taxonID	type	description	language	source
650446796C13FFD08A7BFAECFC82FBA7.taxon	description	(Figs 1, 2)	en	Ermilov, Sergey G., Salavatulin, Vladimir M. (2022): New Species Of Flagellozetes (Cosmogalumna) (Acari, Oribatida, Galumnidae) From Vietnam. Acta Zoologica Academiae Scientiarum Hungaricae 68 (4): 293-304, DOI: 10.17109/AZH.68.4.293.2022, URL: http://dx.doi.org/10.17109/azh.68.4.293.2022
650446796C13FFD08A7BFAECFC82FBA7.taxon	diagnosis	Diagnosis. Body length: 270 – 300. Pteromorph slightly striate; anogenital region and centrodorsal part of notogaster with numerous neural ridges. Lamellar line bifurcate mediobasally; anterior part of S - line dentate. Rostral and lamellar setae setiform, slightly roughened; ro / le = 1.2. Dorsosejugal and postanal porose areas present. Four pairs of rounded porose areas. Median pore absent. Lyrifissure im located between lm and A 1. Epimeral setal formula: 1 - 0 - 1 - 1. Circumpedal carina long. Adanal lyrifissure located close and parallel to anterior part of anal plate. Leg tarsus I with 19 setae (v’ absent).	en	Ermilov, Sergey G., Salavatulin, Vladimir M. (2022): New Species Of Flagellozetes (Cosmogalumna) (Acari, Oribatida, Galumnidae) From Vietnam. Acta Zoologica Academiae Scientiarum Hungaricae 68 (4): 293-304, DOI: 10.17109/AZH.68.4.293.2022, URL: http://dx.doi.org/10.17109/azh.68.4.293.2022
650446796C13FFD08A7BFAECFC82FBA7.taxon	description	Description of adult. Measurements. Small species. Body length: 300 (holotype: female), 270 – 300 (six paratypes: three males and three females); notogaster width: 232 (holotype), 217 – 232 (paratypes). No difference between males and females in body size. Integument (Figs 1 A – C). Body color brown. Surface densely microfoveolate (visible only under high magnification in dissected specimens, 10 × 100). Pteromorph, basal part of prodorsum and narrow region anterior to genital aperture slightly striate. Lateral side of body partially microgranulate. Anogenital region and centrodorsal part of notogaster with numerous neural ridges (without small dense cells); ridges in anogenital region numerous and located on the entire surface (except genital and anal plates); ridges on notogaster medium numerous and occupy the area between setal alveoli lm-lm and porose areas A 1 - A 1. Antiaxial side of leg femora I, II, IV and trochanters III, IV partially tuberculate. Prodorsum (Figs 1 A, C). Rostrum slightly protruding, rounded. Lamellar and sublamellar lines thin, parallel, curving backwards; L - line bifurcate mediobasally; anterior part of S - line with one row of short, distinct teeth. One pair of thin, longitudinal furrows (30) located between mediobasal parts of L - lines. Rostral (20 – 22) and lamellar (17 – 19) setae setiform, slightly roughened; le inserted between branches of L - line. Interlamellar seta (2) setiform, thin, smooth. Bothridial seta (56 – 60) clavate, with long stalk and shorter head rounded distally, sparsely slightly roughened. Exobothridial seta absent. Dorsosejugal porose area (12 – 15 × 4 – 6) oval, transversely oriented, located posterolateral to in. Dorsophragma distinctly elongated longitudinally. Notogaster (Figs 1 A, C, D). Dorsosejugal suture complete, convex medially. With 10 pairs of setal alveoli (microsetae not observed) and four pairs of rounded porose areas (Aa: 13 – 19; A 1: 9 – 15; A 2: 7 – 11; A 3: 11 – 15); Aa located close to pteromorphal hinge, anterior to setal alveolus la. Median pore absent. Opisthonotal gland opening and all lyrifissures distinct: gla located anterolateral to A 2; im between lm and A 1; ip between p 1 and p 2; ips between p 2 and p 3; ih lateral to p 3. Gnathosoma (Figs 2 A – C). Subcapitulum size: 