taxonID	type	description	language	source
556787D3025BFF993FCAFCB7FAF1FEC7.taxon	description	Description (New Zealand). Body length 2.8 – 10.1 mm; not pedunculate. Head. Mandibles with setiferous puncture in scrobe. Labrum slightly emarginate anteriorly (rarely subtruncate). Frontal furrows not extended to posterior margin of eyes. Eyes present; a single setiferous puncture (posteriorly) or two punctures on inner side of each eye (without puncture in Oopterus suavis). Tempora inflated or not. Clypeus with a setiferous puncture on each side. Antennae submoniliform (subfiliform in Oopterus arthurensis, O. ocularius, O. trechoides, and genus Zolus); segment 1 (scape) with a few setae, segment 2 with numerous setae, segments 3 – 11 densely pubescent; segments 2 – 10 with verticillate setae. Mentum: medial tooth present; circular foveae absent; two setae present. Mentum-submentum suture present (absent in Pseudoopterus plicaticollis). Submentum with four setae. Ligula with two fused setae (Oopterus, Zolus, Pseudoopterus (in part )), four setae (two medial setae fused, two lateral setae free in Maungazolus) or six setae (two medial setae fused, four lateral setae free in Pseudoopterus plicaticollis and Synteratus). Palpi: terminal segment conical, glabrous, obtuse at tip; penultimate maxillary segment with numerous long setae; penultimate labial segment with two setae on anterior margin. Thorax. Pronotum with or without lateral groove between lateral margin and laterobasal fovea; usually two setiferous punctures on each side or without puncture (rarely with a single setiferous puncture anteriorly or posteriorly); posterolateral carinae present or absent. Scutellum visible, inserted entirely between elytral bases (invisible, concealed under pronotal base in Synteratus). Legs. Protibiae without outer apical prolongation. Tarsi pubescent dorsally; basal segments 1 + 2 of male protarsi asymmetrical, dentate and dilated on inner side; claws entire ventrally; unguitractor plate invisible between tarsal claws. Elytra. Fused along suture (hindwings vestigial). Basal margin complete (reaching about scutellum) or incomplete (reaching about stria 3 or 4). Discal setiferous punctures present. Stria 1 recurrent at apex (curving back like a hook as in Trechini and Bembidiini (Tachyina )). Subapical seta present (absent in Maungazolus pygmeatus and Oopterus clivinoides). Umbilicate series separated into two major groups (four + five), with posterior group rather continuous. Radial field without fine, dense pubescence. Sutural apices obtuse or rounded. Epipleura twisted (with inner fold or plica) near apex. Abdomen. Apex invisible dorsally. Sterna IV – VI with paired ambulatory setae only. Aedeagus. Basal orifice of aedeagus closed anteriorly (Oopterus, Zolus) or open anteriorly (Maungazolus, Pseudoopterus, Synteratus). Right paramere curved, very long, narrow, untwisted medially (Maungazolus, Oopterus, Pseudoopterus, Synteratus) or rather straight, moderately long and wide, twisted medially (Zolus). Left paramere subtriangular.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30258FF943FCAFE57FA77FD87.taxon	description	Description (New Zealand). Aedeagus with basal orifice closed anteriorly; basal bulb without lateral lobes.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30258FF943FCAFE57FA77FD87.taxon	discussion	Remarks. Two New Zealand genera (Zolus Broun, Oopterus Guérin-Méneville) share the genitalic characters assigned to Zolina by Jeannel (1940; as Oopterini), as well as by Roig-Juñent and Cicchino (2001): aedeagus with basal orifice closed anteriorly; basal bulb without lateral lobes. See Roig-Juñent and Cicchino (2001) for comments on the priority of the subtribal name Zolina over Oopterina. Genus Zolus Sharp, 1886, reinstated Fig. 19 – 23, 66 – 70, 154 – 161	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30258FF943FCAFE57FA77FD87.taxon	type_taxon	Type species. Zolus helmsi Sharp, 1886, by monotypy.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30258FF943FCAFE57FA77FD87.taxon	description	Description. Body length 7.0 – 10.1 mm; not ovate, narrowed around bases of pronotum and elytra. Color mostly dark. Metallic luster absent. Dorsal surface mostly glabrous. Head. Narrow or moderately wide. Mandibles moderately long. Labrum slightly emarginate or subtruncate anteriorly. Eyes convex; a single setiferous puncture posteriorly (two setiferous punctures on inner side of each eye in Z. subopacus). Antennae subfiliform. Tempora not inflated. Mentum tooth entire (acute apically). Ligula with two fused setae. Thorax. Pronotum subquadrate (cordate in Z. wongi); lateral grooves present; setiferous punctures absent on each side (a single puncture present in Z. opacus and Z. unisetosus); posterolateral carinae present; base narrower than apex, about as wide as apex or wider than apex. Scutellum visible, inserted entirely between elytral bases. Legs. Very long (moderately long in Z. subopacus). Segments 1 + 2 of male protarsi with a single row of ventral scales. Elytra. Ovate, subovate, or oblong. Basal margin complete, reaching about scutellum (incomplete, reaching about interval 4 in Z. subopacus). Shoulders well developed, rounded, obtuse or oblique. Scutellar setiferous pore present, inserted at junction of striae 1 + 2. Scutellar striole present (obsolete in Z. subopacus). Striae complete (almost complete in Z. subopacus), consisting of lines; three discal setiferous punctures in stria 3 or on interval 3; recurrent stria directed apically toward stria 5, 6 or 7. Subapical seta present. Apical seta present. Sutural apices rounded. Abdomen. Last visible sternum (sternum VII): male with two apical ambulatory setae; female with four apical ambulatory setae. Aedeagus. Lateral view: dorsal membranous area moderately wide or very wide (narrow in Z. subopacus). Dorsal view (Fig. 15): very wide (moderately wide in Z. subopacus); basal orifice wide (narrow in Z. subopacus), closed anteriorly, moderately or very distant from membranous area; lateral lobes of basal bulb absent. Parameres with four to seven apical setae. Right paramere rather straight (curved in other zoline genera), moderately long and wide, twisted medially (untwisted in other zoline genera).	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30258FF943FCAFE57FA77FD87.taxon	discussion	Remarks. The genus Zolus Sharp, 1886 was previously synonymized with Oopterus. This taxon is resurrected from synonymy here on the basis of the rather straight, moderately long, wide, and medially twisted right paramere of the aedeagus, a character unifying included species and distinguishing Zolus from Oopterus. In addition, Zolus species share the following external characters: rather large, body length 7.0 – 10.1 mm; color dark; eyes convex, antennae subfiliform; pronotum subquadrate, with lateral grooves and posterolateral carinae. The genus Zolus now includes three newly described species, two species previously described in Zolus, and one species originally described in Oopterus. The taxonomic names Zolus helmsi and Z. subopacus are here reinstated to their original combinations.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30257FF943FCAFD94FF24FAE4.taxon	description	Pronotum with a setiferous puncture on each side (anteriorly); head with a single setiferous puncture on inner side of each eye (posteriorly); basal margin of elytra complete, reaching about scutellum; Fig. 159. [Body length 9.2 – 10.1 mm; southwestern South Island (OL, WD)] ........ ................................................................................................. Zolus unisetosus new species	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30257FF943FCAFD94FF24FAE4.taxon	description	Pronotum moderately convex; elytral sides moderately rounded; antennal segment 1 (scape) very long, about 2.5 x longer than its maximum width; Fig. 22 – 23. [South Island] .............. 5	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30256FF963FCAFF37FDB3FE87.taxon	description	Fig. 19, 66, 157	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30256FF963FCAFF37FDB3FE87.taxon	description	Description. Body length 7.1 – 7.7 mm. Head, pronotum, elytra, and abdomen mostly dark brown; apex of head reddish; lateral margins, suture, and apex of elytra reddish; abdomen reddish laterally and apically; antennae, palpi, and legs reddish; femora dark brown. Microsculpture very strong, isodiametric. Iridescence absent. Dull, without metallic luster. Head. Labrum strongly transverse, subtruncate anteriorly. Antennae subfiliform, very long: segment 1 (scape) moderately long, about 2 x longer than its maximum width. Frontal furrows wide, shallow, convergent. Eyes strongly convex; two setiferous punctures on inner side of each eye. Tempora not inflated. Mentum: medial tooth entire, acute apically, moderately shorter than lateral lobes. Paraglossae membranous, prominent, about as long as ligula. Thorax. Pronotum slightly convex, impunctate, wrinkled across base, subquadrate, widest about middle; apex emarginate; anterolateral angles well developed, obtusely rounded; anterior bead incomplete, obsolete medially; sides moderately rounded, slightly sinuate posteriorly; lateral grooves very deep, very long (about as long as adjoining foveae); a single setiferous puncture on each side (posteriorly); posterolateral angles rectangular; posterolateral carinae blunt, very long (longer than adjoining foveae); laterobasal foveae well defined, impunctate, moderately deep, very wide, oblong, prolonged forward, simple; posterior bead absent; base emarginate, about as wide as pronotal apex and elytral base. Legs. Moderately long. Elytra. Moderately convex, subovate, widest about middle. Basal margin incomplete, reaching about interval 4. Shoulders obtuse. Sides moderately rounded. Scutellar striole obsolete. Striae almost complete, feebly impressed, shallow, finely punctate; stria 3 or interval 3 with three setiferous punctures. Recurrent stria long, sharp, directed apically toward stria 6. Intervals depressed. Sutural apices strongly rounded. Abdomen. Last visible sternum (sternum VII): male with two apical ambulatory setae; female with four apical ambulatory setae. Aedeagus. Lateral view (Fig. 66): unusually strongly arcuate, widest medially, narrowed in apical half; base strongly convex dorsally; middle strongly convex dorsally, strongly concave ventrally, with dorsal membranous area narrow (wide in other Zolus species) and short; apex slender, moderately concave dorsally, slightly concave ventrally, with extreme tip unusually slender. Dorsal view: moderately wide, asymmetrical (ostium of membranous area deflected to right); apex straight; basal orifice narrow, closed anteriorly, very distant from membranous area. Parameres with four apical setae.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30256FF963FCAFF37FDB3FE87.taxon	materials_examined	Material examined. 59 specimens (AMNZ, BMNH, JNNZ, LUNZ, MONZ NZAC). Geographic distribution (Fig. 157). South Island: CO, FD, OL, SL.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30256FF963FCAFF37FDB3FE87.taxon	biology_ecology	Ecology. Lowland, montane, subalpine. Epigean. Forests (beech, broadleaf, podocarp). Shaded ground. Nocturnal; hides during the day under logs, fallen trees, and stones. Gregarious. Biology. Seasonality: December – March, August. Predacious (based on mouthpart morphology). Dispersal power. Subapterous (incapable of flight). Moderate runner. Collecting techniques. Lifting logs, fallen trees, and stones; pitfall trapping.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30255FF973FCAFE97FD03FDA7.taxon	description	Fig. 159, 160, 161 Holotype: male (NZAC) labeled: “ NEW ZEALAND OL Mt [= Mount] Aspiring NP [= National Park], Blue Pools walk 350 m - 44.1623 169.2736 7. III. 2017 Larivière, Larochelle (typed) / Wet silver beech forest on river terrace. Under logs and fallen branches. (typed) / HOLOTYPE [male symbol] Zolus unisetosus Larochelle & Larivière, 2017 (red label; typed). ” Paratypes: one female (NZAC) from Cameron Flat, Mount Aspiring National Park (OL) and two females (NZAC) from Mount Kinnaird, Westland National Park (WD), bearing blue paratype labels. Description. Body length 9.2 – 10.1 mm. Head, pronotum, elytra, and abdomen mostly black piceous; abdomen yellow laterally; antennae, palpi, and legs reddish; apical half of femora pale yellow. Microsculpture moderately deep, isodiametric on head, moderately transverse on pronotum, very deep and strongly isodiametric on elytra. Iridescence absent. Very shiny on head and pronotum, dull on elytra, without metallic luster. Head. Labrum strongly transverse, slightly emarginate anteriorly. Antennae subfiliform, very long: segment 1 (scape) very long, about 2.5 x longer than its maximum width. Frontal furrows wide, deep, convergent. Eyes strongly convex; a single setiferous puncture on inner side of each eye (posteriorly). Tempora not inflated. Mentum: medial tooth entire, acute apically, moderately shorter than lateral lobes. Paraglossae membranous, prominent, about as long as ligula. Thorax. Pronotum strongly convex, coarsely punctate across base, wrinkled discally and across base, moderately cordate, widest about middle; apex truncate; anterolateral angles poorly developed, rounded; anterior bead complete; sides strongly rounded, moderately sinuate posteriorly; lateral grooves very deep, very long (longer than adjoining foveae); a single setiferous puncture on each side (anteriorly); posterolateral angles rectangular; posterolateral carinae sharp, very long (longer than adjoining foveae); laterobasal foveae well defined, coarsely punctate, very deep and wide, oblong, prolonged forward, simple; posterior bead absent; base depressed throughout (convex medially in other Zolus species), emarginate, much wider than pronotal apex, about as wide as elytral base. Legs. Very long. Elytra. Moderately convex, ovate, widest about middle. Basal margin complete, reaching about scutellum. Shoulders slightly rounded. Sides strongly rounded. Scutellar striole present. Striae complete, very deep, impunctate; stria 3 or interval 3 with three setiferous punctures. Recurrent stria long, sharp, directed apically toward stria 5. Intervals moderately convex. Sutural apices broadly rounded. Abdomen. Last visible sternum (sternum VII): male with two apical ambulatory setae; female with four apical ambulatory setae. Aedeagus. Lateral view (Fig. 160): strongly arcuate, moderately widened in apical half; base slightly convex dorsally; middle angulate anteriorly and moderately sinuate dorsally, moderately convex ventrally, with dorsal membranous area very wide and long; apex triangular, strongly convex dorsally, moderately convex ventrally, with extreme tip wide and short. Dorsal view: very wide, asymmetrical (ostium of membranous area deflected to right); apex straight; basal orifice wide, closed anteriorly, moderately distant from membranous area. Parameres with six apical setae.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30255FF973FCAFE97FD03FDA7.taxon	materials_examined	Material examined. 12 specimens (CMNZ, NZAC). Geographic distribution (Fig. 161). South Island: OL – Mount Aspiring National Park (Blue Pools Walk; Cameron Flat). WD – Jackson Bay. Westland National Park (Mount Kinnaird). Open Bay Islands.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30255FF973FCAFE97FD03FDA7.taxon	biology_ecology	Ecology. Lowland, montane. Epigean. Wet forests (beech, broadleaf). Shaded. Nocturnal; active at night on trees; hides during the day under logs, fallen branches, and stones. Biology. Seasonality: September, December, August. Predacious (based on mouthpart morphology). Occasionally infested with fungi (Laboulbeniales). Dispersal power. Subapterous (incapable of flight). Fast runner. Regular climber. Collecting techniques. Lifting logs, fallen branches, and stones.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30255FF973FCAFE97FD03FDA7.taxon	discussion	Remarks. This species is based on the Latin uni (one) and setosus (setose), referring to the pronotum with a single setiferous puncture on each side (anteriorly). Zolus unisetosus is morphologically close to Z. wongi. In addition to diagnostic characters of the male genitalia, Z. unisetosus has the following distinguishing features: body length 9.2 – 10.1 mm; eyes strongly convex; pronotum strongly convex, widest about middle, sides strongly rounded, a single setiferous puncture on each side (anteriorly), base depressed throughout; elytra with intervals moderately convex and sutural apices broadly rounded. Zolus unisetosus is restricted to southwestern South Island (OL, WD), while Z. wongi occurs mostly in western areas of the South Island (SD to WD).	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30254FF903FCAFD77FD0FFCA7.taxon	description	Fig. 20, 67, 156 Holotype: male (NZAC) labeled: “ NEW ZEALAND CL (typed) Tapu-Coroglen Saddle 400 m 13 Feb 1979 J. S. Dugdale (hand-written) / under stones in bush tunnel (hand-written) / [male symbol] / HOLOTYPE [male symbol] Zolus kauriensis Larochelle & Larivière, 2017 (red label; typed). ” Paratype: one female (NZAC) from Nihotapu [= Nihotupu], Waitakere Ranges (AK), East side, 29. Nov. 1956, K. A. J. Wise, bearing a blue paratype label. Description. Body length 8.0 – 8.3 mm. Head, pronotum, and elytra mostly piceous black; abdomen mostly dark brown, yellow laterally and apically; lateral margins of elytra reddish; antennae, palpi, and legs entirely reddish. Microsculpture very strong, isodiametric on head, very transverse (with microlines) on pronotum, moderately transverse on elytra. Iridescence absent on head and elytra, very strong on pronotum. Very shiny, without metallic luster. Head. Labrum moderately transverse, slightly emarginate anteriorly. Antennae subfiliform, very long: segment 1 (scape) moderately long, about 2 x longer than its maximum width. Frontal furrows wide, shallow, convergent. Eyes moderately convex; a single setiferous puncture on inner side of each eye (posteriorly). Tempora not inflated. Mentum: medial tooth entire, acute apically, moderately shorter than lateral lobes. Paraglossae membranous, prominent, about as long as ligula. Thorax. Pronotum strongly convex, finely punctate across base, wrinkled (discally, medioapically, and between laterobasal foveae), subquadrate, widest about middle; apex subtruncate; anterolateral angles poorly developed, rounded; anterior bead complete; sides strongly rounded, not sinuate posteriorly; lateral grooves moderately deep, very long (about as long as adjoining foveae); setiferous punctures absent on each side; posterolateral carinae sharp, moderately long (about half as long as adjoining foveae); posterolateral angles obtuse (with extreme tip rounded); laterobasal foveae well defined, finely punctate, moderately deep, very wide, square, prolonged forward, simple; posterior bead absent; base emarginate, much narrower than pronotal apex, about as wide as elytral base. Legs. Very long. Elytra. Moderately convex, oblong, widest before middle. Basal margin complete, reaching about scutellum. Shoulders obtuse. Sides strongly rounded. Scutellar striole present. Striae complete, very deep, impunctate; stria 3 or interval 3 with three setiferous punctures. Recurrent stria long, sharp, directed apically toward stria 5. Intervals slightly to moderately convex from base to apex. Sutural apices obtusely rounded. Abdomen. Last visible sternum (sternum VII): male with two apical ambulatory setae; female with four apical ambulatory setae. Aedeagus. Lateral view (Fig. 67): strongly arcuate, not widened in apical half; base moderately biconvex dorsally; middle strongly convex dorsally, slightly convex ventrally, with dorsal membranous area very wide and long; apex subtriangular, slightly convex dorsally, almost straight ventrally, with extreme tip narrow and long. Dorsal view: very wide, asymmetrical (ostium of membranous area deflected to right); apex straight; basal orifice wide, closed anteriorly, moderately distant from membranous area. Parameres with five apical setae.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30254FF903FCAFD77FD0FFCA7.taxon	materials_examined	Material examined. 2 specimens (NZAC). Geographic distribution (Fig. 156). North Island: AK – Nihotapu, Waitakeres [= Nihotupu, Waitakere Ranges]. CL – Tapu-Coroglen Saddle.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30254FF903FCAFD77FD0FFCA7.taxon	biology_ecology	Ecology. Lowland (hills). Forests (broadleaf-podocarp). Nocturnal; hides during the day under stones. Biology. Seasonality: November, February. Predacious (based on mouthpart morphology). Dispersal power. Subapterous (incapable of flight). Fast runner. Collecting technique. Lifting stones.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30254FF903FCAFD77FD0FFCA7.taxon	discussion	Remarks. This species is named after the geographic range of the kauri tree (Agathis australis) where the beetle occurs, and the Latin suffix - e nsis, denoting a place, locality, or country. Zolus kauriensis is morphologically close to Z. carinatus. In addition to diagnostic characters of the male genitalia, Z. kauriensis has the following distinguishing features: pronotum widest about middle, sides strongly rounded, not sinuate posteriorly, lateral grooves moderately deep, posterolateral angles obtuse and posterolateral carinae moderately long (about half as long as adjoining foveae); elytra oblong, widest before middle, microsculpture moderately transverse, intervals convex. The two species are allopatric: Z. kauriensis appears to be restricted to northern areas of the North Island, while Z. carinatus occurs mostly south of the Central Volcanic Plateau.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30253FF913FCAFC77FE63F984.taxon	description	Fig. 21, 68, 154	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30253FF913FCAFC77FE63F984.taxon	description	Description. Body length 7.3 – 9.2 mm. Head, pronotum, and elytra mostly piceous black; abdomen mostly dark brown, yellow laterally and apically; lateral margins of elytra reddish; antennae, palpi, and legs reddish; femora pale yellow basally. Microsculpture moderately strong and isodiametric on head, very strong and transverse (with microlines) on pronotum and elytra. Iridescence absent on head, very strong on pronotum and elytra. Very shiny, without metallic luster. Head. Labrum moderately transverse, slightly emarginate anteriorly. Antennae subfiliform, very long: segment 1 (scape) moderately long, about 2 x longer than its maximum width. Frontal furrows wide, shallow, convergent. Eyes moderately convex; a single setiferous puncture on inner side of each eye (posteriorly). Tempora not inflated. Mentum: medial tooth entire, acute apically, moderately shorter than lateral lobes. Paraglossae membranous, prominent, about as long as ligula. Thorax. Pronotum strongly convex, finely punctate across base, wrinkled (discally, medioapically, and between laterobasal foveae), subquadrate, widest before middle; apex subtruncate; anterolateral angles poorly developed, rounded; anterior bead complete; sides moderately rounded, moderately sinuate posteriorly; lateral grooves very deep, very long (about as long as adjoining foveae); setiferous punctures absent on each side; posterolateral angles subrectangular; posterolateral carinae sharp, very long (about as long adjoining foveae); laterobasal foveae well defined, finely punctate, very deep and wide, square, prolonged forward, simple; posterior bead absent; base emarginate, much narrower than pronotal apex, about as wide as elytral base. Legs. Very long. Elytra. Moderately convex, ovate, widest about middle (wider at base than in other Zolus species). Basal margin complete, reaching about scutellum. Shoulders obtuse. Sides strongly rounded. Scutellar striole present. Striae complete, moderately deep, impunctate or finely punctate; stria 3 or interval 3 with three setiferous punctures. Recurrent stria long, sharp, directed apically toward stria 5, 6 or 7. Intervals subdepressed. Sutural apices obtusely rounded. Abdomen. Last visible sternum (sternum VII): male with two apical ambulatory setae; female with four apical ambulatory setae. Aedeagus. Lateral view (Fig. 68): strongly arcuate, not widened in apical half; base slightly convex dorsally; middle subparallel, almost straight dorsally, moderately concave ventrally, with dorsal membranous area moderately wide and very long; apex subtriangular, slightly convex dorsally, slightly concave ventrally, with extreme tip narrow and long. Dorsal view: very wide, asymmetrical (ostium of membranous area deflected to right); apex straight; basal orifice wide, closed anteriorly, moderately distant from membranous area. Parameres with five to seven apical setae.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30253FF913FCAFC77FE63F984.taxon	materials_examined	Material examined. 545 specimens (BMNH, CMNH, CMNZ, JNNZ, LUNZ, MONZ, NZAC). Geographic distribution (Fig. 154). North Island: AK, BP, GB, HB, RI, TK, TO, WA, WI, WN, WO.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30253FF913FCAFC77FE63F984.taxon	biology_ecology	Ecology. Lowland, montane, subalpine. Epigean-arboreal. Wet or dry forests (beech, broadleaf, podocarp). Shaded ground. Nocturnal; active at night on trees and logs; hides during the day in fallen branches and in rotten logs. Gregarious. Biology. Seasonality: throughout the year, except June. Predacious (based on mouthpart morphology). Dispersal power. Subapterous (incapable of flight). Fast runner. Occasional climber on trees and logs. Collecting techniques. Breaking rotten branches and logs; pitfall trapping.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30253FF913FCAFC77FE63F984.taxon	discussion	Remark. Examination of the types of Zolus atratus and Z. femoralis revealed them to be conspecific with Z. carinatus.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30252FF933FCAF997FC1EFE64.taxon	description	Fig. 22, 69, 155	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30252FF933FCAF997FC1EFE64.taxon	description	Description. Body length 7.6 – 8.9 mm. Head, pronotum, elytra, and abdomen mostly reddish piceous; base of pronotum reddish; lateral margins and suture of elytra reddish; abdomen reddish laterally and apically; antennae, palpi, and legs reddish; femora pale yellow. Microsculpture obsolete and isodiametric on head, feeble, very transverse (with microlines) on pronotum, moderately transverse (brick-like) on elytra. Iridescence absent on head and elytra, moderately strong on pronotum. Very shiny, without metallic luster. Head. Labrum moderately transverse, subtruncate anteriorly. Antennae subfiliform, very long: segment 1 (scape) very long, about 2.5 x longer than its maximum width. Frontal furrows wide, shallow, convergent. Eyes strongly convex; a single setiferous puncture on inner side of each eye (posteriorly). Tempora not inflated. Mentum: medial tooth entire, acute apically, moderately shorter than lateral lobes. Paraglossae membranous, prominent, about as long as ligula. Thorax. Pronotum moderately convex, finely punctate across base and apex, wrinkled across base, subquadrate, widest about middle; apex subtruncate; anterolateral angles poorly developed, rounded; anterior bead incomplete, obsolete medially; sides moderately rounded, slightly sinuate posteriorly; lateral grooves very deep, very long (longer than adjoining foveae); setiferous punctures absent on each side; posterolateral angles rectangular; posterolateral carinae sharp, very long (longer than adjoining foveae); laterobasal foveae well defined, finely punctate, very deep and wide, square, prolonged forward, simple; posterior bead absent; base emarginate, about as wide as pronotal apex and elytral base. Legs. Very long. Elytra. Moderately convex, subovate, widest before middle. Basal margin complete, reaching about scutellum. Shoulders oblique. Sides moderately rounded. Scutellar striole present. Striae complete, moderately deep, finely punctate; stria 3 or interval 3 with three setiferous punctures. Recurrent stria long, sharp, directed apically toward stria 5. Intervals subdepressed. Sutural apices strongly rounded. Abdomen. Last visible sternum (sternum VII): male with two apical ambulatory setae; female with four apical ambulatory setae. Aedeagus. Lateral view (Fig. 69): strongly arcuate, strongly widened in apical half; base moderately convex dorsally; middle angulate anteriorly and moderately convex dorsally, slightly concave ventrally, with dorsal membranous area moderately wide and very long; apex subtriangular, strongly convex dorsally, almost straight ventrally, with extreme tip wide and rather short. Dorsal view: very wide, asymmetrical (ostium of membranous area deflected to right); apex straight; basal orifice wide, closed anteriorly, moderately distant from membranous area. Parameres with five or six apical setae.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30252FF933FCAF997FC1EFE64.taxon	materials_examined	Material examined. 233 specimens (AMNZ, BMNH, CMNZ, JNNZ, LUNZ, MONZ, NZAC). Geographic distribution (Fig. 155). South Island: BR, MB, NN, SD, WD.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30252FF933FCAF997FC1EFE64.taxon	biology_ecology	Ecology. Lowland, montane, subalpine. Epigean-arboreal. Wet forests (beech, podocarp, broadleaf). Shaded ground. Nocturnal; active at night on mossy logs and trees; hides during the day under logs, fallen branches, and the loose bark of fallen trees. Gregarious. Biology. Seasonality: September – March, May – August. Tenerals: February – March, August. Predacious (based on mouthpart morphology). Dispersal power. Subapterous (incapable of flight). Fast runner. Occasional climber on logs and trees. Collecting techniques. Lifting logs, fallen branches, and the loose bark of fallen trees; pitfall trapping.