taxonID	type	description	language	source
5F30F95FFFFC9030FC8FFE7BFCE75A74.taxon	materials_examined	Material examined. RV Hakuhō Maru KH- 72 - 1 cruise, sta. 54, 1 Ə (CB 10.7 × CL 13.9 mm including pseudorostral tooth), NSMTCr 30899. Remarks. The present male from the station 54 (760 – 777 m depth) agrees well with the specimens from the Timor Sea (535 – 690 m depth) recorded in the second part of this study (Takeda et al., 2022 a: fig. 3 B). Distribution. Deep-sea inhabitants of the West and South Pacific (350 – 935 m): New Zealand, New Caledonia, Vanuatu, eastern Australia, and the South and East China Seas (Ahyong, 2008).	en	Arzivian, Arteen, Alrubaie, Ahmad, Yang, Jessica, Lin, Huiyu, Zhang, Eva, Leong, Rupert (2022): Crabs (Crustacea, Decapoda) from the Seas of East and Southeast Asia Collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 4. South China Sea. Bulletin of the National Museum of Nature and Science. Series A, Zoology 48 (4): 147-191, DOI: 10.50826/bnmnszool.48.4_147, URL: http://dx.doi.org/10.1159/000539401
5F30F95FFFFC9030FF09FCFBFE7A5FE9.taxon	description	(Fig. 1 A) Material examined. RV Hakuhō Maru KH- 72 - 1 cruise, sta. 45, 1 8 (CB 13.1 × CL 15.0 mm including pseudorostral teeth), NSMTCr 30898. Remarks. The female identified here as Mclaydromia colini agrees well with the smaller male from the Sahul Shelf recorded in the third part (Takeda et al., 2022 b: fig. 13 C). The carapace, chelipeds, and ambulatory legs are wholly covered with short, thick, felt-like hairs, and the seemingly bidentate front with small median tooth and prominent pseudorostral tooth on each side. Denudation revealed the carapace anterolateral margin to be armed with two distinct teeth margined with irregular granules. Distribution. Originally reported from New Caledonia at 14 – 62 m depth, and then from the Sahul Shelf, 49 – 52 m depth; now, from the South China Sea at 60 m depth.	en	Arzivian, Arteen, Alrubaie, Ahmad, Yang, Jessica, Lin, Huiyu, Zhang, Eva, Leong, Rupert (2022): Crabs (Crustacea, Decapoda) from the Seas of East and Southeast Asia Collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 4. South China Sea. Bulletin of the National Museum of Nature and Science. Series A, Zoology 48 (4): 147-191, DOI: 10.50826/bnmnszool.48.4_147, URL: http://dx.doi.org/10.1159/000539401
5F30F95FFFFC9036FC80FCD6FEBA5894.taxon	materials_examined	Material examined. RV Hakuhō Maru KH- 72 - 1 cruise, sta. 48, 4 ƏƏ (CB 5.5 × PCL 9.1 mm to 6.1 × 9.6 mm), 4 ovig. 88 (6.7 × 11.0 mm to 7.6 × 11.8 mm), NSMT-Cr 30900. Remarks. The genus Latreillia was extensively revised by Williams (1982) and divided into two genera, Latreillia s. s. with the P 5 propodus fringed with feather-like setae, and a new genus, Eplumura, with the P 5 propodus lacking the conspicuous fringe of setae. The specimen identified as L. pennifera is without doubt referred to Latreillia which comprises L. elegans Roux, 1830 (= L. manningi Williams, 1982 as a synonym) from the Atlantic and Mediterranean Sea, L. williamsi Melo, 1990 from off Brazil, and three Indo-Pacific species, L. valida De Haan, 1839, L. pennifera Alcock, 1900 and L. metanesa Williams, 1982. These species were well figured by Williams (1982) and Castro et al. (2003) based on numerous specimens forming two groups, one with and the other without a dorsal spine on the neck. The present specimen shares the absence of the dorsal spine together with two Atlantic species and L. pennifera from the Indo-West Pacific. In the two Atlantic species, the dactylus of the last leg (P 5) forms a subchela with subdistal spinules of the propodus. Latreillia pennifera is seemingly close to L. valida, but in addition to the distinctive differences mentioned above, the carapace is less stout, with the longer and slender neck, as figured by Alcock (1901: pl. 7 fig. 27) and Castro et al. (2003: fig. 10). Distribution. Known localities are mostly in the Indian Ocean from the Gulf of Martaban and the Mergui Archipelago to South Africa, and otherwise recorded from the Kai Islands, Indonesia, and Mindanao, the Philippines. Its bathymetric range is 37 – 229 m.	en	Arzivian, Arteen, Alrubaie, Ahmad, Yang, Jessica, Lin, Huiyu, Zhang, Eva, Leong, Rupert (2022): Crabs (Crustacea, Decapoda) from the Seas of East and Southeast Asia Collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 4. South China Sea. Bulletin of the National Museum of Nature and Science. Series A, Zoology 48 (4): 147-191, DOI: 10.50826/bnmnszool.48.4_147, URL: http://dx.doi.org/10.1159/000539401
5F30F95FFFFA9036FF2EFD76FCA358C9.taxon	materials_examined	Material examined. RV Hakuhō Maru KH- 72 - 1 cruise, sta. 48, 1 Ə (CB 5.3 mm excluding hepatic tubercles, PCL 6.2 mm), 1 ovig. 8 (6.6 × 8.0 mm), NSMT-Cr 30901. — Sta. 50, 1 ovig. 8 (13.4 × 15.5 mm), NSMT-Cr 30902. Remarks. The genus Latreillopsis was extensively studied, with detailed notes, line drawings and photographs, by Guinot and Richer de Forges (1995), who recognized L. bispinosa Henderson, 1888 (and its new forma L. bispinosa f. trispinosa), L. laciniata Sakai, 1936, L. gracilipes Guinot and Richer de Forges, 1981, L. tetraspinosa Dai and Chen, 1980, and three new species, L. daviei, L. antennata and L. cornuta. Richer de Forges and Ng (2007) subsequently recorded L. bispinosa and L. tetraspinosa, with many specimens from the Philippines and described a new species, L. marivenae from the Philippines, Richer de Forges and Ng (2007) also recorded L. gracilipes from Vanuatu, and L. tetraspinosa from the Solomon Islands, and formally named L. trispinosa, a replacement name of the nomenclaturally incorrect L. bispinosa f. trispinosa Guinot and Richer de Forges, 1995. Castro and Naruse (2014) described L. okala from Hawaii. The present ovigerous female studied here was confirmed as L. bispinosa following the key and many photographs and detailed drawings given by Guinot and Richer de Forges (1995), and the figures and photograph by Ahyong et al. (2009). The most distinctive features are the unarmed pseudorostral spines and the subhepatic protuberance armed with two long spines instead of four spines in the most closely related species, L. tetraspinosa Dai and Chen, 1980. Distribution. Japan, Taiwan, South China Sea, and the Philippines, 20 – 350 m in depth. Chen and Xu (1991) recorded this species from the Nansha Islands in the South China Sea, at 185 m depth.	en	Arzivian, Arteen, Alrubaie, Ahmad, Yang, Jessica, Lin, Huiyu, Zhang, Eva, Leong, Rupert (2022): Crabs (Crustacea, Decapoda) from the Seas of East and Southeast Asia Collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 4. South China Sea. Bulletin of the National Museum of Nature and Science. Series A, Zoology 48 (4): 147-191, DOI: 10.50826/bnmnszool.48.4_147, URL: http://dx.doi.org/10.1159/000539401
5F30F95FFFFA9034FD48FD55FD115AC2.taxon	description	(Fig. 2 A – B) Material examined. RV Hakuhō Maru KH- 72 - 1 cruise, sta. 45, 1 8 (CB 5.1 × CL 6.1 mm), NSMT-Cr 30903. Remarks. In describing the third species of the genus Cosmonotus, C. mclaughlinae, Tavares (2006) suggested that C. grayii recorded by Takeda and Miyake (1970) from Japanese waters may be referable to an unknown species different from both of C. grayii Adams, in Belcher, 1847, and C. genkaiae Takeda and Miyake, 1970. This suggestion stemmed mainly from the fact that the supraorbital border of C. grayii was illustrated as having only one notch by Takeda and Miyake (1970: fig. 1 A) for a specimen from off the Tsushima Islands, north of Kyushu. During this study, we could not re-examine the specimen reported by Takeda and Miyake (1970) and Takeda (1973 b), but examined several additional Japanese specimens as follows: — Shimoda, Sagami Bay (1 8, NSMT-Cr 7913); Suruga Bay (1 Ə, 1 Ə infested by a Sacculina, NSMT-Cr 12054); Kushimoto, Kii Peninsula (2 ƏƏ, 1 8, NSMT-Cr 6388); Tosa Bay (1 Ə, NSMT-Cr 12772); West of Goto Islands (1 Ə, NSMT-Cr 9789); Tsushima Islands (4 ƏƏ, 1 8, NSMT-Cr 979, 980, 982, 983). All of them proved to be referable to C. mclaughlinae, agreeing well with the accounts by Tavares (2006) and the subsequent record by Seo et al. (2018); the main part of the carapace dorsal surface is smooth, but the anterolateral portion is distinctly roughened with minute granules of irregular size, and the carapace midline is rather thickened and ridged for much of its length, somewhat like a lbackbone. z	en	Arzivian, Arteen, Alrubaie, Ahmad, Yang, Jessica, Lin, Huiyu, Zhang, Eva, Leong, Rupert (2022): Crabs (Crustacea, Decapoda) from the Seas of East and Southeast Asia Collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 4. South China Sea. Bulletin of the National Museum of Nature and Science. Series A, Zoology 48 (4): 147-191, DOI: 10.50826/bnmnszool.48.4_147, URL: http://dx.doi.org/10.1159/000539401
5F30F95FFFFA9034FD48FD55FD115AC2.taxon	description	The authorship and publication year of the genus Cosmonotus and the species, C. grayii, were clarified by Clark and Presswell (2001). The description of C. grayii appeared in two papers of same title and content, lShort descriptions of new or little-known decapod Crustacea, z published in l Proceedings of the Zoological Society of London, 15 (1847) and l The Annals and Magazine of Natural History, including Zoology, Botany, and Geology, ser. 2, vol. 2 (1848), z but their actual dates of publication were indicated by them as 29 March and October in 1848, respectively. According to Low et al. (2020), however, the brief explanations of the genus and species by Adams (1847) in Narrative of the Voyage of H. M. S. Samarang are available for the original definition and description of a new genus and a new species, Cosmonotus grayii. Distribution. Takeda and Miyake (1970) provided the synonymy list of C. grayii known to date including the literature recording the specimens from Japanese waters (Yokoya, 1933; Sakai, 1937, 1965). The type locality is Borneo, with a wide geographical range in the Indo-West Pacific from the West Pacific to Western Australia and to the east coast of Africa and the Red Sea, with a bathymetric range of 30 – 212 m. There may be some misidentifications with C. mclaughlinae from the West Pacific, and therefore the specimens from Taiwan, the Philippines, and the South China Sea should be re-examined to determine whether both species are sympatric, parapatric or allopatric in their distributional patterns.	en	Arzivian, Arteen, Alrubaie, Ahmad, Yang, Jessica, Lin, Huiyu, Zhang, Eva, Leong, Rupert (2022): Crabs (Crustacea, Decapoda) from the Seas of East and Southeast Asia Collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 4. South China Sea. Bulletin of the National Museum of Nature and Science. Series A, Zoology 48 (4): 147-191, DOI: 10.50826/bnmnszool.48.4_147, URL: http://dx.doi.org/10.1159/000539401
