identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
5E7F7A4AC750FF9A43ADBD8660FBF82D.text	5E7F7A4AC750FF9A43ADBD8660FBF82D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atheta (Dimetrota) Mulsant & Rey 1873	<div><p>Subgenus Dimetrota Mulsant &amp; Rey, 1873</p><p>Dimetrota Mulsant &amp; Rey, 1873: 165 (type species: Homalota tristicula Mulsant &amp; Rey, 1873 = Homalota cadaverina Brisout de Barneville, 1860).</p><p>Coproceramius Gistel, 1857: 9 (type species: Bolitochara impressifrons Mannerheim, 1830 = Aleochara atramentaria Gyllenhal, 1810).</p><p>Engamota Casey, 1910: 151 (type species: Acrotona absona Casey, 1910).</p><p>Dimetrotina Casey, 1911: 143 (type species: Dimetrotina vaniuscula Casey, 1911).</p><p>Diagnosis. Members of the subgenus Dimetrota can be distinguished from other subgenera of Atheta by the combination of the following characters: lateral regions of body and tibiae with conspicuously long and strong setae; infraorbital carina complete; ligula divided into two lobes in basal half; pronotum with rather narrow and asperate punctation, with conspicuous and long setae in lateral margins, pubescence directed anteriorly in midline; mesocoxal cavities narrowly separated; elytra with asperate punctation; at suture subequal to length of pronotum at middle; abdomen with sparse punctation to apex; abdominal tergite II with 2 macrosetae on each side of midline, III with anterior macrosetae; macrochaetal arrangement of VII 34 (Benick &amp; Lohse, 1974; Yosii &amp; Sawada, 1976; Lohse et al., 1990).</p><p>Key to Korean species of Atheta subgenus Dimetrota</p><p>1. Body length less than 2.0 mm........................................................... A. (Dimetrota) furtiva</p><p>- Body length more than 2.0 mm.......................................................................... 2</p><p>2. Body length about 2.5 mm; posterior margin of abdominal tergite VIII with pointed processes......... Atheta (D.) nigripes</p><p>- Body length more than 2.5 mm; posterior margin of abdominal tergite VIII without processes or with blunt processes...... 3</p><p>3. Antennomere 3 about as long as or slightly longer than 2..................................................... 4</p><p>- Antennomere 3 slightly shorter than 2..................................................................... 9</p><p>4. Antennomere 3 slightly longer than 2; posterior margin of abdominal tergite VIII not modified; posterior margin of male abdominal tergite VIII broadly round; posterior margin of female abdominal tergite VIII subtruncate... A. (D.) photaechonica</p><p>- Antennomere 3 about as long as 2; posterior margin of abdominal tergite VIII modified, not fitting above description...... 5</p><p>5. Antennomeres 4–5 slightly elongate........................................................ A. (D.) aeneipennis</p><p>- Antennomeres 4–5 quadrate to subquadrate................................................................ 6</p><p>6. Antennomeres 7–10 transverse; posterior margin of male abdominal tergite VIII with two blunt processes in lateral regions, crenate in median region.................................................................... A. (D.) altaica</p><p>- Antennomeres 7–10 subquadrate to slightly transverse; posterior margin male abdominal tergite VIII with broad process... 7</p><p>7. Posterior margin of male abdominal tergite VIII crenate......................................... A. (D.) subsericans</p><p>- Posterior margin of male abdominal tergite VIII emarginate in median region and pointed in postero-lateral regions....... 8</p><p>8. Abdominal sternite VIII with 8 macrosetae in male (Fig. 2C), and 7 macrosetae in female (Fig. 2E) on each side of midline................................................................................... A. (D.) machonryongica</p><p>- Abdominal sternite VIII with 10 macrosetae in male (Fig. 5D), and 8 macrosetae in female (Fig. 5F) on each side of midline................................................................................... A. (D.) ovata sp. nov.