identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
C051E46E5BD1472231BDB53EAAADA392.text	C051E46E5BD1472231BDB53EAAADA392.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Morphogenia	<div><p>Morphogenia gen. n.</p><p>Type species.</p><p>Morphogenia struhli, here designated.</p><p>Diagnosis.</p><p>Morphogenia, and its only species, Morphogenia struhli, can be distinguished from all other known jubine genera by the following combination of characters: (1) smoothly convex vertex lacking sulci, and afoveate due to anterolateral shift of vertexal foveae into vicinity of postantennal notches anterior to eyes; (2) absence of V- or Y-shaped gular carina; (3) pronotum with margins smooth, lacking lateral spines, and with a simple, well-defined transverse antebasal sulcus in both sexes; (4) abdomen lacking fovea-like cuticular pockets at bases of tergites V–VII and sternites V–VII (but tergite IV and sternite IV with true mediobasal foveae present).</p><p>Description.</p><p>Body length ~3 mm (Fig. 1). Form relatively flattened and broadened posteriorly, with compact abdomen and elongate legs.</p><p>Head: Approximately triangular (Figs 1, 18), 1.3 × wider than long; without distinct frontal rostrum and lacking prominent, raised antennal tubercles. Vertex smoothly convex, devoid of foveae or sulci, lateral margins incised behind antennal sockets by postantennal notches (region enclosed by dashed line in Fig. 2). Antennae separated by 1/3 maximum head width. Foveae of apparent homology to the vertexal foveae of other Pselaphinae shifted from vertex, situated instead on frontolateral margins, recessed into the top of postantennal notches (Figs 2, 5). Apodemes of tentorium extending from these foveae (Fig. 5) and converging on gular foveae with single opening (Fig. 3). Lateral margins of head smoothly rounded dorsoventrally, lacking ocular mandibular carina. Venter lacking any trace of gular carina (Fig. 3; a medial sutural line can be weakly detected). Antennae (Fig. 4) with 11 antennomeres, with club formed by enlarged antennomeres VIII–XI . Maxillary palpi comprised of five palpomeres, with small triangular palpomeres III and fusiform palpomeres IV (Fig. 3). Maxillary cardos projecting anteriorly, reaching slightly beyond sides of mandibles, with single long setae positioned at apex.</p><p>Thorax: Pronotum (Fig. 6) moderately transverse, 1.4 × wider than head and similar in length, obcordate in shape, approximately semicircular before abrupt constriction in basal quarter. Lateral margins before constriction smoothly rounded, without spines or teeth. Pronotal disc simple and convex, lacking foveae or sulci. Typical, deep and well-impressed antebasal sulcus present, demarcating point of pronotal constriction. Lateral antebasal foveae present, median antebasal fovea absent. Prosternum with lateral procoxal foveae. Mesoventrite with single unpaired median mesoventral fovea, lateral mesoventral foveae and lateral mesocoxal foveae. Metaventrite with lateral metaventral foveae, with median carina from 1/3 segment length to posterior margin.</p><p>Abdomen: Abdomen 2/3 length of elytra (measured along suture). Five tergites ( IV–VIII) evident. Tergites with broad, angularly-projecting paratergites on segments III–VI and smaller paratergites on VII (Fig. 7). Tergite IV longest, 2.7 × tergite V length, with posterior tergites becoming shorter and narrower. Tergite IV with mediobasal foveae present in basal sulcus. Six sternites ( III–VIII) evident; penial plate apparently internalized, not externally visible, with genital aperture formed by contiguous apical margins of tergite and sternite VIII (Figs 11, 12). Apical margin of sternite III entire, uninterrupted by metacoxae. Sternite IV longest, mediobasal foveae present. All tergites and sternites lacking fovea-like cuticular pockets at their bases (Fig. 7 shows bases of tergites V–VII revealed in cleared specimen).</p><p>Elytra: 1.3 × longer than pronotum, broadening gradually until narrowing just before apices. With sinuate transverse basal carina; sutural foveae and single median basal foveae present but largely obscured by arcing of basal carina. Humeri indented by impressed bases of humeral sulci; humeral sulci extending length of elytra, with humeral foveae at base. Sutural striae entire.