taxonID	type	description	language	source
BD97B648DB8E5EE092CD99438639F6EE.taxon	description	Figures 1, 2, 3	en	Julius, Avelinah (2022): Capparis (Capparaceae) in Peninsular Malaysia, including a new species and two new varieties. PhytoKeys 189: 99-127, DOI: http://dx.doi.org/10.3897/phytokeys.189.49367, URL: http://dx.doi.org/10.3897/phytokeys.189.49367
BD97B648DB8E5EE092CD99438639F6EE.taxon	diagnosis	Diagnosis. Vegetatively, this new species resembles Capparis buwaldae Jacobs in having bullate leaves with a long acuminate-caudate apex and distinct reticulation, but C. kenaboiensis differs from C. buwaldae in its terminal (vs. supra-axillary) inflorescences, the absent (vs. present) stem stomata and the smooth (vs. tuberculate) fruit pericarp. The flowers subtended by leaf-like bracts resemble Capparis scortechinii, but are early caducous in the new species.	en	Julius, Avelinah (2022): Capparis (Capparaceae) in Peninsular Malaysia, including a new species and two new varieties. PhytoKeys 189: 99-127, DOI: http://dx.doi.org/10.3897/phytokeys.189.49367, URL: http://dx.doi.org/10.3897/phytokeys.189.49367
BD97B648DB8E5EE092CD99438639F6EE.taxon	materials_examined	Type. Peninsular Malaysia. Negeri Sembilan: Jelebu, Kenaboi FR, Gunung Besar Hantu, road sides towards Lata Kijang, 3 ° 11.42 ' N, 100 ° 59.21 ' E, 459 m alt., 4 Mar 2014 (fl., fr.), Julius et al. FRI 57784 (holotype KEP!; isotypes K!, SAN!, SING!, L!, A!).	en	Julius, Avelinah (2022): Capparis (Capparaceae) in Peninsular Malaysia, including a new species and two new varieties. PhytoKeys 189: 99-127, DOI: http://dx.doi.org/10.3897/phytokeys.189.49367, URL: http://dx.doi.org/10.3897/phytokeys.189.49367
BD97B648DB8E5EE092CD99438639F6EE.taxon	description	Description. Climber 2 - 8 (- 12) m long hanging high up on tree. Twigs straight, pubescent when young and soon glabrous. Stipular thorns retrorse, tips 1 - 2 mm long. Leaves spirally arranged; petioles 8 - 12 x ca. 1 mm, grooved, slender, not thickened, covered with dense, short, silky white hairs; laminas narrowly elliptic, (5.5 -) 10 - 11.5 x (1.5 -) 3.5 - 4 cm, chartaceous, bullate, fresh dark green and glossy above, pale green beneath, brownish-green when dry, base acute, margin revolute and entire, apex long acuminate to caudate, with acumen 1 - 1.5 cm long, glabrous above, hairy beneath, denser on mid-rib and venation; mid-rib sunken above, raised beneath; lateral veins 5 - 7 pairs, arcuate towards the margin; intercostal veins reticulate, distinct above, prominent beneath. Inflorescences axillary or terminal, racemose, rachis slender and flexuous, conferted towards the top as the buds fall off at bottom part and leave scars, velvety, white hairs throughout; bracts leaf-like, elliptic, 10 - 25 x 2 - 5 mm, velvety white hairs on both surfaces, early caducous. Flowers loosely arranged, 10 - 14, buds globose, 4 - 5 x 3 - 5 mm; pedicels 2 - 2.5 cm long, whitish; sepals 4, thin, cucullate, whitish-green, except reddish at base inside, keeled, 2 pairs, lanceolate, 6 - 8 x ca. 5 mm, outer pair larger than inner pair, the outer sepals imbricate, covered with dense, white hairs outside, glabrous inside, the inner pair hairy on keel outside, glabrous inside; petals 4, upper pair pinkish and white along margin and laterally