</p><p>9. Body yellowish brown; antennomere 4 slightly transverse........................................... A. (D.) weisei</p><p>- Body dark brown to black; antennomere 4 about as long as wide............................................... 10</p><p>10. Posterior margin of male abdominal tergite VIII with two pointed processes in lateral regions, emarginate in median region; abdominal sternite VIII with 7 macrosetae in male, and 6 macrosetae in female on each side of midline.................................................................................................... A. (D.) atramentaria</p><p>- Posterior margin of male abdominal tergite VIII with two blunt outer processes in lateral regions and 7–9 blunt inner processes in median region (Fig. 7B); abdominal sternite VIII with 10 macrosetae in male (Fig. 7C), and 7 macrosetae in female (Fig. 7E) on each side of midline................................................................... A. (D.) yamamotoi</p></div>	https://treatment.plazi.org/id/5E7F7A4AC750FF9A43ADBD8660FBF82D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lee, Seung-Gyu;Ahn, Kee-Jeong	Lee, Seung-Gyu, Ahn, Kee-Jeong (2022): Korean species of the Atheta Thomson subgenus Dimetrota Mulsant & Rey (Coleoptera: Staphylinidae: Aleocharinae) with a description of new species. Zootaxa 5138 (4): 401-416, DOI: 10.11646/zootaxa.5138.4.3
5E7F7A4AC753FF9943ADBCA76585FE36.text	5E7F7A4AC753FF9943ADBCA76585FE36.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atheta (Dimetrota) aeneipennis (Thomson 1856)	<div><p>Atheta (Dimetrota) aeneipennis (Thomson, 1856)</p><p>Remarks. Lohse and Smetana (1985) resurrected Atheta (Dimetrota) aeneipennis from the synonymy of A. (D.) picipennis (Mannerheim) . Paśnik (2001) recorded Atheta (Dimetrota) picipennis from North Korea. Later, Smetana (2004) replaced the record of A. (D.) picipennis with A. (D.) aeneipennis . After examining all North Korean specimens of Atheta (Dimetrota) picipennis, we found that the record by Paśnik (2001) is the result of a misidentification of Atheta (Dimetrota) aeneipennis, in agreement with Smetana (2004). This species has not been collected in South Korea.</p></div>	https://treatment.plazi.org/id/5E7F7A4AC753FF9943ADBCA76585FE36	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lee, Seung-Gyu;Ahn, Kee-Jeong	Lee, Seung-Gyu, Ahn, Kee-Jeong (2022): Korean species of the Atheta Thomson subgenus Dimetrota Mulsant & Rey (Coleoptera: Staphylinidae: Aleocharinae) with a description of new species. Zootaxa 5138 (4): 401-416, DOI: 10.11646/zootaxa.5138.4.3
5E7F7A4AC753FF9943ADBDCF644AFD32.text	5E7F7A4AC753FF9943ADBDCF644AFD32.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atheta (Dimetrota) altaica Bernhauer 1901	<div><p>Atheta (Dimetrota) altaica Bernhauer, 1901</p><p>(Fig. 8A)</p><p>Remarks. This species was recorded by Paśnik (2001) from North Korea and no specimen has been collected in South Korea.</p></div>	https://treatment.plazi.org/id/5E7F7A4AC753FF9943ADBDCF644AFD32	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lee, Seung-Gyu;Ahn, Kee-Jeong	Lee, Seung-Gyu, Ahn, Kee-Jeong (2022): Korean species of the Atheta Thomson subgenus Dimetrota Mulsant & Rey (Coleoptera: Staphylinidae: Aleocharinae) with a description of new species. Zootaxa 5138 (4): 401-416, DOI: 10.11646/zootaxa.5138.4.3
5E7F7A4AC753FF9943ADBECB64DEFBAD.text	5E7F7A4AC753FF9943ADBECB64DEFBAD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atheta (Dimetrota) atramentaria (Gyllenhal 1810)	<div><p>Atheta (Dimetrota) atramentaria (Gyllenhal, 1810)</p><p>(Fig. 8B–C)</p><p>Remarks. After detailed comparative studies of Atheta (Dimetrota) atramentaria with its related species, we found that this species is in agreement with the male diagnostic characters including aedeagus of A. (Bessobia) genitilior described by Pace (1998). Therefore, the species may be synonymized under A. (D.) atramentaria . But it is still premature to propose a synonym until thorough comparative and detailed study of the species including type specimens. This species was recorded by Kim et al. (1994) for the first time in Korea and some specimens were collected from mushroom in forest.</p></div>	https://treatment.plazi.org/id/5E7F7A4AC753FF9943ADBECB64DEFBAD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lee, Seung-Gyu;Ahn, Kee-Jeong	Lee, Seung-Gyu, Ahn, Kee-Jeong (2022): Korean species of the Atheta Thomson subgenus Dimetrota Mulsant & Rey (Coleoptera: Staphylinidae: Aleocharinae) with a description of new species. Zootaxa 5138 (4): 401-416, DOI: 10.11646/zootaxa.5138.4.3