5F30F95FFFF89034FC96FB93FB6F5FE9.taxon	materials_examined	Material examined. RV Hakuhō Maru KH- 72 - 1 cruise, sta. 45, left cheliped, NSMT-Cr 30904. Remarks. Only the left cheliped remains in a vial, with a large rounded blotch each on the upper part of the palm outer surface and the basal part of the carpus outer surface. Calappa philargius is one of the more common representatives of the shallow-water species of Calappa in the Indo-West Pacific (Galil, 1997). Distribution. West and South Pacific from Japan to Australia and New Caledonia, the Indian Ocean, and the Red Sea, 30 – 100 m depth.	en	Arzivian, Arteen, Alrubaie, Ahmad, Yang, Jessica, Lin, Huiyu, Zhang, Eva, Leong, Rupert (2022): Crabs (Crustacea, Decapoda) from the Seas of East and Southeast Asia Collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 4. South China Sea. Bulletin of the National Museum of Nature and Science. Series A, Zoology 48 (4): 147-191, DOI: 10.50826/bnmnszool.48.4_147, URL: http://dx.doi.org/10.1159/000539401
5F30F95FFFF99035FF5EFE3BFCB35B2E.taxon	description	(Fig. 1 B) Material examined. RV Hakuhō Maru KH- 72 - 1 cruise, sta. 52, 1 juv. (CB 8.2 mm excluding lateral tubercles, CL 7.4 mm), NSMTCr 30905. Remarks. The present juvenile specimen referred here to Mursia australiensis agrees well with the descriptions and photographs given by the original author (Campbell, 1971), Galil (1993), and Mendoza and Nugroho (2021). The carapace is narrow and convex in both directions; the anterolateral margin is only weakly convex, and the posterolateral margin is strongly convergent toward the lateral end of the carapace posterior margin. The carapace dorsal surface is densely covered with granules and sculptured with three longitudinal ridges at the median gastric, cardiac and intestinal regions and at each branchial region; three ridges are fringed each with a row of bosses; four bosses in a transverse row at the anterior part of the mesogastric and protogastric regions, four in a longitudinal row on the posterior part of the mesogastric, cardiac and intestinal regions, and three in an oblique row on the branchial region. The carapace posterior margin is narrow, and angulated at each lateral end, its median part being produced into an obtuse tubercle that is slightly smaller than the carapace lateral end. Each lateral tubercle of the carapace is sharp, one fourth as wide as the carapace, and weakly directed posterolaterally and obliquely upward. The tubercle at the distal end of the cheliped merus is also sharp, and as long as or slightly longer than the carapace lateral tubercle, with the basal half directed horizontally and distal half directed posterolaterally.	en	Arzivian, Arteen, Alrubaie, Ahmad, Yang, Jessica, Lin, Huiyu, Zhang, Eva, Leong, Rupert (2022): Crabs (Crustacea, Decapoda) from the Seas of East and Southeast Asia Collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 4. South China Sea. Bulletin of the National Museum of Nature and Science. Series A, Zoology 48 (4): 147-191, DOI: 10.50826/bnmnszool.48.4_147, URL: http://dx.doi.org/10.1159/000539401
5F30F95FFFF99035FF5EFE3BFCB35B2E.taxon	distribution	Distribution. Previously known from Australia (southern Queensland and New South Wales, 100 – 136 m), New Caledonia (150 – 320 m) and Indonesia (Sunda Strait and western Java, 234 – 281 m). Otherwise, questionably from Tosa Bay, the Pacific coast of Japan, 280 m (Miyake, 1983, as M. aspera).	en	Arzivian, Arteen, Alrubaie, Ahmad, Yang, Jessica, Lin, Huiyu, Zhang, Eva, Leong, Rupert (2022): Crabs (Crustacea, Decapoda) from the Seas of East and Southeast Asia Collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 4. South China Sea. Bulletin of the National Museum of Nature and Science. Series A, Zoology 48 (4): 147-191, DOI: 10.50826/bnmnszool.48.4_147, URL: http://dx.doi.org/10.1159/000539401
5F30F95FFFF9903AFD42FD95FEFA587C.taxon	description	(Fig. 1 E – F) Material examined. RV Hakuhō Maru KH- 72 - 1 cruise, sta. 42, 1 juv. (CB 5.6 × CL 6.6 mm including posterior tubercle), NSMT-Cr 30906. Remarks. The genus Parilia was extensively revised by Ng et al. (2018) who recognized four species: P. alcocki Wood-Mason, in Wood-Mason and Alcock, 1891, P. major Sakai, 1961, P. ovata Chen, 1984, and P. pattersoni Ng, Devi and Kumar, 2018. The juvenile male examined is somewhat different from the full-grown specimens described by Ng et al. (2018) in having a narrower, longitudinally ovate carapace similar to P. ovata, but agrees with the photograph and comments on the young male recorded by Mendoza and Nugroho (2021). In P. ovata, the chelipeds and ambulatory legs are nearly smooth, and each carapace lateral margin is armed with only a small tubercle, while in the present species the chelipeds and ambulatory legs are thickly covered with minute granules and tubercles, and each carapace lateral margin is armed with three low, but distinct tubercles that are distinct in the juvenile specimen. Distribution. Tamil Nadu, southeastern India (Ng et al., 2018), and off Nias Island in eastern Sumatra (Doflein, 1904, as P. alcocki) and Pelabuhanratu Bay in southwestern Java, 183 – 255 m depth (Mendoza and Nugroho (2021). As Ng et al. (2018) mentioned, the record of P. major from the Gulf of Manaar, southeast India, by Vidhya et al. (2017) may be referred to that of P pattersoni.	en	Arzivian, Arteen, Alrubaie, Ahmad, Yang, Jessica, Lin, Huiyu, Zhang, Eva, Leong, Rupert (2022): Crabs (Crustacea, Decapoda) from the Seas of East and Southeast Asia Collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 4. South China Sea. Bulletin of the National Museum of Nature and Science. Series A, Zoology 48 (4): 147-191, DOI: 10.50826/bnmnszool.48.4_147, URL: http://dx.doi.org/10.1159/000539401
5F30F95FFFF6903AFF7BFEDFFE165D79.taxon	materials_examined	Material examined. RV Hakuhō Maru KH- 72 - 1 cruise, sta. 54, 5 ƏƏ (CB 19.7 × CL 19.9 mm to 28.7 × 30.3 mm), 6 88 (17.2 × 17.4 mm to 19.3 × 19.1 mm), NSMT-Cr 30907. Remarks. Cyrtomaia suhmi is not uncommon in West Pacific waters, and in the present series of study, two specimens having a deformed carapace were already recorded from the depths of 610 – 690 m in the Timor Sea (Takeda et al. 2022 a). Eleven specimens at hand have no distinct morphological abnormalities, and they are preserved with a number of detached ambulatory legs. Distribution. Indo-West Pacific, from Japan to Australia and India through the Philippines and Indonesia, 488 – 1125 m depth.	en	Arzivian, Arteen, Alrubaie, Ahmad, Yang, Jessica, Lin, Huiyu, Zhang, Eva, Leong, Rupert (2022): Crabs (Crustacea, Decapoda) from the Seas of East and Southeast Asia Collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 4. South China Sea. Bulletin of the National Museum of Nature and Science. Series A, Zoology 48 (4): 147-191, DOI: 10.50826/bnmnszool.48.4_147, URL: http://dx.doi.org/10.1159/000539401
5F30F95FFFF6903AFF44FBC5FB6B58B6.taxon	description	(Fig. 3 A) Material examined. RV Hakuhō Maru KH- 72 - 1 cruise, sta. 54, 1 8 (CB 6.6 × PCL 8.5 mm), NSMT-Cr 30908. Remarks. A fully-grown female at hand generally agrees with the original description (Sakai 1932, as Achaeopsis), though the protogastric and epibranchial spines (Fig. 3 A) are not prominent as in the figures represented by Sakai (1932, fig. 3; 1938, fig. 17) and also in the female from the Timor Sea (Takeda et al. 2022 a, fig. 5 C – D). The carapace tuberculation of the present specimen is more distinct than the Timor Sea specimen. Distribution. This species was previously known from Japan, the Kai Islands, and the Timor Sea, 170 – 650 m in depth. This is a new record from the South China Sea. The present bathymetric record, 777 m in the South China Sea, is the deepest known for this species.	en	Arzivian, Arteen, Alrubaie, Ahmad, Yang, Jessica, Lin, Huiyu, Zhang, Eva, Leong, Rupert (2022): Crabs (Crustacea, Decapoda) from the Seas of East and Southeast Asia Collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 4. South China Sea. Bulletin of the National Museum of Nature and Science. Series A, Zoology 48 (4): 147-191, DOI: 10.50826/bnmnszool.48.4_147, URL: http://dx.doi.org/10.1159/000539401
5F30F95FFFF6903AFCD3FD1FFBD85CA1.taxon	description	(Fig. 3 B) Material examined. RV Hakuhō Maru KH- 72 - 1 cruise, sta. 48, 1 Ə (CB 3.4 × PCL 4.3 mm), 1 8 (4.5 × 6.0 mm), NSMT-Cr 30909. Remarks. In the genus Oncinopus, this species is unique in having the combination of the short, triangular, anterolaterally directed rostral lobes separated by a V-shaped hiatus, the interorbital region concealing the basal part of the unretracted eyestalk, the male G 1 being strongly widened subapically, and the female gonopore on a weak elevation and opening more posteriorly than medially. Both male and female specimens in the present collection fully agree with the description by Takeda and Miyake (1969 a) and a recent diagnosis given by Davie (2011). Distribution. Ogasawara Islands (Takeda and Miyake, 1969 a; Takeda, 1973 a; Takeda and Kurata, 1976), the East China Sea (Takeda and Miyake, 1969 a), the Philippines and Indonesia (Griffin and Tranter, 1986), and Western Australia (Davie, 2011); 36 – 200 m depth.	en	Arzivian, Arteen, Alrubaie, Ahmad, Yang, Jessica, Lin, Huiyu, Zhang, Eva, Leong, Rupert (2022): Crabs (Crustacea, Decapoda) from the Seas of East and Southeast Asia Collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 4. South China Sea. Bulletin of the National Museum of Nature and Science. Series A, Zoology 48 (4): 147-191, DOI: 10.50826/bnmnszool.48.4_147, URL: http://dx.doi.org/10.1159/000539401