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30252FF933FCAF997FC1EFE64.taxon	discussion	Remarks. Sharp described Z. helmsi from a pair of specimens, one of which (a male) is here designated as lectotype. The second specimen (a female) is considered to be a paralectotype. These type designations are made to preserve stability of nomenclature in the future. As stated in Broun’s description, the male holotype of Z. labralis (BMNH) only has four setiferous punctures on the labrum; this represents an anomaly. In all other material seen, the labrum displays six setiferous punctures. Examination of the type of Z. labralis revealed it to be conspecific with Z. helmsi.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30250FF8D3FCAFDB7FB47FD07.taxon	description	Fig. 23, 70, 158 Holotype: male (NZAC) labeled: “ [NEW ZEALAND BR] Mt Dewar 1000 m J. C. Watt (typed) / under bark [of] logs (typed) / Paparoa Range Dec. 1969 (typed) / [male symbol] / HOLOTYPE [male symbol] Zolus wongi Larochelle & Larivière, 2017 (red label; typed). ” Paratypes: one female (NZAC) and one male (LUNZ) from the same locality as the holotype, bearing blue paratype labels. Description. Body length 7.0 – 8.3 mm. Head, pronotum, elytra, and abdomen mostly piceous black; abdomen yellow laterally; antennae, palpi, and legs reddish; apical half of femora pale yellow. Microsculpture moderately deep, isodiametric on head, moderately transverse on pronotum, very deep and strongly isodiametric on elytra. Iridescence absent. Very shiny on head and pronotum, dull on elytra, without metallic luster. Head. Labrum strongly transverse, slightly emarginate anteriorly. Antennae subfiliform, very long: segment 1 (scape) very long, about 2.5 x longer than its maximum width. Frontal furrows wide, deep, convergent. Eyes moderately convex; a single setiferous puncture on inner side of each eye (posteriorly). Tempora not inflated. Mentum: medial tooth entire, acute apically, moderately shorter than lateral lobes. Paraglossae membranous, prominent, about as long as ligula. Thorax. Pronotum moderately convex, coarsely punctate across base, wrinkled discally and across base, moderately cordate, widest before middle; apex truncate; anterolateral angles poorly developed, rounded; anterior bead complete; sides moderately rounded, moderately sinuate posteriorly; lateral grooves very deep, very long (longer than adjoining foveae); setiferous punctures absent on each side; posterolateral angles rectangular; posterolateral carinae sharp, very long (longer than adjoining foveae); laterobasal foveae well defined, coarsely punctate, very deep and wide, oblong, prolonged forward, simple; posterior bead absent; base emarginate, much wider than pronotal apex, about as wide as elytral base. Legs. Very long. Elytra. Moderately convex, ovate, widest about middle. Basal margin complete, reaching about scutellum. Shoulders slightly rounded. Sides strongly rounded. Scutellar striole present. Striae complete, very deep, impunctate; stria 3 or interval 3 with three setiferous punctures. Recurrent stria long, sharp, directed apically toward stria 5. Intervals subdepressed. Sutural apices obtusely rounded. Abdomen. Last visible sternum (sternum VII): male with two apical ambulatory setae; female with four apical ambulatory setae. Aedeagus. Lateral view (Fig. 70): strongly arcuate, moderately widened in apical half; base slightly convex dorsally; middle angulate anteriorly and moderately sinuate dorsally, slightly concave ventrally, with dorsal membranous area very wide and long; apex subtriangular, strongly convex dorsally, slightly concave ventrally, with extreme tip wide and long. Dorsal view: very wide, asymmetrical (ostium of membranous area deflected to right); apex straight; basal orifice wide, closed anteriorly, moderately distant from membranous area. Parameres with six or seven apical setae.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30250FF8D3FCAFDB7FB47FD07.taxon	materials_examined	Material examined. 402 specimens (AMNZ, CMNZ, JNNZ, LUNZ, MONZ, NZAC). Geographic distribution (Fig. 158). South Island: BR, MB, MC, MK, NC, NN, SD, WD.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30250FF8D3FCAFDB7FB47FD07.taxon	biology_ecology	Ecology. Lowland, montane, subalpine, alpine. Epigean-arboreal. Wet forests (beech, broadleaf, podocarp), subalpine scrublands, and alpine meadows. Shaded or open ground. Nocturnal; active at night on logs and trees; hides during the day under logs, fallen branches, and stones. Gregarious. Biology. Seasonality: October – August. Tenerals: December (BR), February (NN, SD), March (NC), April (WD). Predacious (based on mouthpart morphology). Occasionally infested with fungi (Laboulbeniales) and mites. Dispersal power. Subapterous (incapable of flight). Fast runner. Regular climber on trees and logs. Collecting techniques. Lifting logs, fallen branches, and stones; pitfall trapping.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30250FF8D3FCAFDB7FB47FD07.taxon	discussion	Remarks. This species is named after Philip Wong (Auckland), a highly skilled health specialist and special acquaintance. Zolus wongi is morphologically close to Z. unisetosus. In addition to diagnostic characters of the male genitalia, Z. wongi has the following distinguishing features: body length 7.0 – 8.3 mm; eyes moderately convex; pronotum moderately convex, widest before middle, sides moderately rounded, setiferous punctures absent on each side, base convex medially; elytra with intervals subdepressed and sutural apices obtusely rounded. Zolus wongi occurs mostly in western areas of the South Island (SD to WD), while Z. unisetosus is restricted to southwestern South Island (OL, WD). Genus Oopterus Guérin-Méneville, 1841 Fig. 24 – 52, 71 – 98, 120 – 148	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30250FF8D3FCAFDB7FB47FD07.taxon	type_taxon	Type species. Oopterus clivinoides Guérin-Méneville, 1841, by monotypy.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30250FF8D3FCAFDB7FB47FD07.taxon	description	Description (New Zealand). Body length 3.4 – 7.9 mm, rarely over 6.5 mm (O. arthurensis, O. ocularius, O. trechoides); not ovate, narrowed around bases of pronotum and elytra. Color dark or pale. Metallic luster present or absent. Dorsal surface mostly glabrous. Head. Narrow or moderately wide. Mandibles moderately long. Labrum slightly emarginate anteriorly (subtruncate in Oopterus palmai). Eyes convex (subdepressed in O. discoideus, O. ocularius, O. quadripunctatus); one or two setiferous punctures on inner side of each eye (without puncture in O. suavis). Antennae submoniliform (subfiliform in O. arthurensis, O. ocularius, O. trechoides). Tempora inflated or not. Mentum tooth entire (usually acute, rarely rounded or subtruncate apically). Ligula with two fused setae. Thorax. Pronotum transverse, subquadrate or cordate; lateral grooves absent (rarely present); usually two setiferous punctures on each side, otherwise a single setiferous puncture or without setiferous puncture; posterolateral carinae usually present; base narrower than apex, as wide as apex or wider than apex. Scutellum visible, inserted entirely between elytral bases. Legs. Moderately long (rarely short or very long). Segments 1 + 2 of male protarsi with a single row of ventral scales. Elytra. Ovate or subovate (oblong in O. nanus). Basal margin incomplete, reaching about stria 4 (rarely complete, reaching about scutellum). Shoulders poorly to well developed, rounded or obtuse (oblique in O. arthurensis, O. trechoides). Scutellar setiferous pore present, inserted at junction of striae 1 + 2. Scutellar striole present or absent. Striae complete or incomplete, consisting of lines (mostly replaced by rows of coarse punctures in O. laeviventris, O. lewisi); usually three discal setiferous punctures in stria or row 3, or on interval 3 (four or five punctures in O. anglemensis, O. nanus, O. quadripunctatus; a single puncture in O. arthurensis; without or with one to three punctures in O. punctatus); recurrent stria directed apically toward stria or row 5, 6 or 7. Subapical seta present (absent in O. clivinoides). Apical seta present. Sutural apices rounded or obtuse. Abdomen. Last visible sternum (sternum VII): male with two apical ambulatory setae (two additional medial ambulatory setae in O. clivinoides, O. marrineri); female with four apical ambulatory setae (two additional medial ambulatory setae in O. clivinoides). Aedeagus. Lateral view: dorsal membranous area narrow (rarely wide). Dorsal view (Fig. 15): moderately or very wide (narrow in O. clivinoides, O. nanus, O. patulus); basal orifice narrow (wide in O. ocularius, O. suavis), closed anteriorly, moderately distant from membranous area (close to membranous area in O. collaris, O. fulvipes, O. patulus); lateral lobes of basal bulb absent. Parameres with two to seven apical setae. Right paramere curved, very long, narrow, untwisted medially.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30250FF8D3FCAFDB7FB47FD07.taxon	discussion	Remarks. In the course of this revision, species previously assigned to the genus Oopterus were found to belong to two subtribes and four genera: Zolina (Oopterus, Zolus reinstated) and Merizodina (Maungazolus new genus, Pseudoopterus reinstated). Three taxa are transferred from Oopterus to Zolus (Oopterus carinatus Broun, 1882; O. helmsi (Sharp, 1886); O. subopacus (Broun, 1915 )), three taxa to Maungazolus (Oopterus pallidipes Broun, 1893; O. parvulus Broun, 1903; O. pygmeatus Broun, 1907), and two taxa to Pseudoopterus (Oopterus plicaticollis Blanchard, 1843; O. latipennis Broun, 1903). The morphological character unifying Oopterus species and distinguishing them from Zolus species is the curved, very long and narrow, medially untwisted right paramere of the aedeagus.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3024BFF893FCAF957FECFF8A7.taxon	description	Fig. 24, 71, 146	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3024BFF893FCAF957FECFF8A7.taxon	description	Description. Body length 4.9 – 5.5 mm. Head, pronotum, elytra, and abdomen mostly reddish piceous; base of pronotum reddish; lateral margins, suture, and apex of elytra reddish; abdomen yellowish laterally and apically; antennae, palpi, and legs reddish; palpi with yellowish tinge; femora pale yellowish, reddish basally. Microsculpture absent. Iridescence absent. Very shiny, without metallic luster. Head. Labrum moderately transverse, slightly emarginate anteriorly. Antennae submoniliform, moderately long: segment 1 (scape) moderately long, about 2 x longer than its maximum width. Frontal furrows wide, deep, convergent. Eyes moderately convex; setiferous punctures absent on inner side of each eye (present in other zoline species). Tempora not inflated. Mentum: medial tooth entire, acute apically, moderately shorter than lateral lobes. Paraglossae membranous, prominent, about as long as ligula. Thorax. Pronotum strongly convex, coarsely punctate across base, unwrinkled, slightly transverse, moderately cordate, widest before middle; apex subtruncate; anterolateral angles poorly developed, obtusely rounded; anterior bead incomplete, obsolete medially; sides strongly rounded, not sinuate posteriorly; lateral grooves moderately deep, very long (about as long as adjoining foveae); setiferous punctures absent on each side; posterolateral angles acute, projected laterally; posterolateral carinae blunt, very long (about as long as adjoining foveae); laterobasal foveae well defined, coarsely punctate, very deep and wide, rounded, not prolonged forward, simple; posterior bead absent; base emarginate, wider than pronotal apex, about as wide as elytral base. Legs. Moderately long. Elytra. Strongly convex, ovate, widest about middle. Basal margin incomplete, reaching about stria 4. Shoulders obtuse. Sides strongly rounded. Scutellar striole present. Stria 1 complete, deep; striae 2 – 7 incomplete, shallow, finely punctate; interval 3 with three setiferous punctures. Recurrent stria long, sharp, directed apically toward stria 5. Subapical seta present. Intervals subdepressed. Sutural apices obtusely rounded. Abdomen. Last visible sternum (sternum VII): male with two apical ambulatory setae; female with four apical ambulatory setae. Aedeagus. Lateral view (Fig. 71): moderately arcuate, strongly widened in apical half; base moderately biconvex dorsally; middle moderately convex dorsally, moderately concave ventrally, with dorsal membranous area moderately wide and very long; apex subtriangular, strongly concave dorsally, almost straight ventrally, with extreme tip unusually wide and short. Dorsal view: narrow, asymmetrical (ostium of membranous area deflected to right); apex straight; basal orifice moderately wide, closed anteriorly, moderately distant from membranous area. Parameres with five apical setae.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3024BFF893FCAF957FECFF8A7.taxon	materials_examined	Material examined. 202 specimens (AMNZ, BMNH, JNNZ, LUNZ, MONZ, NZAC). Geographic distribution (Fig. 146). South Island: FD, OL, SL.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3024BFF893FCAF957FECFF8A7.taxon	biology_ecology	Ecology. Lowland, montane, subalpine. Epigean-arboreal. Forests (beech). Shaded ground. Nocturnal; active at night on mossy trees and logs; hides during the day under the loose bark of fallen trees. Gregarious. Biology. Seasonality: October – March, August. Tenerals: December – March. Predacious (based on mouthpart morphology). Defense mechanism: feigns death when disturbed. Dispersal power. Subapterous (incapable of flight). Moderate runner. Regular climber on trees and logs. Collecting technique. Lifting loose bark of fallen trees.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3024BFF893FCAF957FECFF8A7.taxon	discussion	Remark. Oopterus suavis is the only zoline species without setiferous punctures on the inner side of each eye.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30249FF8B3FCAFF37FD04FE64.taxon	description	Fig. 25, 72, 129	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30249FF8B3FCAFF37FD04FE64.taxon	description	Broun Coll. Brit. Mus. 1922 – 482. (white label with red horizontal line; typed) / Oöpterus puncticeps (hand-written). ” New synonym Pseudoopterus laevicollis: Csiki 1928: 225. Pseudoopterus puncticeps: Csiki 1928: 225. Oopterus laevicollis: Hudson 1934: 177. Oopterus puncticeps: Hudson 1934: 177. Description. Body length 5.9 – 6.4 mm. Head, pronotum, elytra, and abdomen mostly reddish piceous; lateral margins and suture of elytra reddish; abdomen reddish piceous, yellow laterally and apically; antennae, palpi, and legs reddish; femora pale yellow. Microsculpture obsolete and isodiametric on head, strong, moderately transverse on pronotum, very transverse (with microlines) on elytra. Iridescence absent on head and pronotum, very strong on elytra. Very shiny, with metallic luster (aeneous) on elytra. Head. Labrum strongly transverse, slightly emarginate anteriorly. Antennae submoniliform, moderately long: segment 1 (scape) short, stout, about 1.5 x longer than its maximum width. Frontal furrows wide, shallow, convergent. Eyes moderately convex; two setiferous punctures on inner side of each eye. Tempora not inflated. Mentum: medial tooth entire, acute apically, moderately shorter than lateral lobes. Paraglossae membranous, prominent, about as long as ligula. Thorax. Pronotum moderately convex, finely punctate across base, wrinkled discally, subquadrate, widest about middle; apex subtruncate; anterolateral angles poorly developed, rounded; anterior bead complete; sides moderately rounded, not sinuate posteriorly; lateral grooves absent; setiferous punctures absent on each side; posterolateral angles obtuse; posterolateral carinae blunt, very long (about as long as adjoining foveae); laterobasal foveae well defined, finely punctate, very deep and wide, square, prolonged forward, double; posterior bead absent; base emarginate, about as wide as pronotal apex, slightly narrower than elytral base. Legs. Moderately long. Elytra. Moderately convex, subovate, widest about middle. Basal margin complete, reaching about scutellum. Shoulders obtuse. Sides moderately rounded. Scutellar striole obsolete. Striae complete, shallow, finely punctate; stria 3 with three setiferous punctures. Recurrent stria long, sharp, directed apically toward stria 5. Subapical seta present. Intervals subdepressed. Sutural apices obtusely rounded. Abdomen. Last visible sternum (sternum VII): male with two apical ambulatory setae; female with four apical ambulatory setae. Aedeagus. Lateral view (Fig. 72): strongly arcuate, narrowed in apical half; base strongly convex dorsally; middle strongly sinuate dorsally, moderately convex ventrally, with dorsal membranous area moderately wide and long; apex slender, slightly convex dorsally, moderately concave ventrally, with extreme tip unusually slender. Dorsal view: moderately wide, asymmetrical (ostium of membranous area deflected to left); apex deflected to left; basal orifice narrow, closed anteriorly, moderately distant from membranous area. Parameres with three apical setae.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30249FF8B3FCAFF37FD04FE64.taxon	materials_examined	Material examined. 85 specimens (AMNZ, BMNH, CMNZ, JNNZ, LUNZ, MNHN, NZAC). Geographic distribution (Fig. 129). South Island: MC – Banks Peninsula. Port Hills.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30249FF8B3FCAFF37FD04FE64.taxon	biology_ecology	Ecology. Lowland (hills). Epigean-arboreal. Wet or dry forests (broadleaf, podocarp, beech) and scrublands. Nocturnal; active on tree-trunks at night; hides during the day under logs and in leaf litter. Biology. Seasonality: throughout the year, except September, August. Tenerals: November – December, February. Predacious (based on mouthpart morphology). Dispersal power. Subapterous (incapable of flight). Moderate runner. Occasional climber on trees. Collecting techniques. Lifting logs; sifting leaf litter; pitfall trapping.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30249FF8B3FCAFF37FD04FE64.taxon	discussion	Remarks. Bates’ original description was based on a single male. Three specimens, two males and one female, were obtained from the Bates collection in Paris (MNHN); the male bearing Bates’ handwritten identification label is believed to be the holotype. Examination of the type of Oopterus puncticeps revealed it to be conspecific with O. laevicollis.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30248FF843FCAFDB4FBCAFD27.taxon	description	Fig. 26, 73, 142 Holotype: male (NZAC) labeled: “ [NEW ZEALAND FD] Mt. [= Mount] Wilmot 1372 m (hand-written) / Manapouri Exp. Jan 70 J. McBurney (typed) / [male symbol] / HOLOTYPE [male symbol] Oopterus quadripunctatus Larochelle & Larivière, 2017 (red label; typed). ” Paratypes: two females (LUNZ, NZAC) from Percy Saddle (North of), Turret Range (FD), bearing blue paratype labels. Description. Body length 4.5 – 5.4 mm. Head, pronotum, and elytra mostly piceous black; abdomen piceous black to rufous; lateral margins and suture of elytra reddish; antennal segments 1 – 3, palpi, and legs reddish; antennal segments 4 – 11 infuscated red. Microsculpture obsolete and isodiametric on head, strong, moderately transverse on pronotum, very isodiametric on elytra. Iridescence absent. Very shiny on head, moderately shiny on pronotum and elytra, without metallic luster. Head. Labrum strongly transverse, slightly emarginate anteriorly. Antennae submoniliform, moderately long: segment 1 (scape) short, stout, about 1.5 x longer than its maximum width. Frontal furrows wide, deep, convergent. Eyes subdepressed; a single setiferous puncture on inner side of each eye (posteriorly). Tempora moderately inflated (about half as long as eyes). Mentum: medial tooth entire, acute apically, moderately shorter than lateral lobes. Paraglossae membranous, prominent, about as long as ligula. Thorax. Pronotum moderately convex, obsoletely punctate across base, wrinkled discally, subquadrate, moderately cordate, widest before middle; apex truncate; anterolateral angles poorly developed, obtusely rounded; anterior bead complete; sides moderately rounded, moderately sinuate posteriorly; lateral grooves absent; setiferous punctures absent on each side; posterolateral angles obtuse; posterolateral carinae blunt, very long (about as long as adjoining foveae); laterobasal foveae well defined, impunctate, very deep, moderately wide, rounded, not prolonged forward, simple; posterior bead absent; base truncate, much narrower than pronotal apex and elytral base. Legs. Moderately long. Elytra. Moderately convex, subovate, widest about middle. Basal margin complete, reaching about scutellum. Shoulders obtuse. Sides moderately rounded. Scutellar striole obsolete. Striae complete, shallow, finely punctate; interval 3 with four foveate setiferous punctures (without or with 1 – 5 non-foveate punctures in other Oopterus species). Recurrent stria long, sharp, directed apically toward stria 5. Subapical seta present. Intervals depressed. Sutural apices rounded. Abdomen. Last visible sternum (sternum VII): male with two apical ambulatory setae; female with four apical ambulatory setae. Aedeagus. Lateral view (Fig. 73): strongly arcuate, moderately widened in apical half; base rather strongly biconvex dorsally; middle moderately concave dorsally, moderately convex ventrally, with dorsal membranous area very wide and long; apex triangular, straight dorsally, slightly concave ventrally, with extreme tip narrow and long. Dorsal view: narrow, asymmetrical (ostium of membranous area deflected to right); apex straight; basal orifice narrow, closed anteriorly, moderately distant from membranous area. Parameres with four or five apical setae.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30248FF843FCAFDB4FBCAFD27.taxon	materials_examined	Material examined. 176 specimens (AMNZ, JNNZ, LUNZ, NZAC). Geographic distribution (Fig. 142). South Island: FD, OL.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30248FF843FCAFDB4FBCAFD27.taxon	biology_ecology	Ecology. Alpine. Endogean. Grasslands, fellfields, screes. Open ground. Nocturnal; hides during the day under well-embedded stones. Gregarious. Biology. Seasonality: November – February. Tenerals: February (FD). Predacious (based on mouthpart morphology). Occasionally infested with fungi (Laboulbeniales). Dispersal power. Subapterous (incapable of flight). Moderate runner. Collecting technique. Lifting stones.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30248FF843FCAFDB4FBCAFD27.taxon	discussion	Remarks. The name of this species is based on the Latin quadri (four) and punctatus (punctate), referring to the four setiferous punctures on elytral interval 3. Oopterus quadripunctatus is morphologically close to O. discoideus and O. marrisi. In addition to diagnostic characters of the male genitalia, O. quadripunctatus has the following distinguishing features: pronotum subquadrate, widest before middle; elytra moderately convex, microsculpture isodiametric, four foveate setiferous punctures on interval 3. All three species occur in southern areas of the South Island.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30247FF853FCAFCF4FD82FC47.taxon	description	Fig. 27, 74, 125 Holotype: male (NZAC) labeled: “ [NEW ZEALAND FD] Wolfe Flat 600 m- 680 m (typed) / Turret Ra [= Range] (typed) / Manapouri Exp. Jan. 70 J. McBurney (typed) / moss (typed) 70 / 69 (hand-written) / [male symbol] / HOLOTYPE [male symbol] Oopterus discoideus Larochelle & Larivière, 2017 (red label; typed). ” Paratypes: two females (LUNZ, NZAC) from the same locality as the holotype, bearing blue paratype labels. Description. Body length 3.4 – 4.2 mm. Head, pronotum, elytra, and abdomen mostly testaceous; disc of head dark rufous; base and disc of pronotum dark rufous; a dark rufous oblong vitta present on each elytron; abdomen dark rufous medially; antennae, palpi, and femora pale yellow; femora (base), tibiae, and tarsi reddish. Microsculpture absent. Iridescence absent. Very shiny, without metallic luster. Head. Labrum strongly transverse, slightly emarginate anteriorly. Antennae submoniliform, moderately long: segment 1 (scape) moderately long, about 2 x longer than its maximum width. Frontal furrows wide, shallow, convergent. Eyes subdepressed; a single setiferous puncture on inner side of each eye (posteriorly). Tempora moderately inflated (about half as long as eyes). Mentum: medial tooth entire, acute apically, moderately shorter than lateral lobes. Paraglossae membranous, prominent, about as long as ligula. Thorax. Pronotum strongly convex, obsoletely punctate across base, wrinkled discally, strongly transverse, widest about middle; apex subtruncate; anterolateral angles poorly developed, obtusely rounded; anterior bead incomplete, obsolete medially; sides strongly rounded, not sinuate posteriorly; lateral grooves absent; setiferous punctures absent on each side; posterolateral angles obtuse; posterolateral carinae blunt, very long (about as long as adjoining foveae); laterobasal foveae well defined, virtually impunctate, very deep and wide, subrectangular, not prolonged forward, double; posterior bead absent; base subtruncate, about as wide as pronotal apex and elytral base. Legs. Moderately long. Elytra. Strongly convex, ovate, widest about middle. Basal margin complete, reaching about scutellum. Shoulders obtuse. Sides strongly rounded. Scutellar striole present. Striae complete, very deep, coarsely punctate; striae 2 – 7 shallower apically; interval 3 with three setiferous punctures. Recurrent stria long, blunt, directed apically toward stria 5. Subapical seta present. Intervals strongly convex. Sutural apices obtuse. Abdomen. Last visible sternum (sternum VII): male with two apical ambulatory setae; female with four apical ambulatory setae. Aedeagus. Lateral view (Fig. 74): moderately arcuate, strongly widened in apical half; base strongly biconvex dorsally; middle moderately convex dorsally, slightly concave ventrally, with dorsal membranous area moderately wide and very long; apex triangular, straight dorsally, slightly concave ventrally, with extreme tip narrow, long, and curved downward. Dorsal view: narrow, asymmetrical (ostium of membranous area deflected to left); apex straight; basal orifice narrow, closed anteriorly, moderately distant from membranous area. Parameres with five apical setae.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30247FF853FCAFCF4FD82FC47.taxon	materials_examined	Material examined. 8 specimens (JNNZ, LUNZ, NZAC). Geographic distribution (Fig. 125). South Island: FD – Fiordland National Park (Borland Saddle; Turret Range, Percy Stream; Wilmot Pass, Spey River; Wolfe Flat).	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30247FF853FCAFCF4FD82FC47.taxon	biology_ecology	Ecology. Montane. Endogean. Wet forests (beech). Shaded ground. Nocturnal; hides during the day in thick leaf litter and moss. Biology. Seasonality: January – February. Predacious (based on mouthpart morphology). Occasionally infested with fungi (Laboulbeniales). Dispersal power. Subapterous (incapable of flight). Moderate runner. Collecting techniques. Sifting leaf litter and moss.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30247FF853FCAFCF4FD82FC47.taxon	discussion	Remarks. This species name is based on the Latin adjective discoideus (discoid), referring to the discal color-pattern on head, pronotum, and elytra. Oopterus discoideus is morphologically close to O. marrisi. In addition to diagnostic characters of the male genitalia, O. discoideus has the following distinguishing features: body length 3.4 – 4.2 mm; head, pronotum, and elytra mostly testaceous; eyes subdepressed; tempora moderately inflated (about half as long as eyes); pronotum strongly convex, sides strongly rounded, not sinuate posteriorly, laterobasal foveae subrectangular. Both species are known only from the Fiordland region (South Island).	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30246FF863FCAFBD7FC7CFB07.taxon	description	Fig. 28, 75, 133 Holotype: male (NZAC) labeled: “ [NEW ZEALAND FD] Wilmot Pass. 701 m. (hand-written) / Manapouri Exp. Jan 70 (typed) / at night (typed) on moss. (hand-written) / [male symbol] / HOLOTYPE [male symbol] Oopterus marrisi Larochelle & Larivière, 2017 (red label; typed). ” Paratypes: two males (LUNZ, NZAC) from Murchison Mountains (FD), bearing blue paratype labels. Description. Body length 5.7 – 6.6 mm. Head, pronotum, elytra, and abdomen mostly reddish black; base of pronotum reddish medially; lateral margins, suture, and extreme apex of elytra reddish; abdomen bright reddish laterally and apically; antennae, palpi, and legs reddish; apical half of femora yellow. Microsculpture absent. Iridescence absent. Very shiny, with metallic luster (aeneous) on pronotum and elytra. Head. Labrum moderately transverse, slightly emarginate anteriorly. Antennae submoniliform, moderately long: segment 1 (scape) moderately long, about 2 x longer than its maximum width. Frontal furrows wide, deep, convergent. Eyes moderately convex; a single setiferous puncture on inner side of each eye (posteriorly). Tempora not inflated. Mentum: medial tooth entire, acute apically, moderately shorter than lateral lobes. Paraglossae membranous, prominent, slightly longer than ligula. Thorax. Pronotum moderately convex, finely punctate between laterobasal foveae, wrinkled discally, strongly transverse, widest about middle; apex subtruncate; anterolateral angles poorly developed, rounded; anterior bead incomplete, obsolete medially; sides moderately rounded, slightly sinuate posteriorly; lateral grooves absent; setiferous punctures absent on each side; posterolateral angles subrectangular; posterolateral carinae obsolete; laterobasal foveae well defined, virtually impunctate, very deep and wide, oblong, prolonged forward, double; posterior bead absent; base emarginate, wider than pronotal apex, about as wide as elytral base. Legs. Moderately long. Elytra. Strongly convex, ovate, widest about middle. Basal margin complete, reaching about scutellum. Shoulders obtuse. Sides strongly rounded. Scutellar striole present. Striae 1 – 4 complete; striae 5 – 7 incomplete; stria 1 deep; striae 2 – 4 shallow, finely punctate; striae 5 – 7 obsolete; stria 3 with three setiferous punctures. Recurrent stria long, sharp, directed apically toward stria 5. Subapical seta present. Intervals subdepressed. Sutural apices rounded. Abdomen. Last visible sternum (sternum VII): male with two apical ambulatory setae; female with four apical ambulatory setae. Aedeagus. Lateral view (Fig. 75): moderately arcuate, widened in apical half; base slightly convex dorsally; middle slightly convex dorsally, strongly convex ventrally, with dorsal membranous area moderately wide and very long; apex subtriangular, slightly convex dorsally, straight ventrally, with extreme tip narrow and rather long. Dorsal view: narrow, asymmetrical (ostium of membranous area deflected to right); apex straight; basal orifice narrow, closed anteriorly, moderately distant from membranous area. Parameres with five to seven apical setae.