connate at base, lower pair whitish-green and free, elliptic ca. 10 x 3 mm, inside densely, silky tomentose hairy, outside glabrous, except densely hairy at base, ciliate along the margin; stamens 18 - 31, unequal in length, filament whitish, 7 - 20 mm long, glabrous, except hairy at base, anthers dorsifixed, ca. 1.5 mm long, greenish, apex recurved; ovary ellipsoid, ca. 1.5 mm long, greenish, on gynophore 2 - 2.5 cm long, stigma obscure. Fruits young green turning brownish to dark purple when ripe, subglobose to pyriform on slender gynophore, 4.5 - 5 x 5.5 - 6 cm, pulp pinkish or purplish-red. Seeds 1 - 4, sarcotesta fleshy, yellowish, testa thin and black.	en	Julius, Avelinah (2022): Capparis (Capparaceae) in Peninsular Malaysia, including a new species and two new varieties. PhytoKeys 189: 99-127, DOI: http://dx.doi.org/10.3897/phytokeys.189.49367, URL: http://dx.doi.org/10.3897/phytokeys.189.49367
BD97B648DB8E5EE092CD99438639F6EE.taxon	distribution	Distribution. Endemic in Peninsular Malaysia and known only from the type locality (Fig. 1).	en	Julius, Avelinah (2022): Capparis (Capparaceae) in Peninsular Malaysia, including a new species and two new varieties. PhytoKeys 189: 99-127, DOI: http://dx.doi.org/10.3897/phytokeys.189.49367, URL: http://dx.doi.org/10.3897/phytokeys.189.49367
BD97B648DB8E5EE092CD99438639F6EE.taxon	materials_examined	Additional specimens examined. Peninsular Malaysia. Negeri Sembilan: Jelebu, Kenaboi FR, G. Besar Hantu, from Taman Alam Liar towards Trail 1, near pathways, 3 ° 12 ' N, 100 ° 58 ' E, 530 m alt., 4 Mar 2014 (fl., fr.), Julius et al. FRI 57797 (KEP!); road sides towards Lata Kijang, 3 ° 11.42 ' N, 100 ° 59.21 ' E, 459 m alt., 23 May 2019 (fr.), Julius et al. FRI 73545 (KEP!).	en	Julius, Avelinah (2022): Capparis (Capparaceae) in Peninsular Malaysia, including a new species and two new varieties. PhytoKeys 189: 99-127, DOI: http://dx.doi.org/10.3897/phytokeys.189.49367, URL: http://dx.doi.org/10.3897/phytokeys.189.49367
DF1941E5E9C15D46BBDD74518FCB7AD2.taxon	description	Figures 4, 5	en	Julius, Avelinah (2022): Capparis (Capparaceae) in Peninsular Malaysia, including a new species and two new varieties. PhytoKeys 189: 99-127, DOI: http://dx.doi.org/10.3897/phytokeys.189.49367, URL: http://dx.doi.org/10.3897/phytokeys.189.49367
DF1941E5E9C15D46BBDD74518FCB7AD2.taxon	diagnosis	Diagnosis. Recognised in the genus Capparis by the combination of the following characters: the racemes are strictly terminal and compact with flowers subtended by a conspicuous and persistent unguiculate, leaf-like bracts and the smaller [(13 -) 16 - 25 x 2 - 8 mm], greenish, thin leaf-like bracts covered with densely, tomentose, straw hairs on both surfaces soon glabrescent with age and the sepals (inner pair) glabrous along the margin and the filaments and gynophore both entirely glabrous and the petals loosely ciliate along margin.	en	Julius, Avelinah (2022): Capparis (Capparaceae) in Peninsular Malaysia, including a new species and two new varieties. PhytoKeys 189: 99-127, DOI: http://dx.doi.org/10.3897/phytokeys.189.49367, URL: http://dx.doi.org/10.3897/phytokeys.189.49367
