taxonID	type	description	language	source
80F1B83329A25EDBB9FBD30080920C13.taxon	description	Figs 4, 5 A – F	en	Hoch, Hannelore, López, Heriberto, Naranjo, Manuel, Aguín-Pombo, Dora, Oromí, Pedro (2025): Endless forms most wonderful: Four new cavernicolous planthopper species (Hemiptera, Fulgoromorpha, Cixiidae and Meenoplidae) from the Canary Islands. Subterranean Biology 51: 61-101, DOI: 10.3897/subtbiol.51.144111
80F1B83329A25EDBB9FBD30080920C13.taxon	diagnosis	Diagnosis. Cixius palmirandus is similar to the other cavernicolous Cixius species from La Palma, C. palmeros Hoch & Asche, 1993 and C. pinarcoladus Hoch & Asche, 1993 in habitus (degree of troglomorphy), body size, general configuration of the male genital morphology, but differs in several characters: upper portion of frons smooth (vs. pustulate as in C. palmeros and C. pinarcoladus), mesonotum with lateral carinae attaining posterior margin (unlike in C. palmeros), tegmen with Y-vein (Pcu, A 1, Pcu + A 1) complete (vs Y-vein incomplete in C. palmeros and C. pinarcoladus), genital styles with expanded distal portion highly elongate (vs spoon-shaped in C. palmeros and C. pinarcoladus), and aedeagus shaft ventrally with an obtuse ridge which is apically rounded, concave and distinctly curved ventrally (vs apically with an obtuse tip, as in C. palmeros, or directed straight caudally, as in C. pinarcoladus).	en	Hoch, Hannelore, López, Heriberto, Naranjo, Manuel, Aguín-Pombo, Dora, Oromí, Pedro (2025): Endless forms most wonderful: Four new cavernicolous planthopper species (Hemiptera, Fulgoromorpha, Cixiidae and Meenoplidae) from the Canary Islands. Subterranean Biology 51: 61-101, DOI: 10.3897/subtbiol.51.144111
80F1B83329A25EDBB9FBD30080920C13.taxon	description	Description. Habitus. Strongly troglomorphic with compound eyes absent, tegmina, wings and bodily pigmentation strongly reduced. Body length. Male 3.9 mm (n = 1) Colouration. Head, thorax and abdomen stramineous / yellowish, lateral carinae of head and lateral carinae of pronotum in anterior portion slightly darker. Antennae and legs whitish, tegmina translucent with costal vein yellowish, other veins unpigmented. Head. Vertex short and wide, not separated from frons by a transverse carina, i. e., frons continuously rounded into vertex. Vertex laterally near posterior margin of head with two shallowly concave areas. Frons convex, in ventral view ca. twice as wide as medially long, smooth, without median carina, lateral carinae strongly ridged, directed laterally. Frontoclypeal suture highly vaulted. Post- und anteclypeus smooth, without median carina. Post- and anteclypeus together ca. 2.8 × longer than frons medially. Rostrum elongate, well surpassing hind coxae, 2 nd joint longer than 3 rd. Compound eyes and ocelli absent, the former position of the lateral ocelli faintly recognizable by a light roundish spot anteriorly of antennae. Antennae with scape very short, ring-like, pedicel globose, with sensory plaque organs feebly recognizable; antennae shielded anteriorly by lateral margins of frons. Thorax. Pronotum short, ca. 4 × wider than medially long, and 1.5 × wider than maximum width of head; indistinctly tricarinate: median carina obtuse, lateral carinae distinct in anterior portion, diverging laterally, gradually vanishing; posterior margin of pronotum shallowly incised. Mesonotum slightly vaulted, ca. 1.3 × wider than medially long, in midline ca. 3 × the length of pronotum; tricarinate, with carinae obtuse and faintly recognizable, lateral carinae attaining posterior margin. Tegulae vestigial. Tegmina strongly reduced, venation as in Fig. 4. Costal vein in anterior and distal part of tegmen conspicuously wide; „ Y-vein “ (Pcu, A 1, Pcu + A 1) preserved and recognizable. Tegmen ca. 1.6 × longer than maximally wide, attaining, respectively slightly surpassing posterior margin of third abdominal tergite. Longitudinal veins sparsely beset with bases of setae. Wings vestigial. Metatibiae laterally with 3 minute spines, distally with 6 teeth, grouped 5 + 1, lateral tooth longest. First and second metatarsal joints with 4 apical teeth, lateral ones longer than median ones. First metatarsal joint about as long as 2 nd and 3 rd joints together. Pretarsal claws