text	4D54A843FFB51A6F0694FC5DEB5EFEA9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acylophorus tshuapensis Lott	<div><p>Acylophorus tshuapensis Lott, new species</p><p>(Figs 7, 57, 83, 115)</p><p>Description. Length 8mm. Head black. Pronotum chestnut brown. Elytra brown. Abdomen dark and iridescent. Legs pale. Antennae pale with segments II to VIII slightly infuscated. The maxillary palpi are uniformly pale.</p><p>Head somewhat small (pronotum 1.8x wider than head), as long as wide with rounded, not very pronounced temples and antennal insertion close to front margin, but separated by narrow pigmented strip (Fig. 7). Eyes relatively small. Micro-punctures very sparse. Dense short pubescence behind eyes. Two pairs of interocular setae arising from foveate punctures much closer to eyes than each other. A line of five postocular setae plus an additional short seta on hind margin of eye. Maxillary palpi with terminal segment very elongate, densely pubescent, and symmetric, longer than glabrous penultimate segment which is also elongate (Fig. 57). First segment of antenna as long as next three. Segments I to IX elongate, X transverse (Fig. 83).</p><p>Pronotum only slightly transverse (1.1x wider than long) with rounded sides and widest in basal half, front margins somewhat produced at sides. Shining with no micro-punctures. One pair of dorsal setae. One pair of lateral setae. Marginal setae long. Elytra transverse (1.6x wider than long) with pubescence arising from relatively fine asperate punctures. Apical fringe of thick bristles longer than the hairs on the rest of the elytra. Abdominal tergites with long semi-erect pubescence arising from asperate punctures that are denser at the base of each tergite. Hairs longer than marginal fringe of bristles.</p><p>Paramere entire with wide central groove at apex, suggesting two parallel lobes that have become fused together (Fig. 115). Median lobe much longer than paramere and strongly expanded at apex.</p><p>Type material. Holotype 3: “ COLL. MUS. CONGO Tshuapa: Flandria 18-x-1946 Rév. P. Hulstaert / R. DET. UU 5694 / Acylophorus sp. n / HOLOTYPE Acylophorus tshuapensis sp. n. 3 det. DA Lott, 2009” (IRSNB).</p><p>Distribution and bionomics. Known only from the holotype taken in the Democratic Republic of Congo (Fig. 142).</p><p>Comparative notes. Fairly distinct within the A. orientalis species group by virtue of its small eyes, more sinuate front margin to the pronotum and the fused lobes of the paramere. The medial segments of the antennae and the terminal segment of the maxillary palpi are also more elongate than in the other species of the group.</p><p>Etymology. The specific name is the masculine form of an adjective meaning “from Tshuapa”.</p></div>	https://treatment.plazi.org/id/4D54A843FFB51A6F0694FC5DEB5EFEA9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lott, Derek A.	Lott, Derek A. (2010): The species of Acylophorus Nordmann (Coleoptera: Staphylinidae: Staphylininae) in continental sub-Saharan Africa. Zootaxa 2402: 1-51, DOI: 10.5281/zenodo.275907
4D54A843FFB61A6E0694FE3DEC8AFF39.text	4D54A843FFB61A6E0694FE3DEC8AFF39.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acylophorus makhoreae Lott	<div><p>Acylophorus makhoreae Lott, new species</p><p>(Figs 8, 38, 58, 84, 116)</p><p>Description. Length 7mm. Body colour very variable ranging from pale brown to black. Pronotum yellow to red-brown with a dark, diffuse mark covering the disc and sometimes suffused almost to the edges. Abdomen iridescent when dark. Appendages similarly variable in colour, but the terminal segment of the maxillary palpi is always darker than the penultimate segment.</p><p>Head of average size (pronotum 1.75x wider than head), more or less as long as wide with rounded temples not very evident (Fig. 8). Pigmented area of head extending well in front of antennal insertion. Micropunctures extending over much of head, but very sparse away from the front of the head and next to the eyes. Dense short pubescence behind eyes. Two pairs of interocular setae arising from foveate punctures much closer to eyes than each other. Only four postocular setae visible from above on each side. Underside of head sparsely pubescent, depressed at base with gular sutures separate, but very proximate toward base. Right mandible with one sharp median tooth; left mandible lacking sharp tooth (Fig. 38). Maxillary palpi with terminal segment densely pubescent, with rounded angle on outer margin, asymmetric and less elongate than A. orientalis, longer than glabrous penultimate segment which is slightly elongate (Fig. 58). First segment of antenna as long as next five. Segments I to IV elongate, VII to XI transverse (Fig. 84).</p><p>Pronotum only slightly transverse (1.1x wider than long) with rounded sides and widest in basal half. Shining with no micro-punctures. Dorsal, lateral and marginal setae shorter than in A. orientalis . Elytra transverse (1.7x wider than long) with pubescence arising from asperate punctures. Apical fringe of bristles longer than the hairs on the rest of the elytra. Abdominal tergites with evenly spaced, relatively sparse asperate punctures.</p><p>Paramere bilobed, each lobe fairly flat, at least toward apex, pegs concentrated in dense mass at apex (Fig. 116). Median lobe longer than paramere with truncate apex.</p><p>Type material. Holotype 3: “Under plants at a stream edge / ETHIOPIA: Kaffa 28km. S. of Jimma 2,000 m’ xi.1971 / R. O.S. Clark B.M. 1973-450 / HOLOTYPE Acylophorus makhoreae sp. n. 3 det. DA Lott, 2009” (BMNH). Paratypes 13 1Ƥ: “ ETHIOPIA: ILUBADOR 10km. W. Bedelle 0825N 3618E x.1972 1,800 m. / R. O.S. Clark B.M. 1973-450 / PARATYPE Acylophorus makhoreae sp. n. det. DA Lott, 2009” (BMNH); 4: “Under plants &amp; stones at edge stream / ETHIOPIA: Kaffa Belleta F. 2,100 m 40km. S.W. Jimma 1971 / R. O.S. Clark B.M. 1973-450 / PARATYPE Acylophorus makhoreae sp. n. det. DA Lott, 2009” (BMNH); 1Ƥ: “ih red ref. / ETHIOPIA: Kaffa Jimma to Addis Rd. 0749-370 iii.1972. / R. O.S. Clark B.M. 1973-450 / PARATYPE Acylophorus makhoreae sp. n. det. DA Lott, 2009” (BMNH).</p><p>Distribution and bionomics. All material seen so far comes from the south-west of Ethiopia (Fig. 142). I have also seen a female specimen collected at “8,000” feet from the Djem-Djem Forest to the west of Addis Ababa that may belong to this species. Most of the specimens were collected along streams.</p><p>Comparative notes. Fairly distinct within the A. orientalis species group by virtue of the position of the antennal insertion, the broader, more asymmetric terminal segment of the maxillary palpi and the more transverse medial antennal segments. The form of the aedeagus and the arrangement of medial teeth on the mandibles are also useful characters.</p><p>Etymology. Makhore was a legendary queen, who founded the kingdom of Jimma in the area where many of the type specimens were collected. The species name is the genitive case of a Latinised version of her name.</p><p>Discussion. A. makhoreae has been placed in the A. orientalis group, because it lacks dense micropunctures on the head and pronotum. However, the form of the maxillary palpi, the mandibles and the aedeagus and the position of the antennal insertion point all suggest that it may be more closely related to some of the species in the A. densipennis group.</p></div>	https://treatment.plazi.org/id/4D54A843FFB61A6E0694FE3DEC8AFF39	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lott, Derek A.	Lott, Derek A. (2010): The species of Acylophorus Nordmann (Coleoptera: Staphylinidae: Staphylininae) in continental sub-Saharan Africa. Zootaxa 2402: 1-51, DOI: 10.5281/zenodo.275907
4D54A843FFB71A6E0694FE82EAA3FE11.text	4D54A843FFB71A6E0694FE82EAA3FE11.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acylophorus densipennis	<div><p>A. densipennis group</p><p>This group is similar to the A. orientalis group, but the micro-punctures on the head and pronotum are dense and easily visible at low magnifications (x40). In addition the penultimate segment of the maxillary palpi is more triangular and the pigmented area of the head is produced beyond the insertion point of the antennae. The paramere is bilobed or, in one case ( A. densipennis), the lobes are fused to form one lobe.</p></div>	https://treatment.plazi.org/id/4D54A843FFB71A6E0694FE82EAA3FE11	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lott, Derek A.	Lott, Derek A. (2010): The species of Acylophorus Nordmann (Coleoptera: Staphylinidae: Staphylininae) in continental sub-Saharan Africa. Zootaxa 2402: 1-51, DOI: 10.5281/zenodo.275907
4D54A843FFB71A710694FDFAEDB0FE32.text	4D54A843FFB71A710694FDFAEDB0FE32.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acylophorus densipennis Bernhauer	<div><p>Acylophorus densipennis Bernhauer</p><p>(Figs 9, 59, 85, 117, 132)</p><p>Acylophorus densipennis Bernhauer, 1929: 125; Scheerpeltz, 1933: 1466; Cameron, 1950: 43; Herman, 2001: 3028.</p><p>Redescription. Length 5–6mm. Body dark brown to black. Abdominal tergites vaguely iridescent. Legs and palpi pale. Antennae with first segment pale and remainder darkened.</p><p>Head large (pronotum 1.6x wider than head), as long as wide with rounded but unpronounced temples and pigmented area produced well in front of antennal insertion (Fig. 9). Eyes somewhat large. Dorsal surface of head densely covered with micro-punctures. Dense pale pubescence behind eyes. Two pairs of interocular setae arising from foveate punctures much closer to eyes than each other. Five postocular setae just about visible on each side, no additional seta on hind margin of eye. Underside of head strongly depressed at base. Maxillary palpi with terminal segment pubescent, almost angled on outer margin, gently rounded on inner margin and asymmetric, much broader than in A. orientalis, longer than glabrous penultimate segment which is broadly triangular (Fig. 59). First segment of antenna as long as next five. Segments I to II elongate, V to XI transverse (Fig. 85).</p><p>Pronotum slightly transverse (1.1x wider than long) with sides less rounded than in A. orientalis, widest in basal half, covered with dense micro-punctures. One pair of dorsal setae and one pair of lateral setae. Marginal setae long. Elytra transverse (1.5x wider than long) with close, bright yellow pubescence and asperate puncturation finer and denser than in A. orientalis . Apical fringe of bristles very short, shorter than pubescence on other parts of the elytra. Pubescence on elytra and abdominal tergites shorter than in A. orientalis . Punctures on tergites sparser than on elytra.</p><p>Males with genital segment highly modified. Apex of sternite IX deeply incised and styles of tergite nine each with large inward-pointing spine (Fig. 132). Aedeagus also highly modified (Fig. 117) with lobes of paramere fused. Sides of paramere with narrow, almost vertical flange, base lipped. Pegs arranged in rows along the margin well below the apex. Median lobe longer than paramere, but much narrower and acutely pointed at apex.