5F30F95FFFF6903EFCA5F90DFB635BA4.taxon	description	(Figs. 4, 7 A – C) urn: lsid: zoobank. org: act: A 7170734 - 185 C- 402 B- 9 B 18 - A 2 F 28 FD 3 DEBF Material examined. RV Hakuhō Maru KH- 72 - 1 cruise, sta. 50, Ə (CB 8.8 mm excluding lateral spines, PCL 14.3 mm excluding intestinal spine), holotype, NSMT-Cr 30910; Sta. 54, 1 8 (CB 12.3 × PCL 16.9 mm excluding intestinal spine), non-type, NSMT-Cr 30911. Description of holotype. Carapace (Fig. 4 A, B) pyriform, surface thickly tomentose; gastric, cardiac, branchial, intestinal regions welldefined, moderately elevated. Gastric region low, with 4 long spines, 10 tubercles; anterior end with 2 acute tubercles just behind base of pseudorostrum; anterior slope medially with slender spine, low, minute spine on both sides; protogastric spines short, directed anterolaterally, followed by 2, longitudinally lined small tubercles; mesogastric spine largest (though broken), erected upright, with 2 transversely lined low acute tubercles. Cardiac region conical, dorsally elongated into long slender spine (though broken), divided on both lateral sides, each with large conical tubercle, lateral surface bearing slender spine. Branchial region not markedly inflated, longitudinally with 2 long, slender mesobranchial spines of similar size, small spine in front of anterior spine, anteromesially with large protuberance bearing small tubercles; lateral subsurface with 3 slender spines of similar size (though second one broken); metabranchial region with small medial spine just anterior to posterolateral carapace margin. Intestinal region elongated posteriorly into large, conical projection (though broken). Pseudorostrum (Fig. 4 A – D) distinctly divergent (ca. 30 °), straight in lateral view (Fig. 4 C). Pseudorostral spine slender, dorsal surface weakly ridged toward frontal region, bearing short, subdistal spine dorsomesially (distal part broken). Supraorbital eave (Fig. 4 A – B) strongly expanded laterally; anterior angle produced into short, blunt preorbital spine erected uprightly; antorbital angle produced into minute conical tooth, directed laterally. Dorsal orbital hiatus rounded (Fig. 4 B). Postocular cup faintly bilobed (Fig. 4 C); upper postorbital margin broadly concave (in dorsal view), proximally with distinct rounded intercalated lobe; lateral wall consisting of 2 faint lobes, upper lobe with minute medial tubercle; lower postorbital margin sinuous; small but distinct tubercle right behind ventral orbital hiatus. Pterygostomian region moderately inflated, armed with large conical tubercle directed ventrally (Fig. 4 D). Basal antennal article (Fig. 4 D) narrow, distolateral angle produced anterolaterally into acute tooth, visible in dorsal view; lateral margin generally straight; strong triangular tooth near midlength, directed laterally, as high as distolateral spine. Penultimate antennal articles slender. Chelipeds slender, tomentose, sparsely covered with long stick-like setae; merus subcylindrical, distally with sharp dorsal spine; carpus globulous; palm slender, generally subcylindrical, slightly compressed distally; fingers almost half of palm in length, occlusal margins finely dentate, contiguous in distal half. Ambulatory legs with cylindrical articles, surface tomentose, sparsely with long stick-like setae. Merus slender, subcylindrical, distally with sharp dorsal spine. Thoracic sternites 1 – 3 unarmed, sternite 4 posteromesially with small spine medially on pleonal cavity, sternites 5 – 6 with large conical tubercle of similar size, sternite 7 with minute medial spine, sternite 8 unarmed. Pleon with 6 free somites and telson, each with distinct medial tubercle, lateral part of somites 3 – 4 roundly elevated. G 1 (Fig. 7 A – C) shaft stout basally, gently curved laterally, tip sharply pointed, aperture subterminal, without lobular projection. Etymology. Named for the elongate carapace characteristic for this species. Remarks. The present new species is close to Naxioides sahulensis from the Sahul Shelf in the previous paper of this series (Takeda et al., 2022 b). However, in the new species, five dorsal spines on the carapace midline are broader than those of N. sahulensis, in which they are immediately narrowed (Takeda et al., 2022 b, fig. 7 D; otherwise, the hepatic region bears one minute tubercle and one slender lateral spine in the new species, rather than one small tubercle and two lateral spines in N. sahulensis (Takeda et al., 2022 b, fig. 7); the pterygostomian region is armed with a single large conical spine in the new species, but there are small additional spines on both sides in N. sahulensis (Takeda et al., 2022 b, fig. 7 C); the preorbital spine is erect and almost upright in the new species, but only obliquely inclined in N. sahulensis (Takeda et al., 2022 b, fig. 7); the lateral spines on basal antennal article are more anteriorly directed (Takeda et al., 2022 b, fig. 7 D) in N. sahulensis than in the new species. Another close relative, N. tenuirostris (Haswell, 1880), shares the upright preorbital angle, but it is much shorter in N. elongatus sp. nov. The antorbital angle in N. elongatus bears a small conical tubercle directed laterally, rather than the more acuminated, posteriorly directed spine in N. tenuirostris as in Griffin (1966, fig. 5). The anterior margin of the lateral orbital wall is bilobed in the new species, but it is entire in N. tenuirostris as seen in Griffin (1966, fig. 8). The cheliped carpus is unarmed in the new species, whereas in N. tenuirostris, it bears two small spines (Griffin, 1966, fig. 5). In N. tenuirostris, the lateral margins of the male pleon are weakly concave (Griffin, 1966, fig. 6), but not in the new species. The male and female specimens of Naxioides elongatus sp. nov. agree in most respects, but the female has a proportionally slightly wider carapace (Fig. 4 A, E), and the lateral margin of basal antennal article differs slightly from the holotype, with the anteriorly produced distolateral spine (versus anterolaterally) and the slightly anterolaterally directed subproximal marginal spines (versus laterally directed) (Fig. 4 D, H). Unfortunately, the female is badly damaged, missing the P 2 and P 5 on both sides, left P 4, and pleon, with the damaged and somewhat fragile pseudorostral horns, gastric, cardiac, intestinal, pterygostomian spines and thoracic sternites. Given the poor condition of the female, it is not treated as a paratype. The carapace of the holotype (Fig. 4 A – C) is somewhat more elongated than in males either of the two close northern Australian relatives, N. tenuirostris and N. sahulensis from the Torres Strait and the Sahul Shelf, respectively. The young male from the Ogasawara Islands identified as Phalangipus hystrix (Miers, 1886) by Komatsu (2011) and the record of Naxioides tau- rus (Pocock, 1890) from the Philippines, 120 – 124 m at depth, by Richer de Forger and Ng (2013) may be referable to the present new species. Griffin and Tranter (1986) largely revised the genus Chlorinoides Haswell, 1880 (type species: Chlorinoides tenuirostris Haswell, 1880), and transferred most species of Chlorinoides known at that time to the new genus Thacanophrys. However, Prismatopus Ward, 1933, has a priority over Thacanophrys as a senior synonym (Ng et al., 2001; Ng et al., 2008). As regards the systematic position of C. tenuirostris, Griffin and Tranter (1986) mentioned that C. tenuirostris differs significantly from all the other species [of Thacanophrys] and in respect of the G 1, the structure of the orbit and several other features, C. tenuirostris has a much closer relationship to the species of Naxioides than to the species previously grouped with it.	en	Arzivian, Arteen, Alrubaie, Ahmad, Yang, Jessica, Lin, Huiyu, Zhang, Eva, Leong, Rupert (2022): Crabs (Crustacea, Decapoda) from the Seas of East and Southeast Asia Collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 4. South China Sea. Bulletin of the National Museum of Nature and Science. Series A, Zoology 48 (4): 147-191, DOI: 10.50826/bnmnszool.48.4_147, URL: http://dx.doi.org/10.1159/000539401
5F30F95FFFF6903EFCA5F90DFB635BA4.taxon	distribution	Distribution. The description of the new species is based on two specimens from the South China Sea; 132 – 137 and 760 – 777 m depth. This species probably occurs in the Philippines and off Chichi-jima Island, Ogasawara Islands.	en	Arzivian, Arteen, Alrubaie, Ahmad, Yang, Jessica, Lin, Huiyu, Zhang, Eva, Leong, Rupert (2022): Crabs (Crustacea, Decapoda) from the Seas of East and Southeast Asia Collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 4. South China Sea. Bulletin of the National Museum of Nature and Science. Series A, Zoology 48 (4): 147-191, DOI: 10.50826/bnmnszool.48.4_147, URL: http://dx.doi.org/10.1159/000539401
