identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
5836481EBC3DFFCBFF52F2EBF0FCFBD4.text	5836481EBC3DFFCBFF52F2EBF0FCFBD4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hygronemobius duckensis Martins & Pereira	<div><p>Hygronemobius duckensis Martins &amp; Pereira, sp. nov.</p><p>(Figs. 1 A–B, 2, 3A–C, 4A)</p><p>Etymology. The specific name refers to type locality, the Reserva Florestal Adolpho Ducke. This place is a conservation area located in the municipality of Manaus.</p><p>Diagnosis. (i) Pronotum and legs light with darker spots (Figs. 1 A–B; 2A–C); (ii) female tergites with many apparent blackish dots and tergite 4 darker than the others (Fig. 1 B); (iii) male right tegmen with a yellow spot at the apex (Figs. 1 A, 2D); (iv) pseudepiphallic apical lobe reduced, not visible in lateral view (Fig. 3 C); (v) apex of pseudepiphallic sclerite well developed, totally covering the ectophallic parameres in dorsal view (Fig. 3 B); (vi) apex of pseudepiphallus curved ventrally, sub-triangular shaped (Fig. 3 C).</p><p>Description. Holotype, male, measurements (mm): body length, 6.50; pronotum length, 1.30; pronotum width, 1.80; head width, 1.90; length of femur III, 3.90; length of tibia III, 3.10; right tegmen length, 3.40; right tegmen width (dorsal field), 2.40. Head, light yellow with some darker portions: a large dark brown stripe behind the eyes; some dark brown markings on median-apical vertex, fastigium, frons and clypeus; occiput and vertex covered by pubescence and with some long black bristles; black eyes; three ocelli present, central ocellus surrounded by dark brown spot, lateral ocelli partially surrounded by one dark brown spot each; scape light yellow; pedicel light brown; flagellum light brown becoming darker to the apex; anteclypeus whitish; dark brown spot under the eye; gena whitish; mandible dark brown on proximal portion to light yellow on distal; labrum light yellow to light brown; Maxilla whitish, lacinia light brown with whitish apex; maxillary palp: first two palpomere dark brown; third palpomere, proximal portion dark brown, and distal portion whitish; fourth palpomere whitish; fifth palpomere dark brown, whitish on proximal portion and white on apex; labial palp: first palpomere dark brown; second palpomere with dark brown proximal half and whitish distal half; third palpomere dark brown, proximal portion whitish and white on apex. Thorax, pronotum light yellow adorned with diffuse brown spots (Fig. 1 A), presence of fine pubescence and long black bristles covering all its extension; lateral lobe with a large light yellow spot on inferior margin and five light yellow spots aligned in a semi-parabola shaped (Fig. 2 D). Prosternum light brown. Mesosternum dark brown with a yellowish triangle shaped spot in posterior margin (Fig. 1 B). Metasternum yellowish and light brown. Right tegmen, dorsal field dark brown with light yellow spots (Fig. 1 D) situated: 1) proximal portion of basal and harp areas; 2) distal portion of basal area and along of stridulatory file; 3) close proximal portion of cordal area; 4) distal portion of cordal area; 5) covering apical area; 6) large spot between longitudinal vein in harp and Cu1. Lateral field with five veins, the color pattern varying from whitish to dark brown: supero-proximal portion dark brown, supero-distal light brown and inferior portion whitish; three rectangular light yellow spots between M vein and superior vein of lateral field. Hindwings absent. Legs I and II with similar pattern: coxa, trochanter and femur whitish with dark brown diffuse markings; tibia light brown with an oval tympanum on outer face; tarsus light brown, first tarsomere with a central elongated whitish spot. Leg III, femur whitish with four dark brown stripes on external face: three diagonally aligned stripes on superior