identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
677D87E1FFC13A01FF0721C2FDDC11F7.text	677D87E1FFC13A01FF0721C2FDDC11F7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mortoniella Ulmer 1906	<div><p>Genus Mortoniella</p><p>This genus has been treated comprehensively for the neotropics in a series of revisions by Blahnik and Holzenthal (2008, 2011, 2017). The reader is referred to the last of these publications for a discussion of the terminology used here and illustrations of structures. The two new species from Panama fall within the “florica subgroup” of the leroda group, as originally defined by Blahnik and Holzenthal (2008). The subgroup was subsequently merged within the leroda subgroup (Blahnik and Holzenthal 2017), due to uncertainty about the monophyly of the latter group as originally defined. The two subgroups represent the majority of the species of Mortoniella in Central America. Species assigned to the “florica subgroup” are characterized by paired, rounded dorsolateral projections on the phallicata, as well as paired apicoventral projections on the fused inferior appendages and a prominent apicoventral spine on the endophallic membrane.</p></div>	https://treatment.plazi.org/id/677D87E1FFC13A01FF0721C2FDDC11F7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Blahnik, Roger J.;Armitage, Brian J.	Blahnik, Roger J., Armitage, Brian J. (2019): The Trichoptera of Panama. XII. Contributions to the family Glossosomatidae (Insecta: Trichoptera) in Panama. Insecta Mundi 740 (740): 1-17, DOI: 10.5281/zenodo.3676607
677D87E1FFC13A07FF07236BFD3510E5.text	677D87E1FFC13A07FF07236BFD3510E5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mortoniella (Mortoniella) calovebora Blahnik and Armitage 2019	<div><p>Mortoniella (Mortoniella) calovebora Blahnik and Armitage, new species</p><p>Fig. 2</p><p>Diagnosis. This is a rather typical, but distinctive, species of the “florica subgroup.” Mortoniella calovebora is easily distinguished from other described species of the subgroup by the length and asymmetry of the apicoventral lobes of the inferior appendages (Fig. 2C). The asymmetry of the ventral lobes of the inferior appendages seems to be a consistent feature of this species and is a unique character within the genus.</p><p>Description. Length of forewing: male 3.0– 3.3 mm; female 3.2–3.3 mm. Forewing with forks I, II, and III present, hind wing with forks II and III. Spur formula 0:4:4. Overall color (in alcohol) light brown, anastamosis unpigmented, but otherwise not evidently marked.</p><p>Male genitalia. Ventral process of segment VII laterally compressed, short, ventrally projecting, rounded apically, length slightly greater than width at base, process slightly retracted anterobasally. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin rounded dorsally, narrowing ventrally; segment deeply excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by much less than ½ width of segment. Tergum X relatively elongate, lateral margins subparallel, apically with short, subtriangular mesal projection; apicolateral lobes elongate, compressed, subacute, slightly mesally curved; ventrolateral lobes nearly obsolete; ventromesal lobes not evident. Inferior appendages without dorsolateral lobes, but with apicoventral lobes elongate, upturned, and asymmetrically developed, that of the right side narrower apically and more distinctly out-turned, both lobes acute apically. Mesal pockets of inferior appendage with apical processes short and weakly curved. Paramere appendage elongate, narrow, extending about same length as dorsal phallic spine, slightly widened preapically, widened apex with narrow, adpressed, scale-like spines. Dorsal phallic spine, as viewed laterally, undulate in contour, spine distinctly widened on ventral margin at about middle, ventral bulge articulating with dorsal margin of phallicata, apex acute, weakly upturned; in dorsal view (Fig. 2D), lightly widened in middle and preapically, apex acute. Phallicata moderately elongate, dorsal margin with broadly rounded, paired, dorsolateral projections, lateral margin with slightly produced, rounded and compressed projection. Endophallic membrane simple, with single prominent curved ventromesal spine; phallotremal spines absent.