75 – 79 × 60 – 67; subcapitular setae (a: 15 – 19; m, h: 5 – 7) setiform, slightly roughened; a thickest; adoral seta (9 – 11) setiform, slightly barbed. Length of palp: 64 – 67; postpalpal seta (4) spiniform, smooth. Length of chelicera: 86 – 94; cheliceral setae (cha: 26 – 30; chb: 19 – 22) setiform, barbed. Epimeral and lateral podosomal regions (Figs 1 B, C). Epimeral setal formula: 1 - 0 - 1 - 1; all epimeral setae (3 b: 11 – 15; 1 a: 7; 4 a: 4) setiform, thin, smooth. Circumpedal carina long, reaching pedotectum I. Anogenital region (Figs 1 B – D). Genital (g 1: 9 – 11; g 2: 7; g 3 – g 6: 4), aggenital (4), anal (4), and adanal (4) setae setiform, thin, smooth. Anterior edge of genital plate with three setae. Aggenital seta located between genital and anal apertures, closer to genital aperture. Adanal lyrifissure located close and parallel to anterior part of anal plate. Adanal setae ad 1 and ad 2 posterior, ad 3 lateral to anal plate; distance ad 1 – ad 2 shorter than ad 2 – ad 3. Unpaired postanal porose area (7 – 11 × 4 – 6) oval. Legs (Figs 2 D – G). Median claw distinctly thicker than lateral claws, all slightly roughened on dorsal side. Porose area on all femora and on trochanters III, IV present, but poorly visible. Formulas of leg setation and solenidia: I (1 - 4 - 3 - 4 - 19) [1 - 2 - 2], II (1 - 4 - 3 - 4 - 15) [1 - 1 - 2], III (1 - 2 - 1 - 3 - 15) [1 - 1 - 0], IV (1 - 2 - 2 - 3 - 12) [0 - 1 - 0]; famulus lateral to solenidion ω 1; seta v’ on tarsus I absent; s eupathidial; solenidion of tibia IV inserted in anterior part of the segment; homology of setae and solenidia indicated in Table 1.	en	Ermilov, Sergey G., Salavatulin, Vladimir M. (2022): New Species Of Flagellozetes (Cosmogalumna) (Acari, Oribatida, Galumnidae) From Vietnam. Acta Zoologica Academiae Scientiarum Hungaricae 68 (4): 293-304, DOI: 10.17109/AZH.68.4.293.2022, URL: http://dx.doi.org/10.17109/azh.68.4.293.2022
650446796C13FFD08A7BFAECFC82FBA7.taxon	materials_examined	Material examined. Holotype (female) and two paratypes (one male and one female): Vietnam, Dong Nai Province, Dong Nai Biosphere Reserve, Cat Tien National Park, 11 ° 25 ’ 40 ” N, 107 ° 25 ’ 34 ” E, about 130 m a. s. l., bark from Haldina cardifolia (height: 0.5 m) (sample: 1 - 1), 20.06.2021 – 04.07.2021 (V. M. Salavatulin and A. A. Kudrin). Four paratypes (two males and two females): the same as for the holotype but other tree of H. cardifolia (sample: 8 - 1) with coordinates 11 ° 25 ’ 45 ” N, 107 ° 25 ’ 38 ” E. Type deposition. The holotype and six paratypes (preserved in 70 % solution of ethanol with a drop of glycerol) are deposited in the collection of the Tyumen State University Museum of Zoology, Tyumen, Russia.	en	Ermilov, Sergey G., Salavatulin, Vladimir M. (2022): New Species Of Flagellozetes (Cosmogalumna) (Acari, Oribatida, Galumnidae) From Vietnam. Acta Zoologica Academiae Scientiarum Hungaricae 68 (4): 293-304, DOI: 10.17109/AZH.68.4.293.2022, URL: http://dx.doi.org/10.17109/azh.68.4.293.2022
650446796C13FFD08A7BFAECFC82FBA7.taxon	etymology	Etymology. The species name carinodentatus refers to dentate sublamellar line (carina).	en	Ermilov, Sergey G., Salavatulin, Vladimir M. (2022): New Species Of Flagellozetes (Cosmogalumna) (Acari, Oribatida, Galumnidae) From Vietnam. Acta Zoologica Academiae Scientiarum Hungaricae 68 (4): 293-304, DOI: 10.17109/AZH.68.4.293.2022, URL: http://dx.doi.org/10.17109/azh.68.4.293.2022