slender, arolium small. Male genitalia. Genital segment bilaterally symmetrical, in caudal aspect ca. 1.6 × higher than maximally wide, and in lateral aspect ca. 4 × longer ventrally than dorsally, caudal margins laterodorsally slightly expanding into rounded lobes, their margins beset with a cluster of long setae; medioventral process wide at base, distally tapering, tip in ventral view slightly incised, dorsal surface of medioventral process concave. Anal segment tongue-shaped, lateral margins straight, without ventral lobes, parallel from base to level of anal style, distally slightly tapering; anal segment distally of anal style bent ventrally in a ca. 45 ° angle, caudal margin produced into two short rounded lobes. Genital styles narrow in basal third, distally expanding, expanded portion elongate, distally rounded, medially concave; genital styles with dorsal margin of narrow portion and external surface of expanded portion densely beset with setae. Genital styles in repose joining in midline over nearly their whole length ventrally, nearly completely covering the aedeagus. Aedeagus with basal part (shaft) tubular, slender, slightly compressed in basal two thirds, ventrally near base with two small rigid spines directed caudally; shaft ventrally with an obtuse longitudinal ridge which is apically rounded and in upper part slightly curved ventrally. Shaft subapically with a bulbous projection dorsally and right laterally, shaft apically with two sturdy movable spinose processes: left lateral one shallowly S-shaped, tip in repose directed basally, right lateral one slightly shorter than left lateral one, strongly curved and in repose directed right-lateroventrally, its tip pointing right laterally. Distal part of aedeagus (flagellum) in repose bent dorsally, narrow, surpassing midlength of shaft, but not attaining base of shaft; dorsally with a longitudinal ridge which is produced into a short, stout spine at apex, tip of flagellum directed right laterally. Female. Unknown.	en	Hoch, Hannelore, López, Heriberto, Naranjo, Manuel, Aguín-Pombo, Dora, Oromí, Pedro (2025): Endless forms most wonderful: Four new cavernicolous planthopper species (Hemiptera, Fulgoromorpha, Cixiidae and Meenoplidae) from the Canary Islands. Subterranean Biology 51: 61-101, DOI: 10.3897/subtbiol.51.144111
80F1B83329A25EDBB9FBD30080920C13.taxon	etymology	Etymology. The species epithet is an adjective in nominative singular, and a combination from the island of the type locality, La Palma, and the name of the cave, Cueva Honda de Miranda. The gender is masculine.	en	Hoch, Hannelore, López, Heriberto, Naranjo, Manuel, Aguín-Pombo, Dora, Oromí, Pedro (2025): Endless forms most wonderful: Four new cavernicolous planthopper species (Hemiptera, Fulgoromorpha, Cixiidae and Meenoplidae) from the Canary Islands. Subterranean Biology 51: 61-101, DOI: 10.3897/subtbiol.51.144111
80F1B83329A25EDBB9FBD30080920C13.taxon	distribution	Distribution. Known from the type locality in the east of La Palma, municipality of Breña Alta (Fig. 1). Endemic to La Palma.	en	Hoch, Hannelore, López, Heriberto, Naranjo, Manuel, Aguín-Pombo, Dora, Oromí, Pedro (2025): Endless forms most wonderful: Four new cavernicolous planthopper species (Hemiptera, Fulgoromorpha, Cixiidae and Meenoplidae) from the Canary Islands. Subterranean Biology 51: 61-101, DOI: 10.3897/subtbiol.51.144111
5630D00625135C708C4B33D81A2E60BE.taxon	description	Figs 6 A, B, 7 A – G	en	Hoch, Hannelore, López, Heriberto, Naranjo, Manuel, Aguín-Pombo, Dora, Oromí, Pedro (2025): Endless forms most wonderful: Four new cavernicolous planthopper species (Hemiptera, Fulgoromorpha, Cixiidae and Meenoplidae) from the Canary Islands. Subterranean Biology 51: 61-101, DOI: 10.3897/subtbiol.51.144111
