taxonID	type	description	language	source
CE4E0E3CCE3ECE1FC4E64A35EBAD6F38.taxon	materials_examined	Referred material. OREGON. Klamath County: USNM 1207966, Lost River at Stukel Bridge, USNM 1144894, USNM 1469075, USNM 1469082, USNM 1469090, Wood River, south spring source (42.7372 N, 121.9775 W), USNM 1469072, USNM 1469077, USNM 1469078, USNM 1469080, Tecumseh Spring (42.6424 N, 121.9432 W), USNM 1144346, USNM 1190088, Camporee Spring (42.4308 N, 122.0614 W), USNM 883517, USNM 1144348, USNM 1190089, USNM 1207965, USNM 1225874, Harriman Spring, outflow of main spring (42.4673 N, 122.1009 W), USNM 1469076, USNM 1469086, Big Springs at Bonanza (42.1982 N, 121.4004 W), USNM 1469074, USNM 1469088, Lost River, below Harpold Dam (42.1702 N, 121.4530 W).	en	Liu, Hsiu-Ping, Hershler, Robert (2019): A new species and range extensions for three other species of pebblesnails (Lithoglyphidae, Fluminicola) from the upper Klamath basin, California-Oregon. ZooKeys 812: 47-67, DOI: http://dx.doi.org/10.3897/zookeys.812.29205, URL: http://dx.doi.org/10.3897/zookeys.812.29205
CE4E0E3CCE3ECE1FC4E64A35EBAD6F38.taxon	diagnosis	Diagnosis. A large Fluminicola (shell height, 6.5 - 8.4 mm) with a subglobose to ovate-conic shell often having an eroded spire. Fluminicola klamathensis is readily distinguished by its penis, which does not gently taper along its length as in other congeners, but instead abruptly narrows distally and has a well demarcated, short filament. This new species is further differentiated from closely similar and frequently sympatric F. modoci in its generally darker colored shell periostracum, broad central cusps on the central and lateral radula teeth, very short outer wing of the lateral teeth, small number of cusps on the inner and outer marginal teeth, light pigment on the dorsal surface of the penis, large core of internal dark pigment in the distal section of the penis, and the fairly large seminal receptacle.	en	Liu, Hsiu-Ping, Hershler, Robert (2019): A new species and range extensions for three other species of pebblesnails (Lithoglyphidae, Fluminicola) from the upper Klamath basin, California-Oregon. ZooKeys 812: 47-67, DOI: http://dx.doi.org/10.3897/zookeys.812.29205, URL: http://dx.doi.org/10.3897/zookeys.812.29205
CE4E0E3CCE3ECE1FC4E64A35EBAD6F38.taxon	description	Description. Shell (Fig. 5 A-F) subglobose to ovate-conic, spire often eroded in large part, whorls (in specimens having a complete spire), 4.25 - 4.5. Teleoconch whorls low to medium convex, sometimes weakly shouldered. Aperture pyriform; inner lip complete, variably thickened, broadly adnate to parietal wall; columellar shelf narrow or extending over much of umbilical region. Outer lip thin, prosocline. Umbilicus absent or a narrow slit, umbilical region sometimes excavated. Shell white, periostracum brown, fairly thick, sometimes covered with black deposits. Shell measurements and whorl count data are summarized in Table 2. Operculum (Fig. 6 A-C) as for genus; muscle attachment margin little thickened on inner side; rim sometimes present on inner side near outer edge (Fig. 6 B). Radula (Fig. 6 D-H) as for genus; dorsal edge of central teeth concave, lateral cusps 2 - 5, hoe-shaped; basal cusp 1 - 5. Lateral teeth having 2 - 4 cusps on inner side and 3 - 5 cusps on outer side; outer wing slightly longer than length of cutting edge. Inner marginal teeth with 11 - 19 cusps, outer marginal teeth with 11 - 22 cusps. Radula data