taxonID	type	description	language	source
6243878EFFBEA364FF24FDD01DAFFBC1.taxon	materials_examined	Type species: Yezonychus sapporensis Ehara, 1978	en	Beard, Jennifer, Walter, David Evans (2010): New spider mite genus (Prostigmata: Tetranychidae) from Australia & New Zealand, with a discussion of Yezonychus Ehara. Zootaxa 2578: 1-24, DOI: 10.5281/zenodo.197527
6243878EFFBEA364FF24FDD01DAFFBC1.taxon	diagnosis	Diagnosis. Three pairs of prodorsal setae, nine pairs of dorsal opisthosomal setae, seta f 2 absent, setae h 1 – 3 present; empodia elongate with basal stalk split distally into two claw-like prongs (stalk clearly visible in dorsoventral perspective; Fig. 1); tenent hairs elongate, fine. Insertion of posterior dorsal opisthosomal setae h 2 posterior to insertion of setae h 1 (h 2 inserted in commonly observed position). Female with spinneret (suζ) longer than broad. Dorsal setae longer than distance to base of setae in next setal row.	en	Beard, Jennifer, Walter, David Evans (2010): New spider mite genus (Prostigmata: Tetranychidae) from Australia & New Zealand, with a discussion of Yezonychus Ehara. Zootaxa 2578: 1-24, DOI: 10.5281/zenodo.197527
6243878EFFBEA364FF24FDD01DAFFBC1.taxon	discussion	Remarks. Monotypic genus of the “ little web ” or LW-s life-type sensu Saito (1983; 1985; 2009), i. e. mites spin life lines while walking, but have no other use of threads. Yezonychus mites attach their eggs to the tips of leaf hairs, and quiescent stages also rest in similar position.	en	Beard, Jennifer, Walter, David Evans (2010): New spider mite genus (Prostigmata: Tetranychidae) from Australia & New Zealand, with a discussion of Yezonychus Ehara. Zootaxa 2578: 1-24, DOI: 10.5281/zenodo.197527
6243878EFFBEA364FF24FBD21D95F94F.taxon	materials_examined	Material examined. Japan. 7 Ψ, 4 ɗ, deutonymph, protonymph, ex. leaves of Sasa senanensis (Franc. & Sav.) (Poaceae), Sapporo, Hokkaido, 30. vii. 2009, A. Chittenden (QM, all separate slides); 3 Ψ, 4 ɗ, Sapporo, 05. viii. 2008, Y. Saito (QM, 4 slides).	en	Beard, Jennifer, Walter, David Evans (2010): New spider mite genus (Prostigmata: Tetranychidae) from Australia & New Zealand, with a discussion of Yezonychus Ehara. Zootaxa 2578: 1-24, DOI: 10.5281/zenodo.197527
6243878EFFBEA364FF24FBD21D95F94F.taxon	discussion	Remarks. Empodia I – II of both sexes are almost identical, with the empodia of males being slightly smaller than those of the female. Number of setae on legs I – IV: female cx 2, 2, 1, 1; tr 1, 1, 1, 1; fe 9, 7, 3, 3; ge 4, 5, 2, 1; ti 8 (1 + 0), 6, 6, 7; ta 16 (2 + 3), 15 (2 + 3), 11 (1 + 0), 11 (1 + 0); male cx 2, 2 / 1 *, 1, 1; tr 1, 1, 1, 1; fe 9, 6 / 7 *, 3, 3; ge 4, 5, 2, 1; ti 11 (4 + 0), 6, 6, 6 / 7 *; ta 17 (4 + 3), 15 (2 + 3), 10 (1 + 0), 9 * / 10 (1 + 0); deutonymph tr 1, 1, 1, 1; fe 9, 6, 3, 3; ge 4, 5, 2, 1; ti 8 (1 + 0), 6, 6, 4 / 5; ta 16 (2 + 3), 14 (1 + 3), 11 (1), 6; protonymph tr 0, 0, 0, 0; fe 3, 3, 2, 2; ge 4, 4, 2, 1; ti 6 (1 + 0), 5, 5, 5; ta 13 (2 + 3), 10 (1 + 1), 8, 6 [* number setae listed by Ehara (1978)]. We did not examine any larvae, and Ehara (1978) did not provide any morphological details of the immatures. This species is pale yellowish-white to pale greenish-yellow in colour (pers. com. Saito), and females are red in winter (Ehara 1978). Individuals live within woven silken nests.	en	Beard, Jennifer, Walter, David Evans (2010): New spider mite genus (Prostigmata: Tetranychidae) from Australia & New Zealand, with a discussion of Yezonychus Ehara. Zootaxa 2578: 1-24, DOI: 10.5281/zenodo.197527
6243878EFFBEA367FF24F9571C0EFEAC.taxon	materials_examined	Type species: Tribolonychus collyerae Zhang & Martin 2001: 321	en	Beard, Jennifer, Walter, David Evans (2010): New spider mite genus (Prostigmata: Tetranychidae) from Australia & New Zealand, with a discussion of Yezonychus Ehara. Zootaxa 2578: 1-24, DOI: 10.5281/zenodo.197527
