taxonID	type	description	language	source
9A2E144DDC53560E8C1AD97BF089175A.taxon	description	[Japanese name: Tabu-gunbai] Figs 2 B, 4 B, 6 A, 7 B, 9 B, 11 B, 13 B, 15 B, 17 B, 19 B, 21 B, 23 B, 25 B, 27 B, 29 B, 31 B, 34 A, B, 40 D-G	en	Souma, Jun (2022): Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan. Deutsche Entomologische Zeitschrift 69 (2): 219-281, DOI: http://dx.doi.org/10.3897/dez.69.89864, URL: http://dx.doi.org/10.3897/dez.69.89864
9A2E144DDC53560E8C1AD97BF089175A.taxon	materials_examined	Material examined. Holotype of Stephanitis (Norba) vitrea Takeya, 1931 (1 ♀, ELKU) (Fig. 34 A), JAPAN: Ryukyu Islands (northern part): " Yakushima Onoaida-Ambo " [= Yakushima Island, between Onoaida and Anbo (approximate coordinates: 30 ° 16 ' 08.3 " N, 130 ° 36 ' 44.2 " E)], 3. viii. 1929, leg. H. Hori. The labels shown in Fig. 34 A were created after (T. Mita, pers. comm. 2021), but the condition of the specimen matched the photograph of the original description (Takeya 1931). Therefore, the female individual seems to correspond to the holotype of S. (N.) vitrea. Suspected syntype of S. (Stephanitis) propinqua Horvath, 1912 (1 ♀, ELHU) (Fig. 34 B), JAPAN: Kyushu: " カゴシマ " [= Kagoshima-ken (approximate coordinates: 31 ° 35 ' 59.5 " N, 130 ° 32 ' 59.6 " E)], " 7 / 10 " [= 10. vii]. This specimen was labelled as " Stephanitis gratiosa Horv. " (apparently an unpublished name) and " カゴシマグンバイ " (unknown Japanese name). According to the original description (Horvath 1912), the syntype (s) of S. (S.) propinqua was (were) deposited in ELHU and this female individual is the only specimen in Matsumura's collection of ELHU matching the label data of syntype (s) of S. (S.) propinqua. However, the morphological characteristics of the specimen match the original description of S. (N.) aperta (Horvath 1912), but not that of S. (S.) propinqua. Conversely, the general habitus of S. (S.) propinqua recorded from Korea by previous studies (Takeya 1932, 1963) differs from that of all the East Asian species of Stephanitis (present study). Therefore, if there is no syntype of S. (S.) propinqua in other institutions, then S. (S.) propinqua should be synonymised with S. (N.) aperta and a new name should be given to " S. (S.) propinqua " distributed in Korea. Non-types (534 ♂♂ 777 ♀♀ 16 nymphs), JAPAN: Honshu: Ibaraki-ken, Higashiibaraki-gun, Oarai-machi, Isohama-cho, 12. ix. 2020, leg. J. Souma (7 ♂♂ 13 ♀♀ 3 nymphs, TUA); Tochigi Pref., Uchino, Watarase-yusuichi, 26. xi. 2016, leg. S. Maehara (4 ♂♂ 2 ♀♀, TUA); as above but 2. x. 2020 (4 ♂♂ 3 ♀♀, TUA); Chiba-ken, Tateyama-shi, Ohto (approximate coordinates: 34 ° 58 ' 19.8 " N, 139 ° 52 ' 33.9 " E), 22. v. 2021, leg. J. Souma (1 ♀, TUA); Chiba-ken, Tateyama-shi, Shimosanagura, 22. v. 2021, leg. J. Souma (1 ♂ 2 ♀♀, TUA); Chiba-ken, Tateyama-shi, Tateyama, 22. v. 2021, leg. J. Souma (2 ♂♂ 6 ♀♀, TUA); as above but 23. v. 2021 (4 ♀♀, ELKU; 2 ♂♂ 2 ♀♀ 1 nymph, TUA); Chiba-ken, Tateyama-shi, Sunozaki, 23. v. 2021, leg. J. Souma (11 ♂♂ 32 ♀♀, TUA); Chiba-ken, Tateyama-shi, Sunozaki Shrine, 23. v. 2021, leg. J. Souma (1 ♂ 2 ♀♀, TUA); Chiba-ken, Tateyama-shi, Masaki, 23. v. 2021, leg. J. Souma (7 ♂♂ 14 ♀♀, TUA); Chiba-ken, Tateyama-shi, Higashinagata, 24. v. 2021, leg. J. Souma (1 ♂ 1 ♀, TUA); Chiba-ken, Tateyama-shi, Yamogi (approximate coordinates: 34 ° 58 ' 08.7 " N, 139 ° 53 ' 34.4 " E), 24. v. 2021, leg. J. Souma (2 ♂♂ 4 ♀♀, TUA); Chiba-ken, Minamiboso-shi, Chikura-cho, Minamiasai, 24. v. 2021, leg. J. Souma (1 ♀, TUA); Chiba-ken, Kamogawa-shi, Yomogi, 25. v. 2021, leg. J. Souma (1 ♀, TUA); Tokyo Met., Imperial Palace, 5. vi. 1996, leg. A. Saito (7 ♂♂ 7 ♀♀, NSMT); as above but 16. x. 1996, leg. M. Tomokuni (8 ♂♂ 15 ♀♀, NSMT); 19. xi. 1997, leg. M. Tomokuni (9 ♂♂ 6 ♀♀, NSMT); as above but 25. v. 1998 (2 ♀♀, NSMT); as above but 28. v. 1998, leg. M. Tomokuni (4 ♂♂ 3 ♀♀, NSMT); as above but 16. vii. 2009, leg. M. Tomokuni (1 ♀, NSMT); as above but 28. v. 2012, leg. M. Tomokuni (1 ♀, NSMT); as above but 23. vii. 2012, leg. M. Tomokuni (1 ♂ 2 ♀♀, NSMT); as above but 10. ix. 2012, leg. M. Tomokuni (1 ♀, NSMT); Tokyo-to, Minato-ku, Shiba-koen, 8. vii. 2022, leg. J. Souma (4 ♂♂ 1 ♀, TUA); Kanagawa-ken, Sagamihara-shi, Minami-ku, Shimomizo, 15. xi. 2021, leg. J. Souma (9 ♂♂ 8 ♀♀ 3 nymphs, TUA); Kanagawa-ken, Sagamihara-shi, Chuo-ku, Tanashioda, 19. v. 2019, leg. J. Souma (2 ♀♀, TUA); Kanagawa-ken, Sagamihara-shi, Chuo-ku, Tana, 19. v. 2019, leg. J. Souma (22 ♂♂ 16 ♀♀, TUA); as above but 17. xi. 2021 (9 ♂♂ 11 ♀♀, TUA); Kanagawa-ken, Sagamihara-shi, Midori-ku, Oshima, 17. xi. 2021, leg. J. Souma (1 ♂ 3 ♀♀ 1 nymph, TUA); Kanagawa-ken, Zama-shi, Iriyanishi, 1. i. 2022, leg. J. Souma (1 ♂ 14 ♀♀, TUA); Kanagawa-ken, Ebina-shi, Kamiimaizumi, 1. i. 2022, leg. J. Souma (1 ♂ 1 ♀, TUA); Kanagawa-ken, Atsugi-shi, Funako, 29. v. 2017, leg. J. Souma (37 ♂♂ 41 ♀♀, TUA); as above but 30. v. 2017 (32 ♂♂ 30 ♀♀, TUA); as above but 31. v. 2017 (4 ♀♀, ELKU; 52 ♂♂ 55 ♀♀, TUA); as above but 2. vi. 2017 (6 ♂♂ 16 ♀♀, TUA); as above but 5. vi. 2017 (30 ♂♂ 17 ♀♀, TUA); Kanagawa-ken, Atsugi-shi, Nanasawa, 6. vi. 2017, leg. J. Souma (1 ♀, TUA); Kanagawa-ken, Aiko-gun, Aikawa-machi, Nakatsu, 26. v. 2021, leg. J. Souma (3 ♂♂ 11 ♀♀, TUA); as above but 15. xi. 2021 (3 ♀♀, TUA); Kanagawa-ken, Yokohama-shi, Isogo-ku, Negishihachiman Shrine, 31. v. 1999, leg. M. Takakuwa (1 ♀, KPMNH); Kanagawa-ken, Yokohama-shi, Kanazawa-ku, Noukendaimori, 15. vi. 2017, leg. J. Souma (3 ♂♂, TUA); Kanagawa-ken, Yokosuka-shi, Kamoi, 27. vi. 2017, leg. J. Souma (4 ♀♀, TUA); Kanagawa-ken, Chigasaki-shi, Yanagishima, 1. vi. 2019, leg. J. Souma (10 ♂♂ 7 ♀♀, TUA); Kanagawa-ken, Hadano-shi, Tsurumaki, Mt. Azuma, 19. xi. 2021, leg. J. Souma (2 ♂♂ 5 ♀♀, TUA); Kanagawa-ken, Hadano-shi, Soya, Mt. Kobo, 19. xi. 2021, leg. J. Souma (4 ♀♀, TUA); Kanagawa-ken, Ashigarashimo-gun, Manazuru-machi, Manazuru, 23. v. 2021, leg. J. Souma (11 ♂♂ 30 ♀♀, TUA); Niigata-ken, Niigata-shi, Nishi-ku, Igarashi- 2 no-cho, 19. vii. 2015, leg. K. Nakano (2 ♀♀, TUA); as above but 16. vi. 2021, leg. G. Mashima (12 ♂♂ 14 ♀♀, TUA); Niigata-ken, Niigata-shi, Chuo-ku, Sekiya, Nishikaigan Park, 22. x. 2016, leg. K. Nakano (4 ♂♂ 10 ♀♀, TUA); Niigata-ken, Niigata-shi, Chuo-ku, Seigoro, Toyanogata Park, 30. x. 2016, leg. K. Nakano (1 ♂ 4 ♀♀, TUA); Niigata-ken, Niigata-shi, Higashi-ku, Matsuzono, 21. vii. 2019, leg. K. Nakano (2 ♂♂ 1 ♀, TUA); Niigata-ken, Niigata-shi, Nishi-ku, Akatsuka, Sakata, 9. x. 2019, leg. K. Nakano (1 ♂ 3 ♀♀, TUA); Niigata-ken, Niigata-shi, Kita-ku, Nigorikawa, 7. vi. 2020, leg. K. Nakano (5 ♀♀, TUA); Niigata-ken, Niigata-shi, Chuo-ku, Nishifunami-cho, 12. vi. 2021, leg. G. Mashima (12 ♂♂ 17 ♀♀, TUA); Niigata-ken, Niigata-shi, Chuo-ku, Bandaijima, 30. viii. 2021, leg. J. Souma (18 ♂♂ 10 ♀♀, TUA); Shizuoka-ken, Shimoda-shi, Suzaki, Tsumekizaki, 21. vii. 2020, leg. J. Souma (1 ♂, TUA); Shizuoka-ken, Hamamatsu-shi, Nishi-ku, Kamigaya-cho, 16. vi. 2017, leg. J. Souma (15 ♂♂ 33 ♀♀, TUA); Shizuoka-ken, Numazu-shi, Kamikanuki, Higashihongo-cho, 27. xii. 2021, leg. J. Souma (1 ♀, TUA); Yamaguchi-ken, Shimonoseki-shi, Mimosusogawa-cho, 16. ix. 2022, leg. J. Souma (1 ♂ 3 ♀♀, ELKU). Izu Islands (northern part): Izu-Oshima Island: Okada, Minatono-mieru-oka, 4. vi. 2019, leg. Y. Tamadera (19 ♂♂ 37 ♀♀, TUA). Jogashima Island: 4. vi. 2019, leg. J. Souma (11 ♂♂ 9 ♀♀, TUA). Ebisu Island: 21. vii. 2020, leg. J. Souma (3 ♂♂, TUA). Shikoku: Ehime Pref., Kashima, 2. vi. 1971, leg. M. Tomokuni (1 ♂ 1 ♀, NSMT); Tosa, Nakagawa, 19. viii. 1953, leg. G. Yamamoto (1 ♂, ELKU); Kochi Pref., Cape Ashizuri, 7. vi. 1971, leg. M. Tomokuni (1 ♀, NSMT); Kochi-ken, Kochi-shi, Hitsuzan-cho, 30. vi. 2020, leg. J. Souma (1 ♀, TUA). Okinoshima Island (Kochi Prefecture): 11. viii. 1951, leg. T. Esaki (1 ♀, ELKU; 1 ♀, KUM). Kyushu: Prov. Buzen, Kokura, 20. xi. 1951, leg. A. Yamasaki (1 ♂, KUM); Fukuoka, Tachibanayama, 23. vii. 1961, leg. S. Miyamoto (6 ♂♂ 8 ♀♀, KUM); as above but 8. ix. 1961 (1 ♀, KUM); Fukuoka-ken, Fukuoka-shi, Nishi-ku, Motooka, Kyushu University, 23. v. 2020, leg. J. Souma (1 ♀, ELKU); Fukuoka-ken, Itoshima-shi, Tomari, 16. v. 2022, leg. J. Souma (1 ♀, ELKU); as above but 10. vii. 2022 (1 ♀, ELKU); Kumamoto-Pref., Kumamoto-City, Kuwamizuhonmachi, 4. i. 2021, leg. K. Goto (2 ♀♀, ELKU); Oita Pref., Saiki-shi, Yonouzu, Tsurumisaki, 2. vii. 2017, leg. R. Ito (1 ♀, ELKU); Oita Pref., Saiki-shi, Kamiura, Niinameura, 19. vii. 2020, leg. R. Ito (1 ♂ 3 ♀♀, ELKU); Miyazaki Pref., Nichinan-shi, Miyaura, 12. v. 2018, leg. R. Ito (1 ♂ 1 ♀, ELKU); Miyazaki Pref., Hyuga-shi, Okuragahama, 1. vi. 2019, leg. R. Ito (2 ♀♀, ELKU); Kagoshima, 21. v. 1953, leg. S. Miyamoto (1 ♂ 1 ♀, KUM); Oosumi, Izashiki ~ Ootomari, 25. v. 1953, leg. Yoshida (1 ♂, ELKU); Osumi, Sata, 29. v. 1953, leg. Yoshida (1 ♂ 1 ♀, ELKU); Osumi, Sata Cape, 30. v. 1953, leg. I. Hiura (4 ♀♀, KUM); Kagoshima-ken, Kagoshima-shi, Shiroyama-cho, 4. vii. 2017, leg. J. Souma (1 ♂, ELKU); Kagoshima, Minamiosumi-T., Sugiyama-dani Valley, 31. vii. 2017, leg. N. Tsuji (1 ♀, ELKU); Kagoshima Pref., Minamiosumi-cho, Sata, Hetsuka, 30. v. 2020, leg. R. Ito (9 ♂♂ 6 ♀♀, ELKU). Okinoshima Island (Fukuoka Prefecture): 25 - 28. vii. 1958, leg. Hirashima, Murakami & Y. Miyatake (35 ♂♂ 70 ♀♀, ELKU; 1 ♀, KUM). Tsushima Island: Izuhara-machi, Kitazato, Kamisaka, 27. vii. 2022, leg. Y. Uehara (1 ♀, ELKU). Amakusa Islands: Shimoshima Island: Tomioka, 12. ix. 1931, leg. Hori & Cho (2 ♂♂ 4 ♀♀, ELKU; 1 ♀, KUM). Goto Islands: Fukue Island: 1. ix. 1962, leg. S. Miyamoto (1 ♂ 1 ♀, KUM); as above but leg. S. Miyamoto & Kawarabata (1 ♀, ELKU); Inuyamaze, 2. ix. 1962, leg. S. Miyamoto (2 ♂♂ 1 ♀, KUM); as above but (1 ♂ 1 ♀, KUM); Arakawa, 3. ix. 1962, leg. S. Miyamoto (2 ♀♀, KUM). Sakura Island: Kurokami-cho, 27. vii. 2021, leg. Y. Obae (18 ♂♂ 13 ♀♀, TUA). Koshiki Islands: Kamikoshiki Island: Nakano, Mt. Tomeki, 2. v. 2019, leg. N. Kaneko (1 ♂ 4 ♀♀ 1 nymph, TUA). Shimokoshiki Island: Teuchi, 27 - 29. viii. 1960, leg. K. Morimoto (1 ♂ 2 ♀♀, ELKU); 25. v. 1975, leg. Y. Watanabe (1 ♂ 1 ♀, TUA). Ryukyu Islands (northern part): Tanegashima Island: Nishinoomote, 31. viii. 1952, leg. C. Takeya & Y. Hirashima (2 ♂♂, ELKU; 2 ♂♂ 2 nymphs, KUM); Nakatane-cho, Masuda, 10. vii. 2021, leg. T. Saeki (1 ♀, TUA). Yakushima Island: Onoaida, 26. viii. 1952, leg. C. Takeya & Y. Hirashima (3 ♂♂ 1 ♀, ELKU); as above but 27. viii. 1952 (8 ♂♂ 11 ♀♀ 1 nymph, ELKU); Miyanoura, 28. viii. 1952, leg. C. Takeya & Y. Hirashima (1 ♂ 4 ♀♀, ELKU); as above but 18. v. 2022, leg. J. Souma (1 ♀, TUA); Shiratani-unsuikyo, alt. 300 - 600 m, 14. vii. 2017, leg. R. Ito (1 ♂, ELKU); Kurio, Koyojigawa For. Rd., 7. vii. 2021, leg. T. Saeki (1 ♂, ELKU); Kurio, 15. viii. 2021, leg. J. Souma (3 ♂♂, ELKU); Koseda, 17. viii. 2021, leg. J. Souma (1 ♂, ELKU); as above but 19. v. 2022, leg. J. Souma (10 ♂♂ 14 ♀♀, TUA); Funayuki, 17. viii. 2021, leg. J. Souma (1 ♀, ELKU); Hirauchi, 19. viii. 2021, leg. J. Souma (1 ♂ 4 ♀♀, ELKU); Anbo, 20. viii. 2021, leg. J. Souma (9 ♂♂ 14 ♀♀, ELKU); Tabugawa, 18. v. 2022, leg. J. Souma (1 ♂ 3 ♀♀, TUA). Nakanoshima Island: 3 - 13. vi. 1953, leg. S. Miyamoto (1 ♀, ELKU); Okizaki, 5. vii. 2017, leg. J. Souma (2 ♀♀, TUA); as above but 6. vii. 2017 (1 ♀, TUA); Kusuki, 7. vii. 2017, leg. J. Souma (4 ♂♂ 7 ♀♀, TUA). Taira Island: Shuraku, 8 - 10. x. 2016, leg. H. Yoshitake (3 ♀♀, TUA); Higashinohama, 8 - 10. x. 2016, leg. H. Yoshitake (1 ♀, TUA). Akuseki Island: 24. iv. 1971, leg. M. Sakai (2 ♂♂ 3 ♀♀ 4 nymphs, NSMT); 17. vi. 2016, leg. H. Yoshitake (1 ♀, NIAES); Yudomari, 8. vii. 2017, leg. J. Souma (1 ♂ 3 ♀♀, TUA). Seven specimens collected from " Ohto " and " Yamogi " adjacent to the type locality " Sakuna " well match the original description of Stephanitis (Norba) aperta (Horvath 1912). In the present study, the author identified S. (N.) aperta based on these seven individuals. Syntype (s) of S. (N.) aperta exist (s) in the collection of HNHM (D. Redei, pers. comm. 2021).	en	Souma, Jun (2022): Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan. Deutsche Entomologische Zeitschrift 69 (2): 219-281, DOI: http://dx.doi.org/10.3897/dez.69.89864, URL: http://dx.doi.org/10.3897/dez.69.89864
9A2E144DDC53560E8C1AD97BF089175A.taxon	diagnosis	Diagnosis. Stephanitis (Norba) aperta is recognised amongst other species of Stephanitis by a combination of the following characters: head, pronotal disc, marking on hemelytra and ventral surface in various shades of brown (Figs 7 B, 9 B, 11 B, 13 B, 15 B, 17 B, 19 B, 21 B, 23 B); calli dark brown; body in male 2.1 times (in female 2.0 times) as long as maximum width across hemelytra (Figs 2 B, 4 B, 6 A); rostrum not reaching metasternum; pronotum unicarinate (Fig. 25 B); hood pale, shorter than median carina of pronotum, as wide as vertex at widest part, not covering eye, as high as median carina of pronotum at highest part, with posterior margin not extending to middle of pronotal disc; median carina of pronotum with 2 rows of areolae at highest part; pronotal disc opaque; paranotum less erect, narrowed posteriorly, with 3 rows of areolae at widest part, with anterolateral angle protruding anteriad, with outer margin gently curved inwards at posterolateral angle, maximum height shorter than height of eye (Fig. 27 B); apices of hemelytra close to each other in rest; costal area with 3 - 4 rows of areolae at widest part; subcostal area in male with 2 rows (in female with 3 rows) of areolae at widest part; discoidal area with 3 - 4 rows of areolae at widest part; sutural area with 3 - 4 rows of areolae at widest part; hypocostal lamina with a single row of areolae throughout its length; R + M (radiomedial) vein carinate; pygophore elevated at centre of venter, with posterior margin slightly emarginate in middle part (Fig. 29 B); and paramere stout, weakly curved inwards at apex, with outer margin not sinuate in middle part, inner margin nearly straight in basal part (Fig. 31 B).	en	Souma, Jun (2022): Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan. Deutsche Entomologische Zeitschrift 69 (2): 219-281, DOI: http://dx.doi.org/10.3897/dez.69.89864, URL: http://dx.doi.org/10.3897/dez.69.89864