5F30F95FFFF2903EFC86FC23FBAE5F3E.taxon	description	(Fig. 3 C – E) Material examined. RV Hakuhō Maru KH- 72 - 1 cruise, sta. 54, 1 Ə (CB 5.1 mm excluding lateral plates, PCL 8.8 mm), 1 ovig. 8 (6.7 × 11.6 mm), NSMT-Cr 30912; 1 ovig. 8 (7.8 × 12.7 mm), NSMT-Cr 30913; 1 8 (5.0 × 8.3 mm), NSMT-Cr 30914. Remarks. The specimens (Fig. 3 C – E) examined agree well with the original description (Rathbun, 1916), and the accounts (Griffin, 1976; Richer de Forges and Ng, 2009). Oxypleurodon sphaenocarcinoides was already recorded from the Sulu Sea by Takeda et al. (2021) at a depth of 285 – 306 m. Distribution. The Philippines and the South China Sea; 93 – 306 m depth. The present record of O. sphenocarcinoides is the first for the South China Sea, and the bathymetric record of 760 – 777 m is the deepest for the species.	en	Arzivian, Arteen, Alrubaie, Ahmad, Yang, Jessica, Lin, Huiyu, Zhang, Eva, Leong, Rupert (2022): Crabs (Crustacea, Decapoda) from the Seas of East and Southeast Asia Collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 4. South China Sea. Bulletin of the National Museum of Nature and Science. Series A, Zoology 48 (4): 147-191, DOI: 10.50826/bnmnszool.48.4_147, URL: http://dx.doi.org/10.1159/000539401
5F30F95FFFF39023FF66FE04FB735851.taxon	description	(Figs. 5 – 6, 7 D – G) urn: lsid: zoobank. org: act: 19 A 47 CD 2 - 4868 - 4 AD 0 - B 160 - CA 19 FC 43 A 550 Material examined. RV Hakuhō Maru KH- 72 - 1 cruise, sta. 54, Ə (CB 10.7 mm excluding branchial spines, PCL 16.3 mm), holotype, NSMT-Cr 30915; 1 Ə (8.8 × 13.6 mm), 1 8 (10.5 × 15.9 mm), paratypes, NSMT-Cr 30916; 1 ovig. 8 (10.8 × 16.3 mm), paratype, NSMT-Cr 30951; 1 8 (8.3 × 13.0 mm), paratype, NSMT-Cr 30917; 1 8 (12.8 × 18.9 mm), paratype, NSMTCr 30918. Description of holotype. Carapace (Fig. 5 A) pyriform, entirely covered with tomentum (mostly abraded); surface smooth. Gastric region weakly elevated, medially with obtusely pointed conical mesogastric tubercle between low, rounded, protogastric protuberance on each side; cardiac region weakly elevated, with large conical tubercle. Branchial region with 3 distinct, short, conical tubercles, prebranchial region slightly elevated, with 2 tubercles obliquely (mesial one larger than lateral one), metabranchial tubercle shorter than cardiac tubercle; epibranchial spine long, 1.1 times as long as CB excluding spines, slender, directing laterally and obliquely upward (Fig. 5 B – C). Intestinal region low, medially with short, dorsally directed spine on medially convex, posterior carapace margin. Pterygostomian region (Fig. 6 D) not markedly inflated, with 4 low tubercles on gently ridged, pleural suture. Pseudorostral spines (Fig. 5 A, D) slender, 0.7 × PCL, widely separated from bases, weakly outcurved laterally. Supraorbital eave strong, anteriorly produced into subacute, dorsolaterally directed preorbital lobe, with smooth subsurface; postorbital lobe separated from eave by keyholeshaped orbital hiatus; postorbital lobe smooth, compressed laterally (= postorbital plate), subcircular, forming plate-like lateral wall of incomplete orbit. Hepatic margin laterally with dorsally directed elongate lobe (= hepatic plate), incurved distally, lateral surface flattened, smooth, ovoid, with acuminate tip, separated from postorbital plate deep sulcus. Basal antennal article (Fig. 6 D) smooth, surface smooth, distolateral angle produced into short tooth, lateral margin entire, straight, proximally with small acute tubercle sealed by laterally produced, anterolateral angle of buccal cavity (Fig. 7 D). Penultimate article slender, length two-thirds that of ultimate peduncular article. Flagellum not reaching tip of pseudorostral spines. Cheliped (Figs. 5 A – B, 6 D) with few setae and sparse tomentum on lower margin of merus and inner margin of carpus; merus trigonal in crosssection, margins sharply carinate for entire length; dorsal carina irregularly dentate in proximal half, distally with strong, anteriorly directed tooth (Fig. 5 A – B); ventromesial carina entire (Fig. 5 A); ventrolateral carina tri-dentate (Fig. 5 B). Carpus strongly crested on dorsal, mesial, lateral margins. Chela moderately compressed, tapering toward fingertips; palm sharply carinate on upper and lower margins; fingers almost as long as palm, slightly curved distally, occlusal margins weakly dentate, without any prominent tooth subproximally, with slight gap when closed. Ambulatory legs (Fig. 5 A) slender, tomentose. Merus cylindrical, almost one-third length of PCL in P 5. Carpus simple. Propodus subcylindrical. Dactylus slender, faintly falcate, with 0, 4 – 5 denticles on flexor surface in P 2, P 3, respectively; weakly falcate, with 5 or 6 low, rounded tubercles on flexor surfaces in P 4, 5, respectively. Pleon (Fig. 6 D) with six somites and triangular telson; pleomeres 3 – 5 medially ridged; pleomere 6 with large tubercles on both sides, short, medial tubercle anteriorly. G 1 (Fig. 7 D – F) shaft straight, not markedly different from that of S. kotakae (Takeda, 2001). G 2 (Fig. 7 G) short, simple. Notes on paratypes. Erect tubercles on carapace dorsal surface shorter, less distinct in females than in males (Fig. 6 B), reduced in smaller specimens (Fig. 6 C). Pseudorostrum not markedly shorter in females, but less divergent anteriorly when compared with males (Fig. 6 A). Relative length of epibranchial spines against CW 0.7 – 0.9 (N = 4), smaller than in males (1.1, N = 2) (Fig. 6 A). Postorbital, hepatic plates continuous, but the degree of fusion somewhat variable probably sexually and / or ontogenetically (Fig. 6 B – C). Chela slender, palm not so sharply carinate on both upper and lower margins, fingers simply dentate, not gaping when closed in females (Fig. 6 E). Female pleon covered with a thick tomentum in both adolescent and fully grown specimens (Fig. 6 E). Etymology. Named after the RV Hakuhō Maru. Remarks. The genus Samadinia was established by Ng and Richer de Forges (2013). Lee et al. (2021) transferred 26 species previously placed in Rochinia to Samadinia, and subsequently, Richer de Forges et al. (2021) recorded four additional species; S. livermorii (Wood-Mason, in Wood-Mason and Alcock, 1891) was transferred from Scyramathia A. Milne-Edwards, 1880, and three new species, S. jefrii, S. taylorae, and S. yoyoae were described from Indonesian waters. Prior to the present study, Samadinia included 30 species, all from Indo-Pacific waters.	en	Arzivian, Arteen, Alrubaie, Ahmad, Yang, Jessica, Lin, Huiyu, Zhang, Eva, Leong, Rupert (2022): Crabs (Crustacea, Decapoda) from the Seas of East and Southeast Asia Collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 4. South China Sea. Bulletin of the National Museum of Nature and Science. Series A, Zoology 48 (4): 147-191, DOI: 10.50826/bnmnszool.48.4_147, URL: http://dx.doi.org/10.1159/000539401
5F30F95FFFF39023FF66FE04FB735851.taxon	description	The following three species shares with S. hakuhoae sp. nov. the postorbital and strongly upturned hepatic plates that are distinctly separated by a distinct gap: S. galatheae (type locality: off Natal, 535 – 610 m), S. sibogae (Ceram Sea, 924 m), and S. strangeri (Serène and Lohavanijaya, 1973) (South China Sea, 479 m). However, S. galatheae has more rounded carapace with less divergent pseudorostral spines and its postorbital-hepatic plates are fused to one another (Griffin and Tranter, 1986, fig. 11). The posteriorly-produced intestinal spine is very long (see Richer de Forges and Poore, 2008, fig. 2 C). In S. strangeri, the lateral branchial spines are short and similar to the other dorsal spines in length (Lee et al., 2017, fig. 9). In these three species, the G 1 form is also different from that of S. hakuhoe sp. nov. (Fig. 7 A – C vs. Lee et al., 2017, figs. 9, 11 A – D). Distribution. Known only from the type locality, South China Sea, 760 – 777 m depth.	en	Arzivian, Arteen, Alrubaie, Ahmad, Yang, Jessica, Lin, Huiyu, Zhang, Eva, Leong, Rupert (2022): Crabs (Crustacea, Decapoda) from the Seas of East and Southeast Asia Collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 4. South China Sea. Bulletin of the National Museum of Nature and Science. Series A, Zoology 48 (4): 147-191, DOI: 10.50826/bnmnszool.48.4_147, URL: http://dx.doi.org/10.1159/000539401
5F30F95FFFEF9023FD53FD13FCD85C87.taxon	description	(Fig. 8) Material examined. RV Hakuhō Maru KH- 72 - 1 cruise, sta. 42, 1 Ə (CB 36.7 × CL 25.7 mm), NSMT-Cr 30919. Remarks. Takeda (2019) recorded one male and two females of this species as Charybdis (Goniohellenus), with photographs, from off Hong Kong, and mentioned the differences from the closely related congeneric species, C. (A.) truncata (Fabricius, 1798), following Shen (1934) and Leene (1938). The shape of the six frontal lobes is somewhat variable and so more emphasised in the present male (Fig. 8 A), as useful to discriminate C. (A.) hongkongensis from congeners. Distribution. Taiwan; Hong Kong; coasts of China; Sunda Strait, Madura Strait, Banda Sea; Andaman Sea. Bathymetric range is 15 – 60 m as summarized by Takeda (2019), with an exceptional depth record of 397 m by Leene (1938) in the Banda Sea. Chen (1998 a) recorded this species from the Nansha Islands at the depths of 74 and 102 m.	en	Arzivian, Arteen, Alrubaie, Ahmad, Yang, Jessica, Lin, Huiyu, Zhang, Eva, Leong, Rupert (2022): Crabs (Crustacea, Decapoda) from the Seas of East and Southeast Asia Collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 4. South China Sea. Bulletin of the National Museum of Nature and Science. Series A, Zoology 48 (4): 147-191, DOI: 10.50826/bnmnszool.48.4_147, URL: http://dx.doi.org/10.1159/000539401