6243878EFFBEA367FF24F9571C0EFEAC.taxon	diagnosis	Diagnosis. Three pairs of prodorsal setae, nine pairs of dorsal opisthosomal setae, seta f 2 absent, setae h 1 – 3 present; empodia lacking basal stalk, split into three separate prongs (trifid), i. e. two lateral prongs, one smaller dorsomedial prong; tenent hairs elongate, fine. Insertion of posterior dorsal opisthosomal setae h 2 posterior to insertion of setae h 1 (h 2 inserted in commonly observed position). Female with spinneret (suζ) longer than broad.	en	Beard, Jennifer, Walter, David Evans (2010): New spider mite genus (Prostigmata: Tetranychidae) from Australia & New Zealand, with a discussion of Yezonychus Ehara. Zootaxa 2578: 1-24, DOI: 10.5281/zenodo.197527
6243878EFFBEA367FF24F9571C0EFEAC.taxon	discussion	Remarks. Based on the importance of pretarsal morphology to the systematics of the Tetranychidae, this monospecific genus was erected to accommodate T. collyerae, for its unusual empodium which is split into three separate claw-like projections.	en	Beard, Jennifer, Walter, David Evans (2010): New spider mite genus (Prostigmata: Tetranychidae) from Australia & New Zealand, with a discussion of Yezonychus Ehara. Zootaxa 2578: 1-24, DOI: 10.5281/zenodo.197527
6243878EFFBDA367FF24FB481DA9F9E8.taxon	discussion	Remarks. This species is known only from three females collected by E. Collyer from southern beech, Nothofagus sp. (Nothofagaceae) in New Zealand. All three specimens are mounted on one slide (ZQZ 990826; NZAC). Males and immatures of this species are unknown (Zhang & Martin 2001). Number of setae on legs I – IV of female (adjusted from Zhang & Martin 2001): tr 1, 1, 1, 1; fe 9, 5, 3, 2; ge 5, 5, 3, 2; ti 8 (1 + 0), 5, 5, 5; ta 15 (3 + 3), 12 (2 + 3), 7 (1 + 0), 7 (1 + 0). Based on illustrations in Zhang & Martin (2001), tibia and tarsus I-II with same chaetotaxy as female N. tereotus (see Table 1; Fig. 11).	en	Beard, Jennifer, Walter, David Evans (2010): New spider mite genus (Prostigmata: Tetranychidae) from Australia & New Zealand, with a discussion of Yezonychus Ehara. Zootaxa 2578: 1-24, DOI: 10.5281/zenodo.197527
6243878EFFBDA366FF24F9F51EF5F9BF.taxon	materials_examined	Type species: Schizotetranychus cornus Pritchard & Baker 1955, original designation	en	Beard, Jennifer, Walter, David Evans (2010): New spider mite genus (Prostigmata: Tetranychidae) from Australia & New Zealand, with a discussion of Yezonychus Ehara. Zootaxa 2578: 1-24, DOI: 10.5281/zenodo.197527
6243878EFFBDA366FF24F9F51EF5F9BF.taxon	diagnosis	Diagnosis. Three pairs of prodorsal setae, nine pairs of dorsal opisthosomal setae, seta f 2 absent, setae h 1 – 3 present; empodia lacking basal stalk (Figs 2, 3, 16), split into two structures each with three prongs (Figs 2 A – D, 3), often with ventral prong of each pair much thicker than two dorsal prongs; tenent hairs thick with large heads. Insertion of posterior dorsal opisthosomal setae h 2 variable in relation to setae h 1, from obviously anterior to the insertion of h 1 to obviously posterior. Female with spinneret (suζ) broader than long. Live within nests with a thickly woven silken roof, i. e. life type WN sensu Saito (1983; 1985; 2009). opposite the instar in which they first appear (F = female); parentheses indicate pairs of setae. Trochanter Femur Genu Tibia Tarsus Leg I	en	Beard, Jennifer, Walter, David Evans (2010): New spider mite genus (Prostigmata: Tetranychidae) from Australia & New Zealand, with a discussion of Yezonychus Ehara. Zootaxa 2578: 1-24, DOI: 10.5281/zenodo.197527
6243878EFFBDA366FF24F9F51EF5F9BF.taxon	etymology	Etymology. This genus name is derived from the Latin “ neo ” meaning “ spin ”, and from the Latin “ nidulus ”, a masculine diminutive form of “ nidis ” meaning “ nest ”, and refers to the nests built by these mites, with a densely woven web canopy.	en	Beard, Jennifer, Walter, David Evans (2010): New spider mite genus (Prostigmata: Tetranychidae) from Australia & New Zealand, with a discussion of Yezonychus Ehara. Zootaxa 2578: 1-24, DOI: 10.5281/zenodo.197527
