taxonID	type	description	language	source
81311371E6DF568DA8BA8EE75FEC0263.taxon	description	Figs 3, 4, 5	en	Cumming, Royce T., Foley, Evelyn Marie, Hennemann, Frank H., Le Tirant, Stephane, Daawia,, Warikar, Evie Lilly, Yando, Heron, Suhartawan, Bambang, Henze, Katharina, Büscher, Thies H., Bank, Sarah (2025): A deeper look into the diversity of Phyllium leaf insects from Indonesia: seven new species and two unique egg morphologies (Phasmatodea, Phylliidae). ZooKeys 1256: 317-370, DOI: 10.3897/zookeys.1256.162609
81311371E6DF568DA8BA8EE75FEC0263.taxon	description	Description. Female. Coloration. Coloration description is based upon images of the living type specimens (Fig. 3). In the holotype and paratype, the general coloration is pale green throughout. In the holotype, these are numerous colored patches across the dorsal and ventral surfaces (Fig. 3 A) while the paratype is simply pale green throughout and lacks these colored patches. In the holotype, there are brown spots of varying sizes and shapes on the legs, abdomen, and tegmina. Ventral coxae coloration is pale orange (Fig. 3 B). Morphology. Head capsule slightly longer than wide, with a dorsal surface that is smooth, lacking granulation (Fig. 4 B). The posteromedial tubercle is present, singularly lobed, but not very prominent (Fig. 4 B). Frontal convexity broad and ending in a blunted point; there are only a few short setae across the surface. Compound eyes small and only slightly protruding from the head capsule, not bulbous, taking up ~ 1 / 5 of the head capsule lateral margins (Fig. 4 B). Ocelli absent. Antennal field approx. as wide as first antennomere. Antennae consist of nine segments (Fig. 4 C), with the terminal segment not particularly wide or long (slightly shorter than the combined length of the previous two segments combined lengths). Antennomeres IX and the distal 1 / 2 of segment VIII have a rough, fuzzy texture with short, dark setae. The remaining segments are smooth, and sparsely marked with short, transparent setae, none prominent (Fig. 4 C). Thorax. Pronotum with slightly concave anterior margin and lateral margins that anteriorly start wide, angle inward strongly towards the posterior margin which is slightly wider than 1 / 2 of the anterior margin width (Fig. 4 B). The pronotum anterior margin has a prominent rim, the lateral margins are less prominent, and the posterior margin is weakly developed. The pronotum surface is relatively smooth, has a small but prominent sagittal slit in the center and another near the anterior. The remainder of the surface is smooth and marked with a few other furrows (Fig. 4 B). Prosternum and the anterior 1 / 2 of the mesosternum are covered throughout by moderately spaced granulation. The posterior 1 / 2 of the mesosternum and the metasternum are relatively smooth (Fig. 4 G). Mesoprescutum slightly longer than wide, lateral rims lumpy with 8 – 10 variable sized and spaced nodes (3 – 5 of these are somewhat prominent; Fig. 4 B). Mesoprescutum anterior rim distinctly raised and slightly lumpy but not forming a distinct sagittal spine (Fig. 4 B). Mesoprescutum surface wrinkled and lumpy, mostly just with nodes along the sagittal plane, but there are several prominent nodes spread throughout the surface (Fig. 4 B). Mesopleurae begin to diverge ~ ½ of the way along the mesoprescutum, angle prominently away with straight margins (Fig. 4 B). Mesopleurae lateral margins with five or six small spiniform tubercles with interspersed granulation throughout, giving the margin a roughly textured appearance since none are particularly prominent (Fig. 4 B). Face of the mesopleura slightly wrinkled along the lateral margins, and marked with two notable divots, one on the anterior margin and one near the middle (Fig. 4 B). Wings. Tegmina long, reaching the anterior margin of abdominal segment VIII. Tegmina venation; the subcosta (Sc) is the first vein in the forewing, running parallel with the margin for the first 1 / 2, and then bending and running towards the margin. The subcosta runs for ~ ¼ of the tegmina length. The radius (R) spans the anterior ½ of the forewing with two subparallel branched veins; the first radius (R 1) branches ~ ¼ of the way through the wing length and