</p><p>Legs: All pairs of coxae contiguous. Coxae all carinate along length of external face. Procoxal length greater than half femoral length, procoxae strongly conically projecting ventrally. Mesocoxae shorter than procoxae, moderately conically projecting, orientated somewhat posteriorly. Metacoxae transverse-conical, spanning from ventral midline to metaventral margin, and projecting posteriorly. All trochanters short, with negligible separation of coxal apex and femoral base ( “brachysceline” type). Femora and tibiae simple, lacking modifications. Profemora somewhat thickened. Tarsi (Figs 8, 9) 3-segmented with short tarsomeres I; tarsomeres II longest. Metatarsi especially elongate. Tarsi with two claws of equal size.</p><p>Etymology.</p><p>Morphogens are gradient-forming molecules that specify positional information and govern tissue growth during animal development. The generic name acknowledges the pervasive role of morphogens in sculpting organismal morphology. The gender is feminine.</p></div>	https://treatment.plazi.org/id/C051E46E5BD1472231BDB53EAAADA392	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Parker, Joseph	Parker, Joseph (2014): Morphogenia: a new genus of the Neotropical tribe Jubini (Coleoptera, Staphylinidae, Pselaphinae) from the Brazilian Amazon. ZooKeys 373: 57-66, DOI: http://dx.doi.org/10.3897/zookeys.373.6788, URL: http://dx.doi.org/10.3897/zookeys.373.6788
800B365C76A9CDC369AFA47B9D07F738.text	800B365C76A9CDC369AFA47B9D07F738.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Morphogenia struhli	<div><p>Morphogenia struhli sp. n.</p><p>Type material.</p><p>Holotype ♂: "88 1 // Leaf litter, Winkler method. Terra firmé fst. // BRAZIL: Manaus, A.M. INPA/Smithsonian Res. 2°25'S, 59°50'W, R. Didham, i. 1994 // BMNH{E} 2003-84.". Paratypes (2 ♀♀): "89 7 // Leaf litter, Winkler method. Terra firmé fst. // BRAZIL: Manaus, A.M. INPA/Smithsonian Res. 2°25'S, 59°50'W, R. Didham, i. 1994//BMNH{E} 2003-84 // (pink label) 0749". "68 12 // Leaf litter, Winkler method. Terra firmé fst. // BRAZIL: Manaus, A.M. INPA/Smithsonian Res. 2°25'S, 59°50'W, R. Didham, i. 1994 // BMNH{E} 2003-84". All material is deposited in the Coleoptera collection of the Department of Entomology, Natural History Museum, London.</p><p>Description.</p><p>Body length 2.9 mm (Fig. 1). Holotype male somewhat teneral; body colour of female paratypes dark reddish-purple (e.g. Figs 17, 19) with appendages lighter in colour. Dorsal regions shiny, with shallow punctures and sparse setae in most areas.</p><p>Head: Length 0.71 mm from occipital constriction to clypeus; width across widest point (posterior to eyes) 0.55 mm. Margins broadly and smoothly rounded from eyes to base (Figs 1, 18). Margins narrowing anterior to eyes before broadening slightly to clypeus. Setae sparse on vertex and becoming longer and denser towards clypeus. Anterolaterally-shifted vertexal fovea concealed in postantennal notches (region indicated in Fig. 2) by several long, apically-directed setae (Fig. 5). Eyes (Fig. 2) large with ~105 ommatidia, broadly crescent-shaped with shallow ocular canthi at posterior margin. Antennae received by triangular frontolateral excavations with carinate edges (Fig. 2). Antennal (Fig. 4) length 1.18 mm, with scape weakly transverse in dorsal view, pedicel slightly narrower and subquadrate. Antennomeres III–V subequal in width, slightly narrower and shorter than pedical, with segments becoming progressively longer apically; VI and VII transverse and subequal in length; VII wider than VI and approaching width of segment VIII. Antennomeres VIII–X with carinate apical and basal margins, roughly obconical and equal in width. Antennomere VIII slightly longer than preceding three segments, 2/3 longer than wide; antennomeres IX and X 0.7 × length of VIII; XI longest, 1.8 × length of VIII, with carinate basal margin, widest in apical half before tapering to apex. Antennomeres densely covered with apically- directed setae. Apical pseudosegment of maxillary palpus continuous with external face of palpomere IV and pointing slightly mesially.</p><p>Thorax: Pronotum (Fig. 6) length 0.68 mm, width 0.77 mm at widest point. Disc region sparsely setiferous. Lateral margins behind antebasal sulcus with several long, laterally-projecting setae. Prosternum with medial region in front of procoxae with long setae covering setiferous lateral procoxal fovea. Episternal areas with moderately dense, shorter setae orientated somewhat dorsally. Short lateral prosternal carinae extending briefly from sides of procoxal cavities. Mesoventral plate moderately setiferous, meso-epiventral regions largely glabrous. Metasternum with sparse, dorsoapically-pointing aciculate setae.