DF1941E5E9C15D46BBDD74518FCB7AD2.taxon	materials_examined	Type. Peninsular Malaysia. Pahang: Fraser's Hill, 25 Jan 1982 (fl.), Kiew RK 3747 (holotype KLU!).	en	Julius, Avelinah (2022): Capparis (Capparaceae) in Peninsular Malaysia, including a new species and two new varieties. PhytoKeys 189: 99-127, DOI: http://dx.doi.org/10.3897/phytokeys.189.49367, URL: http://dx.doi.org/10.3897/phytokeys.189.49367
DF1941E5E9C15D46BBDD74518FCB7AD2.taxon	description	Description. Climber ca. 1.5 m long or scrambling shrub (height unknown). Twigs straight covered with indumentum of dense, appressed, pale brown hairs soon glabrescent. Stipular thorns retrorse, tips 1 - 2 mm long. Leaves spirally arranged, but in one plane; petioles 10 - 15 x ca. 10 mm long, slender, young densely hairy with brown hairs soon glabrescent with age; lamina narrowly elliptic or oblong-elliptic, 8.5 - 11.5 (- 13.5) x 2.6 - 3.6 cm, chartaceous to subcoriaceous, drying brownish-green, base cuneate, margin flat and entire, apex acuminate and slightly caudate, with acumen 5 - 7 (- 10) mm long, glabrous above, sparsely hairy, denser on mid-rib beneath; mid-rib flat above, raised beneath; lateral veins 4 - 6 pairs, sunken above, distinct beneath, reddish when dry, looping towards the margin; intercostal veins reddish, finely reticulate, obscure above, distinct beneath. Inflorescences terminal, elongated racemes, rachis stout, straight, 3 - 8 (- 10) cm long, densely hairy with pale brown hairs; bracts leaf-like, unguiculate, lanceolate, (13 -) 16 - 25 x 2 - 8 mm, densely hairy with pale brown hairs on both surfaces. Flowers [in bud] many, compact, buds globose, 4 - 7 x 3 - 7 mm; pedicels 4 - 5 mm long, brownish; sepals 4, thin, green, orbicular to ovate, 4 - 6 x 4 - 6 mm, outer pair imbricate, densely hairy with appressed pale brown hairs outside, glabrous inside, inner pair with broadly transparent margin, glabrous on both surfaces; petals 4, pinkish, obovate, 6.5 - 7 x 4.5 - 5.5 mm, glabrous on both surfaces, except long, silky hairs at base of dorsal pair and dense, short silky hairs at base of ventral pair inside, loosely, ciliate along margin; stamens numerous, unequal length, filament pinkish, ca. 5 - 6 mm long, anther dorsifixed, ca. 1.5 x 0.5 mm; ovary ovoid, ca. 1.5 x 2.5 mm, on gynophore ca. 4 mm long, glabrous, stigma obscure. Fruits young green, mature fruit n. v., globose on stout gynophore, ca. 9.5 cm in diameter. Seeds many.	en	Julius, Avelinah (2022): Capparis (Capparaceae) in Peninsular Malaysia, including a new species and two new varieties. PhytoKeys 189: 99-127, DOI: http://dx.doi.org/10.3897/phytokeys.189.49367, URL: http://dx.doi.org/10.3897/phytokeys.189.49367
DF1941E5E9C15D46BBDD74518FCB7AD2.taxon	distribution	Distribution. Endemic in Peninsular Malaysia, Pahang, known only from Fraser's Hill and Cameron Highlands (Fig. 4).	en	Julius, Avelinah (2022): Capparis (Capparaceae) in Peninsular Malaysia, including a new species and two new varieties. PhytoKeys 189: 99-127, DOI: http://dx.doi.org/10.3897/phytokeys.189.49367, URL: http://dx.doi.org/10.3897/phytokeys.189.49367
DF1941E5E9C15D46BBDD74518FCB7AD2.taxon	etymology	Etymology. This new variety is named after Dr Ruth Kiew, collector of type specimen and the Flora of Peninsular Malaysia project co-ordinator and consultant.	en	Julius, Avelinah (2022): Capparis (Capparaceae) in Peninsular Malaysia, including a new species and two new varieties. PhytoKeys 189: 99-127, DOI: http://dx.doi.org/10.3897/phytokeys.189.49367, URL: http://dx.doi.org/10.3897/phytokeys.189.49367
