taxonID	type	description	language	source
533F87C4FF89732026E2FB5BFED8CEC4.taxon	description	Saussure 1899: 626; Kirby 1906: 270; Karny 1912: 8, 38; Chopard 1948: 125; Gorochov 1988: 176, 180; Otte 1997: 32.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF89732026E2FB5BFED8CEC4.taxon	diagnosis	Diagnosis. Body relatively depressed dorso-ventrally (Fig. 12). Fastigium of vertex elongated, simple, spiniform (Figs. 1, 10). Face almost flattened, with distinct tubercles, four of which are larger and spiniform. Pronotum almost covers tegmina in male (Figs. 10, 11). Mesothorax with a large seashell-shaped mesothoracic auricle. Tympanal organ on fore tibia slit shaped. Forelegs strongly developed, large, with very strong and large spines (Figs. 10, 16). Fore femur strongly widened, with a large inner spine on anterio-lateral margin; its inner and outer sides without apical spine ventrally (Figs. 4, 10). Hind tibia without apical spur dorsally (Fig. 7). Male last tergite very short, without or with a shallow small median notch on posterior margin (Figs. 13, 43). Ovipositor shorter than abdomen (Fig. 18). This genus is closely related to Malagasopus gen. nov. and Colossopus Saussure. See Table 1 for comparison of these genera.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF88732426E2FCCBFCDACC96.taxon	description	http: // lsid. speciesfile. org / urn: lsid: Orthoptera. speciesfile. org: TaxonName: 15096	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF88732426E2FCCBFCDACC96.taxon	materials_examined	Type locality. Madagascar, Mozondeva (Morondava). Lectotype male, here designated, in alcohol (MNHN).	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF88732426E2FCCBFCDACC96.taxon	description	Redescription. Male: Body size quite variable (Tab. 2). Fastigium of vertex (Figs. 1, 10, 11) forming a simple short horn, spiniform; almost conical, upper surface flattened with a wide furrow, 2.2 times longer than diameter of eye, its basal part broader than antennal scape. Fastigium of frons (Fig. 11) with a short spiniform slightly downcurved tooth. Pronotum (Figs. 10, 11) large, anterior margin narrow and convex, widened in middle, posterior margin with a distinct median incision; 3.7 times longer than its height, and only 1.1 times longer than its width. Prothoracic auditory spiracle large, partially concealed under pronotum. Opening of mesothoracic auricle large and elliptical; ventral lobe distinct. Fore coxa with an elongate spine, projected forward dorsally. Fore femur (Fig. 10) very muscular, short and broad; with a large inner spine on anterio-lateral margin; inner margin with 5 – 7 strong spines ventrally. Fore tibia (Fig. 10) with very strong, curved 4 inner spines ventrally; its outer side with smaller 4 – 5 spines. Prosternum with 2 long, U-shaped spines. Mid femur with 2 – 5 outer spines ventrally. Hind femur with 4 – 8 ventral spines on outer side. Mid and hind tibia bear ventral spines. Hind tibia without apical spur dorsally (Fig. 7). Tegmina (Figs. 10 – 12) fully concealed under pronotum, their apex visible in some specimens, reaching to half of metanotum. Abdomen (Fig. 12) large, mostly oval, anterior and posterior parts narrow, mid part broad. Last tergite (Fig. 13) short and broad, posterior margin convex with a small median notch. Cercus (Fig. 14) with a short, incurved apical tooth, inner arm very long with a distinct tooth at apex. Subgenital plate (Fig. 15) simple, with a very short styli. Female: As male. Hind margin of pronotum (Fig. 16) with a strong and broadly rounded incision. Tegmina (Fig. 17) strongly reduced, small scale like laterally, not reaching halfway down of mesonotum. Cercus simple, basal half cylindrical, apical half conical. Subgenital plate (Fig. 19) almost triangular with a distinct median carina, its posterior margin with a small median notch. Ovipositor (Fig. 18) slender, slightly upcurved, almost 1.2 – 1.3 times longer than hind femur.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF88732426E2FCCBFCDACC96.taxon	materials_examined	Colour (dry specimens). Body milky brown with distinct transverse dark bands in typical form. Head dark except fastigium of vertex, eyes and posterior margin of occiput. Anterior margin of pronotum with a narrow dark band; posterior part, (mainly of the metazona) dark, remaining part milky brown. Fore tibia light brown with indistinct longitudinal dark stripes. Apical half of femora dark. Abdominal tergites with distinct dark band on posterior margin. Male subgenital plate and cercus milky brown. Ovipositor reddish brown in apical part.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF88732426E2FCCBFCDACC96.taxon	diagnosis	Diagnosis. This species appears to be most closely related to O. janetae sp. nov. based on the structure of head, last tergite of male, female subgenital plate, the variable body size. But this species differs from it by the structure of pronotum, male cercus (with long inner arm) and colouration (body and pronotum with transverse dark bands) in typical form. It is also near to O. brongniarti sp. nov. based on its colouration (with distinct transverse bands) and male cercus with long inner arm. However, the male cercus is different to that of brongniarti (apical arm distinctly pointed with a tooth), as is the male last tergite with a wide median incision on posterior margin.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF88732426E2FCCBFCDACC96.taxon	materials_examined	Material examined. Madagascar, Mozondeva (Morondava), 1891, 2 male nymphs (Lecto- and paralectotype), (leg. M. Grandidier et M. Doulliot), in alcohol; Madagascar, 1867, 2 male nymphs (leg. M. Grandidier); Madagascar (S. O.), Forets du Fiher [en] ana, 1905, 1 male (leg. F. Geay); Prov. de Tulear, Bas Fiher [en] ana, 4 males, 8 females (leg. F. Geay); Madagask., 1 female (leg. M. Grandidier); Madagascar, Ikongo, 1902, 1 male (leg. M. Grandidier); Madagascar Sud, Akiliazivo, 25.11.1963, 2 males, 1 female (leg. D. Wintrebert) (all in MNHN). Madagascar, Morondava, February 2004, 1 male, in dead tree; Madagascar, Toliara Province, Zombitse-Vohibasia National Park, (- 22.60, 44.81) [22 ° 36 ' 0 '' S, 44 ° 48 ' 36 '' E], 17.11.2004, Coll. G. W. Beccaloni, 1 male, 1 female, in forest near park headquarters, under bark of dead tree; Madagascar, Toliara Prov., c. 20 km North of Belo Sur Tsiribihina (on main road north), 19 ° 30 ' 08.72 " S, 44 ° 30 ' 25.17 " E, 15.11.2007, Coll. G. W. Beccaloni, 3 males, 2 females, under bark of dead tree (all in NHMUK). SW Madagascar, Tulear, March 1904, 1 male (leg. S. Voltzkow) (ZMB). Madagascar, Toliara Prov., Lake Ranobe, elev. 30 m, 23 ° 00 ' 629 " S, 43 ° 336 ' E, 2 – 10 May 2003, 1 male nymph (leg. Frontier, Wilderness), grassland, transitional spiny-tamarind forest; Toliara Prov., Fiheranana [Fiherenana], elev. 65 m, 23 ° 13 ' 351 " S, 43 ° 52 ' 853 " E, 5 – 7 August 2003, 1 male nymph (leg. Frontier & Wilderness), leaf litter extraction degraded riparian forest close to water (all in CAS).	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF88732426E2FCCBFCDACC96.taxon	description	Measurements (mm). Lectotype (male) is a nymph. See Table 2 for the range of measurement of the adults.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF88732426E2FCCBFCDACC96.taxon	distribution	Distribution. S. W. of Madagascar (Fig. 110). Toliara Province: Morondava, Toliara, Ikongo, Akiliazivo, Zombitse-Vohibasia N. P., Belo Sur Tsiribihina, Fiherenana, Lake Ranobe. Habitat. This species is known to occur in spiny forest and dry deciduous forest habitats as well as in riparian forest. Individuals have been found resting under the bark of dead trees and one nymph was extracted from leaf litter.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF88732426E2FCCBFCDACC96.taxon	description	Phenology. Adults of this species have been found between November and March, and nymphs in May and August.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF88732426E2FCCBFCDACC96.taxon	discussion	Remarks. The body size is highly variable in this species (see Table 2). Piotr Naskrecki (pers. commun., 2005) believes that the largest males may be ' satellite males'. This variation in size amongst the adults of a species is also seen in some of the other species of Oncodopus (see Table 2).	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF8D732B26E2FADDFF6ACAD8.taxon	description	http: // lsid. speciesfile. org / urn: lsid: Orthoptera. speciesfile. org: TaxonName: 499685	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF8D732B26E2FADDFF6ACAD8.taxon	materials_examined	Type locality. Madagascar, Toliara Province, Andohahela National Park. Holotype male (NHMUK).	