</p><p>Type material. Bernhauer described the species from specimens collected at “Haut-Uele: Moto, Februar, März 1923” and “Ituri: La Moto: Madyu, L. Burgeon”. Three specimens can be recognised as belonging to the type series and one of these is here designated as the lectotype in order to fix the identity of the species. Lectotype 3: “ TYPE / MUSÉE DU CONGO; Ituri: La Moto: Madyu; L. Burgeon / R. DÉT. M 879 / Acyl. densipennis Brh. n. sp. / LECTOTYPE Acylophorus densipennis Bernhauer 3 det. DA Lott, 2009” (MRAC). Paralectotypes 1Ƥ: “ MUSÉE DU CONGO / Haut-Uelé: Moto; -II-III 1923; L. Burgeon / Acylophorus densipennis Bernh. Typus. / Chicago NHMus M. Bernhauer Collection / PARALECTOTYPE Acylophorus densipennis Bernhauer Ƥ det. DA Lott, 2009” (FMNH); 1Ƥ: “ MUSÉE DU CONGO / Ituri; La Moto: Madyu; L. Burgeon / Acylophorus densipennis Brnh. Cotypus. / Chicago NHMus M. Bernhauer Collection / PARALECTOTYPE Acylophorus densipennis Bernhauer 3 det. DA Lott, 2009” (FMNH).</p><p>Further material examined. R.D. CONGO: Bas-Congo: Singa-Karu, 26. ii.1924, 1 Ƥ (IRSNB); Seke- Banga, 13. iv.1924, 1 Ƥ (BMNH); Thysville, N.Leleup, 13. vii.1949, 1 Ƥ (IRSNB); Haut-Uelé: Watsa, L.</p><p>Burgeon, xi.1919, 1 Ƥ (MRAC). R. CONGO: Mayumbe: Suinbi, A.Collart, 4. v.1926, 13 (IRSNB). SIERRA LEONE: Western Area: near Bathurst, W Rossi, 26.v.1991, 133Ƥ (cBord &amp; cLott); Regent, W Rossi, 6- 7. iii.1993, 1 Ƥ (cBord).</p><p>Distribution and bionomics. Known from West Africa and the Congo catchment (Fig. 143). Ecological data are somewhat sparse. Recorded from vegetation at the margins of a marsh and “le détritus de manioc”.</p><p>Comparative notes. Apart from the characters specific to the species group, A. densipennis has larger eyes on a larger head than A. orientalis . Very distinctive within the species group by virtue of the short fringe of apical bristles on the elytra and the highly modified male sexual characters. The bright yellow pubescence on the elytra is a character shared with several other species in the group.</p></div>	https://treatment.plazi.org/id/4D54A843FFB71A710694FDFAEDB0FE32	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lott, Derek A.	Lott, Derek A. (2010): The species of Acylophorus Nordmann (Coleoptera: Staphylinidae: Staphylininae) in continental sub-Saharan Africa. Zootaxa 2402: 1-51, DOI: 10.5281/zenodo.275907
4D54A843FFA81A700694FD80EF4AFC2C.text	4D54A843FFA81A700694FD80EF4AFC2C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acylophorus antennalis Cameron	<div><p>Acylophorus antennalis Cameron</p><p>(Figs 10, 39, 40, 60, 86, 118)</p><p>Acylophorus antennalis Cameron, 1932: 135; Scheerpeltz, 1933: 1466; Herman, 2001: 3026.</p><p>Acylophorus tenuiceps Bernhauer, 1932: 157; Scheerpeltz, 1933: 1467; Herman, 2001: 3037 (new synonym).</p><p>Redescription. Length 5.5– 7mm. Body dark brown to black. Abdomen sometimes iridescent. Legs dark brown or pale with darkened femora. Maxillary palpi pale, sometimes with last segment darker. Antennae pale with middle segments slightly darkened.</p><p>Head large (pronotum 1.6x wider than head), 1.1x longer than wide with evident temples and pigmented area produced well in front of antennal insertion (Fig. 10). Forehead somewhat arched. Eyes relatively small. Covered with dense micro-punctures. Dense pale pubescence behind eyes. Two pairs of interocular setae arising from foveate punctures much closer to eyes than each other. Four postocular setae visible from above on each side, no additional seta on hind margin of eye. Mandibles with one small medial tooth on the right and just a flange on the left (Figs 39 &amp; 40). Maxillary palpi with terminal segment pubescent, more rounded on outer margin than inner margin and asymmetric, sometimes slightly attenuated at apex, longer than glabrous penultimate segment which is triangular (Fig. 60). First segment of antenna as long as next five. Segments I to VI elongate, X transverse (Fig. 86).</p><p>Pronotum relatively narrow with sides less rounded, slightly transverse (1.1x wider than long) and widest toward basal half, covered with dense micro-punctures. One pair of dorsal setae and one pair of lateral setae. Marginal setae short. Elytra only slightly transverse (1.4x to 1.5x wider than long) with close, bright yellow pubescence. Asperate punctures strong as in A. orientalis . Fringe of apical bristles slightly longer than the pubescence on the rest of the elytra. Abdominal tergites with similar pubescence. Punctation on abdominal tergites finer than on elytra and becoming sparser on apical segments.</p><p>Sternite IX of male with apex entire. Paramere of aedeagus bilobed, lobes parallel, pegs confusedly arranged in apical third of each lobe, base lipped as in A. densipennis (Fig. 118). Median lobe longer than paramere, expanded at spoon-shaped apex.</p><p>Type material. Cameron described the species from specimens collected by A. Collart on 17th April 1926 at Lolo Damvu (Mayumbe) in what is now the Republic of Congo. Note that the date of publication is given as 30th June 1932. Two specimens from the type series have been located, one of which is here designated as the lectotype in order to fix the identity of the species. Lectotype Ƥ: “ TYPE / Lolo Damvu; 17-IV-26; A. COLLART / A. antennalis Cam. TYPE / LECTOTYPE Acylophorus antennalis Cameron Ƥ det. DA Lott, 2009” (IRSNB); Paralectotype Ƥ: “Lolo Damvu; 17-IV-26; A. COLLART / M. Cameron Bequest B.M. 1955-147 / A. antennalis Cam. COTYPE / PARALECTOTYPE Acylophorus antennalis Cameron Ƥ det. DA Lott, 2009” (BMNH).</p><p>Acylophorus tenuiceps Bernhauer. The species was described from specimens collected by Burgeon at Yebo (in December 1925) and Moto in the Haut-Uelé area in what is now the Democratic Republic of Congo. Note that the date of publication is given as 16th October 1932. Only one specimen collected from Moto can be unequivocally recognised as belonging to the type series. A further specimen collected from Yebo has a date that does not quite match the details in the original description. However, this is probably due to some kind of mix-up, as the date has been changed on the label and it has been labelled as a type by Bernhauer. Consequently, this Yebo specimen can be regarded as belonging to the type series. A lectotype is here designated in order to fix the identity of the species. Lectotype 3: “ MUSÉE DU CONGO; Haut-Uelé: Moto / 1920 L. Burgeon / tenuiceps Brnh. Acylophorus Cotypus. / Chicago NHMus M. Bernhauer Collection / LECTOTYPE Acylophorus tenuiceps Bernhauer 3 det DA Lott, 2009” (FMNH); Paralectotypes Ƥ: “ MUSÉE DU CONGO Haut-Uele:Yebo / - XII-1920 L. Burgeon / Acylophorus angusticeps Brnh. n. sp. / tenuiiceps Brnh. Acylophorus Typus / PARALECTOTYPE Acylophorus tenuiceps Bernhauer Ƥ det DA Lott, 2009” (FMNH). Bernhauer did not compare his species with A. antennalis, which was only described a few months earlier. The three types are a millimetre larger than the two types of A. antennalis and there are slight differences in the form of the mandibles (Figs 39 &amp; 40) and the last segment of the maxillary palpi. However, there is considerable infra-specific variation in this last character, even in the small amount of material available for the present study and, notwithstanding the absence of males in the type series of A. antennalis, there can be little doubt that the two series are conspecific. A. tenuiceps is, therefore, here synonymised with A. antennalis .</p><p>Further material examined. R.D. CONGO: Stanleyville, L Burgeon, 13 (MRAC); Haut-Uele: Yebo, L Burgeon, i.1926, 13 (MRAC).</p><p>Distribution and bionomics. Only recorded from the Democratic Republic of and the Republic of Congo (Fig. 144). There are no data on habitat.</p><p>Comparative notes. Similar to A. densipennis by virtue of the bright yellow pubescence on the elytra, but generally larger and with smaller eyes. The apical bristles on the elytra are longer and the aedeagus is completely different. Much more similar to A. collarti, from which it is best distinguished by the form of the aedeagus.</p></div>	https://treatment.plazi.org/id/4D54A843FFA81A700694FD80EF4AFC2C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lott, Derek A.	Lott, Derek A. (2010): The species of Acylophorus Nordmann (Coleoptera: Staphylinidae: Staphylininae) in continental sub-Saharan Africa. Zootaxa 2402: 1-51, DOI: 10.5281/zenodo.275907
4D54A843FFA91A730694FBB8EDAAFCE4.text	4D54A843FFA91A730694FBB8EDAAFCE4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acylophorus collarti Cameron	<div><p>Acylophorus collarti Cameron</p><p>(Figs 11, 41, 61, 87, 119)</p><p>Acylophorus collarti Cameron, 1935: 375; Herman, 2001: 3027.</p><p>Redescription. Length 6.5– 7mm. Body generally black, sometimes with dark brown abdomen or more generally dark brown. Leg colour very variable from all pale to black with red tarsi. Maxillary palpi pale with last segment darker. Antennae pale with middle segments slightly darkened to all dark.</p><p>Head large (pronotum 1.6x wider than head), 1.1x longer than wide with evident temples (Fig. 11). Pigmented area of head produced in front of antennal insertion. Forehead somewhat arched. Eyes relatively small. Covered with dense micro-punctures. Dense pale pubescence behind eyes. Two pairs of interocular setae arising from foveate punctures much closer to eyes than each other. Four postocular setae visible on each side, additional seta on hind margin of eye absent. Underside of head strongly depressed at base. Mandibles with one well developed medial tooth in front of a vestigial tooth on the right and just a flange on the left (Fig. 41). Maxillary palpi with terminal segment pubescent, more rounded on outer margin than inner margin and asymmetric, sometimes slightly attenuated at apex, longer than glabrous penultimate segment which is triangular (Fig. 61). First segment of antenna as long as next five. Segments I to IV elongate, IX and X transverse (Fig. 87).</p><p>Pronotum relatively narrow with sides less rounded, almost quadrate (1.05x wider than long) and widest toward basal half, covered with dense micro-punctures. One pair of dorsal setae and one pair of lateral setae. Marginal setae short. Elytra only slightly transverse (1.4x wider than long) with close, bright yellow pubescence. Asperate punctures strong as in A. orientalis . Fringe of apical bristles slightly longer than the pubescence on the rest of the elytra. Abdominal tergites with similar pubescence. Punctation on abdominal tergites finer than on elytra and becoming sparser on apical segments.</p><p>Sternite IX of male with apex entire. Paramere of aedeagus bilobed, lobes slightly divergent, pegs confusedly arranged in apical inner half of each lobe, base lipped as in A. densipennis (Fig. 119). Median lobe not longer than paramere, barely expanded at apex, which is truncate.</p><p>Type material. Cameron described the species from specimens collected by A. Collart on 26th January 1929 at Blukwa (Nizi) in what is now the Democratic Republic of Congo. Two specimens from the type series have been located and one of these is here designated as the lectotype in order to fix the identity of the species. Lectotype 3: “ Paratype / Nizi: Blukwa; 26-I-29; A. COLLART / TYPE / Acylophorus Collarti Cam. TYPE / LECTOTYPE Acylophorus collarti Cameron 3 det. DA Lott, 2009” (IRSNB); Paralectotype 1Ƥ: “ Paratype / Nizi: Blukwa; 26-I-29; A. COLLART / M. Cameron Bequest B.M. 1955-147 / Acyl. Collarti Cam. COTYPE / PARALECTOTYPE Acylophorus collarti Cameron 3 det. DA Lott, 2009” (BMNH).</p><p>Further material examined. CAMEROON: Northwest Province: 3km SW of Bamenda, 1300m, RL Aalbu, 5. ii.1980, 13 (FMNH). R.D. CONGO: Nizi: Blukwa, A Collart, 10.xii.1928 &amp; 25.i.1929, 132Ƥ (IRSNB).</p><p>Distribution and bionomics. Only recorded from the Democratic Republic of Congo and Cameroon (Fig. 144). In Cameroon it was recorded in a stream-side thicket in afromontane forest.</p><p>Comparative notes. Similar to A. densipennis by virtue of the bright yellow pubescence on the elytra, but generally larger and with smaller eyes. The apical bristles on the elytra are longer and the aedeagus is completely different. Much more similar to A. antennalis, from which it is best distinguished by the aedeagus. There may also be differences in the form of the mandibles and the maxillary palpi are narrower, but the extent of intraspecific variation in these characters is not yet understood.</p></div>	https://treatment.plazi.org/id/4D54A843FFA91A730694FBB8EDAAFCE4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lott, Derek A.	Lott, Derek A. (2010): The species of Acylophorus Nordmann (Coleoptera: Staphylinidae: Staphylininae) in continental sub-Saharan Africa. Zootaxa 2402: 1-51, DOI: 10.5281/zenodo.275907