</p><p>Female and immatures. Unknown.</p><p>Holotype male (alcohol). PANAMA: Veraguas: Cuenca 097, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-81.16486&amp;materialsCitation.latitude=8.55038" title="Search Plazi for locations around (long -81.16486/lat 8.55038)">Río Calovebora</a>, 8.55038°N and 81.16486°W, PSPSCD-PNSF-C097-2017-006, 461 m, Malaise trap, 23–27.iv.2017, A. Cornejo, T. Ríos, E. Álvarez, and C. Nieto, (COZEM).</p><p>Paratypes. PANAMA: Veraguas: Cuenca 097, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-81.16486&amp;materialsCitation.latitude=8.55038" title="Search Plazi for locations around (long -81.16486/lat 8.55038)">Río Calovebora</a>, 8.55038°N and 81.16486°W, PSPSCD- PNSF-C097-2017 - 006, 461 m, Malaise trap, 23–27.iv.2017, A. Cornejo, T. Ríos, E. Álvarez, and C. Nieto, 7 males (COZEM) ; ibid., Cuenca 132, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-81.1197&amp;materialsCitation.latitude=8.54513" title="Search Plazi for locations around (long -81.1197/lat 8.54513)">Río Mulaba–Isleta</a>, 8.54513°N and 81.1197°W, PSPSCD-PNSF- C132-2017-015, 412 m, Malaise trap, 2 males and 4 females (COZEM) ; ibid., Río Mulaba–afluente 1 er Brazo, 8.51706°N and 81.1214°W, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-81.1214&amp;materialsCitation.latitude=8.51706" title="Search Plazi for locations around (long -81.1214/lat 8.51706)">Control</a>: PSPSCD-PNSF-C132-2017-008, 770 m, UV light trap, 19.iv.2017, T. Ríos, E. Álvarez, and C. Nieto, 1 male (UMSP) .</p><p>Etymology. This new species is named for the Río Calovebora from which it was first collected. The name is formulated as a noun in apposition.</p></div>	https://treatment.plazi.org/id/677D87E1FFC13A07FF07236BFD3510E5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Blahnik, Roger J.;Armitage, Brian J.	Blahnik, Roger J., Armitage, Brian J. (2019): The Trichoptera of Panama. XII. Contributions to the family Glossosomatidae (Insecta: Trichoptera) in Panama. Insecta Mundi 740 (740): 1-17, DOI: 10.5281/zenodo.3676607
677D87E1FFC73A04FF072056FD3517A1.text	677D87E1FFC73A04FF072056FD3517A1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mortoniella (Mortoniella) yayas Blahnik and Armitage 2019	<div><p>Mortoniella (Mortoniella) yayas Blahnik and Armitage, new species</p><p>Fig. 3</p><p>Diagnosis. This is another species of the “florica subgroup” of Mortoniella . The species is not particularly distinctive, differing from other species of the “subgroup” mostly by its own peculiar combination of characters. It is probably most closely related to M. pectinella Blahnik and Holzenthal, also from Panama, resembling that species in the structure of tergum X, particularly in its prominent, acute lateral lobes and broadly triangular mesal projection. In other details, however, it is quite different, lacking the pectinate apical projections on the paramere appendages of M. pectinella, and in having acute, rather than rounded, apices to the inferior appendages. Other characters useful in diagnosing the new species include the relatively narrow dorsolateral lobes of the phallicata, the narrow, concavely excavated apices of the paramere appendages, and the generally weakly inflected and posteriorly oriented apex of the dorsal phallic spine.</p><p>Description. Length of forewing: male 3.5 mm. Forewing with forks I, II, and III present, hind wing with forks II and III. Spur formula 0:4:4. Overall color (in alcohol) brown, anastamosis unpigmented, but otherwise not evidently marked.</p><p>Male genitalia. Ventral process of segment VII laterally compressed, short, ventrally projecting, rounded apically, length slightly greater than width at base, process not retracted anterobasally. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin slightly rounded and produced dorsally, narrowing ventrally; segment deeply excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by much less than ½ width of segment. Tergum X moderately elongate, lateral margins subparallel, apically with broad, subtriangular mesal projection, subequal in length to apicolateral lobes; apicolateral lobes subtriangular, distinctly sclerotized, compressed, acute apically; ventrolateral lobes nearly obsolete; ventromesal lobes not evident. Inferior appendages without dorsolateral lobes, apicoventral lobes posteriorly projecting, relatively short, narrow, acute apically. Mesal pockets of inferior appendage with apical processes, short and weakly curved. Paramere appendage elongate, narrow, extending about same length as dorsal phallic spine, extreme apex concavely excavated on ventral margin. Dorsal phallic spine, as viewed laterally, undulate in contour, distinctly widened on ventral margin at about middle, ventral bulge articulating with dorsal margin of phallicata, apex acute, posteriorly directed; spine, in dorsal view (Fig. 3D), slightly widened in middle, narrowing apically, apex acute. Phallicata moderately elongate, dorsal margin with relatively narrow, paired, dorsolateral projections, lateral margin without compressed projection, ventral margin, as viewed ventrally (Fig. 3C), with elongate, weakly rounded lateral projections. Endophallic membrane simple, with single prominent curved ventromesal spine; phallotremal spines absent.</p><p>Female and immatures. Unknown.</p><p>Holotype male (alcohol). PANAMA: Cocle: Cuenca 134, Quebrada Las Yayas, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-80.5952&amp;materialsCitation.latitude=8.66168" title="Search Plazi for locations around (long -80.5952/lat 8.66168)">La Pintada</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-80.5952&amp;materialsCitation.latitude=8.66168" title="Search Plazi for locations around (long -80.5952/lat 8.66168)">El Harino</a>, PSPSCD-PNGDOTH-C134-2017-004, 8.66168°N and 80.5952°W, 602 m, 25.iii.2017, UV light trap, E. Álvarez, E. Pérez, and T. Ríos (COZEM).</p><p>Etymology. This new species is named after the Las Yayas stream from which it was collected. The name is formulated as a noun in apposition.</p></div>	https://treatment.plazi.org/id/677D87E1FFC73A04FF072056FD3517A1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Blahnik, Roger J.;Armitage, Brian J.	Blahnik, Roger J., Armitage, Brian J. (2019): The Trichoptera of Panama. XII. Contributions to the family Glossosomatidae (Insecta: Trichoptera) in Panama. Insecta Mundi 740 (740): 1-17, DOI: 10.5281/zenodo.3676607
677D87E1FFC43A04FF07269DFA6911E2.text	677D87E1FFC43A04FF07269DFA6911E2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protoptila Banks 1904	<div><p>Genus Protoptila</p><p>This genus is considered the sister taxon to Mortoniella (Robertson and Holzenthal 2013) . Both genera are characterized by having a pair of short appendages attached to the ventral margin of the phallobase, which engage with a hollowed receptacle on the mesal surface of the inferior appendages or phallic ensemble. This is a unique character, not otherwise found within the subfamily Protoptilinae . The genus has a number of distinctive apomorphies in its male genitalia, most distinctive of which are a ventral extension of sternum VIII beneath segment IX, and the structure of the phallic apparatus, which has an enlarged, blade-like apodeme basodorsally and an elongate and often arched phallicata, which is fused to the phallobase. Additionally, inferior appendages are generally absent or vestigial. Other characters, varying greatly among individual species, include the structure of the lateral lobes of tergum X and the structure of the paramere appendages. Nearly 100 species are currently known, making it the second largest genus in Protoptilinae (behind Mortoniella). Although the individual species are, in general, relatively easy to diagnose, because of the complexity and variability of structures of the genitalia, the genus as a whole has never been treated comparatively or comprehensively. Consequently, it is often difficult to determine relationships among species, or group of species. This is true of the species treated here, each of which is distinctive in its own right, but difficult to compare to other described species.