taxonID	type	description	language	source
4D76240CFFBDFFC662EBFCC3FBFAFAA1.taxon	description	Figures 1 – 8	en	Martínez, Leonel, Kochalka, John A., Cabra-Garcia, Jimmy, Ramírez, Martin (2025): Revealing the identity of Josa chazaliae (Simon, 1897) (Araneae: Anyphaenidae): new species and the highest altitude record for a spider in South America. Zootaxa 5566 (2): 201-242, DOI: 10.11646/zootaxa.5566.2.1, URL: https://doi.org/10.11646/zootaxa.5566.2.1
4D76240CFFBDFFC662EBFCC3FBFAFAA1.taxon	discussion	Note. The syntypes correspond to two different species; the lectotype (MNHP 18396 vchLAM- 243) and one of the paralectotypes belong to Josa chazaliae (MNHP vchLAM- 242) and the other to an undescribed species, which is described below as Josa samaria sp. nov. (MNHP vchLAM- 244). Only records from the additional material examined were mapped, as the type’s label lacks a specific locality within the Sierra Nevada de Santa Marta.	en	Martínez, Leonel, Kochalka, John A., Cabra-Garcia, Jimmy, Ramírez, Martin (2025): Revealing the identity of Josa chazaliae (Simon, 1897) (Araneae: Anyphaenidae): new species and the highest altitude record for a spider in South America. Zootaxa 5566 (2): 201-242, DOI: 10.11646/zootaxa.5566.2.1, URL: https://doi.org/10.11646/zootaxa.5566.2.1
4D76240CFFBDFFC662EBFCC3FBFAFAA1.taxon	materials_examined	Other material examined: COLOMBIA. Magdalena: Ciénaga, San Pedro, Parque Natural Nacional Sierra Nevada de Santa Marta, Loma de la Cebolleta, misc. vegetation, low-medium, manual caught, 3505 m, J. Kochalka leg., 9. V. 1975, 1 ♂ (MUSENUV-Ar- 2671; vchLAM- 245), 1 ♂ (MACN-Ar 46690; vchLAM- 246). Cesar: Valledupar, Parque Nacional Natural Sierra Nevada de Santa Marta (NW Donachuí), Meoyaca, Río Donachuí, 3974 m, [10 ° 47 ′ 39.9 ″ N, 73 ° 36 ′ 03.6 ″ W], 18. I. 1974, 1 ♀ (MUSENUV-Ar- 2672; vchLAM- 247); 3596 m, [10 ° 47 ′ 04.5 ″ N, 73 ° 33 ′ 40.4 ″ W], 2. I. 1973, 2 ♀ (MACN-Ar 46691; vchLAM- 248); Donachuí, Rio Donachuí, top of woody vegetation, 4230 m, [10 ° 47 ′ 50.0 ″ N, 73 ° 37 ′ 34.5 ″ W], 6. I. 1973, 1 ♀ (MUSENUV-Ar- 2673).	en	Martínez, Leonel, Kochalka, John A., Cabra-Garcia, Jimmy, Ramírez, Martin (2025): Revealing the identity of Josa chazaliae (Simon, 1897) (Araneae: Anyphaenidae): new species and the highest altitude record for a spider in South America. Zootaxa 5566 (2): 201-242, DOI: 10.11646/zootaxa.5566.2.1, URL: https://doi.org/10.11646/zootaxa.5566.2.1
4D76240CFFBDFFC662EBFCC3FBFAFAA1.taxon	diagnosis	Diagnosis. Males of Josa chazaliae are similar to those of Josa berlandi sp. nov. by the small conductor with the external region apically bifid and by the short embolus (Figs 7 H – I, compare with Figs 25 A – D, 26 A – C, 27 A – I), but can be differentiated from this species by the conductor widening towards the apex with both apical tips of the external section of the conductor of about the same length, the posterior one curved towards the internal region and with a subapical pronounced bump (Fig. 7 A – D, arrow), median apophysis bifid with its anterior branch tubular and slightly curved (Figs 5 A, 6 A, 7 A – D), and the very small and rugous retrobasal tegular lobe (Fig. 7 A, F). Females are easily distinguishable by the very thin and short ducts of the accessory bulb restricted to the lateral regions of the vulva (Figs 6 D – E, 8 A – D), the swollen, triangular basal sclerotized projections of the epigyne (Figs 2 E, 5 E), the long median septum, and median field oval with deep copulatory openings (Figs 2 E, 5 E – F, 6 D – E, 8 A – F).	en	Martínez, Leonel, Kochalka, John A., Cabra-Garcia, Jimmy, Ramírez, Martin (2025): Revealing the identity of Josa chazaliae (Simon, 1897) (Araneae: Anyphaenidae): new species and the highest altitude record for a spider in South America. Zootaxa 5566 (2): 201-242, DOI: 10.11646/zootaxa.5566.2.1, URL: https://doi.org/10.11646/zootaxa.5566.2.1
4D76240CFFBDFFC662EBFCC3FBFAFAA1.taxon	description	Description. Female (description based on the lectotype, MNHP 18396; vchLAM- 243 and a female MUSENUV- Ar- 2671; vchLAM- 245). (Figs 1 – 2, 4): Total length 7.42. Carapace length 3.70, width 2.56, height 1.64. Clypeus length 0.22. Sternum length 1.88, width 1.29. Legs. Leg I: ti length 1.90 / width 0.51, mt 1.53. II: ti 1.81 / 0.47, mt 1.49. III: ti 1.61 / 0.41, mt 1.57. IV: ti 2.21 / 0.46, mt 2.18. Spines: legs I – II, femur d 1 - 1 - 1, p 2 ap; tibia v 2 - 2 - 2; metatarsus v 2 bas. III, femur d 1 - 1 - 1, p 0 - d 1 - d 1, r 0 - d 1 - d 1; patella r 1 me; tibia v 2 - 2 - 2, p 1 - d 1 - 1 - 0, r 1 - d 1 - 1 - 0, d (r 1 - 1) bristles; metatarsus d 0 - p 1 - 2, v 2 - 0 - 2, p d 1 - 1 - 0 - 1, r d 1 - 1 - 0 - 1. IV = III but femur r d 1 ap; patella d 1 - 1, r 1; metatarsus v 2 - 2 - 2. Abdomen: length 3.75, width 1.86, spiracle – epigastrium 1.26. Body dehydrated, damaged. Color: carapace dark brown, with two wide longitudinal dark bands on sides, narrowed in cephalic region. Ocular area on dark patch with eyes bordered by black rings. Carapace with fringe of white setae on sides. Clypeus brown, low, with some white setae. Chelicerae brown with black patches and fine setae on front. Endites, labium, and sternum dark brown, labium and endites light in apical region. Legs dark brown, with long darker stripes on dorsal edges of femora. All articles covered by white setae. Abdomen dorsally with median spots on dark gray background and abundant white setae; anterior region with bunch of thick setae; venter uniformly dark gray, with light dots. Epigyne: anterior plate