4D54A843FFAA1A720694FCF7EF4AFEA9.text	4D54A843FFAA1A720694FCF7EF4AFEA9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acylophorus micans Lott	<div><p>Acylophorus micans Lott, new species</p><p>(Figs 12, 62, 88, 120)</p><p>Description. Length 4.5– 5mm. Head and elytra black. Pronotum dark brown. Abdominal tergites dark brown and iridescent. Appendages all pale, last segment of maxillary palpi sometimes darker.</p><p>Head of average size (pronotum 1.7x wider than head), 1.1x wider than long with rounded well developed temples and pigmented area produced well in front of antennal insertion (Fig. 12). Eyes relatively small. Dorsal surface of head covered with dense micro-punctures. Dense pale pubescence behind eyes. Two pairs of interocular setae arising from foveate punctures much closer to eyes than each other. Four postocular setae visible on each side, additional seta on hind margin of eye absent. Maxillary palpi with terminal segment pubescent, markedly asymmetric, longer than glabrous penultimate segment which is broadly triangular (Fig. 62). First segment of antenna as long as next six. Segments I to II elongate, IV to X transverse (Fig. 88).</p><p>Pronotum slightly transverse (1.2x wider than long) with sides well rounded, widest in basal half, covered with dense micro-punctures. One pair of dorsal setae and one pair of lateral setae. Marginal setae long. Elytra transverse (1.7x wider than long) with close, bright yellow pubescence arising from asperate punctures finer. Apical fringe of bristles longer than pubescence on other parts of the elytra. Pubescence on abdominal tergites long arising from asperate punctures that are stronger and sparser than on elytra. Abdomen appears wide because of overall small body size.</p><p>Apex of sternite IX broadly rounded. Paramere bilobed, each lobe twisted round side margin of median lobe, pegs confusedly arranged along inner half, basal lip absent (Fig. 120). Median lobe of aedeagus not much longer than paramere.</p><p>Type material. Holotype 3: “ Ivory Coast 7–17km. W. Abidjan I-63 / Coll. W.L. Brown Jr. leg. / HOLOTYPE Acylophorus micans sp. n. 3 det. DA Lott, 2009” (FMNH). Paratypes 1Ƥ: same data as holotype; 131Ƥ (mounted on same card): “Mts. De Cristal Gabon / gabonensis Fvl. / R.I.Sc.N.B. 17.479 Coll. et det. A. Fauvel / PARATYPE Acylophorus micans sp. n. 3 det. DA Lott, 2009” (IRSNB); 2: “Loango Gabon / Coll. et det. A. Fauvel A cylophorus gabonensis Fauv. R.I.Sc.N.B. 17.479 / PARATYPE Acylophorus micans sp. n. det. DA Lott, 2009” (IRSNB). Note that Fauvel gave this species the name gabonensis on his specimen labels, but he never published a description.</p><p>Further material examined. CAMEROON: Missellele near Mt Cameroon, F Zumpt, 1936, 1Ƥ (FMNH).</p><p>Distribution and bionomics. Known so far from Côte d’Ivoire, Cameroon and Gabon (Fig. 143). There are no data on habitat.</p><p>Comparative notes. Somewhat similar to A. densipennis by virtue of the bright yellow pubescence on the elytra, but smaller in overall size and with relatively small eyes and long apical bristles on the elytra. The aedeagus is also completely different. Much smaller than other species in the group with small eyes. Etymology. The specific name is a present participle meaning “glittering” and refers to the iridescent abdomen.</p></div>	https://treatment.plazi.org/id/4D54A843FFAA1A720694FCF7EF4AFEA9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lott, Derek A.	Lott, Derek A. (2010): The species of Acylophorus Nordmann (Coleoptera: Staphylinidae: Staphylininae) in continental sub-Saharan Africa. Zootaxa 2402: 1-51, DOI: 10.5281/zenodo.275907
4D54A843FFAB1A720694FE38EA8AF81C.text	4D54A843FFAB1A720694FE38EA8AF81C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acylophorus allardi Levasseur	<div><p>Acylophorus allardi Levasseur</p><p>(Figs 13, 45, 63, 89, 121, 122)</p><p>Acylophorus allardi Levasseur, 1968: 1400; Herman, 2001: 3026.</p><p>Diagnostic characters. Levasseur’s description contains illustrations of the aedeagus and antenna in addition to a textual description of morphological features. The following section contains additional diagnostic features.</p><p>Length 7–8mm. Head large (pronotum 1.6x wider than head), 1.05x wider than long with rounded but evident temples behind somewhat large eyes (Fig. 13). Head produced in front of antennae insertion point. Covered with dense micro-punctures. Dense pale pubescence behind eyes. Two pairs of interocular setae. Four postocular setae visible on each side, additional seta on hind margin of eye absent. Right mandible with large medial tooth in front of flange, no teeth on left mandible (Fig. 45). Maxillary palpi with terminal segment densely pubescent, markedly asymmetric, longer than very short, triangular, glabrous penultimate segment (Fig. 63). First segment of antenna longer than next four (Fig. 89). Segments I to VIII elongate, X to XI transverse.</p><p>Pronotum weakly transverse (1.1x wider than long) with rounded sides and widest in basal half. Covered with dense micro-punctures. Marginal setae short. Elytra transverse (1.7x wider than long). Typical form with short, bright yellow pubescence, but in a series collected from South Africa, the yellow pubescence is much less shining. Fringe of bristles on hind margin longer than the hairs on the rest of the elytra. Pubescence on abdominal tergites also bright yellow in typical form and relatively short, arising from fine and dense asperate punctures, which are sparser toward apex. In the South African form, the pubescence is slightly longer and less shining.</p><p>Apex of sternite IX entire. Median lobe of aedeagus strongly flattened with a broadly incised apex (Figs 121, 122). Paramere bilobed, each lobe twisted round side margin of median lobe, pegs arranged in rows along each margin, basal lip absent.</p><p>Type material. Syntype 3: “ CONGO BELGE 8.54 D.V. ALLARD / H. KATANGA COLL. LEVASSEUR / LUENA HT LOMAMI / CROTTINS D’ELEPHANT / Acylophorus allardi sp. n. LEVASSEUR DET TYPE ” (MNHN).</p><p>Further material examined. SOUTH AFRICA: KwaZulu-Natal: Pietermaritzburg, Queen Elizabeth Park, 29 O 34.157 ’S 30 O 19.299 ’E, J Janák, 22.xi.2006, 333Ƥ (cJanák &amp; cLott).</p><p>Distribution and bionomics. (Fig. 143). The two type specimens from Katanga were collected from elephant dung, an unusual habitat for Acylophorus . The South African specimens were collected from a stream bank, a more characteristic habitat.</p><p>Comparative notes. Distinguished from most other species in the A. densipennis group by the flattened aedeagus. A. congoensis also has a flattened aedeagus, but the median lobe in A. allardi is more elongate. From species with bright yellow pubescence it can be recognised by its large size, short marginal setae and short pubescence on the elytra and abdominal tergites.</p><p>Discussion. For this species material was available for study from only two localities and they differed in several respects. In South African specimens the pubescence on the elytra and abdomen was a much less brighter yellow and was also slightly shorter. The lobes of the paramere were less twisted around the sides of the median lobe and the shape of the median lobe was also subtly different. At this stage these differences are not considered significant enough to split off the South African series into a separate taxon, but further material is needed to resolve whether the observed variation is intraspecific or interspecific.</p></div>	https://treatment.plazi.org/id/4D54A843FFAB1A720694FE38EA8AF81C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lott, Derek A.	Lott, Derek A. (2010): The species of Acylophorus Nordmann (Coleoptera: Staphylinidae: Staphylininae) in continental sub-Saharan Africa. Zootaxa 2402: 1-51, DOI: 10.5281/zenodo.275907
4D54A843FFAC1A750694FF38EC45F94C.text	4D54A843FFAC1A750694FF38EC45F94C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acylophorus congoensis Cameron	<div><p>Acylophorus congoensis Cameron</p><p>(Figs 14, 42, 64, 90, 123)</p><p>Acylophorus congoensis Cameron, 1932: 134; Scheerpeltz, 1933: 1466; Bernhauer &amp; Paulian, 1942: 364; Cameron, 1950: 43; Herman, 2001: 3027.</p><p>Redescription. Length 7–7.5mm. Body black. Pronotum and less frequently elytra dark brown. Abdomen with very weak iridescence. Legs, antennae and palpi reddish brown, antennae and femora sometimes darkened. Maxillary palpi pale with darker terminal segment.</p><p>Head rounded, large (pronotum 1.6x wider than head), as wide as long with rounded but evident temples behind somewhat large eyes, produced in front of antennal insertion, covered with sparse micro-punctures (Fig. 14). Short dense pubescence behind eyes. Two pairs of short and pale interocular setae arising from foveate punctures much closer to eyes than each other. A line of four postocular setae visible from above on each side. No additional seta on hind margin of eye. Right mandible with large medial tooth in front of flange, no teeth on left mandible (Fig. 42). Maxillary palpi with terminal segment pubescent, more rounded on outer margin than inner margin and asymmetric, longer than glabrous penultimate segment which is triangular (Fig. 64). First segment of antenna as long as next five. Segments I to V elongate, VIII to XI transverse (Fig. 90).