portion and another one longitudinally arranged on inferior portion; tibia light brown, external face with five whitish marks, three of them surrounded the dorsal spurs; apical and dorsal spurs dark brown with whitish base and apex; tarsus whitish: first tarsomere with two brownish rings, one closed to proximal portion and another on distal portion; third tarsomere with a light brown ring on proximal portion and another on distal one. Abdomen with tergites dark brown, tergites I–IV light brownish on central portion corresponding to area covered by tegmina, tergites VII–IX with diffuse light brown marks (Fig. 2 A). Supranal plate whitish to light brown, apex rounded (Fig. 2 E). Cerci light brown. Sternites I–II dark brown with posterior margin whitish, sternites III–VIII dark brown with two whitish triangular spots near median portion. Subgenital plate dark brown with two light brown spots close to distal margin, sub-trapezoidal shaped (Fig. 2 F). Male genitalia (Fig. 3 A–C): triangular; pseudepiphallic sclerite well sclerotized: proximal two-thirds inflated, distal third narrow, curved ventrally and invaginated in central portion; pseudepiphallic parameres as long as the apex of pseudepiphallic sclerite, visible in ventral view; apex of pseudepiphallic sclerite well developed, totally covering the ectophallic parameres in dorsal view; apex of pseudepiphallus curved ventrally, sub-triangular; ectophallic fold small, visible only in ventral view; ectophallic apodemes surpassing, but not crossing the rami; endophallus shell shaped divided in three sclerites, the lateral sclerites being larger than the central one; end of lateral sclerites of endophallus laterally projected.</p><p>Observations in paratypes. Stridulatory file of right tegmen with 85–93 teeth (n=4). Female, body shape and color pattern similar to Holotype, except: tegmina reduced to dark brown lateral scales; abdomen, tergites I–VII with several rounded dark brown spots; tergite IV darker than the other ones; sternites varying from same pattern described for holotype to sternites largely whitish with central and lateral portion dark brown; supranal plate rectangular shaped with distal margin rounded, color dark brown to whitish (Fig. 2 G); subgenital plate whitish, rectangular shaped with rounded distal corners and invaginated on apex (Fig. 2 H); ovipositor (Fig. 2 I), valves light brown on proximal portion to dark brown on median and distal portions, apex of the dorsal valve slightly denticulate on their margins.</p><p>Calling song (26.5–27.4°C, Tab. 0 1, Fig. 4 A): 15 to 30 pulses per chirp (n=90); gradual increase in amplitude and duration from the first to the third/sixth pulse of each chirp; duration of the first pulse of 7.79 ± 1.21 ms (5–10, n=24), duration of the third/sixth pulse of 10.5 ± 0.59 ms (9–11, n=24), and the other pulses with duration of 11.92 ± 0.65 ms (11–13, n=48); chirp duration ranged from 320 to 660 ms (n=90) according to number of pulses per chirp (see Tab. 01); inter-chirp interval ranged from 1300 to 1700 ms (n=90); chirp rate of 39.1 ± 4.01 chirps/min (34–48, n=9); dominant frequency of 6.16 ± 0.11 kHz (6.0–6.3, n=9).</p><p>Habitat and male calling site. Males and females live in litter and were quite abundant in forest at border areas. Males often stridulated on soil in sheltered places as under dry leaves. Few specimens were found stridulating between roots or in litter on dry leaves. All males were recorded during daytime.