6243878EFFBDA366FF24F9F51EF5F9BF.taxon	discussion	Remarks. The dorsal chaetotaxy of this genus is the same as Yezonychus Ehara and Tribolonychus Zhang & Martin, i. e. seta f 2 absent. The empodia of female Neonidulus (a pair of structures each with three prongs) differ to those of Yezonychus (basal stalk, split distally into two claw-like structures) and Tribolonychus (a total of three separate prongs, medial claw dorsal to two lateral claws). The legs of the Australian and New Zealand species of Neonidulus have the same number of setae on tr I – IV, ge I – IV, ti I – IV, but different numbers on the coxae, femora and tarsi. The leg setal count for New Zealand species N. cornus, N. falsicornus and N. brevipilus is: female cx 2, 2, 1, 1; tr 1, 1, 1, 1; fe 9, 5, 2, 0; ge 5, 4, 3, 3; ti 8 (1 + 0), 5, 5, 5; ta 16 (3 + 3), 12 (2 + 3), 7 (1 + 0), 7 (1 + 0); male same as female except ta I with 17 (4 + 3) (Fig. 2 E). The number of setae on cx I – IV, fe I – IV and ta I – IV for N. tereotus is: female cx 2, 1, 1, 1; fe 9, 6, 3, 3; ta 15 (3 + 3), 12 (2 + 3), 8 (1 + 0), 8 (1 + 0); male same as female except ti I 9 (2 + 0) and ta I 17 (5 + 3). The males of the three New Zealand species do not have an enlarged or modified leg I like those present on the Australian species, N. tereotus (Figs 2 E, 15, 17).	en	Beard, Jennifer, Walter, David Evans (2010): New spider mite genus (Prostigmata: Tetranychidae) from Australia & New Zealand, with a discussion of Yezonychus Ehara. Zootaxa 2578: 1-24, DOI: 10.5281/zenodo.197527
6243878EFFBCA360FF24F90F1F5AFE69.taxon	materials_examined	Material examined. Holotype. ɗ. New Zealand, Levin, ex. Elaeocarpus dentatus (Elaeocarpaceae), xii. 1949, K. P. Lamb (USNM, 2 ɗ on slide; A- 27, Lot 50 - 2779). Paratypes. 2 ɗ, 4 Ψ, 2 deutonymphs, 3 protonymphs, 2 larvae, 11 additional immatures (cannot determine stage, slides require remounting) (all in USNM; 13 slides in total).	en	Beard, Jennifer, Walter, David Evans (2010): New spider mite genus (Prostigmata: Tetranychidae) from Australia & New Zealand, with a discussion of Yezonychus Ehara. Zootaxa 2578: 1-24, DOI: 10.5281/zenodo.197527
6243878EFFBCA360FF24F90F1F5AFE69.taxon	discussion	Remarks. Dorsal setae h 1 – 3 inserted in commonly observed positions, in all stages. All stages of this mite are orange to red or carmine in colour with light black irregular streaking or black patches (Manson 1967; Zhang & Martin 2001). The mites live on the underside of leaves of their host plant, Elaeocarpus dentatus (Elaeocarpaceae), inside natural domatia created at the junction between main veins and side veins, and spin dense webbing over the entrance (Manson 1967; see Zhang & Martin 2001: 312 for further details). Manson (1967) listed the number of setae on the legs of the immatures: deutonymph cx 2, 2, 1, 1; fe 5, 3, 2, 0; ge 5, 4, 3, 3; ti 8 (1 + 0), 5, 5, 5; ta 16 (3 + 3), 11 (1 + 3), 7 (1 + 0), 6; protonymph cx 2, 1, 1, 0; fe 3, 3, 2, 0; ge 4, 3, 2, 2; ti 6 (1 + 0), 5, 5, 5; ta 13 (2 + 0), 11 (1 + 0), 6, 4; larva cx 1, 0, 0; fe 3, 3, 2; ge 4, 3, 2; ti 6 (1 + 0), 5, 5; ta 9 (1 + 0), 9 (1 + 0), 4.	en	Beard, Jennifer, Walter, David Evans (2010): New spider mite genus (Prostigmata: Tetranychidae) from Australia & New Zealand, with a discussion of Yezonychus Ehara. Zootaxa 2578: 1-24, DOI: 10.5281/zenodo.197527
6243878EFFBAA360FF24FE621F0AFC19.taxon	materials_examined	Material examined. New Zealand. Ψ, ɗ, deutonymph, protonymph, larva, Banks Peninsula, ex. Rubus cissoides (Rosaceae), 22. i. 2002, N. A. Martin (all on same slide); deutonymph, ex. leaves of R. cissoides in silken shelters, Tapotupotu Bay, 25. ii. 2002, R. C. Henderson (five deutonymphs on one slide); larva, same data (deutonymph, larva, egg on same slide). All material deposited in NZAC.	en	Beard, Jennifer, Walter, David Evans (2010): New spider mite genus (Prostigmata: Tetranychidae) from Australia & New Zealand, with a discussion of Yezonychus Ehara. Zootaxa 2578: 1-24, DOI: 10.5281/zenodo.197527