</p><p>Pronotum relatively narrow with sides less rounded, almost quadrate (1.05x wider than long) and widest toward basal half, covered with sparse micro-punctures. Dorsal, lateral and marginal setae relatively short. Elytra transverse (1.5x wider than long) with short, pale pubescence that is not shining arising from fine, dense asperate punctures. Apical fringe of bristles rather short, not much longer than hairs on rest of elytra. Abdominal tergites with short, sparse pubescence, barely overlapping in centre of tergite III, though longer on apical tergites.</p><p>Paramere bilobed and divergent, twisted round side margin of median lobe, which is flattened and broadly incised at apex (Fig. 123). Pegs confusedly arranged at apex of each lobe.</p><p>Type material. Cameron described the species from specimens collected by A. Collart on 10th December 1928 at Blukwa (Nizi) in what is now the Democratic Republic of Congo. Two syntypes have been located and one of these is here designated as the lectotype in order to fix the identity of the species. Lectotype Ƥ: “ TYPE / Blukwa; 10-XII-28; A. Collart / Acylophorus congoensis Cam. TYPE / LECTOTYPE Acylophorus congoensis Cameron Ƥ det. DA Lott” (IRSNB). Paralectotype Ƥ: “ Paratype / Blukwa; 10-XII-28; A. Collart / M. Cameron Bequest B.M. 1955-147 / Acylophorus congoensis Cam. COTYPE / PARALECTOTYPE Acylophorus congoensis Cameron Ƥ det. DA Lott” (BMNH).</p><p>Further material examined. SUDAN: Didinga District: Nagichot, 6700 feet, GDH Carpenter, xii.1925 - i.1926, 132Ƥ (BMNH). TANZANIA: W Mkusi, Usambara Mts, B Verdcourt, 31. viii.1950, 13 (MMUM).</p><p>Distribution and bionomics. I have seen specimens from Congo, Tanzania and SE Sudan (Fig. 143). Further records in the literature from Cameroon (Bernhauer &amp; Paulian, 1942) and Congo (Cameron, 1950) must be open to question, given the frequency of misidentifications of African Acylophorus in collections. There are no ecological data.</p><p>Comparative notes. Lacks bright yellow pubescence on elytra found in many other species of this species group. Aedeagus flattened, similar to that of A. allardi but less elongate. Distinguishable from A. mareei and A. ziloensis with similarly short pubescence on the abdomen, either by the form of the mandibles or the last segment of the maxillary palpi.</p></div>	https://treatment.plazi.org/id/4D54A843FFAC1A750694FF38EC45F94C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lott, Derek A.	Lott, Derek A. (2010): The species of Acylophorus Nordmann (Coleoptera: Staphylinidae: Staphylininae) in continental sub-Saharan Africa. Zootaxa 2402: 1-51, DOI: 10.5281/zenodo.275907
4D54A843FFAC1A740694F97DED72FA22.text	4D54A843FFAC1A740694F97DED72FA22.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acylophorus ziloensis Levasseur	<div><p>Acylophorus ziloensis Levasseur</p><p>(Figs 15, 43, 44, 65, 91, 124)</p><p>Acylophorus ziloensis Levasseur, 1968: 1403; Herman, 2001: 3038.</p><p>Diagnostic characters. Levasseur’s description contains illustrations of the aedeagus and antenna in addition to a textual description of morphological features. The following section contains additional diagnostic features.</p><p>Length 6–7mm. Black with margins of pronotum and abdominal tergites sometimes vaguely dark brown. Abdominal tergites very weakly iridescent. Appendages reddish brown.</p><p>Head of average size (pronotum 1.7x wider than head), 1.1x wider than long with rounded, but pronounced temples, with pigmented area produced well in front of antennal insertion (Fig. 15). Dorsal surface of head covered with dense micro-punctures. Dense pale pubescence behind eyes. Two pairs of interocular setae arising from foveate punctures much closer to eyes than each other. Four postocular setae visible on each side, additional seta on hind margin of eye absent. Mandibles with well developed medial tooth on right and untoothed on left (Figs 43 &amp; 44). Maxillary palpi with terminal segment pubescent, very elongate, almost symmetric, longer than glabrous penultimate segment which is broadly triangular (Fig. 65). First segment of antenna nearly as long as next five. Segments I to V elongate, IX to X transverse (Fig. 91).</p><p>Pronotum slightly transverse (1.1x wider than long) with sides well rounded, widest in basal half, covered with dense micro-punctures. One pair of dorsal setae and one pair of lateral setae. Marginal setae short. Elytra transverse (1.5x wider than long) with short, pale, but not shining pubescence arising from dense asperate punctures. Apical fringe of bristles much longer than pubescence on other parts of the elytra. Pubescence on abdominal tergites short but overlapping in centre of tergite III, arising from asperate punctures that are fairly evenly distributed across each tergite and stronger and denser than on elytra.</p><p>Apex of sternite IX entire. Paramere bilobed, lobes widely separated, pegs confusedly arranged right at apex, base lipped (Fig. 124). Median lobe of aedeagus slightly longer than paramere.</p><p>Type material. Holotype 3: “KOLWEZI H. KATANGA COLL. LEVASSEUR / Zilo Walaba / CONGO BELGE 4.1959 D.V. ALLARD / Acylophorus ziloensis sp. n. LEVASSEUR DET 1967 HOLOTYPE ” (MNHN).</p><p>Further material examined. R. D. CONGO: Ituri: Djugu, Mt. Aboro 2200 m. N Leleup i.1954, 131Ƥ (IRSNB); Kivu: Kabare, affl. de la Lwiro, 1700 m, N Leleup xi.1951, 3 (IRSNB); Kabare, Lwiro, 2000 / 2200m, N Leleup xi.1953, 1 (IRSNB); Kalehe, Kahuzi, 2200 m, N Leleup 9.viii.1951, 232Ƥ (IRSNB); Kalehe, Kahuzi, 1800 m, P Basilewsky 25. iii.1953, 1 Ƥ (IRSNB); Kalehe, Kahuzi, 2200 m, P Basilewsky 27. iii.1953, 4 Ƥ (IRSNB); Lubero, 2200 m. N. Leleup xii.1951, 2 (IRSNB); Lubero, Katondi, 2200 m. N. Leleup xii.1951, 13 (IRSNB); Lubero, Lubero-Wangi, 2350 m. RPMJ Celis 16. xi.1954, 13 (IRSNB); Lubero, Katondi, 2200 m. N. Leleup 12.xii.1951, 132Ƥ (IRSNB); Mwenga, Luiko, 1900 m. N. Leleup 19. i.1952, 2 Ƥ (IRSNB); Uvira, Bassin Ulindi, 2650 m. N. Leleup 28. ix.1950, 1 Ƥ (IRSNB); Uvira, Mulenge, 2500 m. N. Leleup, viii.1959, 1 Ƥ (IRSNB). RWANDA: Forêt Rugege, 2200m, N. Leleup, iii. 1951, 2350m, P Basilewsky, 20/23.v. 1953132Ƥ (IRSNB).</p><p>Distribution and bionomics. Relatively frequently found in montane forest in eastern Congo and Rwanda (Fig. 144), where it is usually recorded in litter.</p><p>Comparative notes. Similar to A. congoensis in lacking bright yellow pubescence on the elytra. Most easily recognised by the narrow terminal segment of the maxillary palpi and the form of the aedeagus, in particular the incurving tips of the lobes of the paramere.</p></div>	https://treatment.plazi.org/id/4D54A843FFAC1A740694F97DED72FA22	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lott, Derek A.	Lott, Derek A. (2010): The species of Acylophorus Nordmann (Coleoptera: Staphylinidae: Staphylininae) in continental sub-Saharan Africa. Zootaxa 2402: 1-51, DOI: 10.5281/zenodo.275907
4D54A843FFAD1A770694F98DEDAFFBB4.text	4D54A843FFAD1A770694F98DEDAFFBB4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acylophorus mareei Bernhauer	<div><p>Acylophorus mareei Bernhauer</p><p>(Figs 16, 48, 66, 92, 125)</p><p>Acylophorus mareei Bernhauer, 1943: 290; Herman, 2001: 3034.</p><p>Redescription. Length 6–7mm. Body relatively wide and flat, black to dark brown with black to red-brown pronotum. Abdominal tergites iridescent. Legs, antennae, and labrum dark brown, middle segments of antennae sometimes paler. Proximal segments of maxillary palpi gradually paler than terminal segment.</p><p>Head of average size (pronotum 1.7x wider than head), as long as wide with evident, almost angled temples just behind very large eyes, slightly produced in front of antennal insertion, covered with dense micro-punctures (Fig. 16). Dense pale pubescence behind eyes. Two pairs of interocular setae arising from foveate punctures much closer to eyes than each other. Two lines of five postocular setae visible from above with no additional setae on hind margin of eyes. Mandibles with two large medial teeth on either side (Fig.</p><p>48). Maxillary palpi with terminal segment pubescent, more or less symmetric and elongate, longer than glabrous penultimate segment which is short and parallel-sided at apex (Fig. 66). First segment of antenna as long as next four. Segments I to VII elongate, IX to XI transverse (Fig. 92).</p><p>Pronotum slightly transverse (1.1x wider than long) with rounded sides and widest in basal half. Marginal setae short. Elytra transverse (1.6x wider than long) with close but relatively short pubescence. Apical bristles short, about the same length as the hairs on the rest of the elytra. Pubescence on abdominal tergites short, but overlapping comfortably on tergite III. Asperate punctures stronger than on elytra and fairly evenly distributed across each tergite.</p><p>Paramere bilobed, lobes proximate, parallel and narrowed to apex, pegs confusedly arranged right at apex, base lipped (Fig. 125). Median lobe of aedeagus expanded at apex and much longer than paramere.</p><p>Type material. Bernhauer described the species from specimens collected by Capt. Marée in March 1939 at Lokandu in what is now the Democratic Republic of Congo. Two specimens from the type series have been located, but in one the terminal abdominal segments are missing. The complete specimen is here designated as the lectotype in order to fix the identity of the species. Lectotype Ƥ: “ TYPUS A. Maréei / COLL. MUS. CONGO; Lokandu; - III-1939; Capt. Marée / R. DÉT. Q 4674 / Acylophorus maréei Brh. Typ. / LECTOTYPE Acylophorus mareei Bernhauer Ƥ det. DA Lott” (MRAC). Paralectotype Ƥ: “MUS. CONGO / Lokandu; - III- 1939; Capt. Marée / Acylophorus maréei Brh. Typ. / Maréei Brnh. Typus Acylophorus PARALECTOTYPE Acylophorus mareei Bernhauer Ƥ det. DA Lott” (FMNH).</p><p>Further material examined. R.D. CONGO: Bas-Congo: Boma, H Schouteden, ix.1920, 13 (ISRNB). SUDAN: Bahr el Ghazal: Wunatong, 8 O 30’N 28 O 30’E, 19. iii.1955, 17 (BMNH).</p><p>Distribution and bionomics. Only known from Congo and southern Sudan (Fig. 144). I have also seen a male (BMNH) from near Lake Bangweulu, Zambia, that may belong to this species. Unfortunately the aedeagus is missing. There are no ecological data.</p><p>Comparative notes. Very distinctive within the species group by virtue of the large eyes, the two medial teeth on each mandible and the narrow median lobe of the aedeagus. In addition, the short pubescence, which is not bright yellow on the elytra, will distinguish it from several species.</p></div>	https://treatment.plazi.org/id/4D54A843FFAD1A770694F98DEDAFFBB4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lott, Derek A.	Lott, Derek A. (2010): The species of Acylophorus Nordmann (Coleoptera: Staphylinidae: Staphylininae) in continental sub-Saharan Africa. Zootaxa 2402: 1-51, DOI: 10.5281/zenodo.275907