9A2E144DDC53560E8C1AD97BF089175A.taxon	distribution	Distribution. Japan (Honshu; Izu Islands (northern part): Izu-Oshima Island; Jogashima Island; Ebisu Island; Shikoku; Okinoshima Island (Kochi Prefecture); Kyushu; Okinoshima Island (Fukuoka Prefecture); Tsushima Island; Amakusa Islands: Shimoshima Island; Goto Islands: Fukue Island; Sakura Island; Koshiki Islands: Kamikoshiki Island, Shimokoshiki Island; Ryukyu Islands (northern part): Tanegashima Island, Yakushima Island, Nakanoshima Island, Taira Island, Akuseki Island) (Fig. 45) (Horvath 1912; Takeya 1931, 1963; Yamada and Tomokuni 2012; Yamada and Ishikawa 2016; present study). Previous records from China in the 20 th century (Drake and Ruhoff 1965; Jing 1981) do not list the examined specimens and appear to be erroneous. Judging from the photographs, a recent record from Hong Kong (Yiu and Yip 2012) corresponds to another species, as the pronotum has lateral carina. Therefore, the presence of S. (N.) aperta in China remains unconfirmed. The previous record from the central part of the Ryukyu Islands (Miyamoto 1964 b; Azuma and Kinjo 1987; Hayashi 2002) corresponds to Stephanitis (Norba) exigua, S. (N.) hayashii sp. nov. or S. (N.) hiurai. The previous records from the southern part of the Ryukyu Islands and northern Taiwan (Takeya 1963; Miyamoto 1964 a; Azuma and Kinjo 1987; Hayashi 2002) correspond to S. (N.) ishikawai sp. nov., described below. Hundreds of specimens from the central and southern parts of the Ryukyu Islands possessing the unicarinate pronotum were examined, but all of them belong to S. (N.) exigua, S. (N.) hayashii sp. nov., S. (N.) hiurai or S. (N.) ishikawai sp. nov. Therefore, S. (N.) aperta is probably not distributed in the central and southern parts of Ryukyu Islands. Stephanitis (Norba) aperta inhabits the laurilignosa in a temperate climate of Japan proper (Honshu, Shikoku and Kyushu) and its surrounding islands and the northern part of the Izu and Ryukyu Islands, which is in the Palaearctic Region.	en	Souma, Jun (2022): Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan. Deutsche Entomologische Zeitschrift 69 (2): 219-281, DOI: http://dx.doi.org/10.3897/dez.69.89864, URL: http://dx.doi.org/10.3897/dez.69.89864
9A2E144DDC53560E8C1AD97BF089175A.taxon	biology_ecology	Biology. Stephanitis (Norba) aperta feeds on the abaxial surface of leaves of the three known host plants (present study). Adults were collected in almost all seasons (Takeya 1931, 1953; Yasunaga et al. 1993; Yano et al. 2013; present study); nymphs were collected in April, May, August, September and November (present study); the overwintering stage is represented by the adult (present study).	en	Souma, Jun (2022): Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan. Deutsche Entomologische Zeitschrift 69 (2): 219-281, DOI: http://dx.doi.org/10.3897/dez.69.89864, URL: http://dx.doi.org/10.3897/dez.69.89864
75671D62D1035B35AACF1E13DE5DC8EF.taxon	description	[Japanese name: Himetabu-gunbai] Figs 2 C, 4 C, 7 C, 9 C, 11 C, 13 C, 15 C, 17 C, 19 C, 21 C, 23 C, 25 C, 27 C, 29 C, 31 C, 35, 40 H, I	en	Souma, Jun (2022): Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan. Deutsche Entomologische Zeitschrift 69 (2): 219-281, DOI: http://dx.doi.org/10.3897/dez.69.89864, URL: http://dx.doi.org/10.3897/dez.69.89864
75671D62D1035B35AACF1E13DE5DC8EF.taxon	materials_examined	Material examined. Syntype (1 ♀, ELHU) (Fig. 34 C), JAPAN: Honshu: " タテヤマ " [= Chiba-ken, Tateyama-shi, former Tateyama-machi in early 20 th century (current Tateyama, Kamisanagura and Shimosanagura; approximate coordinates: 34 ° 58 ' 50.9 " N, 139 ° 51 ' 27.1 " E)], 11 / VIII 1905 [= 11. viii. 1905], " Matsumra " [sic; = collected by Shonen Matsumura and / or deposited in Matsumura's collection]. As pointed out in a previous study (Tomokuni 1994), this single female syntype corresponds to Stephanitis (Stephanitis) pyrioides (Scott, 1874) and does not match the original description of S. (Norba) exigua (Horvath 1912). Therefore, if the remaining syntypes are present in ELHU and / or HNHM, a lectotype should be designated. Nevertheless, the former curator of HNHM does not know if the syntype of S. (N.) exigua exists in the collection (D. Redei, pers. comm. 2021). Suspected syntypes (2 ♂♂ 1 ♀ 1 nymph, ELHU) (Fig. 35), JAPAN: Ryukyu Islands: " Okinawa " [= Okinawa-ken (a prefecture including a number of islands) or Okinawa Island (an island)], " 6 29 ". These four individuals were labelled with inscriptions of " Stephanitis yaeyamae " (unpublished name) and " Matsum " (collected by Shonen Matsumura and / or deposited in Matsumura's collection). The species epithet of the unpublished name seems to refer to the Yaeyama Islands, the southern part of the Ryukyu Islands, but this morphological species is only distributed in the Daito Islands and the central part of the Ryukyu Islands. The morphological characteristics and locality data of three adult specimens match the original description of S. (N.) exigua (Horvath 1912). However, the collector data of the four specimens are unclear and syntype (s) from " Okinawa " was (were) collected by " Kuroiwa " (Horvath 1912). Therefore, these three adults could correspond to syntypes of S. (N.) exigua. In the present study, the author identified S. (N.) exigua based on three adult individuals. Non-types (106 ♂♂ 189 ♀♀ 7 nymphs), JAPAN: Ryukyu Islands (central part): Okinawa Island: 10. viii. 1957, leg. T. Takara (1 ♀, NSMT); Kin, 14. vi. 1958, leg. T. Takara (2 ♀♀, NSMT); Osato, 8. xi. 1960, leg. K. Yasumatsu (2 ♂♂ 1 ♀, ELKU; 3 ♀♀, KUM); Yona, 14. xi. 1960, leg. K. Yasumatsu (1 ♂, KUM); as above but 19. xi. 1963, leg. H. Hasegawa (1 ♀, KUM); as above but 24. iii. 1964, leg. Y. Miyatake (1 ♀, KUM); as above but 23. xi. 1985, leg. M. Hayashi (2 ♂♂ 1 ♀, TUA); as above but 28. vi. 1984, leg. M. Tomokuni (2 ♀♀, NSMT); Shuri, 2. vi. 1961, leg. O. Nakoshi (1 ♂, KUM); Nago, 22. x. 1963, leg. S. Ueno (1 ♂, KUM); Tamagusuku, 17. xi. 1963, leg. H. Hasegawa (1 ♀, KUM); Kudeken, 20. iii. 1964, leg. Y. Miyatake (2 ♂♂ 4 ♀♀, KUM); Izumi, 22. iii. 1964, leg. T. Shirozu (9 ♂♂ 24 ♀♀, KUM); Izumi-Gogayama, 22. iii. 1964, leg. S. Kimoto (1 ♂, KUM); as above but leg. Y. Miyatake (3 ♂♂ 2 ♀♀, KUM); Shoshi, 23. iii. 1964, leg. S. Kimoto (1 ♂ 1 ♀, KUM); Nago, 23. iii. 1964, leg. Y. Miyatake (1 ♂ 2 ♀♀, KUM); Hiji-Yonahadake, 25. iii. 1964, leg. T. Shirozu (2 ♀♀, KUM); Hiji-gawa, 26. iii. 1964, leg. T. Shirozu (1 ♀, KUM); Chinen, Sefa utaki, 17. ii. 1973, leg. H. Hasegawa (1 ♀, NIAES); Yona, 21. ii. 1973, leg. H. Hasegawa (1 ♀, NIAES); Kunigami-son, Mt. Yonahadake, 29. vi. 1984, leg. M. Tomokuni (3 ♀♀, NSMT); Hanejiokawa, 14. xi. 1985, leg. M. Hayashi (1 ♂ 2 ♀♀, TUA); Kunigami, Mt. Nishime, 19. x. 1987, leg. M. Tomokuni (1 ♀, NSMT); Kunigami, Hama, 20. x. 1987, leg. M. Sakai (1 ♂ 3 ♀♀, NSMT); Kunigami, Ooguni-rindo, 8 km from Yona, alt. 300 m, 21. x. 1987, leg. M. Tomokuni (3 ♂♂ 3 ♀♀, NSMT); Kunigami, Hiji-Hiji Fall, 22. x. 1987, leg. M. Tomokuni (1 ♂, NSMT); Kudeken, Seifa-utaki, 8. x. 1988, leg. M. Sakai (1 ♂ 1 ♀, NSMT); Kunigamison, 10 - 11. x. 1988, leg. K. Konishi (1 ♀, NIAES); Nago City, 12. x. 1988, leg. K. Konishi (1 ♀, NIAES); Nakijin, Uebaru, 23. x. 1990, leg. M. Hayashi et al. (1 ♂, NSMT); Afuso, 3. iv. 1991, leg. M. Hayashi (1 ♂, TUA); Mt. Terukubi, 5. v. 1991, leg. M. Hayashi (1 ♀, TUA); Mt. Yonahadake, 3. iv. 1999, leg. M. Hayashi (1 ♀, TUA); Hedo, 16. ix. 2002, leg. M. Hayashi (1 ♀, TUA); Kisebaru, 10. xii. 2010, leg. M. Hayashi (1 ♀, TUA); Manzamo, 30. iii. 2013, leg. M. Hayashi (1 ♂, TUA); Nago, Katsuyama, Mt. Katsuudake, 9. vi. 2015, leg. H. Yoshitake (1 ♀, NIAES); Motobu, Namizato, Yaedake-sakura-no-mori-koen, 30. iii. 2018, leg. H. Yoshitake (4 ♂♂ 4 ♀♀, NIAES); Kunigami-son, Uka-rindo, 10. xi. 2018, leg. H. Yoshitake (1 ♀, NIAES); Naha-shi, Shuri-sueyoshi-cho, Sueyoshi-koen, 5. i. 2019, leg. H. Yoshitake (1 ♀, NIAES); Yaese Park, 19. i. 2019, leg. H. Shigetoh (4 ♂♂ 7 ♀♀ 1 nymph, TUA); as above but leg. H. Yoshitake (2 ♂♂ 1 ♀, NIAES); Nago-shi, Tanodake, 28. iv. 2019, leg. R. Ito (1 ♂, TUA); Kunigami-gun, Kunigami-son, Sate, 6. v. 2019, leg. R. Ito (1 ♂ 1 ♀, TUA); Nago, Genka Shisen For. Rd., 3. vi. 2019, leg. T. Saeki (1 ♂ 1 ♀, TUA); Uruma-shi, Mt. Ishikawadake, 29. xii. 2019, leg. H. Shigetoh (2 ♂♂ 1 ♀, TUA); Naha-shi, Ohnoyama Park, 8. iii. 2020, leg. J. Souma (4 ♂♂ 5 ♀♀ 3 nymphs); Kunigami-son, Aha, 5. v. 2019, leg. H. Yoshitake (1 ♀, NIAES); as above but 18. iv. 2020 (1 ♂ 1 ♀, NIAES); as above but 18. iv. 2020, leg. H. Shigetoh (3 ♀♀, TUA); Toyomigusuku, 6. xi. 2020, leg. J. Souma (1 ♀, TUA); Tabaru, 6. xi. 2020, leg. J. Souma (1 ♂ 9 ♀♀, TUA); Midorigaoka Park, 10. xi. 2020, leg. J. Souma (5 ♂♂ 4 ♀♀ 1 nymph, TUA); Nanjo-shi, Chinen-joseki, 30. i. 2021, leg. H. Yoshitake (1 ♀, NIAES); Kitanakagusuku-son, Taguchi, 3. ii. 2021, leg. H. Yoshitake (1 ♀, NIAES); Nakagusuku, Noborimata, 3. ii. 2021, leg. H. Yoshitake (1 ♀, NIAES); Okinawa-shi, Yaeshima-koen, 4. ii. 2021, leg. H. Yoshitake (1 ♀, NIAES); Uruma-shi, Enobi, 4. ii. 2021, leg. H. Yoshitake (1 ♀, NIAES); Yomitan-son, Zakimi-joseki, 5. ii. 2021, leg. H. Yoshitake (1 ♀, NIAES); Yonabaru-cho, Untamamori, 20. ii. 2021, leg. H. Yoshitake (1 ♀, NIAES); Nago-shi, Makiya, 1 - 2. vi. 2021, leg. T. Saeki (1 ♀, TUA). Aka Island: 13. viii. 1977, leg. M. Kinjo (1 ♀, NSMT); as above but 14. viii. 1977 (1 ♀, NSMT); as above but 14. viii. 1977, leg. S. Azuma (2 ♂♂ 1 ♀, NSMT); Aka, 4. v. 2021, leg. R. Ito (1 ♂ 3 ♀♀, TUA). Fukaji Island: 3. v. 2021, leg. R. Ito (6 ♂♂ 6 ♀♀, TUA). Geruma Island: 10. viii. 1977, leg. S. Azuma (1 ♀, NSMT); 2. viii. 2019, leg. H. Shigetoh (1 ♀, TUA); 4. v. 2021, leg. R. Ito (4 ♂♂ 2 ♀♀, TUA). Kume Island: Une, Tonnaha-enchi, 27. iii. 2018, leg. H. Yoshitake (1 ♀, NIAES); Daruma-yama, 28. iv. 2018, leg. R. Ito (3 ♂♂ 4 ♀♀, TUA); Shirase-Riv., 29. iv. 2018, leg. R. Ito (1 ♀, TUA); Yamashiro, 22 - 24. vii. 2020, leg. H. Yoshitake (1 ♀, NIAES). Tokashiki Island: near Shuraku, 27. iv. 2019, leg. H. Shigetoh (1 ♂, TUA); Tokashiki, 7. xi. 2020, leg. J. Souma (2 ♂♂ 9 ♀♀, TUA); Aharen, 7. xi. 2020, leg. J. Souma (2 ♂♂ 2 ♀♀, ELKU; 1 ♂ 3 ♀♀, TUA); Mt. Kumichizi, 8. xi. 2020, leg. J. Souma (2 ♂♂ 4 ♀♀ 2 nymphs, TUA); Otani-path, 1. v. 2018, leg. R. Ito (1 ♂ 2 ♀♀, TUA); Tokashiki, Otani road, 30. iv. 2021, leg. R. Ito (1 ♀, TUA); as above but 1. v. 2021 (4 ♂♂ 4 ♀♀, TUA). Tsuken Island: 16. iii. 2019, leg. H. Shigetoh (1 ♀, TUA). Yabuchi Island: 5. iii. 2020, leg. J. Souma (1 ♂ 1 ♀, TUA). Yagaji Island: Gabu, 9. iii. 2020, leg. J. Souma (2 ♂♂ 2 ♀♀, TUA); Sumuide, 17. iv. 2020, leg. H. Shigetoh (1 ♂, TUA). Zamami Island: near Mt. Odake, 20. iii. 2020, leg. H. Shigetoh (1 ♂ 3 ♀♀, TUA); Asa Evacuation Route, 21. iii. 2020, leg. H. Shigetoh (1 ♀, TUA); Ama, near Mt. Bansho, 21. iii. 2020, leg. H. Shigetoh (1 ♂ 1 ♀, TUA); Inazaki-Kaminohama, 21. iii. 2020, leg. H. Shigetoh (1 ♀, TUA); Ama, 3. vi. 2020, leg. H. Shigetoh (1 ♀, TUA); Asa, 2. v. 2018, leg. R. Ito (2 ♂♂ 2 ♀♀, TUA). Daito Islands: Kitadaito Island: Nakano, Daitogu Shrine, 24. xi. 2021, leg. T. Saeki (2 ♀♀, TUA). Minamidaito Island: Ikenosawa, 10. iii. 2013, leg. H. Yoshitake (1 ♂ 1 ♀, NIAES); Zaisho, 6. vii. 2014, leg. H. Ogai (5 ♂♂ 3 ♀♀, TUA); Daito Shrine, 9. ii. 2018, leg. R. Ito (3 ♂♂ 2 ♀♀, TUA). Six specimens from Aka and Geruma islands collected in 1977 were considered to be recorded as " Stephanitis aperta " by a previous study (Azuma and Kinjo 1987).	en	Souma, Jun (2022): Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan. Deutsche Entomologische Zeitschrift 69 (2): 219-281, DOI: http://dx.doi.org/10.3897/dez.69.89864, URL: http://dx.doi.org/10.3897/dez.69.89864
75671D62D1035B35AACF1E13DE5DC8EF.taxon	diagnosis	Diagnosis. Stephanitis (Norba) exigua is recognised amongst other species of Stephanitis by a combination of the following characters: head, pronotal disc, marking on hemelytra and ventral surface in various shades of brown (Figs 7 C, 9 C, 11 C, 13 C, 15 C, 17 C, 19 C, 21 C, 23 C); calli light brown; body in male 2.2 times (in female 2.1 times) as long as maximum width across hemelytra (Figs 2 C, 4 C); rostrum not reaching metasternum; pronotum unicarinate (Fig. 25 C); hood pale, shorter than median carina of pronotum, as wide as vertex at widest part, not covering eye, as high as median carina of pronotum at highest part, with posterior margin not extending to middle of pronotal disc; median carina of pronotum with 2 rows of areolae at highest part; pronotal disc lustrous; paranotum less erect, narrowed posteriorly, with 2 rows of areolae at widest part, with anterolateral angle slightly protruding anteriad, with outer margin gently curved inwards at posterolateral angle, maximum height shorter than height of eye (Fig. 27 C); apices of hemelytra close to each other in rest; costal area with 3 rows of areolae at widest part; subcostal area with 2 rows of areolae at widest part; discoidal area with 3 rows of areolae at widest part; sutural area with 3 - 4 rows of areolae at widest part; hypocostal lamina with a single row of areolae throughout its length; R + M (radiomedial) vein carinate; pygophore elevated at centre of venter, with posterior margin slightly emarginate in middle part (Fig. 29 C); and paramere stout, weakly curved inwards at apex, with outer margin not sinuate in middle part, inner margin nearly straight in basal part (Fig. 31 C).	en	Souma, Jun (2022): Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan. Deutsche Entomologische Zeitschrift 69 (2): 219-281, DOI: http://dx.doi.org/10.3897/dez.69.89864, URL: http://dx.doi.org/10.3897/dez.69.89864