</p></div>	https://treatment.plazi.org/id/677D87E1FFC43A04FF07269DFA6911E2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Blahnik, Roger J.;Armitage, Brian J.	Blahnik, Roger J., Armitage, Brian J. (2019): The Trichoptera of Panama. XII. Contributions to the family Glossosomatidae (Insecta: Trichoptera) in Panama. Insecta Mundi 740 (740): 1-17, DOI: 10.5281/zenodo.3676607
677D87E1FFC43A05FF07235FFCC9101B.text	677D87E1FFC43A05FF07235FFCC9101B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protoptila inflata Blahnik and Armitage 2019	<div><p>Protoptila inflata Blahnik and Armitage, new species</p><p>Fig. 4</p><p>Diagnosis. Protoptila inflata is best diagnosed by the relatively elongate, down-turned lateral lobes of tergum X, greatly inflated apex of the phallus (as viewed laterally), and hooked apices of the paramere appendages.</p><p>Description. Length of forewing: male 2.7–2.8 mm. Forewing with forks I, II, and III present; hind wing with fork II only, wing narrow and acute apically, costal margin with pronounced invagination. Spur formula 0:4:4. Overall color (in alcohol) brown (specimens cleared).</p><p>Male genitalia. Sternum VI process relatively prominent, posteriorly directed, longer than wide, apex subacute. Tergum VIII with row of elongate setae along posterior margin; sternum VIII moderately densely setose, setae elongate, posterior margin distinctly produced, slightly curved upward, extending beyond dorsolateral lobes of tergum X; as viewed ventrally, with ventral projection narrow, forked apically, apices of fork subacute. Segment IX with anterior margin broadly rounded, sternum mesally produced into short, acute process, posterolateral margin of segment IX without processes. Preanal appendages absent. Tergum X, as viewed laterally, with lateral lobes not obviously subsegmented, but somewhat bent or flexed basally, posterior margin of lateral lobe with small sclerotized mesal projection at flexion point, dorsal margin between lobes narrow and continuous, textured with minute spines, apex of mesal projection with short, bilobed, membranous projection; lateral lobes simple, moderately elongate and narrow, downturned, with several short marginal setae, apex narrowed and with ventrally-directed, spine-like projection. Phallobase dorsally with large, laterally compressed apodeme, ventrally with pair of short, rod-like, articulated appendages with membranous apices, appendages fitting into sclerotized pockets on ventral margin of phallobase, ventral margin of each pocket with short, hook-like projection. Phallicata, as viewed laterally, tubular, fused to phallobase, relatively short, dorsally arched, distinctly widened apically on ventral margin; as viewed dorsally, relatively narrow, apical enlargement slightly channeled dorsomesally, bilobed apically. Phallotremal sclerite not evident. Parameres each with “nested” membranous basal structure and relatively short, narrow, rod-like, sclerotized paramere appendage, appendage nearly straight, apex hooked or somewhat spirally curved.</p><p>Female and immatures. Unknown.</p><p>Holotype male (alcohol). PANAMA: Cocle: Cuenca 134, Quebrada Las Yayas, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-80.5952&amp;materialsCitation.latitude=8.66168" title="Search Plazi for locations around (long -80.5952/lat 8.66168)">La Pintada</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-80.5952&amp;materialsCitation.latitude=8.66168" title="Search Plazi for locations around (long -80.5952/lat 8.66168)">El Harino</a>, PSPSCD-PNGDOTH-C134-2017-004, 8.66168°N and 80.5952°W, 602 m, 22-26.iii.2017, Malaise trap, E. Álvarez, E. Pérez, and T. Ríos (COZEM).