sclerotized with deep anterior excavations, median notch long and narrow (Figs 2 E, 5 E, arrows). Median field transverse, large, oval in posterior view, with medial depression (Fig. 8 E – F). Lateral lobes with sclerotized extensions very wide, forming a longitudinal concavity, posterior extensions rounded at base (Figs 2 E, 5 E). Copulatory ducts long, touching with the accessory bulb ducts medially (Fig. 6 E). Spermathecae rounded medially positioned in the epigyne, oriented towards median region (Figs 5 F, 6 E, 8 A – E). Ducts of accessory bulbs long, thin, convergent (Figs 2 E, 5 E – F, 6 D – E, 7). Male (MUSENUV-Ar- 2671; vchLAM- 245) (Figs 3): Total length 4.54. Carapace length 2.34, width 1.61, height 0.72. Clypeus length 0.10. Sternum length 1.20, width 0.79. Legs. Leg I: ti length 2.17 / width 0.26, mt 1.88. II: ti 1.97 / 0.25, mt 1.72. III: ti 1.32 / 0.23, mt 1.20. IV: ti 1.62 / 0.27, mt 1.55. Spines: legs s I – II, femur d 1 - 1 - 1, p 2 ap, r 0 - d 1 - d 1; tibia v 2 - 2 - 2, p 1 - d 1 - 1 - 0, r 1 - d 1 - 1 - 0; metatarsus v 2 - 2 - 0, p d 1 - d 1 - 0, r d 1 - d 1. III – IV, femur d 1 - 1 - 1, p 0 - d 1 - d 1, r 0 - d 1 - d 1; patella r 1 me; tibia v 2 - 2 - 2, p 1 - d 1 - 1 - 0, r 1 - d 1 - 1 - 0, d (r 1 - 1) bristles; metatarsus d 0 - p 1 - 2, v 2 - 2 - 2, p d 1 - 1 - 0 - 1, r d 1 - 1 - 0 - 1. Abdomen: length 2.34, width 1.35, spiracle – epigastrium 0.83. Color: carapace brown, with two faint dark bands on sides, anteriorly covered by white setae. Ocular area brown with eyes bordered in black rings. Clypeus brown, low, with abundant white setae. Chelicerae brown with black stripes and fine setae on front. Endites, labium, and sternum brown; labium and sternum darker. Legs coxae-tarsi brown, with some tenuous dark patches. Articles with a few white and black setae. Abdomen dehydrated, with a pattern of faint spots on a brown background. Dorsum covered with few fine white setae. Venter uniformly brown. Palp: tegulum with small basal retrolateral lobe, very sclerotized, rugous (Figs 5 A, 6 A, 7 A, 8 A – F). Median apophysis large, bifid, anterior tip long, tubular, hook-shaped, posterior branch membranous at base with external border sub-rounded (Figs 5 A, 6 A, 7 A – B). Conductor small, rounded, with projected base, external section wide, anteriorly swollen, bifid, with both tips equal in length, but posterior tip curved towards internal region (Fig. 7 A – B, D); internal section ending as a sharp tip (Fig. 7 D). Paramedian apophysis cup-shaped, small, heavily sclerotized, concave, apically sharp (Figs 5 A, D; 6 A – C; 7 A – E). Embolus short, not describing a complete loop around bulb, tape-shaped at base, coiled, with poorly developed laminar projection; basal process small, flat, retrolateral region pointed in posterior region, prolateral region oval, poorly pronounced into external keel (Figs 5 A – D, 6 A – C, 7). Variation: Males (n = 2): total length: 4.28 – 4.54; carapace length: 2.18 – 2.34, width: 1.57 – 1.61; abdomen length: 2.14 – 2.34, width: 1.19 – 1.35. Females (n = 3): total length: 7.40 – 8.19; carapace length: 3.52 – 3.70, width: 2.47 – 2.56; abdomen length: 3.75 – 4.42, width: 1.86 – 2.64. Fresh specimens look more pigmented.	en	Martínez, Leonel, Kochalka, John A., Cabra-Garcia, Jimmy, Ramírez, Martin (2025): Revealing the identity of Josa chazaliae (Simon, 1897) (Araneae: Anyphaenidae): new species and the highest altitude record for a spider in South America. Zootaxa 5566 (2): 201-242, DOI: 10.11646/zootaxa.5566.2.1, URL: https://doi.org/10.11646/zootaxa.5566.2.1
4D76240CFFBDFFC662EBFCC3FBFAFAA1.taxon	distribution	Distribution. Only known for Sierra Nevada de Santa Marta, Colombia (Fig. 31).	en	Martínez, Leonel, Kochalka, John A., Cabra-Garcia, Jimmy, Ramírez, Martin (2025): Revealing the identity of Josa chazaliae (Simon, 1897) (Araneae: Anyphaenidae): new species and the highest altitude record for a spider in South America. Zootaxa 5566 (2): 201-242, DOI: 10.11646/zootaxa.5566.2.1, URL: https://doi.org/10.11646/zootaxa.5566.2.1
4D76240CFFB2FFC562EBFA2FFBFAFC7D.taxon	description	Figures 9 – 10	en	Martínez, Leonel, Kochalka, John A., Cabra-Garcia, Jimmy, Ramírez, Martin (2025): Revealing the identity of Josa chazaliae (Simon, 1897) (Araneae: Anyphaenidae): new species and the highest altitude record for a spider in South America. Zootaxa 5566 (2): 201-242, DOI: 10.11646/zootaxa.5566.2.1, URL: https://doi.org/10.11646/zootaxa.5566.2.1
4D76240CFFB2FFC562EBFA2FFBFAFC7D.taxon	materials_examined	Type Material. Holotype: COLOMBIA. Magdalena, from Sierra Nevada de Santa Marta, De Dalmas, 1 ♀, (MNHP; vchLAM- 244; paralectotype of Josa chazaliae Simon 1897).	en	Martínez, Leonel, Kochalka, John A., Cabra-Garcia, Jimmy, Ramírez, Martin (2025): Revealing the identity of Josa chazaliae (Simon, 1897) (Araneae: Anyphaenidae): new species and the highest altitude record for a spider in South America. Zootaxa 5566 (2): 201-242, DOI: 10.11646/zootaxa.5566.2.1, URL: https://doi.org/10.11646/zootaxa.5566.2.1
4D76240CFFB2FFC562EBFA2FFBFAFC7D.taxon	etymology	Etymology. The specific epithet refers to the demonym for the Santa Marta people “ Samario / a ”. It is a noun in apposition.	en	Martínez, Leonel, Kochalka, John A., Cabra-Garcia, Jimmy, Ramírez, Martin (2025): Revealing the identity of Josa chazaliae (Simon, 1897) (Araneae: Anyphaenidae): new species and the highest altitude record for a spider in South America. Zootaxa 5566 (2): 201-242, DOI: 10.11646/zootaxa.5566.2.1, URL: https://doi.org/10.11646/zootaxa.5566.2.1