5F30F95FFFEF9026FD48FACDFEED5BFF.taxon	description	(Fig. 9 C – D) Material examined. RV Hakuhō Maru KH- 72 - 1 cruise, sta. 50, 1 Ə (CB 10.7 × CL 7.6 mm), NSMT-Cr 30920. Remarks. Chopra (1935) thoroughly explained the taxonomic status of Charybdis (Goniohellenus) hoplites var. vadorum Alcock, 1899, and as a result, regarded it as a full species with C. (G.) sinensis Gordon, 1930 as its synonym. Charybdis vadorum is a small species, reaching at most about 2.5 cm in the carapace breadth including epibranchial teeth. The carapace surface (Fig. 9 C) is wholly pilose, with granular patches on the regions; most characteristic is the somewhat Y-shaped, thick cardiac region; otherwise, it is noted on close examination that the metagastric transverse ridge and metagastric region are connected by a median longitudinal line of granules. The front (Fig. 9 C – D) is low and six-lobed; the median lobe is inclined laterally, the submedian lobe is subtruncated along the inner margin, with the lateral angle hardly attaining the level of the median lobe; the inner part of the submedian lobe slightly overlaps the median lobe; the lateral lobe is narrow and separated from the submedian lobe by a deep notch. The orbit is large, with the diameter wider than half the frontal length. As seen in Fig. 9 C – D, of the six anterolateral teeth of the carapace, the first (the external orbital tooth) is directed forward and slightly larger than the second and fourth teeth, but slightly smaller than the third and fourth teeth; the last (the epibranchial tooth) is narrow, much longer than the precedings and sharply directed laterally. This species was represented with excellent figures as C. (G.) vadorum by Chopra (1935), Leene (1938), and Wee and Ng (1995). Although Wee and Ng (1995) considered C. philippinensis Ward, 1941 as a doubtful synonym of this species, Padate et al. (2010) examined and redescribed the type specimen. Charybdis philippinensis is referable to the subgenus Charybdis, with the carapace surface naked and devoid of tomentum. Distribution. According to Wee and Ng (1995), this species ranges from Taiwan to the Red Sea through the East and Southeast Asian Sea, India and the Persian Gulf, from depths of 10 to 80 m.	en	Arzivian, Arteen, Alrubaie, Ahmad, Yang, Jessica, Lin, Huiyu, Zhang, Eva, Leong, Rupert (2022): Crabs (Crustacea, Decapoda) from the Seas of East and Southeast Asia Collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 4. South China Sea. Bulletin of the National Museum of Nature and Science. Series A, Zoology 48 (4): 147-191, DOI: 10.50826/bnmnszool.48.4_147, URL: http://dx.doi.org/10.1159/000539401
5F30F95FFFEA9026FF19FC65FBAD5A74.taxon	description	(Fig. 11 D) Material examined. RV Hakuhō Maru KH- 72 - 1 cruise, sta. 45, 1 8 (CB 25.3 mm with epibranchial tubercles, CL 14.5 mm), NSMT-Cr 30928. Remarks. The specimen at hand was first referred to the genus Xiphonectes A. Milne-Edwards, 1873, based on the presence of an acute tubercle at the lateral end of the carapace posterior margin, and identified as X. tuberculosus (A. Milne-Edwards, 1861) owing to the characteristic carapace tuberculation. This species was, however, recently transferred to a new genus Incultus by Koch et al. (2022) as the type species of the genus, together with two congeners, I. alcocki (Nobili, 1905) and I. brockii (De Man, 1887). In this specimen (Fig. 11 D), the lateral tubercles of the carapace posterior margin are stout and directed obliquely outward. This species is characteristic in the carapace ornamentation and armature: the dorsal surface is flattened overall, areolated with a distinct gastric region, a strong ridge running from the epibranchial tubercle toward the gastric region, and several obtuse tubercles arranged symmetrically around the cardiac region. The carapace anterolateral margin is armed with nine teeth including the obtuse external orbital angle and the strong epibranchial tubercle as usual, but the first, second, third and fifth teeth are distinctly smaller than the fourth, sixth and seventh teeth. This species was well characterized with the detailed descriptions and figures by the original author (A. Milne Edwards, 1861) and Crosnier (1962), Stephenson and Rees (1967), Dai and Yang (1991), Apel and Spiridonov (1998), and Koch et al. (2022). Distribution. Sandwich Islands; South China Sea off Hong Kong, 111 m depth; the Philippines, 66 – 81 m depth; Palau Islands, 30 m depth; Indonesia; Sri Lanka, 50 m depth; Persian Gulf; Madagascar, 25 – 65 m depth. Recently, Ng et al. (2019) recorded this species from southwestern coast of India at the depth of 200 m.	en	Arzivian, Arteen, Alrubaie, Ahmad, Yang, Jessica, Lin, Huiyu, Zhang, Eva, Leong, Rupert (2022): Crabs (Crustacea, Decapoda) from the Seas of East and Southeast Asia Collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 4. South China Sea. Bulletin of the National Museum of Nature and Science. Series A, Zoology 48 (4): 147-191, DOI: 10.50826/bnmnszool.48.4_147, URL: http://dx.doi.org/10.1159/000539401
5F30F95FFFEA9026FC83FCDDFB885F13.taxon	description	(Fig. 1 D) Material examined. RV Hakuhō Maru KH- 72 - 1 cruise, sta. 45, 1 8 (CB 9.8 × CL 8.7 mm), NSMT-Cr 30921. Remarks. Lissocarcinus arkati is characterised by having many transverse striae on the carapace dorsal surface (Fig. 1 D). The records of occurrence are few, but the characteristics are sufficiently illustrated by Kemp (1923), Chopra (1935), Sakai (1939), Crosnier (1962), Dai and Xu (1991), Dai and Yang (1991) and Ng et al. (2001). Distribution. Indo-West Pacific, ranging from Japan to Madagascar, with several intervening localities; 4 – 65 m depth. Gordon (1931) recorded this species from Hong Kong.	en	Arzivian, Arteen, Alrubaie, Ahmad, Yang, Jessica, Lin, Huiyu, Zhang, Eva, Leong, Rupert (2022): Crabs (Crustacea, Decapoda) from the Seas of East and Southeast Asia Collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 4. South China Sea. Bulletin of the National Museum of Nature and Science. Series A, Zoology 48 (4): 147-191, DOI: 10.50826/bnmnszool.48.4_147, URL: http://dx.doi.org/10.1159/000539401
5F30F95FFFEB9027FF0CFC5BFBC95F71.taxon	description	(Fig. 9 A – B) Material examined. RV Hakuhō Maru KH- 72 - 1 cruise, sta. 45, 1 Ə (CB 15.1 mm including lateral teeth, CL 10.5 mm including frontal teeth), NSMT-Cr 30923. Remarks. All the chelipeds and ambulatory legs of the male are missing, but the specimen is identifiable as either Lupocycloporus innominatus (Rathbun, 1909) or L. minutus (Shen, 1937), which are often considered as synonymous with, or specifically distinct from, each other. Stephenson (1967) first made a detailed description of Portunus minutus based on a female from the Gulf of Thailand to compensate for the brief original description by Shen (1937), but did not make a comparison with L. innominatus. Recently, Spiridonov (1999) recorded two females from Ambon as P. innominatus and stressed that the determination of the taxonomic status of both species will be possible after reexamination and re-description of the type series. Takeda and Komatsu (2020) recorded a male and an ovigerous female from the Okinawa-jima Island, the Ryukyu Islands, as P. (L.) innominatus, and also three males, a female and an ovigerous female from Taiwan were recorded as L. minutus by Huang and Shih (2021) who mentioned the differences in the shallow front lobes of L. minutus and the G 1 morphology. The shape of the front lobes seems to be similar in both species, and may be subject to individual variation; also, differences in the G 1 are not clear. The dis- tributions of both species are almost sympatric as noted by Takeda and Komatsu (2020). The present report follows them in considering both species to be synonymous. The following account of the carapace is based on the present male examined (Fig. 9 A). The carapace dorsal surface is wholly covered with a thick coat of short setae; each protogastric region is provided with two transverse rows of minute granules along the anterior and median parts, which are united at the lateral end; the protogastric regions of both sides are isolated by a longitudinal median row of minute granules that reaches to the mesogastric region; a patch of minute granules on each side of the metagastric region and the cardiac region is indicated by a pair of low, granulated mounds arranged side by side. The frontal margin is cut into four shallow lobes, the inner pair being about half as wide as the outer pair. The carapace anterolateral margin is gently curved, with nine serrated teeth that are subequal or becoming slightly larger posteriorly; the last (epibranchial tooth) is about two times longer than the eighth tooth, directed laterally and weakly curved anteriorly, with the sharp tip. The basal half of the G 1 is stout, and distal half tapers and strongly curved outwards (Fig. 9 B). Distribution. Lupocycloporus innominatus ranges from east coast of India to Indonesia, New Caledonia and the Ryukyu Islands, and L. minutus is recorded from almost the same area as L. innominatus, at depths of 6 – 33 m. The present male was obtained at 60 m depth.	en	Arzivian, Arteen, Alrubaie, Ahmad, Yang, Jessica, Lin, Huiyu, Zhang, Eva, Leong, Rupert (2022): Crabs (Crustacea, Decapoda) from the Seas of East and Southeast Asia Collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 4. South China Sea. Bulletin of the National Museum of Nature and Science. Series A, Zoology 48 (4): 147-191, DOI: 10.50826/bnmnszool.48.4_147, URL: http://dx.doi.org/10.1159/000539401
5F30F95FFFE89024FE90FE04FE425D26.taxon	description	(Fig. 11 F) Material examined. RV Hakuhō Maru KH- 72 - 1 cruise, sta. 45, 1 Ə infested by a Sacculina (CB 20.5 mm including lateral teeth, CL 17.0 mm including frontal teeth), NSMT-Cr 30924. Remarks. The female at hand (Fig. 11 F), infested by a Sacculina parasite, lacks all the chelipeds and ambulatory legs, but the carapace agrees well with the excellent figures of Lupocyclus philippinensis Semper, in Nauck, 1880, given by Leene (1940), Sakai (1939, 1976), Crosnier (1962), and Osmani et al. (2019). This species is rather well known (Ng et al., 2008). Distribution. Widely distributed in the Indo-West Pacific from Japan to the western Indian Ocean through the East and Southeast Asia, and Australia; 25 – 130 m depth.	en	Arzivian, Arteen, Alrubaie, Ahmad, Yang, Jessica, Lin, Huiyu, Zhang, Eva, Leong, Rupert (2022): Crabs (Crustacea, Decapoda) from the Seas of East and Southeast Asia Collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 4. South China Sea. Bulletin of the National Museum of Nature and Science. Series A, Zoology 48 (4): 147-191, DOI: 10.50826/bnmnszool.48.4_147, URL: http://dx.doi.org/10.1159/000539401