6243878EFFBAA360FF24FE621F0AFC19.taxon	discussion	Remarks. Adult female with insertion of dorsal posterior opisthosomal setae h 2 anterior to insertion of setae h 1 (Fig. 4). Immature stages with insertion of h 2 approximately level with that of h 1 (Figs 5, 6). These mites live on the underside of the leaves of their host plant, Rubus cissoides (Rosaceae), and spin a canopy of dense webbing over depressions in the leaf surface (e. g. dimples made by scale insects), or from prominent leaf veins (Manson 1967; Zhang & Martin 2001). Live mites are similar in appearance to N. cornus (Zhang & Martin 2001).	en	Beard, Jennifer, Walter, David Evans (2010): New spider mite genus (Prostigmata: Tetranychidae) from Australia & New Zealand, with a discussion of Yezonychus Ehara. Zootaxa 2578: 1-24, DOI: 10.5281/zenodo.197527
6243878EFFB8A362FF24FB911A50FA75.taxon	discussion	Remarks. Adult female with insertion of dorsal posterior opisthosomal setae h 2 level with insertion of setae h 1. Immatures are not mentioned in the description by Zhang and Martin (2001). This species lives on the upper and lower surface of the leaves of its host plant, Leionema nudum (Rutaceae), spinning thick webbing at the base of the leaf between the midrib and the edge of the leaf, or over natural depressions in the leaf surface (see Zhang & Martin 2001: 320 for further details). The mites are similar in appearance to N. cornus (Zhang & Martin 2001). All material deposited in NZAC (holotype male ZQZ 990115 - 1 b).	en	Beard, Jennifer, Walter, David Evans (2010): New spider mite genus (Prostigmata: Tetranychidae) from Australia & New Zealand, with a discussion of Yezonychus Ehara. Zootaxa 2578: 1-24, DOI: 10.5281/zenodo.197527
6243878EFFB8A377FF24FA421C07FE4F.taxon	materials_examined	Material examined. Holotype. Ψ. Australia, ex. Weeping Lillypilly, Waterhousea floribunda (Myrtaceae), The University of Queensland St Lucia campus, Brisbane, Queensland, 27 ° 30 ’ S 153 ° 00 ’ E, 11. xi. 2000, D. E. Walter (QM). Paratypes. 10 Ψ, ɗ, 1 deutonymph, same data as holotype (one slide); 2 ɗ, same data as holotype except 12. xii. 2000 (one slide); 8 Ψ, 5 ɗ, deutonymph, 4 protonymphs, 4 larvae, same data as holotype except 05. xii. 2001, J. J. Beard (all on separate slides, QM); 4 Ψ, ɗ, protonymph, larva, same data (all on separate slides, USNM). Other material examined: 10 Ψ, 5 ɗ, pharate Ψ, 9 deutonymphs, 2 pharate deutonymphs, 3 protonymphs, 1 larva, Gap Creek Road, Kenmore, Brisbane, 27 ° 28 ’ 51 ” S 152 ° 55 ’ 39 ” E, 19. xii. 2004, ex. Brush Box, Lophostemon confertus (Myrtaceae), J. J. Beard and P. I. Forster (all separate slides; QM); 4 Ψ, 3 ɗ, deutonymph, protonymph, same data (USNM); 2 Ψ, 2 ɗ, 2 deutonymphs, protonymph, same data (NZAC); 2 Ψ, ɗ, 2 deutonymphs, 2 protonymphs, 2 larvae (separate slides), protonymph 3 larvae (on same slide), ex. Brush Box, Lophostemon confertus, 6 km WNW Orchid Beach, Great Sandy National Park, Fraser Island, 24 ° 56 ’ 56 ” S 153 ° 15 ’ 40 ” E, 04. ix. 2004, P. I. Forster & J. J. Beard (QM).	en	Beard, Jennifer, Walter, David Evans (2010): New spider mite genus (Prostigmata: Tetranychidae) from Australia & New Zealand, with a discussion of Yezonychus Ehara. Zootaxa 2578: 1-24, DOI: 10.5281/zenodo.197527