terminates ~ 2 / 5 of the way through the wing length; the radial sector (Rs) branches ~ 1 / 3 of the way through the wing length and terminates near the distal 3 / 5 of the wing length. There is a weak continuation of the radius following the prominent Rs branching which continues on as a short but distinct R – M crossvein that weakly connects the two veins. The media (M) is bifurcate with both the media anterior (MA) and media posterior (MP) terminating near to the posterior of the tegmina. The cubitus (Cu) is also bifurcate, branching near the posterior ¼ of the wing into the cubitus anterior (CuA) and cubitus posterior (CuP) which both terminate near the wing apex. The first anal vein (1 A) is simple and fuses with the cubitus ~ 1 / 5 of the way through the tegmina length. Alae vestigial nubs. Abdomen. Abdominal segments II through the anterior ½ of IV gradually and uniformly diverging. The posterior ½ of segment IV through the anterior ½ of segment VII are gradually converging. The posterior ½ of segment VII through the apex of the abdomen is converging to a blunted apex. Genitalia. Subgenital plate starts at the anterior margin of tergum VIII, is relatively narrow, and extends half-way onto tergum X. The overall shape is rather narrow, with the apex a blunted point (Fig. 4 F). Gonapophyses VIII are long and narrow, slightly exceeding the apex of abdominal tergum X; gonapophyses IX are mostly obstructed from view (Fig. 4 F). Cerci flat, and somewhat paddle-shaped, with narrow bases broadening out to a wide distal ½ (Fig. 4 F). Legs. Profemoral exterior lobe broad, rounded, arching from end to end, with a width similar to the width of the interior lobe (Fig. 4 D). Margin of profemoral exterior lobe smooth (Fig. 4 D). Profemoral interior lobe ~ 2.5 × as wide as the greatest width of the profemoral shaft, slightly obtusely angled, and marked with several teeth (starting on the proximal end: two large triangular teeth, then a large gap, another large triangular tooth, followed by a smaller finely pointed tooth on the distal end; Fig. 4 D). Mesofemoral exterior lobe approx. a narrow, rounded triangle with the greatest width similar to the mesofemoral shaft width, and the greatest width situated slightly distal to the midlength. Mesofemoral exterior lobe margin smooth, lacking teeth. Mesofemoral interior lobe is approx. the same width as the mesofemoral shaft, with a shape that is gently arching from end to end, with the distal 1 / 2 marked with 6 – 9 small, serrate teeth. Metafemoral interior lobe narrow on the proximal 1 / 2 with the wider distal 1 / 2 arching to the distal end. The distal 1 / 2 of the metafemoral interior lobe is marked with eight or nine small, serrate teeth. Metafemoral exterior lobe is thin and smooth, hugging the metafemoral shaft and lacks dentation. Protibia lacking an exterior lobe (Fig. 4 D). Protibia interior lobe spans the entire length of the protibia and is ~ 2 × the width of the protibia shaft itself. The lobe is roundly triangular with the widest portion slightly situated slightly distal to the midlength. Mesotibiae and metatibiae simple, lacking exterior and interior lobes. Measurements (mm). Holotype, female: body length (including cerci and head, excluding antennae): 97.9, length / width of head: 8.4 / 7.2, antennae: 4.8, pronotum: 5.9, mesonotum: 8.6, length of tegmina: 62.8, greatest width of abdomen: 35.8, profemora: 20.2, mesofemora: 17.9, metafemora: 22.2, protibia: 11.4, mesotibia: 11.9, metatibia: 17.1. Measurements (mm). Paratype, female: body length (including cerci and head, excluding antennae): 91.1, length / width of head: 8.1 / 7.0, antennae: 4.5, pronotum: 6.0, mesonotum: 8.1, length of tegmina: 58.3, greatest width of abdomen: 35.3, profemora: 16.9, mesofemora: 15.8, metafemora: 19.5, protibia: 9.3, mesotibia: 10.4, metatibia: 16.2. Eggs (Fig. 5). Margins rimmed with prominent feather-like pinnae. When viewed laterally, the general shape is ovoid (Fig. 5 B), and in cross-section the egg is a narrow triangle (Fig. 5 E, F). Lateral surfaces flattened and marked with seven longitudinal lines of bald impressions, the lines between appearing like a network of hairy ridges. Micropylar plate slender with relatively straight margins, and