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30246FF863FCAFBD7FC7CFB07.taxon	materials_examined	Material examined. 6 specimens (LUNZ, NZAC). Geographic distribution (Fig. 133). South Island: FD – Fiordland National Park (Borland Valley Bivouac (South of); McKenzie Burn; Murchison Mountains; Wilmot Pass).	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30246FF863FCAFBD7FC7CFB07.taxon	biology_ecology	Ecology. Montane. Endogean. Wet beech forests. Shaded ground. Nocturnal; active at night on moss; hides during the day in thick moss, leaf litter, and under bark. Biology. Seasonality: November – January. Tenerals: November. Predacious (based on mouthpart morphology). Dispersal power. Subapterous (incapable of flight). Moderate runner. Collecting techniques. Sifting moss and leaf litter; lifting bark.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30246FF863FCAFBD7FC7CFB07.taxon	discussion	Remarks. This species is named after our friend and colleague John M. W. Marris (Lincoln University, Lincoln, Christchurch) for his special help and encouragement in our entomological studies. Oopterus marrisi is morphologically close to O. discoideus. In addition to diagnostic characters of the male genitalia, O. marrisi has the following distinguishing features: body length 5.7 – 6.6 mm; head, pronotum, and elytra mostly reddish black; eyes moderately convex; tempora not inflated; pronotum moderately convex, sides moderately rounded, slightly sinuate posteriorly, laterobasal foveae oblong. Both species are known only from the Fiordland region (South Island).	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30245FF873FCAFB17FD58FA87.taxon	description	Fig. 29, 76, 131	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30245FF873FCAFB17FD58FA87.taxon	description	Description. Body length 4.1 – 4.9 mm. Head, pronotum, elytra, and abdomen mostly piceous black; apex and base of head reddish; base of pronotum reddish; lateral margins, suture, and apex of elytra reddish; abdomen reddish laterally and apically; antennae, palpi, and femora pale yellow; femora (base), tibiae, and tarsi reddish. Microsculpture absent. Iridescence absent. Very shiny, with metallic luster (aeneous) on head, pronotum, and elytra. Head. Labrum strongly transverse, slightly emarginate anteriorly. Antennae submoniliform, moderately long: segment 1 (scape) moderately long, about 2 x longer than its maximum width. Frontal furrows wide, deep, convergent. Eyes moderately convex; a single setiferous puncture on inner side of each eye (posteriorly). Tempora not inflated. Mentum: medial tooth entire, acute apically, moderately shorter than lateral lobes. Paraglossae membranous, prominent, about as long as ligula. Thorax. Pronotum strongly convex, obsoletely punctate across base, unwrinkled, strongly transverse, slightly cordate, widest about middle; apex subtruncate; anterolateral angles poorly developed, obtusely rounded; anterior bead incomplete, obsolete medially; sides strongly rounded, slightly sinuate posteriorly; lateral grooves absent; setiferous punctures absent on each side; posterolateral angles obtuse; posterolateral carinae blunt, very long (about as long as adjoining foveae); laterobasal foveae well defined, obsoletely punctate, very deep and wide, subrectangular, not prolonged forward, double; posterior bead absent; base subtruncate, wider than pronotal apex, about as wide as elytral base. Legs. Moderately long. Elytra. Strongly convex, ovate, widest about middle. Basal margin incomplete, reaching about stria 4. Shoulders obtuse. Sides strongly rounded. Scutellar striole present. Striae mostly replaced by rows of coarse punctures; stria 1 complete, very deep; rows 2 – 5 moderately deep, obsolete apically; rows 6 – 7 obsolete basally and apically; interval 3 with three setiferous punctures. Recurrent stria long, sharp, directed apically toward row 5. Subapical seta present. Intervals slightly convex. Sutural apices obtuse. Abdomen. Last visible sternum (sternum VII): male with two apical ambulatory setae; female with four apical ambulatory setae. Aedeagus. Lateral view (Fig. 76): moderately arcuate, widest medially, narrowed in apical half; base moderately biconvex dorsally; middle strongly convex dorsally, strongly concave ventrally, with dorsal membranous area moderately wide and very long; apex triangular, slightly concave dorsally, straight ventrally, with extreme tip wide and long. Dorsal view: narrow, asymmetrical (ostium of membranous area deflected to right); apex straight; basal orifice narrow, closed anteriorly, moderately distant from membranous area. Parameres with five apical setae.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30245FF873FCAFB17FD58FA87.taxon	materials_examined	Material examined. 52 specimens (BMNH, CMNZ, JNNZ, LUNZ, NZAC). Geographic distribution (Fig. 131). South Island: BR, WD.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30245FF873FCAFB17FD58FA87.taxon	biology_ecology	Ecology. Lowland, montane, subalpine. Endogean. Wet forests (podocarp, broadleaf, beech) and scrublands. Shaded ground. Nocturnal; hides during the day under fallen branches, logs, well-embedded stones, and in moss. Gregarious. Biology. Seasonality: October – May. Tenerals: March (WD). Predacious (based on mouthpart morphology). Dispersal power. Subapterous (incapable of flight). Moderate runner. Collecting techniques. Lifting branches, logs and stones; sifting moss; pitfall trapping.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30244FF803FCAFA97FD58FA67.taxon	description	Fig. 30, 77, 140	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30244FF803FCAFA97FD58FA67.taxon	description	(hand-written). ” Oöpterus patulus: Broun 1886: 755. Tarastethus patulus: Sharp 1886: 373. Pseudoopterus patulus: Csiki 1933: 1651. Oopterus patulus: Johns 1974: 301. Description. Body length 5.4 – 6.2 mm. Head, pronotum, and elytra mostly piceous black; apex and base of head reddish; lateral margins, suture, and apex of elytra reddish; abdomen smoky brown, yellow laterally and apically; antennae, palpi, and legs reddish; femora pale yellow. Microsculpture strong, isodiametric on head, moderately transverse on pronotum and elytra. Iridescence absent. Very shiny, with metallic luster (aeneous) on elytra. Head. Labrum moderately transverse, slightly emarginate anteriorly. Antennae submoniliform, moderately long: segment 1 (scape) short, stout, about 1.5 x longer than its maximum width. Frontal furrows wide, shallow, convergent. Eyes moderately convex; a single setiferous puncture on inner side of each eye (posteriorly). Tempora not inflated. Mentum: medial tooth entire, acute apically, moderately shorter than lateral lobes. Paraglossae membranous, prominent, about as long as ligula. Thorax. Pronotum moderately convex, finely punctate across base, wrinkled across apex and base, moderately transverse and cordate, widest about middle; apex truncate; anterolateral angles poorly developed, rounded; anterior bead incomplete, obsolete medially; sides strongly rounded, not sinuate posteriorly; lateral grooves very long (as long as pronotum), moderately deep; setiferous punctures absent on each side; posterolateral angles obtuse; posterolateral carinae blunt, very long (about as long as adjoining foveae); laterobasal foveae well defined, finely punctate, shallow, very wide, subrectangular, somewhat prolonged forward, double; posterior bead absent; base emarginate, about as wide as pronotal apex, slightly narrower than elytral base. Legs. Moderately long. Elytra. Moderately convex, ovate, widest about middle. Basal margin incomplete, reaching about stria 4. Shoulders obtuse. Sides strongly rounded. Scutellar striole obsolete. Striae almost complete, very shallow (almost obsolete), virtually impunctate; interval 3 with three setiferous punctures. Recurrent stria long, sharp, directed apically toward stria 5. Subapical seta present. Intervals subdepressed. Sutural apices obtusely rounded. Abdomen. Last visible sternum (sternum VII): male with two apical ambulatory setae; female with four apical ambulatory setae. Aedeagus. Lateral view (Fig. 77): strongly arcuate, moderately widened in apical half; base slightly convex dorsally; middle slightly concave dorsally, moderately convex ventrally, with dorsal membranous area narrow and very long; apex subtriangular, moderately convex dorsally, moderately concave ventrally, with extreme tip wide, short, and curved downward. Dorsal view: narrow, symmetrical (ostium of membranous area dorsal); apex deflected to left; basal orifice narrow, closed anteriorly, close to membranous area. Parameres with three or four apical setae.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30244FF803FCAFA97FD58FA67.taxon	materials_examined	Material examined. 390 specimens (AMNZ, BMNH, CMNZ, JNNZ, MNHN, MONZ, NZAC). Geographic distribution (Fig. 140). South Island: CO, DN, FD, OL, SC, SL. Stewart Island.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30244FF803FCAFA97FD58FA67.taxon	biology_ecology	Ecology. Lowland, montane, subalpine. Epigean-arboreal. Wet forests (beech, broadleaf). Shaded ground. Nocturnal; active at night on trees and moss; hides during the day under the loose bark of fallen trees (beech) and live trees (Fuchsia), in fallen rotten branches, under logs and stones. Gregarious. Biology. Seasonality: throughout the year, except June. Tenerals: January – March. Predacious (based on mouthpart morphology). Defense mechanism: drops from trees when disturbed. Occasionally infested by fungi (Laboulbeniales). Dispersal power. Subapterous (incapable of flight). Moderate runner. Frequent climber on trees. Collecting techniques. Lifting loose bark of trees; breaking fallen branches; lifting logs and stones.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30243FF813FCAF9B4FD58F927.taxon	description	Fig. 31, 78, 127	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30243FF813FCAF9B4FD58F927.taxon	description	Description. Body length 4.6 – 5.0 mm. Head, pronotum, elytra, and abdomen mostly piceous black; base of head and pronotum reddish; base, lateral margins, suture, and apex of elytra reddish; abdomen yellow laterally and apically; antennae, palpi, and legs pale red; femora pale yellow. Microsculpture mostly absent on head (strong and isodiametric basally), absent on pronotum, strong and very transverse (with microlines) on elytra. Iridescence absent on head and pronotum, very strong on elytra. Very shiny, with metallic luster (aeneous) on elytra. Head. Labrum strongly transverse, slightly emarginate anteriorly. Antennae submoniliform, moderately long: segment 1 (scape) moderately long, about 2 x longer than its maximum width. Frontal furrows wide, deep, convergent. Eyes moderately convex; a single setiferous puncture on inner side of each eye (posteriorly). Tempora not inflated. Mentum: medial tooth entire, acute apically, moderately shorter than lateral lobes. Paraglossae membranous, prominent, about as long as ligula. Thorax. Pronotum moderately convex, coarsely punctate across base, wrinkled discally, strongly transverse, moderately cordate, widest before middle; apex subtruncate; anterolateral angles poorly developed, rounded; anterior bead incomplete, present near anterolateral angles; sides strongly rounded, moderately sinuate posteriorly; lateral grooves moderately to very deep, very long (as long as pronotum); setiferous punctures absent on each side; posterolateral angles rectangular; posterolateral carinae sharp, very long (about as long as adjoining foveae); laterobasal foveae well defined, coarsely punctate, very deep, moderately wide, square, not prolonged forward, simple; posterior bead absent; base emarginate, about as wide as pronotal apex and elytral base. Legs. Moderately long. Elytra. Moderately convex, subovate, widest about middle. Basal margin incomplete, reaching about stria 4. Shoulders moderately rounded. Sides moderately rounded. Scutellar striole obsolete. Striae incomplete, mostly obsolete; stria 1 deep; striae 2 – 4 shallow, virtually impunctate; interval 3 with three setiferous punctures. Recurrent stria long, sharp, directed apically toward stria 5. Subapical seta present. Intervals subdepressed. Sutural apices obtusely rounded. Abdomen. Last visible sternum (sternum VII): male with two apical ambulatory setae; female with four apical ambulatory setae. Aedeagus. Lateral view (Fig. 78): strongly arcuate, moderately widened in apical half; base strongly convex dorsally; middle moderately convex dorsally, rather straight ventrally, with dorsal membranous area moderately wide and rather long; apex subtriangular, subangulate dorsally, slightly concave ventrally, with extreme tip wide and short. Dorsal view: narrow, symmetrical (ostium of membranous area dorsal); apex straight; basal orifice narrow, closed anteriorly, close to membranous area. Parameres with three or four apical setae.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30243FF813FCAF9B4FD58F927.taxon	materials_examined	Material examined. 77 specimens (AMNZ, BMNH, JNNZ, LUNZ, NZAC). Geographic distribution (Fig. 127). North Island: BP, HB, ND, TK, TO, WA, WN.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30243FF813FCAF9B4FD58F927.taxon	biology_ecology	Ecology. Lowland, montane, subalpine. Epigean-arboreal. Wet forests (beech, broadleaf, podocarp). Shaded ground. Nocturnal; active at night on tree trunks; hides during the day under the loose bark of fallen trees, in fallen rotten branches and logs. Gregarious. Biology. Seasonality: throughout the year, except October. Tenerals: December – March. Predacious (based on mouthpart morphology). Dispersal power. Subapterous (incapable of flight). Moderate runner. Occasional climber on trees. Collecting techniques. Lifting loose bark of fallen trees; breaking rotten branches and logs; fogging logs; pitfall trapping.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30241FF833FCAFF34FE0BFD24.taxon	description	Fig. 32, 79, 144	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30241FF833FCAFF34FE0BFD24.taxon	description	Description. Body length 5.1 – 6.0 mm. Head, pronotum, elytra, and abdomen mostly black; lateral margins, suture, and apex of elytra reddish; abdomen yellow laterally and apically; antennae and legs pale red; palpi and femora pale yellow. Microsculpture strong and isodiametric on head, very transverse (with microlines), obsolete on pronotum, feeble on elytra. Iridescence absent on head and pronotum, feeble on elytra. Very shiny, with metallic luster (aeneous) on elytra. Head. Labrum strongly transverse, slightly emarginate anteriorly. Antennae submoniliform, moderately long: segment 1 (scape) short, stout, about 1.5 x longer than its maximum width. Frontal furrows wide, shallow, convergent. Eyes moderately convex; a single setiferous puncture on inner side of each eye (posteriorly). Tempora not inflated. Mentum: medial tooth entire, acute apically, moderately shorter than lateral lobes. Paraglossae membranous, prominent, about as long as ligula. Thorax. Pronotum moderately convex, impunctate, wrinkled discally, slightly transverse, widest about middle; apex truncate; anterolateral angles poorly developed, obtusely rounded; anterior bead complete; sides moderately rounded, not sinuate posteriorly; lateral grooves moderately deep, very long (about as long as adjoining foveae); setiferous punctures absent on each side; posterolateral angles obtuse; posterolateral carinae sharp, very long (about as long as adjoining foveae); laterobasal foveae well defined, impunctate, moderately deep and wide, square, not prolonged forward, double; posterior bead absent; base emarginate, about as wide as pronotal apex and elytral base. Legs. Moderately long. Elytra. Moderately convex, subovate, widest about middle. Basal margin incomplete, reaching about stria 4. Shoulders obtuse. Sides moderately rounded. Scutellar striole absent. Striae incomplete, mostly obsolete, virtually impunctate; stria 1 deep; striae 2 – 4 shallow; interval 3 with three setiferous punctures. Recurrent stria long, sharp, directed apically toward stria 5. Subapical seta present. Intervals subdepressed. Sutural apices obtusely rounded. Abdomen. Last visible sternum (sternum VII): male with two apical ambulatory setae; female with four apical ambulatory setae. Aedeagus. Lateral view (Fig. 79): strongly arcuate, narrowed in apical half; base slightly convex dorsally; middle slightly convex dorsally, moderately convex ventrally, with dorsal membranous area invisible; apex elongate-rounded, slightly concave dorsally and ventrally, with extreme tip unusually wide and short. Dorsal view: narrow, symmetrical (ostium of membranous area dorsal); apex straight; basal orifice narrow, closed anteriorly, close to membranous area. Parameres with three apical setae.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30241FF833FCAFF34FE0BFD24.taxon	materials_examined	Material examined. 52 specimens (BMNH, JNNZ, LUNZ, MONZ, NZAC). Geographic distribution (Fig. 144). North Island: HB, ND, RI, TK, WA, WI, WN.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30241FF833FCAFF34FE0BFD24.taxon	biology_ecology	Ecology. Lowland, montane, subalpine. Epigean-arboreal. Wet forests (beech, broadleaf, podocarp). Shaded ground. Nocturnal; active on trees at night; hides during the day under the loose bark of fallen trees, in fallen rotten branches, in moss growing on logs and trees. Gregarious. Biology. Seasonality: November – June, August. Predacious (based on mouthpart morphology). Dispersal power. Subapterous (incapable of flight). Moderate runner. Regular climber on trees and logs. Collecting techniques. Lifting loose bark of fallen trees; breaking rotten branches; sifting moss.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30241FF833FCAFF34FE0BFD24.taxon	discussion	Remark. Examination of the types of Oopterus nigritulus and O. laevigatus revealed them to be conspecific with O. sobrinus.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30240FFBC3FCAFCF4FD58FCA7.taxon	description	Fig. 33, 80, 123	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30240FFBC3FCAFCF4FD58FCA7.taxon	description	Description. Body length 5.8 – 6.4 mm. Head, pronotum, elytra, and abdomen mostly dark brown; head darker medially; abdomen reddish laterally and apically; antennae, palpi, and legs reddish. Microsculpture obsolete and isodiametric on head, feeble and moderately transverse on pronotum, strong and very transverse (with microlines) on elytra. Iridescence absent on head and pronotum, strong on elytra. Very shiny, without metallic luster. Head. Labrum moderately transverse, slightly emarginate anteriorly. Antennae submoniliform, moderately long: segment 1 (scape) short, stout, about 1.5 x longer than its maximum width. Frontal furrows wide, deep, convergent. Eyes moderately convex; a single setiferous puncture on inner side of each eye (posteriorly). Tempora not inflated. Mentum: medial tooth entire, acute apically, moderately shorter than lateral lobes. Paraglossae membranous, prominent, about as long as ligula. Thorax. Pronotum moderately convex, coarsely punctate across base, wrinkled discally, subquadrate, widest about middle; apex truncate; anterolateral angles poorly developed, obtusely rounded; anterior bead incomplete, obsolete medially; sides moderately rounded, slightly sinuate posteriorly; lateral grooves moderately to very deep, very long (as long as pronotum); a single setiferous puncture on each side (posteriorly); posterolateral angles rectangular; posterolateral carinae sharp, very long (about as long as adjoining foveae); laterobasal foveae well defined, coarsely punctate, moderately deep and wide, oblong, prolonged forward, simple; posterior bead absent; base emarginate, about as wide as pronotal apex and elytral base. Legs. Moderately long. Elytra. Moderately convex, subovate, widest about middle. Basal margin incomplete, reaching about stria 4. Shoulders obtuse. Sides moderately rounded. Scutellar striole present. Striae incomplete; striae 1 – 3 shallow, finely punctate; striae 4 – 7 obsolete, impunctate; stria 3 with three setiferous punctures. Recurrent stria long, sharp, directed apically toward stria 6. Subapical seta present. Intervals subdepressed. Sutural apices obtusely rounded. Abdomen. Last visible sternum (sternum VII): male with two apical ambulatory setae; female with four apical ambulatory setae. Aedeagus. Lateral view (Fig. 80): strongly arcuate, not widened in apical half; base moderately biconvex dorsally; middle subparallel, slightly convex dorsally, moderately convex ventrally, with dorsal membranous area moderately wide and very long; apex triangular, slightly concave dorsally, straight ventrally, with extreme tip wide and long. Dorsal view: narrow, asymmetrical (ostium of membranous area deflected to right); apex straight; basal orifice narrow, closed anteriorly, close to membranous area. Parameres with three apical setae.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30240FFBC3FCAFCF4FD58FCA7.taxon	materials_examined	Material examined. 73 specimens (BMNH, JNNZ, LUNZ, NZAC). Geographic distribution (Fig. 123). South Island: BR, NN.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30240FFBC3FCAFCF4FD58FCA7.taxon	biology_ecology	Ecology. Lowland, montane, subalpine. Epigean-arboreal. Wet forests (beech). Shaded ground. Nocturnal; active at night on trees and logs; hides during the day under loose bark of trees, and in moss growing on logs and trees. Gregarious. Biology. Seasonality: October – May. Tenerals: December (NN). Predacious (based on mouthpart morphology). Dispersal power. Subapterous (incapable of flight). Moderate runner. Regular climber on trees and logs. Collecting techniques. Lifting loose bark of trees; sifting moss.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3027FFFBD3FCAFC77FD58FC27.taxon	description	Fig. 34, 81, 130	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3027FFFBD3FCAFC77FD58FC27.taxon	description	Description. Body length 4.1 – 5.3 mm. Head, pronotum, elytra, and abdomen mostly piceous black; apex of head reddish; lateral margins, suture, and apex of elytra reddish; abdomen reddish laterally and apically; antennae, palpi, and legs pale yellow; base of femora piceous black. Microsculpture absent. Iridescence absent. Very shiny, without metallic luster. Head. Labrum strongly transverse, slightly emarginate anteriorly. Antennae submoniliform, moderately long: segment 1 (scape) very long, about 2.5 x longer than its maximum width. Frontal furrows wide, deep, convergent. Eyes moderately convex; two setiferous punctures on inner side of each eye. Tempora not inflated. Mentum: medial tooth entire, acute apically, moderately shorter than lateral lobes. Paraglossae membranous, prominent, about as long as ligula. Thorax. Pronotum strongly convex, coarsely punctate across base, unwrinkled, strongly transverse, slightly cordate, widest about middle; apex subtruncate; anterolateral angles poorly developed, obtusely rounded; anterior bead incomplete, obsolete medially; sides strongly rounded, slightly sinuate posteriorly; lateral grooves absent; a single setiferous puncture on each side (anteriorly); posterolateral angles obtuse; posterolateral carinae blunt, very long (about as long as adjoining foveae); laterobasal foveae well defined, coarsely punctate, very deep and wide, subrectangular, not prolonged forward, double; posterior bead absent; base emarginate, wider than pronotal apex, about as wide as elytral base. Legs. Moderately long. Elytra. Strongly convex, ovate, widest about middle. Basal margin incomplete, reaching about stria 4. Shoulders rounded. Sides strongly rounded. Scutellar striole present. Striae mostly replaced by rows of coarse punctures; stria 1 complete, very deep; rows 2 – 5 moderately deep, obsolete apically; rows 6 – 7 obsolete basally; interval 3 with three setiferous punctures, close to row 3. Recurrent stria long, sharp, directed apically toward row 5. Subapical seta present. Intervals slightly convex. Sutural apices obtuse. Abdomen. Last visible sternum (sternum VII): male with two apical ambulatory setae; female with four apical ambulatory setae. Aedeagus. Lateral view (Fig. 81): moderately arcuate, moderately widened in apical half; base strongly biconvex dorsally; middle slightly concave dorsally, mostly straight ventrally, with dorsal membranous area very wide and moderately long; apex subtriangular, slightly concave dorsally, straight ventrally, with extreme tip wide and long. Dorsal view: narrow, asymmetrical (ostium of membranous area deflected to right); apex straight; basal orifice narrow, closed anteriorly, moderately distant from membranous area. Parameres with five or six apical setae.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3027FFFBD3FCAFC77FD58FC27.taxon	materials_examined	Material examined. 157 specimens (AMNZ, BMNH, CMNZ, JNNZ, LUNZ, MONZ, NZAC). Geographic distribution (Fig. 130). South Island: BR, MC, NN, WD.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3027FFFBD3FCAFC77FD58FC27.taxon	biology_ecology	Ecology. Lowland, montane, subalpine. Epigean-arboreal. Wet forests (beech, podocarp, broadleaf) and scrublands. Shaded ground. Nocturnal; active at night on mossy logs; hides during the day in moss and leaf litter, as well as under logs. Biology. Seasonality: September, November – June. Tenerals: February – March (BR). Predacious (based on mouthpart morphology). Dispersal power. Subapterous (incapable of flight). Moderate runner. Occasional climber on logs. Collecting techniques. Pitfall trapping; sifting moss and leaf litter; lifting logs.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3027EFFBE3FCAFBF4FAFCFCC7.taxon	description	Fig. 35, 121 Holotype: female (MONZ) labeled: “ [NEW ZEALAND NN] Mt. Balloon [= Balloon Hut, Mount Arthur] IN CAVE J. T. Salmon 24. I. 48 (hand-written) / [female symbol] / HOLOTYPE [female symbol] Oopterus arthurensis Larochelle & Larivière, 2017 (red label; typed). ” Description. Body length 7.9 mm. Head, pronotum, elytra, and abdomen mostly rufous; margins and suture of elytra red; antennae, palpi, and legs testaceous. Microsculpture strong, isodiametric on head, very transverse (with microlines) on pronotum and elytra. Iridescence absent on head, very strong on pronotum and elytra. Very shiny, without metallic luster. Forebody narrow in comparison to elytra (as in O. trechoides). Head. Labrum strongly transverse, slightly emarginate anteriorly. Antennae subfiliform (as in O. ocularius, O. trechoides, and Zolus species), very long: segment 1 (scape) very long, about 2.5 x longer than its maximum width. Frontal furrows wide, deep, subparallel. Eyes slightly convex; two setiferous punctures on inner side of each eye. Tempora strongly inflated (about as long as eyes). Mentum: medial tooth entire, acute apically, moderately shorter than lateral lobes. Paraglossae membranous, prominent, slightly longer than ligula. Thorax. Pronotum slightly convex, finely punctate across base, wrinkled in basal half, subquadrate, widest before middle; apex subtruncate; anterolateral angles well developed, obtusely rounded; anterior bead incomplete, obsolete medially; sides moderately rounded, moderately sinuate posteriorly; lateral grooves absent; a single setiferous puncture on each side (anteriorly); posterolateral angles subrectangular; posterolateral carinae absent; laterobasal foveae well defined, finely punctate, very deep and wide, oblong, prolonged forward, simple; posterior bead absent; base emarginate, about as wide as pronotal apex, much narrower than elytral base. Legs. Very long. Elytra. Moderately convex, subovate, widest about middle. Basal margin incomplete, reaching about stria 4. Shoulders oblique. Sides strongly rounded. Scutellar striole present. Stria 1 complete, deep; striae 2 – 7 incomplete, obsolete; striae 1 – 2 finely punctate; stria 3 with a single setiferous puncture (anteriorly). Recurrent stria long, sharp, directed apically toward stria 6. Subapical seta present. Intervals subdepressed. Sutural apices obtusely rounded. Abdomen. Last visible sternum (sternum VII): male unknown; female with four apical ambulatory setae. Aedeagus. Male unknown.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3027EFFBE3FCAFBF4FAFCFCC7.taxon	materials_examined	Material examined. 1 specimen (MONZ). Geographic distribution (Fig. 121). South Island: NN – Mount Arthur, Balloon Hut.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3027EFFBE3FCAFBF4FAFCFCC7.taxon	biology_ecology	Ecology. Subalpine. A beech forest and red tussock area. Found in a cave. Biology. Seasonality: January. Predacious (based on mouthpart morphology). Dispersal power. Subapterous (incapable of flight). Fast runner (based on leg morphology).	