</p><p>Comments. Hygronemobius duckensis sp. nov. belongs to the benoisti group proposed by Desutter- Grandcolas (1993). Among species of this group H. duckensis sp. nov. has the external and the male genitalia morphology very similar to H. tetraplagion . These two species differ from H. elegans by the body pattern coloration to be light yellow with dark spots and by male genitalia, as follow: i) pseudepiphallic sclerite with many long bristles (Fig. 3 B–C); ii) pseudepiphallic apical lobe surpassing pseudepiphallic apex (lateral view, Fig. 3 C); iii) ectophallic fold greatly reduced, badly or not visible in lateral view (Fig. 3 C). Hygronemobius duckensis sp. nov and H. tetraplagion differ from H. diplagion by the light pronotal lateral lobe with small dark spots, tergites II, III, V and VI lighter than other tergites, pseudepiphallic sclerite bristly and longer than rami, ectophallic apodemes and rami parallel, and distinct separation between pseudepiphallic sclerite and rami by membranous area. Hygronemobius duckensis sp. nov and H. tetraplagion differ from H. benoisti by the light spot without a welldefined dark outline in the postero-inferior portion of pronotal lateral lobe, male tegmina with a light spot at the apex, and pseudepiphallic apical lobe reduced, covering only the apex of pseudepiphallic sclerite (in dorsal view, Fig. 3 B). Hygronemobius duckensis sp. nov. differs from H. tetraplagion only by male genitalia, as follow: pseudepiphallus expanded apically surrounded completely the pseudepiphallic apical lobe in dorsal and posterior views (Fig. 3 B); This expansion is curved ventrally with a sub-triangular shape, so it is not possible to see the ectophallic parameres in lateral view (Fig. 3 C); pseudepiphallic apical lobe is reduced not surpassing the apex of pseudepiphallus, when in lateral view (Fig. 3 C). Regarding calling song of H. tetraplagion, there is discordance between description of song and spectrogram presented by Desutter-Grandcolas (1993, pags. 24 and 25), in spectrogram the song consists in many pulses forming long chirps and in description it is characterized as chirps with three pulses. The calling song of Hygronemobius duckensis sp. nov. (Fig. 4 A) presents a similar pattern presented by Desutter-Grandcolas (1993) in the spectrogram, however with fewer pulses and greater interval between pulses.</p><p>Measurements (mm). Female (n=7): body length, 7.06 ± 0.27 (6.70–7.50); pronotum length, 1.57 ± 0.05 (1.50–1.60); pronotum width, 1.91 ± 0.13 (1.80–2.20); head width, 2.06 ± 0.05 (2.00–2.10); length of femur III, 4.27 ± 0.20 (3.90–4.40); length of tibia III, 3,26 ± 0.18 (3.00–3.50); ovipositor length, 2.64 ± 0.26 (2.10–2.90). Male (n=5): body length, 6.42 ± 0.54 (5.90–7.30); pronotum length, 1.20 ± 0.07 (1.10–1.30); pronotum width, 1.84 ± 0.05 (1.80–1.90); head width, 1.94 ± 0.05 (1.90–2.00); length of femur III, 3.96 ± 0.24 (3.70–4.30); length of tibia III, 3.08 ± 0.11 (2.90–3.20); right tegmen length, 3.30 ± 0.16 (3.10–3.50); right tegmen width (dorsal field), 2.50 ± 0.16 (2.30–2.70).</p><p>Type material. Holotype ♂: BRASIL, AM[azonas], Manaus, R.[eserva] F.[lorestal] Adolpho Ducke, AM-010, 21-24.iv.2011, 02º55’49”S, 59°58’31”W. Coleta ativa. L. P. Martins / 69PROSET (INPA). Holotype condition: genitalia placed in microvial with glycerin. Paratypes: same data of Holotype (4♂, 5♀, INPA). idem / 70PROSET (1♂, INPA). idem / 72PROSET (1♂, INPA). idem / 80PROSET (1♂, INPA). idem 01-03.xi.2010. L. P. Martins &amp; D. Mendes (1♀, MZUSP). idem 15-19.xii.2010 / 21PROSET (1♂, INPA). idem / 22PROSET (1♂, INPA). idem / 23PROSET (1♂, MZUSP). idem 26-28.ii.2011. L. P. Martins (1♂, MZUSP). idem 14-19.x.2011. L. P. Martins &amp; A. Souza (8♂, INPA). idem L-08, 500. 02º55’, 03º01’S, 59°53’, 59º59’W. 15.ix a 20.x.2006. Pitfall 0 3. J. L. P. Souza col (1♀, MZUSP). idem L-04, 4500. Pitfall 10 (2♀, INPA). idem L-06, 500. Pitfall 0 1 (1♀, INPA). idem L- 0 3, 4500. Pitfall 0 6 (1♀, INPA).</p></div>	https://treatment.plazi.org/id/5836481EBC3DFFCBFF52F2EBF0FCFBD4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Martins, Luciano De P.;Pereira, Marcelo R.;Henriques, Augusto L.;Zefa, Edison	Martins, Luciano De P., Pereira, Marcelo R., Henriques, Augusto L., Zefa, Edison (2014): Two new species of Hygronemobius Hebard, 1913 (Orthoptera, Grylloidea, Nemobiinae) from Brazilian Amazon. Zootaxa 3794 (3): 469-480, DOI: 10.11646/zootaxa.3794.3.8