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF8D732B26E2FADDFF6ACAD8.taxon	description	Description. Male (Holotype): Body size average for genus strongly variable in other males (Tab. 2). Fastigium of vertex (Figs. 20, 21) simple, conical, elongated into a short horn; upper surface flattened with a wide furrow; 2.6 times longer than diameter of eye; its basal part broader than antennal scape; its apex slightly downcurved, straight in some males. Fastigium of frons (Fig. 21) with a large spiniform tooth. Face flattened, granulated, with 4 very distinct spiniform teeth. Pronotum (20, 21) large, anterior margin convex, posterior margin with a large incision; pronotum clearly concave in lateral view, three times longer than its height, very slightly longer than its width, in Bekily population pronotum clearly longer than wide; prozona 1.5 times longer than metazona. Prothoracic auditory spiracle large, triangular but with corners rounded, a large portion concealed under lateral lobes of pronotum. Mesothoracic auricle and its opening large, ventral lobe distinct. Prosternum with 2 long, U-shaped spines, reaching to level of ventral margin of fore coxa. Fore coxa with a long dorsal spine projected forward. Fore femur (Fig. 20) with a large inner spine on anterio-lateral margin; with 5 inner spines ventrally, in other males with 5 – 7 inner spines ventrally. Fore tibia (Fig. 20) with 4 very large inner and 4 smaller outer ventral spines. Mid femur with 3 ventral spines on outer side (other males have 2 – 4 spines). Hind femur with 2 ventral spines on outer margin of right leg, left leg with 4 spines (other males have 2 – 5 spines). Fore and mid tibia without dorsal spines. Hind tibia with 5 – 6 dorsal spines on inner margin (other males have 3 – 6 spines); with 4 apical spurs ventrally, inner 2 slightly smaller than outer ones. Tegmina fully concealed under pronotum (Figs. 20, 21), reaching to end of mesonotum (in some males they protrude a little beyond the hind margin of pronotum). Last tergite (Fig. 23) narrow and very wide, with a small median notch. Male cercus (Fig. 24) stout, short and thick; apical arm rounded, with a short spine, slightly curved inwards; inner arm short, pointed with a spine. Subgenital plate (Fig. 25) with long styli, longer than depth of posterior incision of plate. Female: Head as in male. Pronotum (Figs. 26, 27) broader than long and 3 times longer than its height; straight in lateral view; its hind margin distinctly concave; metazona very short, almost 1 / 6 of total length of pronotum; prozona 4.7 times longer than metazona in typical form. Legs as in male. Tegmina (Fig. 27) strongly reduced, scale like laterally, reaching just beyond the middle of mesonotum, in some females fully concealed under pronotum. Subgenital plate (Fig. 29) triangular, broader than long, hind margin with a small median notch. Last tergite very short, with an indistinct median notch. Cercus simple, triangular, pointed at apex. Ovipositor (Fig. 28) slender, gradually upcurved, in some females straighter; slightly longer than hind femur.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF8D732B26E2FADDFF6ACAD8.taxon	materials_examined	Colour (dry specimens). Body dark with dark reddish brown and black in typical form. Head reddish brown, labrum, eye, scapus, lighter. Pronotum dark yellowish brown in male, blackish brown in female. Apical part of fore femur blackish brown, basal and outer parts yellowish with dark spots in male, much darker in female with black. All tibiae yellowish in male, brownish in female. Apical part of mid and hind femora dark with black, basal part yellowish. Tarsi darker than tibiae with blackish brown. Abdomen pale reddish and yellowish brown with transverse black stripes and spots, darker in female. Pronotal and abdominal sternites yellowish milky brown. Subgenital plate and cercus yellowish brown. Ovipositor dark, with black, apical part reddish black.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF8D732B26E2FADDFF6ACAD8.taxon	diagnosis	Diagnosis. This new species is most closely related to O. zonatus judging by the structure of the head, the last tergite of the male, the female subgenital plate, and possibly the variable body size. But the new species differs from it in having a wider pronotum, a shorter inner arm of the male cercus, and its colouration (without distinct transverse band, especially on the pronotum) in the typical form. In some populations the shape of pronotum is also near to O. zonatus, but, it is easily recognizable by the male cercus and colouration. Without the male, it is difficult to separate the females of O. zonatus and this new species except by the colour.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF8D732B26E2FADDFF6ACAD8.taxon	materials_examined	Material examined. Madagascar, Toliara Province, Andohahela National Park: Mangatsiaka parcel, (- 24.83, 46.57) [24 ° 49 ' 48 '' S, 46 ° 34 ' 12 '' E], 30.11.2004, Coll. G. W. Beccaloni, 1 male (Holotype), 1 female; Madagascar, Toliara Province, Andohahela National Park: Tsimelahy parcel, (- 24.93, 46.65) [24 ° 55 ' 48 '' S, 46 ° 38 ' 60 '' E], 1.12.2004, Coll. G. W. Beccaloni, 1 male, 1 female, at base of [a living leaf of] Dypsis decaryi; Madagascar, Toliara Province, littoral forest fragment, nr. Saint Luce, (- 24.78, 47.18) [24 ° 46 ' 48 '' S, 47 ° 10 ' 48 '' E], 2.12.2004, Coll. G. W. Beccaloni, 1 male; Fort Douphin [Dauphin], 1 male (leg. Scott Elliot) (all in NHMUK). Bekily, Red sud de L’ile, January 1932, 3 males, 4 females, 4 male nymphs, January 1933, 1 female (leg. A. Seyrig); Col de Sakaralama, Alluaud, 1901, 1 female; Sud de Morondava, Torev de Be Fasy, 1 female; Madagascar, Rég d’Ankazoabo, 1920, 4 male nymphs, 2 female nymphs (leg. C. Le Barbier); Andohahela [National Park?], 1800 m, 1 female, 1500 m, 1 female; Madagascar, Fénerive, 1902, 1 male (leg. F. Genot); Ambatoloma [Ambatalaona?], 1907, 1 female (leg. Bernard); Madagascar, Forét d’Andampy, 60 km S. O. de Voheinar [Vohémar], Co’te N. E., 1889, 3 male nymphs (leg. M. Grandidier) (all in MNHN). Madagascar, 1 male nymph (ZMB). Ampandrandave, Sud de Madagascar, 12.1931, 3 females (leg. A Seyrig) (MNCN). Madagascar, Toliara Prov., Réserve Spéciale de cap Sainte Marie, 14.9 km 261 ° W. Marovato, elev. 160 m., 13 – 19 Feb. 2002, 25 ° 35 ' 40 " S, 45 ° 8 ' 49 " E, 1 female, 12.3 km 262 ° W. Marovato, elev. 200 m., 11 – 15 Feb. 2002, 25 ° 34 ' 54 " S, 45 ° 10 ' 6 " E, 1 female (leg. Fisher, Griswold et al.), in spiny forest thicket; Toliara Prov., Parc National d’Andohahela, Forét de Manantalingo, 33.6 km 63 ENE Amboasary, 7.6 km. 99 E Hazofotsy, 12 – 16 January 2002, 24 ° 49 ' 1 " S, 45 ° 36 ' 36 " E, elev. 150 m., 1 male (leg. Fisher, Griswold et al.), in spiny forest thicket; Toliara prov., Parc National de Zombitse, 19.8 km 8 E Sakaraha, elev. 770 m., 5 – 9 Feb. 2003, 22 ° 50 ' 36 " S, 44 ° 42 ' 36 ", 1 female (leg. Fisher, Griswold et al.), in tropical dry forest (all in CAS).	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF8D732B26E2FADDFF6ACAD8.taxon	description	Measurements (mm). Holotype (male): Body length: 44; pronotum length: 16.4; width of pronotum: 14.6; fore femur length: 16.1; width of fore femur: 6.8; fore tibia length: 15.9; hind femur length: 18.7. See Table 2 for the range of measurement of the other specimens including females.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF8D732B26E2FADDFF6ACAD8.taxon	etymology	Etymology. This species name is dedicated to Janet Beccaloni who provided considerable help to the second author with collecting work in Madagascar.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF8D732B26E2FADDFF6ACAD8.taxon	distribution	Distribution. Southern and eastern Madagascar (Fig. 110). Toliara Province: Morondava, Toliara, Bekily, Fort Dauphin, Ankazoabo, Andohahela N. P., Marovato, Zombitse-Vohibasia N. P. Toamasina Province (see remarks below): Fénerive, Ambatoloma [Ambatalaona?], Forét d’Andampy. Habitat. This species is known from littoral forest, spiny forest and tropical dry forest. Records for rainforest are dubious (see remarks below). It has been found under dead bark and in the natural cavity found at the base of living leaves of the palm Dypsis decaryi.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF8D732B26E2FADDFF6ACAD8.taxon	description	Phenology. Adult individuals have been found between November and February and nymphs in January.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF8D732B26E2FADDFF6ACAD8.taxon	discussion	Remarks. We regard the three old (1889, 1902, 1907) locality records in the Toamasina Province as dubious since these sites are located in the humid climatic zone (Cornet 1974) which has rainforest vegetation, whereas all other records are from dry forests in arid regions with a long dry season. Very extensive collecting of arthropods since 1992 at more than 300 sites all over Madagascar (Fisher 2017), including many in the humid climatic zone, by a team led by Brian Fisher of the California Academy of Sciences has failed to collect any specimens of O. janetae, or indeed any of the other taxa discussed in our study, in rainforest habitats. Also note that old collecting localities are often difficult to identify in Madagascar due to the large number of synonymous locality names and because geographic coordinates are absent from the data labels. The body size is highly variable in this species as it is in O. zonatus (see the remarks under O. zonatus and Table 2).	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF82732926E2FD18FF6ACCB0.taxon	description	http: // lsid. speciesfile. org / urn: lsid: Orthoptera. speciesfile. org: TaxonName: 499686	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF82732926E2FD18FF6ACCB0.taxon	materials_examined	Type locality. Madagascar, Toliara Province, S. Morondava, Belo Sur Mer. Holotype male (NHMUK).	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF82732926E2FD18FF6ACCB0.taxon	description	Description. Male (Holotype): Body size average for genus (Tab. 2) (in one male body size quite large). Fastigium of vertex (Figs. 30, 31) simple, conical, forming a short horn, upper surface flattened with a narrow longitudinal median furrow, 2.3 times longer than diameter of eye, basal part 1.4 times wider than antennal scape. Fastigium of frons (Fig. 31) with a spiniform slightly downcurved tooth. Face flattened, strongly granulated; each tubercle like a small tooth, of which the lower two are larger. Pronotum (30, 31) long, 3.2 times longer than its height and 1.2 times longer than its width; anterior margin convex; posterior margin with an angular, large median incision; upper surface slightly concave in lateral view; prozona 1.9 times longer than metazona. Prothoracic auditory spiracle large, oval; a small portion of its opening concealed under pronotum. Mesothoracic auricle small, short; its opening almost triangular; ventral lobe short. Prosternum with two long, V-shaped spines, reaching to level of ventral margin of fore coxa; widely separated towards to apex. Fore coxa with a short spine projecting forward. Femora without dorsal spines. Fore femur (Fig. 30) with a large inner spine on anterio-lateral margin; with 6 inner spines ventrally. Fore tibia (Fig. 30) with 5 inner and 5 outer spines ventrally. Mid femur with 2 ventral spines on outer margin. Hind femur with 2 outer spines ventrally (1 – 4 spines in other males). Fore and mid tibia without dorsal spines. Hind tibia with 7 dorsal spines on inner margin (other males