4D54A843FFAE1A790694FB1FECC0FDEC.text	4D54A843FFAE1A790694FB1FECC0FDEC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acylophorus methneri Bernhauer	<div><p>Acylophorus methneri Bernhauer</p><p>(Figs 17, 93)</p><p>Acylophorus methneri Bernhauer, 1917: 48; Scheerpeltz, 1933: 1467; Bernhauer &amp; Chapman, 1939: 74; Scheerpeltz, 1974: 25; Herman, 2001: 3034.</p><p>Description. Length 6.5mm. Body relatively wide and flat, pitchy with paler pronotum. Abdomen with weak iridescence. Legs and palpi pale, antennae mostly infuscate with pale terminal segments.</p><p>Head somewhat small (pronotum 1.8x wider than head) as long as wide, pigmented area produced in front of antennal insertion (Fig. 17). Side of head rounded with temples not very prominent. Covered with dense micro-punctures. Dense pale pubescence behind eyes. Four rather short postocular setae visible from above. Two pairs of short and pale interocular setae arising from shallow punctures less than twice as close to eyes as to each other. Maxillary palpi missing on material available for study. First segment of antenna longer than next four. Segments I to V elongate, VII to XI transverse (Fig. 93).</p><p>Pronotum slightly transverse (1.1x wider than long) with rounded sides and widest in basal half, micropunctures much sparser than on head. Dorsal and lateral setae short. Marginal setae short and reduced in number. Elytra transverse (1.7x wider than long) with short pubescence. Apical fringe of bristles short, slightly longer than hairs on the rest of the elytra. Punctures sparse and less obviously asperate than in other species. Abdominal tergites with short pubescence on basal segments, not overlapping on tergites III, IV and V, becoming longer on apical segments, punctures fine and dense throughout. Form of the aedeagus unknown.</p><p>Type material. Bernhauer described this species from a single specimen, which has been located. This specimen is the holotype by monotypy. Holotype Ƥ: “D. Ostafrika; Methner; Kaguru. v.09. Bach U. / Methneri Bernh. Typus unicus / Chicago NHMus M. Bernhauer Collection / HOLOTYPE Acylophorus methneri Bernhauer Ƥ det. DA Lott, 2009” (FMNH).</p><p>Distribution and bionomics. The type was collected from the banks of a stream at Kaguru, Tanzania (Fig. 143). There are further records in the literature from Kenya (Bernhauer &amp; Chapman, 1939) and Sudan (Scheerpeltz, 1974), but these must be open to question given the frequency of misidentifications in collections.</p><p>Comparative notes. Easily recognised by the extremely short pubescence on the elytra and abdomen and the short setae on the head and pronotum.</p></div>	https://treatment.plazi.org/id/4D54A843FFAE1A790694FB1FECC0FDEC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lott, Derek A.	Lott, Derek A. (2010): The species of Acylophorus Nordmann (Coleoptera: Staphylinidae: Staphylininae) in continental sub-Saharan Africa. Zootaxa 2402: 1-51, DOI: 10.5281/zenodo.275907
4D54A843FFA01A790694FDF8EFC5FC51.text	4D54A843FFA01A790694FDF8EFC5FC51.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acylophorus salifi	<div><p>A. salifi group</p><p>This group shares many characters with the A. orientalis group, but is easily distinguished by the small size of the head in comparison with the pronotum. The pronotum is 1.9x wider than the head or more. The temples are poorly developed and the neck relatively unconstricted. The penultimate segment of the maxillary palpi is short and triangular. The antennae are inserted close to or right on the front margin of the head, but there is a tendency for the genae to be produced forwards at each side of the front margin. The pubescence behind the eyes is sparse and localised. Micro-punctures on the head are absent or very sparse. Unlike in other species groups the empodial setae between the mid- and hind-tarsal claws are very short and hardly visible. The paramere is bilobed.</p></div>	https://treatment.plazi.org/id/4D54A843FFA01A790694FDF8EFC5FC51	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lott, Derek A.	Lott, Derek A. (2010): The species of Acylophorus Nordmann (Coleoptera: Staphylinidae: Staphylininae) in continental sub-Saharan Africa. Zootaxa 2402: 1-51, DOI: 10.5281/zenodo.275907
4D54A843FFA01A780694FBBAEC27FA7E.text	4D54A843FFA01A780694FBBAEC27FA7E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acylophorus salifi Lott	<div><p>Acylophorus salifi Lott, new species</p><p>(Figs 18, 46, 67, 94, 108, 126–128)</p><p>Description. Length 6 –7mm. Head black. Pronotum dark brown to black. Scutellum black. Elytra red-brown to black. Abdomen black, iridescent to varying degrees. Sometimes the body is all brown. Legs red. Antennae pale or dark with segment I pale. Maxillary palpi usually all pale.</p><p>Head much smaller than in A. orientalis (pronotum 2x wider than head), more or less as long as wide with temples barely suggested (Fig. 18). Antennae inserted right on front margin with no pigmented area in front. No micro-punctures visible at 80x magnification. Short pubescence behind eyes localised and much sparser than in A. orientalis . Two pairs of interocular setae arising from foveate punctures much closer to eyes than each other and five postocular setae visible from above on each side. No extra seta by hind margin of eye. Underside of head with sparse pubescence, weakly depressed at base (Fig. 108). Gular sutures continued to base of head, converging toward base but well separated throughout their length. Mandibles short with no medial teeth in front of a wide flange (Fig. 46). Maxillary palpi with terminal segment densely pubescent, markedly asymmetric, longer than short, triangular, glabrous penultimate segment (Fig. 67). First segment of antenna as long as next four (Fig. 94). Segments I to V elongate, VIII to XI transverse.</p><p>Pronotum transverse (1.4x wider than long) with rounded sides and widest in basal half. Shining with no micro-punctures. One pair of dorsal setae. One pair of lateral setae. Marginal setae long. Elytra strongly transverse (1.65x wider than long) with relatively short pubescence arising from fine, asperate punctures. Fringe of bristles on hind margin longer than the hairs on the rest of the elytra. Asperate punctures on abdominal tergites stronger than on elytra, denser toward base.</p><p>Paramere bilobed, each lobe weakly arched and well separated, pegs confusedly arranged in apical half of each lobe (Figs 126–128). Median lobe longer than paramere, barely expanded at apex which is rounded. There is considerable variation in the form of the aedeagus. The lobes of the paramere can be very slightly curving inward toward apex or slightly diverging. The apex of the median lobe can broadly rounded, sometimes with a central point, or more produced.</p><p>Type material. Holotype 3: “ BURKINA Naouri Nazinga, Bge d’Akouzina 11 O 9’N 1 O 36’W tussock DA Lott, 29.x.2003 / HOLOTYPE Acylophorus salifi sp. n. 3 det. DA Lott, 2009”. Paratypes 13 same data as holotype; 1Ƥ: “ BURKINA Naouri Nazinga, Bge de Barka, 11 O 8’N 1 O 37’W tussock DA Lott, 29.x.2003 / PARATYPE Acylophorus salifi sp. n. Ƥ det. DA Lott, 2009”; 1Ƥ: BURKINA Naouri Bge de Kouzougou, mud 11 O 9’N 1 O 32’W, DA Lott, 11.x.2004 / PARATYPE Acylophorus salifi sp. n. Ƥ det. DA Lott, 2009”; 332Ƥ: BURKINA Comoe Bge de Bounouna, 10 O 39’N 4 O 44’W, DA Lott, 20.x.2004 / PARATYPE Acylophorus salifi sp. n. det. DA Lott, 2009”. The holotype and paratypes have been placed in the collections of the BMNH.</p><p>This species was given the manuscript name, A. rufipennis by Bernhauer and a type was labelled accordingly, but he never published a description of the species. The name is now unavailable.</p><p>Further material examined. ANGOLA: Kundueje, R. Sanga, xi.1948, 13 (BMNH); no data, 1Ƥ (BMNH). BURKINA FASO: Comoe: Bge de Bounouna, 10 O 39’N 4 O 44’W, DA Lott, 20. x.2004, 14 (cBord, cJanak &amp; cLott). R.D. CONGO: Ituri: Bunia, Mt. Hoyo, 1200m, N Leleup, iii.1952 2 Ƥ (ISRNB); Katanga: Kundelungu, 1680m – 1750m, N Leleup, 8.iii.1950 – 25.x.1951, 432Ƥ (ISRNB); Kinshasa: cataracts, N Leleup, xii.1952 23 (ISRNB); Kivu: Masisi, Lac Ndalaga, 1800m, N Leleup, vi.1959 1 Ƥ (ISRNB); Masisi, Walikali, 700m, N Leleup, ix.1953 13 (ISRNB); Masisi, Mutakato, 800m, N Leleup, ix.1953 231Ƥ (ISRNB); Uvira, rives basses, Sanghe, 980m, N Leleup, 13.ix.1951 13 (ISRNB). CÔTE D’IVOIRE: Bingerville, J Decelle, ix.1961, 13 (IRSNB). ZAMBIA: R. Zambezi, Victoria Falls, N Leleup, vii.1960 13 (ISRNB); Abercorn, F. de la Mwengo, 1800m, N Leleup, vii.1960, 2 Ƥ (ISRNB). ZIMBABWE: Salisbury, GAK Marshall, vii.1896, 4 (BMNH).</p><p>Distribution and bionomics. A widespread species ranging from Burkina Faso to Angola, but not yet known from East Africa (Fig. 145). One of the more frequently collected species from the Congo catchment, where it has been found in “humus” in gallery forests as well as “roseaux morts” on a riverbank. Nearly all the specimens from Burkina Faso were collected by dissecting tussocks in well vegetated margins of large reservoirs constructed for irrigation or as waterholes for wildlife in the Nazinga Game Reserve.</p><p>Discussion. There are variations in both coloration and the form of the aedeagus (Figs 126–128). Most of this variation occurs between specimens taken from localities in the Congo catchment. Outside this area, most specimens are black with reddish elytra, sometimes with black front and side margins. The variations in the aedeagus occur both between different localities and to some extent within series taken from the same locality. Intermediate forms occur between the extremes of each type of variation, but some populations may be referable to distinct taxa once sufficient material from further localities becomes available for study.</p><p>Etymology. This species is named after Salif Mohammed Ouedraogo of Ouagadougou, driver extraordinaire, who accompanied me on my expeditions in Burkina Faso. The species name is the genitive case of a Latinised version of his name.</p></div>	https://treatment.plazi.org/id/4D54A843FFA01A780694FBBAEC27FA7E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lott, Derek A.	Lott, Derek A. (2010): The species of Acylophorus Nordmann (Coleoptera: Staphylinidae: Staphylininae) in continental sub-Saharan Africa. Zootaxa 2402: 1-51, DOI: 10.5281/zenodo.275907