5630D00625135C708C4B33D81A2E60BE.taxon	diagnosis	Diagnosis. Cixius theseus sp. nov. is similar to the other cavernicolous Cixius species from El Hierro, C. ariadne Hoch & Asche, 1993 and C. nycticolus Hoch & Asche, 1993 in habitus (degree of troglomorphy), body size, general configuration of the male and female morphology, but differs in several characters: Frons 2.3 × wider than medially high and not pustulate (vs. 1.5 × wider and pustulate in C. ariadne); tegmen with Y-vein preserved (vs. reduced in C. ariadne); in the male genitalia: caudal margin of anal segment medially incised (vs. rounded in C. ariadne), genital styles with distal expanded part dorsally rounded (vs. dorsally produced in C. ariadne), basal part of aedeagus (shaft) left laterally with a prominent longitudinal ridge (vs. without such a ridge in C. ariadne), and with bifurcate ventral projection slender (vs. wide in C. ariadne), right lateral subapical spinose process sturdy and in repose curved dorsally (vs. slender and in repose curved basally in C. ariadne) and, most prominently, distal part of aedeagus (flagellum) with a slender spinose process at ca. midlength (vs. without such a spinose process in C. ariadne); in the female genitalia: caudal margin of 7 th sternite medially expanding into an obtusely angulate process (as in C. nycticolus, vs. caudal margin straight in C. ariadne), and wax-secreting field on 9 th tergite medially separated by a narrow, longitudinal, membranous area (vs. wax-secreting field medially not separated but with a distinct median ridge in C. ariadne and C. nycticolus).	en	Hoch, Hannelore, López, Heriberto, Naranjo, Manuel, Aguín-Pombo, Dora, Oromí, Pedro (2025): Endless forms most wonderful: Four new cavernicolous planthopper species (Hemiptera, Fulgoromorpha, Cixiidae and Meenoplidae) from the Canary Islands. Subterranean Biology 51: 61-101, DOI: 10.3897/subtbiol.51.144111
5630D00625135C708C4B33D81A2E60BE.taxon	description	Description. Habitus. Strongly troglomorphic with compound eyes absent, tegmina, wings and bodily pigmentation strongly reduced. Body length. Male 2.7 mm (n = 1). Female 3.3 mm (n = 1). Colouration. Male. Head and thorax incl. legs light yellowish, lateral carinae of head and posterior margin of vertex brownish; antennae whitish; tegmina translucent, whitish, venation white-yellowish, veins beset with brownish setae; legs whitish, distal spines of hind tibiae and of metatarsal joints brownish; abdomen whitish, genital capsule slightly darker, yellowish brown. Female. Head with vertex light yellowish, frons yellowish, medially with a brownish longitudinal stripe; clypeus light brown; antennae yellowish with distinct reddish brown star-shaped sensory plaque organs; pronotum medially, i. e., between lateral carinae, yellowish, lateral portions slightly darker, yellowish brown; mesonotum light yellowish; tegmina translucent, venation whitish, beset with brownish setae; legs yellowish white; abdomen light yellowish, genital segment incl. ovipositor yellowish brown. Head. Vertex short, ca. 3.5 × wider at base than medially long, very faintly separated from frons by an obsolete transverse carina. Frons convex, ca. 2.3 × wider than medially high, lateral carinae distinctly ridged and directed (antero-) laterally; frons smooth, without median carina, not pustulate. Frontoclypeal suture highly vaulted / arched. Post- and anteclypeus smooth, without median carina, together ca. 3.4 × longer than frons. Rostrum elongate, 2 nd joint longest; rostrum relatively shorter in the male: surpassing caudal margin of hind coxae only slightly, in the female with ca. half the length of 3 rd joint. Compound eyes and ocelli absent. Antennae with scape very short, ring-like, pedicel subcylindrical, ca. 1.4 × longer than wide, in the female with distinctly recognizable star-shaped sensory plaque organs. Thorax. Pronotum faintly tricarinate, lateral carinae diverging laterally, gradually vanishing; pronotum ca. 1.8 × wider than maximum width of head, and 4.2 × wider than medially long, posterior margin concave, obtusely angulate. Mesonotum tricarinate, carinae only faintly recognizable, lateral carinae reaching posterior margin, median carina feeble, obtuse, vanishing caudally; mesonotum in the male 1.5 ×, in the female 1.4 × wider than medially long, and in midline 2.3 × longer than length of pronotum. Tegulae small. Tegmina strongly reduced, their caudal margin attaining ca. midlength of 3 rd abdominal tergite; tegmen longer than maximally wide: ca. 1.5 × in the male, and 1.65 × in the female; venation rudimentary, costal vein strong, basal cell closed, „ Y-vein “ (Pcu, A 1, Pcu + A 1) preserved and recognizable, A 1 and Pcu + A 1 very close to posterior