with 5 – 6 spines); with 4 apical spurs ventrally, inner 2 slightly smaller than outer ones; without apical spurs dorsally. All tibia with ventral spines on both margins. Tegmina (Figs. 30, 31) concealed under pronotum, its apex visible from the posterior incision of pronotum in dorsal view, reaching to half of metanotum. Last tergite (Fig. 33) wider than long, posterior margin with a shallow, wide, rounded median incision. Cercus (Fig. 34) stout, apical part with a very small tooth (which is not spiniform) on its ventral margin; inner arm very long, its apex slightly bent backward, pointed with a distinct tooth. Subgenital plate (Fig. 35) almost as long as wide, its styli distinctly longer than depth of median incision. Female: Face as in male. Fastigium of vertex (Fig. 36) short, 1.8 times longer than diameter of eye, its basal part 1.5 times wider than antennal scape. Pronotum (Figs. 36, 37) as wide as long; posterior margin with a large rounded incision; pronotum 2.9 times longer than its height; prozona 3 times longer than metazona. Tegmina (Fig. 37) strongly reduced, scale like laterally, reaching to half of mesonotum. Prosternal spines V-shaped, slightly shorter than that of male. Fore femur (Fig. 36) with 6 – 7 ventral spines on inner margin and a large inner spine on anterio-lateral margin. Tibial spines as in male. Mid femur with 2 outer spines ventrally, in other females with 2 – 4 spines. Hind femur with 2 ventral spines on outer side. Last tergite very short and wide, its posterior incision distinct. Cercus stout, basal part broad, apical third strongly and sharply narrowed, its apex slightly incurved and pointed. Subgenital plate (Fig. 39) almost triangular with a median carina, its apex with a small rounded incision. Ovipositor (Fig. 38) clearly longer than hind femur, slightly upcurved.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF82732926E2FD18FF6ACCB0.taxon	materials_examined	Colour (dry specimens). Body milky brown with distinct transverse dark reddish-brown bands. Head reddish brown. Eye, fastigium of vertex, labrum, maxillary and mandibulary palps slightly lighter. Anterior part of pronotum milky brown, posterior part (metazona) very dark with reddish-brown. In female metazona dark brown. All tibiae milky brown. Apical part of each femur dark, with a large black or reddish-black band. Posterior half of first abdominal tergite in male and the whole of it in the female dark, black or reddish brown; posterior part of each abdominal tergite with transverse dark band, black or reddish brown. All sternites, cerci and subgenital plate yellowish milky brown. Ovipositor milky brown in basal 1 / 3, reddish brown in remaining part.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF82732926E2FD18FF6ACCB0.taxon	diagnosis	Diagnosis. Based on the colour pattern (with distinct transverse dark bands), the male cercus with a long inner arm, and the shape of the ovipositor this new species appears to be most closely related to O. zonatus. It differs from it by the absence of a distinct apical spine on the male cercus, the larger median incision of the last tergite in both sexes, the more pronounced tubercles of the face and the female cercus. It is recognized by the shape of the male cercus and the last tergite.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF82732926E2FD18FF6ACCB0.taxon	materials_examined	Material examined. Madagascar, Toliara Province, Belo Sur Mer (South of Morondava), 20 ° 44 ' 53.63 " S, 43 ° 59 ' 36.97 " E, 22.11.2007, Coll. G. W. Beccaloni, 2 males (including Holotype), 2 females, under bark of dead tree; Madagascar, S. Centr. Madagascar, 1 male [no more data]; Toliara province, Forest near village of Mangily, c. 27 km N. of Toliara, (- 23.15, 43.60) [23 ° 8 ' 60 '' S, 43 ° 36 ' 0 '' E], 11.11.2004, Coll. G. W. Beccaloni, 1 female, under bark of dead tree; Madagascar, Foret de Zombitsy [Zombitse-Vohibasia N. P.], near Sakaraha, 650 m, 16.12.1959, 1 female (leg. E. S. Ross) (all in NHMUK).	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF82732926E2FD18FF6ACCB0.taxon	description	Measurements (mm). Holotype (male): Body length: 40.3; pronotum length: 14.2; width of pronotum: 12.1; fore femur length: 14.5; width of fore femur: 5.7; fore tibia length: 13.3; hind femur length: 15. See Table 2 for the range of measurement of the other specimens including females.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF82732926E2FD18FF6ACCB0.taxon	etymology	Etymology. This species name is dedicated to Charles Brongniart who was the first to discover this group of insects in 1897. He is the author of the genus Oncodopus and the species Oncodopus zonatus and Colossopus redtenbacheri.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF82732926E2FD18FF6ACCB0.taxon	distribution	Distribution. South-west and south-central Madagascar (Fig. 110). Toliara Province: Toliara, Mangily, Foret de Zombitsy [Zombitse-Vohibasia N. P.], Morondava. Habitat. This species is known from spiny forest, dry costal scrub and dry deciduous forest. It has been found under the bark of dead trees.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF82732926E2FD18FF6ACCB0.taxon	description	Phenology. Adults have been found in November and December.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF82732926E2FD18FF6ACCB0.taxon	discussion	Remarks. The body size is variable as in above 2 species (see under the Remarks section for O. zonatus and Table 2).	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF80732E26E2FAFAFE83CF77.taxon	description	http: // lsid. speciesfile. org / urn: lsid: Orthoptera. speciesfile. org: TaxonName: 499687	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF80732E26E2FAFAFE83CF77.taxon	materials_examined	Type locality. Madagascar, Fianarantsoa Prov., South of Ambohimahasoa. Holotype male (NHMUK).	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF80732E26E2FAFAFE83CF77.taxon	description	Description. Male (Holotype): Body size smallest in the genus (Tab. 2). Fastigium of vertex (Figs. 40, 41) simple, almost conical, forming a short horn, upper surface flattened without furrow, twice as long as eye diameter, its basal part 1.5 times wider than antennal scape, its apex slightly downcurved. Fastigium of frons (Fig. 41) with a spiniform tooth. Face flattened, with few indistinct tubercles, with 2 distinct teeth. Pronotum (Figs. 40, 41) slightly longer than wide, 3.8 times longer than its height and 1.1 times longer than its width; anterior margin convex, posterior margin slightly concave; metazona long, only 1.4 times shorter than prozona in middle; pronotum strongly concave in lateral view. Prothoracic auditory spiracle small almost triangular, only a small part visible in lateral view. Mesothoracic auricle small, its inner gap oval, lacking ventral lobe. Prosternum with 2 short, U-shaped spines, almost parallel, not reaching to level of ventral margin of fore coxa. Fore coxa with a short spine projecting forward. Fore femur (Fig. 40) with a large inner spine on anterio-lateral margin; with 5 ventral spines on inner margin (other males have 3 – 5 spines). Fore tibia (Fig. 40) with 4 inner and 4 outer spines ventrally. Mid femur with 1 outer spine ventrally (some males lack this spine). Hind femur without spines. Hind tibia with 2 – 4 dorsal spines on inner margin (other males have 3 – 5 spines); 4 apical spurs ventrally, inner two slightly smaller than outer ones. Tegmina (Figs. 40, 41) fully concealed under pronotum, reaching to mid point of metanotum. Last abdominal tergite (Fig. 43) very short and wide, its posterior margin slightly concave, without notch or incision. Cercus (Fig. 44) short, apical part rounded, with an incurved apical tooth; with 2 inner arms, upper one very short, pointed with a tooth; lower one longer and pointed with a tooth. Subgenital plate (Fig. 45) slightly longer than wide, its styli broad and longer than depth of posterior incision. Female: Head as in male. Pronotum (Figs. 46, 47) 1.2 times wider than long and 2.7 times longer than its width; posterior margin strongly concave; prozona 3.5 times longer than metazona. Ventral half of prothoracic auditory spiracle visible in lateral view. Fore femur (Fig. 46) with 3 – 6 ventral spines on inner margin. Mid and hind femora without or with 1 – 2 ventral spines on outer margin. Hind tibia with 3 – 5 dorsal spines on inner margin. Spines of prosternum reaching to level of ventral margin of fore coxa, in some specimens a little shorter. Tegmina (Fig. 47) strongly reduced, scale like laterally, not reaching to half of mesonotum, largely concealed under pronotum. Last tergite short and wide, its posterior margin convex, without median incision. Subgenital plate (Fig. 49) triangular, with a median carina, posterior margin with a small incision. Cercus triangular, slightly incurved, pointed at apex with an indistinct spine. Ovipositor (Fig. 48) longer than hind femur, slightly upcurved.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF80732E26E2FAFAFE83CF77.taxon	materials_examined	Colour (dry specimens). Body various shades of greenish brown, black and yellowish brown. Face brown in male, blackish brown in female. Dorsal surface of head greenish brown with irregular black pattern. Pronotum greenish brown, with short longitudinal black stripes. Fore femur and tibia milky yellowish brown; dorsal surface of fore femur with distinct black spots, ventral surface completely black including spines; fore tibial spines black. Mid femur as fore femur. Hind femur dark apically, basal part with dark spots. Anterior part of each abdominal tergite except last 2 – 3 ones with black transverse band on both sides, not touching in middle, behind it with a faint band; hind margin of abdominal tergites completely black. All sternites, male cercus and subgenital plate unicolorous milky brown. Female subgenital plate with a black macula in middle. Ovipositor milky brown at base, remaining part dark with black or brownish black.