5F30F95FFFE89024FF51FB8EFBDE5B2E.taxon	description	(Fig. 1 C) Material examined. RV Hakuhō Maru KH- 72 - 1 cruise, sta. 45, 1 8 (CB 11.0 × CL 7.1 mm), NSMT-Cr 30925. Remarks. The female at hand (Fig. 1 C) is not fully mature, but is clearly identifiable with Podophthalmus nacreus, with the hexagonal carapace armed with a strong epibranchial tooth and the long eyestalk having a large cornea. Ng et al. (2017) transferred P. nacreus to the genus Vojmirophthalmus erected by Števčič (2011) to accommodate P. minabensis Sakai, 1961, but P. nacreus as well as P. vigil are generically distinct from V. minabensis by the unusually ornamented eyestalk of the latter species (Sakai (1961: pl. 4 fig. 1; 1976: fig. 207 a, pl. 135 fig. 1) and Huang and Shih (2021: fig. 3 E). Distribution. Indo-West Pacific from Japan through Taiwan, the Philippines, and Indonesia to New Caledonia in the Pacific, and from Western Australia through the Andaman Sea and the Gulf of Martaban to Madagascar, 30 – 126 m depth (Alcock, 1899 b; Leene, 1938, 1940; Sakai, 1939, 1965, 1976; Crosnier and Thomassin, 1974; Moosa, 1981; Hosie, 2012).	en	Arzivian, Arteen, Alrubaie, Ahmad, Yang, Jessica, Lin, Huiyu, Zhang, Eva, Leong, Rupert (2022): Crabs (Crustacea, Decapoda) from the Seas of East and Southeast Asia Collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 4. South China Sea. Bulletin of the National Museum of Nature and Science. Series A, Zoology 48 (4): 147-191, DOI: 10.50826/bnmnszool.48.4_147, URL: http://dx.doi.org/10.1159/000539401
5F30F95FFFE89025FC82FD55FDB65B2E.taxon	description	(Fig. 9 E – F) Material examined. RV Hakuhō Maru KH- 72 - 1 cruise, sta. 45, 1 ovig. 8 (CB 7.0 × CL 6.0 mm), NSMT-Cr 30926; 1 ovig. 8 (6.3 × 4.5 mm), NSMT-Cr 30927. Remarks. The specimens were identified as Thalamita sexlobata characterized by small body size, the pilose carapace surface, the large orbital fossae, and the presence of short transverse ridges and patch of granules behind the epibranchial ridge. The frontal margin is poorly developed, sinuous and seemingly four-lobed, with the outer lobe overlapping half of the inner lobe, as correctly figured by the original author (Miers, 1886, fig. 2 a) and Crosnier (1962, fig. 196). In the present specimens (Fig. 9 E – F), the anterior margin of the outer lobe is shallowly concave at the subterminal part and shallowly separated from the supraorbital angle. In this case, the frontal margin is indistinctly six-lobed rather than four-lobed, being somewhat different from the figures given by Crosnier (1962), Takeda (1989), and Wee and Ng (1995). This condition of the frontal lobes approaching to the six-lobed appearance was already mentioned by Stephenson and Hudson (1957). The carapace anterolateral margin is armed with five anteriorly-directed teeth including the external orbital tooth which is the largest of all; the second is slightly smaller than the first, but larger than the subequal third and fifth teeth; the fourth is apparently smaller than the others, but distinct in the specimens examined (Fig. 9 E). Distribution. Widely distributed in the Indo-West Pacific from the Hawaiian Islands and the northern Ryukyu Islands southwards to Queensland, Australia, and westwards to Madagascar and the Red Sea through the Andaman Islands and the Persian Gulf, 15 – 80 m in depth (Takeda 1989; Apel and Spiridonov, 1998).	en	Arzivian, Arteen, Alrubaie, Ahmad, Yang, Jessica, Lin, Huiyu, Zhang, Eva, Leong, Rupert (2022): Crabs (Crustacea, Decapoda) from the Seas of East and Southeast Asia Collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 4. South China Sea. Bulletin of the National Museum of Nature and Science. Series A, Zoology 48 (4): 147-191, DOI: 10.50826/bnmnszool.48.4_147, URL: http://dx.doi.org/10.1159/000539401
5F30F95FFFE99025FF2CFD88FB29581E.taxon	description	(Fig. 10 C – D) Material examined. RV Hakuhō Maru KH- 72 - 1 cruise, sta. 45, 1 Ə (CB 10.0 × CL 7.0 mm), NSMT-Cr 30929; 1 Ə (6.1 × 4.6 mm), 7 ovig. 88 (6.1 × 4.2 mm to 8.6 × 6.4 mm), 2 88 (6.4 × 4.8 mm; 8.0 × 6.1 mm), NSMT-Cr 30930. Remarks. Takeda and Komatsu (2017) figured the carapace and both chelae of Actumnus forficigerus, which have important features distinguishing this species from its congener. This species is close to A. squamosus (De Haan, 1835) in the general form of the carapace, as being densely covered with short velvety tomentum and sparsely with long silky setae, but differs in the carapace being wider and with the anterolateral margin armed with three stout distinct teeth behind the external orbital angle. Actumnus forficigerus differs remarkably from A. squamosus in having the usual tubercles on the outer surface of the cheliped palm instead of the thick pavement of the tubercles. In the smaller male and all of the females, both chelipeds are stout and covered with some rows of conical granules on the palm outer surfaces, with short and stout fingers, but in the larger male (Fig. 10 C – D), both fingers are rather elongated and weakly curved downwards against the palm. Distribution. Hitherto known only from Japanese waters, from Sagami Bay at the Pacific coast of central Honshu to Amami-Oshima Island in the northern Ryukyu Islands, 35 – 121 m depth (Sakai, 1939, 1965, 1976; Takeda and Miyake, 1969); now, from the southern South China Sea.	en	Arzivian, Arteen, Alrubaie, Ahmad, Yang, Jessica, Lin, Huiyu, Zhang, Eva, Leong, Rupert (2022): Crabs (Crustacea, Decapoda) from the Seas of East and Southeast Asia Collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 4. South China Sea. Bulletin of the National Museum of Nature and Science. Series A, Zoology 48 (4): 147-191, DOI: 10.50826/bnmnszool.48.4_147, URL: http://dx.doi.org/10.1159/000539401
5F30F95FFFE99025FD63FE45FC3B5C09.taxon	description	(Fig. 10 E – F) Material examined. RV Hakuhō Maru KH- 72 - 1 cruise, sta. 45, 1 Ə (CB 5.1 × CL 4.6 mm), NSMT-Cr 30931; 1 8 (7.7 × 5.5 mm), NSMT-Cr 30932. Remarks. The most remarkable feature of Actumnus squamosus is the pavement of tubercles covering the whole outer surface of the palm, as shown by A. Milne-Edwards (1865) and Sakai (1939). Takeda and Komatsu (2017) clearly figured this feature with a photograph of both chelae in comparing the new species, A. tsu- rukaii, with the related species. The present female (Fig. 10 E – F) agrees with the ovigerous female from the Sahul Shelf in having the carapace covered with a thick felt of short setae and in not being strongly convex dorsally, and also in having the characteristic chelipeds reported by Takeda et al. (2022 b). In the specimen examined, however, the tubercles covering the cheliped palm seem to be somewhat loosely spread all over the surface, showing the obtuse polygonal form of various sizes. Distribution. Indo-West Pacific from Japan to India, 10 – 324 m depth.	en	Arzivian, Arteen, Alrubaie, Ahmad, Yang, Jessica, Lin, Huiyu, Zhang, Eva, Leong, Rupert (2022): Crabs (Crustacea, Decapoda) from the Seas of East and Southeast Asia Collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 4. South China Sea. Bulletin of the National Museum of Nature and Science. Series A, Zoology 48 (4): 147-191, DOI: 10.50826/bnmnszool.48.4_147, URL: http://dx.doi.org/10.1159/000539401
5F30F95FFFE9902BFD67FAB6FD815E2C.taxon	description	(Fig. 11 A – B) Material examined. RV Hakuhō Maru KH- 72 - 1 cruise, sta. 45, 1 8 (CB 8.7 × CL 7.0 mm), NSMT-Cr 30933. Remarks. The female specimen examined agrees well with the descriptions and figures given by Gordon (1930, 1931, as Pilumnus), Ng and Tan (1984, as Bathypilumnus), Dai and Yang (1991, as Pilumnus), and Yang and Dai (1994, as Pilumnus). The carapace is robust in appearance, vaulted in both directions, with the ill-defined surface covered with stiff longish setae, the chelipeds are stout, and different in size but similar in the armature, with some rows of equidistant long, curved spiniform tubercles on both carpi and palms (Fig. 11 A – B). In the third part of this study (Takeda et al., 2022 b), Bathypilumnus pugilator (A. Milne-Edwards, 1973) was recorded from the Sahul Shelf, which is one of three congeneric species. In B. sinensis, the cheliped armature is characteristic, somewhat in the form of mushroom-shaped tubercles, and in all the species, the male abdomen is narrow, with the long and straight G 1 different from the so-called sigmoid Pilumnus - type. Distribution. Ng and Tan (1984) mentioned the distributional range from the Laccadive Archipelago to the Gulf of Thailand, Hong Kong, and the South China sea, bathymetrically from 21 to 115 m. Yang and Dai (1994) recorded a female from the Nansha Islands as Pilumnus.	en	Arzivian, Arteen, Alrubaie, Ahmad, Yang, Jessica, Lin, Huiyu, Zhang, Eva, Leong, Rupert (2022): Crabs (Crustacea, Decapoda) from the Seas of East and Southeast Asia Collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 4. South China Sea. Bulletin of the National Museum of Nature and Science. Series A, Zoology 48 (4): 147-191, DOI: 10.50826/bnmnszool.48.4_147, URL: http://dx.doi.org/10.1159/000539401