650446796C13FFD08A7BFAECFC82FBA7.taxon	discussion	Remarks. Flagellozetes (Cosmogalumna) carinodentatus sp. n. differs from all representatives of the subgenus by the presence of a row of teeth on sublamellar line. In having neural ridges on the notogaster and in anogenital region, stria on pteromorph, four pairs of notogastral porose areas, long circumpedal carina, bifurcate lamellar line, 19 setae on leg tarsus I, and in the absence of a median pore, the new species is morphologically most similar to F. (C.) sandori Ermilov et Kalúz, 2019 from Malaysia. However, the new species differs (in addition to the presence of teeth on S) from the latter by the absence (versus presence) of sparse and minute foveolae on the body and stria on the subcapitular mentum, distinctly longer lamellar setae (ro / le = 1.2 versus 1.6), the presence (versus absence) of two prodorsal furrows, and by more numerous neural ridges in anogenital region which are located on entire surface (versus mostly between genital and anal apertures).	en	Ermilov, Sergey G., Salavatulin, Vladimir M. (2022): New Species Of Flagellozetes (Cosmogalumna) (Acari, Oribatida, Galumnidae) From Vietnam. Acta Zoologica Academiae Scientiarum Hungaricae 68 (4): 293-304, DOI: 10.17109/AZH.68.4.293.2022, URL: http://dx.doi.org/10.17109/azh.68.4.293.2022
650446796C17FFDC8A47FB6BFD90FA8D.taxon	description	(Fig. 3)	en	Ermilov, Sergey G., Salavatulin, Vladimir M. (2022): New Species Of Flagellozetes (Cosmogalumna) (Acari, Oribatida, Galumnidae) From Vietnam. Acta Zoologica Academiae Scientiarum Hungaricae 68 (4): 293-304, DOI: 10.17109/AZH.68.4.293.2022, URL: http://dx.doi.org/10.17109/azh.68.4.293.2022
650446796C17FFDC8A47FB6BFD90FA8D.taxon	diagnosis	Diagnosis. Body length: 330 – 352. Anogenital region and centrodorsal part of notogaster with few neural ridges. Lamellar line not bifurcate mediobasally. Rostral and lamellar setae similar in length, setiform, slightly roughened. Dorsosejugal and postanal porose areas present. Four pairs of rounded porose areas. Median pore present. Lyrifissure im located close and anterior to A 1. Epimeral setal formula: 1 - 0 - 1 - 2. Circumpedal carina short. Adanal lyrifissure located close and parallel to median part of anal plate. Leg tarsus I with 20 setae (v’ present).	en	Ermilov, Sergey G., Salavatulin, Vladimir M. (2022): New Species Of Flagellozetes (Cosmogalumna) (Acari, Oribatida, Galumnidae) From Vietnam. Acta Zoologica Academiae Scientiarum Hungaricae 68 (4): 293-304, DOI: 10.17109/AZH.68.4.293.2022, URL: http://dx.doi.org/10.17109/azh.68.4.293.2022
650446796C17FFDC8A47FB6BFD90FA8D.taxon	description	Description of adult. Measurements. Small species. Body length: 352 (holotype: female), 330 (one paratype: male); notogaster width: 255 (holotype), 240 (paratype). Integument (Figs 3 A, B). Body color brown. Surface densely microfoveolate (visible only under high magnification in dissected specimens, 10 × 100). Laterobasal part of prodorsum slightly striate. Lateral side of body partially microgranulate. Anogenital region and centrodorsal part of notogaster with few neural ridges (without small dense cells); ridges in anogenital region located only between aggenital setae; ridges on notogaster occupy the narrow transverse area between and slightly anterior to porose areas A 1 - A 1. Antiaxial side of leg femora I, II, IV and trochanters III, IV partially tuberculate. 1. Foveolate ornamentation (foveolae sparse, small but well visible) on body: A = absent; P = present. 2. Form of bothridial seta: C = clavate (broadly rounded distally); F = fusiform (narrowed distally). 3. One pair of longitudinal furrows on prodorsum: A = absent; P = present. 4. Lamellar line mediobasally: Bi = bifurcate; Si = simple (not bifurcate). 5. Teeth on sublamellar line: A = absent; P = present. 6. Relative length of rostral and lamellar setae: le / ro. 7. Localization of setal alveolus c on pteromorph: An = anterior to alary furrow; Po = posterior to alary furrow. 