covering most of the length of dorsal egg surface. Micropylar cup small and situated on the posterior 1 / 3. Operculum somewhat teardrop shaped due to the triangular egg cross-section, flat, and with the lateral and ventral margins set with a row of the same long feather-like appendages seen along the longitudinal margins of the capsule (dorsal margin of the operculum lacks these pinnae). The operculum surface is marked throughout with irregularly sized pits with smooth, thick rims abutting the thick rim of the neighboring pits rims (Fig. 5 E). The posterior of the egg capsule has similar pits to those found on the operculum but they are slightly larger and fewer (Fig. 5 F). General color tan.	en	Cumming, Royce T., Foley, Evelyn Marie, Hennemann, Frank H., Le Tirant, Stephane, Daawia,, Warikar, Evie Lilly, Yando, Heron, Suhartawan, Bambang, Henze, Katharina, Büscher, Thies H., Bank, Sarah (2025): A deeper look into the diversity of Phyllium leaf insects from Indonesia: seven new species and two unique egg morphologies (Phasmatodea, Phylliidae). ZooKeys 1256: 317-370, DOI: 10.3897/zookeys.1256.162609
81311371E6DF568DA8BA8EE75FEC0263.taxon	description	Measurements including the extended pinnae [mm]. Length (including operculum expansion): 7.1; maximum width of capsule when viewed from lateral aspect 5.1; length of micropylar plate 3.4.	en	Cumming, Royce T., Foley, Evelyn Marie, Hennemann, Frank H., Le Tirant, Stephane, Daawia,, Warikar, Evie Lilly, Yando, Heron, Suhartawan, Bambang, Henze, Katharina, Büscher, Thies H., Bank, Sarah (2025): A deeper look into the diversity of Phyllium leaf insects from Indonesia: seven new species and two unique egg morphologies (Phasmatodea, Phylliidae). ZooKeys 1256: 317-370, DOI: 10.3897/zookeys.1256.162609
81311371E6DF568DA8BA8EE75FEC0263.taxon	etymology	Etymology. Patronym; named to honor Thierry Boislard (Canada), a colleague of the Montreal Insectarium and a good friend to the fourth author.	en	Cumming, Royce T., Foley, Evelyn Marie, Hennemann, Frank H., Le Tirant, Stephane, Daawia,, Warikar, Evie Lilly, Yando, Heron, Suhartawan, Bambang, Henze, Katharina, Büscher, Thies H., Bank, Sarah (2025): A deeper look into the diversity of Phyllium leaf insects from Indonesia: seven new species and two unique egg morphologies (Phasmatodea, Phylliidae). ZooKeys 1256: 317-370, DOI: 10.3897/zookeys.1256.162609
81311371E6DF568DA8BA8EE75FEC0263.taxon	distribution	Distribution. At present only known from the type locality of Malinau, in North Kalimantan, Indonesia (Fig. 2).	en	Cumming, Royce T., Foley, Evelyn Marie, Hennemann, Frank H., Le Tirant, Stephane, Daawia,, Warikar, Evie Lilly, Yando, Heron, Suhartawan, Bambang, Henze, Katharina, Büscher, Thies H., Bank, Sarah (2025): A deeper look into the diversity of Phyllium leaf insects from Indonesia: seven new species and two unique egg morphologies (Phasmatodea, Phylliidae). ZooKeys 1256: 317-370, DOI: 10.3897/zookeys.1256.162609
CE7CFC045E7A562B97920ED659BA3E63.taxon	description	Figs 6, 7, 8, 9, 10	en	Cumming, Royce T., Foley, Evelyn Marie, Hennemann, Frank H., Le Tirant, Stephane, Daawia,, Warikar, Evie Lilly, Yando, Heron, Suhartawan, Bambang, Henze, Katharina, Büscher, Thies H., Bank, Sarah (2025): A deeper look into the diversity of Phyllium leaf insects from Indonesia: seven new species and two unique egg morphologies (Phasmatodea, Phylliidae). ZooKeys 1256: 317-370, DOI: 10.3897/zookeys.1256.162609
CE7CFC045E7A562B97920ED659BA3E63.taxon	description	Description. Female. Coloration. Coloration description is based upon photos of the type material shared with the authors of the live specimens prior to preservation. The general coloration is pale green throughout. The only areas that differ are the antennae that are somewhat orange / tan and some of the more prominent veins of the tegmina which are brown. Morphology. Head capsule longer than wide, with a vertex that is somewhat roughly textured, and marked with minimal granulation along the posterior (Fig. 6 A). The posteromedial tubercle is present, singularly lobed, but not very prominent (Fig. 6 A). Frontal convexity is broad and ending in a blunted point; there are several short setae across the surface. Compound eyes slightly protruding from