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3027EFFBE3FCAFBF4FAFCFCC7.taxon	discussion	Remarks. This species is named after the type locality Mount Arthur (NN) and the Latin suffix - ensis, denoting a place, locality, or country. Oopterus arthurensis is morphologically close to O. corvinki and has the following distinguishing features: body length 7.9 mm; forebody narrow in comparison to elytra; antennae subfiliform, very long; tempora strongly inflated (about as long as eyes); pronotum subquadrate, not cordate, posterolateral angles subrectangular; elytral striae mostly obsolete. The two species are allopatric: O. arthurensis is restricted to the northwestern South Island (NN, Mount Arthur), while O. corvinki occurs in the southwest of the South Island (FD, Gilbert Islands). O. arthurensis, with its darkly colored body and normally developed eyes, is probably not a regular cave-dweller.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3027DFFBF3FCAFC54FD67FB87.taxon	description	Fig. 36, 82, 124 Holotype: male (NZAC) labeled: “ NEW ZEALAND FD (typed) Breaksea So Gilbert Is No 6 May 1982 (hand-written) / C. F. Butcher ex litter r 2 (hand-written) / [male symbol] / HOLOTYPE [male symbol] Oopterus corvinki Larochelle & Larivière, 2017 (red label; typed). ” Paratypes: two males (LUNZ, NZAC) from the same locality as the holotype, bearing blue paratype labels. Description. Body length 4.3 – 4.6 mm. Head reddish, darker medially; pronotum and elytra dark red; abdomen mostly yellow, black anterobasally; lateral margins and a large apical space on each elytron pale yellow; antennae, palpi, and legs reddish; antennal segment 1 (scape) and femora pale yellow. Microsculpture obsolete and isodiametric on head, strong and very transverse (with microlines) on pronotum and elytra. Iridescence absent on head, very strong on pronotum and elytra. Very shiny, with metallic luster (aeneous) on pronotum and elytra. Head. Labrum strongly transverse, slightly emarginate anteriorly. Antennae submoniliform, moderately long: segment 1 (scape) moderately long, about 2 x longer than its maximum width. Frontal furrows wide, shallow, convergent. Eyes slightly convex; two setiferous punctures on inner side of each eye. Tempora not inflated. Mentum: medial tooth entire, acute apically, moderately shorter than lateral lobes. Paraglossae membranous, prominent, about as long as ligula. Thorax. Pronotum moderately convex, impunctate, wrinkled discally, strongly transverse, moderately cordate, widest before middle; apex truncate; anterolateral angles poorly developed, rounded; anterior bead incomplete, obsolete medially; sides strongly rounded, moderately sinuate posteriorly; lateral grooves absent; a single setiferous puncture on each side (anteriorly); posterolateral angles obtuse; posterolateral carinae blunt, very long (about as long as adjoining foveae); laterobasal foveae well defined, impunctate, shallow, very wide, subrectangular, prolonged forward, double; posterior bead absent; base emarginate, about as wide as pronotal apex and elytral base. Legs. Moderately long. Elytra. Strongly convex, ovate, widest about middle. Basal margin incomplete, reaching about stria 4. Shoulders obtuse. Sides strongly rounded. Scutellar striole present. Stria 1 complete; striae 4 – 7 incomplete; striae 1 – 4 moderately deep; striae 5 – 7 shallow; striae 1 – 7 finely punctate; stria 3 with three setiferous punctures. Recurrent stria long, sharp, directed apically toward striae 5 or 6. Subapical seta present. Intervals slightly convex. Sutural apices obtuse. Abdomen. Last visible sternum (sternum VII): male with two apical ambulatory setae; female with four apical ambulatory setae. Aedeagus. Lateral view (Fig. 82): strongly arcuate, narrowed in apical half; base slightly convex dorsally; middle angulate basally and somewhat straight dorsally, moderately convex ventrally, with dorsal membranous area moderately wide and very long; apex subtriangular, straight dorsally, slightly concave ventrally, with extreme tip moderately wide and long. Dorsal view: narrow, asymmetrical (ostium of membranous area deflected to right); apex straight; basal orifice narrow, closed anteriorly, moderately distant from membranous area. Parameres with five apical setae.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3027DFFBF3FCAFC54FD67FB87.taxon	materials_examined	Material examined. 3 specimens (LUNZ, NZAC). Geographic distribution (Fig. 124). South Island: FD – Breaksea Sound, Gilbert Islands.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3027DFFBF3FCAFC54FD67FB87.taxon	biology_ecology	Ecology. Lowland. Wet forest (beech). Found in leaf litter. Biology. Seasonality: May. Predacious (based on mouthpart morphology). Dispersal power. Subapterous (incapable of flight). Moderate runner. Collecting technique. Sifting leaf litter.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3027DFFBF3FCAFC54FD67FB87.taxon	discussion	Remarks. This species is named after our friend and colleague Cor J. Vink (Canterbury Museum, Christchurch) for his special help and encouragement in our entomological studies. Oopterus corvinki is morphologically close to O. arthurensis. In addition to diagnostic characters of the male genitalia, O. corvinki has the following distinguishing features: body length 4.3 – 4.6 mm; antennae submoniliform, moderately long; tempora not inflated; pronotum strongly transverse, moderately cordate, posterolateral angles obtuse; elytral striae mostly visible. The two species are allopatric: O. corvinki is restricted to the southwestern South Island (FD, Gilbert Islands), while O. arthurensis occurs in the northwest of the South Island (NN, Mount Arthur).	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3027CFFB83FCAFB94FCA6FA87.taxon	description	Fig. 37, 83, 139 Holotype: male (NZAC) labeled: “ New Zealand MK (typed) Temple Stream Ohau Valley 11 April 2010 (typed) / Under log, mountain beech forest (typed) / JOHN NUNN (typed) / [male symbol] / HOLOTYPE [male symbol] Oopterus palmai Larochelle & Larivière, 2017 (red label; typed). ” Paratype: one female (NZAC) from the same locality as the holotype, bearing a blue paratype label. Description. Body length 5.3 – 5.7 mm. Head, pronotum, elytra, and abdomen mostly piceous black; apex and base of head reddish; base of pronotum reddish; lateral margins and suture of elytra reddish; abdomen yellowish laterally and apically; antennae, palpi, and legs reddish; antennal segment 1 (scape) pale yellowish; femora piceous black. Microsculpture strong and isodiametric on head, obsolete and moderately transverse on pronotum, strong and very transverse (with microlines) on elytra. Iridescence absent on head and pronotum, very strong on elytra. Very shiny, without metallic luster. Head. Labrum moderately transverse, subtruncate anteriorly. Antennae submoniliform, moderately long: segment 1 (scape) moderately long, about 2 x longer than its maximum width. Frontal furrows wide, deep, subparallel. Eyes moderately convex; two setiferous punctures on inner side of each eye. Tempora not inflated. Mentum: medial tooth entire, acute apically, moderately shorter than lateral lobes. Paraglossae membranous, prominent, about as long as ligula. Thorax. Pronotum moderately convex, impunctate, wrinkled apically and discally, strongly transverse, widest about middle; apex subtruncate; anterolateral angles poorly developed, rounded; anterior bead incomplete, obsolete medially; sides moderately rounded, not sinuate posteriorly; lateral grooves absent; a single setiferous puncture on each side (posteriorly); posterolateral angles subrectangular; posterolateral carinae absent; laterobasal foveae ill-defined, impunctate, shallow, moderately wide, not prolonged forward, double; base emarginate, about as wide as pronotal apex and elytral base. Legs. Moderately long. Elytra. Moderately convex, ovate, widest about middle. Basal margin incomplete, reaching about stria 4. Shoulders obtuse. Sides strongly rounded. Scutellar striole present. Striae 1 – 4 complete, shallow, finely punctate; striae 5 – 7 incomplete, obsolete. Interval 3 with three setiferous punctures. Recurrent stria long, sharp, directed apically toward stria 5. Subapical seta present. Intervals depressed. Sutural apices rounded. Abdomen. Last visible sternum (sternum VII): male with two apical ambulatory setae; female with four apical ambulatory setae. Aedeagus. Lateral view (Fig. 83): strongly arcuate, narrowed in apical half; base strongly biconvex dorsally; middle narrowed from base to apex, moderately convex dorsally, almost straight ventrally, with dorsal membranous area very wide and moderately long; apex triangular, slightly concave dorsally and ventrally, with extreme tip narrow and long. Dorsal view: narrow, asymmetrical (ostium of membranous area deflected to right); apex straight; basal orifice narrow, closed anteriorly, moderately distant from membranous area. Parameres with seven apical setae.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3027CFFB83FCAFB94FCA6FA87.taxon	materials_examined	Material examined. 24 specimens (JNNZ, NZAC). Geographic distribution (Fig. 139). South Island: CO – Rock and Pillar Range, Six-Mile Creek. MK – Lake Ohau. Ohau Range, Ohau Ski Field. Ohau Valley, Temple Stream.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3027CFFB83FCAFB94FCA6FA87.taxon	biology_ecology	Ecology. Montane, subalpine, alpine. Epigean. Dry forests (beech); fellfields. Shaded or open ground. Nocturnal; hides during the day under logs and stones. Gregarious. Biology. Seasonality: December, March – April. Predacious (based on mouthpart morphology). Dispersal power. Subapterous (incapable of flight). Moderate runner. Collecting techniques. Lifting logs and stones.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3027CFFB83FCAFB94FCA6FA87.taxon	discussion	Remarks. This species is named after our friend and colleague Ricardo L. Palma (Museum of New Zealand Te Papa Tongarewa, Wellington) for his special help and encouragement in our entomological studies, and for his many years (1991 – 2016) as curator of entomology at the Museum of New Zealand. Oopterus palmai is morphologically close to O. monticola. In addition to diagnostic characters of the male genitalia, O. palmai has the following distinguishing features: pronotum with sides moderately rounded, not sinuate posteriorly, posterolateral carinae absent, laterobasal foveae ill-defined; elytral striae 5 – 7 poorly developed, incomplete. Both species inhabit southern areas of the South Island: O. palmai occurs in the Central Otago (CO) and Mackenzie (MK) regions, while O. monticola is known only from Central Otago (CO, Rock and Pillar Range).	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3027BFFB93FCAFA94FB6CFA67.taxon	description	Fig. 38, 84, 135 Holotype: male (NZAC) labeled: “ [NEW ZEALAND CO] Rock & Pillar Ra Otago 13. IX. 69 J. S. Dugdale (typed) / moss (typed) 69 / 203 (hand-written) / [male symbol] / HOLOTYPE [male symbol] Oopterus monticola Larochelle & Larivière, 2017 (red label; typed). ” Paratypes: one female (NZAC) and one male (CMNZ) from the same locality as the holotype, bearing blue paratype labels. Description. Body length 4.8 – 5.2 mm. Head, pronotum, and elytra mostly piceous black; abdomen mostly dark brown, yellow laterally and apically; apex of head reddish; lateral margins and suture of elytra reddish; antennae and legs mostly reddish; antennal segments 1 + 2 pale yellow; palpi mostly yellow, strongly infuscated; femora dark brown. Microsculpture strong and isodiametric on head, feeble and moderately transverse on pronotum, obsolete and very transverse (with microlines) on elytra. Iridescence absent on head and pronotum, very strong on elytra. Very shiny, without metallic luster. Head. Labrum strongly transverse, slightly emarginate anteriorly. Antennae submoniliform, moderately long: segment 1 (scape) moderately long, about 2 x longer than its maximum width. Frontal furrows wide, shallow, convergent. Eyes moderately convex; two setiferous punctures on inner side of each eye. Tempora not inflated. Mentum: medial tooth entire, acute apically, moderately shorter than lateral lobes. Paraglossae membranous, prominent, about as long as ligula. Thorax. Pronotum moderately convex, impunctate, wrinkled discally, strongly transverse, widest before middle; apex subtruncate; anterolateral angles poorly developed, rounded; anterior bead incomplete, obsolete medially; sides strongly rounded, slightly sinuate posteriorly; lateral grooves absent; a single setiferous puncture on each side (posteriorly); posterolateral angles subrectangular; posterolateral carinae blunt, very long (about as long as adjoining foveae); laterobasal foveae well defined, impunctate, shallow, moderately wide, oblong, prolonged forward, double; posterior bead absent; base subtruncate, about as wide as pronotal apex and elytral base. Legs. Moderately long. Elytra. Moderately convex, ovate, widest about middle. Basal margin incomplete, reaching about stria 4. Shoulders obtuse. Sides strongly rounded. Scutellar striole present. Striae complete, shallow, finely punctate; striae 5 – 6 shallower; interval 3 with three setiferous punctures. Recurrent stria long, sharp, directed apically toward stria 5. Subapical seta present. Intervals depressed. Sutural apices obtuse. Abdomen. Last visible sternum (sternum VII): male with two apical ambulatory setae; female with four apical ambulatory setae. Aedeagus. Lateral view (Fig. 84): strongly arcuate, strongly widened in apical half; base strongly biconvex dorsally; middle slightly concave dorsally, strongly convex ventrally, with dorsal membranous area very wide and long; apex triangular, slightly biconcave dorsally, moderately concave ventrally, with extreme tip wide and long. Dorsal view: narrow, asymmetrical (ostium of membranous area deflected to right); apex straight; basal orifice narrow, closed anteriorly, moderately distant from membranous area. Parameres with five or six apical setae.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3027BFFB93FCAFA94FB6CFA67.taxon	materials_examined	Material examined. 6 specimens (CMNZ, LUNZ, NZAC). Geographic distribution (Fig. 135). South Island: CO – Rock and Pillar Range (McPhees Rock; Otago Ski Hut).	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3027BFFB93FCAFA94FB6CFA67.taxon	biology_ecology	Ecology. Alpine. Epigean. Herbfields, along seepages and at the edge of a bog. Open ground. Nocturnal; hides during the day in moss. Biology. Seasonality: September, November. Predacious (based on mouthpart morphology). Dispersal power. Subapterous (incapable of flight). Moderate runner. Collecting technique. Sifting moss.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3027BFFB93FCAFA94FB6CFA67.taxon	discussion	Remarks. The name of this species is based on the Latin noun mons, montis (mountain) and the suffix - cola (dweller). Oopterus monticola is morphologically close to O. palmai. In addition to diagnostic characters of the male genitalia, O. monticola has the following distinguishing features: pronotum with sides strongly rounded, slightly sinuate posteriorly, posterolateral carinae blunt and very long, laterobasal foveae well defined, oblong; elytral striae 5 – 7 well developed, complete. Both species occur in the southern South Island: O. monticola is restricted to Central Otago (CO, Rock and Pillar Range), while O. palmai occurs in the Central Otago (CO) and Mackenzie (MK) regions.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3027AFFBA3FCAF9B7FD58FA27.taxon	description	Fig. 39, 85, 145	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3027AFFBA3FCAF9B7FD58FA27.taxon	description	Description. Body length 3.9 – 4.6 mm. Head, pronotum, elytra, and abdomen mostly reddish brown; lateral margins, suture, and apex of elytra reddish; abdomen reddish laterally and apically; antennae, palpi, and legs reddish; femora yellow. Microsculpture strong and isodiametric on head, strong and moderately transverse on pronotum, obsolete and very transverse (with microlines) on elytra. Iridescence absent on head and pronotum, feeble on elytra. Very shiny, without metallic luster. Head. Labrum strongly transverse, slightly emarginate anteriorly. Antennae submoniliform, short: segment 1 (scape) moderately long, about 2 x longer than its maximum width. Frontal furrows wide, shallow, convergent. Eyes slightly convex; a single setiferous puncture on inner side of each eye (posteriorly). Tempora slightly inflated (about one third as long as eyes). Mentum: medial tooth entire, acute apically, moderately shorter than lateral lobes. Paraglossae membranous, prominent, about as long as ligula. Thorax. Pronotum moderately convex, impunctate, unwrinkled, subquadrate, widest before middle; apex subtruncate; anterolateral angles poorly developed, obtusely rounded; anterior bead incomplete, obsolete medially; sides strongly rounded, slightly sinuate or not sinuate posteriorly; lateral grooves absent; two setiferous punctures present on each side; posterolateral angles subrectangular; posterolateral carinae blunt, very long (about as long as adjoining foveae); laterobasal foveae well defined, impunctate, very deep and wide, oblong, prolonged forward, double; posterior bead absent; base emarginate, slightly narrower than pronotal apex and elytral base. Legs. Short. Elytra. Moderately convex, ovate, widest about middle. Basal margin incomplete, reaching about stria 4. Shoulders obtuse. Sides strongly rounded. Scutellar striole absent. Striae incomplete; stria 1 deep; striae 2 – 7 obsolete, impunctate; stria 3 with three setiferous punctures. Recurrent stria long, blunt, directed apically toward stria 5. Subapical seta present. Intervals depressed. Sutural apices obtuse. Abdomen. Last visible sternum (sternum VII): male with two apical ambulatory setae; female with four apical ambulatory setae. Aedeagus. Lateral view (Fig. 85): moderately arcuate, not narrowed in apical half; base almost straight dorsally; middle moderately convex dorsally and angulate posteriorly, straight ventrally, with dorsal membranous area very wide and long; apex subtriangular, straight dorsally, slightly concave ventrally, with extreme tip wide and long. Dorsal view: narrow, asymmetrical (ostium of membranous area deflected to right); apex straight; basal orifice narrow, closed anteriorly, moderately distant from membranous area. Parameres with four apical setae.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3027AFFBA3FCAF9B7FD58FA27.taxon	materials_examined	Material examined. 107 specimens (CMNZ, LUNZ, NZAC) Geographic distribution (Fig. 145). Subantarctic Islands: SN – Penguin Creek. Penguin Rookery. Sinkhole Drain. Sinkhole Flat. Station Point.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3027AFFBA3FCAF9B7FD58FA27.taxon	biology_ecology	Ecology. Lowland. Epigean. Wet peaty forests (Olearia); penguin colonies and their vicinity. Shaded or open ground. Nocturnal; hides during the day in leaf litter and bird nests. Gregarious. Biology. Seasonality: October, December – January. Predacious (based on mouthpart morphology). Dispersal power. Subapterous (incapable of flight). Slow runner. Collecting techniques. Sifting leaf litter; inspecting bird nests.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30279FFBB3FCAF9F7FE6BF8C7.taxon	description	Fig. 40, 86, 122	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30279FFBB3FCAF9F7FE6BF8C7.taxon	description	Description. Body length 4.5 – 5.6 mm. Head, pronotum, elytra, and abdomen mostly piceous black; lateral margins and suture of elytra reddish; abdomen yellowish laterally and apically; antennae and legs reddish; palpi yellowish; femora yellowish, reddish basally. Microsculpture strong and isodiametric on head, moderately transverse, strong on pronotum, feeble on elytra. Iridescence absent. Very shiny, with metallic luster (aeneous) on head, pronotum, and elytra. Head. Labrum moderately transverse, slightly emarginate anteriorly. Antennae submoniliform, short: segment 1 (scape) moderately long, about 2 x longer than its maximum width. Frontal furrows wide, deep, convergent. Eyes moderately convex; two setiferous punctures on inner side of each eye. Tempora strongly inflated (about two-thirds as long as eyes). Mentum: medial tooth entire, acute apically, moderately shorter than lateral lobes. Paraglossae membranous, prominent, about as long as ligula. Thorax. Pronotum strongly convex, finely punctate across base, wrinkled between laterobasal foveae, slightly transverse and cordate, widest before middle; apex subtruncate; anterolateral angles poorly developed, obtusely rounded; anterior bead incomplete, obsolete medially; sides strongly rounded, slightly sinuate posteriorly; lateral grooves absent; two setiferous punctures on each side; posterolateral angles obtuse; posterolateral carinae absent; laterobasal foveae well defined, finely punctate, very deep and wide, oblong, prolonged forward, simple; posterior bead absent; base emarginate, much narrower than pronotal apex, about as wide as elytral base. Legs. Short. Elytra. Strongly convex, ovate, widest about middle. Basal margin incomplete, reaching about stria 4. Shoulders rounded. Sides strongly rounded. Scutellar striole obsolete. Striae incomplete; stria 1 deep; striae 2 – 7 finely punctate, obsolete (striae 2 – 5 shallow medially); stria 3 with three setiferous punctures. Recurrent stria long, sharp, directed apically toward stria 7. Subapical seta absent (present in other Oopterus species). Intervals depressed. Sutural apices rounded. Abdomen. Last visible sternum (sternum VII): male with two apical and two medial ambulatory setae; female with four apical and two medial ambulatory setae. Aedeagus. Lateral view (Fig. 86): strongly arcuate, not widened in apical half; base slightly convex dorsally; middle subparallel, dorsally angulate, moderately concave anteriorly and straight posteriorly, moderately concave ventrally, with dorsal membranous area narrow and short; apex subtriangular, straight dorsally, slightly concave ventrally, with extreme tip wide and long. Dorsal view: narrow, symmetrical (ostium of membranous area dorsal); apex straight; basal orifice narrow, closed anteriorly, moderately distant from membranous area. Parameres with three or four apical setae.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30279FFBB3FCAF9F7FE6BF8C7.taxon	materials_examined	Material examined. 653 specimens (AMNZ, CMNZ, LUNZ, MONZ, NZAC). Geographic distribution (Fig. 122). Stewart Island: Port William. Subantarctic Islands: AN, AU, CA, SN.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30279FFBB3FCAF9F7FE6BF8C7.taxon	biology_ecology	Ecology. Lowland, montane, subalpine, alpine. Epigean. Wet forests (broadleaf); peaty scrublands; tussock areas; pastures; fellfields, supralittoral zone; sea beaches; penguin, shag, and seal colonies. Shaded or open ground. Nocturnal; hides during the day under logs, stones, in leaf litter, moss and mat plants, also under cow dung, in turf, among tussock roots, in logs, and under algae. Gregarious. Biology. Seasonality: October – February, April – May, July. Tenerals: December – April (AU). Predacious: amphipods, caterpillars. Occasionally infested with fungi (Laboulbeniales). Dispersal power. Subapterous (incapable of flight). Slow runner. Collecting techniques. Lifting logs and stones; sifting leaf litter, moss, and mat plants.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30279FFBB3FCAF9F7FE6BF8C7.taxon	discussion	Remarks. Oopterus clivinoides is the only Oopterus species without an elytral subapical seta. This taxon, common in the Subantarctic Islands, is recorded for the first time from Stewart Island (Port William; CMNZ).	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30277FFB53FCAFF37FB27FE07.taxon	description	Fig. 41, 87, 134 Holotype: male (NZAC) labeled: “ NEW ZEALAND HB Mohi Bush Sce. Res. [= Scenic Reserve] 300 m 21. I. 1998 Larochelle and Larivière (typed) / Dry Tawa forest. Under fallen branches. (typed) / [male symbol] / HOLOTYPE [male symbol] Oopterus mohiensis Larochelle & Larivière, 2017 (red label; typed). ” Paratypes: two males (CMNH, NZAC) from the same locality as the holotype, bearing blue paratype labels. Description. Body length 5.0 – 6.1 mm. Head, pronotum, elytra, and abdomen mostly piceous black; apex and base of head reddish; base of pronotum reddish; lateral margins and suture of elytra reddish; abdomen reddish laterally and apically; antennae, palpi, and legs reddish; antennal segments 1 + 2 yellow; femora infuscated. Microsculpture mostly absent on head (isodiametric basally), obsolete and very transverse (with microlines) on pronotum and elytra. Iridescence absent on head, feeble on pronotum and elytra. Very shiny, without metallic luster. Head. Labrum strongly transverse, slightly emarginate anteriorly. Antennae submoniliform, moderately long: segment 1 (scape) moderately long, about 2 x longer than its maximum width. Frontal furrows wide, deep, convergent. Eyes moderately convex; two setiferous punctures on inner side of each eye. Tempora not inflated. Mentum: medial tooth entire, subtruncate apically, moderately shorter than lateral lobes. Paraglossae membranous, prominent, slightly longer than ligula. Thorax. Pronotum moderately convex, coarsely punctate across base, unwrinkled, strongly transverse, widest before middle; apex subtruncate; anterolateral angles poorly developed, rounded; anterior bead incomplete, obsolete medially; sides moderately rounded, slightly sinuate posteriorly; lateral grooves absent; two setiferous punctures on each side; posterolateral angles rectangular; posterolateral carinae blunt, very long (about as long as adjoining foveae); laterobasal foveae well defined, coarsely punctate, very deep and wide, subrectangular, not prolonged forward, double; posterior bead absent; base emarginate, wider than pronotal apex, about as wide as elytral base. Legs. Moderately long. Elytra. Strongly convex, ovate, widest about middle. Basal margin complete, reaching about scutellum. Shoulders obtuse. Sides strongly rounded. Scutellar striole present. Striae incomplete; stria 1 deep; striae 2 – 4 shallow, finely punctate; striae 5 – 7 obsolete; stria 3 with three setiferous punctures. Recurrent stria long, sharp, directed apically toward stria 5. Subapical seta present. Intervals subdepressed. Sutural apices obtusely rounded. Abdomen. Last visible sternum (sternum VII): male with two apical ambulatory setae; female with four apical ambulatory setae. Aedeagus. Lateral view (Fig. 87): strongly arcuate, not narrowed in apical half; base strongly biconvex dorsally; middle rather straight dorsally, moderately convex ventrally, with dorsal membranous area moderately wide and very long; apex subtriangular, rather straight dorsally, straight ventrally, with extreme tip wide and rather long. Dorsal view: narrow, asymmetrical (ostium of membranous area deflected to right); apex straight; basal orifice narrow, closed anteriorly, moderately distant from membranous area. Parameres with four to six apical setae.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30277FFB53FCAFF37FB27FE07.taxon	materials_examined	Material examined. 38 specimens (AMNZ, CMNH, MONZ, NZAC). Geographic distribution (Fig. 134). North Island: HB, TK, WA, WN.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30277FFB53FCAFF37FB27FE07.taxon	biology_ecology	Ecology. Lowland, montane, subalpine, alpine. Epigean. Wet or dry forests (beech, broadleaf); alpine tussock grasslands. Shaded or open ground. Nocturnal; hides during the day in leaf litter and moss, under stones and fallen branches. Gregarious. Biology. Seasonality: December – March. Predacious (based on mouthpart morphology). Dispersal power. Subapterous (incapable of flight). Moderate runner. Collecting techniques. Sifting leaf litter and moss; lifting stones and fallen branches; pitfall trapping.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30277FFB53FCAFF37FB27FE07.taxon	discussion	Remarks. This species is named after the type locality Mohi Bush Scenic Reserve (HB) and the Latin suffix - ensis, denoting a place, locality, or country. Oopterus mohiensis is morphologically close to O. frontalis and O. punctatus. In addition to diagnostic characters of the male genitalia, O. mohiensis has the following distinguishing features: microsculpture obsolete on pronotum and elytra; eyes moderately convex; pronotum with posterolateral angles rectangular and laterobasal foveae coarsely punctate. Oopterus mohiensis and O. frontalis occur in southern areas of the North Island, while O. punctatus is known only from Banks Peninsula and the Port Hills in Christchurch (South Island).	