5E7F7A4AC753FF9943ADB8B7644AFAAA.text	5E7F7A4AC753FF9943ADB8B7644AFAAA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atheta (Dimetrota) furtiva Cameron 1939	<div><p>Atheta (Dimetrota) furtiva Cameron, 1939</p><p>(Fig. 8D)</p><p>Remarks. This species was recorded by Paśnik (2001) from North Korea and no specimen has been collected in South Korea.</p></div>	https://treatment.plazi.org/id/5E7F7A4AC753FF9943ADB8B7644AFAAA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lee, Seung-Gyu;Ahn, Kee-Jeong	Lee, Seung-Gyu, Ahn, Kee-Jeong (2022): Korean species of the Atheta Thomson subgenus Dimetrota Mulsant & Rey (Coleoptera: Staphylinidae: Aleocharinae) with a description of new species. Zootaxa 5138 (4): 401-416, DOI: 10.11646/zootaxa.5138.4.3
5E7F7A4AC753FF9E43ADB9B36160FC8D.text	5E7F7A4AC753FF9E43ADB9B36160FC8D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atheta (Dimetrota) machonryongica Pasnik 2001	<div><p>Atheta (Dimetrota) machonryongica Paśnik, 2001</p><p>(Figs. 1A, 2A–I, 8E)</p><p>Atheta (Dimetrota) machonryongica Paśnik, 2001: 222; Smetana, 2004: 382; Schülke &amp; Smetana, 2015: 532.</p><p>Description. Length 2.5–3.2 mm. Body (Fig. 1A) dark brown to black; head, pronotum and abdomen almost black; elytra and legs paler than other parts. Head. Slightly transverse, approximately 1.1–1.2 times as wide as long, widest across eyes, slightly narrower than pronotum; eyes large and prominent, about 1.5 times as long as temples; gular sutures moderately separated, diverged basally; cervical carina complete. Antennae (Fig. 2A) long and slender; antennomeres 1–3 elongate, 1 longest, 2 about as long as 3, 4–10 quadrate to subquadrate, 11 about as long as 1, about as long as preceding two combined. Mouthparts. Labrum with 9–10 macrosetae on each side of midline; epipharynx with α-sensillum long and setaceous, about 3.0 times as long as ε-sensillum; β- and γ-sensilla short. Mandibles asymmetrical, subtriangular, decurved and pointed apically, about 1.5–1.6 times as long as basal width; very few small denticles present in molar region; right one with small internal tooth, internal margin slightly serrulate; prostheca developed, composed of three portions, second portion slightly longer. Galea and lacinia of maxilla long and slender; maxillary palpus elongate and pubescent; palpomere 1 smallest, 2 about 2.5–2.7 times as long as wide, 3 slightly longer than 2, about 2.6–2.8 times as long as wide, 4 digitiform, filamentous sensilla not reaching to basal half. Prementum with two medial setae narrowly separated; two basal pores close together; several medial pseudopores, lateral pseudopores, 1 setal pore and 2 real pores present on each side of midline; labial palpomere 1 largest, about 1.5–1.7 times as long as wide, γ-setula contiguous with b-seta, 2 shortest, about 1.4–1.6 times as long as wide, 3 more or less dilated apically and about as long as 1, about 3.0–3.5 times as long as wide. Mentum trapezoidal, anterior margin slightly emarginate; v-seta relatively long, close to u-seta. Thorax. Pronotum approximately 1.3–1.4 times as wide as long, widest in apical third to half. Metanotal scutum with 1 long seta and about 2–10 relatively short setae on each side of midline. Mesoventral process more or less pointed at apex, distinctly longer than isthmus and metaventral process combined. Elytra slightly longer and wider than pronotum; elytron approximately 1.6–1.7 times as long as wide; hind wings fully developed, flabellum composed of about 6–8 long setose lobes. Legs. Length ratio of tarsomeres 25:29:32:78 (protarsus); 28:36:37:33:78 (mesotarsus); 44:48:47:42:85 (metatarsus). Abdomen. Surface glossy and densely pubescent, with transverse microsculpture; macrochaetal arrangement of tergites II–VI 02-13-13 (or 23)-13 (or 23)-13 (or 23); male sternite VII with moderately many granular punctures in posterior margin; male tergite VIII (Fig. 2B) with 4 macrosetae on each side of midline, posterior margin with broad process, emarginate in median region and pointed in postero-lateral regions; male sternite VIII (Fig. 2C) with 8 macrosetae on each side of midline, posterior margin convex, with inconspicuous marginal setae; posterior margin of female tergite VIII (Fig. 2D) truncate in median region; female sternite VIII (Fig. 2E) with 7 macrosetae on each side of midline, posterior margin truncate, with conspicuous and moderately long marginal setae, robust minute setae present in median region. Aedeagus. Median lobe (Figs. 2F–G) narrowly ovate, apical process pointed apically in ventral aspect. Apical lobe of paramerites (Fig. 2H) subparallel-sided, with 4 setae; a-seta longest, b-seta slightly longer than c- and d-setae. Spermatheca. Bursa with large umbilicus; duct recurved, deflected at apex (Fig. 2I).