4D54A843FFA11A7A0694FA49EAFCFA49.text	4D54A843FFA11A7A0694FA49EAFCFA49.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acylophorus lomaensis Levasseur	<div><p>Acylophorus lomaensis Levasseur</p><p>(Figs 19, 47, 68, 95, 129, 130)</p><p>Acylophorus lomaensis Levasseur, 1971: 303; Herman, 2001: 3033.</p><p>Diagnostic characters. Levasseur’s description contains illustrations of the aedeagus and antenna in addition to a textual description of morphological features. The following section contains additional diagnostic features as well as variations exhibited by further specimens studied.</p><p>Length varies from 5–7mm. Body often all brown with head dark brown. Abdominal tergites iridescent. Antennae often all pale with apical segments of antennae almost white.</p><p>Head small (pronotum 1.9x wider than head), more or less as long as wide with temples barely suggested (Fig. 19). Antennae inserted right on front margin. Genae well developed and produced at sides of front margin. Micro-punctures extremely sparse. Short pubescence behind eyes localised and very sparse. Two pairs of interocular setae arising from foveate punctures much closer to eyes than each other. Five postocular setae visible from above on each side. No extra seta by hind margin of eye. Underside of head weakly depressed at base. Gular sutures continued to base of head, converging toward base but well separated throughout their length. Mandibles without medial teeth, similar to A. salifi (Fig. 47). Maxillary palpi with terminal segment densely pubescent, markedly asymmetric, longer than short, triangular, penultimate segment, which is glabrous (Fig. 68). First segment of antenna nearly as long as next four (Fig. 95). Segments I to V elongate, VII to XI transverse.</p><p>Pronotum less transverse than in A. salifi (1.2x wider than long) with rounded sides and widest in basal half. Shining with no micro-punctures. Elytra strongly transverse (1.6x wider than long). Fringe of bristles on hind margin longer than the hairs on the rest of the elytra. Asperate punctures on abdominal tergites stronger than on elytra, denser toward base.</p><p>There is some variation in the form of the aedeagus. In most specimens, including the paratype that I have seen, each lobe is weakly arched and proximate and the pegs confusedly arranged in apical half of each lobe (Figs 129 &amp; 130). The median lobe is longer than the paramere and expanded at the apex which is rounded.</p><p>Type material. Paratype 3: “Mission ENS-IFAN aux monts LOMA / Ravin SE du Camp II – 1600 m 8- 1-1966 / tamisage bord ruisseau / Acylophorus lomaensis sp. n. LEVASSEUR DET 1967 PARATYPE ” (MNHN).</p><p>Further material examined. R.D. CONGO: Bas Congo: Kisantu, P. Basilewsky, 29–30. xi.1952 1 Ƥ (ISRNB); Equateur: Coquilhatville, N Leleup, x.1959, 2 Ƥ (ISRNB); Kivu: Kalehe, Bunyakiri, 1050m, N Leleup, ix.1953 131Ƥ (ISRNB); Masisi, Walikali, 700m, N Leleup, ix.1953 13 (ISRNB); Masisi, Mutakato, 800m, N Leleup, ix.1953 231Ƥ (ISRNB); Uvira, rives basses, Sanghe, 980m, N Leleup, 13.ix.1951 13 (ISRNB); Tshuapa: Bikoro, Mabali, Lac Tumba, N Leleup, 29.ix.1955 – ix.1959, 35 (ISRNB); Ikela, riv. Lukendu, N Leleup, viii.1959 13 (ISRNB); Ikela, vill. Besoke, N Leleup, ix.1959 1 Ƥ (ISRNB). R. CONGO: Brazzaville, Forêt Pattes d’Oie, J.P. Adam, x.1960 2 Ƥ (ISRNB).</p><p>Distribution and bionomics. Apart from the type series from Sierra Leone, all material seen so far comes from the Congo catchment (Fig. 146). Habitats range from forest humus to clay lake margins and stream banks.</p><p>Comparative notes. Often smaller than A. salifi with a less transverse pronotum, smaller eyes, more developed genae and a distinctive aedeagus with narrower, more proximate lobes to the paramere.</p><p>Discussion. Specimens from the Congo catchment are smaller (5–6mm) than those from Sierra Leone (6– 7mm).There are variations in the form of the aedeagus (Figs 129 &amp; 130). Some populations may be referable to distinct taxa once sufficient material from further localities becomes available for study.</p></div>	https://treatment.plazi.org/id/4D54A843FFA11A7A0694FA49EAFCFA49	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lott, Derek A.	Lott, Derek A. (2010): The species of Acylophorus Nordmann (Coleoptera: Staphylinidae: Staphylininae) in continental sub-Saharan Africa. Zootaxa 2402: 1-51, DOI: 10.5281/zenodo.275907
4D54A843FFA31A7D0694FA52EC83FC2C.text	4D54A843FFA31A7D0694FA52EC83FC2C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acylophorus minor Lott	<div><p>Acylophorus minor Lott, new species</p><p>(Figs 20, 69, 96, 131)</p><p>Description. Length 5–5.5mm. Body dark brown with pronotum and apical margins of abdominal tergites paler. Abdominal tergites iridescent. Appendages red-brown, apical segments of antennae yellow.</p><p>Head small (pronotum 2x wider than head), more or less as long as wide with temples barely suggested (Fig. 20). Antennae inserted right on front margin with no pigmented area in front. Genae well developed and produced at sides of front margin. No micro-punctures visible at 80x magnification. Short pubescence behind eyes localised and very sparse. Two pairs of interocular setae arising from foveate punctures much closer to eyes than each other and five postocular setae visible from above on each side. No extra seta by hind margin of eye. Underside of head weakly depressed at base. Gular sutures continued to base of head, converging toward base but well separated throughout their length. Mandibles similar to A. salifi . Maxillary palpi with terminal segment densely pubescent, markedly asymmetric, longer than short, triangular, penultimate segment, which is glabrous (Fig. 69). First segment of antenna longer than next four (Fig. 96). Segments I to V elongate, VII to XI transverse.</p><p>Pronotum less transverse than in A. salifi (1.2x wider than long) with rounded sides and widest in basal half. Shining with no micro-punctures. One pair of dorsal setae. One pair of lateral setae. Marginal setae long. Elytra strongly transverse (1.8x wider than long) with pubescence arising from fine, asperate punctures. Fringe of bristles on hind margin longer than the hairs on the rest of the elytra. Asperate punctures on abdominal tergites stronger than on elytra, denser toward base.</p><p>Paramere bilobed, each lobe weakly arched and curved inwards at apex, pegs confusedly arranged in apical half of each lobe (Fig. 131). Median lobe longer than paramere, expanded at apex which is rounded.</p><p>Type material. Holotype 3: “ CÔTE D’IVOIRE Riv. Sassandra / MUSÉUM PARIS 12-1930- IV-1931 P.A. CHAPUIS / Acylophorus Pauliani Brh Typ / Chicago NHMUS M. Bernhauer Collection / HOLOTYPE Acylophorus minor sp. n. 3 det. DA Lott, 2009” (FMNH). Paratypes 13: “K431 OPRS NIGERIA 1951 / PARATYPE Acylophorus minor sp. n. 3 det. DA Lott, 2009” (MMUM); 1Ƥ: “K700 IBADAN NIGERIA 21 4 53 / PARATYPE Acylophorus minor sp. n. 3 det. DA Lott, 2009” (MMUM); 1Ƥ: “K711 IBADAN NIGERIA 14 4 53 / PARATYPE Acylophorus minor sp. n. 3 det. DA Lott, 2009” (MMUM).</p><p>This species was given the manuscript name, A. pauliani by Bernhauer and a type was labelled accordingly, but he never published a description of the species.</p><p>Distribution and bionomics. Only known from four specimens collected in Côte d’Ivoire and Nigeria (Fig. 146). There are no ecological data.</p><p>Comparative notes. Similar to A. lomaensis . Distinguished by the form of the aedeagus.</p><p>Etymology. The specific name is the masculine form of the comparative adjective meaning “lesser” and refers to the relatively small size of the species.</p></div>	https://treatment.plazi.org/id/4D54A843FFA31A7D0694FA52EC83FC2C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lott, Derek A.	Lott, Derek A. (2010): The species of Acylophorus Nordmann (Coleoptera: Staphylinidae: Staphylininae) in continental sub-Saharan Africa. Zootaxa 2402: 1-51, DOI: 10.5281/zenodo.275907
4D54A843FFA41A7D0694FBB8EFB9FB2A.text	4D54A843FFA41A7D0694FBB8EFB9FB2A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acylophorus trigonocephalus	<div><p>A. trigonocephalus group</p><p>This group is characterised by the two terminal pubescent segments on each maxillary palpus, three pairs of interocular setae and additional marginal setae on the pronotum. The paramere can be either bilobed or produced into one single lobe.</p></div>	https://treatment.plazi.org/id/4D54A843FFA41A7D0694FBB8EFB9FB2A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lott, Derek A.	Lott, Derek A. (2010): The species of Acylophorus Nordmann (Coleoptera: Staphylinidae: Staphylininae) in continental sub-Saharan Africa. Zootaxa 2402: 1-51, DOI: 10.5281/zenodo.275907
4D54A843FFA41A7C0694FAB5EF59FB92.text	4D54A843FFA41A7C0694FAB5EF59FB92.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acylophorus trigonocephalus Cameron	<div><p>Acylophorus trigonocephalus Cameron</p><p>(Figs 21, 70, 97, 135)</p><p>Acylophorus trigonocephalus Cameron, 1949: 322; Herman, 2001: 3037.</p><p>Acylophorus leonensis Bordoni, 1994: 312; Herman, 2001: 3033 (new synonym).</p><p>Redescription. Length 6.5mm. Body black with iridescent abdomen, pronotum sometimes slightly paler. I have also seen one specimen that is completely pale brown. Antennae brown to dark brown, first segment often paler. Legs pale, sometimes with darkened femora and tibiae. Palpi all pale.</p><p>Head small in comparison with pronotum (pronotum 1.9x wider than head) more or less as long as wide with well developed temples behind large eyes (Fig. 21). Head pigmented in front of antennal insertion. Micro-punctures very sparse and scarcely detectable at 40x magnification. Short pubescence behind eyes. Three pairs of interocular setae and a line of five postocular setae just visible from above on each side. Maxillary palpi with last two segments pubescent, terminal segment elongate, rounded on both sides, but slightly asymmetric, longer than penultimate segment which is also elongate (Fig. 70). First segment of antenna longer than next five. Segments I to IV elongate, IX to X transverse (Fig. 97).</p><p>Pronotum transverse (1.3x wider than long) with rounded sides, widest in basal half. Shining with no micro-punctures. Marginal setae very long and more numerous than in A. orientalis . One pair of dorsal and one pair of lateral setae. Elytra transverse (1.65x wider than long) with long but sparse pubescence and coarse asperate punctures which are stronger than in A. orientalis . Apical bristles much longer than hairs on the rest of the elytra. Abdominal tergites also with long, sparse pubescence. Punctures much sparser on apical half of each tergite than on basal half. Apical fringe of bristles on each tergite of two distinct lengths, the shorter bristles more numerous than the longer bristles.</p><p>Male sternite IX with apex simple. Aedeagus with bilobed paramere, each lobe narrow, parallel, pegs confusedly arranged toward apex, which has numerous short marginal hairs (Fig. 135). Median lobe with apex barely expanded, only just surpassing paramere.</p><p>Type material. Cameron described A. trigonocephalus from material collected in Côte d’Ivoire. One syntype has been located and is here designated as the lectotype in order to fix the identity of the species. Lectotype Ƥ: “IFAN-1946 TONKUI C.I. 900–1200m A. VILLIERS / FORET PRIM; 20-30-IX / M. Cameron Bequest B.M. 1955-147 / Dr. M. Cameron det., 1948; Acylophorus trigonocephalus Cam. COTYPE / LECTOTYPE Acylophorus trigonocephalus Cameron Ƥ det. DA Lott” (BMNH).</p><p>Acylophorus leonensis Bordoni. Holotype 3: “S. Leone W. Area Sugar Loaf Mt. Rossi 7.3.92 / HOLOTYPUS / HOLOTYPUS Acylophorus leonensis sp. n. Bordoni det. 1992” (cBord). In the original description Bordoni compared his new species to A. rossii, but not to A. trigonocephalus with which it is similar in all respects. Consequently, A. leonensis is here synonymised with A. trigonocephalus .</p><p>Further material examined. R.D. CONGO: Kivu: Kabari, SE Kahuzi, 2080/ 2200m, N Leleup, viii.1951, 1 Ƥ (ISRNB); Mwenga, Luiko, 2050m, N Leleup, 21. i.1952, 1 Ƥ (IRSNB). SIERRA LEONE: Northern Province: Sinikoro, W Rossi, 31. xii.1982, 13 (cBord). ZIMBABWE: Kutsaga near Harare Airport, W Rossi, 18. vi.1997, 13 (cLott).</p><p>Distribution and bionomics. Recorded from Sierra Leone, Côte d’Ivoire, Congo and Zimbabwe, always in small numbers (Fig. 147). Recorded habitats include primary forest, montane forest and the margins of a fast-flowing stream.</p><p>Comparative notes. Easily distinguished from A. orientalis by the characters common to the species group, two pubescent terminal segments on the maxillary palpi, the three pairs of interocular punctures and the larger number of marginal setae. Within the group it can be recognised by a combination of the long setae on the head and pronotum, the large eyes and the moderately transverse elytra. The aedeagus is also distinctive.</p></div>	https://treatment.plazi.org/id/4D54A843FFA41A7C0694FAB5EF59FB92	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lott, Derek A.	Lott, Derek A. (2010): The species of Acylophorus Nordmann (Coleoptera: Staphylinidae: Staphylininae) in continental sub-Saharan Africa. Zootaxa 2402: 1-51, DOI: 10.5281/zenodo.275907