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30276FFB63FCAFE14FEC4FD07.taxon	description	Fig. 42, 88, 126	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30276FFB63FCAFE14FEC4FD07.taxon	description	Description. Body length 5.3 – 6.5 mm. Head, pronotum, elytra, and abdomen mostly piceous black; apex and base of head reddish; base of pronotum reddish; lateral margins and suture of elytra reddish; abdomen reddish laterally and apically; antennae, palpi, and legs reddish; femora infuscated. Microsculpture strong, isodiametric on head, very transverse (with microlines) on pronotum and elytra. Iridescence absent on head, very strong on pronotum and elytra. Very shiny, without metallic luster. Head. Labrum strongly transverse, slightly emarginate anteriorly. Antennae submoniliform, moderately long: segment 1 (scape) moderately long, about 2 x longer than its maximum width. Frontal furrows wide, deep, convergent. Eyes slightly convex; two setiferous punctures on inner side of each eye. Tempora not inflated. Mentum: medial tooth entire, rounded apically, moderately shorter than lateral lobes. Paraglossae membranous, prominent, about as long as ligula. Thorax. Pronotum moderately convex, slightly punctate across base, wrinkled discally, strongly transverse, widest before middle; apex subtruncate; anterolateral angles poorly developed, rounded; anterior bead incomplete, obsolete medially; sides moderately rounded, slightly sinuate posteriorly; lateral grooves absent; two setiferous punctures on each side; posterolateral angles obtuse; posterolateral carinae blunt, very long (about as long as adjoining foveae); laterobasal foveae well defined, finely punctate, very deep and wide, subrectangular, prolonged forward, double; posterior bead absent; base emarginate, wider than pronotal apex, about as wide as elytral base. Legs. Moderately long. Elytra. Strongly convex, ovate, widest about middle. Basal margin complete, reaching about scutellum. Shoulders obtuse. Sides strongly rounded. Scutellar striole present. Stria 1 complete, deep; striae 2 – 7 incomplete; striae 2 – 4 shallow, finely punctate; striae 5 – 7 obsolete; stria 3 with three setiferous punctures. Recurrent stria long, sharp, directed apically toward stria 5. Subapical seta present. Intervals depressed. Sutural apices obtusely rounded. Abdomen. Last visible sternum (sternum VII): male with two apical ambulatory setae; female with four apical ambulatory setae. Aedeagus. Lateral view (Fig. 88): strongly arcuate, strongly widened in apical half; base moderately biconvex dorsally; middle almost straight dorsally, slightly convex ventrally, with dorsal membranous area moderately wide and short; apex subtriangular, almost straight dorsally, straight ventrally, with extreme tip wide and short. Dorsal view: narrow, symmetrical (ostium of membranous area dorsal); apex straight; basal orifice narrow, closed anteriorly, moderately distant from membranous area. Parameres with four or five apical setae.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30276FFB63FCAFE14FEC4FD07.taxon	materials_examined	Material examined. 36 specimens (BMNH, JNNZ, MONZ, NZAC). Geographic distribution (Fig. 126). WI – Palmerston North. WN – Dennan. Kaitoke. Mount Holdsworth. Orongorongo Valley. Wadestown. Wellington. Wilton’s Bush.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30276FFB63FCAFE14FEC4FD07.taxon	biology_ecology	Ecology. Lowland, montane. Epigean. Wet forests (beech, broadleaf, podocarp) and scrublands. Shaded ground. Nocturnal; hides during the day under stones, in leaf litter; also in flood debris along a stream. Biology. Seasonality: September – April, June – July. Predacious (based on mouthpart morphology). Dispersal power. Subapterous (incapable of flight). Moderate runner. Collecting techniques. Lifting stones; sifting leaf litter; pitfall trapping; inspecting flood debris.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30276FFB63FCAFE14FEC4FD07.taxon	discussion	Remark. Because O. frontalis is only known from specimens collected in the Wellington area, Wadestown is the most likely type locality. The Palmerston North record is plausible but not supported by specimen data.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30275FFB73FCAFD14FDCEFC07.taxon	description	Fig. 43, 89, 141 Holotype: male (NZAC) labeled: “ Under logs N. fusca forest Above Long Bay Banks Peninsula [NEW ZEALAND MC] 1. X. 1974 (hand-written) R. M. Emberson (typed) / HOLOTYPE [male symbol] Oopterus punctatus Larochelle & Larivière, 2017 (red label; typed). ” Paratypes: one male (NZAC) and one female (CMNZ) from the same locality as the holotype, bearing blue paratype labels. Description. Body length 5.4 – 6.5 mm. Head, pronotum, elytra, and abdomen mostly dark brown; apex and base of head reddish; base of pronotum reddish; lateral margins and suture of elytra reddish; abdomen reddish laterally and apically; antennal segments 1 – 3, palpi, tibiae, and tarsi reddish; antennal segments 4 – 11 dark brown; femora infuscated. Microsculpture strong, isodiametric on head, very transverse (with microlines) on pronotum and elytra. Iridescence absent on head, very strong on pronotum and elytra. Very shiny, without metallic luster. Head. Labrum strongly transverse, slightly emarginate anteriorly. Antennae submoniliform, moderately long: segment 1 (scape) moderately long, about 2 x longer than its maximum width. Frontal furrows wide, deep, convergent. Eyes slightly convex; two setiferous punctures on inner side of each eye. Tempora moderately inflated (about half as long as eyes). Mentum: medial tooth entire, rounded apically, moderately shorter than lateral lobes. Paraglossae membranous, prominent, about as long as ligula. Thorax. Pronotum moderately convex, virtually impunctate, wrinkled discally, strongly transverse, widest before middle; apex subtruncate; anterolateral angles poorly developed, rounded; anterior bead incomplete, obsolete medially; sides strongly rounded, slightly sinuate posteriorly; lateral grooves absent; two setiferous punctures on each side; posterolateral angles obtuse; posterolateral carinae blunt, very long (about as long as adjoining foveae); laterobasal foveae well defined, virtually impunctate, very deep and wide, subrectangular, not prolonged forward, double; posterior bead absent; base emarginate, wider than pronotal apex, about as wide as elytral base. Legs. Moderately long. Elytra. Strongly convex, ovate, widest about middle. Basal margin complete, reaching about scutellum. Shoulders obtuse. Sides strongly rounded. Scutellar striole obsolete. Striae complete; stria 1 deep, striae 2 – 7 shallow, finely punctate; stria 3 either without setiferous punctures or with one to three punctures. Recurrent stria long, sharp, directed apically toward stria 5. Subapical seta present. Intervals depressed. Sutural apices obtusely rounded. Abdomen. Last visible sternum (sternum VII): male with two apical ambulatory setae; female with four apical ambulatory setae. Aedeagus. Lateral view (Fig. 89): strongly arcuate, strongly widened in apical half; base moderately biconvex dorsally; middle biconvex dorsally, almost straight ventrally, with dorsal membranous area narrow and long; apex subtriangular, moderately convex dorsally, slightly concave ventrally, with extreme tip narrow and long. Dorsal view: narrow, symmetrical (ostium of membranous area dorsal); apex straight; basal orifice narrow, closed anteriorly, moderately distant from membranous area. Parameres with four or five apical setae.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30275FFB73FCAFD14FDCEFC07.taxon	materials_examined	Material examined. 12 specimens (CMNZ, LUNZ, NZAC). Geographic distribution (Fig. 141). South Island: MC – Banks Peninsula (Akaroa; Hinewai Scenic Reserve; Long Bay (above); Otepatutu Scenic Reserve; Port Levy). Port Hills, Christchurch.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30275FFB73FCAFD14FDCEFC07.taxon	biology_ecology	Ecology. Lowland (hills). Epigean. Forests (broadleaf, podocarp). Shaded ground. Nocturnal; hides during the day under logs and stones. Biology. Seasonality: October, February. Tenerals: March. Predacious (based on mouthpart morphology). Dispersal power. Subapterous (incapable of flight). Moderate runner. Collecting techniques. Lifting logs and stones.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30275FFB73FCAFD14FDCEFC07.taxon	discussion	Remarks. The name of this species is derived from the Latin adjective punctatus (punctate), referring to the strong microsculpture giving a “ punctate ” appearance to the head of the beetle. Oopterus punctatus is morphologically close to O. frontalis. In addition to diagnostic characters of the male genitalia, O. punctatus has the following distinguishing features: head, pronotum, elytra, and abdomen mostly dark brown; antennal segments 1 – 3 reddish, segments 4 – 11 dark brown; pronotum with sides strongly rounded, laterobasal foveae virtually impunctate, not prolonged forward; elytral striae complete and scutellar striole obsolete. The two species are allopatric: O. punctatus is known only from Banks Peninsula and the Port Hills in Christchurch (South Island), while O. frontalis occurs in southern areas of the North Island (WI, WN).	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30274FFB03FCAFC14FE26FB47.taxon	description	Fig. 44, 90, 148 Holotype: male (NZAC) labeled: “ [NEW ZEALAND FD] Summit Mt. [= Mount] Barber 1342 m (handwritten) / Manapouri Exp. Jan 70 (typed) / Under stones (typed) / [male symbol] / HOLOTYPE [male symbol] Oopterus trechoides Larochelle & Larivière, 2017 (red label; typed). ” Paratypes: one female (NZAC) and one male (MONZ) from the same locality as the holotype, bearing blue paratype labels. Description. Body length 5.9 – 7.3 mm. Head rufous; pronotum and elytra mostly rufopiceous; pronotum reddish basally; abdomen reddish piceous, yellow laterally and apically; antennae, palpi, and legs yellow. Microsculpture strong and isodiametric on head, obsolete on pronotum, strong and very transverse (with microlines) on elytra. Iridescence absent on head and pronotum, very strong on elytra. Very shiny, without metallic luster. Forebody narrow in comparison to elytra (as in O. arthurensis). Head. Labrum strongly transverse, slightly emarginate anteriorly. Antennae subfiliform (as in O. arthurensis, O. ocularius, and Zolus species), very long: segment 1 (scape) very long, about 2.5 x longer than its maximum width. Frontal furrows wide, deep, convergent. Eyes slightly convex; two setiferous punctures on inner side of each eye. Tempora strongly inflated (about as long as eyes). Mentum: medial tooth entire, acute apically, moderately shorter than lateral lobes. Paraglossae membranous, prominent, slightly longer than ligula. Thorax. Pronotum moderately convex, finely punctate across base, wrinkled in basal half, subquadrate, strongly cordate, widest before middle; apex subtruncate; anterolateral angles poorly developed, obtuse; anterior bead incomplete, obsolete medially; sides strongly rounded, strongly sinuate posteriorly; lateral grooves absent; two setiferous punctures on each side; posterolateral angles acute, projected laterally; posterolateral carinae absent; laterobasal foveae well defined, finely punctate, very deep and wide, oblong, not prolonged forward, simple; posterior bead absent; base emarginate, much narrower than pronotal apex and elytral base. Legs. Very long. Elytra. Moderately convex, subovate, widest about middle. Basal margin incomplete, reaching about stria 4. Shoulders oblique. Sides strongly rounded. Scutellar striole obsolete. Striae complete; striae 1 – 3 shallow, finely punctate; striae 4 – 7 obsolete; stria 3 with three setiferous punctures. Recurrent stria long, blunt, directed apically toward stria 6. Subapical seta present. Intervals subdepressed. Sutural apices obtusely rounded. Abdomen. Last visible sternum (sternum VII): male with two apical ambulatory setae; female with four apical ambulatory setae. Aedeagus. Lateral view (Fig. 90): very strongly arcuate, strongly widened both in basal half and apical half; base straight dorsally; middle strongly concave dorsally, moderately convex ventrally, with dorsal membranous area very wide and long; apex subtriangular, moderately sinuate dorsally, moderately concave ventrally, with extreme tip narrow, long, and sinuate. Dorsal view: moderately wide, asymmetrical (ostium of membranous area deflected to right); apex straight; basal orifice narrow, closed anteriorly, moderately distant from membranous area. Parameres with two apical setae.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30274FFB03FCAFC14FE26FB47.taxon	materials_examined	Material examined. 17 specimens (AMNZ, LUNZ, MONZ, NZAC). Geographic distribution (Fig. 148). South Island: FD – Fiordland National Park (Eglinton Valley, Mistake Creek; Mackinnon Pass; Mount Barber; Mount Grey [= Mount Gray]; Murchison Mountains, Plateau Creek). OL – Liverpool Stream. SL – Longwood Forest, Bald Hill.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30274FFB03FCAFC14FE26FB47.taxon	biology_ecology	Ecology. Subalpine, alpine. Epigean. Mostly alpine fellfields and grasslands; also wet subalpine forests (beech) and screes. Open or shaded ground. Nocturnal; hides during the day under stones. Gregarious. Biology. Seasonality: December – February. Tenerals: January – February. Predacious (based on mouthpart morphology). Dispersal power. Subapterous (incapable of flight). Fast runner. Collecting technique. Lifting stones.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30274FFB03FCAFC14FE26FB47.taxon	discussion	Remarks. The name of this species is derived from Trechus (type genus of the tribe Trechini) and the suffix - oides (similar), referring to the trechine-like body shape of the beetle. Oopterus trechoides is morphologically close to O. sculpturatus and O. taieriensis. In addition to diagnostic characters of the male genitalia, O. trechoides has the following distinguishing features: forebody narrow in comparison to elytra; antennae, palpi, and legs yellow; antennae subfiliform, very long; tempora strongly inflated (about as long as eyes); pronotum subquadrate, strongly cordate; elytra subovate. The three species are allopatric: O. trechoides occurs in the southwest of the South Island (FD, OL, SL), O. sculpturatus is restricted to the northwest of the South Island (NN), and O. taieriensis is found in the southeast of the South Island (DN).	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30273FFB13FCAFAD4FDC0F907.taxon	description	Fig. 45, 91, 143	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30273FFB13FCAFAD4FDC0F907.taxon	description	Description. Body length 5.2 – 6.3 mm. Head, pronotum, elytra, and abdomen mostly piceous black; apex and base of head reddish; base of pronotum reddish; lateral margins, suture (apical half), and apex of elytra reddish; abdomen reddish laterally and apically; antennae, palpi, and legs reddish; femora infuscated. Microsculpture strong, isodiametric on head, very transverse (with microlines) on pronotum and elytra. Iridescence absent on head, very strong on pronotum and elytra. Very shiny, without metallic luster. Head. Labrum strongly transverse, slightly emarginate anteriorly. Antennae submoniliform, moderately long: segment 1 (scape) moderately long, about 2 x longer than its maximum width. Frontal furrows wide, deep, convergent. Eyes slightly convex; two setiferous punctures on inner side of each eye. Tempora not inflated. Mentum: medial tooth entire, rounded apically, moderately shorter than lateral lobes. Paraglossae membranous, prominent, about as long as ligula. Thorax. Pronotum moderately convex, coarsely punctate basally, wrinkled discally, strongly transverse, widest about middle; apex subtruncate; anterolateral angles poorly developed, rounded; anterior bead incomplete, obsolete medially; sides strongly rounded, slightly sinuate posteriorly; lateral grooves absent; two setiferous punctures on each side; posterolateral angles acute, projected laterally; posterolateral carinae blunt, very long (about as long as adjoining foveae); laterobasal foveae well defined, coarsely punctate, very deep and wide, subrectangular, prolonged forward, double; posterior bead absent; base emarginate, wider than pronotal apex, slightly narrower than elytral base. Legs. Moderately long. Elytra. Strongly convex, ovate, widest about middle. Basal margin incomplete, reaching about stria 4. Shoulders strongly rounded. Sides strongly rounded. Scutellar striole present. Stria 1 complete, deep; striae 2 – 7 incomplete; striae 2 – 4 shallow, finely punctate; striae 5 – 7 obsolete; stria 3 with three setiferous punctures. Recurrent stria long, sharp, directed apically toward stria 5. Subapical seta present. Intervals depressed. Sutural apices obtusely rounded. Abdomen. Last visible sternum (sternum VII): male with two apical ambulatory setae; female with four apical ambulatory setae. Aedeagus. Lateral view (Fig. 91): moderately arcuate, not widened in apical half; base moderately biconvex dorsally; middle subparallel, slightly concave dorsally, straight ventrally, with dorsal membranous area moderately wide and long; apex triangular, straight dorsally, straight ventrally, with extreme tip wide and short. Dorsal view: narrow, asymmetrical (ostium of membranous area deflected to right); apex straight; basal orifice narrow, closed anteriorly, moderately distant from membranous area. Parameres with five apical setae.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30273FFB13FCAFAD4FDC0F907.taxon	materials_examined	Material examined. 15 specimens (BMNH, JNNZ, LUNZ, NZAC). Geographic distribution (Fig. 143). South Island: NN – Boulder Lake (track). Cobb Reservoir, Lake Sylvester Road. Cobb Valley, Galena Creek. Lake Sylvester (track). Mount Arthur, Summit. Oparara River (mouth). Westport.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30273FFB13FCAFAD4FDC0F907.taxon	biology_ecology	Ecology. Lowland, montane, subalpine. Endogean. Wet forests (beech). Shaded ground. Nocturnal; hides during the day under well-embedded logs and stones. Biology. Seasonality: November, January – March. Tenerals: March. Predacious (based on mouthpart morphology). Dispersal power. Subapterous (incapable of flight). Moderate runner. Collecting techniques. Lifting logs and stones.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30273FFB13FCAFAD4FDC0F907.taxon	discussion	Remark. Examination of the type of Oopterus sculpturatus var. ovinotatus revealed it to be O. sculpturatus sculpturatus; any differences noted by Broun are taxonomically unimportant and do not warrant the recognition of a subspecies.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30272FFB23FCAF917FA5FF8C7.taxon	description	Fig. 46, 92, 147 Holotype: male (NZAC) labeled: “ NEW ZEALAND DN Taieri Mouth 30 May 2010 (typed) / In flood debris from Taieri River (typed) / J T NUNN (typed) / [male symbol] / HOLOTYPE [male symbol] Oopterus taieriensis Larochelle & Larivière, 2017 (red label; typed). ” Paratype: one female (NZAC) from the same locality as the holotype, bearing a blue paratype label. Description. Body length 4.6 – 5.7 mm. Head, pronotum, elytra, and abdomen mostly piceous black; apex and base of head reddish; base of pronotum reddish; lateral margins and suture of elytra reddish; abdomen yellow laterally and apically; antennae, palpi, and legs reddish. Microsculpture strong, isodiametric on head, moderately transverse on pronotum, very transverse (with microlines) on elytra. Iridescence absent on head, moderately strong on pronotum, very strong on elytra. Very shiny, without metallic luster. Head. Labrum strongly transverse, slightly emarginate anteriorly. Antennae submoniliform, moderately long: segment 1 (scape) moderately long, about 2 x longer than its maximum width. Frontal furrows wide, deep, convergent. Eyes moderately convex; two setiferous punctures on inner side of each eye. Tempora not inflated. Mentum: medial tooth entire, acute apically, moderately shorter than lateral lobes. Paraglossae membranous, prominent, much shorter than ligula. Thorax. Pronotum moderately convex, impunctate, unwrinkled, strongly transverse, widest before middle; apex subtruncate; anterolateral angles poorly developed, rounded; anterior bead incomplete, obsolete medially; sides strongly rounded, slightly sinuate posteriorly; lateral grooves absent; two setiferous punctures on each side; posterolateral angles acute, projected laterally; posterolateral carinae absent; laterobasal foveae well defined, impunctate, moderately deep, very wide, oblong, prolonged forward, double; posterior bead absent; base emarginate, slightly wider than pronotal apex, slightly narrower than elytral base. Legs. Moderately long. Elytra. Moderately convex, ovate, widest about middle. Basal margin incomplete, reaching about stria 4. Shoulders obtuse. Sides strongly rounded. Scutellar striole present. Striae shallow, finely punctate; striae 1 – 4 complete; striae 5 – 7 incomplete, obsolete; stria 3 with three setiferous punctures. Recurrent stria long, sharp, directed apically toward stria 5. Subapical seta present. Intervals depressed. Sutural apices obtuse. Abdomen. Last visible sternum (sternum VII): male with two apical ambulatory setae; female with four apical ambulatory setae. Aedeagus. Lateral view (Fig. 92): moderately arcuate, narrowed in apical half; base slightly convex dorsally; middle angulate anteriorly and moderately convex dorsally, moderately convex ventrally, with dorsal membranous area moderately wide and very long; apex unusually slender, slightly concave dorsally, strongly concave ventrally, with extreme tip unusually slender and curved downward. Dorsal view: narrow, asymmetrical (ostium of membranous area deflected to right); apex straight; basal orifice narrow, closed anteriorly, moderately distant from membranous area. Parameres with three or four apical setae.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30272FFB23FCAF917FA5FF8C7.taxon	materials_examined	Material examined. 26 specimens (JNNZ, NZAC). Geographic distribution (Fig. 147). South Island: DN – Allison Scenic Reserve. Flagstaff Hill. Leith Stream (Dunedin, town belt). Otago Peninsula, Boulder Beach. Taieri River, Taieri Mouth.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30272FFB23FCAF917FA5FF8C7.taxon	biology_ecology	Ecology. Lowland. Epigean. Forests (broadleaf), along streams. Shaded ground. Nocturnal; hides during the day in flood debris and under logs. Biology. Seasonality: January – February, May. Tenerals: March, May. Predacious (based on mouthpart morphology). Dispersal power. Subapterous (incapable of flight). Moderate runner. Collecting techniques. Inspecting flood debris; lifting logs.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30272FFB23FCAF917FA5FF8C7.taxon	discussion	Remarks. This species is named after the type locality Taieri Mouth (DN) and the Latin suffix - ensis, denoting a place, locality, or country. Oopterus taieriensis is morphologically close to O. sculpturatus. In addition to diagnostic characters of the male genitalia, O. taieriensis has the following distinguishing features: pronotum widest before middle, base impunctate, posterolateral carinae absent, laterobasal foveae oblong; elytra moderately convex. The two species are allopatric: O. taieriensis is restricted to the southeastern South Island (DN), while O. sculpturatus occurs in the northwest of the South Island (NN).	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30270FFAC3FCAFF37FDBFFE87.taxon	description	Fig. 47, 93, 138	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30270FFAC3FCAFF37FDBFFE87.taxon	description	Description. Body length 7.4 – 7.8 mm. Head, pronotum, elytra, and abdomen mostly reddish piceous; lateral margins and suture of elytra reddish; abdomen yellow laterally and apically; antennae, palpi, and legs testaceous. Microsculpture strong and isodiametric on head, feeble and moderately transverse on pronotum, obsolete and very transverse (with microlines) on elytra. Iridescence absent on head and pronotum, feeble on elytra. Very shiny, without metallic luster. Head. Labrum strongly transverse, slightly emarginate anteriorly. Antennae subfiliform (as in O. arthurensis, O. trechoides, and Zolus species), very long: segment 1 (scape) very long, about 2.5 x longer than its maximum width. Frontal furrows wide, deep, convergent. Eyes subdepressed; two setiferous punctures on inner side of each eye. Tempora strongly inflated (about as long as eyes). Mentum: medial tooth entire, acute apically, moderately shorter than lateral lobes. Paraglossae membranous, prominent, much longer than ligula. Thorax. Pronotum moderately convex, finely punctate, wrinkled across base, moderately transverse, widest just before middle; apex subtruncate; anterolateral angles poorly developed, obtusely rounded; anterior bead incomplete, obsolete medially; sides moderately rounded, slightly sinuate posteriorly; lateral grooves absent; two setiferous punctures on each side; posterolateral angles rectangular; posterolateral carinae blunt, very long (about as long as adjoining foveae); laterobasal foveae well defined, finely punctate, very deep and wide, square, prolonged forward, double; posterior bead absent; base subtruncate, slightly wider than pronotal apex, slightly narrower than elytral base. Legs. Very long. Elytra. Moderately convex, subovate, widest before middle. Basal margin incomplete, reaching about stria 4. Shoulders obtuse. Sides moderately rounded. Scutellar striole obsolete. Striae complete, shallow, finely punctate; interval 3 with three setiferous punctures. Recurrent stria long, sharp, directed apically toward stria 5. Subapical seta present. Intervals depressed. Sutural apices obtuse. Abdomen. Last visible sternum (sternum VII): male with two apical ambulatory setae; female with four apical ambulatory setae. Aedeagus. Lateral view (Fig. 93): strongly arcuate, moderately widened in apical half; base moderately biconvex dorsally; middle moderately concave dorsally, slightly concave ventrally, with dorsal membranous area very wide and long; apex triangular, straight dorsally, slightly concave ventrally, with extreme tip wide and long. Dorsal view: moderately wide, symmetrical (ostium of membranous area dorsal); apex straight; basal orifice wide, closed anteriorly, moderately distant from membranous area. Parameres with five or seven apical setae.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30270FFAC3FCAFF37FDBFFE87.taxon	materials_examined	Material examined. 15 specimens (AMNZ, BMNH, JNNZ, LUNZ, NZAC). Geographic distribution (Fig. 138). South Island: FD, OL, WD.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30270FFAC3FCAFF37FDBFFE87.taxon	biology_ecology	Ecology. Lowland (hills), montane, subalpine. Endogean. Wet forests (beech). Shaded ground. Nocturnal; hides during the day under well-embedded logs. Biology. Seasonality: December – March. Tenerals: December (FD). Predacious (based on mouthpart morphology). Occasionally infested with fungi (Laboulbeniales). Dispersal power. Subapterous (incapable of flight). Fast runner. Collecting techniques. Lifting logs; pitfall trapping.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3026FFFAD3FCAFE94FD58FE67.taxon	description	Fig. 48, 94, 132	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3026FFFAD3FCAFE94FD58FE67.taxon	description	Description. Body length 4.9 – 5.4 mm. Head, pronotum, elytra, and abdomen mostly reddish brown (forebody slightly paler); abdomen yellow laterally and apically; lateral margins and suture of elytra reddish; antennae, palpi, and legs reddish. Microsculpture strong, isodiametric on head, moderately transverse on pronotum and elytra. Iridescence absent. Very shiny, without metallic luster. Head. Labrum strongly transverse, slightly emarginate anteriorly. Antennae submoniliform, short: segment 1 (scape) moderately long, about 2 x longer than its maximum width. Frontal furrows wide, shallow, convergent. Eyes moderately convex; two setiferous punctures on inner side of each eye. Tempora strongly inflated (about two-thirds as long as eyes). Mentum: medial tooth entire, acute apically, moderately shorter than lateral lobes. Paraglossae membranous, prominent, much longer than ligula. Thorax. Pronotum moderately convex, impunctate, wrinkled across base, slightly transverse, moderately cordate, widest before middle; apex truncate; anterolateral angles poorly developed, obtusely rounded; anterior bead incomplete, obsolete medially; sides strongly rounded, slightly sinuate posteriorly; lateral grooves shallow, very long (about two-thirds as long as pronotum); two setiferous punctures on each side; posterolateral angles obtuse; posterolateral carinae blunt, very long (about as long as adjoining foveae); laterobasal foveae well defined, impunctate, moderately deep and wide, rounded, not prolonged forward, simple; posterior bead absent; base subtruncate, much narrower than pronotal apex, about as wide as elytral base. Legs. Short. Elytra. Moderately convex, ovate, widest about middle. Basal margin incomplete, reaching about stria 4. Shoulders moderately rounded. Sides strongly rounded. Scutellar striole obsolete. Striae complete, shallow, impunctate or finely punctate; stria 3 with three setiferous punctures. Recurrent stria long, sharp, directed apically toward stria 5 or 6. Subapical seta present. Intervals depressed. Sutural apices obtusely rounded. Abdomen. Last visible sternum (sternum VII): male with two apical and two medial ambulatory setae; female with four apical ambulatory setae. Aedeagus. Lateral view (Fig. 94): moderately arcuate, narrowed in apical half; base strongly biconvex dorsally; middle strongly sinuate dorsally, moderately concave ventrally, with dorsal membranous area moderately wide and long; apex subtriangular, moderately concave dorsally, slightly concave ventrally, with extreme tip wide and long. Dorsal view: narrow, asymmetrical (ostium of membranous area deflected to right); apex straight; basal orifice narrow, closed anteriorly, moderately distant from membranous area. Parameres with two or three apical setae.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3026FFFAD3FCAFE94FD58FE67.taxon	materials_examined	Material examined. 201 specimens (AMNZ, CMNZ, MONZ, NZAC). Geographic distribution (Fig. 132). Subantarctic Islands: CA.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3026FFFAD3FCAFE94FD58FE67.taxon	biology_ecology	Ecology. Lowland, montane, subalpine, alpine. Epigean. Supralittoral zone; peaty scrublands; tussock swards; fellfields; mollymawk and penguin colonies. Open or shaded ground. Nocturnal; hides during the day under stones, in leaf litter, under pieces of wood, in / under tussock clumps and moss, and under mat plants. Gregarious. Biology. Seasonality: September – January, May, August. Tenerals: December – January. Predacious (based on mouthpart morphology). Occasionally infested with mites. Dispersal power. Subapterous (incapable of flight). Slow runner. Collecting techniques. Lifting stones and pieces of wood; sifting leaf litter and moss; inspecting vegetation clumps.