6243878EFFB8A377FF24FA421C07FE4F.taxon	diagnosis	Diagnosis. Prodorsum with three pairs of setae, two pairs of eyes; opisthosoma with nine pairs of setae (c 1, c 2, c 3, d 1, d 2, e 1, e 2, f 1, h 1); setae h 2 – 3 present (para-anal setae); two pairs of ps setae (anal setae); ventral setae 2 c absent; entire dorsum plicate, with distinct regular polygonal reticulate pattern medially on prodorsum; peritreme with short distal hook like a golf-club (sometimes hook not apparent, appears as a distal expansion); tarsus I with two pairs of duplex setae, distal and adjacent; tarsus II with one pair duplex setae; female empodia I – IV and male empodia II – IV consisting of a pair of three-pronged structures, each prong thickened basally and strongly tapering distally; male empodium I consisting of a pair of three-pronged structures, each prong short, stout and subequal in length, median prong slightly larger than other prongs (Fig. 16). As for genus in addition to: dorsal setae long, thin, obviously barbed; male leg I enlarged, elongate; male femur I with two adaxial setae and one ventral seta modified into thickened spines; female with spinneret (suζ) obviously swollen; female pregenital striae longitudinal, becoming broken towards setae ag and forming a distinct cresentic pattern between setae ag; male body strongly diamond shaped; female body strongly rounded.	en	Beard, Jennifer, Walter, David Evans (2010): New spider mite genus (Prostigmata: Tetranychidae) from Australia & New Zealand, with a discussion of Yezonychus Ehara. Zootaxa 2578: 1-24, DOI: 10.5281/zenodo.197527
6243878EFFB8A377FF24FA421C07FE4F.taxon	description	Female. Dorsum. (Fig. 7) Body measurements: v 2 – h 1 290 – 320, c 1 – c 1 15 – 26, c 3 – c 3 222 – 264, d 1 – d 1 17 – 26, d 1 – d 2 61 – 73, e 1 – e 1 25 – 33, e 1 – e 2 45 – 61, f 1 – f 1 18 – 31, h 1 – h 1 26 – 51. Prodorsum distinct, with regular polygonal reticulate pattern mesally, fine striate pattern laterally and anteriorly; series of small pores visible in row medad setae sc 1 (6 - 8); opisthosoma transversely striate, with striae forming an arch between setae e 2 – e 2; lateral opisthosomal striae longitudinal. Dorsum with 14 pairs of finely barbed setae, mostly longer than the distance to the next setal base; v 2 32 – 38, sc 1 63 – 73, sc 2 56 – 71, c 1 64 – 85, c 2 50 – 67, c 3 74 – 92, d 1 66 – 78, d 2 47 – 67, e 1 63 – 78, e 2 55 – 73, f 1 56 – 75, h 1 58 – 71, h 2 22 – 43, h 3 31 – 37. Setae h 2 and h 3 usually inserted ventrally. Ve n te r. (Fig. 8) Body finely striate, striae mostly transverse with longitudinal striae between setae 3 a – 4 a and laterad pregenital region; coxal bases III – IV smooth; coxal bases I – II smooth or with light, fine striae; small lobes present on striae between setae 1 a – 3 a, and between setae 4 a – ag; distinct pattern in striae between setae ag – g 1 — female pregenital striae longitudinal, becoming broken towards setae ag and forming a distinct cresentic pattern between setae ag (Fig. 9); genital flap small; all ventral setae thin, smooth: 1 a 21 – 32, 1 b 42 – 64, 1 c 23 – 42, 2 b 23 – 34, 3 a 23 – 32, 3 b 33 – 39, 4 a 22 – 28, 4 b 27 – 37, ag 17 – 35, g 1 23 – 27, g 2 27 – 34, ps 1 13 – 28, ps 2 15 – 21. Gnathosoma. Palp tarsus with short, broad, swollen spinneret (su ζ) (6 – 7 wide, 4 – 7 long) (Fig. 10 A); single solenidion (3 – 5) and two eupathids (ul’ 3 – 5, ul ” 6 – 7) (Fig. 10 A). Peritreme ending in small expansion or short hook (Fig. 10 B). Legs. (Fig. 11) Tarsus I with two duplex setae (solenidia: proximal ω ' 39 – 50, distal ω " 49 – 64), proximal solenidion (ω " 1 26 – 38) and three distal eupathidia (pv’ ζ 12 – 18), tibia I with one solenidion (φ 33 – 49), tarsus II with one duplex seta (solenidion ω " 40 – 54), proximal solenidion (ω " 1 24 – 41) and three distal eupathidia (pv’ ζ 15 – 20), tarsus III with solenidion (ω ’ 31 – 38), tarsus IV with solenidion (ω ’ 30 – 38); all leg setae finely barbed. Number of setae on legs I – IV respectively: coxae 2, 1, 1, 1; trochanters 1, 1, 1, 1; femora 9, 6, 3, 3; genua 5, 4, 3, 3; tibiae 8 (1 + 0), 5, 5, 5; tarsi 15 (3 + 3), 12 (2 + 3), 8 (1 + 0), 8 (1 + 0). Aberrant setal counts: femur I: 1 / 12 females with 8 setae — one dorsal distal seta absent; femur II: 1 / 12 females with 7 setae — extra dorsal seta, 1 / 12 with 5 setae — one ventral proximal seta absent; genu II: 1 / 12 females with 3 setae — one dorsal seta absent; genu III: 1 / 12 females with 