</p><p>Paratypes. Same data as holotype, 1 male (UMSP) .</p><p>Etymology. This new species is named for the inflated apex of the phallus in lateral view. The genus name is feminine and the species name adjectival.</p></div>	https://treatment.plazi.org/id/677D87E1FFC43A05FF07235FFCC9101B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Blahnik, Roger J.;Armitage, Brian J.	Blahnik, Roger J., Armitage, Brian J. (2019): The Trichoptera of Panama. XII. Contributions to the family Glossosomatidae (Insecta: Trichoptera) in Panama. Insecta Mundi 740 (740): 1-17, DOI: 10.5281/zenodo.3676607
677D87E1FFC53A0AFF072117FB841753.text	677D87E1FFC53A0AFF072117FB841753.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protoptila totumas Blahnik and Armitage 2019	<div><p>Protoptila totumas Blahnik and Armitage, new species</p><p>Fig. 5</p><p>Diagnosis. Protoptila totumas is distinctive and unlikely to be confused with any other described species. The most distinctive and usefully diagnostic feature of this new species is the structure of the paramere appendages, which are relatively elongate and narrow and each possesses a corkscrew-like spiral at midlength, subtending and crossing over each other beneath the phallicata, with the apices dorsal to the phallicata. Although the exact structure is somewhat variable in the material examined, the symmetrical arrangement of the appendages, as in Fig. 5E, is probably the most common configura- tion. Other usefully diagnostic features include the structure of tergum X, which has relatively elongate, narrow lateral lobes, each with two acute mesal projections, one in the basal half of the lobe (not readily observed in lateral view), and the other preapically, and also the structure of the phallicata, which is relatively narrow and arched dorsally, as viewed laterally, but is distinctively bulged in its basal half, as viewed dorsally, and has acute apicolateral projections.</p><p>Description. Length of forewing: male 3.2 mm. Forewing with forks I, II, and III present; hind wing with fork II only, wing narrow and acute apically, costal margin with pronounced invagination. Spur formula 0:4:4. Overall color (in alcohol) brown, anastamosis unpigmented, but otherwise not evidently marked.</p><p>Male genitalia. Sternum VI process relatively prominent, posteriorly directed, slightly curved, longer than wide, apex subacute. Tergum VIII with row of elongate setae along posterior margin; sternum VIII densely setose with elongate setae, posterior margin distinctly produced, slightly curved upward, extending nearly as far as dorsolateral lobes of tergum X; as viewed ventrally, with ventral projection narrow, weakly and bluntly forked apically. Segment IX with anterior margin broadly rounded, sternum mesally produced into short, acute process, with single apical seta, posterolateral margin of segment IX without processes. Preanal appendages absent. Tergum X, as viewed laterally, with lateral lobes not obviously subsegmented, each lobe relatively elongate and narrow, downturned, forked preapically, ventral branch forming mesally directed, trianguloid projection with spine-like apex, dorsal branch extending as narrow, apically acute, finger-like projection; as viewed dorsally, with additional acute, mesal projection in basal half, not readily observable in lateral view. Phallobase dorsally with large, laterally compressed apodeme, ventrally with pair of short, rod-like, articulated appendages with membranous apices, appendages fitting into sclerotized pockets on ventral margin of phallobase. Phallicata, as viewed laterally, tubular, fused to phallobase, dorsally arched, slightly widened preapically; as viewed dorsally, with lateral margins distinctly rounded and widened in basal half, narrowing apically, preapically with acute, trianguloid, lateral projections on either side. Phallotremal sclerite forming curved, narrow, tube-like structure, upturned as viewed laterally. Parameres each with membranous basal structure and elongate, narrow, sclerotized paramere appendage, with a helical spiral past midlength; appendages apparently crossing over each other beneath phallicata and converging dorsally, apices relatively straight and acute.