margin of tegmen; tegmina with numerous conspicuous setae along veins. Wings vestigial, very small. Metatibiae laterally with 3 tiny spines, distally with 6 (in the male), and 6 / 7 (in the female) apical teeth, of which the lateral one is largest. First metatarsal joint in both sexes with 4, and second metatarsal joint with 4 (in the male) and ¾ (in the female) distal spines. First metatarsal joint about as long as 2 nd and 3 rd metatarsal joints together. Pretarsal claws slender, arolium small. Male genitalia. Genital segment in caudal aspect slightly higher than wide, medioventral process simple, in ventral aspect obtusely angulate. Anal segment in dorsal aspect rectangular, ca. 2 × longer than wide, with distal portion slightly bent ventrally, lateral margins parallel, distal margin medially incised. Genital styles slender at base, distally expanding dorsally, expanded part medially concave. Aedeagus with basal part (shaft) slender, more or less tubular, left laterally with a prominent, longitudinal, rounded ridge and ventrally with a bifurcate projection directed basally. Shaft subapically on its right side with a sturdy movable spinose process which in repose is curved basally, its tip pointing dorsally. Distal part of aedeagus (flagellum) tubular, in repose bent dorsally and to right side, surpassing midlength of shaft, with a slender, spinose process, in repose directed basally, left laterally at ca. midlength of flagellum; visible part of ejaculatory duct rugose; phallotreme wide, located apically. Female genitalia. 7 th sternite with anterior margin convex, highly vaulted cephally, rounded, caudal / posterior margin medially expanding caudally, expanded portion obtusely angulate. Ovipositor ensiform, slightly curved dorsally, caudally surpassing anal tube with less than 1 / 3 of its total length; anal segment tubular, short, in lateral view ca. 2 × higher than long, lateral margins more or less parallel; 9 th tergite caudally truncate, wax-secreting field distinctly limited, slightly concave, medially separated by a narrow, longitudinal membranous portion.	en	Hoch, Hannelore, López, Heriberto, Naranjo, Manuel, Aguín-Pombo, Dora, Oromí, Pedro (2025): Endless forms most wonderful: Four new cavernicolous planthopper species (Hemiptera, Fulgoromorpha, Cixiidae and Meenoplidae) from the Canary Islands. Subterranean Biology 51: 61-101, DOI: 10.3897/subtbiol.51.144111
5630D00625135C708C4B33D81A2E60BE.taxon	etymology	Etymology. The species epithet is a noun in nominative singular and refers to Theseus, one of the heroes in Greek mythology, friend of Ariadne. The gender is masculine.	en	Hoch, Hannelore, López, Heriberto, Naranjo, Manuel, Aguín-Pombo, Dora, Oromí, Pedro (2025): Endless forms most wonderful: Four new cavernicolous planthopper species (Hemiptera, Fulgoromorpha, Cixiidae and Meenoplidae) from the Canary Islands. Subterranean Biology 51: 61-101, DOI: 10.3897/subtbiol.51.144111
5630D00625135C708C4B33D81A2E60BE.taxon	distribution	Distribution. Adults known only from the type locality (Camino San Salvador), in the laurel forest in the huge landslide of El Golfo, at the northwest of El Hierro (Fig. 1). Endemic to El Hierro.	en	Hoch, Hannelore, López, Heriberto, Naranjo, Manuel, Aguín-Pombo, Dora, Oromí, Pedro (2025): Endless forms most wonderful: Four new cavernicolous planthopper species (Hemiptera, Fulgoromorpha, Cixiidae and Meenoplidae) from the Canary Islands. Subterranean Biology 51: 61-101, DOI: 10.3897/subtbiol.51.144111
1200FC52C16E578E9EFCE7EE4C63642B.taxon	description	Figs 11 A, B, 12 A, B, 13 A – G, 14 A, B	en	Hoch, Hannelore, López, Heriberto, Naranjo, Manuel, Aguín-Pombo, Dora, Oromí, Pedro (2025): Endless forms most wonderful: Four new cavernicolous planthopper species (Hemiptera, Fulgoromorpha, Cixiidae and Meenoplidae) from the Canary Islands. Subterranean Biology 51: 61-101, DOI: 10.3897/subtbiol.51.144111