4D76240CFFB2FFC562EBFA2FFBFAFC7D.taxon	diagnosis	Diagnosis. The female is similar to that of Josa nadineae sp. nov. by the posterior quadrangular notch of the epigyne (Fig. 13 E, compare with Figs 13 E, 14 D) but can be distinguished by the posterior lateral margins on a straight transversal line (vs. oblique in Josa nadineae sp. nov.), and by the copulatory ducts describing two wide posterior coils, and the small spermathecae, anteromedially placed with long fertilization ducts (Fig. 10 A – E).	en	Martínez, Leonel, Kochalka, John A., Cabra-Garcia, Jimmy, Ramírez, Martin (2025): Revealing the identity of Josa chazaliae (Simon, 1897) (Araneae: Anyphaenidae): new species and the highest altitude record for a spider in South America. Zootaxa 5566 (2): 201-242, DOI: 10.11646/zootaxa.5566.2.1, URL: https://doi.org/10.11646/zootaxa.5566.2.1
4D76240CFFB2FFC562EBFA2FFBFAFC7D.taxon	description	Description. Female (holotype, MNHP; vchLAM- 244) (Fig. 9): Total length 7.33. Carapace length 3.41, width 2.36, height 1.35. Clypeus length 0.19. Sternum length 1.79, width 1.31. Legs. Leg I: ti length 1.84 / width 0.42, metatarsus (mt) 1.45. II: ti 1.74 / 0.40, mt 1.39. III: ti 1.59 / 0.38, mt 1.55. IV: ti 2.17 / 0.44, mt 2.41. Spines: legs I – II, femur d 1 - 1 - 1, p 2 ap; tibia v 2 - 2 - 2; metatarsus v 2 bas. III, femur d 1 - 1 - 1, p 0 - d 1 - d 1, r 0 - d 1 - d 1; patella r 1 me; tibia v 2 r- 1 - 2, p 1 - d 1 - 1 - 0 r 1 - d 1 - 1 - 0, d (r 1 - 1) bristles; metatarsus d 0 - p 1 - 2, v 2 - 0 - 2, p d 1 - 1 - 0 - 1, r d 1 - 1 - 0 - 1. IV = III but metatarsus v 2 - 2 - 2. Abdomen: length 4.03, width 2.33, spiracle – epigastrium 0.80. Color: carapace brown, with two wide, longitudinal darker band on sides, narrow in cephalic region. Ocular area on black patch with eyes bordered by black rings. Carapace with lateral region covered by fringe of white setae. Clypeus brown, low with abundant white setae. Chelicerae brown with black patches and fine setae on frontal edge. Endites, labium, and sternum dark brown, labium and endites light in apical region. Sternum with a very thin medial pale stripe. Legs coxae-tarsi dark brown, femora and patellae with longitudinal dark bands. All articles covered by white setae, abundant between coxae. Abdomen with dark brown pattern of spots on a gray background, dorsally with white medial band followed by 4 - 5 faint light chevrons; lateral regions covered by abundant small dots. Abdomen laterally surrounded by abundant white setae, with a thick bunch of setae on anterior region. Venter uniformly covered by dark and light patches. Epigyne: anterior plate sclerotized with deep and rounded anterior excavations, median notch quadrangular, posteriorly positioned (Fig. 10 A, D). Median field transverse, large, oval in posterior view (Fig. 10 C). Lateral lobes with sclerotized extensions wide, forming, posterior extensions rounded at base (Fig. 10 A, C – D). Copulatory ducts long, forming two basal coils, anteriorly medially expanded, joining with the accessory bulb ducts at medial level. Spermathecae small, rounded, medially positioned in the epigyne, oriented towards longitudinal plane. Ducts of accessory bulbs short, wide, convergent (Fig. 10 A – E). Male. Unknown.	en	Martínez, Leonel, Kochalka, John A., Cabra-Garcia, Jimmy, Ramírez, Martin (2025): Revealing the identity of Josa chazaliae (Simon, 1897) (Araneae: Anyphaenidae): new species and the highest altitude record for a spider in South America. Zootaxa 5566 (2): 201-242, DOI: 10.11646/zootaxa.5566.2.1, URL: https://doi.org/10.11646/zootaxa.5566.2.1
4D76240CFFB2FFC562EBFA2FFBFAFC7D.taxon	distribution	Distribution. Only known for Sierra Nevada de Santa Marta, Colombia (Fig. 31).	en	Martínez, Leonel, Kochalka, John A., Cabra-Garcia, Jimmy, Ramírez, Martin (2025): Revealing the identity of Josa chazaliae (Simon, 1897) (Araneae: Anyphaenidae): new species and the highest altitude record for a spider in South America. Zootaxa 5566 (2): 201-242, DOI: 10.11646/zootaxa.5566.2.1, URL: https://doi.org/10.11646/zootaxa.5566.2.1
4D76240CFFB1FFDC62EBFC13FBD7FB31.taxon	description	Figures 11 – 16	en	Martínez, Leonel, Kochalka, John A., Cabra-Garcia, Jimmy, Ramírez, Martin (2025): Revealing the identity of Josa chazaliae (Simon, 1897) (Araneae: Anyphaenidae): new species and the highest altitude record for a spider in South America. Zootaxa 5566 (2): 201-242, DOI: 10.11646/zootaxa.5566.2.1, URL: https://doi.org/10.11646/zootaxa.5566.2.1
4D76240CFFB1FFDC62EBFC13FBD7FB31.taxon	materials_examined	Type material. Holotype: COLOMBIA. Magdalena: Ciénaga, San Pedro, Parque Natural Nacional Sierra Nevada de Santa Marta, Loma de la Cebolleta, dead mats of curly grass, manual caught, 3505 m, [10 ° 54 ′ 09 ″ N, 73 ° 55 ′ 27 ″ W], J. Kochalka leg., 9. V. 1975, 1 ♂ (MUSENUV-Ar- 2678; vchLAM- 249). Paratypes: COLOMBIA. Magdalena: Ciénaga, San Pedro, Parque Natural Nacional Sierra Nevada de Santa Marta, north of Lago Mavaria, manual caught, 4008 m, J. Kochalka leg., 8. V. 1975, 1 ♀ (MUSENUV-Ar- 2679; vchLAM- 251), 1 ♂ 1 ♀ (MUSENUV-Ar- 2680); Serranía Dunanchucua, northern peak of Norte Grande, 3797 m, 5. III. 1975, 1 ♀ (MUSENUV-Ar- 2681); 3900 m, [10 ° 54 ′ 01.8 ″ N, 73 ° 53 ′ 18.6 ″ W], 7. III. 1975, 1 ♂ (MACN-Ar 46692; vchLAM- 250); Río Tucurinca, Lago Arucuina, 4176 m, 12. III. 1975, 1 ♀ (MACN-Ar 46693; vchLAM- 252).	en	Martínez, Leonel, Kochalka, John A., Cabra-Garcia, Jimmy, Ramírez, Martin (2025): Revealing the identity of Josa chazaliae (Simon, 1897) (Araneae: Anyphaenidae): new species and the highest altitude record for a spider in South America. Zootaxa 5566 (2): 201-242, DOI: 10.11646/zootaxa.5566.2.1, URL: https://doi.org/10.11646/zootaxa.5566.2.1