2 setae — one dorsal seta absent; tibia I: 1 / 12 females with 9 (2 + 0) setae — extra solenidion (male chaetotaxy); tarsus I: 1 / 12 females with 17 (5 + 3) setae — two extra solenidia (male chaetotaxy); tarsus III with 9 (1 + 0) setae — extra ventral seta; tarsus IV: 1 / 12 females with 7 (1 + 0) setae — one dorsal seta absent. Colour. Pale olive green with red eyes; eggs are pale yellow to green and rounded. Male. Dorsum. (Fig. 12) Body measurements: v 2 – h 1 238 – 287, c 1 – c 1 13 – 20, c 3 – c 3 173 – 221, d 1 – d 1 13 – 21, d 1 – d 2 46 – 71, e 1 – e 1 23 – 31, e 1 – e 2 32 – 41, f 1 – f 1 11 – 18, h 1 – h 1 22 – 35. Prodorsum distinct, with fine regular polygonal reticulate pattern mesally, fine striate pattern laterally and anteriorly; series of small pores (8) visible in row medad setae sc 1; opisthosoma transversely striate. Dorsum with 14 pairs of barbed setae, mostly longer than the distance to the next setal base; v 2 28 – 46, sc 1 48 – 75, sc 2 60 – 91, c 1 53 – 72, c 2 44 – 73, c 3 68 – 96, d 1 49 – 70, d 2 43 – 63, e 1 42 – 64, e 2 42 – 64, f 1 42 – 57, h 1 43 – 61, h 2 22 – 33, h 3 25 – 37. Setae h 2 and h 3 usually inserted posterodorsally. Ve n te r. Body finely striate with large areas of smooth cuticle around coxal bases; striae between setae 3 a and 4 a with lobes; all ventral setae thin, smooth: 1 a 22 – 30, 1 b 49 – 76, 1 c 30 – 45, 2 b 27 – 39, 3 a 25 – 36, 3 b 33 – 46, 4 a 21 – 32, 4 b 31 – 40, ag 13 – 20, g 1 13 – 17, g 2 14 – 17, ps 1 13 – 16, ps 2 15 – 18. Gnathosoma. Palp tarsus with cone-shaped spinneret (su ζ) smaller than female (3 – 4) with nipple-like tip; single solenidion (4 – 5) and two eupathids (ul’ 4 – 5, ul ” 5 – 7) (Fig. 13). Peritreme ending in small expansion or short hook (see Fig. 10 B). Aedeagus. (Fig. 14) Long, thin and tapering to finger-like tip (see Fig. 14 for measurements, a: 3 – 4, b: 20 – 27, c: 6, d: 47 – 62. Legs. (Figs 15 – 17) Tarsus I with two duplex setae (solenidia: proximal ω ' 39 – 50, distal ω " 47 – 65), three solenidia (proximal – distal 20 – 28, 20 – 30, ω " 1 28 – 37) and three distal eupathidia (pv’ ζ 12 – 17); tibia I with two solenidia (proximal 21 – 29, φ distal 34 – 41); tarsus II with one duplex seta (solenidion ω " 41 – 51), single proximal solenidion (ω " 1 20 – 28) and three distal eupathidia (pv’ ζ 14 – 20); tarsus III with solenidion (ω ’ 26 – 40); tarsus IV with solenidion (ω ’ 26 – 36); all leg setae barbed; femur I with two thickened spine-like setae (l’ 17 – 34, v’ 10 – 19), barbed or smooth (Fig. 17), and one thickened seta (v ” 1 26 – 36). Number of setae on legs I – IV respectively: coxae 2, 1, 1, 1; trochanters 1, 1, 1, 1; femora 9, 6, 3, 3; genua 5, 4, 3, 3; tibiae 9 (2 + 0), 5, 5, 5; tarsi 17 (5 + 3), 12 (2 + 3), 8 (1 + 0), 8 (1 + 0). Aberrant setal counts: femur I: 1 / 13 males with 10 setae — extra spurlike seta, 1 / 13 males with 8 setae – one dorsal seta absent; tarsus I: 3 / 13 males with 16 (5 + 3) setae – one ventral seta absent. Empodia II – IV consisting of a pair of three-pronged structures, each prong thickened basally and strongly tapering distally; empodium I consisting of a pair of three-pronged structures, each prong short, stout and subequal in length, median prong slightly larger than other lateral prongs (Fig. 16). Colour. Yellow-green, red eyes, and legs I and II becoming orange distally. Deutonymph. Dorsum. Body measurements: v 2 – h 1 206 – 271, c 1 – c 1 16 – 21, c 3 – c 3 177 – 210, d 1 – d 1 17 – 22, d 1 – d 2 49 – 60, e 1 – e 1 23 – 33, e 1 – e 2 39 – 46, f 1 – f 1 12 – 21, f 1 – h 1 16 – 24. Prodorsum distinct, with fine regular polygonal reticulate pattern mesally, fine striate pattern laterally and anteriorly, weaker than that of adult. Dorsum with 14 pairs of lightly barbed setae — v 2 27 – 37, sc 1 42 – 60, sc 2 55 – 71, c 1 43 – 66, c 2 37 – 56, c 3 68 – 99, d 1 43 – 68, d 2 37 – 57, e 1 38 – 61, e 2 41 – 64, f 1 38 – 67, h 1 47 – 60, h 2 22 – 34, h 3 22 – 35; setae h 2, h 3 usually ventral. Ve n te r. Venter almost entirely transversely striate, striae between setae 3 a and g 1 with small lobes; ventral setae thin, smooth or lightly barbed: 1 a 19 – 26, 1 b 41 – 50, 1 c 21 – 33, 2 b 20 – 33, 3 a 26 – 32, 3 b 28 – 33, 4 a 19 – 27, 4 b 23 – 28, ag 20 – 27, g 1 15 – 25, ps 1 11 – 18, ps 2 11 – 17. Gnathosoma. The sexually dimorphic spinneret (su ζ), clearly visible in the adult stage, is also apparent in the deutonymph: spinneret broad rounded (5 wide, 5 long) to short thin (2 wide, 3 long); single solenidion (3 – 5) and two eupathids (ul’ 4 – 5, ul ” 5 – 6). Peritreme ending in small expansion or short hook. Legs. (Fig. 18) Tarsus I with two duplex setae (solenidia: proximal ω ' 28 – 40, distal ω " 45 – 51), single proximal solenidion (21 – 28), three distal eupathidia (pv’ ζ 13 – 15); tibia I with solenidion (φ 31 – 38); tarsus II with one duplex seta (solenidion ω " 38 – 44), three distal eupathidia (pv’ ζ 13 – 15); tarsus III with single solenidion (ω ’ 24 – 31); all leg setae lightly barbed except a few smooth setae distally on tarsi. Number of setae on legs I – IV respectively: coxae 2, 1, 1, 1; trochanters 1, 1, 1, 0; femora 6, 3, 2, 2; genua 5, 4, 3, 2; tibiae 8 (1 + 0), 5, 5, 5; tarsi 15 (3 + 3), 11 (1 + 3), 8 (1 + 0), 7. Protonymph. Dorsum. (Fig. 19) Body measurements: v 2 – h 1 155 – 223, c 1 – c 1 17 – 25, c 3 – c 3 145 – 203, d 1 – d 1 17 – 22, d 1 – d 2 42 – 47, e 1 – e 1 21 – 26, e 1 – e 2 22 – 32, f 1 – f 1 11 – 18, f 1 – h 1 12 – 24. Prodorsum distinct, with fine regular polygonal reticulate pattern mesally, fine striate pattern laterally and anteriorly, weaker than that of adult. Dorsum with 14 pairs of lightly barbed setae — v 2 23 – 31, sc 1 38 – 50, sc 2 41 – 70, c 1 43 – 57, c 2 30 – 43, c 3 64 – 86, d 1 37 – 57, d 2 36 – 50, e 1 32 – 51, e 2 32 – 58, f 1 31 – 52, h 1 34 – 54, h 2 20 – 33, h 3 22 – 32; setae h 2, h 3 usually ventral. Ve n te r. Venter mostly transversely striate, striae with small lobes; ventral setae thin, smooth or lightly barbed: 1 a 17 – 24, 1 b 38 – 43, 1 c 20 – 26, 2 b 21 – 24, 3 a 24 – 29, 3 b 22 – 30, g 1 17 – 22, ps 1 8 – 16, ps 2 7 – 15. Gnathosoma. The sexually dimorphic spinneret (su ζ), clearly visible in the adult stage, is also apparent to a lesser degree in the protonymph: spinneret broad rounded (4 wide, 3 long) to thin (2 – 3 wide, 4 long); single solenidion (3 – 4) and two eupathids (ul’ 4, ul ” 5 – 6). Peritreme ending in small expansion or short hook, like a golf-club. Legs. (Fig. 20) Tarsus I with two duplex setae (solenidia: proximal ω ' 27 – 33, distal ω " 39 – 51) and three distal eupathidia (pv’ ζ 11 – 18); tibia I with single solenidion (φ 27 – 31); tarsus II with one duplex seta (solenidion ω " 31 – 42) and three distal eupathidia (pv’ ζ 11 – 18); all leg setae lightly barbed except a few smooth setae distally on tarsi. Number of setae on legs I – IV respectively: coxae 2, 1, 1, 0; trochanters 0, 0, 0, 0; femora 3, 3, 2, 2; genua 4, 3, 2, 2; tibiae 6 (1 + 0), 5, 5, 5; tarsi 13 (2 + 3), 11 (1 + 3), 7, 5. Larva. Dorsum. (Fig. 21) Body measurements: v 2 – h 1 120 – 187, c 1 – c 1 22 – 29, c 3 – c 3 127 – 145, d 1 – d 1 16 – 24, d 1 – d 2 29 – 36, e 1 – e 1 17 – 25, e 1 – e 2 21 – 26, f 1 – f 1 6 – 12, f 1 – h 1 9 – 15. Prodorsum distinct, with small patch of fine regular polygonal reticulate pattern mesally, fine striate pattern laterally and anteriorly, weaker than that of adult. Dorsum with 14 pairs of lightly barbed setae — v 2 21 – 28, sc 1 30 – 42, sc 2 31 – 61, c 1 37 – 50, c 2 24 – 34, c 3 47 – 76, d 1 36 – 46, d 2 30 – 42, e 1 32 – 41, e 2 37 – 52, f 1 30 – 46, h 1 45 – 64, h 2 33 – 41, h 3 42 – 52; setae h 2, h 3 usually ventral. Ve n te r. Venter mostly transversely striate, striae with lobes; ventral setae thin, smooth or lightly barbed: 1 a 16 – 24, 1 b 36 – 40, 3 a 25 – 33, ps 1 7 – 13, ps 2 9 – 