75671D62D1035B35AACF1E13DE5DC8EF.taxon	distribution	Distribution. Japan (Ryukyu Islands (central part): Okinawa Island, Aka Island, Fukaji Island, Geruma Island, Kume Island, Tokashiki Island, Tsuken Island, Yabuchi Island, Yagaji Island, Zamami Island; Daito Islands: Kitadaito Island, Minamidaito Island) (Fig. 45); China (Horvath 1912; Drake 1937; Takeya 1963; Miyamoto 1964 a, 1964 b; Azuma and Kinjo 1987; Yamada and Tomokuni 2012; Yamada and Ishikawa 2016; Nakatani 2021; present study). The previous record from Honshu (Horvath 1912) is a misidentification of Stephanitis (Stephanitis) pyrioides. Hundreds of specimens from Honshu possessing the unicarinate pronotum were examined, but all of them belong to S. (Norba) aperta, S. (N.) mendica or S. (S.) tabidula. Therefore, S. (N.) exigua is probably not distributed in Honshu. The previous records from the southern part of the Ryukyu Islands and northern Taiwan (Takeya 1963; Miyamoto 1964 a, 1964 b, 1964 c; Azuma and Kinjo 1987; Hayashi 2002) correspond to S. (Norba) ishikawai sp. nov., described below. Hundreds of specimens from the southern part of the Ryukyu Islands possessing the unicarinate pronotum were examined, but all of them belong to S. (N.) ishikawai sp. nov. Therefore, S. (N.) exigua is probably not distributed in the southern part of the Ryukyu Islands. In Japan, S. (N.) exigua inhabits the laurilignosa in a subtropical climate of the central part of the Ryukyu Islands, which is in the Oriental Region.	en	Souma, Jun (2022): Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan. Deutsche Entomologische Zeitschrift 69 (2): 219-281, DOI: http://dx.doi.org/10.3897/dez.69.89864, URL: http://dx.doi.org/10.3897/dez.69.89864
75671D62D1035B35AACF1E13DE5DC8EF.taxon	biology_ecology	Biology. Stephanitis (Norba) exigua feeds on the abaxial surface of leaves of the three host plants in Japan (present study). In Japan, adults were collected in almost all seasons (Miyamoto 1964 a, 1964 b; present study); nymphs were collected in January, March and November (present study).	en	Souma, Jun (2022): Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan. Deutsche Entomologische Zeitschrift 69 (2): 219-281, DOI: http://dx.doi.org/10.3897/dez.69.89864, URL: http://dx.doi.org/10.3897/dez.69.89864
FC87BF49D8BB5D41BF0CA8D0818B70C6.taxon	description	[Japanese name: Hayashi-gunbai] Figs 2 D, 4 D, 7 D, 9 D, 11 D, 13 D, 15 D, 17 D, 19 D, 21 D, 23 D, 25 D, 27 D, 29 D, E, 31 D, E, 41 A-D	en	Souma, Jun (2022): Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan. Deutsche Entomologische Zeitschrift 69 (2): 219-281, DOI: http://dx.doi.org/10.3897/dez.69.89864, URL: http://dx.doi.org/10.3897/dez.69.89864
FC87BF49D8BB5D41BF0CA8D0818B70C6.taxon	materials_examined	Additional material examined (27 nymphs). JAPAN: Ryukyu Islands (central part): Kakeroma Island: Osai, 3. xi. 2020, leg. J. Souma (3 nymphs, ELKU). Aguni Island: Higashi, 6. iii. 2020, leg. J. Souma (6 nymphs, TUA); as above but 7. iii. 2020 (1 nymph, TUA); as holotype (6 nymphs, ELKU). Tokashiki Island: Tokashiki, 9. xi. 2020, leg. J. Souma (7 nymphs, ELKU). Yagaji Island: Gabu, 9. iii. 2020, leg. J. Souma (2 nymphs, ELKU; 2 nymphs, TUA). All 27 nymphs recorded above are in poor condition and are thus not described in the present study.	en	Souma, Jun (2022): Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan. Deutsche Entomologische Zeitschrift 69 (2): 219-281, DOI: http://dx.doi.org/10.3897/dez.69.89864, URL: http://dx.doi.org/10.3897/dez.69.89864
FC87BF49D8BB5D41BF0CA8D0818B70C6.taxon	diagnosis	Diagnosis. Stephanitis (Norba) hayashii sp. nov. is recognised amongst other species of Stephanitis by a combination of the following characters: head, pronotal disc, marking on hemelytra and ventral surface in various shades of brown (Figs 7 D, 9 D, 11 D, 13 D, 15 D, 17 D, 19 D, 21 D, 23 D); calli light brown; body in male 2.1 times (in female 2.0 times) as long as maximum width across hemelytra (Figs 2 D, 4 D); rostrum reaching metasternum; pronotum unicarinate (Fig. 25 D); hood pale, shorter than median carina of pronotum, wider than vertex at widest part, incompletely covering eye, as high as median carina of pronotum at highest part, with posterior margin extending to middle of pronotal disc; median carina of pronotum with 2 rows of areolae at highest part; pronotal disc opaque; paranotum more erect, slightly narrowed posteriorly, with 3 rows of areolae at widest part, with anterolateral angle protruding anteriad, with outer margin gently curved inwards at posterolateral angle, maximum height longer than height of eye (Fig. 27 D); apices of hemelytra close to each other in rest; costal area with 3 - 4 rows of areolae at widest part; subcostal area in male with 2 rows (in female with 3 rows) of areolae at widest part; discoidal area with 3 - 4 rows of areolae at widest part; sutural area with 3 - 4 rows of areolae at widest part; hypocostal lamina with a single row of areolae throughout its length; R + M (radiomedial) vein carinate; pygophore elevated at centre of venter, with posterior margin slightly emarginate in middle part (Fig. 29 D, E); and paramere slender, weakly curved inwards at apex, with outer margin not sinuate in middle part, inner margin slightly curved inwards in basal part (Fig. 31 D, E).	en	Souma, Jun (2022): Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan. Deutsche Entomologische Zeitschrift 69 (2): 219-281, DOI: http://dx.doi.org/10.3897/dez.69.89864, URL: http://dx.doi.org/10.3897/dez.69.89864
FC87BF49D8BB5D41BF0CA8D0818B70C6.taxon	description	Description. Male. Head, pronotal disc, marking on hemelytra and ventral surface various shades of brown; calli light brown; eye dark red; areolae of pronotum and hemelytron transparent; hood pale; pronotal disc opaque; pubescence on body yellowish (Figs 2 D, 7 D, 11 D, 15 D, 19 D, 21 D). Body 2.1 times as long as maximum width across hemelytra (Fig. 2 D). Head (Figs 7 D, 11 D, 19 D, 25 D) glabrous; pair of frontal spines close at apices, not reaching apex of clypeus; median spine as long as frontal spines, reaching bases of frontal spines; pair of occipital spines longer than median spine, reaching middle part of eyes; antenniferous tubercles obtuse, slightly curved inwards; clypeus smooth. Compound eye round in dorsal view. Antenna densely covered with pubescence; segment I cylindrical; segment II cylindrical, shortest amongst antennal segments; segment III longest amongst antennal segments; segment IV cylindrical, longer than segment I. Bucculae closed at anterior ends, with 3 rows of areolae throughout length. Rostrum reaching metasternum. Pronotum (Figs 7 D, 11 D, 25 D, 27 D) unicarinate, 1.4 times as long as maximum width across paranota, sparsely covered with pubescence. Pronotal disc coarsely punctate. Hood shorter than median carina of pronotum, wider than vertex at widest part, incompletely covering eye, as high as median carina of pronotum at highest part, posterior margin extending to middle of pronotal disc, 4 rows of areolae at highest part, dorsal margin arched. Median carina straight, extending to apex of posterior process, 2 rows of areolae at highest part, dorsal margin arched. Calli smooth. Paranotum more erect, slightly narrowed posteriorly, 3 rows of areolae at widest part, anterolateral angle protruding anteriad, outer margin gently curved inwards at posterolateral angle, maximum height longer than height of eye. Posterior process triangular, obtuse at apex. Hemelytron (Fig. 15 D) 2.4 times as long as maximum width, extending beyond apex of abdomen, sparsely covered with pubescence; maximum width across hemelytra 1.7 times as wide as maximum width across paranota; apices close in rest; costal area with 3 - 4 rows of areolae at widest part; subcostal area with 2 rows of areolae at widest part; discoidal area with 3 - 4 rows of areolae at widest part; sutural area with 3 - 4 rows of areolae at widest part; hypocostal lamina with a single row of areolae throughout its length; C (costal), R + M (radiomedial) and Cu (cubital) veins carinate. Thoracic pleura (Fig. 11 D) smooth in anterior part, coarsely punctate in posterior part. Ostiolar peritreme oblong. Sternal laminae (Fig. 19 D) lower than bucculae; pro- and mesosternal laminae open in both anterior and posterior ends; metasternal laminae as high as mesosternal laminae, open at anterior ends, fused each other at posterior ends. Legs (Fig. 2 D) smooth, densely covered with pubescence; femora thickest at middle. Abdomen oblong in dorsal and ventral views. Pygophore (Figs 21 D, 29 D, E) compressed dorsoventrally, semicircular in ventral view, elevated at centre of venter, with posterior margin slightly emarginate in middle part, covered with pubescence. Paramere (Fig. 31 D, E) slender, expanded in middle part, slightly curved inwards at apex, outer margin not sinuate in middle part, inner margin weakly curved inward in basal part, covered with pubescence in middle part of outer and inner margins. Measurements (n = 20). Body length with hemelytra 2.9 - 3.2 mm; maximum width across hemelytra 1.4 - 1.5 mm; length of antennal segments I to IV 0.2 mm, 0.1 mm, 1.2 mm and 0.6 mm, respectively; pronotal length 1.2 - 1.3 mm; pronotal width across paranota 0.8 - 0.9 mm; hemelytral length 2.2 - 2.5 mm; maximum width of hemelytron 1.0 - 1.1 mm. Female. General habitus very similar to that of male (Figs 4 D, 9 D, 13 D, 17 D, 23 D), except for the following characters: body 2.0 times as long as maximum width across hemelytra; antennal segment III shorter than in male; pronotum 1.5 times as long as maximum width across paranota; hood wider than in male; hemelytron 2.3 times as long as its maximum width; maximum width across hemelytra 1.9 times as much as maximum width across paranota; subcostal area wider than in male, with 3 rows of areolae at widest part; and apical part of abdomen pentagonal in ventral view. Measurements (n = 20). Body length with hemelytra 3.1 - 3.4 mm; maximum width across hemelytra 1.6 - 1.7 mm; length of antennal segments I to IV 0.2 mm, 0.1 mm, 1.0 mm and 0.6 mm, respectively; pronotal length 1.3 - 1.4 mm; pronotal width across paranota 0.9 mm; hemelytral length 2.4 - 2.5 mm; maximum width of hemelytron 1.0 - 1.1 mm.	en	Souma, Jun (2022): Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan. Deutsche Entomologische Zeitschrift 69 (2): 219-281, DOI: http://dx.doi.org/10.3897/dez.69.89864, URL: http://dx.doi.org/10.3897/dez.69.89864
FC87BF49D8BB5D41BF0CA8D0818B70C6.taxon	distribution	Distribution. Japan (Ryukyu Islands (central part): Amami-Oshima Island, Kakeroma Island, Yoron Island, Okinawa Island, Aguni Island, Fukaji Island, Kouri Island, Senaga Island, Tokashiki Island, Yagaji Island) (Fig. 46) (Miyamoto 1964 b; Yamada and Ishikawa 2016; present study). Stephanitis (Norba) hayashii sp. nov. inhabits the laurilignosa in a subtropical climate of the central part of the Ryukyu Islands, which is in the Oriental Region.	en	Souma, Jun (2022): Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan. Deutsche Entomologische Zeitschrift 69 (2): 219-281, DOI: http://dx.doi.org/10.3897/dez.69.89864, URL: http://dx.doi.org/10.3897/dez.69.89864
FC87BF49D8BB5D41BF0CA8D0818B70C6.taxon	etymology	Etymology. This new species is named in honour of Masami Hayashi, a Japanese heteropterist who collected part of paratypes and taught the author how to conduct fieldwork.	en	Souma, Jun (2022): Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan. Deutsche Entomologische Zeitschrift 69 (2): 219-281, DOI: http://dx.doi.org/10.3897/dez.69.89864, URL: http://dx.doi.org/10.3897/dez.69.89864
FC87BF49D8BB5D41BF0CA8D0818B70C6.taxon	biology_ecology	Biology. Stephanitis (Norba) hayashii sp. nov. feeds on the abaxial surface of leaves of the two known host plants (present study). Adults were collected from March to May and in January, August and November (Miyamoto 1964 b; present study); nymphs were collected in March and November (present study).	en	Souma, Jun (2022): Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan. Deutsche Entomologische Zeitschrift 69 (2): 219-281, DOI: http://dx.doi.org/10.3897/dez.69.89864, URL: http://dx.doi.org/10.3897/dez.69.89864
64316192E12352CEAB0E137EB21EC898.taxon	description	[Japanese name: Hiura-gunbai] Figs 2 E, F, 4 E, F, 7 E, F, 9 E, F, 11 E, F, 13 E, F, 15 E, F, 17 E, F, 19 E, 21 E, 23 E, 25 E, 28 A, 29 F, 31 F, 36 A, 36 B, 41 E-G	en	Souma, Jun (2022): Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan. Deutsche Entomologische Zeitschrift 69 (2): 219-281, DOI: http://dx.doi.org/10.3897/dez.69.89864, URL: http://dx.doi.org/10.3897/dez.69.89864