the head capsule, not bulbous, taking up ~ ¼ of the head capsule lateral margins (Fig. 6 A). Ocelli absent. Antennal fields slightly wider than the first antennomere width. Antennae consist of ten segments, with the terminal segment approx. the same length as the preceding 2 ½ segments’ lengths combined (Fig. 10 A). Antennomeres I – VIII are smooth, and sparsely marked with short setae, the terminal two antennomeres are covered in short, dense setae, giving these segments a fuzzy appearance (Fig. 10 A). Stridulatory file of antennomere III has 33 – 35 teeth, and the stridulatory ridge has 33 or 34 teeth (Fig. 10 B). Thorax. Pronotum with slightly concave anterior margin and lateral margins that anteriorly start wide, angle inward strongly to the posterior margin which is ~ ½ the width of the anterior margin (Fig. 6 A). The pronotum anterior margin and the lateral margins have prominent rims, while the posterior margin is less prominent. The pronotum surface is relatively smooth, with only a prominent pit in the center and a distinct furrow anterior to the center (Fig. 6 A). Prosternum and the anterior 1 / 3 of the mesosternum are covered with irregularly spaced granulation; the remainder of the mesosternum and the metasternum are slightly wrinkled but lack nodes. Mesoprescutum slightly longer than wide, lateral rims with six or seven small tubercles (Fig. 6 E). Mesoprescutum anterior rim prominently raised into a distinct and finely pointed sagittal spine (Fig. 6 C). Mesoprescutum surface slightly lumpy and only slightly raised along the sagittal crest, which has several haphazardly located nodes along its length, not perfectly aligned along the plane (Fig. 6 E). Mesopleurae narrow for the anterior 1 / 3 until they begin to diverge and angle prominently away with nearly straight margins (Fig. 6 E). Mesopleuron lateral margin with six or seven medium sized nodes, mostly situated on the anterior 1 / 2, and the posterior 1 / 2 only has some minimal granulation (Fig. 6 E). Face of the mesopleuron slightly wrinkled (Fig. 6 E). Wings. Tegmina long, reaching the anterior margin of abdominal segment VIII. Tegmina venation; the subcosta (Sc) is the first vein in the forewing, running parallel with the margin for the first 1 / 2, and then bending and running towards the margin. The subcosta runs for ~ ¼ of the tegmina length. The radius (R) spans the central portion of the forewing with two subparallel (slightly diverging) branched veins; the first radius (R 1) branches ~ ¼ of the way through the wing length and terminates ~ 1 / 3 of the way through the wing length; the radial sector (Rs) branches ~ 1 / 3 of the way through the wing length and terminates near the distal 1 / 3 of the wing length. There is a weak continuation of the radius following the prominent Rs branching which continues on as a short but distinct R – M crossvein that weakly connects the two veins. The media (M) is bifurcate with both the media anterior (MA) and media posterior (MP) terminating near to the posterior of the tegmina. The cubitus (Cu) is also bifurcate, branching near the posterior ¼ of the wing into the cubitus anterior (CuA) and cubitus posterior (CuP) which both terminate near the wing apex. The first anal vein (1 A) is simple and fuses with the cubitus ~ ¼ of the way through the tegmina length. Alae vestigial, only small nubs. Abdomen. Abdominal segments II through the anterior ½ of IV gradually and uniformly diverging. The posterior ½ of segment IV through the anterior ½ of segment VII are only slightly diverging to the widest point of the abdomen. Abdominal segment VII is rounded ca 90 degrees with posterior margins angled almost directly inward where they meet the notably narrower segment VIII. Segments VIII – X have straight, converging margins ending in a broad rounded apex (Fig. 6 G). Genitalia. Subgenital plate starts at the anterior margin of tergum VIII, is moderately broad, and extends ¾ of the way onto tergum X. The shape is somewhat tiered into approximately three widths as it converges to a finely pointed apex (Fig. 6 F). Gonapophyses VIII are long and not particularly broad, exceeding the apex of the abdominal tergum X slightly; gonapophyses IX are obstructed from view (Fig. 6 