8. Number of notogastral porose areas. 9. Notogastral area with neural ridges: La = large, mostly entire surface; Me = medium-sized, broad quadrangular area between setal alveoli lm-lm and porose areas A 1 - A 1 or slightly larger; Sm = small, narrowly transverse area between and slightly anterior to porose areas A 1 - A 1. 10. Ornamentation of pteromorph: A = absent; Gr = heavily granulate; St = slightly or heavily striate (or with ridges). 11. Localization of notogastral lyrifissure im: Re = im between lm and A 1 (or between lm and lp) and equally removed from them; Cl = im close and anterior to A 1. 12. Median pore: A = absent; P = present. 13. Epimeral formula. 14. Length of circumpedal carina: L = long (reaching pedotectum I); M = medium length, directed to setal alveolus 3 b but slightly not reaching it, or only reaching level of localization of acetabulum IV. 15. Ornamentation of genital plate: A = absent; St = striate. 16. Number of genital setae on anterior edge of genital plate. 17. Aggenital seta: A = absent; P = present; Re = ag equally removed from genital and anal apertures; Cl = ag closer to genital aperture; Not = not applicable (if ag absent). 18. Reticulate ornamentation in anogenital region: A = absent; P = present. 19. Anogenital area with neural ridges: La = large, mostly entire surface including adanal region; Me = medium-sized, only between genital and anal apertures; Sm = small, only between aggenital setae; Not = not applicable (if neural ridges absent). 20. Ornamentation of anal plate: A = absent; St = striate. 21. Localization of adanal lyrifissure: Re = removed from anal aperture; Cl = close to anal aperture, parallel; An = close to level of the anterior half of anal plate; Mi = close to level of the middle of anal plate; Not = not applicable (if iad removed from anal aperture). 22. Number of setae on leg tarsus I. 23. Body length. Prodorsum (Figs 3 A, C). Rostrum broadly rounded. Lamellar and sublamellar lines parallel, curving backwards; L - line slightly thickened, not bifurcate basally; S - line thin. Rostral and lamellar setae similar in length (28 – 30), setiform, slightly roughened; le inserted lateral to L - line. Interlamellar seta (2) setiform, thin, smooth. Bothridial seta (45 – 49) clavate, with long stalk and shorter head rounded distally, sparsely slightly roughened. Exobothridial seta absent. Dorsosejugal porose area (15 – 19 × 7) oval, transversely oriented, located posterolateral to in. Dorsophragma distinctly elongated longitudinally. Notogaster (Figs 3 A, C, D). Dorsosejugal suture complete, nearly straight medially. With 10 pairs of setal alveoli (microsetae not observed) and four pairs of rounded porose areas (Aa: 17 – 19; A 1, A 3: 11; A 2: 7); Aa located close to pteromorphal hinge, anterior to setal alveolus la. Median pore present, located posterior to virtual line connected areas A 2. Opisthonotal gland opening and all lyrifissures distinct: gla located lateral to A 1; im close and anterior to A 1; ip between p 1 and p 2; ips between p 2 and p 3; ih anterior to p 3. Gnathosoma. Mostly similar to F. (C.) carinodentatus sp. n. except some sizes. Subcapitulum size: 86 × 79; a: 19; m, h: 11; adoral seta: 15; length of palp: 79; length of chelicera: 105; cha: 34; chb: 22. Epimeral and lateral podosomal regions (Figs 3 B, C). Epimeral setal formula: 1 - 0 - 1 - 2; all epimeral setae (3 b: 19 – 22; 1 a: 11; 4 a, 4 b: 7) setiform, thin, smooth. Circumpedal carina comparatively short, reaching of level of localization of acetabulum IV. Anogenital region (Figs 3 B – D). Genital (g 1: 15; g 2: 11; g 3 – g 6: 7), aggenital (7), anal (7), and adanal (7) setae setiform, thin, smooth. Anterior edge of genital plate with three setae. Aggenital seta equally removed from genital and anal apertures. Adanal lyrifissure located close and parallel to median part of anal plate. Adanal setae ad 1 and ad 2 posterior, ad 3 lateral to anal plate; distance ad 1 – ad 2 shorter than ad 2 – ad 3. Unpaired postanal porose area (18 – 22 × 7) oval. Legs. Mostly similar to F. (C.) carinodentatus sp. n., but tarsus I with 20 setae (v’ present), and famulus inserted between solenidia ω 1 and ω 2.	