4D54A843FFA51A7F0694FB7DEFE4FDC9.text	4D54A843FFA51A7F0694FB7DEFE4FDC9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acylophorus setiger Lott	<div><p>Acylophorus setiger Lott, new species</p><p>(Figs 22, 71, 98, 136)</p><p>Description. Length 6mm. Body brown, head darker. Legs pale. Antennae pale with middle segments darker. Palpi all pale.</p><p>Head of average size (pronotum 1.7x wider than head), 1.1x wider than long with well developed temples behind large eyes (Fig. 22). Pigmented area of head extends in front of antennal insertion. Micro-punctures very sparse and concentrated toward front of head. Short pubescence behind eyes. Three pairs of interocular setae and a line of four postocular setae visible from above on each side. Maxillary palpi with two terminal segments pubescent, terminal segment very elongate, slightly asymmetric, longer than penultimate segment, which is also elongate (Fig. 71). First segment of antenna as long as next five. Segments I to III elongate, VII to X transverse (Fig. 98).</p><p>Pronotum slightly transverse (1.2x wider than long) with sides well rounded, widest in basal half. Shining with no micro-punctures. One pair of dorsal setae and one pair of lateral setae. Marginal setae long and more numerous than in A. orientalis . Elytra very transverse (1.9x wider than long) with sparse pubescence and coarse asperate punctures which are stronger than in A. orientalis . Apical bristles much longer than hairs on the rest of the elytra. Abdominal tergites also with long, sparse pubescence. Punctures much sparser on apical half of each tergite than on basal half. Apical fringe of bristles on each tergite of two distinct lengths, the shorter bristles more numerous than the longer bristles, as in A. trigonocephalus .</p><p>Male sternite IX with apex entire. Aedeagus with simple paramere, pegs arranged around margin with one or two scattered inside (Fig. 136). Median lobe with apex barely expanded, only just surpassing paramere.</p><p>Type material. Holotype 3: “ Paratype / Mioko Fernando Poo 1700–2000m 8.XII.51 / De Keyser Lopdome et A. Villiers / PARATYPE / M. Cameron Bequest B.M. 1955-147 / Acylophorus brevipennis Cam. Cotype / HOLOTYPE Acylophorus setiger sp. n. 3 det. DA Lott, 2009” (BMNH).</p><p>Note that Cameron gave this species a manuscript name brevipennis and labelled types accordingly, but he never published a description.</p><p>Distribution and bionomics. Only known so far from a single specimen taken on the island of Bioko (formerly known as Fernando Po) in Equatorial Guinea (Fig. 147). There are no data on habitat.</p><p>Comparative notes. Similar to A. trigonocephalus, but with short, very transverse elytra and distinctive aedeagus with paramere that is not bilobed. The paler colour may also be diagnostic.</p><p>Etymology. The specific name is the masculine form of an adjective meaning “bristly” and refers to the long setae on the body.</p></div>	https://treatment.plazi.org/id/4D54A843FFA51A7F0694FB7DEFE4FDC9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lott, Derek A.	Lott, Derek A. (2010): The species of Acylophorus Nordmann (Coleoptera: Staphylinidae: Staphylininae) in continental sub-Saharan Africa. Zootaxa 2402: 1-51, DOI: 10.5281/zenodo.275907
4D54A843FFA61A7F0694FDDDEF4AF951.text	4D54A843FFA61A7F0694FDDDEF4AF951.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acylophorus usambarae Bernhauer	<div><p>Acylophorus usambarae Bernhauer</p><p>(Figs 23, 72, 99)</p><p>Acylophorus usambarae Bernhauer, 1917: 48; Scheerpeltz, 1933: 1467; Herman, 2001: 3038.</p><p>Redescription. Length 7mm. Body brown with head darker. Legs and palpi pale, antennae brown with paler terminal segments.</p><p>Head large (pronotum 1.5x wider than long) slightly transverse (1.1x wider than long), temples well developed (Fig. 23). Area in front of antennal insertion pigmented. Micro-punctures very sparse. Short dense pubescence behind eyes. Three pairs of interocular setae and a line of five postocular setae visible from above on each side. Maxillary palpi with last two segments densely pubescent, terminal segment much more rounded on outer margin than inner margin, so markedly asymmetric, longer than penultimate segment which is also elongate (Fig. 72). First segment of antenna as long as next five. Segments I to V elongate, VIII to XI transverse (Fig. 99).</p><p>Pronotum transverse (1.2x wider than long) with rounded sides and widest at centre. Shining with no micro-punctures. One pair of dorsal and one pair of lateral setae. Marginal setae long and more numerous than in A. orientalis . Elytra transverse (1.6x wider than long) with long pubescence, asperate punctures stronger but less dense than in A. orientalis . Apical fringe of bristles much longer than hairs on the rest of the elytra. Scutellum markedly less strongly punctured than elytra. Abdominal tergites with very long and semi-erect pubescence, asperate punctures much weaker and sparser than on elytra. Punctures on apical tergites much sparser than on proximal segments. Apical fringe of bristles on each tergite of equal length. The form of the aedeagus is unknown.</p><p>Type material. Bernhauer described this species from a single specimen, which has been located. This specimen is the holotype by monotypy. Holotype Ƥ: “West Usambara; II 12 Methner / D. Ostafrika; Methner / Usambarae Bernh. Typus unic / Chicago NHMus M. Bernhauer Collection / HOLOTYPE Acylophorus usambarae Bernhauer Ƥ det. DA Lott, 2009” (FMNH).</p><p>Distribution and bionomics. Only known from the unique type collected in the Usambara Mountains, Tanzania (Fig. 147).</p><p>Comparative notes. Easily recognisable within the species group by the long and outstanding hairs on the abdomen.</p></div>	https://treatment.plazi.org/id/4D54A843FFA61A7F0694FDDDEF4AF951	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lott, Derek A.	Lott, Derek A. (2010): The species of Acylophorus Nordmann (Coleoptera: Staphylinidae: Staphylininae) in continental sub-Saharan Africa. Zootaxa 2402: 1-51, DOI: 10.5281/zenodo.275907
4D54A843FFA61A7E0694F8A5EC22FB4C.text	4D54A843FFA61A7E0694F8A5EC22FB4C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acylophorus grandis Bernhauer	<div><p>Acylophorus grandis Bernhauer</p><p>(Figs 24, 73, 100, 134)</p><p>Acylophorus grandis Bernhauer, 1929: 124; Scheerpeltz, 1933: 1466; Herman, 2001: 3031.</p><p>Redescription. Length 9.5–10.5mm. Body black. Legs, antennae and palpi dark brown. Head somewhat large (pronotum 1.6x wider than pronotum) more or less as wide as long, temples rounded, with pigmented area of head extending in front of antennal insertion (Fig. 24). Micro-punctures very sparse. Short pubescence behind eyes. Three pairs of interocular setae. A line of five postocular setae visible from above on each side. Maxillary palpi with two terminal segments pubescent, terminal segment very elongate, slightly asymmetric, longer than penultimate segment, which is also elongate (Fig. 73). First segment of antenna as long as next five (Fig. 100). Segments I to III elongate, X transverse.</p><p>Pronotum almost as long as broad (1.03x wider than long) with rounded sides and widest in basal half. Shining with no micro-punctures. One pair of dorsal and one pair of lateral setae. Marginal setae very long and more numerous than in A. orientalis . Elytra transverse (1.75x wider than long) with long pubescence, punctures normal, not asperate, less dense than A. orientalis . Apical fringe of bristles shorter than hairs on rest of elytra. Hind margins more diagonal than in other species, so forming a deeper “V”. Abdominal tergites also with long pubescence, with slightly asperate punctures much sparser on apical half of tergites than on basal half. Apical fringe of bristles on each tergite of equal length.</p><p>Male sternite IX with apex entire. Aedeagus with paramere bilobed, each lobe short and expanded medially, base lipped (Fig. 134). Medial lobe slightly longer than paramere, somewhat flattened with truncate apex.</p><p>Type material. Bernhauer described the species from specimens collected by Gérard at Niemba – Kalembe in 1918. Two specimens from the type series have been located and one of these is here designated as the lectotype in order to fix the identity of the species. Lectotype 3: “ TYPE / MUSÉE DU CONGO; MANYEMA 1918 Niemba – Kalembe; Dr. Gérard / R. DÉT. O 879 / Acylophorus grandis Bernh. Typ. / LECTOTYPE Acylophorus grandis Bernhauer 3 det. DA Lott ” (MRAC). Paralectotype Ƥ: “ MUSÉE DU CONGO / MANYEMA 1918 Niemba - Kalembe / Dr. Gérard / grandis Brh. Typ / Acylophorus grandis Brh. Typus./ Chicago NHMus M. Bernhauer Collection / PARALECTOTYPE Acylophorus grandis Bernhauer Ƥ det. DA Lott” (FMNH).</p><p>Further material examined. R.D. CONGO: Haut-Uelé: Watsa, L. Burgeon, 1922, 13 (MRAC).</p><p>Distribution and bionomics. Known from two localities in the Democratic Republic of Congo (Fig. 147). There are no ecological data.</p><p>Comparative notes. Instantly recognisable by the large size, long setae, quadrate pronotum, simple punctures and the distinctive aedeagus.</p></div>	https://treatment.plazi.org/id/4D54A843FFA61A7E0694F8A5EC22FB4C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lott, Derek A.	Lott, Derek A. (2010): The species of Acylophorus Nordmann (Coleoptera: Staphylinidae: Staphylininae) in continental sub-Saharan Africa. Zootaxa 2402: 1-51, DOI: 10.5281/zenodo.275907