</p><p>Abdomen: Dorsal abdominal length (with segments contracted) 0.57 mm. Tergite lengths: IV= 0.32 mm, V = 0.12 mm, VI = 0.10 mm. Apical margin of sternite IV with male-specific medially positioned small, round tubercle covered in small setae. Male tergite VIII (Fig. 11) with basal margin of sclerite weakly convex; with uniform moderately dense distribution of long setae. Sternite VIII (Fig. 12) with basal margin of sclerite strongly convex; mediobasal region with sparse, short setae becoming much denser towards apical margin; lateral areas of sternite with much longer setae of moderate density. Apical margin of sternite VIII medially depressed to receive corresponding apex of tergite VIII.</p><p>Elytra and flight wings: Elytra with scattered, short setae. Elytral length along suture 0.87 mm; width at widest point 0.5 mm. Full flight wings present.</p><p>Legs: Femora brownish-red, slightly lighter than body; tibiae and tarsi lighter than femora, yellowish (Fig. 1). Tarsi of increasing relative length: protarsi 0.5 × protibial length; mesotarsi 0.6 × protibial length; metatarsi 0.7 × metatibial length. Protarsomeres II 1.7 × tarsomere III length (Fig. 8). Mesotarsomeres II 1.8 × tarsomere III length. Metatarsomeres II especially long (Fig. 9), 2.3 × tarsomere III length. All tarsal claws with inner spine present on ventral face (Fig. 10). Pro- and mesotarsi with additional spines on ventral side of tarsomeres II and III (Fig. 8).</p><p>Aedeagus: Length 0.43 mm, width 0.27 mm at widest point (Fig. 13). Asymmetric, dorsoventrally relatively flattened, with large spherical and very weakly sclerotized basal bulb. Lacking obvious remnants of parameres. Complex, more heavily sclerotized medioapical piece with “hooked” apophysis extending apically from right side of basal bulb.</p><p>Sexual dimorphism.</p><p>Female dimensions similar to male. Males with large crescent shaped eyes (Fig. 16); female eyes very small and approximately oval, consisting of ~12 ommatidia (Fig. 17). Vertex of female head (Fig. 19) broader at base than that of male (Fig. 18). Antennal club ( VIII–XI) twice as long as preceding segments combined in male (Fig. 18), only 1/3 as long as preceding segments combined in female (Fig. 19). Medial sexual patch on sternite IV absent in female, and flight wings lacking. Female tergite VIII (Fig. 14) with basal margin of sclerites shallowly concave; with moderate density of long setae and two pairs of much longer setae. Female sternite VIII (Fig. 15) with basal margin of sclerite weakly convex; covered in moderate density of short setae and several pairs of much longer setae. Apex with internally-projecting cuticular protuberance.</p><p>Etymology .</p><p>The type species of Morphogenia is named in honour of Dr. Gary Struhl, developmental biologist, whose Drosophila studies have yielded great insights into the genetic control of animal development.</p><p>Biology.</p><p>Morphogenia was collected from rainforest litter-the typical habitat of jubines. Like other pselaphines, jubines are most likely predators of soil microarthropods. The spines on the ventral face of the pro- and mesotarsi of Morphogenia struhli (Fig. 8) may serve a raptorial function. Prior to photography, a filament protruding from the mouthparts of one female paratype was removed. A subsequent survey of jubine specimens in a range of genera have revealed similar filaments, which appear to be dried, threadlike residues of glutinous secretions from the enlarged maxillary cardos that are characteristic of the tribe. Such structures suggest the evolution of a novel mode of feeding or prey capture in Jubini (J. Parker &amp; C. Carlton unpublished observations).</p></div>	https://treatment.plazi.org/id/800B365C76A9CDC369AFA47B9D07F738	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Parker, Joseph	Parker, Joseph (2014): Morphogenia: a new genus of the Neotropical tribe Jubini (Coleoptera, Staphylinidae, Pselaphinae) from the Brazilian Amazon. ZooKeys 373: 57-66, DOI: http://dx.doi.org/10.3897/zookeys.373.6788, URL: http://dx.doi.org/10.3897/zookeys.373.6788