1200FC52C16E578E9EFCE7EE4C63642B.taxon	diagnosis	Diagnosis. Meenoplus skotinophilus is similar in general appearance and degree of troglomorphy to Meenoplus claustrophilus from La Palma, but differs from this species by the distally stronger reduced tegmina and lighter overall pigmentation. From the other two cavernicolous Meenoplus species on El Hierro (M. cancavus und M. charon), it differs distinctly by its degree of troglomorphy (compound eyes present, tegmina and wings well developed, tegmina surpassing tip of abdomen). While the general configuration of the male and female genitalia is similar in all four species (M. claustrophilus, M. skotinophilus, M. cancavus und M. charon), Meenoplus skotinophilus differs from these by the following characters of the male genitalia: ventrocaudal lobes of anal segment with median tips subacute and well separated (vs rounded and nearly apposed in the other species), aedeagus with apical margins of phallotreme angulate (vs rounded in the other species), and of the female genitalia: ventral valvulae distally with a beak-shaped, sturdy and acute tip pointing mediad (vs bearing a minute tip), and with proximal portion broadly lobate and finely serrate (vs rounded and smooth in the other species).	en	Hoch, Hannelore, López, Heriberto, Naranjo, Manuel, Aguín-Pombo, Dora, Oromí, Pedro (2025): Endless forms most wonderful: Four new cavernicolous planthopper species (Hemiptera, Fulgoromorpha, Cixiidae and Meenoplidae) from the Canary Islands. Subterranean Biology 51: 61-101, DOI: 10.3897/subtbiol.51.144111
1200FC52C16E578E9EFCE7EE4C63642B.taxon	description	Description. Habitus. Troglomorphies weakly defined except for compound eyes and pigmentation, tegmina and wings well developed, in repose surpassing the tip of the abdomen. In general appearance, intermediate between epigean Meenoplidae and strongly troglomorphic species, such as e. g., Meenoplus cancavus Remane & Hoch, 1988 and M. charon Hoch & Asche, 1993. Body length. Male 2.8 – 2.9 mm (n = 3). Female 3.2 – 3.5 mm (n = 6). Colouration. Head, pro- and mesonotum as well as male and female genitalia yellowish-brown, otherwise thorax und pregenital abdomen white. Compound eyes red. Tegmina translucent, pale stramineous in males, light brown in females; venation as well as areas along veins and between sensory pits yellowish-brown. Wings hyaline, venation yellowish-brown. Legs stramineous, apical spines of tibia and tarsal joints I – II, light brown. Head. Vertex very short, ca. 12 times wider than medially long, distinctly separated from frons by a ridged transverse carina. Frons strongly convex, anterior portion bulbous, about as wide as medially high, at and below level of antennae with a short row of small and hardly visible sensory pits irregular in number (4 – 6). Lateral carinae of frons foliately ridged and directed anterolaterad, anteriorly with a distinct row of large sensory pits; lateral lamelliform carinae continuing onto postclypeus. Frons smooth, without median carina, postclypeus shallowly, anteclypeus steeply vaulted. Compound eyes small, lateral ocelli vestigial, median frontal ocellus strongly reduced, ist former position marked by a light circular spot at the lower portion of the frontal bulbous area. Scape short, ring-like, pedicel cylindrical, ca. 1.7 times longer than wide. Thorax. Pronotum medially about 3.5 times the length of vertex, posterior margin obtusely angulate; pronotum weakly tricarinate, median carina very feeble. Tegulae, tegmina and wings well developed; tegmina distally surpassing tip of abdomen with ca. 1 / 5 their total length. Tegmen with rows of sensory pits along the distal part of the costal vein, along ScP + R (+ Ma), RP (+ MA) and along PCu, A 1, and their common stem PCu + A 1 („ Y-vein “). Metatibiae laterally unarmed, with 8 apical teeth. First metatarsal joint with 7 apical teeth, second metatarsal joint with 6 apical teeth. Male genitalia. Genital segment in lateral aspect ventrally ca. 3 times as long as dorsally. Anal segment distally produced into two ventrocaudal lobes which converge medially, their median tips subacute and well separated from each other. Genital styles slender, narrow throughout, apically rounded, gently curved dorsad. Aedeagus tubular, stout, with phallotrema apically and dorsocaudally exposed, apical margins dorsally bluntly angulate. Female genitalia. Strongly reduced, ventral valvifers produced into a rounded lobe; ventral valvulae with distal portion „ bird-head-shaped “, i. e., caudally rounded and medially with an acute tip pointing mediad, and with proximal