</p><p>Type material. Holotype, ♂, labeled as in Fig. 8E.</p><p>Material examined. SOUTH KOREA: Chungbuk Prov.: 1 ex., Danyang-gun, Mt. Sobaeksan, Cheongdong, 7–9 v 1999, US Hwang, HJ Kim, FIT . Chungnam Prov.: 5 exx., Buyeo-gun, Naesan-myeon, Mt. Wolmyeongsan, Geumjisa, 3 v–1 vi 2000, US Hwang, HJ Kim, FIT; 5 exx., Nonsan-si, Beolgok-myeon, Mt. Daedunsan, Surak-valley, 2–10 V 2000, US Hwang, SJ Park, HJ Kim, FIT. Gangwon Prov.: 1 ex., Hwacheon-gun Mt. Hwaaksan, 11–12 ix 1998, KL You, KJ Ahn, mushroom; 5 exx., Inje-gun, Sangnam-myeon, Bangtaesan, 18 viii 2000, MH Kim, mushroom; 4 exx., Pyeongchang-gun, Jinbu-myeon, Dongsan-ri, Mt. Odaesan, Sangwonsa, 30 iv–4 vi 2001, KJ Ahn, SJ Park, MS Kim, MJ Jeon, FIT; 12 exx., same data as former except for ‘ 4–22 vi 2001 ’; 6 exx., same data as former except for ‘ 22 vi–16 viii 2001, SJ Park, CW Shin ’; 1 ex., same data as former except for ‘ 16 viii–15 ix 2001 ’; 2 ex., same data as former except for ‘ 18 vi–22 vii 2004, SJ Park, KM Yang, DH Lee, FIT’; 4 exx., same data as former except for ‘ 18 vi 2004, SJ Park, FIT’; 1 ex., same data as former except for ‘ 30 viii 2006, YH Kim’; 2 exx., same data as former except for ‘Woljeongsa, 22 viii–20 x 2000, KJ Ahn’; 1 ex., same data as former except for ‘Bukdaesa, 23 viii 2000, MH Kim, mushroom’; 4 exx., Pyeongchang-gun, Mt. Odaesan, Jeokmyeolbogung, 7–9 x 1999, US Hwang, MS Kim, FIT; 2 exx., Pyeongchang-gun, Cheondong-ri, Mt. Sambangsan, 13 vii–15viii 2001, KJ Ahn, SJ Park, SW Shin, FIT in Pinus forest; 6 exx., Inje-gun, Mt. Jeombongsan, Gombaeryeong, 23–30 viii 1999, US Hwang, bait trap. Gyeongnam Prov.: 6 exx., Habcheon-gun, Haeinsa, 4 x 1986; Gyeonggi Prov.: 2 exx., Seoulsi, Jungnang-gu, Mt. Yongmasan, 1 vi 1986, YS Kim; 1 ex., Yongmunsan, Jogaegol, 31 v 1986. Jeonnam Prov.: 2 exx., Gurye-gun, Mt. Jirisan, Piagol, 24 v 2000, HJ Kim, MJ Jeon, sifting’ .</p><p>Distribution. Korea (South, North).</p><p>Remarks. This species was described by Paśnik (2001) from North Korea and is recorded here for the first time in South Korea, and some specimens were collected from mushroom in forest.</p></div>	https://treatment.plazi.org/id/5E7F7A4AC753FF9E43ADB9B36160FC8D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lee, Seung-Gyu;Ahn, Kee-Jeong	Lee, Seung-Gyu, Ahn, Kee-Jeong (2022): Korean species of the Atheta Thomson subgenus Dimetrota Mulsant & Rey (Coleoptera: Staphylinidae: Aleocharinae) with a description of new species. Zootaxa 5138 (4): 401-416, DOI: 10.11646/zootaxa.5138.4.3
5E7F7A4AC754FF9E43ADBF6B644AFC7B.text	5E7F7A4AC754FF9E43ADBF6B644AFC7B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atheta (Dimetrota) nigripes (Thomson 1856)	<div><p>Atheta (Dimetrota) nigripes (Thomson, 1856)</p><p>Remarks. This species was recorded by Paśnik (2001) from North Korea and no specimen has been collected from South Korea.</p></div>	https://treatment.plazi.org/id/5E7F7A4AC754FF9E43ADBF6B644AFC7B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lee, Seung-Gyu;Ahn, Kee-Jeong	Lee, Seung-Gyu, Ahn, Kee-Jeong (2022): Korean species of the Atheta Thomson subgenus Dimetrota Mulsant & Rey (Coleoptera: Staphylinidae: Aleocharinae) with a description of new species. Zootaxa 5138 (4): 401-416, DOI: 10.11646/zootaxa.5138.4.3