are from USNM 144346, USNM 1468970. Snout, cephalic tentacles grey or black, pigment light around eyespots; pallial roof, visceral coil usually light brown; foot variably pigmented dorsally, sole pale. Ctenidial filaments 33 - 36 (N = 5), broadly triangular. Glandular oviduct and associated structures shown in Figure 7 A, B. Coiled oviduct circular, anterior arm kinked, posterior arm sometimes having small accessory pouches containing sperm. Bursa copulatrix large, ovate or globular, partly overlapped by albumen gland. Bursal duct narrow, much shorter than bursa. Seminal receptacle medium-sized, pouch-like, partly overlapped by albumen gland. Albumen gland having small pallial component. Capsule gland slightly shorter than albumen gland. Genital aperture a small, terminal pore. Penis (Figs 7 C, D, 8 A) large, base rectangular, often having a distinct, lobe-like swelling along inner edge distally (Fig. 7 D); distal end of penis blunt, with short, narrow, filament. Distal section of penis having dense core of internal black pigment; dorsal surface pale or lightly pigmented. Penial duct near inner edge, narrow, nearly straight.	en	Liu, Hsiu-Ping, Hershler, Robert (2019): A new species and range extensions for three other species of pebblesnails (Lithoglyphidae, Fluminicola) from the upper Klamath basin, California-Oregon. ZooKeys 812: 47-67, DOI: http://dx.doi.org/10.3897/zookeys.812.29205, URL: http://dx.doi.org/10.3897/zookeys.812.29205
CE4E0E3CCE3ECE1FC4E64A35EBAD6F38.taxon	etymology	Etymology. The species name is an adjectival geographic epithet referring to the distribution of this pebblesnail in the (upper) Klamath River basin.	en	Liu, Hsiu-Ping, Hershler, Robert (2019): A new species and range extensions for three other species of pebblesnails (Lithoglyphidae, Fluminicola) from the upper Klamath basin, California-Oregon. ZooKeys 812: 47-67, DOI: http://dx.doi.org/10.3897/zookeys.812.29205, URL: http://dx.doi.org/10.3897/zookeys.812.29205
CE4E0E3CCE3ECE1FC4E64A35EBAD6F38.taxon	distribution	Distribution. Large, spring-influenced habitats in the UKL.	en	Liu, Hsiu-Ping, Hershler, Robert (2019): A new species and range extensions for three other species of pebblesnails (Lithoglyphidae, Fluminicola) from the upper Klamath basin, California-Oregon. ZooKeys 812: 47-67, DOI: http://dx.doi.org/10.3897/zookeys.812.29205, URL: http://dx.doi.org/10.3897/zookeys.812.29205
ACD2E4CBD6C514A0E13E04C72040EF0F.taxon	distribution	Distribution. North Fork Umpqua River drainage and Rogue River basin north of Little Butte Creek, Oregon (Hershler et al. 2017).	en	Liu, Hsiu-Ping, Hershler, Robert (2019): A new species and range extensions for three other species of pebblesnails (Lithoglyphidae, Fluminicola) from the upper Klamath basin, California-Oregon. ZooKeys 812: 47-67, DOI: http://dx.doi.org/10.3897/zookeys.812.29205, URL: http://dx.doi.org/10.3897/zookeys.812.29205
ACD2E4CBD6C514A0E13E04C72040EF0F.taxon	materials_examined	Referred material. OREGON. Klamath County. USNM 1144900, USNM 1469091, Harriman Spring, outflow of main spring (42.4673 N, 122.1009 W),	en	Liu, Hsiu-Ping, Hershler, Robert (2019): A new species and range extensions for three other species of pebblesnails (Lithoglyphidae, Fluminicola) from the upper Klamath basin, California-Oregon. ZooKeys 812: 47-67, DOI: http://dx.doi.org/10.3897/zookeys.812.29205, URL: http://dx.doi.org/10.3897/zookeys.812.29205
