taxonID	type	description	language	source
673287C8FFE21A7EFC504B78FB0FFBC2.taxon	description	(Figs 3 A; 4 A)	en	Gilgado, José D., Ortuño, Vicente M. (2023): Polydesmid millipedes in the MSS of the northern Iberian Peninsula (Diplopoda, Polydesmida, Polydesmidae). Zoosystema 45 (15): 451-461, DOI: 10.5252/zoosystema2023v45a15
673287C8FFE21A7EFC504B78FB0FFBC2.taxon	materials_examined	MATERIAL EXAMINED. — Spain • 1 ♂, 3 ♀; Burgos, Pineda de la Sierra, Sierra de Mencilla in Sierra de la Demanda, beech forest; 42 ° 11 ’ 55 ” N, 03 ° 18 ’ 41 ” W; 1460 m a. s. l.; 22. VI. 2019; V. M. Ortuño, J. D. Gilgado & E. Ledesma leg.; MSS; UAH • 1 ♂; same data; MNHN • 1 ♂; same data; 18. VI. 2018; UAH.	en	Gilgado, José D., Ortuño, Vicente M. (2023): Polydesmid millipedes in the MSS of the northern Iberian Peninsula (Diplopoda, Polydesmida, Polydesmidae). Zoosystema 45 (15): 451-461, DOI: 10.5252/zoosystema2023v45a15
673287C8FFE21A7EFC504B78FB0FFBC2.taxon	discussion	REMARKS This species was previously known only from two records in the Balneario de Panticosa (Ceuca 1968) and a mixed forest near San Juan de la Peña, both in Huesca (Serra et al. 1996; Gilgado et al. 2015). Both localities are in the Pyrenean region, and the present record extends the range of the species to the north of the Iberian System. The species could therefore also occur in other nearby mountains such as the Basque Mountains, or the Cantabrian Range.	en	Gilgado, José D., Ortuño, Vicente M. (2023): Polydesmid millipedes in the MSS of the northern Iberian Peninsula (Diplopoda, Polydesmida, Polydesmidae). Zoosystema 45 (15): 451-461, DOI: 10.5252/zoosystema2023v45a15
673287C8FFE01A7FFEE74C5CFC89F983.taxon	description	(Figs 3 B; 4 B)	en	Gilgado, José D., Ortuño, Vicente M. (2023): Polydesmid millipedes in the MSS of the northern Iberian Peninsula (Diplopoda, Polydesmida, Polydesmidae). Zoosystema 45 (15): 451-461, DOI: 10.5252/zoosystema2023v45a15
673287C8FFE01A7FFEE74C5CFC89F983.taxon	materials_examined	MATERIAL EXAMINED. — Spain • 1 ♀; Palencia, Piedras Luengas, Hoyo de los Lobos, Las Lombas, Forest; 43 ° 02 ’ 42 ” N, 04 ° 28 ’ 16 ” W; 1300 m a. s. l.; 20. VI. 2016; V. M. Ortuño, J. D. Gilgado & E. Ledesma leg.; MSS; UAH • 1 ♂; same data; 25. VI. 2019; UAH • 1 ♂; Palencia, Sierra de Peña Labra, Piedras Luengas, bare scree; 43 ° 02 ’ 21 ” N, 04 ° 26 ’ 55 ” W; 1520 m a. s. l.; 21. VI. 2018; V. M. Ortuño, J. D. Gilgado & E. Ledesma leg.; MSS; UAH • 3 ♂; Cantabria, Pesaguero, Sierra Mediana, Collado de la Cruz de las Cabezuelas, forest; 43 ° 04 ’ 19 ” N, 04 ° 28 ’ 18 ” W; 1050 m a. s. l.; 27. VI. 2017; V. M. Ortuño, J. D. Gilgado & E. Ledesma leg; MSS; UAH • 1 ♂, 1 ♀; same data; MNHN • 5 ♂, 3 ♀; same data; 31. X. 2018; UAH • 33 ♂, 7 ♀; same data; 21. VI. 2018; UAH • 9 ♂, 6 ♀; same data; 22. VI. 2019; UAH • 121 ♂, 20 ♀; same data; 1. IX. 2021; UAH.	en	Gilgado, José D., Ortuño, Vicente M. (2023): Polydesmid millipedes in the MSS of the northern Iberian Peninsula (Diplopoda, Polydesmida, Polydesmidae). Zoosystema 45 (15): 451-461, DOI: 10.5252/zoosystema2023v45a15