64316192E12352CEAB0E137EB21EC898.taxon	materials_examined	Material examined. Holotype (1 ♀, ELKU) (Fig. 36 A), JAPAN: Ryukyu Islands (central part): " Amami Is. Naze " [= Amami-Oshima Island, Naze (approximate coordinates: 28 ° 23 ' 13.6 " N, 129 ° 29 ' 38.2 " E)], 4. v. 1960, leg. T. Shibata. Holotype of Stephanitis (Norba) hiurai takaranis Takeya, 1963 (1 ♀, ELKU) (Fig. 36 B), JAPAN: Ryukyu Islands (central part): " Tokara Is. Takarajima " [= Takara Island (approximate coordinates: 29 ° 08 ' 38.0 " N, 129 ° 12 ' 27.8 " E)], 5. vii. 1960, leg. Y. Hama. Paratype of Stephanitis (Norba) hiurai takaranis Takeya, 1963 (1 ♀, ELKU), JAPAN: Ryukyu Islands (central part): " Tokara Is. Takarajima " [= Takara Island (approximate coordinates: 29 ° 08 ' 38.0 " N 129 ° 12 ' 27.8 " E)], 5. vii. 1960, leg. Y. Hama. The original description of S. (N.) hiurai and S. (N.) hiurai takaranis (Takeya 1963) states that their type specimens are deposited in OMNH, but these holotypes and paratype are now deposited in ELKU. Non-types collected at type locality (4 ♂♂ 6 ♀♀), JAPAN: Ryukyu Islands (central part): Amami-Oshima Island: Amami-shi, Naze, Kaneku (approximate coordinates: 28 ° 22 ' 27.1 " N, 129 ° 29 ' 43.6 " E), 30. iv. 2022, leg. J. Souma (3 ♂♂ 5 ♀♀, TUA); Amami-shi, Naze, Asani (approximate coordinates: 28 ° 24 ' 03.1 " N, 129 ° 29 ' 25.3 " E), 30. iv. 2022, leg. J. Souma (1 ♂ 1 ♀, TUA). Non-types (50 ♂♂ 104 ♀♀ 16 nymphs), JAPAN: Ryukyu Islands (central part): Takara Island: 26. v- 1. vi. 1953, leg. S. Miyamoto (1 ♂ 2 ♀♀, ELKU); 27. iv. 1971, leg. M. Sakai (2 ♀♀ 1 nymph, NSMT). Kikai Island: Hyakunodai, 9. iii. 2019, leg. H. Kojima (1 ♂ 3 ♀♀, TUA); between Keraji and Aden, 9. iii. 2019, leg. H. Kojima (9 ♀♀, TUA). Amami-Oshima Island: Hatsuno, 11. xi. 1962, leg. Y. Miyatake (1 ♂, ELKU); as above but 5. x. 1988, leg. M. Tomokuni (1 ♂ 3 ♀♀, NSMT); Uragami, 31. x. 1966, leg. Y. Miyatake (3 ♂♂ 2 ♀♀, KUM); Santaro-toge Pass, 2. xi. 1966, leg. Y. Miyatake (1 ♀, KUM); as above but 30. vi. 2000, leg. Y. Nakatani (1 ♂, NIAES); Yuwan, 3. xi. 1966, leg. Y. Miyatake (5 ♂♂ 10 ♀♀, KUM); Naze, 21. iv. 1971, leg. M. Sakai (3 ♀♀ 1 nymph, NSMT); Naze-shi, Akazaki, 2. xi. 1984, leg. M. Tomokuni (1 ♂ 5 ♀♀, NSMT); Sumiyo-son, Kawauchi, Chuo-rindo, alt. 200 m, 3. x. 1988, leg. M. Tomokuni (5 ♀♀, NSMT); Mt. Yuwandake, 4. x. 1988, leg. M. Tomokuni (1 ♀, NSMT); Uken-son, Mt. Yuwan-dake, 23. iii. 2019, leg. Y. Hisasue (1 ♀, TUA); Shinokawa, 2. xi. 2020, leg. J. Souma (3 ♂♂ 1 ♀, ELKU; 5 ♂♂ 5 ♀♀, TUA); as above but 4. xi. 2020 (1 ♂ 1 ♀ 8 nymphs, TUA); Konase, 2. xi. 2020, leg. J. Souma (5 ♂♂ 5 ♀♀ 1 nymph, TUA); Sokaru, 5. xi. 2020, leg. J. Souma (1 ♀, TUA); Uken-son, Yuwan, 28. iv. 2022, leg. J. Souma (1 ♂ 2 ♀♀, TUA); Setouchi-cho, Amurogama, 29. iv. 2022, leg. J. Souma (3 ♀♀, TUA); Uken-son, Ikegachi, 30. iv. 2022, leg. J. Souma (1 ♂, TUA); Amami-shi, Sumiyo-cho, Aoku, 30. iv. 2022, leg. J. Souma (1 ♂ 1 ♀, TUA); Amami-shi, Sumiyo-cho, Santaro Pass, 1. v. 2022, leg. J. Souma (1 ♂ 5 ♀♀ 4 nymphs, TUA); Amami-shi, Kasari-cho, Manya, 3. v. 2022, leg. J. Souma (2 ♂♂ 2 ♀♀, TUA); Asani, Akazaki Park, 29. vi. 2022, leg. S. Imada (1 ♀, TUA). Kakeroma Island: Shokazu, 3. xi. 2020, leg. J. Souma (3 ♀♀, TUA); Kachiyuki, 3. xi. 2020, leg. J. Souma (8 ♂♂ 10 ♀♀ 1 nymph, TUA). Tokunoshima Island: Asahigaoka, 11. xi. 1966, leg. Y. Miyatake (2 ♀♀, KUM); Amagi-cho, Yonama, 2. x. 1988, leg. M. Tomokuni (2 ♂ 1 ♀, NSMT). Okinoerabu Island: Mt. Koshiyama, 8. vii. 2019, leg. Y. Tamadera (1 ♂ 3 ♀♀, TUA); Oyama Botanical Park, 23 - 26. vi. 2022, leg. S. Imada (3 ♀♀, TUA). Okinawa Island: Yona, 19. x. 1963, leg. S. Miyamoto (2 ♂♂, KUM); as above but leg. Y. Hirashima (1 ♀, KUM); as above but 25 - 27. v. 1974, leg. M. Sato (2 ♀♀, NSMT); Izumi-Gogayama, 22. iii. 1964, leg. S. Kimoto (1 ♂, KUM); as above but leg. Y. Miyatake (1 ♀, KUM); Aha, 26. xii. 1973, leg. M. Hamakawa (1 ♂ 1 ♀, NSMT); Kunigami, Mt. Yonaha, 20. x. 1987, leg. M. Sakai (1 ♀, NSMT); Kunigami, Ooguni-rindo, 8 km from Yona, alt. 300 m, 21. x. 1987, leg. M. Tomokuni (1 ♂, NSMT); Okuni-Rindo, 16 - 21. iv. 1997, leg. T. Ishikawa (1 ♀, TUA); Oku For. Rd., 12. ix. 2005, leg. M. Hayashi (1 ♀, TUA).	en	Souma, Jun (2022): Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan. Deutsche Entomologische Zeitschrift 69 (2): 219-281, DOI: http://dx.doi.org/10.3897/dez.69.89864, URL: http://dx.doi.org/10.3897/dez.69.89864
64316192E12352CEAB0E137EB21EC898.taxon	diagnosis	Diagnosis. Stephanitis (Norba) hiurai is recognised amongst other species of Stephanitis by a combination of the following characters: head, pronotal disc, marking on hemelytra and ventral surface in various shades of brown (Figs 7 E, F, 9 E, F, 11 E, F, 13 E, F, 15 E, F, 17 E, F, 19 E, 21 E, 23 E); calli light brown; body in male 2.0 - 2.1 times (in female 1.8 - 2.0 times) as long as maximum width across hemelytra (Figs 2 E, F, 4 E, F); rostrum not reaching metasternum; pronotum unicarinate (Fig. 25 E); hood pale, shorter than median carina of pronotum, wider than vertex at widest part, incompletely covering eye, as high as median carina of pronotum at highest part, with posterior margin not extending to middle of pronotal disc; median carina of pronotum with 2 rows of areolae at highest part; pronotal disc lustrous; paranotum less erect, narrowed posteriorly, 3 rows of areolae at widest part, anterolateral angle protruding anteriad, outer margin gently curved inwards at posterolateral angle, maximum height longer than height of eye (Fig. 28 A); apices of hemelytra close to each other in rest; costal area with 4 rows of areolae at widest part; subcostal area in male with 2 rows (in female with 3 rows) of areolae at widest part; discoidal area with 3 rows of areolae at widest part; sutural area with 3 - 4 rows of areolae at widest part; hypocostal lamina with a single row of areolae throughout its length; R + M (radiomedial) vein in male not carinate (in female carinate); pygophore elevated at centre of venter, posterior margin slightly emarginate in middle part (Fig. 29 F); and paramere stout, weakly curved inwards at apex, with outer margin not sinuate in middle part, inner margin nearly straight in basal part (Fig. 31 F).	en	Souma, Jun (2022): Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan. Deutsche Entomologische Zeitschrift 69 (2): 219-281, DOI: http://dx.doi.org/10.3897/dez.69.89864, URL: http://dx.doi.org/10.3897/dez.69.89864
64316192E12352CEAB0E137EB21EC898.taxon	distribution	Distribution. Japan (Ryukyu Islands (central part): Takara Island, Kikai Island, Amami-Oshima Island, Kakeroma Island, Tokunoshima Island, Okinoerabu Island, Okinawa Island) (Fig. 46) (Takeya 1963; Miyamoto 1964 a, 1964 b, 1964 c; Hayashi 2002; Yamada and Tomokuni 2012; Yamada and Ishikawa 2016; present study). The previous record from Ishigaki Island, the southern part of the Ryukyu Islands (Miyamoto 1964 a, 1964 b), corresponds to Stephanitis (Norba) ishikawai sp. nov., described below. Hundreds of specimens from the southern part of the Ryukyu Islands possessing an unicarinate pronotum were examined, but all of them belonged to S. (N.) ishikawai sp. nov. Therefore, S. (N.) hiurai is probably not distributed in the southern part of the Ryukyu Islands. Stephanitis (Norba) hiurai inhabits the laurilignosa in a subtropical climate of the central part of the Ryukyu Islands, which is in the Oriental Region.	en	Souma, Jun (2022): Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan. Deutsche Entomologische Zeitschrift 69 (2): 219-281, DOI: http://dx.doi.org/10.3897/dez.69.89864, URL: http://dx.doi.org/10.3897/dez.69.89864
64316192E12352CEAB0E137EB21EC898.taxon	biology_ecology	Biology. Stephanitis (Norba) hiurai feeds on the abaxial surface of leaves of Machilus thunbergii (present study. Adults were collected in almost all seasons (Takeya 1963; Miyamoto 1964 a, 1964 b; present study). Nymphs were collected in April and November (present study).	en	Souma, Jun (2022): Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan. Deutsche Entomologische Zeitschrift 69 (2): 219-281, DOI: http://dx.doi.org/10.3897/dez.69.89864, URL: http://dx.doi.org/10.3897/dez.69.89864
3A2B74D8666753F7AD7C78A31C6283F9.taxon	description	[Japanese name: Ishikawa-gunbai] Figs 3 A, 5 A, 8 A, 10 A, 12 A, 14 A, 16 A, 18 A, 19 F, 21 F, 23 F, 25 F, 28 B, 30 A-C, 32 A-C, 41 H, I	en	Souma, Jun (2022): Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan. Deutsche Entomologische Zeitschrift 69 (2): 219-281, DOI: http://dx.doi.org/10.3897/dez.69.89864, URL: http://dx.doi.org/10.3897/dez.69.89864
3A2B74D8666753F7AD7C78A31C6283F9.taxon	materials_examined	Additional material examined. Non-types (13 ♂♂ 8 ♀♀ 13 nymphs), JAPAN: Ryukyu Islands: (southern part): Iriomote Island: Shirahama-Hoshidate, 8. iii. 1964, leg. Y. Miyatake (1 nymph); Ohara, Fusatoruba, 3. iii. 2002, leg. T. Ishikawa (4 nymphs, TUA). Yonaguni Island: as holotype (1 nymph). TAIWAN: Taipei City: Taihoku [= Taipei], 16. iv. 1917, leg. M. Maki (2 ♂♂, ELKU); as above but 15. i. 1927, leg. R. Takahashi (1 ♀, ELKU); as above but 2. iv. 1930, leg. C. Takeya (1 ♂ 1 ♀, ELKU); as above, but 10. ii. 1932, leg. R. Takahashi (4 ♂♂ 2 ♀♀, ELKU; 1 ♂, KUM); Da'an District, 5. x. 2021, leg. Y. - J. Tsai (3 ♂♂ 1 ♀ 7 nymphs, NMNS); as above but 22. xi. 2021 (2 ♂♂ 3 ♀♀, NMNS). Twelve specimens collected from Taiwan in the early 20 th century were recorded as " Stephanitis aperta " or " Stephanitis exigua " by Takeya (1931, 1963). The Taiwanese specimens are most similar to Stephanitis (Norba) ishikawai sp. nov. in morphological characteristics and were provisionally identified as pertaining to the new species in the present study. All 13 nymphs recorded above are in poor condition and are thus not described here.	en	Souma, Jun (2022): Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan. Deutsche Entomologische Zeitschrift 69 (2): 219-281, DOI: http://dx.doi.org/10.3897/dez.69.89864, URL: http://dx.doi.org/10.3897/dez.69.89864
3A2B74D8666753F7AD7C78A31C6283F9.taxon	diagnosis	Diagnosis. Stephanitis (Norba) ishikawai sp. nov. is recognised amongst other species of Stephanitis by a combination of the following characters: head, pronotal disc, marking on hemelytra and ventral surface in various shades of brown (Figs 8 A, 10 A, 12 A, 14 A, 16 A, 18 A, 19 F, 21 F, 23 F); calli light brown; body in male 2.1 times (in female 2.0 times) as long as maximum width across hemelytra (Figs 3 A, 5 A); rostrum reaching metasternum; pronotum unicarinate (Fig. 25 F); hood pale, shorter than median carina of pronotum, wider than vertex at widest part, incompletely covering eye, as high as median carina of pronotum at highest part, posterior margin extending to middle of pronotal disc; median carina of pronotum with 2 rows of areolae at highest part; pronotal disc opaque; paranotum less erect, slightly narrowed posteriorly, with 3 rows of areolae at widest part, anterolateral angle protruding anteriad, outer margin gently curved inwards at posterolateral angle, maximum height longer than height of eye (Fig. 28 B); apices of hemelytra close to each other in rest; costal area with 3 - 4 rows of areolae at widest part; subcostal area in male with 2 rows (in female with 3 rows) of areolae at widest part; discoidal area with 3 - 4 rows of areolae at widest part; sutural area with 3 - 4 rows of areolae at widest part; hypocostal lamina with a single row of areolae throughout its length; R + M (radiomedial) vein carinate; pygophore elevated at centre of venter, posterior margin slightly emarginate in middle part (Fig. 30 A-C); and paramere stout, weakly curved inwards at apex, with outer margin not sinuate in middle part, inner margin nearly straight in basal part (Fig. 32 A-C).	en	Souma, Jun (2022): Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan. Deutsche Entomologische Zeitschrift 69 (2): 219-281, DOI: http://dx.doi.org/10.3897/dez.69.89864, URL: http://dx.doi.org/10.3897/dez.69.89864
3A2B74D8666753F7AD7C78A31C6283F9.taxon	description	Description. Male. Head, pronotal disc, marking on hemelytra and ventral surface in various shades of brown; calli light brown; eye dark red; areolae of pronotum and hemelytron transparent; hood pale; pronotal disc opaque; pubescence on body yellowish (Figs 3 A, 8 A, 12 A, 16 A, 19 F, 21 F). Body 2.1 times as long as maximum width across hemelytra (Fig. 3 A). Head (Figs 8 A, 12 A, 19 F, 25 F) glabrous; a pair of frontal spines close to each other at apices, not reaching apex of clypeus; median spine as long as frontal spines, reaching bases of frontal spines; pair of occipital spines longer than median spine, reaching middle part of eyes; antenniferous tubercles obtuse, slightly curved inwards; clypeus smooth. Compound eye round in dorsal view. Antenna densely covered with pubescence; segment I cylindrical; segment II cylindrical, shortest among antennal segments; segment III longest amongst antennal segments; segment IV cylindrical, longer than segment I. Bucculae closed at anterior ends, with 3 rows of areolae throughout length. Rostrum reaching metasternum. Pronotum (Figs 8 A, 12 A, 25 F, 28 B) unicarinate, 1.3 times as long as maximum width across paranota, sparsely covered with pubescence. Pronotal disc coarsely punctate. Hood shorter than median carina of pronotum, wider than vertex at widest part, incompletely covering eye, as high as median carina of pronotum at highest part, posterior margin extending to middle of pronotal disc, 4 rows of areolae at highest part, dorsal margin arched. Median carina straight, extending to apex of posterior process, 2 rows of areolae at highest part, dorsal margin arched. Calli smooth. Paranotum less erect, slightly narrowed posteriorly, with 3 rows of areolae at widest part, anterolateral angle protruding anteriad, outer margin gently curved inwards at posterolateral angle, maximum height longer than height of eye. Posterior process triangular, obtuse at apex. Hemelytron (Fig. 16 A) 2.4 times as long as its maximum width, extending beyond apex of abdomen, sparsely covered with pubescence; maximum width across hemelytra 1.6 times as much as maximum width across paranota; apices close to each other in rest; costal area with 3 - 4 rows of areolae at widest part; subcostal area with 2 rows of areolae at widest part; discoidal area with 3 - 4 rows of areolae at widest part; sutural area with 3 - 4 rows of areolae at widest part; hypocostal lamina with a single row of areolae throughout its length; C (costal), R + M (radiomedial) and Cu (cubital) veins carinate. Thoracic pleura (Fig. 12 A) smooth in anterior part, coarsely punctate in posterior part. Ostiolar peritreme oblong. Sternal laminae (Fig. 19 F) lower than bucculae; pro- and mesosternal