420644878A529D760853DD7C3DF2918C.taxon	distribution	Distribution. Several springs in the Goose Lake basin, California-Oregon (Hershler and Frest 1996; Hershler 1999; Hershler et al. 2007).	en	Liu, Hsiu-Ping, Hershler, Robert (2019): A new species and range extensions for three other species of pebblesnails (Lithoglyphidae, Fluminicola) from the upper Klamath basin, California-Oregon. ZooKeys 812: 47-67, DOI: http://dx.doi.org/10.3897/zookeys.812.29205, URL: http://dx.doi.org/10.3897/zookeys.812.29205
420644878A529D760853DD7C3DF2918C.taxon	materials_examined	Referred material. OREGON. Klamath County. USNM 1190095, USNM 1207964, Wood River, south spring source (42.7372 N, 121.9775 W), USNM 1144333, USNM 1190094, Tecumseh Spring (42.6424 N, 121.9432 W), USNM 874187, USNM 874935, USNM 1144336, USNM 1144337, USNM 1144942, USNM 1190086, Barkley Spring (42.3822 N, 121.8111 W), USNM 1190102, Klamath River, east of Boyle Power Station (42.0934 N, 122.0964 W), USNM 1190087, Ouxy Spring (42.3989 N, 121.8235 W), USNM 1144390, USNM 1144923, USNM 1190100, Brown Spring (42.4951 N, 121.2956 W), USNM 1190093, Spring, Klamath Fish Hatchery (42.6519 N, 121.9479 W), USNM 1190096, Reservation Spring (42.7023 N, 121.9629 W), USNM 1144407, USNM 1190098, Spring, Williamson River campground (42.6584 N, 121.8499 W), USNM 1190097, Spring Creek, headspring (42.6690 N, 121.8860 W), USNM 1144411, USNM 1190090, Crystal Spring (42.5736 N, 122.0823 W), USNM 1144966, USNM 1190121, Big Springs at Bonanza (42.1982 N, 121.4004 W), USNM 1144418, USNM 1185800, Casebeer Spring, ouflow at Gerber Dam Road (42.2056 N, 121.0592 W), USNM 1144454, USNM 1190123, Duncan Spring, north complex (42.0416 N, 121.0689 W), USNM 1190123, Lost River at Big Springs City Park, Bonanza (42.1976 N, 121.4002 W), USNM 114469, USNM 1190124, Gwinn Spring Creek at Gwinn Spring Creek Road (42.0063 N, 120.9545 W), USNM 1020714, USNM 1144673, USNM 1144925, USNM 1154376, Link River under US 97 / OR 140 bridge (42.2185 N, 121.7892 W). USNM 1190137, Link River, above Link River dam (42.2341 N, 121.8036 W), USNM 1144494, USNM 1144986, USNM 1207967, Lost River, Stevenson County Park (42.1831 N, 121.5994 W), USNM 1190139, Lost River, below Harpold Dam (42.1702 N, 121.4530 W), USNM 1144503, USNM 1190101, Sprague River at Beatty Gap (42.4476 N, 121.2377 W), USNM 1144515, USNM 1190099, Williamson River, Klamath County park (42.5705 N, 121.8791 W), USNM 1144520, USNM 1190140, Upper Klamath Lake, south of Modoc Point (42.4373 N, 121.8672 W). Lake County: USNM 1144565, USNM 1185798, Spring southeast of Slash Spring, Yocum Valley (42.0174 N, 120.7316 W).	en	Liu, Hsiu-Ping, Hershler, Robert (2019): A new species and range extensions for three other species of pebblesnails (Lithoglyphidae, Fluminicola) from the upper Klamath basin, California-Oregon. ZooKeys 812: 47-67, DOI: http://dx.doi.org/10.3897/zookeys.812.29205, URL: http://dx.doi.org/10.3897/zookeys.812.29205
388C55BDCFE402018D4237BD72A19EFC.taxon	distribution	Distribution. Sacramento River headwater region, upper reaches of the McCloud and Rogue River drainages, California-Oregon (Hershler et al. 2007; Hershler et al. 2017).	en	Liu, Hsiu-Ping, Hershler, Robert (2019): A new species and range extensions for three other species of pebblesnails (Lithoglyphidae, Fluminicola) from the upper Klamath basin, California-Oregon. ZooKeys 812: 47-67, DOI: http://dx.doi.org/10.3897/zookeys.812.29205, URL: http://dx.doi.org/10.3897/zookeys.812.29205