5E7F7A4AC754FF9143ADB80360F1FE6D.text	5E7F7A4AC754FF9143ADB80360F1FE6D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atheta (Dimetrota) ovata Lee & Ahn 2022	<div><p>Atheta (Dimetrota) ovata Lee &amp; Ahn, sp. nov.</p><p>(Figs. 1B, 3A–F, 4A–H, 5A–G, 6A–D)</p><p>Description. Length 2.5–3.2 mm. Body (Fig. 1B) surface glossy, densely pubescent with microsculpture. Body dark brown to black; antennae, elytra and legs paler than other parts, brownish. Head. Slightly transverse (Fig. 4A), approximately 1.1–1.2 times as wide as long, widest across eyes, slightly narrower than pronotum; eyes large and prominent, about 1.5–1.6 times as long as temples; gular sutures moderately separated, more or less diverged basally; cervical carina complete. Antennae (Fig. 4B) long and slender; antennomeres 1–3 elongate, 1 longest, 2 about as long as 3, 4–10 quadrate to subquadrate, 11 about as long as 1, about as long as preceding two combined. Mouthparts. Labrum (Fig. 3A) with 10–11 macrosetae on each side of midline; epipharynx (Fig. 3B) with α-sensillum long and setaceous, about 3.0 times as long as ε-sensillum; β- and γ-sensilla short. Mandibles (Figs. 3C–D) asymmetrical, subtriangular, decurved and pointed apically, about 1.5–1.6 times as long as basal width; very few small denticles present in molar region; right one (Fig. 3C) with small internal tooth, internal margin slightly serrulate; prostheca developed, composed of three portions, second portion slightly longer. Galea and lacinia of maxilla (Fig. 3E) long and slender; maxillary palpus elongate and pubescent; palpomere 1 smallest, 2 about 2.6–2.8 times as long as wide, 3 slightly longer than 2, about 2.5–2.7 times as long as wide, 4 digitiform, filamentous sensilla not reaching to basal half. Prementum (Fig. 3F) with two medial setae very narrowly separated; two basal pores contiguous, less than 1.0 times width of basal pore; several medial pseudopores, lateral pseudopores, 1 setal pore and 2 real pores present on each side of midline; labial palpomere 1 largest, about 1.5–1.6 times as long as wide, γ-setula contiguous with b-seta, 2 shortest, about 1.6–1.8 times as long as wide, 3 more or less dilated apically and slightly shorter than 1, about 2.5–3.0 times as long as wide. Mentum (Fig. 3F) trapezoidal, anterior margin slightly emarginate; v-seta relatively long, close to u-seta. Thorax. Pronotum (Fig. 4C) approximately 1.3–1.4 times as wide as long, widest in apical third to half. Prosternum as in Fig. 4D. Metanotal scutum (Fig. 4E) with 1 long seta and about 3–4 short setae on each side of midline. Mesoventral process (Fig. 4F) distinctly pointed at apex, longer than isthmus and metaventral process combined. Scutellum as in Fig. 4G. Elytra slightly longer and wider than pronotum; elytron (Fig. 4H) approximately 1.6 times as long as wide; hind wings fully developed, flabellum (Fig. 4E) composed of about 6–7 long setose lobes. Legs. Length ratio of tarsomeres 22:26:30:76 (protarsus); 30:35:38:35:68 (mesotarsus); 45:44:43:46:96 (metatarsus). Abdomen. Surface glossy and densely pubescent, with transverse and reticulate microsculpture (Fig. 5C); macrochaetal arrangement of tergites II–VI 02-13 (or 23)-23-23-23; male sternites III–VI with many small pores, VII with several small pores in anterior region; male tergite VIII (Fig. 5A) with 4 macrosetae on each side of midline, posterior margin (Fig. 5B) with broad process, slightly emarginate in median region and slightly angled in postero-lateral margins; male sternite VIII (Fig. 5D) with 10 macrosetae on each side of midline, posterior margin with inconspicuous marginal setae; posterior margin of female tergite VIII (Fig. 5E) truncate in median region; female sternite VIII (Fig. 5F) with 8 macrosetae, posterior margin (Fig. 5G) broadly rounded, with conspicuous and long marginal setae, minute setae present in median region. Aedeagus. Median lobe (Figs. 6A–B) narrowly ovate, apical process subtriangular and convergent at apex, apex slightly swollen and globular in ventral aspect. Apical lobe of paramerites (Fig. 6C) subparallel-sided, with 4 setae; b-seta longest, distinctly longer than other setae short and subequal in length, c- and d-setae close together. Spermatheca. Bursa elongate, with slender umbilicus; duct recurved, deflected at apex (Fig. 6D).</p><p>Type material. Holotype, ♂, labeled as follows: ‘ KOREA: Gangwon Prov., Pyeongchang-gun, Jinbu-myeon, Dongsan-ri, Mt. Odaesan, Sangwonsa, 22 VI–16 VIII 2001, S.-J. Park, C.-W. Shin, ex FIT’ . Paratypes, 9 exx., same data as holotype.