laminae open in both anterior and posterior ends; metasternal laminae as high as mesosternal laminae, open at anterior ends, fused each other at posterior ends. Legs (Fig. 3 A) smooth, densely covered with pubescence; femora thickest at middle. Abdomen oblong in dorsal and ventral views. Pygophore (Figs 21 F, 30 A-C) compressed dorsoventrally, semicircular in ventral view, elevated at centre of venter, with posterior margin slightly emarginate in middle part, covered with pubescence. Paramere (Fig. 32 A-C) stout, expanded in middle part, weakly curved inwards at apex, outer margin not sinuate in middle part, inner margin nearly straight in basal part, covered with pubescence in middle part of outer and inner margins. Measurements (n = 20). Body length with hemelytra 2.9 - 3.2 mm; maximum width across hemelytra 1.4 - 1.6 mm; length of antennal segments I to IV 0.2 mm, 0.1 mm, 1.2 mm and 0.7 - 0.8 mm, respectively; pronotal length 1.2 - 1.3 mm; pronotal width across paranota 0.8 - 1.0 mm; hemelytral length 2.2 - 2.5 mm; maximum width of hemelytron 0.9 - 1.1 mm. Female. General habitus very similar to that of male (Figs 4 D, 9 D, 13 D, 17 D, 23 D), except for the following characters: body 2.0 times as long as maximum width across hemelytra; antennal segment III shorter than in male; pronotum 1.4 times as long as maximum width across paranota; hood wider than in male; hemelytron 2.3 times as long as maximum width; maximum width across hemelytra 1.7 times as much as maximum width across paranota; subcostal area wider than in male, with 3 rows of areolae at widest part; and apical part of abdomen pentagonal in ventral view. Measurements (n = 20). Body length with hemelytra 3.0 - 3.4 mm; maximum width across hemelytra 1.5 - 1.7 mm; length of antennal segments I to IV 0.2 mm, 0.1 mm, 1.0 mm and 0.7 - 0.8 mm, respectively; pronotal length 1.2 - 1.4 mm; pronotal width across paranota 0.9 - 1.0 mm; hemelytral length 2.3 - 2.5 mm; maximum width of hemelytron 1.0 - 1.1 mm.	en	Souma, Jun (2022): Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan. Deutsche Entomologische Zeitschrift 69 (2): 219-281, DOI: http://dx.doi.org/10.3897/dez.69.89864, URL: http://dx.doi.org/10.3897/dez.69.89864
3A2B74D8666753F7AD7C78A31C6283F9.taxon	distribution	Distribution. Japan (Ryukyu Islands (southern part): Miyako Island, Irabu Island, Kurima Island, Ogami Island, Ishigaki Island, Iriomote Island, Yonaguni Island); Taiwan (northern part) (Fig. 47) (Takeya 1931, 1963; Maa 1957; Miyamoto 1964 a, 1964 b, 1964 c; Takara and Azuma 1972; Azuma and Kinjo 1987; Yasunaga et al. 1993; Hayashi 2002; Yamada and Tomokuni 2012; Zheng and Lin 2013; Yamada and Ishikawa 2016; present study). Judging from the photographs, living individuals identified as " Stephanitis (Norba) aperta " or " S. (N.) exigua " in previous studies (Yasunaga et al. 193; Yamada and Tomokuni 2012; Zheng and Lin 2013) corresponded to the new species. Stephanitis (Norba) ishikawai sp. nov. inhabits the laurilignosa in a subtropical climate of the southern part of the Ryukyu Islands and northern Taiwan, which is in the Oriental Region.	en	Souma, Jun (2022): Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan. Deutsche Entomologische Zeitschrift 69 (2): 219-281, DOI: http://dx.doi.org/10.3897/dez.69.89864, URL: http://dx.doi.org/10.3897/dez.69.89864
3A2B74D8666753F7AD7C78A31C6283F9.taxon	etymology	Etymology. The new species is named in honour of Tadashi Ishikawa, a Japanese heteropterist who collected part of paratypes and taught the author how to describe new species.	en	Souma, Jun (2022): Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan. Deutsche Entomologische Zeitschrift 69 (2): 219-281, DOI: http://dx.doi.org/10.3897/dez.69.89864, URL: http://dx.doi.org/10.3897/dez.69.89864
3A2B74D8666753F7AD7C78A31C6283F9.taxon	biology_ecology	Biology. Stephanitis (Norba) ishikawai sp. nov. feeds on the abaxial surface of leaves of the three host plants (present study). Adults were collected in almost all seasons (Miyamoto 1964 a, 1964 b, 1964 c; Takara and Azuma 1972; Yasunaga et al. 1993; Yamada and Tomokuni 2012; present study); nymphs were collected in March and November (present study).	en	Souma, Jun (2022): Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan. Deutsche Entomologische Zeitschrift 69 (2): 219-281, DOI: http://dx.doi.org/10.3897/dez.69.89864, URL: http://dx.doi.org/10.3897/dez.69.89864
6D10E055B4255DE3AC49B8540B033EFB.taxon	description	[Japanese name: Yabunikkei-gunbai] Figs 3 B, 5 B, 8 B, 10 B, 12 B, 14 B, 16 B, 18 B, 20 A, 22 A, 24 A, 26 A, B, 28 C, 30 D, 32 D, 42 A, B	en	Souma, Jun (2022): Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan. Deutsche Entomologische Zeitschrift 69 (2): 219-281, DOI: http://dx.doi.org/10.3897/dez.69.89864, URL: http://dx.doi.org/10.3897/dez.69.89864
6D10E055B4255DE3AC49B8540B033EFB.taxon	materials_examined	Material examined. Non-types (186 ♂♂ 291 ♀♀ 9 nymphs), JAPAN: Honshu: Chiba-ken, Tateyama-shi, Shimosanagura, 22. v. 2021, leg. J. Souma (19 ♂♂ 16 ♀♀ 1 nymph, TUA); Chiba-ken, Tateyama-shi, Tateyama, 22. v. 2021, leg. J. Souma (8 ♂♂ 1 ♀ 5 nymphs); as above but 23. v. 2021 (7 ♂♂ 7 ♀♀, TUA); Chiba-ken, Tateyama-shi, Sunozaki, 23. v. 2021, leg. J. Souma (1 ♂ 3 ♀♀); Chiba-ken, Tateyama-shi, Yamogi (approximate coordinates: 34 ° 58 ' 08.9 " N, 139 ° 53 ' 33.4 " E), 24. v. 2021, leg. J. Souma (3 ♂♂ 1 nymph, ELKU; 1 ♂ 4 ♀♀, TUA); Kanagawa-ken, Yokohama-shi, Kanazawa-ku, Noukendaimori. 15. vi. 2017, leg. J. Souma (45 ♂♂ 61 ♀♀, TUA); Kanagawa-ken, Yokosuka-shi, Kamoi, 27. vi. 2017, leg. J. Souma (1 ♂ 3 ♀♀, ELKU; 38 ♂♂ 98 ♀♀, TUA). Jogashima Island: 4. vi. 2019, leg. J. Souma (7 ♂♂ 13 ♀♀, TUA). Shikoku: Kochi Pref., Ashizuri-misaki, 29. v. 1999, leg. T. Befu (1 ♂, NSMT); Kochi Pref., Mt. Oodo, 3. vi. 1971, leg. M. Tomokuni (2 ♂♂ 2 nymphs, NSMT). Kyushu: Chikuzen, Fukuoka, 27. vi. 1931, leg. C. Takeya (1 ♂, ELKU; 5 ♂♂ 5 ♀♀, KUM); Chikuzen, Aburayama, 6. vii. 1952, leg. C. Takeya (4 ♀♀, KUM); Fukuoka, Atagoyama, 26. vi. 1959, leg. Y. Miyatake (1 ♂, KUM); as above but 4. viii. 1961, leg. S. Miyamoto (1 ♀, KUM); Fukuoka, Hirao, 4. vii. 1959, leg. Y. Miyatake (4 ♂♂ 4 ♀♀, KUM); Fukuoka-ken, Itoshima-shi, Shimakeya, 14. vi. 2021, leg. J. Souma (6 ♂♂ 11 ♀♀, ELKU); Oita Pref., Saiki-shi, Kamiura, Niinameura, 19. vii. 2020, leg. R. Ito (1 ♂ 5 ♀♀, ELKU); Miyazaki Pref., Takanabe-cho, Mochida, Omarugawa, 12. v. 2019, leg. R. Ito (13 ♂♂ 8 ♀♀, ELKU); Miyazaki Pref., Hyuga-shi, Okuragahama, 1. vi. 2019, leg. R. Ito (4 ♂♂ 6 ♀♀, ELKU); Kagoshima-ken, Kagoshima-shi, Shiroyama-cho, 4. vii. 2017, leg. J. Souma (4 ♀♀, TUA); Kagoshima Pref., So-shi, Sueyoshi-cho, Minaminogo, 8. vi. 2019, leg. R. Ito (1 ♀, ELKU); Kagoshima Pref., Kanoya-shi, Aira-cho, Kamimyo, 7. vi. 2020, leg. R. Ito (1 ♀, ELKU). Nokonoshima Island: 27. vi. 1987, leg. S. Miyamoto (2 ♀♀, KUM). Ryukyu Islands (northern and central parts): Yakushima Island: Tabugawa, 18. v. 2022, leg. J. Souma (13 ♂♂ 12 ♀♀, TUA); Kusugawa, 18. v. 2022, leg. J. Souma (5 ♂♂ 6 ♀♀, TUA); Koseda, 19. v. 2022, leg. J. Souma (3 ♂♂ 1 ♀, TUA). Amami-Oshima Island: Amami-shi, Sumiyo-cho, Ishihara, 2. v. 2022, leg. J. Souma (4 ♂♂ 2 ♀♀, TUA); Amami-shi, Kasari-cho, Manya, 3. v. 2022, leg. J. Souma (2 ♂♂ 12 ♀♀, TUA). Okinawa Island: 20. iv. 1958, leg. T. Takara (2 ♂♂ 2 ♀♀, NSMT). Eight adult specimens collected from " Yamogi ", which is adjacent to one of the type localities, " Sakuna ", match the original description of Stephanitis (Norba) mendica (Horvath 1912). The author identified S. (N.) mendica based on these eight adults in the present study. Syntype (s) of S. (N.) mendica exist in the collection of HNHM (D. Redei, pers. comm. 2021). Four specimens collected from Okinawa Island were recorded as " Stephanitis fasciicarina " by Takara and Hidaka (1960).	en	Souma, Jun (2022): Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan. Deutsche Entomologische Zeitschrift 69 (2): 219-281, DOI: http://dx.doi.org/10.3897/dez.69.89864, URL: http://dx.doi.org/10.3897/dez.69.89864
6D10E055B4255DE3AC49B8540B033EFB.taxon	diagnosis	Diagnosis. Stephanitis (Norba) mendica is recognised amongst other species of Stephanitis by a combination of the following characters: head, pronotal disc, marking on hemelytra and ventral surface in various shades of brown (Figs 8 B, 10 B, 12 B, 14 B, 16 B, 18 B, 20 A, 22 A, 24 A); calli light brown; body in male 2.2 times (in female 2.0 times) as long as maximum width across hemelytra (Figs 3 B, 5 B); rostrum reaching metasternum; pronotum unicarinate or tricarinate (Fig. 26 A, B); hood pale, shorter than median carina of pronotum, wider than vertex at widest part, incompletely covering eye, higher than median carina of pronotum at highest part, with posterior margin extending to middle of pronotal disc; median carina of pronotum with 2 rows of areolae at highest part; pronotal disc opaque; paranotum more erect, not narrowed posteriorly, with 3 rows of areolae at widest part, with anterolateral angle protruding anteriad, with outer margin angularly curved inwards at posterolateral angle, maximum height longer than height of eye (Fig. 28 C); apices of hemelytra close to each other in rest; costal area with 3 - 4 rows of areolae at widest part; subcostal area in male with 2 rows (in female with 2 - 3 rows) of areolae at widest part; discoidal area with 3 rows of areolae at widest part; sutural area with 3 - 4 rows of areolae at widest part; hypocostal lamina with a single row of areolae throughout its length; R + M (radiomedial) vein not carinate; pygophore elevated at centre of venter, with posterior margin slightly emarginate in middle part (Fig. 30 D); and paramere stout, weakly curved inwards at apex, with outer margin not sinuate in middle part, inner margin nearly straight in basal part (Fig. 32 D).	en	Souma, Jun (2022): Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan. Deutsche Entomologische Zeitschrift 69 (2): 219-281, DOI: http://dx.doi.org/10.3897/dez.69.89864, URL: http://dx.doi.org/10.3897/dez.69.89864
6D10E055B4255DE3AC49B8540B033EFB.taxon	distribution	Distribution. Japan (Honshu; Jogashima Island; Shikoku; Kyushu; Nokonoshima Island; Ryukyu Islands (northern and central parts): Yakushima Island, Amami-Oshima Island, Okinawa Island) (Fig. 47) (Horvath 1912; Takeya 1931, 1963; Yamada and Tomokuni 2012; Yamada and Ishikawa 2016; present study). Judging from the description and illustration provided by Jing (1981), Chinese individuals identified as Stephanitis (Norba) mendica differ from Japanese specimens in the structure of the pronotum, suggesting that they pertain to another species. Stephanitis (N.) mendica inhabits the laurilignosa in a temperate climate of Japan proper (Honshu, Shikoku and Kyushu) and its surrounding islands, which are in the Palaearctic Region.	en	Souma, Jun (2022): Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan. Deutsche Entomologische Zeitschrift 69 (2): 219-281, DOI: http://dx.doi.org/10.3897/dez.69.89864, URL: http://dx.doi.org/10.3897/dez.69.89864
6D10E055B4255DE3AC49B8540B033EFB.taxon	biology_ecology	Biology. Stephanitis (Norba) mendica feeds on the abaxial surface of leaves of Cinnamomum yabunikkei (present study). This lace bug occurs only around " Tsuyu " (rainy season in Japan) (present study) and seems to be univoltine; adults were collected from April to August (Yamada and Tomokuni 2012; Okochi 2019; Souma 2021 d; present study); nymphs were collected in May and June (present study); the overwintering stage is unknown.	en	Souma, Jun (2022): Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan. Deutsche Entomologische Zeitschrift 69 (2): 219-281, DOI: http://dx.doi.org/10.3897/dez.69.89864, URL: http://dx.doi.org/10.3897/dez.69.89864
4B04CE33F5E15588B8763093F75F6561.taxon	description	[Japanese name: Yamakobashi-gunbai] Figs 2 A, 4 A, 7 A, 9 A, 11 A, 13 A, 15 A, 17 A, 19 A, 21 A, 23 A, 25 A, 27 A, 29 A, 31 A, 33, 40 A-C	en	Souma, Jun (2022): Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan. Deutsche Entomologische Zeitschrift 69 (2): 219-281, DOI: http://dx.doi.org/10.3897/dez.69.89864, URL: http://dx.doi.org/10.3897/dez.69.89864
4B04CE33F5E15588B8763093F75F6561.taxon	materials_examined	Material examined. Non-types collected at type locality (36 ♂♂ 34 ♀♀ 2 nymphs), JAPAN: Honshu: Kanagawa-ken, Sagamihara-Shi, Midori-ku, Yoshino (approximate coordinates: 35 ° 37 ' 54.8 " N, 139 ° 10 ' 11.0 " E), 9. viii. 2017, leg. J. Souma (2 ♂♂, ELKU; 30 ♂♂ 28 ♀♀, TUA); Kanagawa-ken, Sagamihara-Shi, Midori-ku, Naramoto For. Rd. (approximate coordinates: 35 ° 37 ' 48.6 " N, 139 ° 09 ' 55.5 " E), 4. ix. 2020, leg. J. Souma (2 ♂♂ 2 ♀♀, ELKU; 2 ♂♂ 2 ♀♀ 2 nymphs, TUA). 64 adult individuals recorded above, matching the original description of the species (Horvath 1912), were collected from the type locality, " Kanagawa " [= Honshu, Kanagawa-ken (a prefecture) or Kanagawa-ku (a ward) of Yokohama-shi (a city) of Kanagawa-ken]. Identification of S. (S.) ambigua in the present work is mainly based on these 64 adults. A total of 8 - 10 syntypes of S. (S.) ambigua exist in the collection of HNHM (D. Redei, pers. comm. 2021). Non-types (26 ♂♂ 22 ♀♀ 1 abdomen missing 2 nymphs), JAPAN: Honshu: " 群馬 " [= Gunma-ken (approximate coordinates: 36 ° 32 ' 43.2 " N, 139 ° 01 ' 07.5 " E)], " 20 / 5 / 914 " [= 20. v. 1914], " 武井氏 " [= leg. Takei] (2 ♂♂ 2 ♀♀, ELHU) (Fig. 33); Nagano, Ohya, 8. viii. 1959, leg. S. Miyamoto (15 ♂♂ 10 ♀♀ 1 abdomen missing 2 nymphs, KUM); Nagano, Onasawa, 21. viii. 1962, leg. Y. Miyatake (5 ♂♂, KUM); Nagano, Arayasu, 7. vii. 1966, leg. Y. Miyatake (4 ♂♂ 9 ♀♀, KUM). Shikoku: Kagawa-ken, Takamatsu-shi, Nishiueta-cho, Donguri Land, Fujio Shrine, 4. vi. 2020, leg. Y. Waki (2 ♀♀, TUA). Kyushu: Bungo, Sobosan, 20. vii. 1930, leg. Fujino & Yasumatsu (1 ♀, ELKU). Four specimens deposited in ELHU were labeled with an inscription of " Matsumura " (deposited in Matsumura's collection).	en	Souma, Jun (2022): Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan. Deutsche Entomologische Zeitschrift 69 (2): 219-281, DOI: http://dx.doi.org/10.3897/dez.69.89864, URL: http://dx.doi.org/10.3897/dez.69.89864