673287C8FFE01A7FFEE74C5CFC89F983.taxon	discussion	REMARKS The species has a wide distribution in western Europe, namely in Germany, Switzerland, the United Kingdom and France, including the northern part of the Pyrenees, and several other countries (Kime & Enghoff 2011). This species was reported in Spain for the first time by Mauriès (1971), in the province of Lerida, in the Catalonian Pyrenees, and according to Kime & Enghoff (2011) the species had not been recorded in the Atlantic zone of Spain. Djusrvoll (2019) states that this species occurs in the Atlantic zone, in the provinces of Alava and Asturias, but he provides no capture information or bibliographic references. The present record from Pesaguero confirms the occurence of the species in the Atlantic region of Spain. However, it is not clear whether this region was part of its native distribution range, whether it has been recently colonized due to natural dispersal of the species (Hauser & Voigtländer 2009) or whether it is the result of anthropogenic introduction events (it has been introduced in North America, see for example Shelley (2002) or Kime & Enghoff (2011 )). Interestingly, Polydesmus angustus was found in syntopy with P. coriaceus, the most common polydesmid in the present study.	en	Gilgado, José D., Ortuño, Vicente M. (2023): Polydesmid millipedes in the MSS of the northern Iberian Peninsula (Diplopoda, Polydesmida, Polydesmidae). Zoosystema 45 (15): 451-461, DOI: 10.5252/zoosystema2023v45a15
673287C8FFE01A7FFF0448B8FB17FD9D.taxon	description	(Figs 3 C; 5 A-C)	en	Gilgado, José D., Ortuño, Vicente M. (2023): Polydesmid millipedes in the MSS of the northern Iberian Peninsula (Diplopoda, Polydesmida, Polydesmidae). Zoosystema 45 (15): 451-461, DOI: 10.5252/zoosystema2023v45a15
673287C8FFE01A7FFF0448B8FB17FD9D.taxon	materials_examined	MATERIAL EXAMINED. — Spain • 2 ♂; Cantabria, Barrio, monte Casanzo, Llobango, forested scree; 43 ° 03 ’ 40 ” N, 04 ° 39 ’ 42 ” W; 1300 m a. s. l.; 23. VI. 2019; V. M. Ortuño, J. D. Gilgado & E. Ledesma leg.; MSS; UAH • 1 ♂; same data; MNHN • 2 ♂; same data; 27. VI. 2017; UAH.	en	Gilgado, José D., Ortuño, Vicente M. (2023): Polydesmid millipedes in the MSS of the northern Iberian Peninsula (Diplopoda, Polydesmida, Polydesmidae). Zoosystema 45 (15): 451-461, DOI: 10.5252/zoosystema2023v45a15
673287C8FFE01A7FFF0448B8FB17FD9D.taxon	discussion	REMARKS This species was recently described from nine specimens collected in a cave in Teverga, and in Vega de Enol, both in Asturias Province (Djursvoll 2019). The present record in Cantabria province is 35 km southeast from the closest locality, so it only slightly extends the known range of the species. The presence of Polydesmus asturiensis in a cave and the MSS, indicate at least a partially subterranean lifestyle. The morphology of the gonopod of the captured specimens corresponds to the original description (Djursvoll 2019), but the exomere seems to be slightly longer and more twisted. However, observation with different angles of tilt may cause different impressions (Fig. 5 A-C).	en	Gilgado, José D., Ortuño, Vicente M. (2023): Polydesmid millipedes in the MSS of the northern Iberian Peninsula (Diplopoda, Polydesmida, Polydesmidae). Zoosystema 45 (15): 451-461, DOI: 10.5252/zoosystema2023v45a15