DF1941E5E9C15D46BBDD74518FCB7AD2.taxon	materials_examined	Additional specimens examined. Peninsular Malaysia. Pahang: one-way road to Fraser's Hill, 25 Jan 1982 (fl.), Kiew RK 1116 (KEP!); Fraser's Hill, 1 Aug 1993 (fl.), A. Zainuddin AZ 4636 (KEP!); Fraser's Hill, Jalan kecil to Gap, 19 Jun 1930 (fl.), Kalong FMS 22381 (KEP!); Fraser's Hill, Richmond, 21 Apr 1955 (fr.), Purseglove P 4248 (L- image!, barcode L 1856045) Cameron Highlands, 13 Apr 1930 (fr.), Henderson SFN 23551 (KEP!, SING!).	en	Julius, Avelinah (2022): Capparis (Capparaceae) in Peninsular Malaysia, including a new species and two new varieties. PhytoKeys 189: 99-127, DOI: http://dx.doi.org/10.3897/phytokeys.189.49367, URL: http://dx.doi.org/10.3897/phytokeys.189.49367
3C109AC557765752B847663FF9039F68.taxon	description	Figures 6, 7, 8	en	Julius, Avelinah (2022): Capparis (Capparaceae) in Peninsular Malaysia, including a new species and two new varieties. PhytoKeys 189: 99-127, DOI: http://dx.doi.org/10.3897/phytokeys.189.49367, URL: http://dx.doi.org/10.3897/phytokeys.189.49367
3C109AC557765752B847663FF9039F68.taxon	diagnosis	Diagnosis. Recognised in the genus Capparis by the combination of the following characters: the leaves broadly ovate-elliptic, relatively large (13 - 16 x 5.5 - 8.5 cm) and leaves drying reddish-brown, rarely pale green with pale yellow and rarely dark red venation, the tertiary venation obscure, the inflorescence strictly terminal with flowers arranged racemosely and becoming crowded at the distal part of the inflorescence, the stamens 30 - 40 and the fruit globose with ca. 11 cm in diameter.	en	Julius, Avelinah (2022): Capparis (Capparaceae) in Peninsular Malaysia, including a new species and two new varieties. PhytoKeys 189: 99-127, DOI: http://dx.doi.org/10.3897/phytokeys.189.49367, URL: http://dx.doi.org/10.3897/phytokeys.189.49367
3C109AC557765752B847663FF9039F68.taxon	materials_examined	Type. Peninsular Malaysia. Selangor: Kuala Kubu Bharu to Fraser's Hill Road, along the road side, 17 Apr 1971 (fl.), Mahmud Sidek s. n. (holotype KLU!).	en	Julius, Avelinah (2022): Capparis (Capparaceae) in Peninsular Malaysia, including a new species and two new varieties. PhytoKeys 189: 99-127, DOI: http://dx.doi.org/10.3897/phytokeys.189.49367, URL: http://dx.doi.org/10.3897/phytokeys.189.49367
3C109AC557765752B847663FF9039F68.taxon	description	Description. Woody climber 9 - 12 m long. Twigs straight, covered with velvety, ferruginous hairs and glabrescent with age. Stipular thorns recurved downwards, tips 1.5 - 3 mm long. Leaves spirally arranged; petioles 15 - 20 x 2 - 3 mm, stout, incrassate or thickened, velvety hairy becoming glabrous; lamina broadly ovate sometimes ovate-elliptic, 13 - 16 x 5.5 - 8.5 cm, coriaceous to subcoriaceous, surface rugose and usually reddish-brown when dry, rarely pale green, base cuneate or occasionally rounded, margin flat and entire, apex acute or shortly cuspidate, with acumen ca. 4 mm long, glabrous on both surfaces; mid-rib flat above, raised beneath; lateral veins 5 - 7 (- 8) pairs, flat above, distinct beneath, pale yellow when dry, looping towards the margin; intercostal