taxonID	type	description	language	source
560A8789FF84FFDFFC02FAD3FDA8FC7F.taxon	type_taxon	Type genus. Ochlesis Stebbing, 1910.	en	Ariyama, Hiroyuki (2024): Curidia japonica sp. nov., the First Species of the Family Ochlesidae from the Northwest Pacific (Crustacea: Amphipoda). Species Diversity 29 (2): 199-207, DOI: 10.12782/specdiv.29.199, URL: http://dx.doi.org/10.12782/specdiv.29.199
560A8789FF84FFDFFC02FAD3FDA8FC7F.taxon	diagnosis	Diagnosis. Body compressed laterally. Head short, tall, partially enveloped by pereonite 1; rostrum well developed. Pereonites raised into thin keels or formed into dorsal plaques. Urosomites free. Antennae poorly developed, short; accessory flagellum absent, both flagella reduced, with ≤ 5 articles. Mouthparts grouped conically. Upper lip narrow, tip not incised. Mandibles with lacinia mobilis in left, accessory blades absent, molar small; palp 3 - articulated, article 1 elongate. Lower lip without inner lobes, outer lobes thin. Maxilla 1 with small inner plate; outer plate subconi- © 2024 The Japanese Society of Systematic Zoology. This is an open access article distributed under a Creative Commons Attribution 4.0 License (CC BY, https: // creativecommons. org / licenses / by / 4.0 /), which permits unrestricted use, distribution, and cal, with mostly-fused robust setae; palp tiny, 1 - articulated. Maxilla 2 elongate. Maxilliped with very thin plates, palp absent or 1 – 2 - articulated. Coxae 1 – 3 narrow, coxae 1 and 4 shorter than coxae 2 – 3. Gnathopods feeble, simple; gnathopod 1 ischium, carpus and propodus elongate, dactylus very short, usually with several feather setae; gnathopod 2 carpus and propodus elongate, merus and carpus produced posterodistally, dactylus very short, usually with robust seta. Pereopods 3 – 7 stout, bases of pereopods 5 – 7 expanded posteriorly or posterodistally. Uropod 1 biramous, outer ramus subequal to or slightly shorter than inner ramus; uropod 2 biramous, outer ramus shorter than inner ramus; uropod 3 biramous or uniramous, in biramous uropod 3, outer ramus usually shorter than inner ramus. Telson entire, tip acute or rounded.	en	Ariyama, Hiroyuki (2024): Curidia japonica sp. nov., the First Species of the Family Ochlesidae from the Northwest Pacific (Crustacea: Amphipoda). Species Diversity 29 (2): 199-207, DOI: 10.12782/specdiv.29.199, URL: http://dx.doi.org/10.12782/specdiv.29.199
560A8789FF84FFDFFC02FAD3FDA8FC7F.taxon	discussion	Remarks. On the shape of the gnathopod 2, J. L. Barnard and Karaman (1991) stated “ ... simple, otherwise ... carpo- or merochelate. ” However, it should be unified to “ simple ”, because the shape is quite different from the true “ carpochelate ” (ex. Grandidierella Coutière, 1904) and “ merochelate ” (ex. Aora KrØyer, 1845 and Aoroides Walker, 1898) (see Ariyama and Kohtsuka 2022).	en	Ariyama, Hiroyuki (2024): Curidia japonica sp. nov., the First Species of the Family Ochlesidae from the Northwest Pacific (Crustacea: Amphipoda). Species Diversity 29 (2): 199-207, DOI: 10.12782/specdiv.29.199, URL: http://dx.doi.org/10.12782/specdiv.29.199
560A8789FF85FFDFFE83FAB1FAEAFDF6.taxon	type_taxon	Type species. Curidia debrogania Thomas, 1983, by original designation.	en	Ariyama, Hiroyuki (2024): Curidia japonica sp. nov., the First Species of the Family Ochlesidae from the Northwest Pacific (Crustacea: Amphipoda). Species Diversity 29 (2): 199-207, DOI: 10.12782/specdiv.29.199, URL: http://dx.doi.org/10.12782/specdiv.29.199
560A8789FF85FFDFFE83FAB1FAEAFDF6.taxon	diagnosis	Diagnosis. Lateral cephalic lobe acutely projected anteroventrally. Pereonites raised into thin keels. Dorsal margins of pereonite 7 and pleonites 1 – 2 usually projected posteriorly. Dorsal margin of pleonite 3 with median hump. Posterolateral margin of pleonite 3 bearing pointed tooth. Antenna 1, peduncular articles 1 – 3 progressively shortened, articles 1 and 2 each with acute process posterodistally, flagellar article 1 elongate; antenna 2, peduncular article 4 bearing posterodistal process. Mandibular palp long, visible from outside. Maxilliped palp 1 - articulated, with apical seta. Included taxa. Nine species: C. andreae Coleman and Heinz, 2010; C. debrogania; C. japonica sp. nov.; C. knoxi Lowry and Myers, 2003; C. magellanica Coleman and J. L. Barnard, 1991 b; C. monicae Ortiz, Lalana, and Varela, 2007; C. nunoi Winfield and Ortiz, 2013; C. ramonae Lowry and Myers, 2003; and C. wakabarae Souza-Filho and Serejo, 2008.	en	Ariyama, Hiroyuki (2024): Curidia japonica sp. nov., the First Species of the Family Ochlesidae from the Northwest Pacific (Crustacea: Amphipoda). Species Diversity 29 (2): 199-207, DOI: 10.12782/specdiv.29.199, URL: http://dx.doi.org/10.12782/specdiv.29.199