673287C8FFE01A70FC4E4CBDFB8FFBC2.taxon	description	(Figs 3 D; 4 C)	en	Gilgado, José D., Ortuño, Vicente M. (2023): Polydesmid millipedes in the MSS of the northern Iberian Peninsula (Diplopoda, Polydesmida, Polydesmidae). Zoosystema 45 (15): 451-461, DOI: 10.5252/zoosystema2023v45a15
673287C8FFE01A70FC4E4CBDFB8FFBC2.taxon	materials_examined	MATERIAL EXAMINED. — Spain • 8 ♂, 8 ♀, 9 juv; León, Boca de Huérgano, Portilla de la Reina, Calar de San Martino, Forested scree; 43 ° 04 ’ 60 ” N, 04 ° 51 ’ 31 ” W; 1410 m a. s. l., 28. VI. 2017.; V. M. Ortuño, J. D. Gilgado & E. Ledesma leg.; MSS; UAH • 2 ♂, 2 ♀, 51 juv.; same data; 20. VI. 2018; UAH • 2 ♀, 1 juv.; Cantabria, Barrio, monte Casanzo, Llobango, forested scree; 43 ° 03 ’ 40 ” N, 04 ° 39 ’ 42 ” W; 1300 m a. s. l.; 19. VI. 2018; V. M. Ortuño, J. D. Gilgado & E. Ledesma leg.; MSS; UAH • 2 ♂, 1 ♀; same data; 23. VI. 2019; UAH • 5 ♂, 1 ♀; Palencia, Piedras Luengas, Hoyo de los Lobos, Las Lombas, forest; 43 ° 02 ’ 42 ” N, 04 ° 28 ’ 16 ” W; 1300 m a. s. l.; 16. XI. 2015; V. M. Ortuño, J. D. Gilgado & E. Ledesma leg.; UAH • 1 ♂, 2 ♀; Palencia, Piedras Luengas, Sierra de Peña Labra, bare scree; 43 ° 02 ’ 21 ” N, 04 ° 26 ’ 55 ” W; 1520 m a. s. l.; 27. VI. 2017; V. M. Ortuño, J. D. Gilgado & E. Ledesma leg.; MSS; UAH • 3 ♂, 6 ♀; same data; 21. VI. 2018; UAH • 2 ♀; Cantabria, Vega de Liébana, Dobres, Cucayo, below Pico Palanca, scree with trees beside river Requejada; 43 ° 02 ’ 55 ” N, 04 ° 38 ’ 08 ” W; 1150 m a. s. l.; 27. VI. 2017; V. M. Ortuño, J. D. Gilgado & E. Ledesma leg.; MSS • 2 ♂, 3 ♀, 7 juv.; same data; 19. VI. 2018; UAH • 3 ♂, 2 ♀, 3 juv.; same data; 23. VI. 2019; UAH • 1 ♀, 1 juv.; León, Boca de Huérgano, Llánaves de la Reina, Boquerón de Tarna, scree; 43 ° 03 ’ 23 ” N, 04 ° 46 ’ 03 ” W; 1710 m a. s. l., 28. VI. 2017; V. M. Ortuño, J. D. Gilgado & E. Ledesma leg.; MSS; UAH • 1 ♂, 2 ♀; 1 juv; same data; 20. VI. 2018; UAH • 1 ♂; Cantabria, Pesaguero, Sierra Mediana, Collado de la Cruz de las Cabezuelas, forest; 43 ° 04 ’ 19 ” N, 04 ° 28 ’ 18 ” W; 1050 m a. s. l.; 22. VI. 2019; V. M. Ortuño, J. D. Gilgado & E. Ledesma leg.; MSS, UAH • 4 ♀; same data; 1. IX. 2021; UAH • 21 ♂, 69 ♀, 96 juv.; Huesca, Longás, Sierra de Santo Domingo, Portillo de Longás, bare scree; 42 ° 26 ’ 45 ” N, 00 ° 55 ’ 60 ” W; 1200 m a. s. l.; 2. IX. 2015; V. M. Ortuño, J. D. Gilgado, E. Ledesma & E. Cuesta leg.; MSS; UAH • 8 ♂, 5 ♀, 8 juv.; same data; 21. XII. 2015; UAH • 8 ♂, 4 ♀, 1 juv; same data; 13. IV. 2016; UAH • 9 ♂, 18 ♀, 2 juv.; same data; 25. VII. 2016; UAH • 118 ♂, 101 ♀, 337 juv.; Huesca, Longás, Sierra de Santo Domingo, Portillo de Longás, forested scree; 42 ° 26 ’ 55 ” N, 00 ° 56 ’ 11 ” W; 1220 m a. s. l.; 2. IX. 2015; V. M. Ortuño, J. D. Gilgado, E. Ledesma & E. Cuesta leg.; MSS; UAH • 18 ♂, 5 ♀, 3 juv.; same data; 21. XII. 2015; UAH • 6 ♂, 2 juv.; same data; 13. IV. 2016; UAH • 59 ♂, 59 ♀, 