portion broadly lobate, with median margin straight and finely serrate, apposed. Molecular identification. Mitochondrial COI barcode sequences of 635 pb were obtained for three individuals of Meenoplus skotinophilus (specimen codes BC 1267, BC 1268 and BC 1269). These individuals have identical barcode sequences, so no interpopulation genetic divergence has been detected. Either in BOLD or GenBank, no matches with similarity values greater than 85 % were detected, so the genetic information that we supply is actually new for the genetic lineage of the group of species that may belong Meenoplus skotinophilus. The sequences were deposited in GenBank (accession numbers PQ 530856, PQ 530856 y PQ 530856), with the following base composition: AATGAGCCAGATTAATAGGTATAACAAGAAGAATAATTATTCGAATTGAATTAATACAACCTGGTTCAATAATTAAAAATGATCAAATTTATAACTCAATTGTTACATCACATGCATTCATTATAATTTTTTTTTCAGTTATACCCATCCTAATCGGTGGATTTGGAAATTGACTTGTACCTCTAATGATTGGAGCACCTGATATAGCATTCCCACGAATAAACAATATAAGCTTCTGAATATTACCTCCATCACTAATACTATTAATTTTCAGTTCATTTTCAGGTTCAGGTACAGGTACAGGATGAACAATTTATCCACCATTATCAAGAATTCCTGCACATTCTGGCCCATCTACTGACTTATCTATCTTTTCCCTTCATATAGCAGGTGTAAGATCAATTCTAGGAGCAATTAATTTCATTTCAACTATTATTAATATACGACCTAAAATAATAACAATAGAAAAAATACCCCTATTTTGCTGATCAATTTTCATTACAGCAATTTTACTTCTTCTATCATTACCTATTCTTGCAGGAGCAATTACTATACTATTAACTGATCGAAACTTTAATACATCATTTTTTGATCCAACAGGAGGAGGAGACCCTATTTTATATCAACATTTATTT	en	Hoch, Hannelore, López, Heriberto, Naranjo, Manuel, Aguín-Pombo, Dora, Oromí, Pedro (2025): Endless forms most wonderful: Four new cavernicolous planthopper species (Hemiptera, Fulgoromorpha, Cixiidae and Meenoplidae) from the Canary Islands. Subterranean Biology 51: 61-101, DOI: 10.3897/subtbiol.51.144111
1200FC52C16E578E9EFCE7EE4C63642B.taxon	etymology	Etymology. The species epithet is an adjective in nominative singular and a combination of the Greek words „ skótos “ (= darkness) and „ phílos “ (= friend). The gender is masculine.	en	Hoch, Hannelore, López, Heriberto, Naranjo, Manuel, Aguín-Pombo, Dora, Oromí, Pedro (2025): Endless forms most wonderful: Four new cavernicolous planthopper species (Hemiptera, Fulgoromorpha, Cixiidae and Meenoplidae) from the Canary Islands. Subterranean Biology 51: 61-101, DOI: 10.3897/subtbiol.51.144111
1200FC52C16E578E9EFCE7EE4C63642B.taxon	distribution	Distribution. The species is kown only from the type locality, Cueva de Guinea, municipality of Frontera (Fig. 1). Endemic to El Hierro.	en	Hoch, Hannelore, López, Heriberto, Naranjo, Manuel, Aguín-Pombo, Dora, Oromí, Pedro (2025): Endless forms most wonderful: Four new cavernicolous planthopper species (Hemiptera, Fulgoromorpha, Cixiidae and Meenoplidae) from the Canary Islands. Subterranean Biology 51: 61-101, DOI: 10.3897/subtbiol.51.144111
C929567264845AB084562A52FF6CCEFD.taxon	description	Figs 8 A, B, 9, 10 A – G	en	Hoch, Hannelore, López, Heriberto, Naranjo, Manuel, Aguín-Pombo, Dora, Oromí, Pedro (2025): Endless forms most wonderful: Four new cavernicolous planthopper species (Hemiptera, Fulgoromorpha, Cixiidae and Meenoplidae) from the Canary Islands. Subterranean Biology 51: 61-101, DOI: 10.3897/subtbiol.51.144111
C929567264845AB084562A52FF6CCEFD.taxon	diagnosis	Diagnosis. In general appearance and in the overall configuration of male and female genital structures Tachycixius gomerobscurus sp. nov. ressembles T. crypticus and T. retrusus from Tenerife, but differs in the following characters: shape of vertex: vertex short, anterior margin very shallowly rounded (vs strongly convex towards frons in T. crypticus and T. retrusus); colouration of tegmina: less vividly coloured than in T. crypticus and T. retrusus; reduction of hind wings much stronger than in T. crypticus and T. retrusus; male genitalia: caudal margin of anal segment medially strongly concave (vs shallowly concave in T. crypticus and straight in T. retrusus); shaft of aedeagus with 3 subapical movable spines (vs. 2 such spines in T. crypticus and T. retrusus); female genitalia: 9 th tergite medioventrally deeply incised, membranous excavation acutely triangular (vs 9 th tergite medioventrally only shallowly incised, membranous excavation dorsally shallowly rounded in T. crypticus and T. retrusus); 9 th tergite with wax-secreting field medially with a short, but distinct median ridge (vs without such a ridge in T. crypticus and T. retrusus).	en	Hoch, Hannelore, López, Heriberto, Naranjo, Manuel, Aguín-Pombo, Dora, Oromí, Pedro (2025): Endless forms most wonderful: Four new cavernicolous planthopper species (Hemiptera, Fulgoromorpha, Cixiidae and Meenoplidae) from the Canary Islands. Subterranean Biology 51: 61-101, DOI: 10.3897/subtbiol.51.144111