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3026EFFAE3FCAFDB4FD8BFD47.taxon	description	Fig. 49, 95, 120 Holotype: male (NZAC) labeled: “ [NEW ZEALAND SI] Mt. [= Mount] Anglem, 3000 ’ Stewart Island (typed) / 28. I. 62 G. Kuschel (typed) / [male symbol] / HOLOTYPE [male symbol] Oopterus anglemensis Larochelle & Larivière, 2017 (red label; typed). ” Paratypes: two females (AMNZ, NZAC) from the same locality as the holotype, bearing blue paratype labels. Description. Body length 6.0 – 6.5 mm. Head, pronotum, elytra, and abdomen mostly black; base of pronotum reddish; lateral margins and suture of elytra reddish; abdomen reddish laterally and apically; antennae, palpi, and legs reddish. Microsculpture strong, isodiametric on head, very transverse (with microlines) on pronotum and elytra. Iridescence absent on head, very strong on pronotum and elytra. Very shiny, without metallic luster. Head. Labrum strongly transverse, slightly emarginate anteriorly. Antennae submoniliform, moderately long: segment 1 (scape) moderately long, about 2 x longer than its maximum width. Frontal furrows wide, deep, convergent. Eyes slightly convex; two setiferous punctures on inner side of each eye. Tempora moderately inflated (about half as long as eyes). Mentum: medial tooth entire, acute apically, moderately shorter than lateral lobes. Paraglossae membranous, prominent, slightly longer than ligula. Thorax. Pronotum moderately convex, impunctate, wrinkled discally, strongly transverse and cordate, widest before middle; apex subtruncate; anterolateral angles poorly developed, rounded; anterior bead incomplete, obsolete medially; sides strongly rounded, strongly sinuate posteriorly; lateral grooves absent; two setiferous punctures on each side; posterolateral angles subrectangular; posterolateral carinae absent; laterobasal foveae ill-defined, impunctate, shallow, very wide, prolonged forward, double; posterior bead absent; base subtruncate, much narrower than pronotal apex, slightly narrower than elytral base. Legs. Moderately long. Elytra. Moderately convex, subovate, widest about middle. Basal margin incomplete, reaching about stria 4. Shoulders obtuse. Sides moderately rounded. Scutellar striole obsolete. Striae complete, shallow, finely punctate; stria 3 with four or five setiferous punctures. Recurrent stria long, blunt, directed apically toward stria 5. Subapical seta present. Intervals subdepressed. Sutural apices obtuse. Abdomen. Last visible sternum (sternum VII): male with two apical ambulatory setae; female with four apical ambulatory setae. Aedeagus. Lateral view (Fig. 95): moderately arcuate, not widened in apical half; base moderately biconvex dorsally; middle subparallel, almost straight dorsally, slightly concave ventrally, with dorsal membranous area moderately wide and very long; apex subtriangular, slightly biconcave dorsally, slightly concave ventrally, with extreme tip narrow and long. Dorsal view: narrow, asymmetrical (ostium of membranous area deflected to right); apex straight; basal orifice narrow, closed anteriorly, moderately distant from membranous area. Parameres with four or five apical setae.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3026EFFAE3FCAFDB4FD8BFD47.taxon	materials_examined	Material examined. 48 specimens (AMNZ, CMNZ, LUNZ, NZAC). Geographic distribution (Fig. 120). Stewart Island: Freds Camp. Freshwater. Mount Anglem. Oban. Ocean Beach. Pegasus Creek. Port Pegasus. Port William. Table Hill. Twilight Bay.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3026EFFAE3FCAFDB4FD8BFD47.taxon	biology_ecology	Ecology. Lowland, montane, subalpine, alpine. Epigean. Wet forests (broadleaf, podocarp). Shaded ground. Nocturnal; hides during the day under logs, as well as in moss and leaf litter. Gregarious. Biology. Seasonality: January – February. Tenerals: February. Predacious (based on mouthpart morphology). Dispersal power. Subapterous (incapable of flight). Moderate runner. Collecting techniques. Lifting logs; sifting moss and leaf litter.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3026EFFAE3FCAFDB4FD8BFD47.taxon	discussion	Remarks. This species is named after the type locality Mount Anglem (Stewart Island) and the Latin suffix - ensis, denoting a place, locality, or country. Oopterus anglemensis is morphologically close to O. nanus. In addition to diagnostic characters of the male genitalia, O. anglemensis has the following distinguishing features: body length 6.0 – 6.5 mm; pronotum strongly cordate, laterobasal foveae ill- defined; legs moderately long; elytra moderately convex, subovate, sides moderately rounded. The two species are allopatric: O. anglemensis is restricted to Stewart Island, while O. nanus occurs in southern areas of the South Island (CO, OL).	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3026DFFAF3FCAFCD4FC3CFCA7.taxon	description	Fig. 50, 96, 136 Holotype: male (NZAC) labeled: “ [NEW ZEALAND CO] Old Man Ra CO 1615 m 26 Feb 74 J. S. Dugdale (hand-written) / ex Celmisia haastii (hand-written) / HOLOTYPE [male symbol] Oopterus nanus Larochelle & Larivière, 2017 (red label; typed). ” Paratypes: two females (LUNZ, NZAC) from the same locality as the holotype, bearing blue paratype labels. Description. Body length 3.9 – 5.2 mm. Head, pronotum, elytra, and abdomen mostly black; abdomen reddish piceous laterally and apically; antennae, palpi, and legs reddish. Microsculpture strong and isodiametric on head, obsolete and moderately transverse on pronotum, feeble and isodiametric on elytra. Iridescence absent. Very shiny, without metallic luster. Head. Labrum strongly transverse, slightly emarginate anteriorly. Antennae submoniliform, short: segment 1 (scape) moderately long, about 2 x longer than its maximum width. Frontal furrows wide, deep, convergent. Eyes moderately convex; two setiferous punctures on inner side of each eye. Tempora moderately inflated (about half as long as eyes). Mentum: medial tooth entire, acute apically, moderately shorter than lateral lobes. Paraglossae membranous, prominent, slightly longer than ligula. Thorax. Pronotum moderately convex, impunctate, wrinkled discally, strongly transverse, widest before middle; apex subtruncate; anterolateral angles poorly developed, rounded; anterior bead incomplete, obsolete medially; sides strongly rounded, not sinuate posteriorly; lateral grooves absent; two setiferous punctures on each side; posterolateral angles subrectangular; posterolateral carinae absent; laterobasal foveae well defined, impunctate, shallow, very wide, rectangular, not prolonged forward, double; posterior bead absent; base emarginate, much narrower than pronotal apex, slightly narrower than elytral base. Legs. Short. Elytra. Subdepressed, oblong, widest about middle. Basal margin incomplete, reaching about stria 4. Shoulders obtuse. Sides subparallel. Scutellar striole obsolete. Striae complete, shallow, finely punctate; stria 3 with four unusually large setiferous punctures. Recurrent stria long, sharp, directed apically toward stria 5, 6 or 7. Subapical seta present. Intervals depressed. Sutural apices obtuse. Abdomen. Last visible sternum (sternum VII): male with two apical ambulatory setae; female with four apical ambulatory setae. Aedeagus. Lateral view (Fig. 96): strongly arcuate, narrowed in apical half; base moderately biconvex dorsally; middle strongly convex dorsally, slightly convex ventrally, with dorsal membranous area very wide and moderately long; apex slender, strongly concave dorsally, slightly concave ventrally, with extreme tip wide, unusually long. Dorsal view: narrow, asymmetrical (ostium of membranous area deflected to right); apex straight; basal orifice narrow, closed anteriorly, moderately distant from membranous area. Parameres with four to six apical setae.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3026DFFAF3FCAFCD4FC3CFCA7.taxon	materials_examined	Material examined. 196 specimens (AMNZ, JNNZ, LUNZ, NZAC). Geographic distribution (Fig. 136). South Island: CO, OL.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3026DFFAF3FCAFCD4FC3CFCA7.taxon	biology_ecology	Ecology. Alpine. Epigean. Fellfields; tussock grasslands. Open ground. Nocturnal; hides during the day under stones and in mat plant litter. Gregarious. Biology. Seasonality: November – April. Tenerals: January – February. Predacious (based on mouthpart morphology). Dispersal power. Subapterous (incapable of flight). Slow runner. Collecting techniques. Lifting stones; sifting plant litter.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3026DFFAF3FCAFCD4FC3CFCA7.taxon	discussion	Remarks. The species name is based on the Latin noun nanus, denoting a dwarf, referring to the small size of the beetle. Oopterus nanus is morphologically close to O. anglemensis. In addition to diagnostic characters of the male genitalia, O. nanus has the following distinguishing features: body length 3.9 – 5.2 mm; pronotum not cordate, laterobasal foveae well defined, rectangular; elytra subdepressed, oblong, sides subparallel. The two species are allopatric: O. nanus occurs in southern areas of the South Island (CO, OL), while O. anglemensis is restricted to Stewart Island.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3026CFFA83FCAFC74FC9FFBA7.taxon	description	Fig. 51, 97, 128 Holotype: male (NZAC) labeled: “ [NEW ZEALAND FD] Mt. [= Mount] Barber 1350 m Wilmot Pass (hand-written) / Wilmot Pass 300 m- 630 m (typed) / Manapouri Exp. Jan 70 A. C. Eyles (typed) / Mats (typed) 70 / 88 (hand-written) / [male symbol] / HOLOTYPE [male symbol] Oopterus garnerae Larochelle & Larivière, 2017 (red label; typed). ” Paratype: one male (NZAC) from Lake Hauroko (FD), bearing a blue paratype label. Description. Body length 4.1 – 6.2 mm. Head, pronotum, elytra, and abdomen mostly dark reddish (forebody slightly paler); disc of head darker; lateral margins, suture, and apex of elytra reddish; abdomen yellow laterally and apically; antennae, palpi, and legs reddish; antennal segments 1 + 2 pale yellow; femora mostly pale yellow, reddish basally. Microsculpture strong and isodiametric on head, moderately transverse on pronotum, strong and very transverse (with microlines) on elytra. Iridescence absent on head and pronotum, very strong on elytra. Very shiny, without metallic luster. Head. Labrum strongly transverse, slightly emarginate anteriorly. Antennae submoniliform, moderately long: segment 1 (scape) moderately long, about 2 x longer than its maximum width. Frontal furrows wide, deep, convergent. Eyes slightly convex; two setiferous punctures on inner side of each eye. Tempora moderately inflated (about half as long as eyes). Mentum: medial tooth entire, acute apically, moderately shorter than lateral lobes. Paraglossae membranous, prominent, slightly longer than ligula. Thorax. Pronotum strongly convex, impunctate, unwrinkled, strongly transverse and cordate, widest before middle; apex subtruncate; anterolateral angles poorly developed, rounded; anterior bead incomplete, obsolete medially; sides strongly rounded, strongly sinuate posteriorly; lateral grooves absent; two setiferous punctures on each side; posterolateral angles subrectangular; posterolateral carinae blunt, short (shorter than adjoining foveae); laterobasal foveae well defined, impunctate, moderately deep, very wide, oblong, prolonged forward, double; posterior bead absent; base emarginate, much narrower than pronotal apex, slightly narrower than elytral base. Legs. Moderately long. Elytra. Strongly convex, ovate, widest about middle. Basal margin incomplete, reaching about stria 4. Shoulders obtuse. Sides strongly rounded. Scutellar striole obsolete. Striae shallow, finely punctate; stria 1 complete; striae 2 – 7 incomplete; stria 3 with three setiferous punctures. Recurrent stria long, sharp, directed apically toward stria 5, 6 or 7. Subapical seta present. Intervals subdepressed. Sutural apices obtusely rounded. Abdomen. Last visible sternum (sternum VII): male with two apical ambulatory setae; female with four apical ambulatory setae. Aedeagus. Lateral view (Fig. 97): moderately arcuate, narrowed in apical half; base slightly convex dorsally; middle angulate anteriorly and slightly convex dorsally, slightly convex ventrally, with dorsal membranous area moderately wide and long; apex triangular, straight dorsally, slightly concave ventrally, with extreme tip narrow and long. Dorsal view: narrow, symmetrical (ostium of membranous area dorsal); apex straight; basal orifice narrow, closed anteriorly, moderately distant from membranous area. Parameres with four to six apical setae.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3026CFFA83FCAFC74FC9FFBA7.taxon	materials_examined	Material examined. 38 specimens (JNNZ, NZAC). Geographic distribution (Fig. 128). South Island: FD, SL.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3026CFFA83FCAFC74FC9FFBA7.taxon	biology_ecology	Ecology. Montane, subalpine, alpine. Epigean. Wet forests (beech); alpine grasslands. Shaded or open ground. Nocturnal; hides during the day in moss and tussock litter. Biology. Seasonality: October – February, April – June. Predacious (based on mouthpart morphology). Dispersal power. Subapterous (incapable of flight). Moderate runner. Collecting techniques. Sifting moss and tussock litter; pitfall trapping.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3026CFFA83FCAFC74FC9FFBA7.taxon	discussion	Remarks. This species is named after our friend and colleague Beulah Garner (The Natural History Museum, London) for facilitating our study of the type material of Carabidae, especially the Broun Collection. Oopterus garnerae is morphologically close to O. nunni. In addition to diagnostic characters of the male genitalia, O. garnerae has the following distinguishing features: head, pronotum, elytra, and abdomen mostly dark reddish; tempora moderately inflated (about half as long as eyes); pronotum strongly convex and cordate, posterolateral carinae blunt and short; elytra strongly convex. The two species are allopatric: O. garnerae occurs in the southwest of the South Island (FD, SL), while O. nunni is found in the southeast of the South Island (DN, SL).	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3026BFFA93FCAFB74FE28FAE7.taxon	description	Fig. 52, 98, 137 Holotype: male (NZAC) labeled: “ NEW ZEALAND SL Catlins SFP [= Catlins State Forest Park], Tawanui Campground 100 m 12. I. 1999 Larochelle and Larivière (typed) / Wet beech forest – along stream banks – in leaf litter. (typed) / [male symbol] / HOLOTYPE [male symbol] Oopterus nunni Larochelle & Larivière, 2017 (red label; typed). ” Paratypes: one female (NZAC) and one male (MONZ) from the same locality as the holotype, bearing blue paratype labels. Description. Body length 4.9 – 5.8 mm. Head, pronotum, elytra, and abdomen mostly piceous black; disc of head paler; lateral margins, suture, and apex of elytra reddish yellow; abdomen reddish piceous, yellow laterally and apically; antennae, palpi, and legs reddish; femora pale yellow. Microsculpture strong and isodiametric on head, obsolete and very transverse (with microlines) on pronotum and elytra. Iridescence absent on head, very strong on pronotum and elytra. Very shiny, without metallic luster. Head. Labrum strongly transverse, slightly emarginate anteriorly. Antennae submoniliform, moderately long: segment 1 (scape) moderately long, about 2 x longer than its maximum width. Frontal furrows wide, deep, convergent. Eyes slightly convex; two setiferous punctures on inner side of each eye. Tempora slightly inflated (about one third as long as eyes). Mentum: medial tooth entire, acute apically, moderately shorter than lateral lobes. Paraglossae membranous, prominent, much longer than ligula. Thorax. Pronotum moderately convex, impunctate, wrinkled discally, strongly transverse, widest before middle; apex emarginate; anterolateral angles moderately developed, rounded; anterior bead incomplete, obsolete medially; sides strongly rounded, slightly sinuate posteriorly; lateral grooves absent; two setiferous punctures on each side; posterolateral angles subrectangular; posterolateral carinae absent; laterobasal foveae well defined, impunctate, moderately deep, very wide, oblong, not prolonged forward, double; posterior bead absent; base emarginate, much narrower than pronotal apex, slightly narrower than elytral base. Legs. Moderately long. Elytra. Moderately convex, ovate, widest about middle. Basal margin incomplete, reaching about stria 4. Shoulders rounded. Sides strongly rounded. Scutellar striole obsolete. Striae incomplete; striae 1 – 4 shallow, finely punctate; striae 5 – 7 obsolete; stria 3 with three setiferous punctures. Recurrent stria long, sharp, directed apically toward stria 5. Subapical seta present. Intervals depressed. Sutural apices obtusely rounded. Abdomen. Last visible sternum (sternum VII): male with two apical ambulatory setae; female with four apical ambulatory setae. Aedeagus. Lateral view (Fig. 98): moderately arcuate, not widened in apical half; base strongly biconvex dorsally; middle slightly concave dorsally, moderately convex ventrally, with dorsal membranous area moderately wide and long; apex subtriangular, slightly concave dorsally, moderately concave ventrally, with extreme tip slender and curved downward. Dorsal view: narrow, symmetrical (ostium of membranous area dorsal); apex straight; basal orifice narrow, closed anteriorly, moderately distant from membranous area. Parameres with four to six apical setae.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3026BFFA93FCAFB74FE28FAE7.taxon	materials_examined	Material examined. 75 specimens (JNNZ, MONZ, NZAC). Geographic distribution (Fig. 137). South Island: DN, SL.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3026BFFA93FCAFB74FE28FAE7.taxon	biology_ecology	Ecology. Lowland. Endogean. Wet forests (beech, broadleaf, podocarp), along streams. Shaded ground. Nocturnal; hides during the day in leaf litter, under well-embedded fallen branches, logs, and stones. Gregarious. Biology. Seasonality: December – March, July. Tenerals: February (DN). Predacious (based on mouthpart morphology). Dispersal power. Subapterous (incapable of flight). Moderate runner. Collecting techniques. Treading leaf litter into water; lifting fallen branches, logs, and stones; pitfall trapping.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3026BFFA93FCAFB74FE28FAE7.taxon	discussion	Remarks. This species is named after our friend and colleague John Nunn (Dunedin) for his contribution to the building of important reference collections of New Zealand carabids. Oopterus nunni is morphologically close to O. garnerae. In addition to diagnostic characters of the male genitalia, O. nunni has the following distinguishing features: head, pronotum, elytra, and abdomen mostly piceous black; tempora slightly inflated (about one third as long as eyes); pronotum moderately convex, not cordate, posterolateral carinae absent; elytra moderately convex. The two species are allopatric: O. nunni is found in the southeast of the South Island (DN, SL), while O. garnerae occurs in the southwest of the South Island (FD, SL).	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3026AFFAA3FCAFA37FC3CFA07.taxon	description	Description (New Zealand). Aedeagus with basal orifice open anteriorly; basal bulb with lateral lobes unequal in length.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3026AFFAA3FCAFA37FC3CFA07.taxon	discussion	Remarks. Three New Zealand genera (Synteratus Broun, Pseudoopterus Csiki, Maungazolus new genus) share the genitalic characters assigned to Merizodina by Jeannel (1940) and Roig-Juñent and Cicchino (2001): aedeagus with basal orifice open anteriorly; basal bulb with lateral lobes unequal in length. The genera Synteratus and Pseudoopterus (taxon previously synonymized with Oopterus) are transferred from subtribe Zolina to subtribe Merizodina. The presence of the latter subtribe is confirmed here for New Zealand. Genus Synteratus Broun, 1909 Fig. 53, 99, 153	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3026AFFAA3FCAFA37FC3CFA07.taxon	type_taxon	Type species. Synteratus ovalis Broun, 1909, by monotypy.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3026AFFAA3FCAFA37FC3CFA07.taxon	description	Description. Body length 3.0 – 3.5 mm; ovate, not narrowed around bases of pronotum and elytra (as opposed to other zoline genera). Color mostly dark. Metallic luster present. Dorsal surface mostly glabrous. Head. Narrow. Mandibles moderately long. Labrum slightly emarginate anteriorly. Eyes subdepressed; a single setiferous puncture on inner side of each eye (posteriorly). Antennae submoniliform. Tempora inflated. Mentum tooth entire (truncate apically). Ligula with six setae (two medial setae fused, four lateral setae free). Thorax. Pronotum transverse, trapezoidal; lateral grooves absent; setiferous punctures absent on each side; posterolateral carinae absent; base wider than apex. Scutellum invisible, concealed under pronotal base. Legs. Short. Segments 1 + 2 of male protarsi with a single row of ventral scales. Elytra. Oblong. Basal margin complete, reaching about scutellar striole. Shoulders well developed, angular. Scutellar setiferous pore present, inserted at junction of striae 1 + 2. Scutellar striole present. Striae complete, consisting of lines; two or three discal setiferous punctures in stria 3; recurrent stria directed apically toward stria 5. Subapical seta present. Apical seta present. Sutural apices obtusely rounded. Abdomen. Last visible sternum (sternum VII): male with two apical ambulatory setae; female with four apical ambulatory setae. Aedeagus. Lateral view: dorsal membranous area narrow. Dorsal view (Fig. 18): moderately wide; basal orifice wide, open anteriorly, moderately distant from membranous area; lateral lobes of basal bulb widely distant (moderately distant in Maungazolus and Pseudoopterus). Parameres with two or three apical setae. Right paramere curved, very long, narrow, untwisted medially.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3026AFFAA3FCAFA37FC3CFA07.taxon	discussion	Remarks. The genus Synteratus is transferred from the subtribe Zolina to the subtribe Merizodina on the basis of the aedeagus with basal orifice open anteriorly and basal bulb with lateral lobes unequal in length. The morphological characters distinguishing this genus from Pseudoopterus and Maungazolus are: body ovate, not narrowed around bases of pronotum and elytra; eyes subdepressed; pronotum trapezoidal, without lateral setiferous punctures; scutellum invisible, concealed under pronotal base; aedeagus (in dorsal view) with basal lobes widely open behind.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30269FFA43FCAFA17FEA9FC24.taxon	description	Fig. 53, 99, 153	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30269FFA43FCAFA17FEA9FC24.taxon	description	Description. Body length 3.0 – 3.5 mm. Head reddish; pronotum and elytra mostly reddish piceous; lateral margins, suture and apex of elytra reddish; abdomen mostly reddish piceous, reddish laterally and apically; antennae, palpi, and legs reddish; femora mostly pale yellow, reddish basally. Microsculpture strong and isodiametric on head, feeble and isodiametric on pronotum, moderately transverse on elytra. Iridescence absent on head and pronotum, moderately strong on elytra. Very shiny, with metallic luster (aeneous) on head, pronotum, and elytra. Head. Labrum moderately transverse, slightly emarginate. Antennae submoniliform, short: segment 1 (scape) short, stout, about 1.5 x longer than its maximum width. Frontal furrows wide, shallow, convergent. Eyes subdepressed; a single setiferous puncture on inner side of each eye (posteriorly). Tempora slightly inflated (about one third as long as eyes). Mentum: medial tooth entire, truncate apically, moderately shorter than lateral lobes. Paraglossae membranous, prominent, about as long as ligula. Thorax. Pronotum moderately convex, finely punctate across base, wrinkled discally, strongly transverse, subtriangular, widest basally; apex subtruncate; anterolateral angles poorly developed, obtusely rounded; anterior bead incomplete, obsolete medially; sides strongly rounded, slightly sinuate posteriorly; lateral grooves absent; setiferous punctures absent on each side; posterolateral angles obtusely rounded; laterobasal foveae ill-defined, finely punctate, shallow, very wide, not prolonged forward, double; posterior bead absent; base emarginate, much wider than pronotal apex, about as wide as elytral base. Legs. Short. Elytra. Strongly convex, oblong, widest basally. Basal margin complete, reaching about scutellar striole. Shoulders angular. Sides strongly rounded. Scutellar striole present. Striae complete, shallow, finely punctate; stria 3 with two or three setiferous punctures. Recurrent stria long, blunt. Intervals depressed. Abdomen. Last visible sternum (sternum VII): male with two apical ambulatory setae; female with four apical ambulatory setae. Aedeagus. Lateral view (Fig. 99): very strongly arcuate, narrowed in apical half; base strongly biconvex dorsally; middle strongly convex dorsally, slightly convex ventrally, with dorsal membranous area narrow anteriorly, moderately wide posteriorly, very long; apex slender, strongly convex dorsally, strongly concave ventrally, with extreme tip slender and curved downward. Dorsal view: moderately wide, asymmetrical (ostium of membranous area deflected to right); basal orifice wide, open anteriorly, moderately distant from membranous area. Parameres with two or three apical setae.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30269FFA43FCAFA17FEA9FC24.taxon	materials_examined	Material examined. 187 specimens (AMNZ, CMNZ, JNNZ, LUNZ, NZAC). Geographic distribution (Fig. 153). Subantarctic Islands: SN.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30269FFA43FCAFA17FEA9FC24.taxon	biology_ecology	Ecology. Lowland. Endogean. Wet peaty forests (Olearia). Shaded ground. Nocturnal; hides during the day in leaf litter, under fallen branches and logs, and in rotten pieces of wood. Gregarious. Biology. Seasonality: September – October, December – March. Tenerals: December, March, July. Predacious (based on mouthpart morphology). Occasionally infested with mites; regularly infested with fungi (Laboulbeniales). Dispersal power. Subapterous (incapable of flight). Slow runner. Collecting techniques. Sifting leaf litter; lifting branches and logs; breaking rotten branches.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30269FFA43FCAFA17FEA9FC24.taxon	type_taxon	Type species. Oopterus plicaticollis Blanchard, 1843, designated by Larochelle & Larivière (2001: 73).	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30269FFA43FCAFA17FEA9FC24.taxon	description	Description. Body length 3.7 – 5.7 mm; not ovate, narrowed around bases of pronotum and elytra. Color dark or pale. Metallic luster present or absent. Dorsal surface mostly glabrous. Head. Narrow or moderately wide. Mandibles moderately long. Labrum slightly emarginate. Eyes convex; two setiferous punctures on inner side of each eye (a single puncture posteriorly in P. otiraensis). Antennae submoniliform. Tempora inflated. Mentum tooth bifid (entire in other zoline genera). Mentum-submentum suture present as in other zoline genera (absent in P. plicaticollis). Ligula with two fused setae (with six setae in P. plicaticollis; two medial setae fused, four lateral setae free). Thorax. Pronotum cordate; lateral grooves absent (present in P. latipennis); two setiferous punctures on each side; posterolateral carinae present, absent, or obsolete; base narrower than apex or about as wide as apex. Scutellum visible, inserted entirely between elytral bases. Legs. Short or moderately long. Segments 1 + 2 of male protarsi with two rows of ventral scales (with a single row in other zoline genera). Elytra. Ovate or subovate. Basal margin complete, reaching about scutellum. Shoulders poorly or well developed, rounded. Scutellar setiferous pore present, inserted at junction of striae 1 + 2. Scutellar striole present or absent. Striae complete or incomplete, consisting of lines (replaced by rows of coarse punctures in P. otiraensis). Three discal setiferous punctures in stria or row 3; recurrent stria directed apically toward stria or row 5 or 7. Subapical seta present. Apical seta present. Sutural apices acute, obtuse, or rounded. Abdomen. Last visible sternum (sternum VII): male with two apical ambulatory setae (two additional medial ambulatory setae in P. plicaticollis); female with four apical ambulatory setae. Aedeagus. Lateral view: dorsal membranous area moderately wide. Dorsal view (Fig. 16): moderately wide; basal orifice wide, open anteriorly, moderately distant from membranous area; lateral lobes of basal bulb moderately distant (as in Maungazolus). Parameres with three or four apical setae. Right paramere curved, very long, narrow, untwisted medially.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30269FFA43FCAFA17FEA9FC24.taxon	discussion	Remarks. The genus Pseudoopterus Csiki, 1928 was synonymized with Oopterus by Jeannel (1940) without justification. This taxon is resurrected from synonymy here on the basis of the bifid mentum tooth, the two ventral rows of scales on segments 1 + 2 of the male protarsi, and the aedeagus with its dorsal membranous area moderately wide and moderately distant from the basal bulb. The genus Pseudoopterus now includes two newly described species as well as two species previously described in Oopterus.