4D76240CFFB1FFDC62EBFC13FBD7FB31.taxon	etymology	Etymology. The specific name is after the Canadian arachnologist Nadine Dupérré, in recognition of her many contributions to the spider’s taxonomy.	en	Martínez, Leonel, Kochalka, John A., Cabra-Garcia, Jimmy, Ramírez, Martin (2025): Revealing the identity of Josa chazaliae (Simon, 1897) (Araneae: Anyphaenidae): new species and the highest altitude record for a spider in South America. Zootaxa 5566 (2): 201-242, DOI: 10.11646/zootaxa.5566.2.1, URL: https://doi.org/10.11646/zootaxa.5566.2.1
4D76240CFFB1FFDC62EBFC13FBD7FB31.taxon	diagnosis	Diagnosis. Males resemble those of Josa andesiana (Berland, 1913) by the short and tape-shaped embolus (Fig. 13 E−F, compare with Dupérré, 2023: 139, fig. 16 A – C; Poy. et. al. 2023: 6, fig. 3 E). It can be distinguished from the latter species by the following combination of characters: laminar projection of the embolus larger (Figs 13 A – D, 14 A – B, 15 B, I), bifid external section of the conductor with both branches interrupted by a deep groove (Fig. 15 A – B), posterior branch of the conductor protruded, with many small spicules and longer than the anterior branch, which is laminar and apically irregular with small reticulations, bifid median apophysis with small scales on the external edge (Fig. 15 F), and pronounced retrolateral basal lobe (Figs 13 A, 14 A). The females are distinguished by the spermathecae converging on the median line, touching each other, the obliquely oriented posterior borders of the epigyne and the copulatory ducts anteriorly arched, widening towards the anterior part of vulva (Figs 13 E – F, 14 E – F, 16).	en	Martínez, Leonel, Kochalka, John A., Cabra-Garcia, Jimmy, Ramírez, Martin (2025): Revealing the identity of Josa chazaliae (Simon, 1897) (Araneae: Anyphaenidae): new species and the highest altitude record for a spider in South America. Zootaxa 5566 (2): 201-242, DOI: 10.11646/zootaxa.5566.2.1, URL: https://doi.org/10.11646/zootaxa.5566.2.1
4D76240CFFB1FFDC62EBFC13FBD7FB31.taxon	description	Description. Male (holotype, MUSENUV-Ar- 2678; vchLAM- 249) (Fig. 11): Total length 5.75. Carapace length 2.95, width 2.13, height 1.10. Clypeus length 0.13. Sternum length 1.45, width 0.98. Legs. Leg I: ti length 1.79 / width 0.36, mt 1.39. II: ti 1.70 / 0.35, mt 1.46. III: ti 1.72 / 0.38, mt 1.75. IV: ti 2.74 / 0.44, mt 2.59. Spines: legs I – II, femur d 1 - 1 - 1, p 2 ap, r 0 - d 1 - d 1; tibia v 2 - 2 - 2, p 1 - d 1 - 1 - 0, r 1 - d 1 - 1 - 0; metatarsus v 2 - 0 - 0, p d 1 - 0. III – IV, femur d 1 - 1 - 1, p 0 - d 1 - d 1, r 0 - d 1 - d 1; patella r 1 me; tibia v 2 - 2 - 2, p 1 - d 1 - 1 - 0, r 1 - d 1 - 1 - 0, d (r 1 - 1) bristles; metatarsus d 0 - p 1 - 2, v 2 - 2 - 2, p d 1 - 1 - 0 - 1, r d 1 - 1 - 0 - 1. Abdomen: length 2.81, width 1.56, spiracle – epigastrium 1.01. Color: carapace brown, marginally darker, with two wide black degraded bands, converging and narrow anteriorly. Ocular area dark brown with eyes bordered on a black patch. Lateral regions of carapace upholstered with small fine white setae. Clypeus brown, low with some white setae. Chelicerae brown with fine setae and dark patches on frontal edge. Endites light brown, with faint medial dark patch. Labium brown reddish, anteriorly lighter. Sternum brown with faint black patches. Legs uniformly brown, covered by white setae, mainly between coxae. Abdomen dorsum with a pattern of black spots on a brown background, surrounded by a thin band, extended to medial region; dorsum with an anterior medial dark patch with posterior extensions, followed by irregular spots. Abdomen covered with fine white setae; anterior region with a dense bunch of setae, marginal ring covered by black setae. Venter uniformly brown with light and dark dots, dark patches surrounding the spinnerets base. Palp: tegulum with a basal small lobe, with granulose texture (Figs 13 A, 14 A, 15 A, G). Median apophysis long, laminar, ending in a very short hook-shaped tip, with many denticles (Fig. 15 F, see square). Conductor small, rounded, ventral section apically bifid, with many narrow furrows, anterior tip apically laminar with several very small denticles, posterior tip with small granules, posterior tip slightly longer than anterior one; base of the conductor with a sharp, spine-shaped projection (Fig. 15 A, B). Paramedian apophysis cup-shaped, concave (Figs 13 A – B, 14 A – B, 15 A – B, D, F). Embolus tape-shaped, not describing a loop, very short, with large laminar projection (Fig. 15 B, G, I), apically twisted; basal process small, flat, sclerotized, with poorly pronounced external keel and medial depression (Figs 13 A – D, 14 A – C, 15). Female (paratype, MUSENUV-Ar- 2679; vchLAM- 251). (Fig. 12): Total length 7.02. Carapace length 3.19, width 2.23, height 1.17. Clypeus length 0.13. Sternum length 1.59, width 1.06. Legs. Leg I: ti length 1.61 / width 0.39, mt 1.21. II: ti 1.63 / 0.34, mt 1.26. III: ti 1.36 / 0.37, mt 1.56. IV: ti 1.95 / 0.38, mt 1.50. Spines: legs I – II, femur d 1 - 1 - 1, p 2 ap; tibia v 2 - 2 - 2; metatarsus v 2 bas. III – IV, femur d 1 - 1 - 1, p 0 - d 1 - d 1, r 0 - d 1 - d 1; patella r 1 me; tibia v 2 - 2 - 2, p 1 - d 1 - 1 - 0, r 1 - d 1 - 1 - 0, d (r 1 - 1) bristles; metatarsus d 0 - p 1 - 2, v 2 - 0 - 2, p d 1 - 1 - 0 - 