4B04CE33F5E15588B8763093F75F6561.taxon	diagnosis	Diagnosis. Stephanitis (Stephanitis) ambigua is recognised amongst other species of Stephanitis by a combination of the following characters: head, pronotal disc, marking on hemelytra and ventral in various shades of brown (Figs 7 A, 9 A, 11 A, 13 A, 15 A, 17 A, 19 A, 21 A, 23 A); calli light brown; body 1.8 times as long as maximum width across hemelytra (Figs 2 A, 4 A); rostrum reaching metasternum; pronotum tricarinate (Fig. 25 A); hood pale, shorter than median carina of pronotum, wider than maximum width of head across eyes, completely covering eye, slightly higher than median carina of pronotum at highest part, with posterior margin extending to middle of pronotal disc; median carina of pronotum with 3 rows of areolae at highest part; pronotal disc opaque; paranotum more erect, not narrowed posteriorly, with 4 rows of areolae at widest part, with anterolateral angle protruding anteriad, with outer margin angularly curved inward at posterolateral angle, maximum height longer than height of eye (Fig. 27 A); apices of hemelytra separated from each other in rest; costal area with 5 rows of areolae at widest part; subcostal area with 2 rows of areolae at widest part; discoidal area with 4 rows of areolae at widest part; sutural area with 3 rows of areolae at widest part; hypocostal lamina with a single row of areolae throughout its length; R + M (radiomedial) vein carinate; pygophore flat at centre of venter, with posterior margin emarginate in middle part (Fig. 29 A); and paramere stout, weakly curved inwards at apex, with outer margin sinuate in middle part, inner margin not curved in basal part (Fig. 31 A).	en	Souma, Jun (2022): Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan. Deutsche Entomologische Zeitschrift 69 (2): 219-281, DOI: http://dx.doi.org/10.3897/dez.69.89864, URL: http://dx.doi.org/10.3897/dez.69.89864
4B04CE33F5E15588B8763093F75F6561.taxon	distribution	Distribution. Japan (Honshu; Shikoku; Kyushu) (Fig. 45); China; Korea (Esaki and Takeya 1931; Takeya 1932, 1963; Drake 1938, 1948; Yamada and Tomokuni 2012; Yamada and Ishikawa 2016). A record from Taiwan (Drake 1948) does not list any examined specimen and appears to be erroneous. In Japan, Stephanitis (Stephanitis) ambigua inhabits deciduous broad-leaved forests in the temperate climatic zone of Japan proper (pertaining to the Palaearctic Region).	en	Souma, Jun (2022): Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan. Deutsche Entomologische Zeitschrift 69 (2): 219-281, DOI: http://dx.doi.org/10.3897/dez.69.89864, URL: http://dx.doi.org/10.3897/dez.69.89864
4B04CE33F5E15588B8763093F75F6561.taxon	biology_ecology	Biology. Stephanitis (Stephanitis) ambigua feeds on the abaxial surface of leaves of the three host plants in Japan (present study). In Japan, adults were collected in almost all seasons (Takeya 1953; Yamada and Tomokuni 2012; Yano et al. 2013; Maehara 2014; present study), whereas nymphs were collected in August and September (present study). The overwintering stage is the adult (Maehara 2014). One of the natural enemies of this lace bug is Stethoconus japonicus Schumacher, 1917 (Hemiptera, Heteroptera, Miridae) (Maehara 2014).	en	Souma, Jun (2022): Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan. Deutsche Entomologische Zeitschrift 69 (2): 219-281, DOI: http://dx.doi.org/10.3897/dez.69.89864, URL: http://dx.doi.org/10.3897/dez.69.89864
6FDD30BF5F045A71AEC66BC6030C719E.taxon	description	[Japanese name: Kusu-gunbai] Figs 3 C, D, 5 C, D, 8 C, D, 10 C-E, 12 C, D, 14 C-E, 16 C, D, 18 C, D, 20 B, 22 B, 24 B, 26 C, D, 28 D, 30 E, F, 32 E, F, 37 A, 38, 42 C-G	en	Souma, Jun (2022): Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan. Deutsche Entomologische Zeitschrift 69 (2): 219-281, DOI: http://dx.doi.org/10.3897/dez.69.89864, URL: http://dx.doi.org/10.3897/dez.69.89864
6FDD30BF5F045A71AEC66BC6030C719E.taxon	materials_examined	Material examined. Non-types collected at type locality of S. (S.) tabidula Horvath, 1912 (77 ♂♂ 69 ♀♀ 8 nymphs), JAPAN: Honshu: Kanagawa-ken, Sagamihara-shi, Chuo-ku, Tanashioda (approximate coordinates: 35 ° 32 ' 03.6 " N, 139 ° 20 ' 57.7 " E), 19. v. 2019, leg. J. Souma (43 ♂♂ 22 ♀♀ 8 nymphs, TUA); as above but 25. v. 2019 (4 ♂♂, ELKU; 22 ♂♂ 31 ♀♀, TUA); as above but 25. ii. 2020, leg. J. Souma (1 ♂ 12 ♀♀, TUA); as above but 17. xi. 2021 (3 ♂♂ 3 ♀♀, TUA); " Mt. Ohokusu " [= Kanagawa-ken, Yokosuka-shi, Ashina, Mt. Ogusu (approximate coordinates: 35 ° 15 ' 00.8 " N, 139 ° 37 ' 40.6 " E)], 7. viii. 1973, leg. K. B. S. (1 ♂, KPMNH); " 三崎 " [= Kanagawa-ken, Miura-shi, Misaki (approximate coordinates: 35 ° 08 ' 31.5 " N, 139 ° 36 ' 55.1 " E)], " 21 - 18 / 4 / 1911 " [= 18 - 21. iv. 1911], " Matsumra " (collected by Shonen Matsumura and / or deposited in Matsumura's collection) (1 ♂ 1 ♀, ELHU) (Fig. 38); Kanagawa-ken, Oiso-machi, Terasaka, Nagayama, 9. vi. 1991, leg. M. Enju (1 ♂, TUA); Kanagawa Pref., Ashigarashimo-gun, Manazuru-hantou, 11. xi. 2000, leg. S. Nagashima (1 ♂, TUA). The type locality of Stephanitis (Stephanitis) tabidula is " Kanagawa " [= Honshu, Kanagawa-ken (a prefecture) or Kanagawa-ku (a ward) of Yokohama-shi (a city) of Kanagawa-ken] and the 146 adult individuals recorded above match the original description of the species (Horvath 1912). The author identified S. (S.) tabidula based on the 146 adults in the present study. Paratypes of S. (S.) fasciicarina Takeya, 1931 collected at type locality (40 ♂ 45 ♀♀, ELKU; 1 ♀, KUM) (Fig. 37 A), JAPAN: Kyushu: " Chikuzen Akama " [= Fukuoka-ken, Munakata-shi, Akama (approximate coordinates: 33 ° 48 ' 26.1 " N, 130 ° 35 ' 31.2 " E)], 31. v. 1931, leg. C. Takeya. The male holotype of S. (S.) fasciicarina is currently missing (Souma 2021 b). However, the labels shown in Fig. 37 A were created after the original description (T. Mita, pers. comm. 2021). As mentioned in the section below, some of the non-types were labelled as " paratype ". Therefore, since labelling errors are suspected to have occurred for some of the type specimens, one of the 40 male paratypes collected at the type locality recorded above may correspond to the holotype. Paratypes of S. (S.) fasciicarina (59 ♂♂ 46 ♀♀, ELKU), JAPAN: Honshu: " Nagato Shimonoseki " [= Yamaguchi-ken, Shimonoseki-shi (approximate coordinates: 33 ° 57 ' 09.1 " N, 130 ° 55 ' 16.8 " E)], 22. viii. 1930, leg. K. Yasumatsu (8 ♂♂ 5 ♀♀). Kyushu: " Fukuoka " [= Fukuoka-ken (a prefecture) or Fukuoka-shi (a city) of Fukuoka-ken (approximate coordinates: 33 ° 35 ' 06.7 " N, 130 ° 22 ' 43.4 " E)], 14. v. 1930, leg. C. Takeya (7 ♂♂ 1 ♀♀); as above but 21. v. 1930 (6 ♂♂ 7 ♀♀); as above, but 22. v. 1930 (21 ♂♂ 14 ♀♀); as above, but 30. vi. 1930 (1 ♂ 1 ♀); as above but 7. vii. 1930 (16 ♂♂ 18 ♀♀). Although the labels of the 105 specimens were created after (T. Mita, pers. comm. 2021), their locality data match that of the original description of S. (S.) fasciicarina (Takeya 1931). Non-types collected at type locality of S. (S.) fasciicarina (4 ♀♀, TUA), JAPAN: Kyushu: Fukuoka-ken, Munakata-shi, Akama (approximate coordinates: 33 ° 48 ' 26.1 " N, 130 ° 35 ' 31.2 " E), 26. ix. 2021, leg. J. Souma. Non-types collected at type locality of S. (S.) kyushuana Drake, 1948 (1 ♀, TUA), JAPAN: Kyushu: Fukuoka-ken, Kitakyushu-shi, Moji-ku, Motokiyotaki (approximate coordinates: 33 ° 56 ' 18.9 " N, 130 ° 57 ' 42.7 " E), 16. ix. 2022, leg. J. Souma. Non-types (518 ♂♂ 653 ♀♀ 60 nymphs), JAPAN: Honshu: Miyagi-ken, Oshika-gun, Onagawa-cho, Kirigasaki, 20. ix. 2020, leg. H. Konno (2 ♂♂ 3 ♀♀, TUA); Miyagi-ken, Miyagi-gun, Rifu-cho, Akanuma, Tanbazawa, 22. ix. 2020, leg. H. Konno (1 ♀, TUA); Ibaraki, Choshi, 15. vi. 1981, leg. M. Miyazaki (13 ♂♂ 24 ♀♀ 1 nymph, NIAES); Tochigi Pref., Uchino, Watarase-yusuichi, 29. v. 2001, leg. S. Maehara (2 ♂♂ 1 ♀, TUA); as above but 5. xi. 2012 (1 ♂, TUA); Tochigi, Simotsuke City, Motoyoshida, 12. x. 2013, leg. S. Maehara (2 ♀♀, TUA); as above, but 28. xi. 2016 (1 ♂ 2 ♀♀); as above but 12. xi. 2019 (1 ♂ 2 ♀♀, TUA); as above, but 15. x. 2020 (4 ♂♂ 1 ♀, TUA); Saitama Pref., Nihongi Pass, 3. vii. 1984, leg. M. Hayashi et al. (1 ♂, TUA); Saitama-ken, Iruma-gun, Moroyama-machi, Takinoiri, Shobo-goya, 28. v. 2021, leg. J. Souma (2 ♂♂ 4 ♀♀, TUA); Saitama-ken, Iruma-gun, Moroyama-machi, Takinoiri, Kanikusa-hashi, 28. v. 2021, leg. J. Souma (8 ♂♂ 9 ♀♀, TUA); Tokyo, 22. viii. 1949, leg. R. Takahashi (1 ♂ 1 ♀, ELKU); Chiba, Tateyama, Susaki, 11. x. 2001, leg. M. Tomokuni (5 ♂♂ 5 ♀♀, NSMT); Tokyo, Kiyosumi-koen, 24. vi. 1972, leg. H. Hasegawa (11 ♂♂ 22 ♀♀, NIAES); Niigata-ken, Kitakanbara-gu, Seiro-machi, Mano, 14 - 15. vi. 2013, leg. K. Nakano (1 ♂, TUA); Niigata-ken, Murakami-shi, Kashio, 13. vi. 2015, leg. K. Nakano (1 ♂ 4 ♀♀, TUA); as above but 1. x. 2016, leg. K. Nakano (1 ♂ 4 ♀♀, TUA); Niigata-ken, Niigata-shi, Nishi-ku, Igarashi- 2 no-cho, 19. vii. 2015, leg. K. Nakano (1 ♀, TUA); Niigata-ken, Kariwa-gun, Kariwa-mura, Takiya, 13. vi. 2021, leg. G. Mashima (1 ♂ 1 ♀, TUA); Niigata-ken, Kashiwazaki-shi, Miyagawa, 13. vi. 2021, leg. G. Mashima (2 ♂♂, TUA); Ishikawa, Sekidoyama, 2. viii. 1960, leg. T. Hidaka (1 ♀, ELKU); Fukui Pref., Otomi, 12. vii. 1981, leg. O. Kishimoto (1 ♀, KUM); Fukui Pref., Mt. Aoba, 20. ix. 1981, leg. O. Kishimoto (1 ♀, KUM); Mino, Utsumi, 21. x. 1952, leg. Yasumatu (1 ♂ 1 ♀, ELKU); Izu, Kuren, 16. x. 1952, leg. Yasumatsu (3 ♀♀, ELKU); Izu, Suzaki, 30. ix. 1980, leg. M. Tomokuni (7 ♂♂ 7 ♀♀, NSMT); Izu, Ohno-yama, nr. Ohsawa, 1. x. 1980, leg. M. Tomokuni (3 ♀♀, NSMT); Shizuoka-ken, Hamamatsu-shi, Nishi-ku, Kamigaya-cho, 16. vi. 2017, leg. J. Souma (1 ♀, TUA); Aichi-ken, Kitashitara-gun, Shitara-cho, Taguchi, 17. vi. 2017, leg. J. Souma (1 ♀, TUA); Mie Pref., Oosugidani, 11. viii. 1970, leg. M. Tomokuni (1 ♂, NSMT); Mie Pref., Ise-shi, Shimogu, 1. xi. 2000, leg. T. Ishikawa (1 ♀, TUA); Kyoto Pref., Kurama-Kibune, 12. vii. 1970, leg. M. Tomokuni (1 ♀, NSMT); Osaka Pref., Higashi-Sumiyoshi, Nagai, 2. vi. 1985, leg. T. Kishimoto (1 ♀, TUA); Kobe, 29. v. 1941, leg. Kurosa (1 ♂, ELKU); Tajima, Sekinomiya, 8. vii. 1952, leg. R. Morimoto (7 ♂♂ 6 ♀♀, ELKU); Hyogo-ken, Kobe-shi, Suma-ku, Higashisuma, 8. x. 2022, leg. J. Souma (3 ♂♂ 3 ♀♀, TUA); Yamato, Nara, 14. x. 1951, leg. C. Takeya (3 ♂♂ 3 ♀♀, ELKU); as above but 16. x. 1951 (23 ♂♂ 30 ♀♀, ELKU; 1 ♂ 1 ♀, KUM); Prov. Kii, Wakayama, 15. x. 1951, leg. S. Goto (2 ♀♀, ELKU); Wakayama, Nachi-Irokawa, 2. ix. 1998, leg. leg. S. Gotoh (1 ♂, NSMT); Okayama-ken, Okayama-shi, Kita-ku, Heidan, 20. v. 2022, leg. J. Souma (1 ♀, TUA); Yamaguchi-ken, Yamaguchi-shi, Atou, Shinome, 25. x. 2008, leg. M. Hayashi (1 ♀, TUA); Yamaguchi-ken, Shimonoseki-shi, Hon-machi, 24. ix. 2022, leg. J. Souma (6 ♂♂ 3 ♀♀, TUA). Tashiro Island: Shikishima, near Manga Island, 10. xi. 2019, leg. H. Konno (1 ♂ 1 ♀, TUA). Sado Island: Kitaikari, 26. viii. 2021, leg. J. Souma (2 ♂♂ 1 ♀, TUA); Kubota, 27. viii. 2021, leg. J. Souma (2 ♀♀, TUA); Ishida, 27. viii. 2021, leg. J. Souma (2 ♂♂ 8 ♀♀, TUA); Anyoji, 28. viii. 2021, leg. J. Souma (1 ♀, TUA); Kujikawachi, 29. viii. 2021, leg. J. Souma (1 ♀, TUA); Kawasaki, 29. viii. 2021, leg. J. Souma (3 ♂♂ 4 ♀♀, TUA). Awa Island: Uramura, Uchiura, 3. x. 2021, leg. J. Souma (9 ♂♂ 7 ♀♀, TUA). Izu Islands (northern part): Izu-Oshima Island: Kasamatsu, 4. x. 2002, leg. M. Tomokuni (1 ♂♂ 2 ♀♀, NSMT). Awaji Island: Fukura, 29. ix. 1972, leg. M. Tomokuni & M. Sakai (1 ♀, NSMT); Mt. Mikuma, 30. ix. 1972, leg. M. Tomokuni & M. Sakai (1 ♀, NSMT). Oki Islands: Dogo Island: Saigo-cho, Mt. Daimanji, 12. ix. 1984, leg. M. Tomokuni (3 ♀♀, NSMT). Shikoku: Tokushima-ken, Tokushima-shi, Bizan-cho, Mosukegahara, 2. x. 2022, leg. J. Souma (1 ♂ 1 ♀, TUA); " Sanuki Wada-mura " [= Kagawa-ken, Kannonji-shi, Toyohama-cho, Wada], 10. vi. 1930, leg. M. Hanada (2 ♂ 3 ♀♀, ELKU; 1 ♀, KUM); Tosa, Yamakita-mura, 10. xi. 1951, leg. K. Yasumatsu (3 ♂♂ 2 ♀, ELKU); Ehime P., Matsuyama C., 25. x. 1972, leg. H. Hasegawa (11 ♂♂ 7 ♀♀ 4 nymphs, NIAES). Kyushu: as paratype collected at type locality (1 nymph, ELKU); Fukuoka, Hirao, 22. x. 1931, leg. Fujino & Hashimoto (1 ♀, ELKU); as above but 16. x. 1932, leg. T. Shrozu (1 ♂ 1 ♀, ELKU); as above but 3. vi. 1951, leg. Matsuda (1 ♀, ELKU); as above but 21. vi. 1957, leg. Y. Miyatake (1 ♀, KUM); Chikuzen, Inunakiyama, 25. x. 1931, leg. K. Yasumatsu (1 ♀, KUM); as above, but 15. vi. 1969, leg. S. Miyamoto (1 ♀, KUM); Chikuzen, Tsutsugatake, 6. xii. 1931, leg. C. Takeya (1 ♂ 14 ♀♀, ELKU); Tikuzen, Kyudai-Kasuya-Ensyurin, 20 - 22. vi. 1944, leg. S. Ito (1 ♀, ELKU); Buzen, Hikosan, 12. vii. 1948, leg. S. Miyamoto (1 ♀, KUM); as above, but 5. viii. 1951, leg. C. Takeya (5 ♂♂ 9 ♀♀, ELKU); as above, but 6. viii. 1951, leg. C. Takeya (8 ♂♂ 15 ♀♀, ELKU); as above but 15. vii. 1958, leg. Y. Miyatake (2 ♂♂ 1 ♀, KUM); as above, but 6. vi. 1959, leg. Y. Miyatake (1 ♂, KUM); as above but 7. vi. 1959, leg. Y. Miyatake (1 ♀, KUM); as above, but 14. vi. 1959 (3 ♀♀, KUM); as above, but 14. vi. 1959, leg. K. Yasumatsu (1 ♀, ELKU); Chikugo, Korasan, 25. vii. 1951, leg. C. Takeya (3 ♂♂ 3 ♀♀, ELKU); as above, but 7. viii. 1951, leg. S. Miyamoto (1 ♂ 1 ♀, ELKU); as above, but 1. ix. 1951, leg. C. Takeya (1 ♂ 3 ♀♀, ELKU); as above but 26. v. 1953 (4 ♂♂ 5 ♀♀, ELKU); Fukuoka, Fukuoka, Korasan, 25. vii. 1951, leg. S. Miyamoto (1 ♂ 3 ♀♀, KUM); as above but 7. viii. 1951 (2 ♀♀, KUM); as above, but 11. vi. 1961 (3 ♀, KUM); Chikuzen, Fukuoka, 27. vii. 