veins obscure. Inflorescences terminal, flowers arranged racemosely and becoming crowded at the distal part of the inflorescence rachis slender and straight; bracts caducous. Flowers [in bud] 4 - 7, buds globose, 1 - 1.8 x 1.3 - 1.8 cm; pedicels 3 - 3.5 cm long, swollen towards apex, velvety with ferruginous hairs; sepals 4, thick, whitish-green, orbicular, 1.2 - 1.5 x 1.2 cm, outer pair of sepals imbricate, covered with dense, velvety ferruginous hairs outside, glabrous inside, inner pair of sepals glabrous on both surfaces, except silky hairs at base outside; petals 4, cream or white with pink- or dark purple at base; stamens 30 - 40, unequal in length, anthers yellow; ovary green. Fruits young shiny and green, mature fruit n. v., globose to sub-globose on a stout gynophore, sometimes beaked when young, ca. 9 x 11 cm. Seeds (3 -) 12 - 15, sarcotesta whitish, testa thin and whitish-cream.	en	Julius, Avelinah (2022): Capparis (Capparaceae) in Peninsular Malaysia, including a new species and two new varieties. PhytoKeys 189: 99-127, DOI: http://dx.doi.org/10.3897/phytokeys.189.49367, URL: http://dx.doi.org/10.3897/phytokeys.189.49367
3C109AC557765752B847663FF9039F68.taxon	distribution	Distribution. Endemic in Peninsular Malaysia, Negeri Sembilan, Selangor and Pahang (Fig. 6).	en	Julius, Avelinah (2022): Capparis (Capparaceae) in Peninsular Malaysia, including a new species and two new varieties. PhytoKeys 189: 99-127, DOI: http://dx.doi.org/10.3897/phytokeys.189.49367, URL: http://dx.doi.org/10.3897/phytokeys.189.49367
3C109AC557765752B847663FF9039F68.taxon	etymology	Etymology. The new variety is named after Dr Richard Chung Cheng Kong, Curator of the Kepong Herbarium (KEP) and Project Leader of Flora of Malaysia under 10 thand 11 th Malaysian Plans.	en	Julius, Avelinah (2022): Capparis (Capparaceae) in Peninsular Malaysia, including a new species and two new varieties. PhytoKeys 189: 99-127, DOI: http://dx.doi.org/10.3897/phytokeys.189.49367, URL: http://dx.doi.org/10.3897/phytokeys.189.49367
3C109AC557765752B847663FF9039F68.taxon	materials_examined	Additional specimens examined. Peninsular Malaysia. Selangor: Ulu Gombak FR, 259 m (850 ft) alt., 19 Aug 1964 (fr.), Mohd. Kasim & Mahmud Sidek 624 (KLU!); Gombak Forest, 16 th mile, followed path to stream and beyond, 427 m (1400 ft) alt., 12 Jun 1963 (fl.), M. E. D. Poore 1189 (KLU!); Kepong Plantation Field 9 J, hillslope, 10 Jan 1934 (fl.), Abdul Hamid 33570 (KEP!); Ulu Langat, 30 Mar 1960 (fl., fr.), Gadoh KL 2072 (KEP! 2 sheets); Genting Sempah, 11 Dec 1970 (fl., fr.), Kochummen FRI 16263 (KEP!). Pahang: Fraser's Hill, near gate at gap, roadside, 1067 m (3500 ft) alt., 28 Feb 1979 (fl., fr.), Kochummen FRI 18471 (KEP!); road to Fraser's Hill, climber on Mahang tree beside the road, 914 m (3000 ft) alt., 26 Aug 1981 (fr.), K. M. Wong FRI 32242 (KEP!). Negeri Sembilan: Jelebu, Kenaboi FR, G. Besar Hantu, near Kg. London area, 3 ° 11.00 ' N, 102 ° 00.76 ' E, 500 m alt., 6 May 2010 (fr.), Julius et al. FRI 64006 (KEP!).	en	Julius, Avelinah (2022): Capparis (Capparaceae) in Peninsular Malaysia, including a new species and two new varieties. PhytoKeys 189: 99-127, DOI: http://dx.doi.org/10.3897/phytokeys.189.49367, URL: http://dx.doi.org/10.3897/phytokeys.189.49367