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF80732E26E2FAFAFE83CF77.taxon	diagnosis	Diagnosis. This species is the smallest member of the genus. It is unique in the shape of the male pronotum and cercus, the last tergite in both sexes, the ovipositor and colouration. It appears to be most closely related to O. soalalaensis sp. nov. judging by the shape of posterior margin of pronotum and the last tergite of the male, the tridentate male cercus and its small body size. It differs from it by the general structure of pronotum in both sexes, the male cercus, the fore tibia with 4 ventral spines on both sides (O. soalalaensis has 5 spines), the shape of its shorter ovipositor, its colouration and its distribution.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF80732E26E2FAFAFE83CF77.taxon	materials_examined	Material examined. Madagascar, Fianarantsoa Prov., beside R. N. 7 south of Ambohimahasoa, (c. - 21.17, 47.25) [21 ° 17 ' S, 47 ° 25 ' E], 13.11.2004, Coll. G. W. Beccaloni, 1 male (Holotype), 1 female, under a stone; Antananarivo, (- 18.92, 47.52) [18 ° 55 ' 12 '' S, 47 ° 31 ' 12 '' E], on R. N. 7 to the south of the city, 12.11.2004, Coll. G. W. Beccaloni, 2 females, under a stone, (all in NHMUK). Madagascar, Pic de Ivohibe, 2200 m, 8.11.1950, 3 males, 1 female, 1 female nymph (leg. A. R.); Madagascar-centre, Plateau Soaindrana, 2090 m, Andringitra-Ambalavao, 15.1.1958, 1 female (leg. R. Paulian); Andringitra centre, plat. Andohariana, 2000 – 2100 m, 9.11 – 10.12.1970, 1 female (all in MNHN).	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF80732E26E2FAFAFE83CF77.taxon	description	Measurements (mm). Holotype (male): Body length: 30.9; pronotum length: 9.5; width of pronotum: 8.2; fore femur length: 7.9; width of fore femur: 3.6; fore tibia length: 7.5; hind femur length: 9.9. See Table 2 for the range of measurement of the other specimens including females.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF80732E26E2FAFAFE83CF77.taxon	etymology	Etymology. This species name is dedicated to Henri Louis Frédéric de Saussure, who was the second person to study these insects (in 1899). Saussure is the author of the genus Colossopus and the species C. grandidieri.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF80732E26E2FAFAFE83CF77.taxon	distribution	Distribution. Central and south-central Madagascar (Fig. 110). Antananarivo Province: Antananarivo, Soavinandriana. Fianarantsoa Province: Ambohimahasoa, Andringitra, Ambalavao, Ivohibe. Habitat. This species occurs in seasonally dry habitats on the Central Plateau of Madagascar which are largely grassland today but which were probably sclerophyllous woodland in the past. It is known to occur up to 2200 metres — the highest altitude of any species in this group. It has been found under stones.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF80732E26E2FAFAFE83CF77.taxon	description	Phenology. Adults have been found between November and January and one nymph (instar unknown) in November.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF87733226E2F8BBFE05C889.taxon	description	http: // lsid. speciesfile. org / urn: lsid: Orthoptera. speciesfile. org: TaxonName: 499688 Type locality. Madagascar, Mahajanga, Soalala Reserve. Holotype male (MNHN). Description. Male (Holotype): Body size small for genus (Tab. 2). Fastigium of vertex (Figs. 50, 51) simple, elongated into a short horn, upper surface flattened, 1.7 times longer than eye diameter, its basal part 1.5 times wider than antennal scape. Fastigium of frons (Fig. 51) with a small spiniform tooth. Face convex, not distinctly flattened, with tubercles. Pronotum (Figs. 50, 51) almost saddle shaped, 1.1 times longer than wide and 3.1 times longer than its height; depression of typical sulcus deep and distinct; anterior margin slightly convex, posterior margin concave in middle; prozona 1.5 times longer than metazona. Tegmina (Fig. 50, 51) almost concealed under pronotum, only the apex visible from above; reaching to a little beyond the middle of the metanotum. Prothoracic auditory spiracle large, oval; its dorsal half concealed under lateral lobe of pronotum. Mesothoracic auricle small, without ventral lobe; its dorsal part concealed under lateral lobe of pronotum. Prosternum with two, U-shaped spines, almost parallel, not reaching to level of ventral margin of fore coxa. Fore coxa with a spine projecting forward. Fore femur (Fig. 50) with a large inner spine on anterio-lateral margin and 6 ventral spines on inner margin. Fore tibia (Fig. 50) with 5 inner and 5 outer spines ventrally. Mid and hind femora with 2 ventral spines on outer margin apically. Hind tibia with 6 dorsal spines on inner side (5 – 6 spines in other males); with 4 apical spurs ventrally, all of which almost equal in length. Last abdominal tergite (Fig. 53) short and broad, posterior margin convex without median notch (other males have a very indistinct median notch). Cercus (Fig. 54) stout, with 3 arms; apical arm short, roundly curved inwards, pointed with a spine; basal inner arm short and broad, pointed with a tooth; median inner arm larger, near to ventral surface of cercus, pointed at apex with a large tooth. Subgenital plate (Fig. 55) longer than wide, its styli longer than depth of posterior median incision. Female: Head as in male. Fastigium of vertex (Figs. 56, 57) 1.3 times wider than antennal scape. Pronotum (Fig. 56, 57) short and wide, saddle shaped, 1.2 times wider than long; anterior margin convex, posterior margin broadly concave; prozona 3 times longer than metazona. Tegmina (Fig. 56) reduced, like a very small scale placed on anterio-lateral margin of mesonotum. Only a small portion of prothoracic auditory spiracle covered by lateral lobe of pronotum. Mesothoracic auricle without ventral lobe, not covered by pronotum. Legs and their spines as in male. Last abdominal tergite very short and broad, posterior margin without median notch. Cercus short conical, its apex slightly incurved. Subgenital plate (Fig. 59) triangular, with distinct median carina; its apex with a small notch, hind margin curved upwards in posterior view. Ovipositor (Fig. 58) distinctly longer than hind femur, slightly shorter than abdomen, almost straight, apical part very slightly upcurved.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF87733226E2F8BBFE05C889.taxon	materials_examined	Colour (dry specimens). Body various shades of light brown. Upper portion of face yellowish brown, lower part darker with reddish brown; labrum black; upper surface of head greenish brown; eye darker. Pronotum slightly reddish brown with some short longitudinal stripes; posterior half of metazona greenish brown in male. Legs yellowish brown; mid and hind tibia of male reddish brown. Abdomen slightly darker reddish brown, with indistinct transverse dark bands in male, greenish brown in female. All sternites reddish brown in male and concolorous in female. Basal part of ovipositor greenish brown, turning to reddish brown in apical part.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF87733226E2F8BBFE05C889.taxon	diagnosis	Diagnosis. This species has the following unique characters: the shape of its pronotum in both sexes, the morphology of the male cercus, the shape of the ovipositor and its colour. Features indicating it is closely related to O. saussurei sp. nov. are the tridentate male cercus, the small body size, and the last tergite without median incision. However, this new species differs from O. saussurei by the fore tibia with 5 inner and 5 outer spines ventrally (O. saussurei has 4 spine), the shape of the pronotum, features of the male cercus, the shape of its longer ovipositor and its colour. The fore tibia has a similar structure to that of O. brongniarti sp. nov., having 5 ventral spines on both sides. But, all other characters are very different from it.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF87733226E2F8BBFE05C889.taxon	materials_examined	Material examined. Madagascar, Mahajanga, Soalala Reserve nat. VIII, 3 males (including Holotype), 1 female nymph; Namoroka, 9.1952, 1 male nymph; Madagascar centre, Rég. de Bétafo d’Ambositra, et de Miandrivazo, 1905, 1 female (leg. Bauet) (all in MNHN).	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF87733226E2F8BBFE05C889.taxon	description	Measurements (mm). Holotype (male): Body length: 33.6; pronotum length: 10.6; width of pronotum: 9.4; fore femur length: 8.8; width of fore femur: 3.7; fore tibia length: 9.4; hind femur length: 13. See Table 2 for the range of measurement of the other specimens including female.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF87733226E2F8BBFE05C889.taxon	etymology	Etymology. The species name is derived from “ Soalala ” is the type locality of this new species.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF87733226E2F8BBFE05C889.taxon	distribution	Distribution. Central and western Madagascar (Fig. 110). Mahajanga Province: Soalala, Namoroka. Fianarantsoa-Toliara Province: Betafo, between Ambositra and Miandrivazo. Habitat. The region in which this species occurs contains dry deciduous forest. Living specimens have not been studied.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF87733226E2F8BBFE05C889.taxon	description	Phenology. A nymph of this species was found in September. The exact collecting dates of the adults were not recorded on the labels.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF9B733326E2FB41FBCEC8A4.taxon	materials_examined	Type species. Malagasopus desutterae sp. nov., here designated.