</p><p>Material examined. SOUTH KOREA: Chungbuk Prov.: 3 exx., Buyeo-gun, Oesan-myeon, Gaedeok-ri, Mt. Wolmyeongsan, 1 vi 2000, US Hwang, HJ Kim, sifting; 1 ex., Danyang-gun, Mt. Sobaeksan, Cheongdong, 7–9 v 1999, US Hwang, HJ Kim, sifting. Chungnam Prov.: 1 ex., Gongju-si, Banpo-myeon, Sangsin-ri, Mt. Gyeryongsan, 21 v 2000, MS Kim, near stream. Gangwon Prov.: 1 ex., Hongcheon-gun, Naechon-myeon, Mt. Baekamsan, Garyeong fall, 25 v–20 vi 2002, KJ Ahn, SJ Park, JS Park, FIT; 2 exx., Hongcheon-gun, Nae-myeon, Mt. Gyebangsan, Unduryeong, N37° 42.49.9′ E128° 26.40.3′, 1100 m, 11 v 2007, TK Kim, YH Kim, fungus on log; 5 exx., Injegun, Mt. Jeombongsan, Gombaeryeong, 23–30 viii 1999, US Hwang, bait trap; 1 ex., Jeongseon-gun, Gohan-eup, Mt. Hambaeksan, 13 vii 1999, US Hwang, mushroom; 2 exx., Mt. Seoraksan, 23 viii 1996, T. Pierre, mushroom; 21 exx., Pyeongchang-gun, Cheondong-ri, Mt. Sambangsan, 13 vii–15 viii 2001, KJ Ahn, SJ Park, CW Shin, FIT in Pinus forest; 67 exx., Pyeongchang-gun, Jinbu-myeon, Dongsan-ri, Mt. Odaesan, Sangwonsa, 30 iv–4 vi 2001, KJ Ahn, SJ Park, MS Kim, MJ Jeon, FIT; 161 exx., same data as former except for ‘ 4 vi–22 vi 2001 ’; 9 exx., same data as former except for ‘ 18 viii 2000, MH Kim, entirely rotten mushroom ( Boletaceae)’; 5 exx., same data as former except for ‘ 22 viii 2000, KJ Ahn, JH Ahn’; 152 exx., same data as former except for ‘ 22 vi–16 viii 2001, SJ Park, CW Shin, FIT’; 15 exx., same data as former except for ‘ 16 viii–15 ix 2001 ’; 5 exx., same data as former except for ‘ 15 ix–14 xi 2001, KJ Ahn, CW Shin, FIT’; 9 exx., same data as former except for ‘ 21 iv–18 v 2002, SJ Park, CW Shin, FIT’; 17 exx., same data as former except for ‘ 18 v–23 vi 2002, SJ Park, JS Park, FIT’; 2 exx., same data as former except for ‘ 23 vi 2002, SJ Park, JS Park, mushroom’; 2 exx., same data as former except for ‘ 13 vii 2004, SM Choi, mushroom’; 6 exx., same data as former except for ‘ 18 vi 2004, SJ Park, FIT’; 27 exx., same data as former except for ‘ 18 vi–22 vii 2004, SJ Park, KM Yang, DH Lee, FIT’; 2 exx., same data as former except for ‘37°47′8.3″ E128°33′54.0″ 880 m, 10 ix 2009, TK Kim, YH Kim, leaf litter’; 17 exx., same data as former except for ‘ N37°47′3.4″ E128°33′44.6″ 930 m, 12 VI 2012, YH Kim, SG Lee, YG Ban, JC Lim, mushroom’; 2 exx., same data as former except for ‘Bukdaesa, 23 viii 2000, MH Kim, mushroom’; 2 exx., same data as former except for ‘Namdae jijangam, 12 ix 2007, HW Kim, YH Kim, mushroom’; 7 exx., Pyeongchang-gun, Jinbu-myeon, Mt. Odaesan, Woljeongsa, 22 viii–20 x 2000, KJ Ahn, FIT; 4 exx., Pyeongchang-gun, Mt. Odaesan, Jeokmyeolbogung, 7–9 vii 1998, KL You, HJ Lim, FIT; 1 ex., Pyeongchang-gun, Bangrim-myeon, Ungyo 2-ri, Mt. Baekdeoksan, 12 vii–16 viii 2001, KJ Ahn, SJ Park, CW Shin, FIT; 1 ex., Taebaek-si, Mt. Taebaeksan, Baekdansa, 16 vii 1999, US Hwang, HJ Kim, sifting; 1 ex., Yangyang-gun, Seo-myeon, Osaek-ri, Hangyeryeong, 16 viii 2000, MH Lim, mushroom; 3 exx., Yangyang-gun, Seo-myeon, Osaek-ri, Mt. Seoraksan, Osaekyaksu, 31 vii–15 ix 2002, SJ Park, CW Shin, JS Park, FIT; 3 exx., same data as former except for ‘Osaekyaksu, 20 vii 2004, SJ Park, KM Yang, KJ Ahn, mushroom’; 2 exx., Yeongwol-gun, Suju-myeon, Mt. Baekdeoksan, Gwaneumsa, 13 vii–15 viii 2001, KJ Ahn, SJ Park, CW Shin, FIT. 2 exx., Yeongwol-gun, Yeongwol-eup, Mt. Taehwasan, 14 viii 2001, MH Kim, mushroom ( Boletaceae). Jeonbuk Prov.: 3 exx., Jeongeup-si, Mt. Naejangsan, Naejangsa, Geumseon valley, 15–24 vi 2000, US Hwang, HJ Kim, FIT.</p><p>Distribution. Korea (South).</p><p>Remarks. This species is very similar to Atheta (Dimetrota) machonryongica, but can be distinguished by the characters provided in the key and the different shape and structure of aedeagus and spermatheca. Most specimens were collected by FIT and from mushroom in forest.</p><p>Etymology. Named from Latin ovata meaning “ovate”, which refers to the shape of median lobe of aedeagus.</p></div>	https://treatment.plazi.org/id/5E7F7A4AC754FF9143ADB80360F1FE6D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lee, Seung-Gyu;Ahn, Kee-Jeong	Lee, Seung-Gyu, Ahn, Kee-Jeong (2022): Korean species of the Atheta Thomson subgenus Dimetrota Mulsant & Rey (Coleoptera: Staphylinidae: Aleocharinae) with a description of new species. Zootaxa 5138 (4): 401-416, DOI: 10.11646/zootaxa.5138.4.3