F). Cerci flat, slightly broadening to the apical 1 / 3 into a somewhat blade-like end, with a slightly granular surface (Fig. 6 F). Legs. Profemoral exterior lobe broad and arching end to end, with a width slightly narrower than the width of the interior lobe (Fig. 6 A). Margin of the profemoral exterior lobe slightly granular (Fig. 6 A). Profemoral interior lobe slightly more than 2 × as wide as the greatest width of the profemoral shaft, approximately right angled, and marked with four large, triangular teeth with looping gaps between them, arranged in a two-wide gap-two pattern (Fig. 6 A). Mesofemoral interior and exterior lobes approx. as wide as the mesofemoral shaft width. Mesofemoral exterior lobe with two small, distally pointing teeth on the distal ½ of the lobe with a wide gap between them. Mesofemoral exterior lobe is somewhat angled, not as smoothly arching as in the interior lobe. Mesofemoral interior lobe with six small, distally pointing teeth on the distal 2 / 3 of the lobe, with the teeth somewhat arranged into pairs. Metafemoral interior lobe arcs end to end, with the proximal 1 / 3 notably thinner and smooth and slightly widening out to the distal 2 / 3 which is wider and armed with seven or eight dulled, small teeth. Metafemoral exterior lobe lacks dentation, is thinner than the shaft width, and runs parallel with the shaft throughout its length. Protibia exterior lacking a lobe (Fig. 6 A). Protibiae interior lobe spans the entire length of the protibiae (although the distal end is very thin and not prominent) and is ~ 1.5 × the width of the protibiae shaft itself. The lobe is roundly triangular with the widest portion near the middle, and the proximal end more distinct than the distal end. Mesotibiae and metatibiae simple, lacking exterior and interior lobes. Measurements (mm). Holotype, female: body length (including cerci and head, excluding antennae): 85.9, length / width of head: 8.3 / 6.8, antennae: 4.4, pronotum: 5.9, mesonotum: 8.2, length of tegmina: 54.0, greatest width of abdomen: 36.7, profemora: 14.9, mesofemora: 13.5, metafemora: 17.1, protibia: 9.8, mesotibia: 10.5, metatibia: 15.0. Male. Coloration. Coloration based upon the dead, dried type specimen which is somewhat discolored (Fig. 7). Overall coloration pale green / yellow throughout. The compound eyes are brownish-red, and the antennae are darker / gray. Protibia interior lobe and margin of the profemoral interior lobe with several brown / tan markings Morphology. Head capsule slightly longer than wide, with a vertex that is slightly lumpy and marked throughout with a few small, widely distributed nodes (Fig. 7 A). The posteromedial tubercle is singularly pointed, small, and not notably raised above the head capsule (Fig. 7 A, F). Frontal convexities stout and bluntly pointed with a few sparse setae. Compound eyes large and bulbous, occupying ~ 2 / 5 of the head capsule lateral margins (Fig. 7 A). There are three distinct ocelli raised above the capsule and located between the compound eyes (Fig. 7 A). Antennae (including the scapus and pedicellus) consist of 25 segments (Fig. 7 A), all segments except the scapus and pedicellus and terminal five segments are covered in numerous setae where most are as long as the antenna segment is wide. The terminal five segments are covered in dense, short setae and the scapus and pedicellus are nearly completely bare with only a few sparse setae. Thorax. Pronotum with slightly convex anterior margin and straight lateral and posterior margins. Posterior margin is ~ ½ the width of the anterior margin. The anterior margin is well-developed, the lateral margins are moderately formed, and the posterior margin is weakly formed (Fig. 7 D). Face of the pronotum is marked by a distinct pit in the center, a sagittal furrow on the anterior ½, and slight perpendicular furrows originating from the central pit. The pronotum surface is only slightly lumpy but lacking distinct granulation (Fig. 7 D). Prosternum surface is lumpy with small nodes. Mesosternum surface anterior 1 / 3 marked heavily with granulation, the remainder of the mesosternum surface is wrinkled but lacks notable nodes. Metasternum surface mostly wrinkled throughout, and the anterior margin central