5836481EBC3BFFC6FF52F046F022FC28.text	5836481EBC3BFFC6FF52F046F022FC28.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hygronemobius dialeucus Martins & Pereira	<div><p>Hygronemobius dialeucus Martins &amp; Pereira, sp. nov.</p><p>(Figs. 1 C–D; 3D–F; 4B; 5)</p><p>Etymology. The specific name refers to the white markings on the pronotum, abdomen and legs (from Greek dialeukos = marked with white).</p><p>Diagnosis. (i) Head brown without white marks (Figs. 1 C–D, 5A); (ii) pronotum black with white marks on the dorsal disc: anterior margin, a pair of dots; posterior margin, in female, a stripe slightly incurved on the lateral lobes (Fig. 1 D), in male, three dots, the lateral ones bigger (Figs. 1 C, 5A); (iii) femora I and II with an elongated white dot dorsally (Figs. 1 C–D, 5A); (iv) tibia III with an elongated white dot above the last inner dorsal spur (Fig. 1 C–D); (v) femur III with two white dots on the inner face, the distal bigger than the proximal (Figs. 1 C–D, 5A); (vi) first tarsomeres of legs I, II and III with a dorsal elongated white spot (Fig. 1 C–D); (vii) tergite IV of females with a transversal white band (Fig. 1 D); (viii) terminal tergites with a white band or two spots, comprising laterodistal portion of tegite IX, basal portion of supranal plate and cerci adjacent area (Figs. 1 C–D, 5A, E, G); (ix) pseudepiphallic sclerite sub-trapezoidal, posterior margin twice wider than the anterior margin (Fig. 3 E); (x) pseudepiphallic apical lobes longer than half length of pseudepiphallic sclerite (Fig. 3 E); (xi) infero-posterior angle of pseudepiphallic sclerite rounded, in lateral view (Fig. 3 F); (xii) setae present only on pseudepiphallic apical lobes.</p><p>Description. Holotype, male, measurements (mm): body length, 6.0; pronotum length, 1.3; pronotum width, 2.0; head width, 2.0; length of femur III, 4.1; length of tibia III, 3.2; right tegmen length, 2.8; right tegmen width (dorsal field), 3.2. General color of the body black with some white markings. Head brown, except: vertex dark brown but the adjacent area to eyes light brown; occiput light brown; gena light brown; black eyes; scape e pedicel light brown; flagellum with the firsts antennomeres light brown becoming darker towards the apex; frons with an inverted dark brown T-shaped spot; clypeus, lateral portion of the postclypeus dark brown and center-basal portion of the anteclypeus whitish; labrum light brown, basal portion dark brown with a whitish stripe at the apex; Maxilla light brown, lacinia with whitish apex; maxillary palp: first two palpomere dark brown; third palpomere whitish with proximal portion dark brown; fourth palpomere whitish; fifth palpomere dark brown with whitish apex and third proximal portion light yellow; labial palp: first palpomere dark brown; second palpomere dark brown on proximal half and whitish on distal half; third palpomere from whitish to dark brown with apex whitish. Thorax, pronotum black with five white marks on the dorsal disc (Fig. 5 A): anterior margin, a pair of dots; posterior margin, three dots, the lateral ones bigger; lateral lobe evenly black (Fig. 5 C); pronotum with black fine pubescence and strong bristles covering all its extension. Dorsal field of right tegmen dark brown with light yellow spots (Fig. 5 D) situated: 1) close to insertion of tegmen on metanotum; 2) on junction of anal veins; 3) distal portion of cordal area; 4) continuous band between Cu1 and M veins until external portion of apical area; 5) large spot between longitudinal vein in harp and Cu1. Lateral field of right tegmen with five veins; color pattern: a long dark yellow