4D54A843FFA71A410694FB58EB0FFE32.text	4D54A843FFA71A410694FB58EB0FFE32.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acylophorus bafutensis Levasseur	<div><p>Acylophorus bafutensis Levasseur</p><p>(Figs 25, 49, 74, 101, 137)</p><p>Acylophorus bafutensis Levasseur, 1968: 1401; Herman, 2001: 3026.</p><p>Diagnostic characters. Levasseur’s description contains illustrations of the aedeagus and antenna in addition to a textual description of morphological features. The following section contains additional diagnostic features.</p><p>Length 7mm. Body black. Abdomen and tibiae very weakly iridescent. Appendages dark brown with paler tarsi.</p><p>Head of average size (pronotum 1.7x wider than head), 1.1x wider than long, temples well developed (Fig. 25). Micro-punctures sparse. Short dense pubescence behind eyes. Three pairs of interocular punctures. Setae very short, postocular setae barely visible. Mandibles with one medial tooth on right and simply flanged on left (Fig.49). Maxillary palpi with two terminal segments densely pubescent, terminal segment broad, slightly asymmetric, longer than penultimate segment (Fig. 74). First segment of antenna nearly as long as next five (Fig. 101). Segments I to V elongate, VIII to X transverse.</p><p>Pronotum transverse (1.15x wider than long) with rounded sides and widest at centre. Shining with no micro-punctures. One pair of dorsal punctures and one pair of lateral punctures. Marginal punctures present, but setae very short. Elytra transverse (1.6x wider than long) with sparse pubescence arising from fine, weakly asperate punctures. Apical fringe of bristles longer than hairs on rest of elytra. Scutellum markedly less strongly punctured than elytra. Abdominal tergites with sparse pubescence. Punctures on apical areas of tergites much sparser than on basal areas. Apical fringe of bristles on each tergite missing in centre (possibly deciduous or abraded as associated crenulations are present).</p><p>Aedeagus with bilobed paramere, each lobe with pegs confusedly arranged in distal half (Fig. 137). Median lobe shorter than paramere, emarginate at apex.</p><p>Type material. Paratype Ƥ: “Ft de BAFUT NGUEMBA 16.VIII.65 / Muséum Paris CAMEROUN B. de Miré / Acylophorus bafutensis n. sp. LEVASSEUR DET 1967 PARATYPE ” (MNHN).</p><p>Distribution and bionomics. Only known from three specimens collected in the Bafut Forest, Cameroon (Fig. 147). There are no ecological data.</p><p>Comparative notes. Easily recognised within the species group by the short setae and pubescence.</p></div>	https://treatment.plazi.org/id/4D54A843FFA71A410694FB58EB0FFE32	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lott, Derek A.	Lott, Derek A. (2010): The species of Acylophorus Nordmann (Coleoptera: Staphylinidae: Staphylininae) in continental sub-Saharan Africa. Zootaxa 2402: 1-51, DOI: 10.5281/zenodo.275907
4D54A843FF981A410694FD9DECE0FCEA.text	4D54A843FF981A410694FD9DECE0FCEA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acylophorus kambuiensis	<div><p>A. kambuiensis group</p><p>This group is characterised by the semi-erect pubescence covering the pronotum and most of the head. In addition the terminal segment of the maxillary palpi is glabrous and the apex of each style of tergite IX is devoid of bristles and thus easily visible. Like the A. orientalis group, there are two pairs of interocular setae. The paramere is produced into a single lobe.</p></div>	https://treatment.plazi.org/id/4D54A843FF981A410694FD9DECE0FCEA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lott, Derek A.	Lott, Derek A. (2010): The species of Acylophorus Nordmann (Coleoptera: Staphylinidae: Staphylininae) in continental sub-Saharan Africa. Zootaxa 2402: 1-51, DOI: 10.5281/zenodo.275907
4D54A843FF981A430694FCF5ECFBF8BC.text	4D54A843FF981A430694FCF5ECFBF8BC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acylophorus kambuiensis Bordoni	<div><p>Acylophorus kambuiensis Bordoni</p><p>(Figs 26, 75, 102, 138)</p><p>Acylophorus kambuiensis Bordoni, 1994: 312 .</p><p>Acylophorus kumbuiensis: Herman, 2001: 3032 (misspelling).</p><p>Diagnostic characters. Bordoni’s description contains illustrations of the aedeagus, head, maxillary palpi and posterior tarsi in addition to a textual description of morphological features. The following section contains additional diagnostic features as well as variations exhibited by further specimens studied.</p><p>Length 8–9mm (The figure in the original description of 10 mm for overall body length does not agree with my own measurements of type material). Body black, pronotum and hind margins of tergites brown. Abdomen very weakly iridescent. Legs pale. Antennae dark with apical segments conspicuously pale. The maxillary palpi are all pale. Labrum black.</p><p>Head somewhat large (pronotum 1.6x wider than head), slightly transverse (1.1x wider than long) with well developed temples and antennal insertion close to front margin with narrow pigmented strip in front (Fig. 26). Pubescence extending over most of head leaving only a central glabrous strip free of punctures. Two pairs of interocular setae. Five postocular setae visible, the posterior puncture and seta displaced forward and out of line. No additional short seta on hind margin of eye. Maxillary palpi with terminal segment elongate, glabrous, asymmetric and attenuated at apex (Fig. 75). First segment of antenna as long as next five. Segments I to VIII elongate, X transverse (Fig. 102).</p><p>Pronotum only slightly transverse (1.2x wider than long) with rounded sides and widest in basal half. Covered with somewhat sparse semi-erect pubescence. Dorsal setae absent. One pair of lateral setae. Marginal setae shorter than in A. orientalis . Elytra transverse (1.55x wider than long) with pubescence arising from strong, somewhat close asperate punctures. Apical fringe of thick bristles shorter than the hairs on the rest of the elytra. Abdominal tergites with long, somewhat outstanding pubescence arising from asperate punctures that are much denser at the base of each tergite than the apical half.</p><p>Paramere simply lobed, pegs confusedly arranged at apex with blank area right at the tip in the centre (Fig. 138). Median lobe with rather narrow, very flat apex, only just surpassing paramere.</p><p>Type material. Paratype 3: “S. Leone W. area Sugar Loaf Mt. Rossi 7.3.92 / PARATYPUS Acylophorus kambuiensis sp. n. Bordoni det. 1992” (cBord).</p><p>This species was given the manuscript name, A. leonensis, by Cameron. A type was labelled accordingly, but he never published a description.</p><p>Further material examined. SIERRA LEONE: Eastern Province: near Kenema, Kambui Hills, W Rossi, 27–29. x.1989, 13 (teneral) (cLott); Western Area: Freetown, no date, 3Ƥ (BMNH). Distribution and bionomics. All records so far come from Sierra Leone (Fig. 148), where it has been found on the banks of streams.</p><p>Comparative notes. Distinguished from A. lualabaensis by the larger eyes, the pattern of the pubescence on the pronotum and the form of the aedeagus.</p></div>	https://treatment.plazi.org/id/4D54A843FF981A430694FCF5ECFBF8BC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lott, Derek A.	Lott, Derek A. (2010): The species of Acylophorus Nordmann (Coleoptera: Staphylinidae: Staphylininae) in continental sub-Saharan Africa. Zootaxa 2402: 1-51, DOI: 10.5281/zenodo.275907
4D54A843FF9B1A4D0694F914EF4AFF39.text	4D54A843FF9B1A4D0694F914EF4AFF39.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acylophorus lualabaensis Levasseur	<div><p>Acylophorus lualabaensis Levasseur</p><p>(Figs 27, 50, 76, 103, 139)</p><p>Acylophorus lualabaensis Levasseur, 1968: 1404; Herman, 2001: 3033.</p><p>Diagnostic characters. Levasseur’s description contains illustrations of the aedeagus and antenna in addition to a textual description of morphological features. The following section contains additional diagnostic features.</p><p>Length 9mm. Body black with slightly paler. Abdomen very weakly iridescent. Legs pale with tibiae darkened. Antennae black with apical segments conspicuously paler. Maxillary palpi all pale. Labrum dark brown with pale margins.</p><p>Head large (pronotum 1.5x wider than head), 1.05x wider than long, temples well developed behind relatively small eyes (Fig. 27). Pubescence extending over most of head leaving only a central glabrous strip free of punctures. Two pairs of interocular setae. Five postocular setae visible, the posterior puncture and seta displaced forward and out of line. No additional short seta on hind margin of eye. Mandibles with three medial teeth on right and two on left (Fig. 50). Maxillary palpi with terminal segment elongate, glabrous, asymmetric and attenuated at apex (Fig. 76). First segment of antenna almost as long as next five (Fig. 103). Segments I to IX elongate, X transverse.</p><p>Pronotum slightly transverse (1.1x wider than long) with rounded sides and widest in basal half. Somewhat sparse semi-erect pubescence extending over most of surface except for central longitudinal glabrous strip and apical band. Dorsal setae absent. One pair of lateral setae. Marginal setae apparently short and reduced in number, though setae are possibly abraded in material available for study. Elytra transverse (1.6x wider than long) with pubescence arising from somewhat close asperate punctures. Apical fringe of thick bristles shorter than the hairs on the rest of the elytra. Abdominal tergites with long, somewhat outstanding pubescence arising from asperate punctures that are much denser at the base of each tergite than in the apical half.</p><p>Paramere simply lobed, pegs confusedly arranged at apex (Fig. 139). Median lobe narrowed at rounded apex, easily surpassing paramere.</p><p>Type material. Holotype 3: “KOLWEZI H. KATANGA COLL LEVASSEUR / Zilo Walaba / CONGO BELGE 4.1959 D.V. ALLARD / Acylophorus lualabaensis n. sp. LEVASSEUR DET 1967 HOLOTYPE / 17” (MNHN).</p><p>Distribution and bionomics. Only known from a single specimen collected in the Democratic Republic of Congo (Fig. 148). There are no ecological data.</p><p>Comparative notes. Similar to A. kumbuiensis, but distinguished by the smaller eyes and the form of the aedeagus.</p></div>	https://treatment.plazi.org/id/4D54A843FF9B1A4D0694F914EF4AFF39	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lott, Derek A.	Lott, Derek A. (2010): The species of Acylophorus Nordmann (Coleoptera: Staphylinidae: Staphylininae) in continental sub-Saharan Africa. Zootaxa 2402: 1-51, DOI: 10.5281/zenodo.275907
4D54A843FF941A4D0694FECAEB6DFCE8.text	4D54A843FF941A4D0694FECAEB6DFCE8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acylophorus rufipennis Cameron	<div><p>Acylophorus rufipennis Cameron</p><p>Acylophorus rufipennis Cameron, 1959: 117; Herman, 2001: 3037.</p><p>Cameron described this species in a posthumous publication from two specimens collected by A. Machado from the River Luachimo in Angola. The description is not detailed enough to recognise the species without examination of type material. Two specimens collected from Angola and labelled as types of A. rufipennis are present in the Cameron collection at the BMNH, but the locality on the labels does not match the type locality and they differ markedly in colour and eye size from the description given by Cameron. No matching specimens from Angola can be found in his collection. The two specimens labelled as types are in fact a pale colour form of A. salifi . The true identity of Cameron’s species will remain a matter of speculation, unless the original type material is located. The name is therefore considered to be a nomen dubium.</p></div>	https://treatment.plazi.org/id/4D54A843FF941A4D0694FECAEB6DFCE8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lott, Derek A.	Lott, Derek A. (2010): The species of Acylophorus Nordmann (Coleoptera: Staphylinidae: Staphylininae) in continental sub-Saharan Africa. Zootaxa 2402: 1-51, DOI: 10.5281/zenodo.275907