5E7F7A4AC75BFF9143ADBE76644AFD69.text	5E7F7A4AC75BFF9143ADBE76644AFD69.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atheta (Dimetrota) photaechonica Pasnik 2001	<div><p>Atheta (Dimetrota) photaechonica Paśnik, 2001</p><p>(Fig. 8F)</p><p>Remarks. This species was described by Paśnik (2001) from North Korea and no specimen has been collected in South Korea.</p></div>	https://treatment.plazi.org/id/5E7F7A4AC75BFF9143ADBE76644AFD69	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lee, Seung-Gyu;Ahn, Kee-Jeong	Lee, Seung-Gyu, Ahn, Kee-Jeong (2022): Korean species of the Atheta Thomson subgenus Dimetrota Mulsant & Rey (Coleoptera: Staphylinidae: Aleocharinae) with a description of new species. Zootaxa 5138 (4): 401-416, DOI: 10.11646/zootaxa.5138.4.3
5E7F7A4AC75BFF9143ADBF72644AFC75.text	5E7F7A4AC75BFF9143ADBF72644AFC75.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atheta (Dimetrota) subsericans Cameron 1939	<div><p>Atheta (Dimetrota) subsericans Cameron, 1939</p><p>(Fig. 8G)</p><p>Remarks. This species was recorded by Paśnik (2001) from North Korea and no specimen has been collected in South Korea.</p></div>	https://treatment.plazi.org/id/5E7F7A4AC75BFF9143ADBF72644AFC75	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lee, Seung-Gyu;Ahn, Kee-Jeong	Lee, Seung-Gyu, Ahn, Kee-Jeong (2022): Korean species of the Atheta Thomson subgenus Dimetrota Mulsant & Rey (Coleoptera: Staphylinidae: Aleocharinae) with a description of new species. Zootaxa 5138 (4): 401-416, DOI: 10.11646/zootaxa.5138.4.3
5E7F7A4AC75BFF9143ADB80E67C5FA7D.text	5E7F7A4AC75BFF9143ADB80E67C5FA7D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atheta (Dimetrota) weisei Bernhauer 1907	<div><p>Atheta (Dimetrota) weisei Bernhauer, 1907</p><p>(Fig. 8H)</p><p>Remarks. Atheta (Dimetrota) weisei was recorded by Paśnik (2001) for the first time in North Korea and by Cho et al. (2002) for the first time in South Korea. But, a habitus photograph of A. (D.) weisei provided by the paper of Cho et al. is incorrect identification of Gyrophaena species.</p><p>While studying Korean Athetini, we found that this species is similar to Atheta (Tetropla) nigritula Gravenhorst (1802) . After studying the syntype of A. (D.) weisei and the lectotype of A. (T.) nigritula, we suspect that the character difference between these two species is insufficient to separate as different species. However, a more thorough comparative study including minute characters of specimens from broader distributional areas should be conducted to propose a synonym, A. (D.) weisei under A. (T.) nigritula . Almost Korean specimens were found on mushrooms and some were collected from FIT or bait trap.</p></div>	https://treatment.plazi.org/id/5E7F7A4AC75BFF9143ADB80E67C5FA7D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lee, Seung-Gyu;Ahn, Kee-Jeong	Lee, Seung-Gyu, Ahn, Kee-Jeong (2022): Korean species of the Atheta Thomson subgenus Dimetrota Mulsant & Rey (Coleoptera: Staphylinidae: Aleocharinae) with a description of new species. Zootaxa 5138 (4): 401-416, DOI: 10.11646/zootaxa.5138.4.3
5E7F7A4AC75BFF9043ADBA0661D3FA05.text	5E7F7A4AC75BFF9043ADBA0661D3FA05.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atheta (Dimetrota) yamamotoi Sawada 1989	<div><p>Atheta (Dimetrota) yamamotoi Sawada, 1989</p><p>(Figs. 1C, 7A–I)</p><p>Atheta (Dimetrota) yamamotoi Sawada, 1989: 292; Smetana, 2004: 376; Schülke &amp; Smetana, 2015: 534.</p><p>Description. Length 2.7–3.5 mm. Body (Fig. 1C) dark brown to black; head almost black; elytra brown; basal segments of antennae and legs yellowish brown. Head. Subquadrate, approximately 1.0–1.1 times as wide as long, widest across eyes, slightly narrower than pronotum; eyes large and prominent, about 1.5 times as long as temples; gular sutures moderately separated, diverged basally; cervical carina complete. Antennae (Fig. 7A) long and slender; antennomeres 1–3 elongate, 1 longest, 2 slightly longer than 3, 4–5 slightly elongate, 6–10 quadrate to subquadrate, 11 longer than wide, slightly shorter than the preceding two combined. Mouthparts. Labrum transverse, slightly emarginate in anterior margin, with ε-sensillum and 9–10 macrosetae on each side of midline; epipharynx with α-sensillum long and setaceous, more than 3.0 times as long as