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30266FFA63FCAFF37FAC9FF67.taxon	description	Fig. 54, 100, 151 Holotype: male (NZAC) labeled: “ NEW ZEALAND WD Arthur’s Pass NP, Kellys Creek x Otira River [= Kelly Creek near Otira River Junction] 4248 S 17134 E 350 m 5. III. 2009 Larochelle and Larivière (typed) / Floodplain Podoc. - brleaf for. [= Podocarp-broadleaf forest] (Fuchsia-Horopito dom. [= dominant]): wet clayish soil. Under large, deeply embedded stones. (typed) / HOLOTYPE [male symbol] Pseudoopterus otiraensis Larochelle & Larivière, 2017 (red label; typed). ” Paratypes: one female (NZAC) and one male (AMNZ) from the same locality as the holotype, bearing blue paratype labels. Description. Body length 3.7 – 4.6 mm. Head and pronotum reddish, dark brown medially; elytra and abdomen dark brown; lateral margins and apex of elytra widely reddish; abdomen reddish laterally and apically; antennae, palpi, and legs reddish; antennal segments 1 + 2 and femora pale yellowish. Microsculpture absent. Iridescence absent. Very shiny, without metallic luster. Head. Labrum strongly transverse, slightly emarginate anteriorly. Antennae submoniliform, short: segment 1 (scape) short, stout, about 1.5 x longer than its maximum width. Frontal furrows wide, shallow, subparallel. Eyes slightly convex; a single setiferous puncture on inner side of each eye (posteriorly). Tempora moderately inflated (about half as long as eyes). Mentum: medial tooth bifid, moderately shorter than lateral lobes. Paraglossae membranous, prominent, about as long as ligula. Thorax. Pronotum strongly convex, coarsely punctate across base, wrinkled discally, strongly transverse, slightly cordate, widest before middle; apex subtruncate; anterolateral angles poorly developed, obtusely rounded; anterior bead incomplete, obsolete medially; sides strongly rounded, moderately sinuate posteriorly; lateral grooves absent; two setiferous punctures on each side; posterolateral angles acute, projected laterally; posterolateral carinae blunt, very long (about as long as adjoining foveae); laterobasal foveae well defined, coarsely punctate, very deep and wide, square, not prolonged forward, simple; posterior bead absent; base emarginate, about as wide as pronotal apex and elytral base. Legs. Short. Elytra. Strongly convex, ovate, widest about middle. Shoulders obtuse. Sides strongly rounded. Scutellar striole present. Striae replaced by rows of coarse punctures, deep; row 1 complete; rows 2 – 7 obsolete apically; row 3 with three setiferous punctures. Recurrent stria long, blunt, directed apically toward row 7. Intervals subdepressed. Sutural apices obtuse. Abdomen. Last visible sternum (sternum VII): male with two apical ambulatory setae; female with four apical ambulatory setae. Aedeagus. Lateral view (Fig. 100): moderately arcuate, slightly widened in apical half; base moderately convex dorsally; middle slightly convex dorsally, rather straight ventrally, with dorsal membranous area moderately wide, very long; apex subtriangular, almost straight dorsally, slightly concave ventrally, with extreme tip wide and long. Dorsal view: moderately wide, asymmetrical (ostium of membranous area deflected to right); apex straight; basal orifice wide, open anteriorly, moderately distant from membranous area. Parameres with three apical setae.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30266FFA63FCAFF37FAC9FF67.taxon	materials_examined	Material examined. 11 specimens (AMNZ, CMNZ, LUNZ, NZAC). Geographic distribution (Fig. 151). South Island: BR – Greymouth. WD – Arthur’s Pass National Park (Kelly Creek; Kelly Creek near Otira River Junction). Fox Glacier.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30266FFA63FCAFF37FAC9FF67.taxon	biology_ecology	Ecology. Lowland, montane. Endogean. Forests (broadleaf, podocarp) and scrublands. Shaded ground. Nocturnal; hides during the day under well-embedded stones. Gregarious. Biology. Seasonality: November – January, March, May. Predacious (based on mouthpart morphology). Dispersal power. Subapterous (incapable of flight). Slow runner. Collecting techniques. Lifting stones; pitfall trapping.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30266FFA63FCAFF37FAC9FF67.taxon	discussion	Remarks. This species is named after the type locality “ Kelly Creek near Otira River Junction ” (WD) and the Latin suffix - ensis, denoting a place, locality, or country. In addition to diagnostic characters of the male genitalia, Pseudoopterus otiraensis has the following distinguishing features: a single setiferous puncture on inner side of each eye (posteriorly); elytra without microsculpture, striae replaced by rows of coarse punctures. This species occurs in western areas of the South Island (BR, WD).	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30265FFA73FCAFEB7FE4CFDA4.taxon	description	Fig. 55, 101, 152 Oopterus plicaticollis Blanchard, 1843: pl. 2, fig. 15 (redescribed in 1853: 44). Type locality: Auckland Islands [AU]. Oopterus laticollis Kirsch, in Kiesenwetter and Kirsch 1877: 159. Type locality: Auckland Islands [AU]. Synonymized by Gourlay 1950: 181. Pseudoopterus plicaticollis: Csiki 1928: 225. Combination reinstated Pseudoopterus laticollis: Csiki 1928: 225. Oopterus plicaticollis: Hudson 1934: 177. Oopterus aucklandicus Brookes, 1951: 23. Type locality: Mount Raynal (Flat Topped Mountain), Auckland Islands [AU]. Synonymized by Johns 1974: 298. Description. Body length 4.1 – 4.5 mm. Head, pronotum, elytra, and abdomen mostly dark brown; margins of pronotum reddish; lateral margins, suture, and apex of elytra reddish; abdomen reddish laterally and apically; antennae, palpi, and legs reddish; femora pale yellow. Microsculpture strong, isodiametric. Iridescence absent. Very shiny, with metallic luster (aeneous) on head, pronotum, and elytra. Head. Labrum strongly transverse, slightly emarginate anteriorly. Antennae submoniliform, short: segment 1 (scape) moderately long, about 2 x longer than its maximum width. Frontal furrows wide, shallow, convergent. Eyes strongly convex; two setiferous punctures on inner side of each eye. Tempora moderately inflated (about half as long as eyes). Mentum: medial tooth bifid, moderately shorter than lateral lobes. Mentum-submentum suture absent (present in other zoline taxa). Ligula with six setae (two medial setae fused, four lateral setae free; as opposed to two fused setae only in other Pseudoopterus species). Paraglossae membranous, prominent, about as long as ligula. Thorax. Pronotum strongly convex, impunctate, coarsely wrinkled discally (finely wrinkled in other Pseudoopterus species), strongly transverse, slightly cordate, widest before middle; apex subtruncate; anterolateral angles poorly developed, rounded; anterior bead incomplete, obsolete medially; sides strongly rounded, slightly sinuate posteriorly; lateral grooves absent; two setiferous punctures on each side; posterolateral angles subrectangular; posterolateral carinae obsolete; laterobasal foveae well defined, impunctate, moderately deep, very wide, oblong, prolonged forward, simple; posterior bead absent; base emarginate, much narrower than pronotal apex, about as wide as elytral base. Legs. Short. Elytra. Moderately convex, ovate, widest about middle. Shoulders oblique. Sides strongly rounded. Scutellar striole absent. Striae complete, deep, impunctate or very finely punctate; stria 3 with three large setiferous punctures (small punctures in other Pseudoopterus species). Recurrent stria long, blunt, directed apically toward stria 5 or 7. Intervals slightly convex. Sutural apices obtuse. Abdomen. Last visible sternum (sternum VII): male with two apical and two medial ambulatory setae; female with four apical ambulatory setae. Aedeagus. Lateral view (Fig. 101): moderately arcuate, moderately widened in apical half; base strongly convex dorsally; middle moderately sinuate dorsally, strongly convex ventrally, with dorsal membranous area moderately wide, very long; apex subtriangular, slightly convex dorsally, moderately concave ventrally, with extreme tip narrow and long. Dorsal view: moderately wide, asymmetrical (ostium of membranous area deflected to right); apex deflected to the left; basal orifice wide, open anteriorly, moderately distant from membranous area. Parameres with three or four apical setae.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30265FFA73FCAFEB7FE4CFDA4.taxon	materials_examined	Material examined. 173 specimens (CMNZ, LUNZ, MONZ, NZAC). Geographic distribution (Fig. 152). Subantarctic Islands: AU.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30265FFA73FCAFEB7FE4CFDA4.taxon	biology_ecology	Ecology. Lowland, montane, subalpine, alpine. Epigean. Supralittoral zone; peaty wet forests (Olearia, Metrosideros), tussock grassfields; herbfields; fellfields. Open ground. Nocturnal; hides during the day in leaf litter, under logs and stones, and in peat. Gregarious. Biology. Seasonality: November, January – February, April, June, August. Tenerals: February. Predacious (based on mouthpart morphology). Occasionally infested with fungi (Laboulbeniales). Dispersal power. Subapterous (incapable of flight). Slow runner. Collecting techniques. Sifting leaf litter; lifting logs and stones; pitfall trapping; inspecting peat.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30265FFA73FCAFEB7FE4CFDA4.taxon	discussion	Remark. Pseudoopterus plicaticollis is the only New Zealand zoline species lacking a mentum – submentum suture.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30264FFA23FCAFD77FB55FCC4.taxon	description	Fig. 56, 102, 150	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30264FFA23FCAFD77FB55FCC4.taxon	description	Description. Body length 4.5 – 5.5 mm. Head, pronotum, elytra, and abdomen mostly piceous black; apex and base of head reddish; base of pronotum reddish; lateral margins and suture of elytra reddish; apex of elytra with two large reddish apical spots; abdomen yellow laterally and apically; antennae, palpi, and legs testaceous; base of femora reddish. Microsculpture absent on head and pronotum, strong and very transverse (with microlines) on entire elytra. Iridescence absent on head and pronotum, very strong on elytra. Very shiny, without metallic luster. Head. Labrum strongly transverse, slightly emarginate anteriorly. Antennae submoniliform, moderately long: segment 1 (scape) moderately long, about 2 x longer than its maximum width. Frontal furrows wide, deep, convergent. Eyes slightly convex; two setiferous punctures on inner side of each eye. Tempora moderately inflated (about two-thirds as long as eyes). Mentum: medial tooth bifid, moderately shorter than lateral lobes. Paraglossae membranous, prominent, slightly longer than ligula. Thorax. Pronotum moderately convex, coarsely punctate across base, wrinkled discally, strongly transverse and cordate, widest before middle; apex emarginate; anterolateral angles well developed, rounded; anterior bead incomplete, obsolete medially; sides strongly rounded, strongly sinuate posteriorly; lateral grooves moderately deep, very long (as long as pronotum); two setiferous punctures on each side; posterolateral angles acute, projected laterally; posterolateral carinae blunt, very long (about as long as adjoining foveae); laterobasal foveae well defined, coarsely punctate, very deep and wide, oblong, prolonged forward, simple; posterior bead absent; base emarginate, much narrower than pronotal apex and elytral base. Legs. Moderately long. Elytra. Moderately convex, subovate, widest about middle. Shoulders moderately rounded. Sides strongly rounded. Scutellar striole absent. Striae shallow, finely punctate; stria 1 complete; striae 2 – 7 incomplete; stria 3 with three setiferous punctures. Recurrent stria long, sharp, directed apically toward stria 7. Intervals subdepressed. Sutural apices obtusely rounded. Abdomen. Last visible sternum (sternum VII): male with two apical ambulatory setae; female with four apical ambulatory setae. Aedeagus. Lateral view (Fig. 102): moderately arcuate, slightly widened in apical half; base moderately convex dorsally; middle angulate anteriorly and slightly convex dorsally, moderately concave ventrally, with dorsal membranous area moderately wide, very long; apex subtriangular, almost straight dorsally, straight ventrally, with extreme tip wide and long. Dorsal view: moderately wide, asymmetrical (ostium of membranous area deflected to right); apex straight; basal orifice wide, open anteriorly, moderately distant from membranous area. Parameres with three or four apical setae.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30264FFA23FCAFD77FB55FCC4.taxon	materials_examined	Material examined. 82 specimens (AMNZ, BMNH, CMNZ, JNNZ, LUNZ, MONZ, NZAC). Geographic distribution (Fig. 150). South Island: BR, CO, MC, NN, OL, WD.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30264FFA23FCAFD77FB55FCC4.taxon	biology_ecology	Ecology. Lowland, montane, subalpine, alpine. Epigean-arboreal. Forests (beech, podocarp, broadleaf) and scrublands, along streams and seepages; herbfields. Shaded or open ground. Nocturnal; active at night on moss and trees; hides during the day under stones and in moss. Gregarious. Biology. Seasonality: October – March, May. Predacious (based on mouthpart morphology). Dispersal power. Subapterous (incapable of flight). Moderate runner. Occasional climber on trees. Collecting techniques. Lifting stones; sifting moss.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30264FFA23FCAFD77FB55FCC4.taxon	discussion	Remark. Examination of the types of Oopterus probus and O. latifossus revealed them to be conspecific with Pseudoopterus latipennis. Pseudoopterus fi ordlandensis new species Fig. 57, 103, 149 Holotype: male (NZAC) labeled: “ NEW ZEALAND FD Takahe Valley Murchison Mts [= Mountains] 11 Dec 1972 A. C. Eyles (typed) / Litter (typed) 72 / 271 (hand-written) / JOHN NUNN (typed) / [male symbol] / HOLOTYPE [male symbol] Pseudoopterus fiordlandensis Larochelle & Larivière, 2017 (red label; typed). ” Paratype: one female (NZAC) from the same locality as the holotype, bearing a blue paratype label.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30264FFA23FCAFD77FB55FCC4.taxon	description	Description. Body length 5.3 – 5.7 mm. Head, pronotum, elytra, and abdomen mostly piceous black; base of pronotum reddish; lateral margins and suture of elytra reddish; extreme apex of elytra with two small reddish apical spots; abdomen yellow laterally and apically; antennae, palpi, and legs reddish; femora pale yellow. Microsculpture absent on head, pronotum, and in basal half of elytra, feeble and very transverse (with microlines) in apical half of elytra. Iridescence absent on head, pronotum, and in basal half of elytra, feeble in apical half of elytra. Very shiny, without metallic luster. Head. Labrum strongly transverse, slightly emarginate anteriorly. Antennae submoniliform, moderately long: segment 1 (scape) moderately long, about 2 x longer than its maximum width. Frontal furrows wide, deep, convergent. Eyes slightly convex; two setiferous punctures on inner side of each eye. Tempora moderately inflated (about as long as two-thirds of eyes). Mentum: medial tooth bifid, moderately shorter than lateral lobes. Paraglossae membranous, prominent, slightly longer than ligula. Thorax. Pronotum moderately convex, coarsely punctate across base, wrinkled discally, strongly transverse, moderately cordate, widest before middle; apex emarginate; anterolateral angles well developed, rounded; anterior bead incomplete, obsolete medially; sides strongly rounded, slightly sinuate posteriorly; lateral grooves absent; two setiferous punctures on each side; posterolateral angles rectangular; posterolateral carinae absent; laterobasal foveae well defined, coarsely punctate, very deep and wide, oblong, prolonged forward, simple; posterior bead absent; base emarginate, much narrower than pronotal apex and elytral base. Legs. Moderately long. Elytra. Strongly convex, subovate, widest about middle. Shoulders moderately rounded. Sides strongly rounded. Scutellar striole present. Striae shallow, finely punctate; striae 1 – 7 complete; stria 3 with three setiferous punctures. Recurrent stria long, sharp, directed apically toward stria 7. Intervals depressed. Sutural apices acute. Abdomen. Last visible sternum (sternum VII): male with two apical ambulatory setae; female with four apical ambulatory setae. Aedeagus. Lateral view (Fig. 103): moderately arcuate, slightly widened in apical half; base moderately convex dorsally; middle moderately convex dorsally, strongly concave ventrally, with dorsal membranous area moderately wide and long; apex subtriangular, slightly convex dorsally, straight ventrally, with extreme tip wide and long. Dorsal view: moderately wide, asymmetrical (ostium of membranous area deflected to right); apex straight; basal orifice wide, open anteriorly, moderately distant from membranous area. Parameres not seen (lost during dissection).	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30264FFA23FCAFD77FB55FCC4.taxon	materials_examined	Material examined. 4 specimens (JNNZ, LUNZ, NZAC). Geographic distribution (Fig. 149). South Island: FD – Fiordland National Park (Murchison Mountains; Murchison Mountains, Takahe Valley).	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30264FFA23FCAFD77FB55FCC4.taxon	biology_ecology	Ecology. Montane. Epigean. Forests (beech). Shaded ground. Nocturnal; hides during the day in leaf litter. Biology. Seasonality: December. Predacious (based on mouthpart morphology). Dispersal power. Subapterous (incapable of flight). Moderate runner. Collecting techniques. Sifting leaf litter; pitfall trapping.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30264FFA23FCAFD77FB55FCC4.taxon	discussion	Remarks. The name of this species is based on the geographic region of Fiordland where the beetle occurs, and the Latin suffix - ensis, denoting a place, locality, or country. Pseudoopterus fiordlandensis is morphologically close to P. latipennis. In addition to diagnostic characters of the male genitalia, P. fiordlandensis has the following distinguishing features: pronotum lacking lateral grooves and posterolateral carinae, posterolateral angles rectangular; elytra strongly convex, microsculpture present in apical half only, striae 1 – 7 complete, extreme apex with two small reddish spots. The two species are allopatric: P. fiordlandensis is known only from the Fiordland region (South Island), while P. latipennis occurs both in western and southern areas of the South Island (NN to OL – CO). Genus Maungazolus, new Fig. 58 – 65, 104 – 110, 112 – 119	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30264FFA23FCAFD77FB55FCC4.taxon	type_taxon	Type species. Oopterus pallidipes Broun, 1893, by present designation.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30264FFA23FCAFD77FB55FCC4.taxon	description	Description. Body length 2.8 – 4.4 mm; not ovate, narrowed around bases of pronotum and elytra. Color dark or pale. Metallic luster absent (present in M. septempunctatus). Dorsal surface mostly glabrous. Head. Moderately wide. Mandibles moderately long. Labrum slightly emarginate anteriorly. Eyes convex; two setiferous punctures on inner side of each eye. Antennae submoniliform. Tempora inflated or not. Mentum tooth entire (acute, rounded, truncate, or slightly emarginate apically). Ligula with four setae (two medial setae fused, two lateral setae free). Thorax. Pronotum cordate; lateral grooves absent (present in M. septempunctatus); two setiferous punctures on each side; posterolateral carinae absent or obsolete; base narrower than apex. Scutellum visible, inserted entirely between elytral bases. Legs. Short (moderately long in M. priestleyensis). Segments 1 + 2 of male protarsi with a single row of ventral scales. Elytra. Ovate or subovate. Basal margin incomplete (reaching about stria 3 or stria 4) or complete (reaching about scutellum). Shoulders poorly or well developed, rounded or obtuse. Scutellar setiferous pore present, inserted at junction of striae 1 + 2. Scutellar striole present or absent. Striae complete or incomplete, consisting of lines (replaced by rows of coarse punctures in M. parvulus); three discal setiferous punctures in stria or row 3 (four punctures in M. pygmeatus; seven punctures in M. septempunctatus); recurrent stria directed apically toward stria or row 5. Subapical seta present (absent in M. pygmeatus). Apical seta present. Sutural apices obtuse or rounded. Abdomen. Last visible sternum (sternum VII): male with two apical ambulatory setae (with two additional medial ambulatory setae in M. septempunctatus); female with four apical ambulatory setae. Aedeagus. Lateral view: dorsal membranous area very wide (moderately wide or narrow in other zoline genera). Dorsal view (Fig. 17): very narrow (wide in other zoline genera, rarely moderately narrow in Oopterus (part )); basal orifice narrow, open anteriorly, very close to membranous area (distant from membranous area in other zoline genera, rarely moderately close); lateral lobes of basal bulb moderately distant. Parameres with two to five apical setae. Right paramere curved, very long, narrow, untwisted medially.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30264FFA23FCAFD77FB55FCC4.taxon	discussion	Remarks. The generic name is derived from the Maori noun maunga, mountain and Zolus, the type genus of the tribe Zolini. The morphological characters unifying species of Maungazolus are: mentum tooth entire (not bifid); segments 1 + 2 of male protarsi with a single row of ventral scales; aedeagus with the dorsal membranous area very wide and very close to the basal bulb. The new genus includes five newly described species as well as three species previously described in Oopterus.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30260FFDC3FCAFC97FE00F9C4.taxon	description	Fig. 58, 104, 116	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30260FFDC3FCAFC97FE00F9C4.taxon	description	Description. Body length 3.0 – 4.2 mm. Head, pronotum, elytra, and abdomen mostly piceous black; extreme apex of elytra reddish; abdomen reddish laterally and apically; antennae, palpi, and legs reddish; antennal segments 1 + 2 and femora pale yellow, femora slightly contrasting with tibiae. Microsculpture obsolete on head, absent on pronotum and elytra. Iridescence absent. Very shiny, without metallic luster. Head. Labrum strongly transverse, slightly emarginate anteriorly. Antennae submoniliform, short: segment 1 (scape) short, stout, about 1.5 x longer than its maximum width. Frontal furrows wide, deep, convergent. Eyes slightly convex; two setiferous punctures on inner side of each eye. Tempora moderately inflated (about half as long as eyes). Mentum: medial tooth entire, acute apically, moderately shorter than lateral lobes. Paraglossae membranous, prominent, slightly longer than ligula. Thorax. Pronotum moderately or strongly convex, usually coarsely punctate (rarely finely punctate) across base, wrinkled discally, moderately transverse and cordate, widest before middle; apex subtruncate; anterolateral angles poorly developed, rounded; anterior bead incomplete, obsolete medially; sides strongly rounded, not sinuate posteriorly; lateral grooves absent; two setiferous punctures on each side; posterolateral angles subrectangular; posterolateral carinae absent; laterobasal foveae well defined, usually coarsely punctate (rarely finely punctate or impunctate), very deep, moderately wide, rounded or oblong, not prolonged forward, simple; posterior bead absent; sub-basal transverse impression deep, coarsely punctate (with 8 to 10 punctures); basal transverse impression deep, coarsely punctate (with 8 to 10 punctures); base emarginate, much narrower than pronotal apex, about as wide as elytral base. Legs. Short. Elytra. Strongly convex, subovate, widest about middle. Basal margin complete, reaching about scutellum. Shoulders rounded or obtuse. Sides strongly rounded. Scutellar striole obsolete. Stria 1 complete; striae 2 – 7 incomplete; striae 1 – 4 shallow, finely punctate; striae 5 – 7 obsolete; stria 3 with four setiferous punctures. Recurrent stria long, sharp, directed apically toward stria 5. Subapical seta absent (present in other Maungazolus species). Intervals depressed. Sutural apices obtuse. Abdomen. Last visible sternum (sternum VII): male with two apical ambulatory setae; female with four apical ambulatory setae. Aedeagus. Lateral view (Fig. 104): moderately arcuate, very strongly widened in apical half; base strongly biconvex dorsally; middle moderately concave dorsally, strongly convex ventrally, with dorsal membranous area very wide and long; apex subtriangular, unusually short, moderately convex dorsally, very concave ventrally, with extreme tip wide, unusually short and turned downward. Dorsal view: very narrow, almost thin, asymmetrical (ostium of membranous area deflected to right); apex straight; basal orifice narrow, very close to membranous area. Parameres with three or four apical setae.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30260FFDC3FCAFC97FE00F9C4.taxon	materials_examined	Material examined. 141 specimens (AMNZ, BMNH, JNNZ, LUNZ, MONZ, NZAC). Geographic distribution (Fig. 116). South Island: BR, CO, FD, MC, MK, OL, SL, WD.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30260FFDC3FCAFC97FE00F9C4.taxon	biology_ecology	Ecology. Subalpine, alpine. Epigean. Mostly alpine fellfields and herbfields; also subalpine shrublands. Open or shaded ground. Nocturnal; active at night on moss; hides during the day under stones, in leaf litter and moss. Gregarious. Biology. Seasonality: October – November, January – March, May. Tenerals: October (SL), January (FD, WD). Predacious (based on mouthpart morphology). Occasionally infested with fungi (Laboulbeniales). Dispersal power. Subapterous (incapable of flight). Slow runner. Occasional climber on moss. Collecting techniques. Lifting stones; sifting leaf litter and moss.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30260FFDC3FCAFC97FE00F9C4.taxon	discussion	Remarks. Examination of the types of Oopterus basalis and O. minor revealed them to be conspecific with M. pygmeatus. This is the most widespread and most variable of all Maungazolus species (color; microsculpture of head; convexity; posterolateral angles and laterobasal foveae of pronotum; basal margin and shoulders of elytra). In addition, M. pygmeatus is the only Maungazolus species lacking the elytral subapical seta.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3021FFFDD3FCAF957FA5FF8A7.taxon	description	Fig. 59, 105, 118 Holotype: male (NZAC) labeled: “ [NEW ZEALAND SI] Table Hill Stewart I. (typed) / 2000 ’ 16. II. 68 J. McBurney (typed) / Mat plants (typed) 68 / 55 (hand-written) / [male symbol] / HOLOTYPE [male symbol] Maungazolus septempunctatus Larochelle & Larivière, 2017 (red label; typed). ” Paratypes: one female (NZAC) and one male (CMNZ) from the same locality as the holotype, bearing blue paratype labels. Description. Body length 3.1 – 4.4 mm. Head, pronotum, and elytra mostly piceous black; lateral margins, suture, and extreme apex of elytra reddish; abdomen mostly reddish piceous, reddish laterally and apically; antennae, palpi, and legs reddish; antennal segment 1 (scape) and femora bright yellow, femora strongly contrasting with tibiae. Microsculpture strong and isodiametric on head, absent on pronotum and elytra. Iridescence absent. Very shiny, with metallic luster (aeneous) on head, pronotum, and elytra. Forebody narrow in comparison to elytra (wide in other Maungazolus species). Head. Labrum strongly transverse, slightly emarginate anteriorly. Antennae submoniliform, short: segment 1 (scape) moderately long, about 2 x longer than its maximum width. Frontal furrows wide, shallow, convergent. Eyes moderately convex; two setiferous punctures on inner side of each eye. Tempora not inflated. Mentum: medial tooth entire, acute apically, moderately shorter than lateral lobes. Paraglossae membranous, prominent, slightly longer than ligula. Thorax. Pronotum strongly convex, coarsely punctate across base, unwrinkled, slightly transverse, strongly cordate, widest before middle; apex subtruncate; anterolateral angles poorly developed, rounded; anterior bead incomplete, obsolete medially; sides strongly rounded, strongly sinuate posteriorly; lateral grooves shallow, very long (about as long as adjoining foveae; absent in other Maungazolus species); two setiferous punctures on each side; posterolateral angles rectangular; posterolateral carinae absent; laterobasal foveae ill-defined, coarsely punctate, shallow to moderately deep, moderately wide, not prolonged forward, simple; posterior bead absent; sub-basal transverse impression well developed, coarsely punctate (with about 8 punctures); basal transverse impression well developed, coarsely punctate (with about 10 punctures); base emarginate, much narrower than pronotal apex, about as wide as elytral base. Legs. Short. Elytra. Strongly convex, ovate, widest about middle. Basal margin incomplete, reaching about stria 3. Shoulders rounded. Sides strongly rounded. Scutellar striole present. Striae complete, finely punctate; stria 1 deep; striae 2 – 7 shallow; stria 3 with seven setiferous punctures. Recurrent stria long, blunt, directed apically toward stria 5. Subapical seta present. Intervals subdepressed. Sutural apices obtusely rounded. Abdomen. Last visible sternum (sternum VII): male with two apical and two medial ambulatory setae (without medial setae in other Maungazolus species); female with four apical ambulatory setae. Aedeagus. Lateral view (Fig. 105): moderately arcuate, not widened in apical half; base slightly convex dorsally; middle slightly convex dorsally, mostly straight ventrally, with dorsal membranous area very wide and long; apex subtriangular, strongly concave dorsally, vaguely slightly convex, with extreme tip wide and short. Dorsal view: very narrow, almost thin, asymmetrical (ostium of membranous area deflected to right); apex straight; basal orifice narrow, very close to membranous area. Parameres with two or three apical setae.