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF9B733326E2FB41FBCEC8A4.taxon	description	Description and Diagnosis. Body slightly depressed dorso-ventrally (Fig. 62). Head wide. Fastigium of vertex (Figs. 2, 60) relatively elongated, with 3 short spiniform arms all of which are slightly directed upwards. Face almost flattened, with distinct tubercles, four of which larger, spiniform. In M. desutterae a large portion of the male tegmina is covered by the pronotum and only the apical parts are visible from above (Fig. 60), while in the other species, M. meridianus, a large apical portion of the male tegmina is visible from above (Fig. 70). Mesothorax with a large seashell-shaped mesothoracic auricle. Tympanal organ on fore tibia slit shaped. Forelegs (Figs. 60, 66) strongly developed, large, with very strong and large spines. Fore femur (Figs. 5, 60) strongly widened, with a large inner spine on anterio-lateral margin; its outer and inner sides without apical spine ventrally. Hind tibia without apical spur dorsally (Fig. 8). Last male tergite (Fig. 63) long, with a narrow median incision on posterior margin. Ovipositor (Fig. 68) slightly shorter than abdomen. For more details, see the descriptions of M. desutterae sp. nov. and M. meridianus sp. nov. below. This new genus appears to be closely related to Oncodopus judging by the structure of pronotum, fore femur, hind tibia and by the slightly protruding male tegmina in M. desutterae. It differs from Oncodopus in the tridentate fastigium of its vertex (one simple, elongated spine in Oncodopus), protruding male tegmina (almost covered by pronotum in Oncodopus), the elongated hind margin of the last male tergite (very short in Oncodopus) and the wider head especially in the female. It shares with Colossopus the tridentate fastigium of the vertex, the elongated hind margin of the last male tergite and the strongly protruding tegmina of the male in one species, M. meridianus. It differs from Colossopus in the morphology of the fastigium of the vertex (with very short arms all of which are directed slightly upwards whilst in Colossopus the longer 2 lateral arms are parallel and directed slightly downwards and the longer middle arm is upcurved), the shape of pronotum, the structure of the fore femur (the new genus lacks an apical spine on both sides ventrally whilst Colossopus possesses such spines), the hind tibia without apical spur dorsally (Colossopus, has an apical spur dorsally on inner side), the shorter ovipositor and the wider head especially in female.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF9B733326E2FB41FBCEC8A4.taxon	etymology	Etymology. The generic name is derived from “ Malagasy ” meaning Madagascan.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF9A733126E2FD60FC49CC6A.taxon	description	http: // lsid. speciesfile. org / urn: lsid: Orthoptera. speciesfile. org: TaxonName: 499690	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF9A733126E2FD60FC49CC6A.taxon	materials_examined	Type locality. S. Madagascar, Beheloka. Holotype male (MNHN).	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF9A733126E2FD60FC49CC6A.taxon	description	Description. Male (Holotype): Head wide, 1.2 times longer than its width. Fastigium of vertex (Figs. 2, 60, 61) short and broad, 1.2 times narrower than eye, 1.5 times wider than antennal scape; triarmed, its arms short. Face slightly convex, almost flattened, with 2 rows of vertical tubercles, and 2 spiniform tubercles placed both sides of frons. Fastigium of frons (Fig. 61) with a short, distinct spiniform tooth. Pronotum (Figs. 60, 61) slightly longer than wide; depression of typical sulcus very deep and distinct; anterior margin convex, posterior margin with a rounded median incision; prozona 1.5 times longer than metazona in middle, 2.8 times longer than its height. Tegmina (Figs. 60, 61) slightly protrude from under pronotum, only their apex visible from above; reaching slightly beyond of posterior margin of metanotum. Prothoracic auditory spiracle fully concealed under lateral lobe of pronotum. Mesothoracic auricle small, dorsal half covered by lateral lobe of pronotum, its inner gap elliptical; without ventral lobe. Fore femur (Figs. 5, 60) with a small inner spine on anterio-lateral margin, 4 ventral spines on inner side, no apical spine ventrally, outer side without ventral spines. Fore tibia (Fig. 60) with 4 strong inner and 4 smaller outer spines ventrally. Mid and hind femora without spines. Hind tibia with 6 dorsal spines on outer side; without apical spur dorsally (Fig. 8); with 4 apical spurs ventrally, inner 2 slightly shorter than outer ones. Prosternum with 2, U-shaped spines, not reaching to level of ventral margin of fore coxa. Fore coxa with a forward projecting dorsal spine. Last abdominal tergite (Fig. 63) long and broad, 2.6 times wider than long, posterior margin elongated with a very narrow small median incision, like a split. Cercus (Fig. 64) short and broad, apical part strongly incurved; with 2 arms, dorsal arm short and narrow, pointed with a sharp tooth, ventral arm long, broad with a larger apical tooth. Subgenital plate (Fig. 65) almost twice as long as its width, with a distinct median carina; posterior margin with a wide triangular median incision; its styli small, almost rectangular, longer than posterior incision of plate. Female: Head very wide, almost rounded in frontal view, only 1.1 times longer than its width. Fastigium of vertex (Figs. 66, 67) short and broad; triarmed, all arms directed slightly upwards; as wide as diameter of eye and almost twice as wide as antennal scape. Fastigium of frons (Fig. 67) with a short spiniform tooth. Pronotum (Figs. 66, 67) wider than long, typical sulcus very distinct like a deep depression; anterior margin convex, posterior margin concave; almost twice as long as its height; prozona 2.5 times longer than metazona. Tegmina strongly reduced, like small scales, covered by pronotum. Prothoracic auditory spiracle almost rounded, only a small part visible in lateral view. Mesothoracic auricle like an oval bowl, without ventral lobe. Forelegs as in male but larger. Mid and hind legs as in male but hind tibia with very small 5 dorsal spines on inner side. Prosternum with 2, Ushaped spines. Last abdominal tergite very narrow and broad, with a distinct narrow median incision which separates the hind margin into 2 rounded lobes. Cercus conical, short and broad, its apex slightly incurved. Subgenital plate (Fig. 69) short and broad, median part swollen, 2.6 times wider than long, triangular but hind margin with a large deep incision. Ovipositor (Fig. 68) sword shaped, slightly upcurved, 1.5 times longer than hind femur and slightly shorter than abdomen.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF9A733126E2FD60FC49CC6A.taxon	materials_examined	Colour (dry specimens). Body various shades of brown in male and milky brown in female. Lower margin of frons and mandibles dark, reddish brown. Labrum light, yellowish brown. Spines of fore femur and basal parts of fore tibial spines black. Dorsal surface of fore femur with black stripes and black spots; outer sides of mid and hind femora with irregular, large black spots and stripes. Pronotum with 2 median and 2 lateral short black longitudinal stripes in prozona; metazona with several dark brown spots. Visible part of male tegmina with several small black spots. Abdominal tergites reddish brown in male and milky brown in female, with indistinct irregular transverse dark stripes. Basal third of ovipositor milky brown, apical two thirds dark.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF9A733126E2FD60FC49CC6A.taxon	diagnosis	Diagnosis. This new species shows its close affinity to M. meridianus sp. nov. by the structure of the legs, the last male tergite, the subgenital plate and the cercus. It differs from it in the shape of male pronotum with longer metazona (4.5 mm in the middle), the length of the male tegmina (only hind margin visible from above, almost fully covered by pronotum), the structure of the male cercus, the number of spines on the fore tibia (4 inner and 4 outer spines ventrally in this species, and 5 inner and 5 outer spines in M. meridianus sp. nov.). Although it has a somewhat similar pronotum and tegmina to Oncodopus spp. the generic characters given in the table 1 and under the genus Malagasopus show that these taxa are very different.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF9A733126E2FD60FC49CC6A.taxon	materials_examined	Material examined. Madagascar sud, Beheloka, April 1963, 1 male (Holotype), 2 males nymph, 2 females nymph (MNHN). Madagascar, Toliara Prov., Mahafaly Plateau, 6.2 km 74 ENE Itampolo, elev. 80 m, 21 – 25 Feb. 2002, 24 ° 39 ' 13 " S, 43 ° 59 ' 48 " E, 1 female (leg. Fisher, Griswold et al.), in spiny forest thicket (CAS).	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF9A733126E2FD60FC49CC6A.taxon	description	Measurements (mm). Holotype (male): Body length: 31.2; pronotum length: 11.9; width of pronotum: 10.3; fore femur length: 8.2; width of fore femur: 3.4; fore tibia length: 8.4; hind femur length: 12.4. See Table 3 for the measurement of the female.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF9A733126E2FD60FC49CC6A.taxon	etymology	Etymology. This new species name is dedicated to our colleague Dr. Laure Desutter-Grandcolas from the MNHN Paris in recognition of her very important contributions to Orthoptera research. She very kindly assisted M. Ünal during his research visits to the MNHN and loaned the specimens of this new species as well as many others.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF9A733126E2FD60FC49CC6A.taxon	distribution	Distribution. South-west of Madagascar (Fig. 111). Toliara Province: Beheloka, NE Itampolo. Habitat. This species is known from spiny forest. Living specimens have not been studied.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF9A733126E2FD60FC49CC6A.taxon	description	Phenology. Adults have been found in February and nymphs in April.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF98733726E2FA7EFD19CBF9.taxon	description	http: // lsid. speciesfile. org / urn: lsid: Orthoptera. speciesfile. org: TaxonName: 499691	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF98733726E2FA7EFD19CBF9.taxon	materials_examined	Type locality. Madagascar, Toliara, Raharizonina. Holotype male (MNHN).	