en	Ermilov, Sergey G., Salavatulin, Vladimir M. (2022): New Species Of Flagellozetes (Cosmogalumna) (Acari, Oribatida, Galumnidae) From Vietnam. Acta Zoologica Academiae Scientiarum Hungaricae 68 (4): 293-304, DOI: 10.17109/AZH.68.4.293.2022, URL: http://dx.doi.org/10.17109/azh.68.4.293.2022
650446796C17FFDC8A47FB6BFD90FA8D.taxon	materials_examined	Material examined. Holotype (female): Vietnam, Dong Nai Province, Dong Nai Biosphere Reserve, Cat Tien National Park, 11 ° 25 ’ 40 ” N, 107 ° 25 ’ 34 ” E, about 130 m a. s. l., bark from Ochrocarpos siamensis (height: 0.5 m) (sample: 3 - 1), 20.06.2021 – 04.07.2021 (V. M. Salavatulin and A. A. Kudrin). One paratype (male): the same as for the holotype but tree of Haldina cardifolia (sample: 8 - 3) with coordinates 11 ° 25 ’ 45 ” N, 107 ° 25 ’ 38 ” E and other height (20 m). Type deposition. The holotype and one paratype (preserved in 70 % solution of ethanol with a drop of glycerol) are deposited in the collection of the Tyumen State University Museum of Zoology, Tyumen, Russia.	en	Ermilov, Sergey G., Salavatulin, Vladimir M. (2022): New Species Of Flagellozetes (Cosmogalumna) (Acari, Oribatida, Galumnidae) From Vietnam. Acta Zoologica Academiae Scientiarum Hungaricae 68 (4): 293-304, DOI: 10.17109/AZH.68.4.293.2022, URL: http://dx.doi.org/10.17109/azh.68.4.293.2022
650446796C17FFDC8A47FB6BFD90FA8D.taxon	etymology	Etymology. The species name pseudoareticulatus refers to the similarity of the new species and Flagellozetes (Cosmogalumna) areticulatus (Ermilov, Sandmann, Klarner, Widyastuti & Scheu, 2015).	en	Ermilov, Sergey G., Salavatulin, Vladimir M. (2022): New Species Of Flagellozetes (Cosmogalumna) (Acari, Oribatida, Galumnidae) From Vietnam. Acta Zoologica Academiae Scientiarum Hungaricae 68 (4): 293-304, DOI: 10.17109/AZH.68.4.293.2022, URL: http://dx.doi.org/10.17109/azh.68.4.293.2022
650446796C17FFDC8A47FB6BFD90FA8D.taxon	discussion	Remarks. In having neural ridges on the notogaster and in anogenital region, a median pore, four pairs of notogastral porose areas, short circumpedal carina, and the absence of stria on pteromorph, Flagellozetes (Cosmogalumna) pseudoareticulatus sp. n. is morphologically most similar to F. (C.) areticulatus (Ermilov, Sandmann, Klarner, Widyastuti et Scheu, 2015) from the Oriental region. However, the new species differs from the latter by the comparatively long lamellar seta, ro = le (versus short, ro / le = 2.7), the localization of notogastral lyrifissure im (close and anterior to A 1 versus between lm and lp, equally removed from them) and adanal seta ad 3 (lateral to iad versus anterolateral to iad), the position of aggenital seta (equally distanced from genital and anal apertures versus located closer to genital aperture), and by comparatively small area with neural ridges in anogenital region and on the notogaster (versus areas with ridges clearly larger).	en	Ermilov, Sergey G., Salavatulin, Vladimir M. (2022): New Species Of Flagellozetes (Cosmogalumna) (Acari, Oribatida, Galumnidae) From Vietnam. Acta Zoologica Academiae Scientiarum Hungaricae 68 (4): 293-304, DOI: 10.17109/AZH.68.4.293.2022, URL: http://dx.doi.org/10.17109/azh.68.4.293.2022