1, r d 1 - 1 - 0 - 1. Abdomen: length 3.63, width 2.69, spiracle – epigastrium 1.53. Color pattern as in male, but more pigmented, abdominal spots more defined. Sternum and labium more pigmented. Legs with defined longitudinal dark bands on dorsal edge, and many dots on ventral one. Epigyne: anterior plate sclerotized with deep and rounded anterior excavations, laterally elongated, median depression small, quadrangular, posteriorly positioned (Figs 13 E, 14 D). Median field transverse, large, oval in posterior view with medial depression (Fig. 16 E – F). Lateral lobes with sclerotized extensions wide, apically rounded (Fig. 16 E – F). Copulatory ducts long, with a very wide posterior coil, laterally expanded in anterior side, joining with the accessory bulb ducts at medial level. Spermathecae small, rounded, in median position, oriented towards median septum (Fig. 14 E). Ducts of accessory bulbs long, thin, convergent (Figs 13 E – F, 14 E – F, 16). Variation: Males (n = 3): total length: 5.40 – 5.75; carapace length: 2.47 – 2.97, width: 1.84 – 2.13; abdomen length: 2.66 – 2.81, width: 1.56 – 1.69. Females (n = 4): total length: 7.02 – 7.49; carapace length: 3.19 – 3.79, width: 2.23 – 2.67; abdomen length: 3.63 – 3.72, width: 1.25 – 2.69. Some specimens present body and abdominal spot patterns more pigmented.	en	Martínez, Leonel, Kochalka, John A., Cabra-Garcia, Jimmy, Ramírez, Martin (2025): Revealing the identity of Josa chazaliae (Simon, 1897) (Araneae: Anyphaenidae): new species and the highest altitude record for a spider in South America. Zootaxa 5566 (2): 201-242, DOI: 10.11646/zootaxa.5566.2.1, URL: https://doi.org/10.11646/zootaxa.5566.2.1
4D76240CFFB1FFDC62EBFC13FBD7FB31.taxon	distribution	Distribution. Only known from Sierra Nevada de Santa Marta, Colombia (Fig. 31).	en	Martínez, Leonel, Kochalka, John A., Cabra-Garcia, Jimmy, Ramírez, Martin (2025): Revealing the identity of Josa chazaliae (Simon, 1897) (Araneae: Anyphaenidae): new species and the highest altitude record for a spider in South America. Zootaxa 5566 (2): 201-242, DOI: 10.11646/zootaxa.5566.2.1, URL: https://doi.org/10.11646/zootaxa.5566.2.1
4D76240CFFA8FFD462EBFADFFB5CFDE5.taxon	description	Figs 17 – 22	en	Martínez, Leonel, Kochalka, John A., Cabra-Garcia, Jimmy, Ramírez, Martin (2025): Revealing the identity of Josa chazaliae (Simon, 1897) (Araneae: Anyphaenidae): new species and the highest altitude record for a spider in South America. Zootaxa 5566 (2): 201-242, DOI: 10.11646/zootaxa.5566.2.1, URL: https://doi.org/10.11646/zootaxa.5566.2.1
4D76240CFFA8FFD462EBFADFFB5CFDE5.taxon	materials_examined	Type material. Holotype: COLOMBIA. Magdalena: Ciénaga, Parque Nacional Natural Sierra Nevada de Santa Marta, Quebrada El Chorro, southern branch of the lake, manual caught, 3797 m, [10 ° 47 ′ 00 ″ N, 73 ° 32 ′ 04 ″ W], J. Kochalka leg., 5. III. 1975, 1 ♂ (MUSENUV-Ar- 2682; vchLAM- 260). Paratypes: same locality data as the holotype 1 ♀ (MUSENUV-Ar- 2683; vchLAM- 261), 1 ♂ 3 ♀ (MUSENUV-Ar- 2684); Parque Nacional Natural Sierra Nevada de Santa Marta, Serranía Dunanchucua, northern peak of Norte Grande, 3992 m, 7. III. 1975, 1 ♀ (MUSENUV-Ar- 2695), Tip of Pico Norte Grande de la Cuchilla Cesarguarniaga, 4398 m, 2. II. 1974, 1 ♂ 3 ♀ (MUSENUV-Ar- 2687); Serranía Dunanchucua, northern peak of Norte Grande, 3992 m, 7. III. 1975, 1 ♂ (MACN-Ar 46696; vchLAM- 262), 1 ♂ (MUSENUV-Ar- 2695), 4236 m, 1 ♀ (MUSENUV-Ar- 2690); Quebrada El Chorro, north of Río Sevillano, 3718 m, 13. II. 1974, 1 ♂ (MUSENUV-Ar- 2692); Río Tucurinca, Lago del Rio Tucurinca, 4785 m, 16. III. 1975, 1 ♀ (MUSENUV-Ar- 2693); East tip of Pico Cristobal Colón, 5275 m, [10 ° 49 ′ 59.7 ″ N, 73 ° 40 ′ 08.6 ″ W], 12. I. 1974, 1 ♀ (MUSENUV-Ar- 2696); East tip of NW Pico Cristobal Colón, almost at superior peak, under loose rocks, on a nearly vertical cliff, surrounded by glacial ice, 5500 m, [10 ° 50 ′ 10.3 ″ N, 73 ° 41 ′ 11.0 ″ W], 10. I. 1974, 1 imm. (MUSENUV-Ar- 2697); Lago Arucuina, Río Tucurinca, 4176 m, 12. III. 1975, 1 ♀ (MACN-Ar 46697; vchLAM- 263); Lago del Rio Tucurinca, 4785 m, 16. III. 1975, 1 ♂ (MUSENUV-Ar- 2698), 1 ♂ (MUSENUV-Ar- 2699); South slope of Pico Tiaroma, 1 ♂ (MUSENUV-Ar- 2700). César: Valledupar, Parque Nacional Natural Sierra Nevada de Santa Marta, Donachui, Río Donachui, top of woody vegetation, manual caught, 4230 m, [10 ° 47 ′ 50.0 ″ N, 73 ° 37 ′ 34.5 ″ W], J. Kochalka leg., 6. I. 1973, 1 ♀ (MUSENUV-Ar- 2685); Meoyaca, Río Donachuí, 3596 m, [10 ° 47 ′ 04.5 ″ N, 73 ° 33 ′ 40.4 ″ W], 2. I. 1973, 1 ♀ (MUSENUV-Ar- 2689); same locality as before, but Naboba zone, top of Meoyaca, Rio Donachuí, northern extension, under rocks, at the lower edge of the Pico La Reina / Pico Ojeda Glacier, 4800 m, [10 ° 49 ′ 08.0 ″ N, 73 ° 37 ′ 58.6 ″ W], 9. I. 1973, 1 ♂ (MUSENUV-Ar- 2691); same locality, but Donachuí, Río Donachuí, north tributary of Lago Mavaria, 4008 m, 8. III. 1973, 1 ♂ (MUSENUV-Ar- 2694). Ciénaga, 5 km N from Santa Marta, Sierra Nevada de Santa Marta, S. W. Páramo, 3657 m, P. Dalington coll., 2. II. 1929, 1 ♂, 2 ♀ (MCZ-IZ 170963, 170964, 170965).	en	Martínez, Leonel, Kochalka, John A., Cabra-Garcia, Jimmy, Ramírez, Martin (2025): Revealing the identity of Josa chazaliae (Simon, 1897) (Araneae: Anyphaenidae): new species and the highest altitude record for a spider in South America. Zootaxa 5566 (2): 201-242, DOI: 10.11646/zootaxa.5566.2.1, URL: https://doi.org/10.11646/zootaxa.5566.2.1