C929567264845AB084562A52FF6CCEFD.taxon	description	Description. Habitus. In general appearance resembling Tachycixius crypticus Hoch & Asche, 1993 and T. retrusus Hoch & Asche, 1993 from Tenerife, although less vividly coloured; weakly troglomorphic (i. e. hypogeomorphic, see Deharveng and Bedos 2018): compound eyes present, but small, tegmina covering most of the abdomen but not attaining tip of anal tube in the male, respectively tip of ovipositor in the female; hind wings strongly reduced, vestigial. Body length. Male 3.8 – 4.05 mm (n = 4). Female 4.3 – 5.2 mm (n = 6). Colouration. Vertex, frons and head laterally light yellowish, with lateral carinae of vertex and frons and posterior margin of vertex slightly darker; antennae (pedicel) whitish; compound eyes reddish-dark brown; pro- and mesonotum light yellowish, otherwise thorax incl. Legs whitish, tips of lateral and distal spines of tibia, as well as distal spines of first and second metatarsal joints dark brown. Tegmina translucent, light yellowish, venation whitish, bases of setae along veins and margin of tegmen slightly darker, brownish; tegmen with anterior portion of triangle formed by cubitus posterior (CuP) and posterior margin of tegmen profusely brownish, and three irregularly limited, faint fuscous transverse bands: one at level of tegmen midlength, one at level of pterostigma, and one more or less parallel to distal margin (see remarks). Abdomen incl. genitalia light yellowish, or yellowish-brown, respectively. Head. Vertex short, about 3.5 × wider at base than medially long, anteriorly rounded, with a faint median carina, areolets small, slightly concave, medially separated by an obtuse median carina; vertex indistinctly separated from frons by an obsolete transverse carina. Frons 1.2 × wider than medially high, with a distinct median carina, lateral carinae foliately produced laterally. Post- and anteclypeus with an obtuse median carina; together ca. 1.5 × longer than frons. Frontoclypeal suture strongly arched. Compound eyes present, compared to epigean Tachycixius species reduced in relative size, pigmented, lateral ocelli distinct, median frontal ocellus rudimentary. Rostrum elongate, well surpassing hind coxae, in males attaining anterior margin of 9 th abdominal segment, in females attaining level of caudal margin of 9 th tergite. Antennae with scape short, ring-like, and pedicel cylindrical, ca. 1.4 × longer than wide, with sensory plaque organs arranged in several rows. Thorax. Pronotum tricarinate, lateral carinae ridged and diverging laterally near posterior margin of pronotum; pronotum short, medially ca. 1.8 × longer than vertex, and 1.7 × wider than maximum width of head (incl. compound eyes), posterior margin concave, medially forming an obtuse angle. Mesonotum tricarinate, carinae faint, lateral carinae attaining posterior margin of mesonotum; mesonotum 1.2 × wider than medially long, and medially 3.2 × longer than pronotum. Tegulae small. Tegmina distally reduced, in both sexes attaining caudal margin of anal segment, ca. 2.5 × longer than maximally wide; venation well developed, variable among specimens (in all specimens studied except for 1 female, the CuP vein does not connect to posterior margin of tegmen as is the case for most Cixiidae, but merges with PCu + A 1 (= common stem of Y-vein, see Fig. 6, arrow), see remarks; basal cell closed, pterostigma faintly recognizable; veins beset with or accompanied by numerous conspicuous bases of setae, on distal margin also between veins. Hind wings very small, vestigial, not surpassing posterior margin of metanotum. Metatibiae laterally in the majority of specimens studied with 3 small spines (variation: configurations 4 / 3 and 3 / 2 were observed in one female each), distally with 6 sturdy spines (in one female with 7 spines on one leg) (arranged in a