5F30F95FFFE79028FD64F988FBCA5C6F.taxon	description	(Fig. 12) Material examined. RV Hakuhō Maru KH- 72 - 1 cruise, sta. 42, 1 8 (CB 5.3 × CL 3.3 mm), NSMT-Cr 30934. Remarks. The genus Ceratoplax is comprised of nine species (Ng et al., 2008; Ng and Clark, 2015), each of which is superficially similar with smooth and glabrous carapace surfaces. Ng and Clark (2015) finely depicted the specific characters among C. truncatifrons Rathbun, 1914, C. fulgida Rathbun, 1914, and a new species named C. margarita, with many photographs and illustrations including the type specimens of Rathbun`s two species. According to Ng and Clark (2015), in C. marginata, the carapace posterolateral margin is more distinctly converging towards the carapace posterior margin, and the third maxilliped merus is auriculiform at its antero-external angle similar to that of C. fulgida, but different from the prominently elongated antero-external angle of the merus in C. truncata. It is impossible at present to apply the sex-associated characters with the female at hand, and the differences of the carapace shape may be subtle for the species identification as shown at the photographic angles (Fig. 12 A – B). In the present female, however, the antero-external corner of the third maxilliped merus is sharply developed as in the figure given by Ng and Clark (2015, fig. 4 C). Distribution. Ng and Clark (2015) overlooked Takeda (1989) who recorded this species with fine drawings from the Oshima Passage in Amami-Oshima Island, the northern Ryukyu Islands, at the depth of 45 m. This species is otherwise recorded from the type locality (Badian Island, off western Samar, the Philippines, 58 m depth) by Rathbun (1914), the Sula Islands (east of Sulawesi, Indonesia, 22 m depth) by Tesch (1918), and the Nansha Islands (South China Sea, 97 – 206 m depth) by Chen (1998 b). The specimens from Indonesia collected by the Siboga Expedition were re-examined and photographed by Ng and Clark (2015).	en	Arzivian, Arteen, Alrubaie, Ahmad, Yang, Jessica, Lin, Huiyu, Zhang, Eva, Leong, Rupert (2022): Crabs (Crustacea, Decapoda) from the Seas of East and Southeast Asia Collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 4. South China Sea. Bulletin of the National Museum of Nature and Science. Series A, Zoology 48 (4): 147-191, DOI: 10.50826/bnmnszool.48.4_147, URL: http://dx.doi.org/10.1159/000539401
5F30F95FFFE49029FD77FA95FE3B5B46.taxon	description	(Fig. 11 C) Material examined. RV Hakuhō Maru KH- 72 - 1 cruise, sta. 50, 1 Ə (CB 7.5 × CL 7.7 mm), NSMT-Cr 30935. Remarks. The male examined after ecdysis is wholly semitransparent due to decalcification (Fig. 11 C), agreeing well with the detailed description, photographs and figures by Chia and Ng (2000) who examined the type specimens and synonymized Gonatonotus crassimanus Haswell, 1880 and Eumedonus villosus Rathbun, 1918 with this species. The front may be somewhat variable individually in the depth and width of the median incision, with V-shaped notch in this specimen just like the holotype represented by Chia and Ng (2000: fig. 1). Distribution. Known from Japan, the East China Sea, the Philippines, and eastern and western Australia, 40 – 138 m depth.	en	Arzivian, Arteen, Alrubaie, Ahmad, Yang, Jessica, Lin, Huiyu, Zhang, Eva, Leong, Rupert (2022): Crabs (Crustacea, Decapoda) from the Seas of East and Southeast Asia Collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 4. South China Sea. Bulletin of the National Museum of Nature and Science. Series A, Zoology 48 (4): 147-191, DOI: 10.50826/bnmnszool.48.4_147, URL: http://dx.doi.org/10.1159/000539401
5F30F95FFFE59029FF50FDCDFDC55D5E.taxon	materials_examined	Material examined. RV Hakuhō Maru KH- 72 - 1 Cruise, sta. 45, 1 Ə (CB 5.2 × CL 5.1 mm), 1 8 (6.0 × 5.7 mm), NSMT-Cr 30936. Remarks. The present young specimens agree well with the specimens (3 ƏƏ, 1 8) from the Sahul Shelf recorded in the third part of this study (Takeda et al., 2002 b). Distribution. Chia and Ng (2000) recorded many specimens from Indonesia, Singapore, Thailand, and Australia, 9 – 54 m depth.	en	Arzivian, Arteen, Alrubaie, Ahmad, Yang, Jessica, Lin, Huiyu, Zhang, Eva, Leong, Rupert (2022): Crabs (Crustacea, Decapoda) from the Seas of East and Southeast Asia Collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 4. South China Sea. Bulletin of the National Museum of Nature and Science. Series A, Zoology 48 (4): 147-191, DOI: 10.50826/bnmnszool.48.4_147, URL: http://dx.doi.org/10.1159/000539401
5F30F95FFFE59029FF47FBC5FB6E59C4.taxon	description	(Fig. 10 A – B) Material examined. RV Hakuhō Maru KH- 72 - 1 cruise, sta. 45, 1 Ə (CB 8.0 × CL 6.1 mm), NSMT-Cr 30937, 1 8 (7.5 × 5.1 mm) infested by a Sacculina, NSMT-Cr 30938. Remarks. The male at hand (Fig. 10 A – B) agrees well with the specimens from the Sahul Shelf recorded by Takeda et al. (2002 b) in the setation of the carapace, chelipeds and ambulatory legs with stiff setae of various sizes and the spination of the carapace anterolateral margins and chelipeds. Distribution. Widely distributed in the whole Indo-West Pacific at depths of 5 to 100 m.	en	Arzivian, Arteen, Alrubaie, Ahmad, Yang, Jessica, Lin, Huiyu, Zhang, Eva, Leong, Rupert (2022): Crabs (Crustacea, Decapoda) from the Seas of East and Southeast Asia Collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 4. South China Sea. Bulletin of the National Museum of Nature and Science. Series A, Zoology 48 (4): 147-191, DOI: 10.50826/bnmnszool.48.4_147, URL: http://dx.doi.org/10.1159/000539401
5F30F95FFFE59029FD6FFE43FC415AA9.taxon	materials_examined	Material examined. RV Hakuhō Maru KH- 72 - 1 cruise, sta. 45, 2 ƏƏ (CB 5.3 × CL 4.0 mm; 6.7 × 4.8 mm with left gill chamber deformed by bopyrid parasite), 1 8 (5.5 × 4.1 mm), 2 juv., NSMT-Cr 30939. Remarks. The present specimens agree well with those (2 ƏƏ, 6 ovig. 88) from the Sahul Shelf recorded in the third part of this study (Takeda et al., 2022 b). Distribution. Known from the Philippines, South China Sea, Singapore, and Indonesia, 12 – 78 m in depth. Galil and Takeda (1988) recorded three females obtained from mangrove sponges in Singapore.	en	Arzivian, Arteen, Alrubaie, Ahmad, Yang, Jessica, Lin, Huiyu, Zhang, Eva, Leong, Rupert (2022): Crabs (Crustacea, Decapoda) from the Seas of East and Southeast Asia Collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 4. South China Sea. Bulletin of the National Museum of Nature and Science. Series A, Zoology 48 (4): 147-191, DOI: 10.50826/bnmnszool.48.4_147, URL: http://dx.doi.org/10.1159/000539401
5F30F95FFFE5902FFD40FB33FE5C5894.taxon	description	(Fig. 13) Material examined. RV Hakuhō Maru KH- 72 - 1 cruise, sta. 42, 1 Ə (CB 5.6 × CL 4.7 mm), NSMT-Cr 30940. Remarks. The male at hand is safely identified as Xestopilumnus cultripollex, which is known only by the original description (Ng and Dai, 1997) based on five males and one subadult female from the vicinity of Hong Kong Island. All known specimens, including the present male, are small and not distinctive in the general appearance of the carapace, chelipeds and ambulatory legs, but the wholly smooth carapace, chelipeds and ambulatory legs is of distinct generic value in the genus Glabropilumnus Balss, 1933 and genera such as Gorgonariana, Lentilumnus, Serenolumnus, Xlumnus separated from Glabropilumnus by Galil and Takeda (1988). The carapace dorsal surface (Fig. 13 A – B) is smooth and glabrous without areolar indication, and evenly convex in both directions; the front prominently extends forward, with transverse margin and without median incision, slightly more than one-third as wide as carapace; the carapace anterolateral margin (Fig. 13 A – B) is narrowly edged and cut into three lobes; first lobe is isolated from the less prominent external orbital angle with a short distance, and fringed with minute granules; the second lobe is small, but tooth-like and subacute at the tip, separated from the first lobe by a small but distinct notch; the third lobe obscurely isolated from the second lobe is obscurely convex and weakly convergent toward the carapace posterolateral corner. Both chelipeds (Fig. 13 D – E) are different in size and shape; in the right, or larger cheliped in this male, the palm is stout, fairly bulged, but the left cheliped palm is compressed, with sharp lower margin. The ambulatory legs are all slender (Fig. 13 A, F). A small discrepancy is the absence of the fourth anterolateral lobe of the carapace, but the variation of the armature was shown with fine illustrations in the original description. Distribution. Hitherto known from Hong Kong, and now from the southern South China Sea; subtidal to 36 m depth.	en	Arzivian, Arteen, Alrubaie, Ahmad, Yang, Jessica, Lin, Huiyu, Zhang, Eva, Leong, Rupert (2022): Crabs (Crustacea, Decapoda) from the Seas of East and Southeast Asia Collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 4. South China Sea. Bulletin of the National Museum of Nature and Science. Series A, Zoology 48 (4): 147-191, DOI: 10.50826/bnmnszool.48.4_147, URL: http://dx.doi.org/10.1159/000539401
5F30F95FFFE3902FFF36FD50FDA65FCE.taxon	description	(Fig. 14 G – H) Material examined. RV Hakuhō Maru KH- 72 - 1 Cruise, sta. 42, 1 8 (CB 5.5 × CL 4.8 mm), NSMT-Cr 30941. Remarks. The genus Camatopsis was revised by Ng and Castro (2016) who referred the forma A and B of the Siboga specimens defined by Tesch (1918) to one known species C. rubida, and three new species C. minor, C. leptomeru s and Microtopsis teschi, and furthermore, amended the records of C. rubida from Japan and Taiwan (Yokoya, 1933; Sakai, 1939, 1965, 1976; Takeda, 1973 b; Fang, 1991; Hsueh and Huang, 2002; Komai et al., 2012) to those of a new species C. thula. Camatopsis thula is most characteristic in having the shortest and stoutest ambulatory meri among its congeners. The specimen at hand (Fig. 14 G – H) is female, so that the Gl is not available for definite identification, but the ambulatory legs are slenderer than those of congeners, with the overall appearance agreeing well with the photographs of C. leptomerus given by Ng and Castro (2016) who examined numerous specimens from many localities. Distribution. The type locality is the Bohol Sea in the Philippines, 627 – 645 m in depth, and the other localities recorded by Ng and Castro (2016) are in Taiwan (310 – 346 m), the Philippines (10 – 1,260 m), Malaysia, Indonesia (216 m), Papua New Guinea (80 – 447 m), Vanuatu (83 – 394 m), and New Caledonia (200 – 700 m).	en	Arzivian, Arteen, Alrubaie, Ahmad, Yang, Jessica, Lin, Huiyu, Zhang, Eva, Leong, Rupert (2022): Crabs (Crustacea, Decapoda) from the Seas of East and Southeast Asia Collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 4. South China Sea. Bulletin of the National Museum of Nature and Science. Series A, Zoology 48 (4): 147-191, DOI: 10.50826/bnmnszool.48.4_147, URL: http://dx.doi.org/10.1159/000539401