4D76240CFFA8FFD462EBFADFFB5CFDE5.taxon	etymology	Etymology. The specific epithet alludes to the high altitudes of the type locality in Sierra Nevada de Santa Marta. It is a noun in genitive.	en	Martínez, Leonel, Kochalka, John A., Cabra-Garcia, Jimmy, Ramírez, Martin (2025): Revealing the identity of Josa chazaliae (Simon, 1897) (Araneae: Anyphaenidae): new species and the highest altitude record for a spider in South America. Zootaxa 5566 (2): 201-242, DOI: 10.11646/zootaxa.5566.2.1, URL: https://doi.org/10.11646/zootaxa.5566.2.1
4D76240CFFA8FFD462EBFADFFB5CFDE5.taxon	diagnosis	Diagnosis. Males can be distinguished from those of other species of Josa by the following combination of characters: bifid external section of the conductor with anterior branch shorter than posterior, which is long and sharp (Figs 19 A – B, 20 A – B, 21 A – B, D, H); embolus describing a loop around basal process, with a thin and long laminar projection (Figs 20 C, 21 H – I), and bifid median apophysis with many small denticles on external edge (Fig. 21 F, see inset). Females of this species are distinguished from those of other species of Josa by the shape of the posterior sclerotized projections of lateral lobes, which are narrow, handle-like, arched, and with deep rounded anterior depressions (Figs 19 E, 20 D), the median field visible in dorsal view (Figs 19 E, 22 E – F), small, elongated spermathecae, laterally positioned at the median region of the epigyne (Figs 19 E – F, 20 E – F, 22).	en	Martínez, Leonel, Kochalka, John A., Cabra-Garcia, Jimmy, Ramírez, Martin (2025): Revealing the identity of Josa chazaliae (Simon, 1897) (Araneae: Anyphaenidae): new species and the highest altitude record for a spider in South America. Zootaxa 5566 (2): 201-242, DOI: 10.11646/zootaxa.5566.2.1, URL: https://doi.org/10.11646/zootaxa.5566.2.1
4D76240CFFA8FFD462EBFADFFB5CFDE5.taxon	description	Description. Male (holotype, MUSENUV-Ar- 2682; vchLAM- 260) (Fig. 17): Total length 8.89. Carapace length 4.35, width 3.29, height 1.07. Clypeus length 0.12. Sternum length 2.31, width 1.58. Legs. Leg I: ti length 3.84 / width 0.49, mt 3.20. II: ti 3.82 / 0.52, mt 3.13. III: ti 3.12 / 0.45, mt 3.29. IV: ti 3.95 / 0.54, mt 4.11. Spines: legs I – II, femur d 1 - 1 - 1, p 0 - 1 d 2 ap, r 0 - d 1 - d 1; tibia v 2 - 2 - 2; metatarsus v 2 - 0 - 0. III – IV, femur d 1 - 1 - 1, p 0 - d 1 - d 1, r 0 - d 1 - d 1; patella r 1 me; tibia v 2 - 2 - 2, p 1 - d 1 - 1 - 0, r 1 - d 1 - 1 - 0, d (r 1 - 1) bristles; metatarsus d 0 - p 1 - 2, v 2 - 2 - 2, p d 1 - 1 - 0 - 1, r d 1 - 1 - 0 - 1. Abdomen: length 4.31, width 2.52, spiracle – epigastrium 1.72. Color: carapace brown, with two wide longitudinal discontinuous dark brown stripes, extending to fovea and marginally covered by abundant white setae; eyes bordered by black rings, triangular band emerging from posterior side of PLE and PME. Clypeus brown, low with abundant white setae. Chelicerae brown with black spots and fine setae on frontal edge. Endites light brown, light at anterior side. Labium and sternum dark brown. Legs coxae light brown, femora-tarsi brown with some dark patches, patellae less pigmented. All articles with black dots, stripes, and covered by few white and black setae, abundant white setae between coxae. Abdomen with pattern of black patches on lighter brown background, sub-rounded by thin band; anterior region of dorsum medially with a wide longitudinal dark band extended to submedial region, followed by five wide and very short chevrons, lateral regions with many dots. Abdomen covered with few fine white setae, anterior region with a dense bunch of setae. Venter dark gray with many light patches. Palp: tegulum without tegular lobe. Median apophysis very wide, bifid, with anterior hook-shaped tip, posterior region with a denticulated edge (Fig. 21 F). Conductor large, rounded, external section apically bifid (Figs. 19 A – B, 20 A – B, 21 A – B, D), with posterior tip long, sharp, anterior tip shorter with small apical denticles, dorsal section sharp (Fig. 21 B). Paramedian apophysis short, heavily sclerotized, sharp, narrow. Embolus apically filiform, with a narrow laminar projection extending to subapical side; basal process small, reticulate, posterior region rounded, with pronounced external keel (Figs 19 E – F, 20 E – F, 21). Female (paratype, MUSENUV-Ar- 2683; vchLAM- 261). (Fig. 18): Total length 11.18. Carapace length 5.98, width 4.16, height 1.72. Clypeus length 0.28. Sternum length 2.90, width 2.20. Legs. Leg I: ti length 3.43 / width 0.70, mt 2.86. II: ti 4.12 / 0.65, mt 3.26. III: ti 3.24 / 0.63, mt 3.49. IV: ti 4.39 / 0.60, mt 3.97. Spines: legs I – II, femur d 1 - 1 - 1, p 0 - d 1 - d 1, r 0 - d 1 - d 1; tibia v 2 - 2 - 2; metatarsus v 2 bas. III, femur d 1 - 1 - 1, p 0 - d 1 - d 1, r 0 - d 1 - d 1; patella r 1 me; tibia v p 1 - 2 - 2, p 1 - d 1 - 1 - 0, r 1 - d 1 - 1 - 0, d (r 1 - 1) bristles; metatarsus d 0 - p 1 - 2, v 2 - 2 - 2, p d 1 - 1 - 0 - 1, r d 1 - 1 - 0 - 1, IV = III but femur r 1 d. Abdomen: length 6.08, width 3.96, spiracle – epigastrium 2.15. Color pattern as in male, but more pigmented. Lateral regions of abdomen with many dots. Epigyne: median field transverse, wide, oval in posterior view (Fig. 22 E – F). Lateral lobes with sclerotized extensions wide, rounded, with small, rounded concavities at anterior side. Spermathecae small, oval, lateromedially placed. Copulatory ducts long, thin, coiled along longitudinal axis, oriented in parallel, anterior region rounded (Figs 20 E, 21 A – B). Ducts of accessory bulbs long, convergent (Figs 19 E – F, 