area is additionally marked with granulation. Mesoprescutum longer than wide, with lateral margins that are slightly converging to the posterior margin which is only slightly narrower than the anterior margin (Fig. 7 D). Lateral margins of the mesoprescutum with five or six moderately formed tubercles of a somewhat uniform size (Fig. 7 D). Mesoprescutum surface wrinkled and slightly raised along the sagittal plane which is marked with four or five distinct nodes and the remainder of the surface has slight granulation (Fig. 7 D, F). Mesoprescutum anterior rim moderately formed with a distinct sagittal spine, and the remainder of the rim surface is slightly wrinkled (Fig. 7 D, F). Mesopleurae begin on the anterior mesoprescutum margin, begin very narrow, and diverge slowly at a gradually increasing angle from the anterior to the posterior but are never notably wide throughout their length (Fig. 7 D). Mesopleuron lateral margin with four or five small tubercles and a few small nodes interspersed throughout the length except for the posterior 1 / 3 of the margin which is relatively smooth (Fig. 7 D). Mesopleuron face moderately wrinkled and marked by a distinct pit near the center. Wings. Tegmina moderate length, extending ¾ of the way onto abdominal segment IV. Tegmina wing venation: the subcosta (Sc) is the first vein, is simple, and terminates ~ 1 / 3 of the way through the overall wing length. The radius (R) spans nearly the entire length of the tegmina with the first radius (R 1) branching ~ 1 / 3 of the way through the tegmina length and terminates on the margin slightly less than ½ through the length. There is also a second radius (R 2) that originates near the middle, and a third radius (R 3) which originates ~ 3 / 5 of the way through the wing length. The radial sector runs towards the wing apex, but near the terminal 1 / 3 angles towards the margin, slightly away from the apex. The media (M) spans the entire length of the tegmina running side by side along the radius / radial sector for most of the length, with a first media posterior (MP 1) branching off near the midlength of the tegmina and running angled towards the cubitus, a second media posterior (MP 2) branches off ~ 3 / 5 of the way through the length, and the media anterior (MA) runs straight to the tegmina apex. The cubitus (Cu) cuts across the tegmina to the margin ~ 1 / 3 of the way through the length and runs along the edge of the tegmina where the media posterior veins fuse with it and as the cubitus reaches the apex it fades. The first anal (1 A) vein terminates upon reaching the cubitus ~ 1 / 3 of the way through the tegmina length. Alae well-developed in an oval fan configuration, long, reaching to the middle of abdominal segments VIII. Ala wing venation hidden from view due to the alae being folded in the type specimen. Abdomen. Lateral margins of abdominal segment II parallel, III diverging slightly, IV diverging at a more prominent angle for the anterior 2 / 3 and then slightly less strongly for the posterior 1 / 3, V slightly diverging, VI diverges slightly for the anterior 2 / 3 to the widest point of the abdomen, then converges on the posterior 1 / 3, followed by all preceding segments converging strongly at first and then more gradually towards the apex (segment X), which is broad and rounded. Genitalia. Poculum broad and ends with a flat, blunted apex that slightly passes the anterior margin of abdominal segment X with a margin that is nearly straight (Fig. 7 E). Cerci long, slender, and nearly flat, with subparallel margins, with ~ 2 / 3 of their length extending from under abdominal segment X. The cerci surfaces are slightly granular and there are numerous short setae along the margins (Fig. 7 E). Vomer broad and stout with straight sides evenly converging to the apical hook which is thick and has a singular point (Fig. 7 E). Legs. The profemoral exterior lobe arcs end to end and is narrow, approx. the same width as the profemoral shaft at its widest. The profemoral exterior lobe margin is slightly granular (Fig. 7 A). The profemoral interior lobe is obtusely triangular and at its greatest width it is ~ 2 × the greatest width of the profemoral shaft. The profemoral interior lobe is ornamented