ε-sensillum; β- and γ-sensilla short, blunt at apex. Mandibles asymmetrical, subtriangular, decurved and pointed apically, about 1.6 times as long as basal width; moderately very few small denticles present in molar region; right one with small internal tooth, internal margin slightly serrulate; prostheca developed, composed of three portions. Lacinia of maxilla moderately long; maxillary palpus elongate and pubescent; palpomere 1 smallest, 2 about 2.7–2.8 times as long as wide, 3 slightly longer than 2, about 2.7–2.9 times as long as wide, 4 digitiform, filamentous sensilla almost reaching to basal half. Prementum with two medial setae narrowly separated; two basal pores close together, 1.0–2.0 times of basal width; several medial pseudopores, lateral pseudopores, 1 setal pore and 2 real pores present on each side of midline; labial palpomere 1 largest, about 1.4–1.6 times as long as wide, γ-setula contiguous with b-seta, 2 shortest, about 1.2–1.4 times as long as wide, 3 more or less dilated apically and about as long as 1, about 3.0 times as long as wide. Mentum trapezoidal, anterior margin slightly emarginate; v-seta closer to u-seta than w-seta. Thorax. Pronotum approximately 1.3–1.4 times as wide as long, widest at middle. Metanotal scutum with 1 long seta and about 5–8 relatively short setae on each side of midline. Mesoventral process slightly pointed at apex, longer than isthmus and metaventral process combined; length ratio of mesoventral process, isthmus and metaventral process 12:5:3. Elytra slightly longer and wider than pronotum; elytron approximately 1.6–1.7 times as long as wide; hind wings fully developed, flabellum composed of about 7–10 long setose lobes. Legs. Length ratio of tarsomeres 29:34:37:90 (protarsus); 32:41:43:41:88 (mesotarsus); 49:54:51:51:111 (metatarsus). Abdomen. Surface glossy and densely pubescent, with transversely reticulate microsculpture; macrochaetal arrangement of tergites II–VI 02-13-13 (or 23)-13 (or 23)-13 (or 23); male tergite VIII (Fig. 7B) with 4 macrosetae on each side of midline, posterior margin with two blunt outer processes in postero-lateral margins and about 7–9 blunt inner processes, outer processes slightly longer than inner processes; male sternite VII with numerous granular punctures in posterior region; male sternite VIII (Fig. 7C) with about 10 macrosetae on each side of midline, posterior margin convex, with inconspicuous marginal setae; posterior margin of female tergite VIII (Fig. 7D) truncate, slightly emarginate in median region; female sternite VIII (Fig. 7E) with 7 macrosetae on each side of midline, posterior margin rounded, with conspicuous marginal setae, minute setae present in median region. Aedeagus. Median lobe (Figs. 7F–G) ovate, basal region of apical process constricted in ventral aspect. Apical lobe of paramerites (Fig. 7H) elongate, parallel-sided, with 4 setae; a-seta longest, the other setae subequal in length. Spermatheca. S-shaped; bursa elongate, with relatively large umbilicus (Fig. 7I).</p><p>Material examined. SOUTH KOREA: Chungbuk Prov.: 8 exx. (two on slide), Yeongdong-gun, Sangchonmyeon, Mulhan-ri, Mt. Minjujisan, N36˚03'35.22" E127˚52'31.26", 518 m, 13 v 2010, SG Lee, decaying persimmon . Chungnam Prov.: 3 exx. (one on slide), Daejeon-si, Mt. Gyeryongsan, Sutonggol, 21 iv 1998, KJ Ahn, HJ Kim, HJ Lim, KL Yu, bait trap . Jeonbuk Prov.: 2 exx., Jinan-gun, Mt. Unjangsan, 18 v 1995, YB Cho . Jeonnam Prov.: Suncheon-si, Seonamsa, 1 v 1988, JC Paik .</p><p>Distribution. Korea (South) and Japan.</p><p>Remarks. Atheta (Dimetrota) yamamotoi is very similar to A. (Atheta) transfuga, but can be distinguished by the male sternite VIII (Fig. 7C) with about 10 macrosetae on each side of midline; female sternite VIII (Fig. 7E) with 7 macrosetae on each side of midline. This species is recorded here for the first time in Korea.</p></div>	https://treatment.plazi.org/id/5E7F7A4AC75BFF9043ADBA0661D3FA05	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lee, Seung-Gyu;Ahn, Kee-Jeong	Lee, Seung-Gyu, Ahn, Kee-Jeong (2022): Korean species of the Atheta Thomson subgenus Dimetrota Mulsant & Rey (Coleoptera: Staphylinidae: Aleocharinae) with a description of new species. Zootaxa 5138 (4): 401-416, DOI: 10.11646/zootaxa.5138.4.3