16. Gnathosoma. Palp tarsus with rounded spinneret (3 – 4) (su ζ), single solenidion (3 – 4) and single eupathid (ul’ 3 – 4). Peritreme ending in small expansion or short hook like golf club. Legs. (Fig. 22) Tarsus I with one set of duplex setae (solenidion ω ' 43 – 56) and two distal eupathids; tibia I with single solenidion (φ 20 – 31); tarsus II with one duplex seta (solenidion ω " 34 – 44) and two distal eupathids; all leg setae lightly barbed except a few smooth setae distally on tarsi. Number of setae on legs I – III respectively: coxae 1, 0, 0; trochanters 0, 0, 0; femora 3, 3, 2; genua 4, 3, 2; tibiae 6 (1 + 0), 5, 5; tarsi 9 (1 + 2), 9 (1 + 2), 5.	en	Beard, Jennifer, Walter, David Evans (2010): New spider mite genus (Prostigmata: Tetranychidae) from Australia & New Zealand, with a discussion of Yezonychus Ehara. Zootaxa 2578: 1-24, DOI: 10.5281/zenodo.197527
6243878EFFB8A377FF24FA421C07FE4F.taxon	discussion	Remarks. Ontogeny for leg setae and solenidia for nymphs and adult female given in Table 1. Individuals of N. tereotus feed on the underside of the leaves of their host plants, are dorsoventrally flattened, and spin strong, thick, sheets of webbing to form their nests, i. e. they have the life type known as the woven nest (WN) type sensu Saito (1983; 1985; 2009; pers. com. Saito 2009). Thick silk sheeting is woven along the midrib on the underside of the leaves, from the top of the midrib down to the surface of the leaf blade, forming a tent, that can be as long as 40 mm along the midrib. Females also begin new nests in natural depressions present on the leaf blade, weaving thick silken covers over the depression, usually forming a circular nest. Faecal material appears to be placed amongst the feeding areas within the nests, whereas the eggs are kept away from such areas (pers. com. Saito 2009). Individual females are able to start new nests, but it appears that males are unable to do so. Males use their enlarged forelegs to guard quiescent female deutonymphs prior to mating them, and males have been observed to pick up quiescent female deutonymphs and move them to a new position (pers. obs. D. Walter). We have not observed males attempt to defend the nest when it is broken open, nor male-to-male aggression behaviour like that displayed by males of Stigmaeopsis miscanthi (Saito), also a life type WN species (Saito 1995; Saito & Sahara 1999). Feeding within the nests causes the part of the leaf blade directly under the webbing to turn yellow-orange, and then brown in older nests. This discolouration can be clearly seen from the other side of the leaves, and as it persists long after the mites have died off, even in herbarium specimens (pers. obs. J. Beard), it is a good indication that the mites are present or have been in the past. While there is a broad range in the length of setae for this species, the range is not related to differences in the host plant, as setal lengths are equally variable on both recorded host plant species. The spinneret, suζ, in this species is sexually dimorphic, being enlarged in females and reduced in males. This morphological difference in the spinneret is also apparent in the deutonymphs and to a lesser extent in the protonymphs. The range in lengths of setae on the deutonymphs is also explained by differences in sexes, as female deutonymphs are larger than male deutonymphs. An examination of plant specimens located in the Queensland Herbarium (BRI) indicates the presence of this mite on both recorded hosts, most commonly on W. floribunda, since the earliest record in 1932. Etymology. The name is derived from the Greek word “ tereo ” meaning “ to watch over or guard ”, in reference to body form of the male.	en	Beard, Jennifer, Walter, David Evans (2010): New spider mite genus (Prostigmata: Tetranychidae) from Australia & New Zealand, with a discussion of Yezonychus Ehara. Zootaxa 2578: 1-24, DOI: 10.5281/zenodo.197527