band between R vein and superior vein of lateral field; proximal and supero-distal portions dark brown, remaining areas lighter. Hindwings absent. Prosternum dark brown. Mesosternum dark brown with posterior third light brown. Metasternum dark brown. Legs I and II with similar pattern: coxae and trochanteres dark brown; femora dark brown to black with an elongated white dot on dorsal face (Fig. 5 A); tibiae dark brown, tibia I with an oval tympanum on outer face; tarsi dark brown, first tarsomere with a dorsal elongated white spot. Leg III: coxa and trochanter dark brown; femur III black, knee and proximal area of lighter color, two white dots on the inner face, the distal bigger than the proximal (Fig. 5 A); tibia III black with an elongated white spot close the last inner dorsal spur (Fig. 5 A); tarsus III dark brown, first tarsomere with a dorsal elongated white spot. Abdomen, tergites black; a continuous band comprising latero-distal portion of tegite IX, proximal half of supranal plate and cerci adjacent area (Fig. 5 A); supranal plate with apex rounded (Fig. 5 E); sternites I–IV dark brown, sternites V–VII dark brown in central portion and black in laterals, sternite VIII evenly black (Fig. 5 B); subgenital plate black with distal and lateral margins rounded; cerci light brown. Male genitalia (Fig. 3 D–F): pseudepiphallic sclerite sub-trapezoidal, distal margin twice wider than the proximal margin; pseudepiphallic apical lobes well developed, longer than half length of pseudepiphallic sclerite; numerous smalls setae presents only on pseudepiphallic apical lobes; pseudepiphallic median lobes with distal margin rounded, visible in lateral view; ectophallic fold large, as long as pseudepiphallic sclerite, and as high as pseudepiphallic median lobes (in lateral view, Fig. 3 F); ectophallic apodeme crossing and surpassing rami (Fig. 3 E); endophallus divided in three sclerites, the lateral sclerites triangular and larger than the central one (Fig. 3 D–E).</p><p>Observations in paratypes. Stridulatory file of right tegmen with 80–95 teeth (n=5). White band on terminal tergites not continuous forming two spots separated by a narrow black stripe (Fig. 1 C). Female, body shape and color pattern similar to Holotype, except: tegmina reduced to black lateral scales (Fig. 1 D); posterior margin of pronotum with a wide white stripe slightly incurved on lateral lobes (Fig. 1 D); tergite IV with a wide transversal white band (Fig. 1 D); supranal plate dark brown (Fig. 5 G); subgenital plate from dark to light brown, sub-rounded with a central invagination on apex (Fig. 5 H); all sternites dark brown; prosternum dark brown; mesosternum dark brown with posterior half light brown; metasternum dark brown; ovipositor, valves dark brown, proximal two-thirds of ventral valves light brown (Fig. 5 I); apex of the dorsal valves slightly denticulate on their margins (Fig. 5 I).</p><p>Calling song (25.5–28°C, Tab. 0 1, Fig. 4 B): 5 pulses per chirp, rarely chirps with 4 pulses; chirp duration with 5 pulses of 91 ± 7 ms (83–99, n=40); inter-chirp interval of 272 ± 29 ms (220–330, n=40); chirp rate of 160 ± 15,67 chirps/min (140–176, n=4); pulse duration of 8.13 ± 0.43 ms (7–9, n=30); dominant frequency of 7.3 ± 0.39 kHz (6.8–7.7, n=4).</p><p>Habitat and male calling site. Males and females live in litter and were collected only inside the forest. Males were seen stridulating on soil under dry leaves during daytime.