row, lateral one strongest); metabasitarsus distally with 7 – 8 (bilaterally and individually variable), 2 nd metatarsal joint distally with 7 – 8 spines (bilaterally and individually variable), each of the median 4 bearing one macroseta. Metabasitarsus slightly longer than 2 nd and 3 rd metatarsal joints together. Pretarsal claws short, stout, arolium small. Male genitalia. Genital segment in caudal view ca. 1.3 × higher than wide, and in lateral view ventrally ca. 4.6 × longer than dorsally, caudal margins laterodorsally produced into 2 rounded lobes directed laterocaudally; medioventral process slender, triangular, dorsal surface with a faint median ridge. Anal segment elongate, narrow, in distal third bent ventrocaudally, in dorsal view ca. 2 × longer than wide at base, lateral margins in dorsal view more or less parallel, caudally of anal style converging, distal margin broadly rounded, ventral portion distally of anal style in caudal aspect strongly vaulted, with caudal margin medially concave; anal style elongate, slender. Genital styles narrow at base, distal third expanding dorsally, expanded portion caudally rounded, dorsally with an obtuse angle, medially concave. Aedeagus. Basal part of aedeagus (shaft) in proximal half wide, with three more or less compressed velum-like projections: one left laterally, extending from base to ca. midlength of shaft, one ventrally which is wide at base, narrowing at ca. midlength of shaft and extending from base almost to apex, and one right laterally which is broadly rounded and directed right laterocaudally. Shaft in distal half on right side with a compressed lobe extending right laterally, and subapically with 3 sturdy movable spinose processes: one left laterally, in repose curved dorsally, one ventrally, in repose directed basally, its tip pointing left laterally, and one right laterally, which is double-S-shaped, in repose curved basally and to right side. Distal portion of aedeagus (flagellum) well surpassing midlength of shaft, medially almost rectangularly bent and directed right laterally; without any spinose processes; distal portion of flagellum on ventral side expanding into a lobate, rounded protrusion, visible part of ejaculatory duct rugose, phallotreme wide, exposed dorsally. Female genitalia. Seventh sternite subtriangular, anterior margin broadly rounded, caudal margin medially straight; ovipositor ensiform, slightly curved dorsally, caudally slightly surpassing anal segment; anal segment tubular, short, stout, dorsoventrally only slightly depressed; ninth tergite caudally truncate, wax-secreting field indistinctly limited, shallowly concave, with a short, but distinct median ridge; 9 th tergite medioventrally deeply incised, membranous excavation acutely triangular, attaining dorsal third of 9 th tergite.	en	Hoch, Hannelore, López, Heriberto, Naranjo, Manuel, Aguín-Pombo, Dora, Oromí, Pedro (2025): Endless forms most wonderful: Four new cavernicolous planthopper species (Hemiptera, Fulgoromorpha, Cixiidae and Meenoplidae) from the Canary Islands. Subterranean Biology 51: 61-101, DOI: 10.3897/subtbiol.51.144111
C929567264845AB084562A52FF6CCEFD.taxon	etymology	Etymology. The species epithet is an adjective in nominative singular and a combination of Gomera and oscuro, the Spanish word for dark, probably used to create the toponym of this shadowy location inside the laurel forest. The gender is masculine.	en	Hoch, Hannelore, López, Heriberto, Naranjo, Manuel, Aguín-Pombo, Dora, Oromí, Pedro (2025): Endless forms most wonderful: Four new cavernicolous planthopper species (Hemiptera, Fulgoromorpha, Cixiidae and Meenoplidae) from the Canary Islands. Subterranean Biology 51: 61-101, DOI: 10.3897/subtbiol.51.144111
C929567264845AB084562A52FF6CCEFD.taxon	distribution	Distribution. Known only from the type locality, Reventón Oscuro, municipality of Hermigua, in Garajonay National Park (Fig. 1). Endemic to La Gomera.	en	Hoch, Hannelore, López, Heriberto, Naranjo, Manuel, Aguín-Pombo, Dora, Oromí, Pedro (2025): Endless forms most wonderful: Four new cavernicolous planthopper species (Hemiptera, Fulgoromorpha, Cixiidae and Meenoplidae) from the Canary Islands. Subterranean Biology 51: 61-101, DOI: 10.3897/subtbiol.51.144111