</p><p>Female and immatures. Unknown.</p><p>Holotype male (alcohol). PANAMA: Chiriquí: Cuenca 102, afluente de Río Colorado, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.684074&amp;materialsCitation.latitude=8.884717" title="Search Plazi for locations around (long -82.684074/lat 8.884717)">Mount Totumas Biological Reserve</a>, 8.884717°N and 82.684077°W, 1950 m, 11–17.i.2016, Malaise trap, B. Armitage and J. Dietrich (COZEM).</p><p>Paratypes. Same locality as holotype, 7–13.xii.2015, Malaise trap, J. Dietrich, 2 males (COZEM, UMSP) .</p><p>Etymology. This new species is named after the Mount Totumas Cloud Forest and Biological Reserve from which it was collected. The name is formulated as a noun in apposition.</p></div>	https://treatment.plazi.org/id/677D87E1FFC53A0AFF072117FB841753	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Blahnik, Roger J.;Armitage, Brian J.	Blahnik, Roger J., Armitage, Brian J. (2019): The Trichoptera of Panama. XII. Contributions to the family Glossosomatidae (Insecta: Trichoptera) in Panama. Insecta Mundi 740 (740): 1-17, DOI: 10.5281/zenodo.3676607
677D87E1FFCA3A0BFF0726CFFE1F1606.text	677D87E1FFCA3A0BFF0726CFFE1F1606.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protoptila rambala Blahnik and Armitage 2019	<div><p>Protoptila rambala Blahnik and Armitage, new species</p><p>Fig. 6</p><p>Diagnosis. The most useful diagnostic features of Protoptila rambala include its relatively short paramere appendages, with broad brush-like apices (Fig. 6B); the structure of the lateral lobes of tergum X (Fig. 6A), which are more or less jointed in their basal half and have their apices shaped somewhat like a bird’s head (with a downward-pointed, acute “beak”); and the forked ventromesal apex of segment IX, which is nearly as long as the subtending and also apically forked ventral projection of sternum VIII (Fig. 6C).</p><p>Description. Length of forewing: male 2.5 mm. Forewing with forks I, II, and III present; hind wing with fork II only, wing narrow and acute apically, costal margin with pronounced invagination. Spur formula 0:4:4. Color apparently brown (specimens completely cleared, in alcohol).</p><p>Male genitalia. Sternum VI process relatively prominent, posteriorly directed, longer than wide, apex subacute. Tergum VIII with row of setae along posterior margin; sternum VIII densely setose with relatively elongate setae, posterior margin distinctly produced, slightly curved upward, extending nearly as far as dorsolateral lobes of tergum X; as viewed ventrally, with ventral projection relatively narrow, weakly and bluntly forked apically. Segment IX with anterior margin broadly rounded, sternum mesally produced into forked process, slightly wider than and extending nearly as far as ventral lobe of sternum VIII, posterolateral margin of segment IX with weakly sclerotized, broadly rounded, dorsolateral lobe on either side. Preanal appendages absent. Tergum X, as viewed laterally, with lateral lobes jointed (apparently subsegmented), basal lobe short, only slightly longer than width, apical lobe broadly rounded basally, apically narrowed and appearing like a bird’s head, with downward pointing, acute apical “beak,” apex (head) of lobe with small sensilla; tergum with bilobed membranous mesal projection between lateral lobes. Phallobase dorsally with large, laterally compressed apodeme, ventrally with pair of short, rod-like, articulated appendages with membranous apices, appendages fitting into sclerotized pockets on ventral margin of phallobase. Phallicata, as viewed laterally, narrow, tubular, fused to phallobase, projecting nearly straight, apparently with widened lateral flanges (difficult to see). Endophallic membrane narrow and membranous, phallotremal sclerite rather indistinct. Parameres each with membranous basal structure and relatively short sclerotized paramere appendage, appendage somewhat rod-like, broadened apically into brush-like structure with narrow spines.