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF98733726E2FA7EFD19CBF9.taxon	description	Description. Male (Holotype): Head wide, 1.3 times longer than its width. Fastigium of vertex (Figs. 70, 71) with tridentate, lateral 2 denticles very small, but probably not developed sufficiently (its normal form is probably as in M. desutterae, but it may has been damaged during immature stages); its basal part 1.5 times narrower than eye, 1.3 times wider than antennal scape. Face slightly convex, with distinct tubercles. Fastigium of frons with a small spiniform tooth. Pronotum (Figs. 70, 71) slightly longer than wide, 3.1 times longer than its height; anterior margin convex, posterior margin with a strong median incision; depression of typical sulcus wide and shallow; prozona 3.4 times longer than metazona in middle. Tegmina (Figs. 70, 71) strongly protrude from under pronotum, a large portion visible from above, reaching to slightly beyond posterior margin of metanotum. Prothoracic auditory spiracle concealed under lateral lobe of pronotum. Mesothoracic auricle small, inner gap elliptical, ventral lobe distinct. Fore femur (Fig. 70) with a small inner spine on anterio-lateral margin; with 5 ventral spines on inner side, no apical spine ventrally; outer side of fore femur without ventral spines. Fore tibia (Fig. 70) with 5 large inner and 5 smaller outer spines ventrally. Mid and hind femora without spines. Hind tibia with 5 dorsal spines on outer side, without dorsal apical spur; with 4 apical spurs ventrally, inner 2 slightly shorter than outer ones. Prosternum with 2 U-shaped spines, not reaching to level of ventral margin of fore coxa. Fore coxa with a dorsal spine projecting forward. Last abdominal tergite long and broad, 2.2 times longer than wide, posterior margin elongated with a small, very narrow median incision, like a split. Cercus short and broad, strongly incurved apically; with 2 incurved arms, dorsal arm short and broad, pointed with a distinct tooth, ventral arm longer, pointed with a larger tooth. Subgenital plate (Fig. 75) 1.3 times longer than wide, with a distinct median carina; posterior margin with a narrow triangular median incision; its styli small and rounded, almost as long as depth of posterior incision of plate. Female: Unknown.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF98733726E2FA7EFD19CBF9.taxon	materials_examined	Colour (dry specimens). Body light brown with some darker shades. Lower margin of frons and mandibles dark reddish brown. Spines of forelegs dark. Wing membrane between veins of apical part of tegmina dark. Hind margins of abdominal tergites with transverse light milky brown bars.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF98733726E2FA7EFD19CBF9.taxon	diagnosis	Diagnosis. This new species is similar to M. desutterae sp. nov. in the structure of its legs, last male tergite, subgenital plate and cercus. It different from M. desutterae in the shape of male pronotum with shorter metazona (1.9 mm in the middle), the length of the male tegmina (a large part visible from above, not fully covered by pronotum), structure of the male cercus, the spine number of the fore tibia (it has 5 inner and 5 outer spines ventrally whilst M. desutterae has 4 inner and 4 outer spines). Whilst the pronotum and tegmina of M. meridianus are similar to Colossopus spp. there are many other differences (see the generic characters given in the Table 1 and under the genus Malagasopus).	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF98733726E2FA7EFD19CBF9.taxon	materials_examined	Material examined. Madagascar, Sud Quest, Tulear, P. K. 860, Raharizonina, 7.1959, 1 male (Holotype) (MNHN).	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF98733726E2FA7EFD19CBF9.taxon	description	Measurements (mm). Holotype (male): Body length: 33.1; pronotum length: 10.6; width of pronotum: 9.9; fore femur length: 8.9; width of fore femur: 3.3; fore tibia length: 9.4; hind femur length: 13.6.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF98733726E2FA7EFD19CBF9.taxon	etymology	Etymology. The new species name “ meridianus ” means southern in Latin. This species in common with many of the other species in this paper, is found in South Madagascar.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF98733726E2FA7EFD19CBF9.taxon	distribution	Distribution. South-west of Madagascar (Fig. 111). Toliara Province: S. E. Toliara, Raharizonina. Habitat. The region in which this species occurs has spiny forest. Living specimens have not been studied.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF98733726E2FA7EFD19CBF9.taxon	description	Phenology. This species has been found in July as an adult.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF98733726E2FA7EFD19CBF9.taxon	discussion	Remarks. Lateral denticles of fastigium of vertex are very small and not in natural forms. Possibly they may have been damaged during immature stages.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF9E733726E2FC38FE42CDA1.taxon	description	Karny 1912: 8, 39; Kirby 1906: 270; Chopard 1948: 125; Gorochov 1988: 176; Otte 1997: 27; Ünal & Beccaloni 2008: 34.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF9E733726E2FC38FE42CDA1.taxon	diagnosis	Diagnosis. Body relatively depressed dorso-ventrally (Fig. 78). Fastigium of vertex distinctly elongated, with 3 long spiniform arms (Figs. 3, 76). Facet almost flattened, with distinct tubercles four of which larger, spiniform. Male tegmina wavy, protruding from under pronotum and almost half of them visible from above (Figs. 76, 77). Tympanal organ on fore tibia slit shaped. Mesothorax with a large seashell shaped mesothoracic auricle. Forelegs (Fig. 76, 83) strongly developed, large, with very large and robust spines. Fore femur thickened, with a large inner spine on anterio-lateral margin; its outer and inner sides armed with a large apical spine ventrally (Figs. 6, 76). Hind tibia with an apical spur dorsally on inner side (Fig. 9). Male last tergite long, posterior margin elongated backward, with a distinct narrow median incision (Fig. 80). Ovipositor longer than abdomen (Fig. 85). This genus is most closely related to Malagasopus gen. nov. and Oncodopus Brongniart. See Table 1 for comparison of these genera.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF9D733B26E2FEE8FE03CA44.taxon	description	http: // lsid. speciesfile. org / urn: lsid: Orthoptera. speciesfile. org: TaxonName: 15119	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF9D733B26E2FEE8FE03CA44.taxon	materials_examined	Type locality. Madagascar. Lectotype male (MHNG).	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF9D733B26E2FEE8FE03CA44.taxon	description	Redescription. Male: Body size large (Tab. 4). Fastigium of vertex (Figs. 3, 76) forming a long horn with 3 arms, upper 2 arms short and equal, lower one long, placed between the upper arms, slightly downcurved; its basal part 1.7 times narrower than diameter of eye and 1.1 times wider than antennal scape. Fastigium of frons (Fig. 77) with a simple, spiniform tooth. Face strongly flattened with distinct irregular tubercles. Pronotum (Figs. 76, 77) large, almost as long as wide; 3.8 times longer than high; anterior margin convex, posterior margin with a deep angular median incision; prozona 2.8 times longer than metazona. Tegmina (Figs. 76, 77) protruding from under pronotum, reaching 2 / 3 of the way down metanotum; its upper surface wavy. Prothoracic auditory spiracle (Fig. 79) almost rounded, its dorsal half concealed under lateral lobe of pronotum in lateral view. Mesothoracic auricle (Fig. 79) long, its inner gap oval without or with an indistinct ventral lobe. Prosternum with 2 long spines, distinctly V-shaped. Fore femur (Fig. 76) with a large inner spine on anterio-lateral margin; with a large inner and a large outer apical spine ventrally (in one male with an additional ventral spine on outer side apically); with 4 – 5 additional ventral spines on inner side. Fore tibia (Fig. 76) with 5 very large inner and 6 – 7 smaller outer spines ventrally. Mid femur with 2 – 4 and hind femur with 2 – 4 ventral spines on outer side. Hind tibia with 5 – 7 dorsal spines on inner side, with a single apical spur on inner side dorsally (Fig. 9); with 4 apical spurs ventrally, inner two shorter than outer ones. Last abdominal tergite (Fig. 80) broad, posterior part elongated backward, with a deep narrow median incision. Cercus (Fig. 81) long, narrow and slender, apical third sharply narrowed with a large, slightly incurved apical tooth. Subgenital plate (Fig. 82) almost as long as wide; its styli long, much longer than depth of posterior incision. Female: Head as in male, relatively slightly larger. Fastigium of vertex (Figs. 83, 84) as wide as half eye diameter, its basal part 1.1 times wider than antennal scape. Fastigium of frons (Fig. 84) with a simple, large, slightly downcurved tooth. Pronotum (Figs. 83, 84) short and broad, anterior margin convex, posterior margin broadly concave; prozona 1.4 times narrower and 3.9 times longer than metazona. Tegmina strongly reduced, like a very small scale. Dorsal half of prothoracic auditory spiracle covered by pronotum. Mesothoracic auricle not covered by pronotum. Fore femur (Fig. 83) with a large inner spine on anterio-lateral margin; with a large inner and a large outer apical spine ventrally; with 5 – 6 additional inner spines ventrally. Fore tibia (Fig. 83) with 5 inner and 6 – 7 outer strong spines ventrally. Mid femur with 3 and hind femur with 1 – 3 outer spines ventrally. Hind tibia with 6 – 8 dorsal spines on inner side, with a single inner apical spur dorsally; with 4 apical spurs ventrally, inner 2 slightly shorter than outer ones. Last tergite very short and broad, posterior margin with a narrow, median incision. Cercus conical, pointed at apex like a spine. Subgenital plate (Fig. 86) short and broad, with a wide median carina, its lateral lobes with transverse ridges. Ovipositor (Fig. 85) almost straight, 2 – 2.3 times longer than hind femur and 1.5 – 1.6 times longer than abdomen; apical part of ventral valve with small teeth.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF9D733B26E2FEE8FE03CA44.taxon	materials_examined	Colour (dry specimens). Body dark, black and dark brown. Face dark reddish brown; labrum light brown; mandibles and upper surface of head black. Pronotum black with some irregular dark brown spots. Male tegmina, mesonotum and metanotum in both sexes yellowish milky brown. Forelegs reddish brown and reddish black. Mid and hind tibia yellowish milky brown. Abdominal tergites black or dark reddish brown in some specimens. Sternites yellowish milky brown in male, darker in female. Ovipositor black.