5F30F95FFFE3902DFD58FFFBFDCD587C.taxon	description	(Fig. 14 A – F) Material examined. RV Hakuhō Maru KH- 72 - 1 cruise, sta. 52, 1 Ə (CB 10.1 × CL 8.9 mm), NSMT-Cr 30942. Remarks. The carapace (Fig. 14 A – B) of the present specimen of Camatopsis rubida is subtrapezoidal and high, differing slightly in the pro- portional shape according to the photographing angles. The carapace dorsal surface, chelipeds and ambulatory legs are wholly covered with longish fine setae (Fig. 14 A – F). In the male examined, the right cheliped (Fig. 14 E) is larger, but slender, leaving a narrow gape between both fingers along their basal halves. In the small left cheliped (Fig. 14 F), the palm and fingers are slenderer, being armed with small irregular teeth on both cutting edges and with a sharp spine at distal one third of immovable cutting edge. Both cheliped fingers are apparently different from the chelipeds of developing or mature males. The known species of the genus Camatopsis are morphologically close to each other, with some degree of variation, and it is difficult to evaluate some records of C. rubida. According to Ng and Castro (2016) who illustrated the G 1 s of all the Camatopsis species, the records of C. rubida from the Gulf of Thailand by Rathbun (1910) and Indonesia by Serène (1964) still remain unclear, but the records of C. rubida from the South China Sea by Zarenkov (1972) and Chen (1998 b) show clearly the correct identification, with the illustration of the G 1. Recently, Mendoza et al. (2021) recorded C. rubida from Indonesian waters, with fine photographs. Ng and Castro (2016) recorded numerous specimens from many localities, with the complete literature, many photographs and the detailed geographical information. Distribution. The type locality is the Andaman Sea (349 m), and Ng and Castro (2016) recorded many specimens from the Andaman Sea coast of Thailand (36 – 53 m), Western Australia (101 – 407 m), Sumatra (179 – 660 m), Papua New Guinea (195 – 422 m), Queensland coast of Australia (56 – 695 m), New Caledonia (170 – 460 m), and Fiji (234 – 361 m). Chen`s record (1998 b) from the Nansha Islands is based on specimens obtained from depths of 113 to 224 m.	en	Arzivian, Arteen, Alrubaie, Ahmad, Yang, Jessica, Lin, Huiyu, Zhang, Eva, Leong, Rupert (2022): Crabs (Crustacea, Decapoda) from the Seas of East and Southeast Asia Collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 4. South China Sea. Bulletin of the National Museum of Nature and Science. Series A, Zoology 48 (4): 147-191, DOI: 10.50826/bnmnszool.48.4_147, URL: http://dx.doi.org/10.1159/000539401
5F30F95FFFE1902DFF54FD3BFD9E5F3E.taxon	description	(Fig. 11 E) Material examined. RV Hakuhō Maru KH- 72 - 1 cruise, sta. 42, 1 Ə (CB 7.8 × CL 6.6 mm), NSMT-Cr 30943. Remarks. The genus Retropluma was extensively studied by De Saint Laurent (1989) on six species then known, and later a new species was added by McLay (2006 a). The most remarkable characteristic common to seven known species is that the last pair of the thoracic legs is extremely reduced. Even if all of the species are rather rare, R. denticulata is well known by the records from the West Pacific by Yokoya (1933), Sakai (1934, 1939, 1976), Zarenkov (1968), De Saint Laurent (1989), Chen and Xu (1991), Dai and Yang (1991), and Takeda (2001). In R. denticulata (Fig. 11 E), the rostrum is triangular, with a wide basal part; both of two transverse carinae of the carapace dorsal surface are wholly distinct between the carapace lateral margins of both sides. McLay (2006 a) included the new species, R. laurentae, which was preoccupied by a fossil crab and renamed R. solomonensis by McLay (2006 b), in the key to six species prepared by De Saint Laurent (1989). Distribution. Japan, South China Sea, Philippines, and Solomon Islands; 70 – 286 m depth.	en	Arzivian, Arteen, Alrubaie, Ahmad, Yang, Jessica, Lin, Huiyu, Zhang, Eva, Leong, Rupert (2022): Crabs (Crustacea, Decapoda) from the Seas of East and Southeast Asia Collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 4. South China Sea. Bulletin of the National Museum of Nature and Science. Series A, Zoology 48 (4): 147-191, DOI: 10.50826/bnmnszool.48.4_147, URL: http://dx.doi.org/10.1159/000539401
5F30F95FFFDA9016FD56FBCDFC0C5C86.taxon	materials_examined	Material examined. RV Hakuhō Maru KH- 73 - 2 cruise, sta. 36 (Shelf off Swatow, 22 ° 08.3 ′ N, 117 ° 44.6 ′ E, 80 m depth), 3 m beam trawl, 10 March, 1973, 1 juv. 8 (CB 3.4 × PCL 5.9 mm), NSMT-Cr 30944.	en	Arzivian, Arteen, Alrubaie, Ahmad, Yang, Jessica, Lin, Huiyu, Zhang, Eva, Leong, Rupert (2022): Crabs (Crustacea, Decapoda) from the Seas of East and Southeast Asia Collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 4. South China Sea. Bulletin of the National Museum of Nature and Science. Series A, Zoology 48 (4): 147-191, DOI: 10.50826/bnmnszool.48.4_147, URL: http://dx.doi.org/10.1159/000539401
5F30F95FFFDA9016FD56FAD6FBD55FCF.taxon	materials_examined	Material examined. RV Hakuhō Maru KH- 73 - 2 cruise, sta. 41 (southeast of Hong Kong, 22 ° 15.3 ′ N, 115 ° 28.2 ′ E to 22 ° 15.7 ′ N, 115 ° 28.9 ′ E, 55 m depth), 3 m beam trawl, 18 March, 1973, 2 ƏƏ (CB 15.0 × CL 28.2 mm; 18.0 × 28.2 mm), NSMT-Cr 30945.	en	Arzivian, Arteen, Alrubaie, Ahmad, Yang, Jessica, Lin, Huiyu, Zhang, Eva, Leong, Rupert (2022): Crabs (Crustacea, Decapoda) from the Seas of East and Southeast Asia Collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 4. South China Sea. Bulletin of the National Museum of Nature and Science. Series A, Zoology 48 (4): 147-191, DOI: 10.50826/bnmnszool.48.4_147, URL: http://dx.doi.org/10.1159/000539401
5F30F95FFFDB9017FD56FE15FB8B5B0C.taxon	materials_examined	Material examined. RV Hakuhō Maru KH- 73 - 2 cruise, sta. 36 (Shelf off Swatow, 22 ° 08.3 ′ N, 117 ° 44.6 ′ E, 80 m depth), 3 m beam trawl, 10 March, 1973, 2 ƏƏ (CB 8.5 × CL 6.5 mm; 9.8 × 7.1 mm), 2 ovig. 88 (6.9 × 5.6 mm; 10.3 × 7.6 mm), NSMT-Cr 30949.	en	Arzivian, Arteen, Alrubaie, Ahmad, Yang, Jessica, Lin, Huiyu, Zhang, Eva, Leong, Rupert (2022): Crabs (Crustacea, Decapoda) from the Seas of East and Southeast Asia Collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 4. South China Sea. Bulletin of the National Museum of Nature and Science. Series A, Zoology 48 (4): 147-191, DOI: 10.50826/bnmnszool.48.4_147, URL: http://dx.doi.org/10.1159/000539401
5F30F95FFFDB9017FF19FB88FD895CA1.taxon	materials_examined	Material examined. RV Hakuhō Maru KH- 73 - 2 cruise, sta. 44 (southeast of Hong Kong, 21 ° 41.2 ′ N, 117 ° 31.1 ′ E to 22 ° 42.7 ′ N, 117 ° 33.4 ′ E, 415 – 437 m depth), 3 m beam trawl, 19 March, 1973, 1 ovig. 8 (CB 39.0 excluding brachial tubercles × PCL 29.5 mm), NSMT-Cr 30947.	en	Arzivian, Arteen, Alrubaie, Ahmad, Yang, Jessica, Lin, Huiyu, Zhang, Eva, Leong, Rupert (2022): Crabs (Crustacea, Decapoda) from the Seas of East and Southeast Asia Collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 4. South China Sea. Bulletin of the National Museum of Nature and Science. Series A, Zoology 48 (4): 147-191, DOI: 10.50826/bnmnszool.48.4_147, URL: http://dx.doi.org/10.1159/000539401
5F30F95FFFDB9017FF19FAF0FC9059C4.taxon	materials_examined	Material examined. RV Hakuhō Maru KH- 73 - 2 cruise, sta. 41 (southeast off Hong Kong; 22 ° 15.3 ′ N, 115 ° 28.2 ′ E to 22 ° 15.7 ′ N, 115 ° 28.9 ′ E, 55 m depth), 3 m beam trawl, 18 March, 1973, 8 ƏƏ (CB 36.2 mm including epibranchial tubercles × 20.5 mm to 39.0 × 21.1 mm), 8 88 (34.2 × 18.5 mm to 41.0 × 23.1 mm), 5 ovig. 88 (34.4 × 18.5 mm to 42.4 × 23.7 mm), NSMT-Cr 30948.	en	Arzivian, Arteen, Alrubaie, Ahmad, Yang, Jessica, Lin, Huiyu, Zhang, Eva, Leong, Rupert (2022): Crabs (Crustacea, Decapoda) from the Seas of East and Southeast Asia Collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 4. South China Sea. Bulletin of the National Museum of Nature and Science. Series A, Zoology 48 (4): 147-191, DOI: 10.50826/bnmnszool.48.4_147, URL: http://dx.doi.org/10.1159/000539401
5F30F95FFFDB9017FF19FFD3FDD45D79.taxon	materials_examined	Material examined. RV Hakuhō Maru KH- 73 - 2 cruise, sta. 41 (southeast of Hong Kong, 22 ° 15.3 ′ N, 115 ° 28.2 ′ E to 22 ° 15.7 ′ N, 115 ° 28.9 ′ E, 55 m depth), 3 m beam trawl, 18 March, 1973, 1 Ə (CB 10.5 × CL 11.0 mm), NSMT-Cr 30946. Remarks. The male at hand has been preserved in 75 % ethanol for almost 50 years, but brick red specks are clearly preserved on the carapace and chelipeds. The dispersed pattern of the specks of good and variable sizes agrees with the colored figure given by Adams and White (1849: pl. 12 fig. 2) and the line drawing by Ovaere (1987: fig. 1 A), differing from the pattern of U. parahaematosticta Galil, 2005, in which the smaller spots are crowded at the carapace submedian, anterolateral and posterolateral parts, without larger spots on the chelipeds. The color pattern of this species was also cleary shown by Poore et al. (2008). Low et al. (2020) noted that the original description of this species should be referred to Adams (1847) who wrote the brief notes on the coloration of Leucosia haematosticta prior to the publication of Adams and White (1849).	en	Arzivian, Arteen, Alrubaie, Ahmad, Yang, Jessica, Lin, Huiyu, Zhang, Eva, Leong, Rupert (2022): Crabs (Crustacea, Decapoda) from the Seas of East and Southeast Asia Collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 4. South China Sea. Bulletin of the National Museum of Nature and Science. Series A, Zoology 48 (4): 147-191, DOI: 10.50826/bnmnszool.48.4_147, URL: http://dx.doi.org/10.1159/000539401