</p><p>Female and immatures. Unknown.</p><p>Holotype male (alcohol). PANAMA: Bocas del Toro: Quebrada Rambala, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.15469&amp;materialsCitation.latitude=8.91627" title="Search Plazi for locations around (long -82.15469/lat 8.91627)">Rambala Jungle Lodge</a>, 3.74 km SSE Rambala, 8.91627°N and 82.15469°W, 120 m, 21-31.xii.2016, Malaise trap, Eric Carlson (COZEM).</p><p>Paratypes. Same locality as holotype, 6-12.ii.2017, Malaise trap, Eric Carlson, 1 male (COZEM).</p><p>Etymology. This new species is named after Quebrada Rambala from whence it was collected. The name is formulated as a noun in apposition.</p><p>New Country Records</p></div>	https://treatment.plazi.org/id/677D87E1FFCA3A0BFF0726CFFE1F1606	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Blahnik, Roger J.;Armitage, Brian J.	Blahnik, Roger J., Armitage, Brian J. (2019): The Trichoptera of Panama. XII. Contributions to the family Glossosomatidae (Insecta: Trichoptera) in Panama. Insecta Mundi 740 (740): 1-17, DOI: 10.5281/zenodo.3676607
677D87E1FFCB3A0BFF072764FD9616D1.text	677D87E1FFCB3A0BFF072764FD9616D1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Culoptila costaricensis Flint 1974	<div><p>Culoptila costaricensis Flint, 1974</p><p>Material examined (alcohol). PANAMA, Chiriquí Province, Cuenca 108, tributary of Quebrada Grande, Boquete, Valle Escondido, 8.78291 ° N and 82.44579°W, 1253 m, UV light trap, B. Armitage and T. Arefina-Armitage, 11 March 2018, 1 male (COZEM).</p><p>Distribution. Costa Rica, Panama.</p></div>	https://treatment.plazi.org/id/677D87E1FFCB3A0BFF072764FD9616D1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Blahnik, Roger J.;Armitage, Brian J.	Blahnik, Roger J., Armitage, Brian J. (2019): The Trichoptera of Panama. XII. Contributions to the family Glossosomatidae (Insecta: Trichoptera) in Panama. Insecta Mundi 740 (740): 1-17, DOI: 10.5281/zenodo.3676607
677D87E1FFCB3A0BFF0727BDFD9617AE.text	677D87E1FFCB3A0BFF0727BDFD9617AE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mortoniella opinionis Blahnik and Holzenthal 2008	<div><p>Mortoniella opinionis Blahnik and Holzenthal, 2008</p><p>Material examined (alcohol). PANAMA, Chiriquí Province, Cuenca 108, Volcan Baru National Park, Quebrada Cascante, PSPSCD-PNVB-C108-2017-018, 8.84939°N and 82.49349°W, 1947 m, Malaise trap, T. Ríos, E. Álvarez, and E. Pérez, 5-8 June 2017, 1 male (COZEM).</p><p>Distribution. Costa Rica, Panama.</p></div>	https://treatment.plazi.org/id/677D87E1FFCB3A0BFF0727BDFD9617AE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Blahnik, Roger J.;Armitage, Brian J.	Blahnik, Roger J., Armitage, Brian J. (2019): The Trichoptera of Panama. XII. Contributions to the family Glossosomatidae (Insecta: Trichoptera) in Panama. Insecta Mundi 740 (740): 1-17, DOI: 10.5281/zenodo.3676607
677D87E1FFCB3A0BFF072692FD961084.text	677D87E1FFCB3A0BFF072692FD961084.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protoptila spirifera Flint 1974	<div><p>Protoptila spirifera Flint, 1974</p><p>Material examined (alcohol). PANAMA, Chiriquí Province, Cuenca 102, La Amistad International Park, near Mount Totumas, high altitude bog, 8.89044°N and 82.66684°W, 2075 m, UV light trap, J. Dietrich, 16 May 2015, 1 male (COZEM) .</p><p>Distribution. Costa Rica, Panama.</p></div>	https://treatment.plazi.org/id/677D87E1FFCB3A0BFF072692FD961084	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Blahnik, Roger J.;Armitage, Brian J.	Blahnik, Roger J., Armitage, Brian J. (2019): The Trichoptera of Panama. XII. Contributions to the family Glossosomatidae (Insecta: Trichoptera) in Panama. Insecta Mundi 740 (740): 1-17, DOI: 10.5281/zenodo.3676607