388C55BDCFE402018D4237BD72A19EFC.taxon	materials_examined	Referred material. CALIFORNIA. Siskiyou County: USNM 1207974, Spring on Close Butte (41.9884 N, 122.3229 W), USNM 1190109, Spring northwest of Copco Reservoir (41.9873 N, 122.3275 W), USNM 1145066, USNM 1190108, Fall Creek above Copco Road bridge (41.9834 N, 122.3623 W). OREGON. Jackson County: USNM 1144324, USNM 1144325, USNM 1144326, USNM 1144898, USNM 1145117, USNM 1190128, USNM 1243229, USNM 1254453, Fredenburg Spring (42.1669 N, 122.3268 W), USNM 1207971, Spring Creek north of Schoolhouse Meadow (42.0357 N, 122.3397 W), USNM 1144903, USNM 1190105, USNM 1207969, Spring brook below Schoolhouse Meadow (42.0288 N, 122.3374 W), USNM 1144342, USNM 1144943, USNM 1144365, USNM 1144366, USNM 1144484, USNM 1190114, Keene Creek, east of bridge on Mill Creek Road (42.1046 N, 122.4136 W), USNM 1144368, USNM 1144718, USNM 1144946, USNM 1190103, Rattlesnake Spring (42.0625 N, 122.3389 W), USNM 1190104, Shoat Spring, source (42.0466 N, 122.3360 W), USNM 1207968, Shoat Springs, outflow near source (42.0456 N, 122.3367 W), USNM 1207970, Spring channel above Schoolhouse Meadow, adjacent to cabin ruins (42.0327 N, 122.3379 W), USNM 1144536, USNM 1144537, USNM 1144941, USNM 1144942, USNM 1190118, Spring north of Hyatt Reservoir (42.2064 N, 122.4498 W), USNM 1144540, USNM 1144541, USNM 1144907, USNM 1144993, USNM 1190131, Nameless Spring, outflow (42.2183 N, 122.3087 W), USNM 1190129, Bluejay Spring (42.1810 N, 122.3368 W), USNM 1144587, USNM 1190115, Spring, Chinquapin Mountain (42.1409 N, 122.4268 W), USNM 1190116, Spring along Beaver Creek Road (42.1467 N, 122.4165 W), USNM 1190117, Spring, Craine Prairie (42.1754 N, 122.4086 W), USNM 1190111, Spring north of Soda Spring (42.1025 N, 122.3684 W), USNM 1190119, Spring, west side of Burnt Creek Road (42.1761 N, 122.4911 W), USNM 1144655, USNM 1145079, USNM 1190130, Jenny Creek Spring (42.2034 N, 122.3443 W). Klamath County: USNM 1469081, USNM 1469089, Wood River, south spring source (42.7372 N, 121.9775 W), USNM 1190138, Spring west of Klamath River (42.0257 N, 122.1351 W), USNM 1144392, USNM 1144951, USNM 1190127, Tiger Lily Spring (42.6156 N, 122.0935 W), USNM 1144393, USNM 1190092, Four Mile Spring (42.6331 N, 122.0778 W), USNM 1469079, Spring, Klamath Fish Hatchery (42.6519 N, 121.9479 W), USNM 1469083, Reservation Spring (42.7023 N, 121.9629 W), USNM 1469084, Spring Creek, headspring (42.6690 N, 121.8860 W), USNM 1144414, USNM 1144965, USNM 1190091, Short Creek, headspring (42.7000 N, 122.0776 W), USNM 1144416, USNM 1207975, Rainbow Springs (42.3239 N, 122.2040 W), USNM 1190134, Cold Creek, south of Lake of the Woods (42.3434 N, 122.2083 W), USNM 1144463, USNM 1190133, Spring along western edge of Buck Lake (42.2670 N, 122.1995 W), USNM 1190132, Johnson Creek (42.2401 N, 122.2399 W), USNM 1144468, USNM 1190136, Spring (northernmost), Denny Creek (42.3552 N, 122.0286 W), USNM 1144887, USNM 1144984, USNM 1190135, Spring (southernmost), Denny Creek (42.3324 N, 122.0221 W). Lake County: USNM 1190125, Spring, Holmes Meadow (42.1761 N, 120.8350 W), USNM 1144564, USNM 1185799, Blonde Spring (42.4149 N, 120.7467 W).	en	Liu, Hsiu-Ping, Hershler, Robert (2019): A new species and range extensions for three other species of pebblesnails (Lithoglyphidae, Fluminicola) from the upper Klamath basin, California-Oregon. ZooKeys 812: 47-67, DOI: http://dx.doi.org/10.3897/zookeys.812.29205, URL: http://dx.doi.org/10.3897/zookeys.812.29205