1951, leg. C. Takeya (6 ♂♂ 7 ♀♀, ELKU); Chikuzen, Narutake-yama, 19. viii. 1951, leg. Y. Hirashima (1 ♀, ELKU); as above but 8. vi. 1961, leg. C. Takeya (1 ♀, ELKU); Fukuoka, Mt. Kumado, 23. viii. 1952, leg. S. Miyamoto (4 ♂♂ 3 ♀♀, ELKU; 9 ♂♂ 6 ♀♀ 5 nymphs, KUM); Aburayama, 28. viii. 1952, leg. C. Takeya (1 ♀, ELKU); as above, but 5. v. 1961, leg. S. Miyamoto (5 ♂♂ 9 ♀♀ 17 nymphs, KUM); as above, but 11. v. 1961, leg. S. Miyamoto (16 ♂♂ 11 ♀♀, KUM); as above, but 4. vi. 1961, leg. S. Miyamoto (1 ♀, KUM); Fukuoka, Magarifuchi, 23. ix. 1952, leg. C. Takeya (6 ♂♂ 4 ♀♀, ELKU); as above, but 16. vii. 1961, leg. S. Miyamoto (3 ♂♂ 4 ♀♀, KUM); Chikuzen, Mt. Kosho, 14. vi. 1953, leg. C. Takeya (1 ♀, ELKU); Chikugo, Himeharu, 3. vi. 1954, leg. Gyotoku (3 ♂♂ 3 ♀♀, ELKU); Fukuoka, Sasayama, ix. 1954, leg. S. Miyamoto (1 ♂ 2 ♀♀, KUM); Fukuoka, Gokoku Shrine, 5. ix. 1959, leg. S. Miyamoto (39 ♂♂ 36 ♀♀, KUM); as above, but 24. vii. 1961 (9 ♂♂ 8 ♀♀, KUM); Fukuoka, Kashii, 6. ix. 1959, leg. S. Miyamoto (1 ♀, KUM); as above, but 2. xi. 1961 (1 ♂, KUM); Fukuoka, Nogochi, 16. vi. 1961, leg. S. Miyamoto (13 ♂♂ 9 ♀♀ 3 nymphs, KUM); Fukuoka, Tachibanayama, 23. vii. 1961, leg. S. Miyamoto (2 ♂♂ 7 ♀♀, KUM); Fukuoka, 24. vii. 1961, leg. S. Miyamoto (2 ♂♂ 6 ♀♀, KUM); Fukuoka, Atagoyama, 4. viii. 1961, leg. S. Miyamoto (1 ♂ 1 ♀, KUM); Fukuoka, Hakozaki, Kyushu University, 12. viii. 1961, leg. S. Miyamoto (134 ♂♂ 124 ♀♀ 14 nymphs, KUM); Fukuoka, Kanetake, 13. ix. 1969, leg. S. Miyamoto (5 ♂♂ 12 nymphs, KUM); Fukuoka, Kamado Shrine, 27. ix. 1977, leg. S. Miyamoto (2 ♂♂, KUM); as above, but 5. xi. 1977 (1 ♀, KUM); Fukuoka, Tenmangu, 6. x. 1977, leg. S. Miyamoto (1 ♀, KUM); Fukuoka-ken, Munakata-shi, Yamada, 28. viii. 2017, leg. J. Souma (1 ♀, TUA); Fukuoka-ken, Fukuoka-shi, Nishi-ku, Motooka, Kyushu University, 23. v. 2020, leg. J. Souma (2 ♂♂ 1 ♀, ELKU); as above, but 21. ix. 2020 (3 ♂♂ 1 ♀, ELKU); Fukuoka-ken, Fukuoka-shi, Nishi-ku, Kusaba, 24. v. 2020, leg. J. Souma (1 ♂, ELKU); Fukuoka-ken, Itoshima-shi, Shimanokita, 24. v. 2020, leg. J. Souma (14 ♂♂ 28 ♀♀, ELKU); Fukuoka-ken, Fukuoka-shi, Sawara-ku, Itaya, Mt. Sefuri, 30. v. 2020, leg. S. Yagi (1 ♀, ELKU); Fukuoka-ken, Dazaifu-shi, Kitadani, Mt. Homan, 4. vi. 2020, leg. J. Souma (7 ♂♂ 11 ♀♀, ELKU; 1 ♂ 4 ♀♀, TUA); Fukuoka-ken, Kasuya-gun, Sasaguri-machi, Tsubakuro, 5. vi. 2020, leg. J. Souma (2 ♂♂ 1 ♀, TUA); Fukuoka-ken, Itoshima-shi, Nijofukui, Nijo Forest Park, 20. vi. 2020, leg. S. Chuman (4 ♂♂ 2 ♀♀, TUA); Fukuoka-ken, Tagawa-gun, Soeda-machi, Mt. Hikosan, 21. vi. 2020, leg. J. Souma (3 ♀♀, TUA); as above but 20. vi. 2020, leg. S. Yagi (1 ♀, ELKU); Fukuoka-ken, Itoshima-shi, Tomari, 29. v. 2022, leg. J. Souma (1 ♀, ELKU); as above, but 10. vii. 2022, leg. J. Souma (1 ♂, TUA); Saga Pref., Fujitsu-gun, Tara-cho, Oura, Ushioro, alt. 100 m, 12. vii. 2021, leg. M. Nishida (1 ♂, ELKU); Hizen, Kunimiyama, 16. vi. 1950, leg. T. Shrozu (1 ♂ 2 ♀♀, ELKU); Higo, Yatsushiro, 8. x. 1953, leg. C. Takeya (1 ♀, ELKU); Kumamoto, Yuyama, Mt. Ichifusa, 5. viii. 1985, leg. M. Miyazaki (3 ♀♀, NIAES); as above, but 6. viii. 1985 (2 ♂♂ 4 ♀♀, NIAES); Bungo, Tsukumi, 29. vii. 1951, leg. R. Matsuda (2 ♂♂ 1 ♀, ELKU); Bungo, Hachiya, 11. x. 1951, leg. S. Nakao (1 ♀, KUM); Oita Pref., Saiki-shi, Ume, Minamitabaru, Takadoya-jinja, 26. v. 2019, leg. R. Ito (1 ♂, TUA); Oita Pref., Saeki-shi, Ume, Kitagawa dam, 25 - 26. v. 2019, leg. N. Tsuji & S. Imada (1 ♀, TUA); Hyuga, Takaoka, 25. viii. 1952, leg. A. Ema (6 ♂♂ 10 ♀♀, ELKU); Hyuga, Mt. Kirishima, 12. ix. 1952, leg. Nakao & Ogata (1 ♂, ELKU); Miyazaki Pref., Hinokage-cho, Mt. Tansuke-dake, 1. vi. 2019, leg. R. Ito (1 ♀, TUA); Miyazaki-ken, Higashimorokata-gun, Aya-cho, Takeno, Omoridake For. Rd., 8. x. 2021, leg. T. Saeki (1 ♂, ELKU); Kagoshima Pref., Mianmiosumi-cho, Sata, Hetsuka, 30. v. 2020, leg. R. Ito (1 ♀, TUA). Nokonoshima Island: 27. vi. 1987, leg. S. Miyamoto (1 ♀, KUM). Tsushima Island: Izuhara-Sasutoge, 7. vi. 1941, leg. T. Shirozu (1 ♀, ELKU); Izuhara, 31. x. 1962, leg. S. Miyamoto (1 ♀, ELKU; 1 ♂, KUM); Mt. Ariake, 12. vii. 1968, leg. S. Miyamoto & A. Nakanishi (1 ♀, KUM); Observatory, 28. viii. 1988, leg. S. + K. M. (1 ♀, KUM); Kuda, 29. viii. 1988, leg. S. + K. M. (1 ♂ 1 ♀, KUM); Kamiagata-cho, Sago, 6. ix. 2017, leg. J. Souma (5 ♂♂ 4 ♀♀, TUA); as above but 8. ix. 2017 (3 ♀♀, TUA); Izuhara-cho, Tsutsu, Mount Tatera, 7. ix. 2017, leg. J. Souma (1 ♂, TUA); Kamiagata-machi, near Mt. Konoki-yama, 8. ix. 2017, leg. T. Ishikawa (11 ♂♂ 8 ♀♀ 3 nymphs, TUA); Mitsushima-machi, Kechi, 8. ix. 2017, leg. J. Souma (12 ♂♂ 17 ♀♀, TUA). Iki Island: Gonoura-cho, Katabaruhure, Takenotsuji, 16 - 18. vii. 2016, leg. R. Ito (1 ♂, TUA). Amakusa Islands: Shimoshima Island: " Kakuyama " [= Miyajidake-machi, Mt. Kakuyama], 18. ix. 1931, leg. H. Hori (1 ♂ 1 ♀, ELKU). Goto Islands: Fukue Island: Miiraku-Kahara, 4. viii. 1933, leg. T. Shirozu (2 ♀, KUM). Ryukyu Islands (northern part): Yakushima Island: Kosugidani, 23. viii. 1952, leg. C. Takeya & Y. Hirashima (2 ♂♂ 5 ♀♀, ELKU); Koseda, 21. ix. 2013, leg. N. Tsuji (1 ♀, ELKU). Eight specimens from " Sanuki Wada-mura " and " Kakuyama " were labelled as " paratype " of S. (S.) fasciicarina. However, their labels were created subsequently (T. Mita, pers. comm. 2021) and the locality data of these eight individuals do not match those of the original description of S. (S.) fasciicarina (Takeya 1931). Therefore, the eight specimens do not seem to have paratype status. Four specimens from " Tachibanayama " were identified as " S. (N.) aperta possessing lateral carina " in a previous study (Takeya 1963).	en	Souma, Jun (2022): Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan. Deutsche Entomologische Zeitschrift 69 (2): 219-281, DOI: http://dx.doi.org/10.3897/dez.69.89864, URL: http://dx.doi.org/10.3897/dez.69.89864
6FDD30BF5F045A71AEC66BC6030C719E.taxon	diagnosis	Diagnosis. Stephanitis (Stephanitis) tabidula is recognised amongst other species of Stephanitis by a combination of the following characters: head, calli, pronotal disc, marking on hemelytra and ventral surface in various shades of brown (Figs 8 C, D, 10 C-E, 12 C, D, 14 C-E, 16 C, D, 18 C, D, 20 B, 22 B, 24 B); body in male 2.1 times (in female 2.0 times) as long as maximum width across hemelytra (Figs 3 C, D, 5 C, D); rostrum not reaching metasternum; pronotum unicarinate or tricarinate (Fig. 26 C, D); hood pale, shorter than median carina of pronotum, as wide as or wider than vertex at widest part, not or incompletely covering eye, as high as median carina of pronotum at highest part, with posterior margin not extending to middle of pronotal disc; median carina of pronotum with 2 rows of areolae at highest part; pronotal disc opaque; paranotum more erect, slightly narrowed posteriorly, with 3 rows of areolae at widest part, anterolateral angle protruding anteriad, outer margin gently curved inwards at posterolateral angle, maximum height longer than height of eye (Fig. 28 D); apices of hemelytra close to each other in rest; costal area with 3 - 5 rows of areolae at widest part; subcostal area in male with 2 rows (in female with 2 - 3 rows) of areolae at widest part; discoidal area with 3 - 4 rows of areolae at widest part; sutural area with 3 - 4 rows of areolae at widest part; hypocostal lamina with a single row of areolae throughout its length; R + M (radiomedial) vein carinate; pygophore elevated at centre of venter, with posterior margin slightly emarginate in middle part (Fig. 30 E, F); and paramere stout, strongly curved inwards at apex, with outer margin not sinuate in middle part, inner margin nearly straight in basal part (Fig. 32 E, F).	en	Souma, Jun (2022): Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan. Deutsche Entomologische Zeitschrift 69 (2): 219-281, DOI: http://dx.doi.org/10.3897/dez.69.89864, URL: http://dx.doi.org/10.3897/dez.69.89864
6FDD30BF5F045A71AEC66BC6030C719E.taxon	distribution	Distribution. Japan (Honshu; Izu Islands (northern part): Izu-Oshima Island; Tashiro Island; Sado Island; Awa Island; Awaji Island; Oki Islands: Dogo Island; Shikoku; Kyushu; Nokonoshima Island; Tsushima Island; Iki Island; Amakusa Islands: Shimoshima Island; Goto Islands: Fukue Island; Ryukyu Islands (northern part): Yakushima Island) (Fig. 48); southern Korea (Horvath 1912; Takeya 1931, 1963; Takara and Hidaka 1960; Miyamoto 1976; Tomokuni 1985; Yasunaga et al. 1993; Tomokuni and Hayashi 2006; Yamada and Tomokuni 2012; Yamada and Ishikawa 2016; Ahn et al. 2018; present study). Judging from the description and illustration (Jing 1981), the Chinese population differs from the Japanese population in the structure of the pronotum and seems to correspond to another species. According to the present author's examination, the specimens previously recorded from Okinawa Island (the central part of the Ryukyu Islands) (Takara and Hidaka 1960) correspond to Stephanitis (Norba) mendica. Judging from the photographs and specimens, the previous records from the southern part of the Izu Islands (Takeya 1963; Tomokuni and Ishikawa 2002; Yamada and Tomokuni 2012) correspond to S. (S.) tomokunii sp. nov., described below. In Japan, S. (S.) tabidula inhabits the laurilignosa in a temperate climate of Japan proper (Honshu, Shikoku and Kyushu) and its surrounding islands and the northern part of Izu and Ryukyu Islands, which is located in the Palaearctic Region.	en	Souma, Jun (2022): Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan. Deutsche Entomologische Zeitschrift 69 (2): 219-281, DOI: http://dx.doi.org/10.3897/dez.69.89864, URL: http://dx.doi.org/10.3897/dez.69.89864
6FDD30BF5F045A71AEC66BC6030C719E.taxon	biology_ecology	Biology. Stephanitis (Stephanitis) tabidula feeds on the abaxial surface of leaves of the five host plants in Japan (present study). In Japan, adults were collected in almost all seasons (Takeya 1931; Miyamoto 1976; Tomokuni 1981, 1985; Yasunaga et al. 1993; Tomokuni and Hayashi 2006; Yano et al. 2013; Okochi 2019; present study); nymphs were collected from May to July and in September and October (present study); the overwintering stage is the adult (present study).	en	Souma, Jun (2022): Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan. Deutsche Entomologische Zeitschrift 69 (2): 219-281, DOI: http://dx.doi.org/10.3897/dez.69.89864, URL: http://dx.doi.org/10.3897/dez.69.89864
180143158D8F524CA21EDEE37007513E.taxon	description	[Japanese name: Tosaka-gunbai] Figs 3 E, 5 E, 6 B, 8 E, 10 F, 12 E, 14 F, 16 E, 18 E, 20 C, 22 C, 24 C, 26 E, 28 E, 30 G, 32 G, 37 B, 42 H-J	en	Souma, Jun (2022): Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan. Deutsche Entomologische Zeitschrift 69 (2): 219-281, DOI: http://dx.doi.org/10.3897/dez.69.89864, URL: http://dx.doi.org/10.3897/dez.69.89864
180143158D8F524CA21EDEE37007513E.taxon	materials_examined	Material examined. Lectotype (1 ♂, ELHU) (Fig. 37 B), JAPAN: Honshu: Gifu [= Honshu, Gifu-ken (approximate coordinates: 35 ° 26 ' 05.5 " N, 136 ° 46 ' 16.6 " E)]. Non-types (157 ♂♂ 169 ♀♀ 2 nymphs), JAPAN: Honshu: Saitama-ken, Iruma-gun, Moroyama-machi, Takinoiri, Kanikusa-hashi, 28. v. 2021, leg. J. Souma (3 ♀♀, TUA); Tokyo-to, Hachioji-shi, Uratakao-machi, Kogesawa-rindo, 6. ix. 2016, leg. J. Souma (11 ♂♂ 7 ♀♀, TUA); as above but 12. v. 2017 (1 ♂ 1 ♀, TUA); as above 22. v. 2017, leg. Y. Kato (1 ♀, TUA); Kanagawa-ken, Sagamihara-shi, Minami-ku, Asamizodai, 4. vi. 2017, leg. J. Souma (2 ♂♂ 1 ♀, TUA); Kanagawa-ken, Sagamihara-shi, Minami-ku, Shimomizo, 10. ix. 2020, leg. J. Souma (1 ♀, TUA); Kanagawa-ken, Atsugi-shi, Funako, 12. v. 2016, leg. J. Souma (1 ♂, TUA); as above but 7. iii. 2017 (1 ♀, TUA); as above, but 5. v. 2017 (23 ♂♂ 18 ♀♀, TUA); as above but 6. v. 2017 (20 ♂♂ 9 ♀♀, TUA); as above, but 7. v. 2017, leg. H. Shigetoh (1 ♂, TUA); as above but 11. v. 2017 (43 ♂♂ 52 ♀♀, TUA); as above, but 31. v. 2017 (4 ♀♀, ELKU); as above but 5. vi. 2017 (1 ♀, TUA); Kanagawa-ken, Atsugi-shi, Nanasawa, 6. vi. 2017, leg. J. Souma (8 ♂♂ 5 ♀♀, TUA); Kanagawa-ken, Isehara-shi, Oyama, 31. v. 2017, leg. Y. Yamada (1 ♂, TUA); Yamanashi-ken, Hokuto-shi, Takane-cho, Kiyosato, 5. ix. 2022, leg. J. Souma (1 ♂, TUA); Nagano-ken, Matsumoto-shi, Azumi, 7. ix. 2022, leg. J. Souma (2 ♂♂ 1 ♀, TUA); Hiroshima Pref., Kitahiroshima, Nakaso, 23. vii. 2022, leg. Y. Uehara (1 ♂, TUA); as above but leg. H. Hashimoto (1 ♂, TUA). Sado Island: Chigusa, 28. viii. 2021, leg. J. Souma (1 ♂ 2 ♀♀, TUA); Kamiyokoyama, 28. viii. 2021, leg. J. Souma (3 ♂♂ 5 ♀♀, TUA); Kujikawachi, 29. viii. 2021, leg. J. Souma (2 ♀♀, TUA); Noura, 29. viii. 2021, leg. J. Souma (3 ♀♀, TUA). Shikoku: Kagawa-ken, Takamatsu-shi, Nishiueta-cho, Fujio, 28. iv. 2020, leg. Y. Waki (1 ♂, TUA); Kagawa-ken, Takamatsu-shi, Nishiueta-cho, Donguri Land, 23. v. 2021, leg. Y. Waki (1 ♀, TUA); Kochi-ken, Kochi-shi, Shigekura, 2. vii. 2020, leg. J. Souma (2 ♂♂ 1 ♀ 2 nymphs, ELKU). Kyushu: Fukuoka-ken, Kasuya-gun, Sasaguri-machi, Tsubakuro, 5. vi. 2020, leg. N. Tsuji (5 ♂♂ 9 ♀♀, ELKU); Fukuoka Pref., Tagawa, Soeda, Mt. Hiko-san, Takanosubaru, 28. v. 2022, leg. Y. Uehara (1 ♂ 1 ♀, TUA); Oita-ken, Kusu-gun, Kokonoe-machi, 12. vii. 2017, leg. S. Imada (1 ♀, ELKU); Kumamoto Pref., Aso, Minamiaso, Kain, 12. vi. 2022, leg. H. Hashimoto (1 ♀, TUA); Kumamoto-ken, Kuma-gun, Mizukami-mura, Mt. Ichifusa-yama, 27. vii. 2022, leg. S. Inoue (1 ♀, TUA); Oita-ken, Hita-shi, Maetsue-machi, Ono, 21. viii. 2022, leg. J. Souma (1 ♂ 1 ♀, TUA); Oita-ken, Hita-shi, Maetsue-machi, Yugi, 21. viii. 2022, leg. J. Souma (1 ♂, TUA). Ryukyu Islands (northern part): Yakushima Island: Kosugidani, 23. viii. 1952, leg. C. Takeya & Y. Hirashima (2 ♂♂ 3 ♀♀, ELKU); as above, but 29. vii. 1963, leg. T. Okada (1 ♀, KUM); Hananoego, 24. viii. 1952, leg. C. Takeya & Y. Hirashima (17 ♂♂ 19 ♀♀, ELKU; 1 ♀, KUM); as above, but 26. x. 1979, leg. S. Makihara (4 ♂♂ 7 ♀♀, NSMT); Onoaida, 20. viii. 2021 (3 ♂♂ 5 ♀♀, ELKU).	en	Souma, Jun (2022): Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan. Deutsche Entomologische Zeitschrift 69 (2): 219-281, DOI: http://dx.doi.org/10.3897/dez.69.89864, URL: http://dx.doi.org/10.3897/dez.69.89864