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF9D733B26E2FEE8FE03CA44.taxon	diagnosis	Diagnosis. The shape of fastigium of the vertex and the forelegs indicates that this species is closely related to C. redtenbacheri. It differs from it in the shapes of the pronotum, the male cercus and the ovipositor, in the ratios of ovipositor / hind femur and ovipositor / abdomen, in its distinctly darker body colour and also its larger size. It differs from C. parvicavus sp. nov. by its very larger body size, dark colouration, widely open prothoracic auditory spiracle, the shape of male pronotum, and the presence of ventral spines on the hind femur.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF9D733B26E2FEE8FE03CA44.taxon	materials_examined	Material examined. Madagascar, 3 females (leg. M. Grandidier) (Paralectotypes); Prov. De Tulear, Bas Fiherana [Fiherenana], 1906, 1 male, 2 females (leg. F. Geay); Madagascar (S. O.), Forest du Fiherana [Fiherenana], 1905, 1 male, 1 male nymph (leg. F. Geay); Madagascar sud, 14.5. 1963, 1 female nymph (leg. D. Wintrebert); Madagascar, sud-wat, Efeda, 29.4. 1963, 1 female nymph (leg. D. Wintrebert); Madagascar, Fianarantsoa, Anpamaherana [Ampamaherana], station forestire, 1 female; Madagascar, Tulear, Manombo, 10.5. 1956, 1 male (all in MNHN). Madagascar, S. Centr. Mdg., 2 males; Toliara Province, Forest near village of Mangily, c. 27 km N of Toliara (- 23.15, 43.60) [23 ° 8 ' 60 '' S, 43 ° 36 ' 0 '' E], 19.11. 2004, Coll. G. W. Beccaloni, 2 males, 1 female (all in NHMUK). 1 female (det. Karny), without locality label (NMW). Madagascar, Toliara Prov., Forét de Tsinjoriaky 6.2 km 84 E Tsifota, elev. 70 m, 6 – 10 Mar. 2002, 22 ° 48 ' 8 " S, 43 ° 25 ' 14 " E, 1 female (leg. Fisher, Griswold et al.), in spiny forest thicket; Madagascar, Toliara Prov., Lake Ranobe, elev. 20 m, 23 ° 02 ' 524 " S, 43 ° 37 ' 214 " E, 13 – 21 May 2003, 1 male, 1 female (leg. Frontier, Wilderness), spiny forest; Madagascar, Toliara Prov., Mahafaly Plateau, 6.2 km 74 ENE Itampolo, elev. 80 m, 21 – 25 Feb. 2002, 24 ° 39 ' 13 " S, 43 ° 59 ' 48 " E, 1 female (leg. Fisher, Griswold et al.), in spiny forest thicket (CAS). In addition, 2 adult females were photographed but not collected in spiny forest near Mangily, north of Toliara, Toliara Province by Martin Bader on 15.3.2015 (photographs and video footage were examined by the authors). They were found during the day hiding under bark attached to dead trees.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF9D733B26E2FEE8FE03CA44.taxon	description	Measurements (mm). Lectotype (male): Body length: 53; pronotum length: 14; width of pronotum: 12.5; fore femur length: 16.9; width of fore femur: 6.2; fore tibia length: 14; hind femur length: 18. See Table 4 for the range of measurement of the other specimens including female.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF9D733B26E2FEE8FE03CA44.taxon	distribution	Distribution. South-west and south-central Madagascar (Fig. 111). Toliara Province: Toliara, Fiherenana, Mangily, Efeda, Manombo, Tsifota, Lake Ranobe, NE Itampolo. Fianarantsoa Province: Anpamaherana [Ampamaherana]. Habitat. This species was collected in spiny forest. The specimens collected by the second author were found under loose bark still attached to dead trees.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF9D733B26E2FEE8FE03CA44.taxon	description	Phenology. This species has been collected or observed between February and May: in November and March as adults, and in April and May as nymphs.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF9D733B26E2FEE8FE03CA44.taxon	discussion	Remarks. The record for Anpamaherana [Ampamaherana] is dubious since this locality is located in the subhumid bioclimatic zone (Cornet 1974), rather than the subarid zone like all of the other collecting localities for this species (see remarks for O. janetae). Life history. C. grandidieri has been bred in captivity by several hobbyists in Europe and to our knowledge it is the only one of the species discussed in this paper that has been studied in this respect. What follows is a compilation of information from several people who have reared this species.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF9D733B26E2FEE8FE03CA44.taxon	description	First instar nymphs (Fig. 121) measure ca. 8 mm in body length (Tänzler 2012), are dark brown / black, and lack the turquoise markings on their hind- and midlegs which third instar and older nymphs and adults have. Third instar and older nymphs have pale markings on their pronota, legs and tergites (Figs. 122 – 123), whereas, in adults these markings are replaced by black (Figs. 124, 126). Nymphs take 4 – 5 months to reach adulthood and adults can live for about six months (Tänzler 2012). It is unclear how many nymphal instars there are. C. grandidieri is nocturnal and first and second instar nymphs hide singly in holes in dead bark during the day (Guillermo González, pers. commun., 2017). Older nymphs and adults will rest together during the day in groups, but smaller individuals may be eaten by larger ones (Tänzler 2012). In their natural habitat adults of several species, including C. grandidieri, have been observed to rest by day as male-female pairs, never in groups of more than two individuals (see Discussion), so the aggregations seen in captivity may be the result of a lack of harbourages. Adult males and females have very similar colouration, except that the labrum of the male is yellow-orange (Fig. 127), whilst that of the female is orange-red (Fig. 125). When disturbed adults and larger nymphs rear up, open their mandibles and raise and spread their forelegs (Fig. 125). Adult males stridulate loudly when in this defensive position (George Beccaloni, pers. obs.; Tänzler 2012; Guillermo González, pers. commun., 2017), whereas the adult females do not make any audible sound (Martin Bader, pers. commun., 2017; Guillermo González, pers. commun., 2017). If an aggressor approaches closely enough the insect will attempt to grab it with its forelegs and bite it. The threat display of only one other species, O. brongniarti, has been observed (Fig. 115). It was an adult female and the defensive display was the same as that of the female C. grandidieri (Martin Bader, pers. commun., 2017).	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF92733926E2FDA3FC15CB97.taxon	description	http: // lsid. speciesfile. org / urn: lsid: Orthoptera. speciesfile. org: TaxonName: 15117	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF92733926E2FDA3FC15CB97.taxon	materials_examined	Type locality. Madagascar. Lectotype male (in alcohol) (MNHN), here designated.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF92733926E2FDA3FC15CB97.taxon	description	Redescription. Body size is intermediate between the other two members of the genus (Table 4). Fastigium of vertex (Fig. 87, 88) with three-pronged horn; upper 2 arms short and equal, slightly upcurved; lower one longer and situated between the upper arms; 1.4 times narrower than diameter of eye; its basal part 1.1 times wider than antennal scape. Fastigium of frons (Fig. 88) with a large spiniform tooth. Face almost flattened with distinct tubercles some of which are like short spines. Pronotum (Fig. 87, 88) slightly longer than wide, 3.4 times longer than its height; anterior margin convex, posterior margin with a deep angular median incision; prozona 1.5 times longer than metazona. Tegmina (Fig. 87, 88) wavy, clearly visible from above, reaching 4 / 5 of the way down metanotum. Prothoracic auditory spiracle (Fig. 90) almost rounded, almost fully covered by lateral lobes of pronotum. Mesothoracic auricle (Fig. 90) oval, visible in lateral view, longer than wide, without ventral lobe; its inner gap long and elliptical. Prosternum with two, V-shaped spines, their apices not reaching to level of ventral margin of fore coxa. Fore coxa with a short dorsal spine. Fore femur (Figs. 6, 87) with a large inner spine on anterio-lateral margin; with a large inner and a large outer apical spine ventrally; with 5 – 6 additional ventral spines on inner side. Fore tibia (Fig. 87) with 5 inner and 4 – 7 outer spines ventrally, inner ones larger. Mid femur with 2 – 4 and hind femur with 1 – 3 outer spines ventrally. Hind tibia with 5 – 7 dorsal spines on inner side, with a single inner apical spur dorsally; with 4 apical spurs ventrally, inner two slightly shorter than outer ones. Last abdominal tergite (Fig. 91) twice as wide as its length, posterior margin with a deep narrow incision. Cercus (Fig. 92) short, basal part broad, end of basal part with a distinct projection on inner margin; apical part narrower and slightly incurved, apex blunt, slightly rounded, with a small apical tooth. Subgenital plate (Fig. 93) almost as long as its width, its styli slightly longer than depth of posterior incision of plate. Female: Head as in male. Fastigium of vertex (Figs. 94, 95) 1.4 times narrower than diameter of eye, 1.3 times wider than antennal scape. Pronotum (Figs. 94, 95) with long, narrow prozona and wide, short metazona; prozona 1.4 times narrower and 4.3 times longer than metazona; anterior margin convex, posterior margin strongly concave. Tegmina strongly reduced to very small scale-like projections. Prothoracic auditory spiracle almost concealed by lateral lobe of pronotum. Mesothoracic auricle smaller than that of male, its inner gap narrow elliptical, not covered by pronotum. Fore tibia (Fig. 94) with 5 – 6 ventral spines on outer side; mid femur with 1 – 4 and hind femur with 1 – 4 outer spines ventrally; hind tibia with 6 – 7 dorsal spines on inner side, with an inner apical spur dorsally and 4 apical spurs ventrally. Last abdominal tergite broad and narrow, posterior margin with a small distinct median incision. Cercus short, conical, its apical part slightly incurved, pointed at apex. Subgenital plate (Fig. 97) 3.5 times broader than long, with a broad rectangular median carina; lateral lobes with many transverse ridges. Ovipositor (Fig. 96) straight, 1.5 – 1.6 times longer than hind femur and slightly longer than abdomen; apical part of ventral valve with small teeth.