20 E – F, 22). Variation: Males (n = 5): total length: 7.51 – 11.35; carapace length: 3.75 – 4.35, width: 2.72 – 3.29; abdomen length: 4.31 – 5.38, width: 1.59 – 2.62. Females (n = 5): total length: 11.18 – 11.35; carapace length: 5.88 – 5.98, width: 4.06 – 4.16; abdomen length: 5.38 – 6.08, width: 3.65 – 3.96. The pigmentation and abdominal spots are variable across specimens, the carapace can be darker, and the abdominal spots more defined. The anterior tip of apical region of the external section of the conductor seems broken in some specimens, resulting in an entire and sharp conductor tip.	en	Martínez, Leonel, Kochalka, John A., Cabra-Garcia, Jimmy, Ramírez, Martin (2025): Revealing the identity of Josa chazaliae (Simon, 1897) (Araneae: Anyphaenidae): new species and the highest altitude record for a spider in South America. Zootaxa 5566 (2): 201-242, DOI: 10.11646/zootaxa.5566.2.1, URL: https://doi.org/10.11646/zootaxa.5566.2.1
4D76240CFFA8FFD462EBFADFFB5CFDE5.taxon	distribution	Distribution. Only known from Sierra Nevada de Santa Marta, Colombia (Fig. 31).	en	Martínez, Leonel, Kochalka, John A., Cabra-Garcia, Jimmy, Ramírez, Martin (2025): Revealing the identity of Josa chazaliae (Simon, 1897) (Araneae: Anyphaenidae): new species and the highest altitude record for a spider in South America. Zootaxa 5566 (2): 201-242, DOI: 10.11646/zootaxa.5566.2.1, URL: https://doi.org/10.11646/zootaxa.5566.2.1
4D76240CFFA8FFD462EBFADFFB5CFDE5.taxon	biology_ecology	Ecological comments. When the high-altitude specimens were collected at 5500 masl, and at several other sites nearby, it was noted that this species was not only present but also abundant and active, a very fast runner on the rocks and snow. The specimens found at 5500 masl were collected from beneath rocks, and the specimen MUSENUV-Ar- 2696, found at an altitude of 5275 meters, was caught during the day, near noon, running on the surface of a large rock surrounded by snow and glacial ice; there were no loose rocks for shelter. This species appears to be very abundant in the Páramo of the Sierra Nevada de Santa Marta, which may explain the presence of some specimens at high altitudes. However, systematic field sampling is necessary to determine if there are established populations at altitudes above 5000 meters or if these are merely stochastic occurrences.	en	Martínez, Leonel, Kochalka, John A., Cabra-Garcia, Jimmy, Ramírez, Martin (2025): Revealing the identity of Josa chazaliae (Simon, 1897) (Araneae: Anyphaenidae): new species and the highest altitude record for a spider in South America. Zootaxa 5566 (2): 201-242, DOI: 10.11646/zootaxa.5566.2.1, URL: https://doi.org/10.11646/zootaxa.5566.2.1
4D76240CFFA0FFEC62EBFD6BFBC8F905.taxon	description	Figures 23 – 29	en	Martínez, Leonel, Kochalka, John A., Cabra-Garcia, Jimmy, Ramírez, Martin (2025): Revealing the identity of Josa chazaliae (Simon, 1897) (Araneae: Anyphaenidae): new species and the highest altitude record for a spider in South America. Zootaxa 5566 (2): 201-242, DOI: 10.11646/zootaxa.5566.2.1, URL: https://doi.org/10.11646/zootaxa.5566.2.1
4D76240CFFA0FFEC62EBFD6BFBC8F905.taxon	materials_examined	Type material. Holotype: COLOMBIA. Magdalena: Ciénaga, Parque Nacional Natural Sierra Nevada de Santa Marta, Quebrada El Chorro, Rama Sur del Lago, manual caught, 3797 m, [10 ° 47 ′ 00 ″ N, 73 ° 32 ′ 04 ″ W], J. Kochalka leg., 5. III. 1975, 1 ♂ (MUSENUV-Ar- 2675; vchLAM- 253). Paratypes: same data as the holotype 1 ♂ 4 ♀ (MUSENUV- Ar- 2674); Lake of Río Tucurinca, manual caught, 4785 m, J. Kochalka leg., 16. III. 1975, 1 ♀ (MUSENUV-Ar- 2676; vchLAM- 255), 2 ♀ (MUSENUV-Ar- 2677), 1 ♂ (MACN-Ar 46694; vchLAM- 254); norther peak of Norte Grande, 3900 m, [10 ° 54 ′ 01.8 ″ N, 73 ° 53 ′ 18.6 ″ W], 7. III. 1975, 1 ♀ (MACN-Ar 46695; vchLAM- 256).	en	Martínez, Leonel, Kochalka, John A., Cabra-Garcia, Jimmy, Ramírez, Martin (2025): Revealing the identity of Josa chazaliae (Simon, 1897) (Araneae: Anyphaenidae): new species and the highest altitude record for a spider in South America. Zootaxa 5566 (2): 201-242, DOI: 10.11646/zootaxa.5566.2.1, URL: https://doi.org/10.11646/zootaxa.5566.2.1
4D76240CFFA0FFEC62EBFD6BFBC8F905.taxon	etymology	Etymology. The specific epithet is a patronymic in honor of the late French arachnologist Lucien Berland, for his many contributions to arachnology including the Ecuadorian Anyphaenidae. It is a noun in possessive substantive.	en	Martínez, Leonel, Kochalka, John A., Cabra-Garcia, Jimmy, Ramírez, Martin (2025): Revealing the identity of Josa chazaliae (Simon, 1897) (Araneae: Anyphaenidae): new species and the highest altitude record for a spider in South America. Zootaxa 5566 (2): 201-242, DOI: 10.11646/zootaxa.5566.2.1, URL: https://doi.org/10.11646/zootaxa.5566.2.1
4D76240CFFA0FFEC62EBFD6BFBC8F905.taxon	diagnosis	Diagnosis. Males are easily distinguished from other species by the bifid external section of the conductor, with beak-shaped anterior tip, longer than the posterior one, which is sharp and thin, elongated subapical protrusion (Figs 25 A, 26 A, 27 A – B, D), median apophysis bifid, laminar, projected, with the anterior tip longer than the posterior one (Fig. 27 B, F, H), tegulum prolaterobasally projected (Figs 25 A, 26 A, 27 A). Females are distinguished by a bilobed median field folded toward the median notch, wide lateral lobes with basally rounded posterior sclerotized projections, and an arch-shaped narrow median notch (Figs 25 E – F).	en	Martínez, Leonel, Kochalka, John A., Cabra-Garcia, Jimmy, Ramírez, Martin (2025): Revealing the identity of Josa chazaliae (Simon, 1897) (Araneae: Anyphaenidae): new species and the highest altitude record for a spider in South America. Zootaxa 5566 (2): 201-242, DOI: 10.11646/zootaxa.5566.2.1, URL: https://doi.org/10.11646/zootaxa.5566.2.1