560A8789FF85FFDFFE83FAB1FAEAFDF6.taxon	discussion	Remarks. Curidia monicae may be a junior synonym of C. debrogania because of small morphological differences (see the below key) and closeness of the localities (both distributed in the Caribbean Sea and / or the Gulf of Mexico).	en	Ariyama, Hiroyuki (2024): Curidia japonica sp. nov., the First Species of the Family Ochlesidae from the Northwest Pacific (Crustacea: Amphipoda). Species Diversity 29 (2): 199-207, DOI: 10.12782/specdiv.29.199, URL: http://dx.doi.org/10.12782/specdiv.29.199
560A8789FF85FFD8FC5EF849FAECF7E8.taxon	materials_examined	Material examined. Holotype: female, 1.4 mm (OMNH- Ar- 12915), southwestern coast of Kamishima Island, Toba City, Mie Prefecture, 34 ° 32 ′ 41 ″ N, 136 ° 58 ′ 47 ″ E, 2 m depth, among red algae Corallina pilulifera Postels and Ruprecht and Amphiroa beauvoisii J. V. Lamouroux, 25 July 2014. Paratypes: male, 1.1 mm (OMNH-Ar- 12916), same locality, 2 m depth, among a bryozoan Bugula neritina (Linnaeus, 1758), same date; female, 1.5 mm (OMNH-Ar- 12917, damaged), Oura in Hidaka, Wakayama Prefecture, 33 ° 55.5 ′ N, 135 ° 04 ′ E, 1.5 m depth, among a red alga Corallina aberrans (Yendo) K. R. Hind and G. W. Saunders, 31 July 2016; male, 1.2 mm (OMNH-Ar- 12918), same locality, 2 m depth, among a red alga Gelidium elegans Kützing, same date; male, 1.1 mm (OMNH-Ar- 12919), same locality, 4.5 m depth, among a red alga G. elegans, same date; male, 1.3 mm (OMNH-Ar- 12920), Hikigawa in Shirahama, Wakayama Prefecture, 33 ° 33 ′ N, 135 ° 27 ′ E, 7 m depth, among a red alga G. elegans, 2 August 2003; male, 1.0 mm (OMNH-Ar- 12921), same locality, 3 m depth, among a red alga G. elegans, same date. Type locality. Kamishima Island, Toba City, Mie Prefecture, Japan.	en	Ariyama, Hiroyuki (2024): Curidia japonica sp. nov., the First Species of the Family Ochlesidae from the Northwest Pacific (Crustacea: Amphipoda). Species Diversity 29 (2): 199-207, DOI: 10.12782/specdiv.29.199, URL: http://dx.doi.org/10.12782/specdiv.29.199
560A8789FF85FFD8FC5EF849FAECF7E8.taxon	description	Description. Based on holotype, female, 1.4 mm (OMNH-Ar- 12915); paratype, male 1.2 mm (OMNH-Ar- 12918), for upper lip, maxilla 1 and maxilliped; and paratype, male, 1.3 mm (OMNH-Ar- 12920) for mandibles. Head (Figs 1, 2). Rostrum strongly projected ventrally. Eyes oval, partially covered with pereonite 1. Antenna 1 stout; length ratio of peduncular articles 1 – 3 (excluding processes) 1: 0.65: 0.2, lengths of posterodistal processes on articles 1 and 2 ca. 0.4 and 0.6 times of article proper, respectively; flagellum with 3 articles, article 1 long, article 3 minute, posterodistal part of article 1 and distal ends of articles 2, 3 bearing many aesthetascs. Antenna 2 also stout, but shorter than antenna 1, with ratio of lengths of peduncular articles 3 – 5 1: 1.55: 1.7, posterodistal process of peduncular article 4 very short, 15 % length of article 5; flagellum with 3 articles, article 1 long, article 3 minute, distal end of each article setose. Upper lip triangular, tip pointed. Mandibles slender, incisors and left lacinia mobilis without denticles; length ratio of palp articles 1 – 3 1: 1.3: 1.55, article 3 with numerous fine setae on lateral surface and 3 thick setae on tip. Lower lip, outer lobes with narrow mandibular process. Maxilla 1, inner plate narrowed distally, tip without setae; outer plate bearing 5 mostly-fused robust setae, tip pleated; palp minute, with a long seta. Maxilla 2, outer and