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3021FFFDD3FCAF957FA5FF8A7.taxon	materials_examined	Material examined. 11 specimens (CMNZ, JNNZ, NZAC). Geographic distribution (Fig. 118). Stewart Island: Mount Anglem. Table Hill.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3021FFFDD3FCAF957FA5FF8A7.taxon	biology_ecology	Ecology. Alpine. Epigean. Herbfields. Open ground. Nocturnal; hides during the day in mat plants. Gregarious. Biology. Seasonality: February. Predacious (based on mouthpart morphology). Dispersal power. Subapterous (incapable of flight). Slow runner. Collecting technique. Sifting mat plants.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3021FFFDD3FCAF957FA5FF8A7.taxon	discussion	Remarks. The species name is derived from the Latin words septem (seven) and punctatus (punctate), referring to the presence of seven setiferous punctures in elytral stria 3. In addition to diagnostic characters of the male genitalia, Maungazolus septempunctatus has the following distinguishing features: metallic luster present on head, pronotum, and elytra; forebody narrow in comparison to elytra; seven setiferous punctures in elytral stria 3. Maungazolus septempunctatus is restricted to Stewart Island, while other Maungazolus species occur on the South Island and in southern areas of the North Island.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3021DFFDF3FCAFF37FA65FE07.taxon	description	Fig. 60, 106, 115 Holotype: male (NZAC) labeled: “ [NEW ZEALAND BR] Mt Priestly [= Mount Priestley] 915 m J. G. McBurney (typed) / 10 Dec 1969 Litter (typed) 69 / 241 (hand-written) / TENERAL (typed) / [male symbol] / HOLOTYPE [male symbol] Maungazolus priestleyensis Larochelle & Larivière, 2017 (red label; typed). ” Paratypes: one female (NZAC) from the same locality as the holotype and one male (MONZ) from Mount Dewar (BR), bearing blue paratype labels. Description. Body length 4.0 – 4.3 mm. Head, pronotum, and elytra mostly dark reddish (forebody slightly paler); lateral margins, suture, and extreme apex of elytra pale reddish; abdomen mostly reddish piceous, reddish laterally and apically; antennae, palpi, and legs reddish; femora pale yellow, slightly contrasting with tibiae. Microsculpture absent. Iridescence absent. Very shiny, without metallic luster. Head. Labrum strongly transverse, slightly emarginate anteriorly. Antennae submoniliform, moderately long: segment 1 (scape) moderately long, about 2 x longer than its maximum width. Frontal furrows wide, deep, convergent. Eyes slightly convex; two setiferous punctures on inner side of each eye. Tempora moderately inflated (about half as long as eyes). Mentum: medial tooth entire, acute apically, moderately shorter than lateral lobes. Paraglossae membranous, prominent, slightly longer than ligula. Thorax. Pronotum strongly convex, coarsely punctate across base, virtually unwrinkled, strongly transverse and cordate, widest before middle; apex subtruncate; anterolateral angles poorly developed, rounded; anterior bead incomplete, obsolete medially; sides strongly rounded, strongly sinuate posteriorly; lateral grooves absent; two setiferous punctures on each side; posterolateral angles subrectangular; posterolateral carinae absent; laterobasal foveae well defined, coarsely punctate, very deep, moderately wide, rounded, not prolonged forward, simple; posterior bead absent; sub-basal transverse impression well developed, coarsely punctate (with 10 to 12 punctures); basal transverse impression well developed, coarsely punctate (with about 10 punctures); base emarginate, much narrower than pronotal apex, about as wide as elytral base. Legs. Moderately long (short in other Maungazolus species). Elytra. Strongly convex, subovate, widest about middle. Basal margin complete, reaching about scutellum. Shoulders rounded. Sides strongly rounded. Scutellar striole absent. Stria 1 complete; striae 2 – 7 incomplete; striae 1 – 4 deep, coarsely punctate; striae 5 – 7 obsolete; stria 3 with three setiferous punctures. Recurrent stria long, sharp, directed apically toward stria 5. Subapical seta present. Intervals 1 – 3 slightly convex, 4 – 7 depressed. Sutural apices obtusely rounded. Abdomen. Last visible sternum (sternum VII): male with two apical ambulatory setae; female with four apical ambulatory setae. Aedeagus. Lateral view (Fig. 106): moderately arcuate, strongly widened in apical half; base moderately convex dorsally; middle biconvex dorsally, strongly convex ventrally, with dorsal membranous area very wide and long; apex subtriangular, strongly convex dorsally, slightly convex ventrally, with extreme tip wide and very long. Dorsal view: very narrow, almost thin, asymmetrical (ostium of membranous area deflected to right); apex straight; basal orifice narrow, very close to membranous area. Parameres with two or three apical setae.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3021DFFDF3FCAFF37FA65FE07.taxon	materials_examined	Material examined. 5 specimens (MONZ, NZAC). Geographic distribution (Fig. 115). South Island: BR – Mount Dewar. Mount Priestley.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3021DFFDF3FCAFF37FA65FE07.taxon	biology_ecology	Ecology. Montane. Epigean. Wet forest (beech). Shaded ground. Nocturnal; hides during the day in leaf litter and moss. Biology. Seasonality: December. Predacious (based on mouthpart morphology). Dispersal power. Subapterous (incapable of flight). Moderate runner. Collecting techniques. Sifting leaf litter and moss.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3021DFFDF3FCAFF37FA65FE07.taxon	discussion	Remarks. This species is named after the type locality, Mount Priestley (BR) and the Latin suffix - ensis, denoting a place, locality, or country. Maungazolus priestleyensis is morphologically close to M. acutus and M. parvulus. In addition to diagnostic characters of the male genitalia, M. priestleyensis has the following distinguishing features: pronotum with posterolateral angles subrectangular; elytra with basal margin complete, reaching about scutellum; body length 4.0 – 4.3 mm. Maungazolus priestleyensis is so far known from Mount Dewar and Mount Priestley (BR) only, while M. parvulus and M. acutus are more broadly distributed on the South Island in the northwest (BR, NN) and southwest (FD), respectively.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3021CFFD83FCAFE17FDCCFD87.taxon	description	Fig. 61, 114	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3021CFFD83FCAFE17FDCCFD87.taxon	description	Description. Body length 2.9 – 3.1 mm. Head yellowish, infuscated discally; pronotum and elytra yellowish, dark brown discally; elytra with suture reddish, sides and apex widely yellowish; abdomen mostly dark brown, yellowish laterally and apically; antennal segments 1 + 2, palpi, and legs yellowish; antennal segments 3 – 11 reddish. Microsculpture absent. Iridescence absent. Very shiny, without metallic luster. Head. Labrum strongly transverse, slightly emarginate anteriorly. Antennae submoniliform, short: segment 1 (scape) short, stout, about 1.5 x longer than its maximum width. Frontal furrows wide, deep, convergent. Eyes moderately convex; two setiferous punctures on inner side of each eye. Tempora slightly inflated (about one third as long as eyes). Mentum: medial tooth entire, rounded apically, moderately shorter than lateral lobes. Paraglossae membranous, prominent, slightly longer than ligula. Thorax. Pronotum strongly convex, coarsely punctate across base and finely punctate apically, unwrinkled, strongly transverse and cordate, widest before middle; apex subtruncate; anterolateral angles poorly developed, rounded; anterior bead incomplete, obsolete medially; sides strongly rounded, not sinuate posteriorly; lateral grooves absent; two setiferous punctures on each side; posterolateral angles acute, projected laterally; posterolateral carinae obsolete; laterobasal foveae well defined, coarsely punctate, moderately deep and wide, rounded, not prolonged forward, simple; posterior bead absent; sub-basal transverse impression well developed, coarsely punctate (with about 10 punctures); basal transverse impression well developed, coarsely punctate (with about 10 punctures); base subtruncate, much narrower than pronotal apex, about as wide as elytral base. Legs. Short. Elytra. Strongly convex, ovate, widest about middle. Basal margin incomplete, reaching about stria 4. Shoulders rounded. Sides strongly rounded. Scutellar striole obsolete. Striae replaced by rows of coarse punctures, very deep; row 1 complete; rows 2 – 7 incomplete; row 3 with three setiferous punctures. Recurrent stria long, blunt, directed apically toward row 5. Subapical seta present. Intervals moderately convex. Sutural apices obtuse. Abdomen. Last visible sternum (sternum VII): male with two apical ambulatory setae; female with four apical ambulatory setae. Aedeagus. Not seen (no specimen available for dissection).	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3021CFFD83FCAFE17FDCCFD87.taxon	materials_examined	Material examined. 4 specimens (BMNH, NZAC). Geographic distribution (Fig. 114). South Island: BR – Punakaiki. NN – Westport.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3021CFFD83FCAFE17FDCCFD87.taxon	biology_ecology	Ecology. Lowland. Epigean. Wet forests (beech). Shaded ground. Nocturnal; hides during the day in moss, and under logs and stones. Biology. Seasonality: October. Tenerals: March (BR). Predacious (based on mouthpart morphology). Dispersal power. Subapterous (incapable of flight). Slow runner. Collecting techniques. Sifting moss; lifting logs and stones.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3021CFFD83FCAFE17FDCCFD87.taxon	discussion	Remark. Oopterus parvulus was described by Broun (1903: 610) based on two male syntypes mounted together on a white rectangular label (BMNH); the male on the left is selected as the lectotype, while the male on the right becomes a paralectotype. These type designations are made to preserve stability of nomenclature in the future.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3021BFFD93FCAFD97FDD2FC87.taxon	description	Fig. 62, 107, 112 Holotype: male (NZAC) labeled: “ NEW ZEALAND FD Takahe Valley Murchison Mts [= Mountains] 11 Dec 1972 A. C. Eyles (typed) / Litter (typed) 72 / 271 (hand-written) / [male symbol] / HOLOTYPE [male symbol] Maungazolus acutus Larochelle & Larivière, 2017 (red label; typed). ” Paratypes: one female (NZAC) and one male (AMNZ) from the same locality as the holotype, bearing blue paratype labels. Description. Body length 2.8 – 3.5 mm. Head, pronotum, and elytra mostly piceous black; elytra with suture reddish, sides and apex narrowly pale reddish; abdomen mostly reddish black, reddish laterally and apically; antennae, palpi, and legs pale reddish; femora mostly bright yellow, strongly contrasting with tibiae, reddish basally. Microsculpture feeble and isodiametric on head, absent on pronotum and elytra. Iridescence absent. Very shiny, without metallic luster. Head. Labrum strongly transverse, slightly emarginate anteriorly. Antennae submoniliform, short: segment 1 (scape) short, stout, about 1.5 x longer than its maximum width. Frontal furrows wide, deep, convergent. Eyes moderately convex; two setiferous punctures on inner side of each eye. Tempora slightly inflated (about one third as long as eyes). Mentum: medial tooth entire, acute apically, moderately shorter than lateral lobes. Paraglossae membranous, prominent, slightly longer than ligula. Thorax. Pronotum strongly convex, coarsely punctate across base, wrinkled discally, strongly transverse, moderately cordate, widest before middle; apex subtruncate; anterolateral angles poorly developed, rounded; anterior bead incomplete, obsolete medially; sides strongly rounded, slightly sinuate posteriorly; lateral grooves absent; two setiferous punctures on each side; posterolateral angles unusually acute, projected laterally; posterolateral carinae absent; laterobasal foveae well defined, coarsely punctate, very deep, moderately wide, rounded, not prolonged forward, simple; posterior bead absent; sub-basal transverse impression well developed, coarsely punctate (with about 10 punctures); basal transverse impression well developed, coarsely punctate (with about 8 punctures); base emarginate, much narrower than pronotal apex, about as wide as elytral base. Legs. Short. Elytra. Strongly convex, ovate, widest about middle. Basal margin incomplete, reaching about stria 4. Shoulders rounded. Sides strongly rounded. Scutellar striole present. Striae finely punctate; stria 1 complete, deep; striae 2 – 4 incomplete, shallow; striae 5 – 7 obsolete; stria 3 with three setiferous punctures. Recurrent stria long, sharp, directed apically toward stria 5. Subapical seta present. Intervals depressed. Sutural apices obtuse. Abdomen. Last visible sternum (sternum VII): male with two apical ambulatory setae; female with four apical ambulatory setae. Aedeagus. Lateral view (Fig. 107): slightly arcuate, moderately widened in apical half; base mostly straight dorsally; middle strongly convex dorsally, mostly straight ventrally, with dorsal membranous area very wide and long; apex subtriangular, slightly convex dorsally, straight ventrally, with extreme tip wide and short. Dorsal view: very narrow, almost thin, asymmetrical (ostium of membranous area deflected to right); apex straight; basal orifice narrow, very close to membranous area. Parameres with four apical setae.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3021BFFD93FCAFD97FDD2FC87.taxon	materials_examined	Material examined. 79 specimens (AMNZ, JNNZ, LUNZ, NZAC). Geographic distribution (Fig. 112). South Island: FD.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3021BFFD93FCAFD97FDD2FC87.taxon	biology_ecology	Ecology. Lowland, montane, subalpine, alpine. Epigean-arboreal. Wet subalpine forests (beech); alpine herbfields and fellfields. Shaded or open ground. Nocturnal; active at night on trees; hides during the day in leaf litter, tussock litter, moss, and under stones. Gregarious. Biology. Seasonality: December – February, April. Tenerals: November. Predacious (based on mouthpart morphology). Dispersal power. Subapterous (incapable of flight). Slow runner. Occasional climber on trees. Collecting techniques. Sifting leaf litter, tussock litter, and moss; lifting stones.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3021BFFD93FCAFD97FDD2FC87.taxon	discussion	Remarks. The name of this species is derived from the Latin adjective acutus (acute), referring to the shape of the posterolateral angles of the pronotum. Maungazolus acutus is morphologically close to M. parvulus. In addition to diagnostic characters of the male genitalia, M. acutus has the following distinguishing features: antennae entirely pale reddish; head, pronotum, and elytra mostly piceous black; elytra with sides and apex narrowly pale reddish, striae present, finely punctate, and intervals depressed; mentum with medial tooth acute apically. The two species are allopatric: M. acutus is known only from the Fiordland region (southwestern South Island), while M. parvulus occurs in the northwest of the South Island (BR, NN).	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3021AFFDA3FCAFC97FDB0FBA7.taxon	description	Fig. 63, 108, 113	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3021AFFDA3FCAFC97FDB0FBA7.taxon	description	Description. Body length 3.8 – 4.1 mm. Head reddish, infuscated discally; pronotum pale reddish, brownish discally; elytra reddish, brownish discally, sides and apex widely pale yellow; abdomen mostly dark reddish, pale reddish laterally and apically; antennae, palpi, and legs testaceous; femora pale yellow, strongly contrasting with tibiae. Microsculpture absent. Iridescence absent. Very shiny, without metallic luster. Head. Labrum strongly transverse, slightly emarginate anteriorly. Antennae submoniliform, short: segment 1 (scape) short, stout, about 1.5 x longer than its maximum width. Frontal furrows wide, deep, convergent. Eyes moderately convex; two setiferous punctures on inner side of each eye. Tempora not inflated. Mentum: medial tooth entire, truncate apically, moderately shorter than lateral lobes. Paraglossae membranous, prominent, slightly longer than ligula. Thorax. Pronotum strongly convex, coarsely punctate across base and finely punctate apically, wrinkled discally, slightly transverse, moderately cordate, widest about middle; apex subtruncate; anterolateral angles poorly developed, rounded; anterior bead incomplete, obsolete medially; sides strongly rounded, not sinuate posteriorly; lateral grooves absent; two setiferous punctures on each side; posterolateral angles rectangular; posterolateral carinae obsolete; laterobasal foveae well defined, coarsely punctate, moderately deep and wide, rounded, not prolonged forward, simple; posterior bead absent; sub-basal transverse impression replaced by two foveolae; basal transverse impression well developed, coarsely punctate (with about 10 punctures); base emarginate, much narrower than pronotal apex, about as wide as elytral base. Legs. Short. Elytra. Strongly convex, ovate, widest about middle. Basal margin incomplete, reaching about stria 4. Shoulders rounded. Sides strongly rounded. Scutellar striole present. Striae coarsely punctate; stria 1 complete, very deep; striae 2 – 4 incomplete, moderately deep; striae 5 – 7 obsolete; stria 3 with three setiferous punctures. Recurrent stria long, sharp, directed apically toward stria 5. Subapical seta present. Intervals moderately convex. Sutural apices rounded. Abdomen. Last visible sternum (sternum VII): male with two apical ambulatory setae; female with four apical ambulatory setae. Aedeagus. Lateral view (Fig. 108): strongly arcuate, very strongly widened in apical half; base moderately convex dorsally; middle moderately sinuate dorsally, strongly convex ventrally, with dorsal membranous area very wide and long; apex triangular, slightly concave dorsally, moderately concave ventrally, with extreme tip narrow and short. Dorsal view: very narrow, almost thin, asymmetrical (ostium of membranous area deflected to right); apex straight; basal orifice narrow, very close to membranous area. Parameres with two apical setae.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3021AFFDA3FCAFC97FDB0FBA7.taxon	materials_examined	Material examined. 149 specimens (BMNH, JNNZ, LUNZ, MONZ, NZAC). Geographic distribution (Fig. 113). South Island: NN.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3021AFFDA3FCAFC97FDB0FBA7.taxon	biology_ecology	Ecology. Montane, subalpine, alpine. Epigean-arboreal. Wet forests (beech); alpine tussock grasslands and fellfields. Shaded (usually) or open ground. Nocturnal; active at night on mossy, logs, and trees; hides during the day in moss and leaf litter. Gregarious. Biology. Seasonality: November – March. Tenerals: March, May. Predacious (based on mouthpart morphology). Dispersal power. Subapterous (incapable of flight). Slow runner. Occasional climber on trees. Collecting techniques. Sifting moss and leaf litter.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D3021AFFDA3FCAFC97FDB0FBA7.taxon	discussion	Remark. Broun (1887: 604) cited Oopterus pallidipes from Mount Arthur, NN, without providing any description. This comment and Broun’s annotation of his personal copy of his manual suggest Mount Arthur, NN, as the type locality.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30219FFDB3FCAFB77FB39FA07.taxon	description	Fig. 64, 109, 117 Holotype: male (NZAC) labeled: “ [NEW ZEALAND MB] Fell Pk. [= Mount Fell] 1296 m (typed) / Richmond Ra. [= Range] (typed) / 13 March 69 J. C. Watt (typed) / Litter (typed) 69 / 113 (hand-written) / [male symbol] / HOLOTYPE [male symbol] Maungazolus ranatungae Larochelle & Larivière, 2017 (red label; typed). ” Paratypes: one female (NZAC) and one male (LUNZ) from the same locality as the holotype, bearing blue paratype labels. Description. Body length 2.9 – 3.4 mm. Head and pronotum dark brown; elytra yellowish, dark brown discally, base dark brown medially, sides and apex widely pale yellow; abdomen mostly reddish piceous, reddish laterally and apically; antennae, palpi, and legs reddish; antennal segments 1 + 2 yellow; femora bright yellow, strongly contrasting with tibiae. Microsculpture absent. Iridescence absent. Very shiny, without metallic luster. Head. Labrum strongly transverse, slightly emarginate anteriorly. Antennae submoniliform, short: segment 1 (scape) short, stout, about 1.5 x longer than its maximum width. Frontal furrows wide, deep, convergent. Eyes slightly convex; two setiferous punctures on inner side of each eye. Tempora moderately inflated (about half as long as eyes). Mentum: medial tooth entire, slightly emarginate apically, moderately shorter than lateral lobes. Paraglossae membranous, prominent, slightly longer than ligula. Thorax. Pronotum strongly convex, coarsely punctate across base, unwrinkled, moderately transverse and cordate, widest before middle; apex subtruncate; anterolateral angles poorly developed, rounded; anterior bead incomplete, obsolete medially; sides strongly rounded, not sinuate posteriorly; lateral grooves absent; two setiferous punctures on each side; posterolateral angles subrectangular; posterolateral carinae absent; laterobasal foveae well defined, coarsely punctate, very deep, moderately wide, rounded, not prolonged forward, simple; posterior bead absent; sub-basal transverse impression replaced by two foveolae; basal transverse impression well developed, coarsely punctate (with 10 to 12 punctures); base emarginate, much narrower than pronotal apex, about as wide as elytral base. Legs. Short. Elytra. Strongly convex, subovate, widest about middle. Basal margin incomplete, reaching about stria 4. Shoulders rounded. Sides strongly rounded. Scutellar striole present. Striae finely punctate; stria 1 complete, very deep; striae 2 – 5 incomplete, shallow; striae 6 – 7 obsolete; stria 3 with three setiferous punctures. Recurrent stria long, blunt, directed apically toward stria 5. Subapical seta present. Intervals subdepressed. Sutural apices obtusely rounded. Abdomen. Last visible sternum (sternum VII): male with two apical ambulatory setae; female with four apical ambulatory setae. Aedeagus. Lateral view (Fig. 109): rather strongly arcuate, not widened in apical half; base moderately convex dorsally; middle slightly concave dorsally, slightly convex ventrally, with dorsal membranous area very wide and long; apex subtriangular, moderately convex dorsally, slightly concave ventrally, with extreme tip wide and short. Dorsal view: very narrow, almost thin, asymmetrical (ostium of membranous area deflected to right); apex straight; basal orifice narrow, very close to membranous area. Parameres with two apical setae.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30219FFDB3FCAFB77FB39FA07.taxon	materials_examined	Material examined. 49 specimens (LUNZ, NZAC). Geographic distribution (Fig. 117). North Island: WN – Tararua Range (Dundas Ridge Hut; Otaki Forks, Judd Ridge). South Island: MB – Mount Fell. Mount Johnson. NN – Abel Tasman National Park, Moa Park. SD – Mount Stokes.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30219FFDB3FCAFB77FB39FA07.taxon	biology_ecology	Ecology. Montane, alpine. Epigean. Mostly forests (beech); tussock grasslands. Shaded or open ground. Nocturnal; hides during the day in leaf litter and tussock litter. Gregarious. Biology. Seasonality: October – December, March. Predacious (based on mouthpart morphology). Dispersal power. Subapterous (incapable of flight). Slow runner. Collecting techniques. Sifting leaf litter and tussock litter.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30219FFDB3FCAFB77FB39FA07.taxon	discussion	Remarks. This species is named after Dhahara Ranatunga (Auckland War Memorial Museum, Auckland) for her special help in our entomological studies. Maungazolus ranatungae is morphologically close to M. tararuaensis. In addition to diagnostic characters of the male genitalia, M. ranatungae has the following distinguishing features: head and pronotum dark brown; elytra yellowish, dark brown discally; abdomen mostly reddish piceous; mentum tooth slightly emarginate apically; pronotum moderately transverse and cordate, coarsely punctate across base; elytra with striae finely punctate and intervals subdepressed. M. ranatungae occurs in the south of the North Island (WN) and northern areas of the South Island (MB, NN, SD), while M. tararuaensis is only found in the south of the North Island (WN). M. ranatungae is the only zoline species known from both North and South Islands.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30218FFD43FCAFA17FA5FF9C7.taxon	description	Fig. 65, 110, 119 Holotype: male (NZAC) labeled: “ NEW ZEALAND WN Dennan Tararua FP [= Forest Park] 4 Jun 1984 (hand-written) / J. Nunn (typed) / [male symbol] / HOLOTYPE [male symbol] Maungazolus tararuaensis Larochelle & Larivière, 2017 (red label; typed). ” Paratype: one male (NZAC) from Field’s Hut [= Field Hut], Tararua FP [= Forest Park] (WN), bearing a blue paratype label. Description. Body length 3.1 – 3.3 mm. Head, pronotum, elytra, and abdomen mostly testaceous; disc of head, pronotum, elytra, and abdomen black; suture of elytra reddish to black; antennae, palpi, and legs reddish; antennal segments 1 + 2 and femora bright yellow, femora strongly contrasting with tibiae. Microsculpture absent. Iridescence absent. Very shiny, without metallic luster. Head. Labrum strongly transverse, slightly emarginate anteriorly. Antennae submoniliform, short: segment 1 (scape) short, stout, about 1.5 x longer than its maximum width. Frontal furrows wide, deep, convergent. Eyes slightly convex; two setiferous punctures on inner side of each eye. Tempora moderately inflated (about half as long as eyes). Mentum: medial tooth entire, acute apically, moderately shorter than lateral lobes. Paraglossae membranous, prominent, slightly longer than ligula. Thorax. Pronotum strongly convex, finely punctate across base, wrinkled discally, strongly transverse and cordate, widest before middle; apex subtruncate; anterolateral angles poorly developed, rounded; anterior bead incomplete, obsolete medially; sides strongly rounded, not sinuate posteriorly; lateral grooves absent; two setiferous punctures on each side; posterolateral angles subacute; posterolateral carinae obsolete; laterobasal foveae well defined, virtually impunctate, very deep, moderately wide, oblong, not prolonged forward, simple; posterior bead absent; sub-basal transverse impression replaced by two foveolae; basal transverse impression poorly developed, virtually impunctate (with about 10 obsolete punctures); base emarginate, much narrower than pronotal apex, about as wide as elytral base. Legs. Short. Elytra. Strongly convex, ovate, widest about middle. Basal margin incomplete, reaching about stria 4. Shoulders rounded. Sides strongly rounded. Scutellar striole obsolete. Striae coarsely punctate; stria 1 complete, very deep; striae 2 – 7 incomplete; striae 2 – 5 moderately deep; striae 6 – 7 obsolete; stria 3 with three setiferous punctures. Recurrent stria long, sharp, directed apically toward stria 5. Subapical seta present. Intervals moderately convex. Sutural apices rounded. Abdomen. Last visible sternum (sternum VII): male with two apical ambulatory setae; female unknown. Aedeagus. Lateral view (Fig. 110): strongly arcuate, very strongly widened in apical half; base moderately convex dorsally; middle strongly convex dorsally, slightly convex ventrally, with dorsal membranous area very wide and long; apex subtriangular, slightly concave dorsally, moderately concave ventrally, with extreme tip wide and short. Dorsal view: very narrow, almost thin, asymmetrical (ostium of membranous area deflected to right); apex straight; basal orifice narrow, very close to membranous area. Parameres with four or five apical setae.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30218FFD43FCAFA17FA5FF9C7.taxon	materials_examined	Material examined. 3 specimens (JNNZ, NZAC). Geographic distribution (Fig. 119). North Island: WN – Tararua Range (Dennan; Field Hut).	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30218FFD43FCAFA17FA5FF9C7.taxon	biology_ecology	Ecology. Montane. Forests (beech). Epigean. Shaded ground. Nocturnal; hides during the day in leaf litter. Biology. Seasonality: January, June. Predacious (based on mouthpart morphology). Dispersal power. Subapterous (incapable of flight). Slow runner. Collecting technique. Sifting leaf litter.	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
556787D30218FFD43FCAFA17FA5FF9C7.taxon	discussion	Remarks. This species is named after the Tararua Range (WN) where the beetle occurs, and the Latin suffix - ensis, denoting a place, locality, or country. Maungazolus tararuaensis is morphologically close to M. ranatungae. In addition to diagnostic characters of the male genitalia, M. tararuaensis has the following distinguishing features: head, pronotum, and elytra mostly testaceous; disc of head, pronotum, elytra, and abdomen black; mentum tooth acute apically; pronotum strongly transverse and cordate, finely punctate across base; elytra with striae coarsely punctate and intervals moderately convex. Maungazolus tararuaensis is only found in the south of the North Island (WN), while M. ranatungae occurs both in the south of the North Island (WN) and in northern areas of the South Island (MB, NN, SD).	en	Larochelle, Larivière, Marie-Claude, Larochelle, Larivière (2017): Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 2017 (594): 1-110, DOI: 10.5281/zenodo.5169575