4D76240CFFA0FFEC62EBFD6BFBC8F905.taxon	description	Description. Male (holotype, MUSENUV-Ar- 2675; vchLAM- 253) (Fig. 23): Total length 4.18. Carapace length 2.35, width 1.73, height 0.89. Clypeus length 0.15. Sternum length 1.20, width 0.83. Legs. Leg I: ti length 1.79 / width 0.31, mt 1.45. II: ti 1.72 / 0.28, mt 1.43. III: ti 1.35 / 0.24, mt 1.35. IV: ti 1.71 / 0.30, mt 1.73. Spines: legs I – II, femur d 1 - 1 - 1, p 2 ap, r 0 - d 1 - d 1; tibia v p 1 - 2 - 2, p 1 - d 1 - 1 - 0, r 1 - d 1 - 1 - 0; metatarsus v 2 - 2 - 0, p d 1 - 0. III, femur d 1 - 1 - 1, p 0 - d 1 - d 1, r 0 - d 1 - d 1; patella r 1 me; tibia v 0 - 2 - 2, p 1 - d 1 - 1 - 0, r 1 - d 1 - 1 - 0, d (r 1 - 1) bristles; metatarsus d 0 - p 1 - 2, v 2 - 0 - 2, p d 1 - 1 - 0 - 1, r d 1 - 1 - 0 - 1. IV = III but tibia v 2 - 2 - 2, metatarsus v 2 - 2 - 2. Abdomen: length 1.92, width 1.01, spiracle – epigastrium 0.67. Color: carapace dark brown, with cervical groves spotted in black and converging in the large black patch on fovea. Lateral region with two poorly defined dark bands on sides, covered by abundant white setae. Ocular area on a black patch, eyes bordered in black rings. Clypeus brown, low with abundant white setae. Chelicerae brown with black stripes and fine setae on frontal edge. Endites, labium, and sternum brown with some dark spots; labium more pigmented. Legs coxae-tarsi brown, femora with dark longitudinal bands, tibia-metatarsi with rings of white setae; tarsi uniformly brown. Articles with abundant white setae. Abdomen dehydrated, with a pattern of faint light spots on a dark gray background, covered by abundant white setae. Venter uniformly dark gray. Palp: tegulum with very developed median hematodochae (Figs 25 A, 26 A, 27 A). Median apophysis very large, bifid, with anterior branch long, laminar, with projected prolateral tip, posterior branch membranous at base and with the external border sub-rounded (Figs 25 A – B, 26 A – C, 27 A, B, D, F). Conductor small, rounded, with projected base, external section apically wide and bifid, with a swollen, elongated projection; anterior tip of external section large, curved, and longer than posterior; posterior tip short and sharp; internal section ending as a sharp tip, converging with the posterior tip of external section (Figs 25 A – B, 26 A – C, 27 A, B, D, F). Paramedian apophysis large, cup-shaped, heavily sclerotized, concave, apically acuminated (Fig. 27 C, E) Embolus short, tape-shaped at base, coiled, with poorly developed laminar projection, apically filiform; basal process small, flat, retrolateral region pointed in posterior region, prolateral region oval, poorly pronounced external keel (Figs 25 A – D, 26 A – C, 27). Female (paratype, MUSENUV-Ar- 2676; vchLAM- 255). (Fig. 24): Total length 4.09. Carapace length 1.73, width 1.31, height 0.67. Clypeus length 0.08. Sternum length 1.06, width 0.71. Legs. Leg I: ti length 0.97 / width 0.23, mt 0.70. II: ti 0.90 / 0.21, mt 0.69. III: ti 0.71 / 0.21, mt 0.66. IV: ti 1.12 / 0.22, mt 1,01. Spines: legs I – II, femur d 1 - 1 - 1, p 2 ap; tibia v 2 - 2 - 2; metatarsus v 2 bas. III – IV, femur d 1 - 1 - 1, p 0 - d 1 - d 1, r 0 - d 1 - d 1; patella r 1 me; tibia v p 1 - 2 - 2, p 1 - d 1 - 1 - 0, r 1 - d 1 - 1 - 0, d (r 1 - 1) bristles; metatarsus d 0 - p 1 - 2, v 2 - 0 - 2, p d 1 - 1 - 0 - 1, r d 1 - 1 - 0 - 1. Abdomen: length 2.53, width 1.54, spiracle – epigastrium 0.80. Color pattern as in male, but dark brown, abdominal and legs spots more defined. Sternum and labium darker. Epigyne: anterior plate sclerotized with accentuated anterior excavations, median excavation long and narrow (Figs 25 E, 26 D). Median field totally divided in two sub-rounded section interrupted by a large medial depression (Fig. 28 E – F). Lateral lobes with sclerotized extensions wide, swollen and apically rounded (Fig. 25 E). Copulatory ducts short joining with the accessory bulb ducts at medial level. Spermathecae rounded in median position oriented towards lateral region. Ducts of accessory bulbs long, thin, convergent (Figs 25 E – F, 26 E – F, 28). Variation: Males (n = 3): total length: 4.08 – 4.18; carapace length: 2.10 – 2.35, width: 1.59 – 1.73; abdomen length: 1.87 – 1.93, width: 1.01 – 1.25. Females (n = 5): total length: 3.89 – 4.09; carapace length: 1.73 – 2.10, width: 1.31 – 1.59; abdomen length: 1.93 – 2.53, width: 1.25 – 1.54.	en	Martínez, Leonel, Kochalka, John A., Cabra-Garcia, Jimmy, Ramírez, Martin (2025): Revealing the identity of Josa chazaliae (Simon, 1897) (Araneae: Anyphaenidae): new species and the highest altitude record for a spider in South America. Zootaxa 5566 (2): 201-242, DOI: 10.11646/zootaxa.5566.2.1, URL: https://doi.org/10.11646/zootaxa.5566.2.1
4D76240CFFA0FFEC62EBFD6BFBC8F905.taxon	distribution	Distribution. Only known from Sierra Nevada de Santa Marta Colombia (Fig. 31).	en	Martínez, Leonel, Kochalka, John A., Cabra-Garcia, Jimmy, Ramírez, Martin (2025): Revealing the identity of Josa chazaliae (Simon, 1897) (Araneae: Anyphaenidae): new species and the highest altitude record for a spider in South America. Zootaxa 5566 (2): 201-242, DOI: 10.11646/zootaxa.5566.2.1, URL: https://doi.org/10.11646/zootaxa.5566.2.1