inner plates with 10 and 6 distal setae, respectively. Maxilliped with narrow inner plate bearing apical short robust seta and several distal and many dorsodistal setae; outer plate broad, with 2 median and 6 subdistal setae; palp narrow, apical seta very long, ca. 1.7 – 1.8 times length of palp proper. Pereon (Figs 1, 3). Dorsal margins smooth, pereonite 1 strongly projected anterodorsally; pereonite 7 posterodorsal margin not projected, posteroventral corner bluntly projected. Coxae of gnathopods and pereopods, and bases and ischia of pereopods 5 – 7 double-structured. Gnathopod 1 slender; coxa small, curved anteriorly, anteroproximal corner swollen, ventral margin rounded; basis proximal half widened; ischium ca. 0.65 times length of basis; carpus and propodus almost same length, posterodistal margin of propodus with minute projection and a feather seta and 3 slender setae; dactylus very short, with 3 feather setae and 2 slender and 1 robust setae. Gnathopod 2 shorter than gnathopod 1; coxa elongate, ca. 1.65 times (estimated) length of coxa 1, anteroproximal corner weakly swollen; basis wide, lateral surface bearing many minute setae; ischium short; carpus posterodistal corner produced, tip with 3 setae; propodus narrow, shorter than carpus, bearing several setae distally; dactylus triangular, ca. 0.2 times as long as propodus, tip with robust seta forming chela. Pereopod 3 far larg- er than gnathopod 2, coxa elongate, ca. 1.05 times (estimat- ed) length of coxa 2, anteroproximal corner swollen; basis wide, posterolateral surface bearing many minute setae; merus wide, projected anterodistally; carpus with 1 + 2 robust setae posteriorly; propodus ca. 1.35 times length of carpus, bearing 1 + 1 + 2 + 2 posterior robust setae; dactylus with unguis. Pereopod 4 slightly smaller than pereopod 3; coxa short, ca. 0.6 times (estimated) length of coxa 3, anteroproximal corner inflated anteriorly and anterodistal corner strongly swollen distally, posteroproximal corner weakly inflated; basis – dactylus almost same as those of pereopod 3, but carpus with 1 + 1 + 2 posterior robust setae. Pereopod 5 almost same length as pereopod 4; coxa short, anterodistal corner swollen anteriorly and posterodistal corner inflated distally; basis expanded posteriorly, with 2 rounded posteroproximal projections and large posterodistal lobe; merus wide, posterodistal corner projected; carpus with 1 + 2 robust setae anteriorly; propodus ca. 1.4 times length of carpus, bearing 1 + 1 + 2 + 2 anterior robust setae; dactylus with unguis. Pereopod 6 slightly longer than pereopod 5; coxa narrow and short, posterodistal corner swollen; basis expanded posteriorly, with 1 triangular and 1 rounded posteroproximal projections and large posterodistal lobe; ischium – dactylus almost same as those of pereopod 5, but carpus and propodus with 1 + 1 + 2 and 1 + 1 + 1 + 2 + 2 anterior robust setae, respectively. Pereopod 7 a little longer than pereopod 6; coxa narrow and short, posteroproximal corner swollen; basis expanded posteriorly, with angular posteroproximal projection and large posterodistal lobe; ischium – dactylus almost same as those of pereopod 6, but propodus bearing 1 + 2 + 1 + 1 + 2 anterior robust setae. Pleon (Figs 1, 4). Dorsal margins with posterior hump in pleonites 1 and 2, and median hump in pleonite 3; posteroventral corners of pleonites 1 and 2 with acute and triangular projections, respectively, posteroventral corner of pleonite 3 without projection, posterolateral margins bearing rounded projection each on pleonites 1, 2 and upward pointed tooth on pleonite 3, respectively. Pleopods gradually shortened; peduncles each with 2 coupling hooks; outer rami longer than inner rami, outer and inner rami with 6 and 5 articles, respectively. Uropod 1 slender; peduncle with 5 lateral and 1 mediodistal robust setae; outer ramus ca. 0.75 times length of peduncle, bearing 4 lateral and 2 medial robust setae; inner ramus 1.25 times as long as outer ramus, with 2 lateral and 2 medial robust setae. Uropod 2 about 0.8 times (estimated) length of uropod 1; peduncle with 1 lateral