</p><p>Comments. Hygronemobius dialeucus sp. nov. belongs to the amoenus group proposed by Desutter- Grandcolas (1993). The male genitalia of Hygronemobius dialeucus sp. nov. is more similar to that of H. amoenus differing by: pseudepiphallic apical lobes apex longer and thinner (Fig. 3 D–E); infero-posterior angle of pseudepiphallic sclerite rounded (Fig. 3 F). Among species of amoenus group, position and shape of white spots are very important to determining the species, wherefore H. torquatus differs from H. dialeucus sp. nov. by having: dorsal disc of pronotum with a large transversal white stripe near its anterior margin and posterior margin without white markings; only a white dot on femur III; in females, tergites II–III with two pairs of lateral white dots and tergite IV black; H. albolineatus differs from H. dialeucus sp. nov. by having: two longitudinal white stripes running dorsally from fastigium to the posterior margin of pronotum, and in females reaching the tergite IX; a whitish dot behind the eyes; palpi without white markings; tibiae and tarsi without white dots; central portion of posterior margin of pronotum black; tegmina apex whitish. H. boreus differs from H. dialeucus sp. nov. by having: two longitudinal white stripes running dorsally from fastigium to the posterior margin of pronotum; central portion of posterior margin of pronotum black; in females, tergite II with a transversal white stripe, tergite III with white spots on lateral margins and tergite IV with central portion black; tibiae and tarsi without white dots. H. amoenus differs from H. dialeucus sp. nov. by having: head with two longitudinal thin white stripes, from fastigium to occiput; and two longitudinal white stripes along lateral margins of dorsal disc of pronotum. Calling song of H. amoenus, H. albolineatus and H. dialeucus sp. nov. are characterized by brief chirps with 3 pulses in H. amoenus, usually 4 pulses in H. albolineatus (Desutter-Grandcolas 1993) and usually 5 pulses in H. dialeucus sp. nov. (Fig. 4 B).</p><p>Measurements (mm). Male (n=10, excluding holotype). body length, 6.18 ± 0.43 (5.50–6.80); pronotum length, 1.28 ± 0.11 (1.20–1.50); pronotum width, 1.91 ± 0.06 (1.80–2.00); head width, 1.93 ± 0.05 (1.90–2.00); length of femur III, 3.78 ± 0.20 (3.50–4.10); length of tibia III, 3.12 ± 0.14 (2.90–3.30); right tegmen length, 2.95 ± 0.16 (2.70–3.20); right tegmen width (dorsal field), 2.42 ± 0.04 (2.40–2.50). Female (n=5): body length, 5.98 ± 0.41 (5.40–6.30); pronotum length, 1.42 ± 0.08 (1.30–1.50); pronotum width, 1.86 ± 0.05 (1.80–1.90); head width, 2.00 ± 0.10 (1.90–2.10); length of femur III, 4.08 ± 0.18 (3.90–4.30); length of tibia III, 3.36 ± 0.18 (3.10–3.60); ovipositor length, 2.60 ± 0.07 (2.50–2.70).</p><p>Type material. Holotype ♂: BRASIL, AM[azonas], Manaus, R.[eserva] F.[lorestal] Adolpho Ducke, AM-010, 26-31.viii.2011, 02º55’49”S, 59°58’31”W. Coleta ativa. L. P. Martins &amp; V. Linard / 127PROSET (INPA). Holotype condition: detached left leg III; genitalia placed in microvial with glycerin; all parts are maintained in holotype's tube. Paratypes: same data of Holotype (1♂, 1♀, INPA). idem 23-28.ix.2011 / 144PROSET (1♂, INPA). idem 26- 31.i.2012. L. P. Martins &amp; K. Soares (2♂, MZUSP). idem 14-19.x.2011. L. P. Martins &amp; A. Souza / 152PROSET (1♂, INPA). idem / 153PROSET (1♂, INPA). idem / 154PROSET (1♂, INPA). idem (3♂, 3♀, INPA). idem 21- 24.iv.2011. L. P. Martins (1♀, INPA). idem 01-15.vii.2011 (2♂, 1♀, INPA). idem 23-28.ix.2011. L. P. Martins &amp; V. Linard (2♀, MZUSP). idem 01-03.xi.2010. L. P. Martins &amp; D. Mendes (1♂, INPA). idem / 05PROSET (1♂, INPA). idem 26-28.ii.2011 / 49PROSET (1♂, INPA). idem (1♀, INPA). idem 15-19.xi.2013. L. P. Martins &amp; L. G. Da Silva (3♂, INPA).</p></div>	https://treatment.plazi.org/id/5836481EBC3BFFC6FF52F046F022FC28	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Martins, Luciano De P.;Pereira, Marcelo R.;Henriques, Augusto L.;Zefa, Edison	Martins, Luciano De P., Pereira, Marcelo R., Henriques, Augusto L., Zefa, Edison (2014): Two new species of Hygronemobius Hebard, 1913 (Orthoptera, Grylloidea, Nemobiinae) from Brazilian Amazon. Zootaxa 3794 (3): 469-480, DOI: 10.11646/zootaxa.3794.3.8