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF92733926E2FDA3FC15CB97.taxon	materials_examined	Colour (dry specimens). Body with different shades of brown, mostly milky and light brown. Face darker, reddish brown; clypeus and labrum yellowish brown. Metazona of pronotum darker with reddish brown, prozona light brown. Fore tibia, anterior part of fore femur, knees of mid and hind legs and tarsi reddish brown, remaining parts of legs milky brown. Male tegmina yellowish brown. Abdominal tergites reddish brown, with a very narrow black stripe on hind margin in female. Basal part of ovipositor milky brown, its apical part light reddish brown.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF92733926E2FDA3FC15CB97.taxon	diagnosis	Diagnosis. This species differs from C. grandidieri in the structure of the pronotum, male cercus, shorter ovipositor, the ratios of ovipositor / hind femur and ovipositor / abdomen, clearly lighter body colour and smaller size. It differs from C. parvicavus sp. nov. by the shape of the male pronotum, prothoracic auditory spiracle (which has a wide opening), the presence of ventral spines on the hind femur, and the ratios of ovipositor / hind femur and ovipositor / abdomen.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF92733926E2FDA3FC15CB97.taxon	materials_examined	Material examined. Madagascar, 1867, 1 male (Lectotype), 1 female (Paralectotype) (leg. M. Grandidier) (MNHN). Toliara Province, Andohahela National Park: Tsimelahy parcel, (- 24.93, 46.65) [24 ° 55 ' 48 '' S, 46 ° 38 ' 60 '' E], 1.12.2004, Coll. leg. G. W. Beccaloni, 1 male, 1 female (NHMUK). Madagascar, Toliara Prov., Parc Nat. d’Andohahela, Forét d’Ambohibory, 1.7 km 61 ENE Tsimelahy, 36.1 km 308 NW Tolagnaro, 16 – 20.1.2002, 24 ° 55 ' 48 " S, 46 ° 38 ' 44 " E, 2 males, 1 female (Fisher, Griswold et al.), tropical dry forest; Castlot 0 0 6432 [locality unknown], 3 males, 3 females, 1 male nymph (CAS).	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF92733926E2FDA3FC15CB97.taxon	description	Measurements (mm). Lectotype (male): Body length: 40; pronotum length: 10.6; width of pronotum: 10; fore femur length: 12.9; width of fore femur: 4.6; fore tibia length: 10.3; hind femur length: 14.4. See Table 4 for the range of measurement of the other specimens including females.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF92733926E2FDA3FC15CB97.taxon	distribution	Distribution. South-eastern Madagascar (Fig. 111). Toliara Province: Toliara, Andohahela. Habitat. This species has been found in transitional tropical dry forest. The specimens collected by the second author were found under loose dead bark.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF92733926E2FDA3FC15CB97.taxon	description	Phenology. Adults have been found in December and January.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF90733F26E2FBFEFD43CD26.taxon	description	http: // lsid. speciesfile. org / urn: lsid: Orthoptera. speciesfile. org: TaxonName: 499692 Type locality. Madagascar, Toliara Prov., W. Marovato. Holotype male (CAS). Description. Male (Holotype): Body size smallest in the genus (Tab. 4). Fastigium of vertex (Figs. 98, 99) with three-pronged horn; above 2 arms equal and shorter, directed slightly upwards; lower one placed between the above arms, longer and slightly directed downwards; 1.5 times narrower than diameter of eye; its basal part as wide as antennal scape. Fastigium of frons (Fig. 99) with a large simple spiniform tooth. Face slightly convex, with tubercles some of which look like short spiniform teeth. Pronotum (Figs. 98, 99) slightly longer than wide, 3 times longer than its height; anterior margin convex, posterior margin with a shallow rounded median incision; prozona 2.3 times longer than metazona. Tegmina (Figs. 98, 99) wavy, clearly protruding from under pronotum, reaching beyond middle half of metanotum (in other male reaching to end of metanotum). Prothoracic auditory spiracle (Fig. 101) almost closed except for a small rounded hole which is visible in lateral view. Mesothoracic auricle (Fig. 101) visible in lateral view, its inner gap long, oval, with a long ventral lobe. Prosternum with two long, V-shaped spines. Fore coxa with a short dorsal spine. Fore femur (Fig. 98) with a large inner spine on anterio-lateral margin; with a large inner and a large outer apical spine ventrally; with 5 additional ventral spines on inner side (in the other male, inner side of fore femur with 6 ventral spines). Fore tibia (Fig. 98) with 5 inner and 6 smaller outer spines ventrally (in other male outer side of fore tibia with 4 – 5 ventral spines). Mid femur with 2 ventral spines on outer side (in other male with 1 spine); hind femur without ventral spines. Hind tibia with 6 – 8 dorsal spines on inner side; with a single apical spur on inner side dorsally; with 4 apical spurs ventrally, inner 2 shorter than outer ones. Last abdominal tergite (Fig. 102) large, elongated backwards, with a small and narrow median incision. Cercus (Fig. 103) small, almost 1.5 mm, basal part broad, end of basal part with a small tooth on inner side; apical part narrow, slightly incurved (in other male apical part more incurved); with a very distinct incurved apical tooth. Subgenital plate (Fig. 104) slightly longer than wide, its styli longer than depth of posterior incision of plate. Female: Head as in male, including the ratios. Pronotum (Figs. 105, 106) slightly longer than wide (in other female slightly wider than long); prozona 4.6 times longer than metazona; anterior margin convex, posterior margin concave. No tegmina. Prothoracic auditory spiracle almost closed except for a small rounded hole. Mesothoracic auricle as in male, not covered by pronotum; its ventral lobe smaller than that of female. Fore femur (Fig. 105) with a large inner spine on anterio-lateral margin; with a large inner and a large outer apical spine ventrally; with 4 – 5 additional ventral spines on inner side. Fore tibia (Fig. 105) with 5 inner and 5 outer spines ventrally; mid and hind legs as in male. Last abdominal tergite very broad and narrow, posterior margin with a distinct median incision which separates the posterior margin into two rounded lobes. Cercus narrow, longer than last abdominal tergite; its apex slightly incurved. Subgenital plate (Fig. 108) short and broad, hind margin narrowed and elongated backwards; with a large, almost rectangular median carina, lateral parts of plate with distinct transverse ridges. Ovipositor (Fig. 107) long and straight, slightly more than twice as long as hind femur and 1.4 – 1.5 times longer than abdomen; apical part of ventral valve with small teeth.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF90733F26E2FBFEFD43CD26.taxon	materials_examined	Colour (dry specimens). Body milky and light brown with dark brown transverse bands. Face light brown, its tubercles with black or dark brown tips. Legs milky brown, their spines with black tips; Pronotum light brown hind margin with a large dark brown transverse band. Male tegmina milky brown. Hind margin of each abdominal tergite with a thin black transverse band. Basal third of ovipositor milky brown, apical two thirds dark brown.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF90733F26E2FBFEFD43CD26.taxon	diagnosis	Diagnosis. This species is similar to C. redtenbacheri in its colouration, male cercus and subgenital plate. It differs from it in that its prothoracic auditory spiracle opening is reduced to a small hole, that the male pronotum has a shallow posterior incision, in the absence of ventral spines on the hind femur, in the ratios of ovipositor / hind femur and ovipositor / abdomen, and in its smaller size. The ratios of ovipositor / hind femur and ovipositor / abdomen are similar to C. grandidieri, but it differs from C. grandidieri by its much smaller size, its colouration (milky brown in the new species, distinctly darker in C. grandidieri), its prothoracic auditory spiracle (with a small hole in the new species, and a large opening in C. grandidieri), in the absence of ventral spines on the hind femur, and that the male pronotum has a shallow posterior incision.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF90733F26E2FBFEFD43CD26.taxon	materials_examined	Material examined. Madagascar, Toliara Prov., Réserve Spéciale de cap Sainte Marie, 12.3 km 262 ° W Marovato, elev. 200 m, 11 – 15 Feb. 2002, 25 ° 34 ' 54 " S, 45 ° 10 ' 6 " E, 2 males (including Holotype), 2 females (leg. Fisher, Griswold et al.), in spiny forest thicket (CAS).	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF90733F26E2FBFEFD43CD26.taxon	description	Measurements (mm). Holotype (male): Body length: 34.2; pronotum length: 9.4; width of pronotum: 8.7; fore femur length: 10.4; width of fore femur: 4.3; fore tibia length: 9.2; hind femur length: 11.4. See Table 4 for the range of measurements of the other type specimens.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF90733F26E2FBFEFD43CD26.taxon	etymology	Etymology. The species name is derived from the Latin words “ parvus ” and “ cavum ” meaning small and hole respectively. This refers to the structure of the prothoracic auditory spiracle which has a very small opening	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF90733F26E2FBFEFD43CD26.taxon	distribution	Distribution. Only known from the southern tip of Madagascar (Fig. 111). Toliara Province: Marovato. Habitat. This species was found in spiny forest.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
533F87C4FF90733F26E2FBFEFD43CD26.taxon	description	Phenology. Adults were collected in February.	en	Ünal, Mustafa, Beccaloni, George W. (2017): Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov. Zootaxa 4341 (2), DOI: 10.11646/zootaxa.4341.2.2