180143158D8F524CA21EDEE37007513E.taxon	diagnosis	Diagnosis. Stephanitis (Stephanitis) takeyai is recognised amongst other species of Stephanitis by a combination of the following characters: head, calli, pronotal disc, marking on hemelytra and ventral surface black (Figs 8 E, 10 F, 12 E, 14 F, 16 E, 18 E, 20 C, 22 C, 24 C); body in male 2.1 times (in female 1.9 times) as long as maximum width across hemelytra (Figs 3 E, 5 E, 6 B); rostrum reaching metasternum; pronotum tricarinate (Fig. 26 E); hood dark, longer than median carina of pronotum, wider than maximum width of head across eyes, completely covering eye, higher than median carina of pronotum at highest part, with posterior margin extending near posterior margin of pronotal disc; median carina of pronotum with 2 rows of areolae at highest part; pronotal disc opaque; paranotum more erect, not narrowed posteriorly, 3 rows of areolae at widest part, anterolateral angle slightly protruding anteriad, outer margin gently curved inwards at posterolateral angle, maximum height longer than height of eye (Fig. 28 E); apices of hemelytra close to each other in rest; costal area with 3 - 4 rows of areolae at widest part; subcostal area in male with 3 rows (in female with 4 rows) of areolae at widest part; discoidal area with 3 - 4 rows of areolae at widest part; sutural area with 3 rows of areolae at widest part; hypocostal lamina with a single row of areolae throughout its length; R + M (radiomedial) vein carinate; pygophore elevated at centre of venter, with posterior margin slightly emarginate in middle part (Fig. 30 G); and paramere stout, weakly curved inwards at apex, with outer margin cuspidate in middle part, inner margin slightly curved inwards in basal part (Fig. 32 G).	en	Souma, Jun (2022): Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan. Deutsche Entomologische Zeitschrift 69 (2): 219-281, DOI: http://dx.doi.org/10.3897/dez.69.89864, URL: http://dx.doi.org/10.3897/dez.69.89864
180143158D8F524CA21EDEE37007513E.taxon	distribution	Distribution. Japan (Honshu; Sado Island; Oki Islands: Dogo Island; Shikoku; Kyushu; Ryukyu Islands (northern part): Yakushima Island) (Fig. 48); Austria; Belgium; Canada; Czech Republic; France; Great Britain; Germany; Hungary; Italy; Netherlands; Poland; Portugal; Slovakia; Switzerland; U. S. A. (Takeya 1963; Yamada and Tomokuni 2012; Vetek et al. 2012; Aukema et al. 2013; Barta and Biden 2016; Yamada and Ishikawa 2016; Grosso-Silva et al. 2020; present study). Stephanitis (Stephanitis) takeyai is native to Japan, but it has invaded many countries in Europe and North America (Yamada and Tomokuni 2012). The previous record from India (Drake and Ruhoff 1965) is an error resulting from the confusion between the type locality of Tingis globulifera Walker, 1873 [= Cochlochila (Physodictyon) bullita (Stal, 1873)] described from India and T. globulifera Matsumura, 1905 [= S. (S.) takeyai Drake & Maa, 1955] described from Japan (cf. Drake and Maa 1955). The previous records from Amami-Oshima Island, the central part of the Ryukyu Islands (Takeya 1963), are misidentifications of S. (S.) pyrioides and a nymph pertaining to another genus. In Japan, S. (S.) takeyai inhabits the deciduous broad-leaved forest in a temperate climate of Japan proper (Honshu, Shikoku and Kyushu) and its surrounding islands, which is in the Palaearctic Region.	en	Souma, Jun (2022): Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan. Deutsche Entomologische Zeitschrift 69 (2): 219-281, DOI: http://dx.doi.org/10.3897/dez.69.89864, URL: http://dx.doi.org/10.3897/dez.69.89864
180143158D8F524CA21EDEE37007513E.taxon	biology_ecology	Biology. Stephanitis (Stephanitis) takeyai feeds on the abaxial surface of leaves of the various host plants in Japan (present study). In Japan, this lace bug is trivoltine (Tsukada 1994, 2008); adults were collected in almost all seasons (Takeya 1953; Tomokuni 1981, 1985; Ichita 1989; Yasunaga et al. 1993; Tsukada 1994, 2008; Yamada and Tomokuni 2012; Yano et al. 2013; Maehara 2014; Ito and Sasaki 2018; present study); nymphs were collected from April to October (Tsukada 1994, 2008; Maehara 2014; present study); the overwintering stage is the egg, but third-generation adults are found until March of the following year (Tsukada 1994; present study).	en	Souma, Jun (2022): Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan. Deutsche Entomologische Zeitschrift 69 (2): 219-281, DOI: http://dx.doi.org/10.3897/dez.69.89864, URL: http://dx.doi.org/10.3897/dez.69.89864
0D22FADAAA615F7887CD9E7B072E63BA.taxon	description	[Japanese name: Tomokuni-gunbai] Figs 3 F, 5 F, 8 F, 10 G, 12 F, 14 G, 16 F, 18 F, 20 D, 22 D, 24 D, 26 F, 28 F, 30 H, 32 H, 39, 42 K, L	en	Souma, Jun (2022): Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan. Deutsche Entomologische Zeitschrift 69 (2): 219-281, DOI: http://dx.doi.org/10.3897/dez.69.89864, URL: http://dx.doi.org/10.3897/dez.69.89864
0D22FADAAA615F7887CD9E7B072E63BA.taxon	diagnosis	Diagnosis. Stephanitis (Stephanitis) tomokunii sp. nov. is recognised amongst other species of Stephanitis by a combination of the following characters: head, pronotal disc, marking on hemelytra and ventral surface in various shades of brown (Figs 8 F, 10 G, 12 F, 14 G, 16 F, 18 F, 20 D, 22 D, 24 D); calli dark brown; body in male 2.3 times (in female 2.1 times) as long as maximum width across hemelytra (Figs 3 F, 5 F); rostrum not reaching metasternum; pronotum tricarinate (Fig. 26 F); hood pale, shorter than median carina of pronotum, as wide as vertex at widest part, not covering eye, as high as median carina of pronotum at highest part, posterior margin not extending to middle of pronotal disc; median carina of pronotum with 1 - 2 rows of areolae at highest part; pronotal disc; paranotum less erect, narrowed posteriorly, 3 rows of areolae at widest part, anterolateral angle slightly protruding anteriad, outer margin gently curved inwards at posterolateral angle, maximum height shorter than height of eye (Fig. 28 F); apices of hemelytra close to each other in rest; costal area with 3 rows of areolae at widest part; subcostal area in male with 2 rows (in female with 2 - 3 rows) of areolae at widest part; discoidal area with 3 - 4 rows of areolae at widest part; sutural area with 3 - 4 rows of areolae at widest part; hypocostal lamina with a single row of areolae throughout its length; R + M (radiomedial) vein carinate; pygophore elevated at centre of venter, posterior margin slightly emarginate in middle part (Fig. 30 H); and paramere stout, weakly curved inward at apex, with outer margin not sinuate in middle part, inner margin nearly straight in basal part (Fig. 32 H).	en	Souma, Jun (2022): Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan. Deutsche Entomologische Zeitschrift 69 (2): 219-281, DOI: http://dx.doi.org/10.3897/dez.69.89864, URL: http://dx.doi.org/10.3897/dez.69.89864
0D22FADAAA615F7887CD9E7B072E63BA.taxon	description	Description. Male. Head, pronotal disc, marking on hemelytra and ventral surface in various shades of brown; calli dark brown; eye dark red; areolae of pronotum and hemelytron transparent; hood pale; pronotal disc opaque; pubescence on body yellowish (Figs 3 F, 8 F, 12 F, 16 F, 20 D, 22 D). Body 2.3 times as long as maximum width across hemelytra (Fig. 3 F). Head (Figs 8 F, 12 F, 20 D, 26 F) glabrous; pair of frontal spines close to each other at apices, not reaching apex of clypeus; median spine as long as frontal spines, reaching bases of frontal spines; pair of occipital spines longer than median spine, reaching middle part of eyes; antenniferous tubercles obtuse, slightly curved inwards; clypeus smooth. Compound eye round in dorsal view. Antenna densely covered with pubescence; segment I cylindrical; segment II cylindrical, shortest amongst antennal segments; segment III longest amongst antennal segments; segment IV cylindrical, longer than segment I. Bucculae closed to each other at anterior ends, with 2 rows of areolae throughout its length. Rostrum not reaching metasternum. Pronotum (Figs 8 F, 12 F, 26 F, 28 F) unicarinate, 1.3 times as long as maximum width across paranota, sparsely covered with pubescence. Pronotal disc coarsely punctate. Hood shorter than median carina of pronotum, as wide as vertex at widest part, not covering eye, as high as median carina of pronotum at highest part, posterior margin not extending to middle of pronotal disc, 4 rows of areolae at highest part, dorsal margin slightly arched. Median carina straight, extending to apex of posterior process, 1 - 2 rows of areolae at highest part, dorsal margin arched. Calli smooth. Paranotum less erect, narrowed posteriorly, 3 rows of areolae at widest part, with anterolateral angle slightly protruding anteriad, outer margin gently curved inwards at posterolateral angle, maximum height shorter than height of eye. Posterior process triangular, obtuse at apex. Hemelytron (Fig. 16 F) 2.5 times as long as its maximum width, extending beyond apex of abdomen, glabrous; maximum width across hemelytra 1.6 times as much as maximum width across paranota; apices close to each other in rest; costal area with 3 rows of areolae at widest part; subcostal area with 2 rows of areolae at widest part; discoidal area with 3 - 4 rows of areolae at widest part; sutural area with 3 - 4 rows of areolae at widest part; hypocostal lamina with a single row of areolae throughout its length; C (costal), R + M (radiomedial) and Cu (cubital) veins carinate. Thoracic pleura (Fig. 12 F) smooth in anterior part, coarsely punctate in posterior part. Ostiolar peritreme oblong. Sternal laminae (Fig. 20 D) lower than bucculae; pro- and mesosternal laminae open in both anterior and posterior ends; metasternal laminae as high as mesosternal laminae, open at anterior ends, fused each other at posterior ends. Legs (Fig. 3 F) smooth, densely covered with pubescence; femora thickest at middle. Abdomen oblong in dorsal and ventral views. Pygophore (Figs 22 D, 30 H) compressed dorsoventrally, semicircular in ventral view, elevated at centre of venter, posterior margin slightly emarginate in middle part, covered with pubescence. Paramere (Fig. 32 H) stout, expanded in middle part, weakly curved inwards at apex, outer margin not sinuate in middle part, inner margin nearly straight in basal part, covered with pubescence in middle part of outer and inner margins. Measurements (n = 20). Body length with hemelytra 3.1 - 3.4 mm; maximum width across hemelytra 1.4 - 1.5 mm; length of antennal segments I to IV 0.2 mm, 0.1 mm, 1.2 - 1.3 mm and 0.6 mm, respectively; pronotal length 1.2 - 1.4 mm; pronotal width across paranota 0.8 - 0.9 mm; hemelytral length 2.4 - 2.6 mm; maximum width of hemelytron 1.0 - 1.1 mm. Female. General habitus very similar to that of male (Figs 5 F, 10 G, 14 G, 18 F, 24 D) except for the following characters: body 2.1 times as long as maximum width across hemelytra; antennal segment III shorter than in male; pronotum 1.4 times as long as maximum width across paranota; hood wider than in male; maximum width across subcostal area wider than in male, with 2 - 3 rows of areolae at widest part; and apical part of abdomen pentagonal in ventral view. Measurements (n = 20). Body length with hemelytra 3.3 - 3.6 mm; maximum width across hemelytra 1.5 - 1.7 mm; length of antennal segments I to IV 0.2 mm, 0.1 mm, 1.0 - 1.1 mm and 0.6 mm, respectively; pronotal length 1.3 - 1.5 mm; pronotal width across paranota 0.9 - 1.0 mm; hemelytral length 2.5 - 2.7 mm; maximum width of hemelytron 1.0 - 1.2 mm.	en	Souma, Jun (2022): Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan. Deutsche Entomologische Zeitschrift 69 (2): 219-281, DOI: http://dx.doi.org/10.3897/dez.69.89864, URL: http://dx.doi.org/10.3897/dez.69.89864
0D22FADAAA615F7887CD9E7B072E63BA.taxon	distribution	Distribution. Japan (Izu Islands (southern part): Miyake Island, Hachijo Island) (Fig. 48) (Takeya 1963; Tomokuni and Ishikawa 2002; Yamada and Tomokuni 2012; present study). Stephanitis (Stephanitis) tomokunii sp. nov. inhabits the laurilignosa in a temperate climate of the southern part of the Izu Islands, which is in the Palaearctic Region.	en	Souma, Jun (2022): Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan. Deutsche Entomologische Zeitschrift 69 (2): 219-281, DOI: http://dx.doi.org/10.3897/dez.69.89864, URL: http://dx.doi.org/10.3897/dez.69.89864
0D22FADAAA615F7887CD9E7B072E63BA.taxon	etymology	Etymology. The new species is named in honour of Masaaki Tomokuni, a Japanese heteropterist who collected some of the paratype specimens.	en	Souma, Jun (2022): Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan. Deutsche Entomologische Zeitschrift 69 (2): 219-281, DOI: http://dx.doi.org/10.3897/dez.69.89864, URL: http://dx.doi.org/10.3897/dez.69.89864
0D22FADAAA615F7887CD9E7B072E63BA.taxon	biology_ecology	Biology. Stephanitis (Stephanitis) tomokunii sp. nov. feeds on the abaxial surface of leaves of Machilus thunbergii (present study). Adults were collected in May, July and August (Takeya 1963; Tomokuni and Ishikawa 2002; present study); the nymph and overwintering stage are unknown.	en	Souma, Jun (2022): Integrative taxonomy of the Lauraceae - feeding species of the genus Stephanitis (Hemiptera, Heteroptera, Tingidae) from Japan. Deutsche Entomologische Zeitschrift 69 (2): 219-281, DOI: http://dx.doi.org/10.3897/dez.69.89864, URL: http://dx.doi.org/10.3897/dez.69.89864