49 juv.; same data; 25. VII. 2016; UAH • 4 ♂, 3 ♀, 2 juv.; Burgos, Pineda de la Sierra, Sierra de Mencilla in Sierra de la Demanda, bare scree in Glaciar Circus; 42 ° 11 ’ 21 ” N, 03 ° 18 ’ 37 ” W; 1750 m a. s. l.; 22. VI. 2019; V. M. Ortuño, J. D. Gilgado & E. Ledesma leg.; MSS; UAH • 1 ♀; Burgos, Pineda de la Sierra, Sierra de Mencilla in Sierra de la Demanda, beech forest; 42 ° 11 ’ 55 ” N, 03 ° 18 ’ 41 ” W; 1460 m a. s. l.; 18. VI. 2018; V. M. Ortuño, J. D. Gilgado & E. Ledesma leg.; MSS; UAH • 92 ♂, 62 ♀; same data; 22. VI. 2019; UAH • 18 ♂, 16 ♀; same data; 3. VIII. 2020; UAH • 10 ♂, 15 ♀, 2 juv.; Zaragoza, Añón de Moncayo, Sierra del Moncayo, Collado Bellido, bare scree; 41 ° 46 ’ 1 ” N, 01 ° 47 ’ 01 ” W; 1540 m a. s. l.; 12. III. 2014; V. M. Ortuño, J. D. Gilgado, E. Ledesma & E. Cuesta leg.; MSS; UAH • 41 ♂, 31 ♀, 36 juv.; same data; 2. VII. 2014; UAH • 3 ♂, 2 ♀; same data; MNHN • 2 ♂; Zaragoza, Tarazona, Sierra del Moncayo, Barranco de Castilla, beech forest; 41 ° 48 ’ 18 ” N, 01 ° 50 ’ 37 ” W; 1420 m a. s. l.; V. M. Ortuño, J. D. Gilgado, E. Ledesma & E. Cuesta leg.; MSS; UAH • 9 ♂, 2 ♀, 6 juv.; same data; 2. VII. 2014; UAH.	en	Gilgado, José D., Ortuño, Vicente M. (2023): Polydesmid millipedes in the MSS of the northern Iberian Peninsula (Diplopoda, Polydesmida, Polydesmidae). Zoosystema 45 (15): 451-461, DOI: 10.5252/zoosystema2023v45a15
673287C8FFE01A70FC4E4CBDFB8FFBC2.taxon	discussion	REMARKS This species is very common in the northern Iberian Peninsula, and in some Atlantic regions of Europe such as France, Belgium and the United Kingdom. It occurs in different habitat types and is not rare in caves, probably due to its preference for humid places (Kime & Enghoff 2011). This also agrees with its presence in the MSS in forested screes, as shown in the present study and in Eusébio et al. (2021).	en	Gilgado, José D., Ortuño, Vicente M. (2023): Polydesmid millipedes in the MSS of the northern Iberian Peninsula (Diplopoda, Polydesmida, Polydesmidae). Zoosystema 45 (15): 451-461, DOI: 10.5252/zoosystema2023v45a15
673287C8FFEE1A71FF1B4D9DFE14F8A1.taxon	description	(Figs 3 E; 5 D-F)	en	Gilgado, José D., Ortuño, Vicente M. (2023): Polydesmid millipedes in the MSS of the northern Iberian Peninsula (Diplopoda, Polydesmida, Polydesmidae). Zoosystema 45 (15): 451-461, DOI: 10.5252/zoosystema2023v45a15
673287C8FFEE1A71FF1B4D9DFE14F8A1.taxon	materials_examined	MATERIAL EXAMINED. — Spain • 2 ♂, 1 ♀; Burgos, Pineda de la Sierra, Sierra de Mencilla in Sierra de la Demanda, beech forest; 42 ° 11 ’ 55 ” N, 03 ° 18 ’ 41 ” W; 1460 m a. s. l.; 18. VI. 2018; V. M. Ortuño, J. D. Gilgado & E. Ledesma leg.; MSS; UAH • 39 ♂, 1 ♀; same data; 22. VI. 2019 • 3 ♂; same data; 3. VIII. 2020; UAH • 2 ♂, 1 ♀; same data; MNHN.	en	Gilgado, José D., Ortuño, Vicente M. (2023): Polydesmid millipedes in the MSS of the northern Iberian Peninsula (Diplopoda, Polydesmida, Polydesmidae). Zoosystema 45 (15): 451-461, DOI: 10.5252/zoosystema2023v45a15