and 3 medial robust setae; outer ramus ca. 0.85 times length of peduncle, bearing 2 lateral and 2 medial robust setae; inner ramus longer and wider than outer ramus, with 1 lateral and 1 medial robust setae. Uropod 3 about 0.8 times (estimated) length of uropod 2; peduncle without robust setae; outer ramus ca. 0.65 times as long as peduncle, lanceolate, bare; inner ramus ca. 1.7 times length as long as outer ramus, with 2 dorsal and 2 medial setae. Telson roundish triangular, tip acute (damaged). Sexually dimorphic characters in males. Based on paratype, 1.3 mm (OMNH-Ar- 12920), and paratype, 1.2 mm (OMNH-Ar- 12918) only for antenna 2. Antenna 1 (Fig. 4 M 1 - RA 1), length ratio of peduncular articles 1 – 3 (excluding processes) 1: 0.55: 0.2, lengths of posterodistal processes on articles 1 and 2 ca. 0.3 and 1.2 times of article proper, respectively, peduncular article 2 shorter and posterodistal process of article 2 longer than those of female; flagellum with 4 articles, article 1 long, article 4 minute, posterior surface of each article bearing numerous aesthetascs. Antenna 2 (Fig. 4 M 2 - A 2) more slender than that of female, with ratio of lengths of peduncular articles 3 – 5 1: 1.8: 2.1, posterodistal process of peduncular article 4 short, 25 % length of article 5; flagellum with 4 articles, article 1 long, article 4 minute. Coxa 3 (Fig. 4 M 1 - C 3) long, anteroproximal lobe smaller than that of female. Coxa 4 (Fig. 4 M 1 - C 4) short, anteroproximal and anterodistal lobes smaller than those of female, posteroproximal lobe absent. Gnathopods (Fig. 4 M 1 - G 1, G 2) almost same as those of female. Shape changes in coxae. Based on paratype, female, 1.5 mm (OMNH-Ar- 12917) (Fig. 5). Shapes of coxae 2 – 6 changing probably with growth: coxa 2 posteroproximal corner projected; coxa 3 anteroproximal lobe larger, posteroproximal corner projected; coxa 4 anteroproximal lobe longer, posterodistal corner projected; coxa 5 anterodistal lobe longer, posteroproximal corner projected; coxa 6 anteroproximal corner weakly projected. Coloration of recently fixed specimen. Based on paratype, male, 1.2 mm (OMNH-Ar- 12918) (Fig. 6). Head and pereonites white, light orange internal organs seen through, dorsal margins of pereonites each with dark-red mark; eyes red. Pleonites and urosomites white and partly pale orange, dorsal margins of pleonites 1 – 3 each with small dark-red mark. Antenna 1 peduncular articles 2, 3 and flagellar article 1 orange, other parts white. Antenna 2 peduncular article 5 and flagellar article 1 dark brown, other parts white. Mouthparts including mandibular palp white. Pereopods and uropods white and pale orange.	en	Ariyama, Hiroyuki (2024): Curidia japonica sp. nov., the First Species of the Family Ochlesidae from the Northwest Pacific (Crustacea: Amphipoda). Species Diversity 29 (2): 199-207, DOI: 10.12782/specdiv.29.199, URL: http://dx.doi.org/10.12782/specdiv.29.199
560A8789FF85FFD8FC5EF849FAECF7E8.taxon	discussion	Remarks. Curidia japonica sp. nov. can be clearly distinguished from its congeners by the very short posterodistal process of the antenna 2 peduncular article 4. Length of the process in this new species is 15 % and 25 % length of article 5 in female and male, respectively, whereas in the other Curidia species, lengths of the process are ca. 0.45 – 0.95 times length of article 5 in females and ca. 0.45 – 1.1 times length of article 5 in males. Habitat. 1.5 – 7 m depth, among red algae and bryozoans.	en	Ariyama, Hiroyuki (2024): Curidia japonica sp. nov., the First Species of the Family Ochlesidae from the Northwest Pacific (Crustacea: Amphipoda). Species Diversity 29 (2): 199-207, DOI: 10.12782/specdiv.29.199, URL: http://dx.doi.org/10.12782/specdiv.29.199
560A8789FF85FFD8FC5EF849FAECF7E8.taxon	distribution	Distribution. Japan: Mie and Wakayama Prefectures.	en	Ariyama, Hiroyuki (2024): Curidia japonica sp. nov., the First Species of the Family Ochlesidae from the Northwest Pacific (Crustacea: Amphipoda). Species Diversity 29 (2): 199-207, DOI: 10.12782/specdiv.29.199, URL: http://dx.doi.org/10.12782/specdiv.29.199