673287C8FFEE1A71FF1B4D9DFE14F8A1.taxon	discussion	REMARKS This species is known from the French Pyrenees and northern Spain (provinces of Biscay, Navarre and Gipuzkoa) (Kime & Enghoff 2011; Djursvoll 2019). The present record extends the known distribution of the species to the south and east. It also occurs at a higher altitude than previous records, according to Kime & Enghoff (2011) up to 1300 m a. s. l. As Djursvoll (2019) points out, the gonopod morphology of P. racovitzai may resemble that of Polydesmus inconstans Latzel, 1884, as drawn by Demange (1981). The drawing in the original description of P. racovitzai (Brölemann 1910: plate V, figs 32, 33) shows a shorter and softly curved tip of exomerite, while it is longer and almost describing a straight angle after the lateral tooth in the drawing of Demange (1981: 125, figs 170 - 171). Our specimens agree with the gonopod morphology of P. racovitzai in Demange (1981), and we are including three images of it (Fig. 5 D-F).	en	Gilgado, José D., Ortuño, Vicente M. (2023): Polydesmid millipedes in the MSS of the northern Iberian Peninsula (Diplopoda, Polydesmida, Polydesmidae). Zoosystema 45 (15): 451-461, DOI: 10.5252/zoosystema2023v45a15
673287C8FFEE1A71FF064859FEA4FBC2.taxon	description	(Figs 3 F; 4 D)	en	Gilgado, José D., Ortuño, Vicente M. (2023): Polydesmid millipedes in the MSS of the northern Iberian Peninsula (Diplopoda, Polydesmida, Polydesmidae). Zoosystema 45 (15): 451-461, DOI: 10.5252/zoosystema2023v45a15
673287C8FFEE1A71FF064859FEA4FBC2.taxon	materials_examined	MATERIAL EXAMINED. — Spain • 1 ♂, 1 ♀; Huesca, Longás, Sierra de Santo Domingo, Portillo de Longás, bare scree; 42 ° 26 ’ 45 ” N, 00 ° 55 ’ 60 ” W; 1200 m a. s. l.; 2. IX. 2015; V. M. Ortuño, J. D. Gilgado, E. Ledesma & E. Cuesta leg.; MSS; UAH • 2 ♀; same data; 25. VII. 2016; UAH • 7 ♂, 2 ♀, 16 juv.; Zaragoza, Luesia, Sierra de Luesia, Galapatizos gully, alluvial debris; 42 ° 24 ’ 48 ” N, 01 ° 00 ’ 36 ” W; 790 m a. s. l.; 2. IX. 2015; V. M. Ortuño, J. D. Gilgado, E. Ledesma & E. Cuesta leg.; MSS; UAH • 4 ♂, 6 ♀, 26 juv.; same data; 27. VI. 2016; UAH • 3 ♂, 3 juv.; Zaragoza, Luesia, Arba de Luesia river, alluvial debris; 42 ° 24 ’ 21 ” N, 01 ° 01 ’ 10 ” W; 770 m a. s. l.; 2. IX. 2015; V. M. Ortuño, J. D. Gilgado, E. Ledesma & E. Cuesta leg.; MSS; UAH • 2 ♂; same data; 21. XII. 2015; UAH • 2 ♂, 2 ♀; same data; MNHN • 1 ♂, 1 ♀, 2 juv.; same data; 25. VII. 2016; UAH.	en	Gilgado, José D., Ortuño, Vicente M. (2023): Polydesmid millipedes in the MSS of the northern Iberian Peninsula (Diplopoda, Polydesmida, Polydesmidae). Zoosystema 45 (15): 451-461, DOI: 10.5252/zoosystema2023v45a15
673287C8FFEE1A71FF064859FEA4FBC2.taxon	discussion	REMARKS This species is considered as troglophile. It has been recorded in Italy, Spain, including the Balearic and Canary Islands, and Algeria. Its range is wide in continental Spain, but there are few records, probably due to lack of sampling (Kime & Enghoff 2011).	en	Gilgado, José D., Ortuño, Vicente M. (2023): Polydesmid millipedes in the MSS of the northern Iberian Peninsula (Diplopoda, Polydesmida, Polydesmidae). Zoosystema 45 (15): 451-461, DOI: 10.5252/zoosystema2023v45a15