on the distal ½ with four serrate teeth arranged as small tooth-large tooth-wide gap-large tooth-small tooth (Fig. 7 A). Mesofemoral exterior lobe and interior lobe are of similar shapes and widths, both arching end to end but are slightly wider on the distal 2 / 3. Both lobes at their widest are approx. as wide as the mesofemoral shaft width. The only notable difference between these two lobes is that the interior lobe has five small teeth on the distal 1 / 2. The mesofemoral exterior lobe is unornamented. Metafemoral exterior lobe lacks dentition and has a straight, thin margin along the metafemoral shaft. Metafemoral interior lobe is approx. as wide as the metafemoral shaft width, arcs end to end, but is thinner on the proximal 1 / 3, the distal 1 / 3 is marked with six small serrate teeth. Protibiae lacking exterior lobe, interior lobe mostly situated in the middle of the shaft with the distal end without lobe and the proximal end very thin. The protibial interior lobe is a small, rounded triangle with the widest portion just distal to the midlength (Fig. 7 A). Meso- and metatibiae simple, lacking lobes completely. Measurements of paratype male [mm]. Length of body (including cerci and head, excluding antennae) 65.4, length / width of head 4.4 / 3.8, antenna 26.2, pronotum 3.5, mesonotum 4.9, length of tegmina 24.5, length of alae 42.4, greatest width of abdomen 20.2, profemora 10.4, mesofemora 10.0, metafemora 11.0, protibiae 6.5, mesotibiae 7.2, metatibiae 10.8. Eggs (Figs 8, 9). The overall egg shape is difficult to discern fully due to the long and dense pinnae, but appears to be somewhat reniform, with the ventral surface protruding slightly and the dorsal surface slightly curved inward. The entire egg capsule is covered by spatulate pinnae with short chorionic outgrowths (Fig. 9 B) and hollow, columnar pinnae densely covering the entire egg surface (Fig. 9 D). These two types are somewhat interspersed, but the roughly textured spatulate pinnae are more prominent along the anterior and posterior margins, while the smoother columnar pinnae are more prominent on the flat surfaces. The operculum is nearly circular and rimmed by the roughly textured spatulate pinnae which are slightly shorter than the pinnae on the remainder of the egg capsule. The micropylar plate is mostly obstructed from view but appears to be thin and only situated in the middle of the egg capsule. The capsule surface below the dense pinnae appears to be rather smooth and paler in color than the dark brown pinnae covering the surface.	en	Cumming, Royce T., Foley, Evelyn Marie, Hennemann, Frank H., Le Tirant, Stephane, Daawia,, Warikar, Evie Lilly, Yando, Heron, Suhartawan, Bambang, Henze, Katharina, Büscher, Thies H., Bank, Sarah (2025): A deeper look into the diversity of Phyllium leaf insects from Indonesia: seven new species and two unique egg morphologies (Phasmatodea, Phylliidae). ZooKeys 1256: 317-370, DOI: 10.3897/zookeys.1256.162609
CE7CFC045E7A562B97920ED659BA3E63.taxon	description	Measurements including the extended pinnae [mm]. Length (including operculum expansion): 6.0 – 6.6; maximum width of capsule when viewed from lateral aspect 4.8 – 5.5; length of micropylar plate 2.8 – 3.4.	en	Cumming, Royce T., Foley, Evelyn Marie, Hennemann, Frank H., Le Tirant, Stephane, Daawia,, Warikar, Evie Lilly, Yando, Heron, Suhartawan, Bambang, Henze, Katharina, Büscher, Thies H., Bank, Sarah (2025): A deeper look into the diversity of Phyllium leaf insects from Indonesia: seven new species and two unique egg morphologies (Phasmatodea, Phylliidae). ZooKeys 1256: 317-370, DOI: 10.3897/zookeys.1256.162609
CE7CFC045E7A562B97920ED659BA3E63.taxon	etymology	Etymology. Patronym; named to honor Victor Cayabyab, a collaborator of the Montreal Insectarium for the last 30 years and a very good friend to the fourth author.	en	Cumming, Royce T., Foley, Evelyn Marie, Hennemann, Frank H., Le Tirant, Stephane, Daawia,, Warikar, Evie Lilly, Yando, Heron, Suhartawan, Bambang, Henze, Katharina, Büscher, Thies H., Bank, Sarah (2025): A deeper look into the diversity of Phyllium leaf insects from Indonesia: seven new species and two unique egg morphologies (Phasmatodea, Phylliidae). ZooKeys 1256: 317-370, DOI: 10.3897/zookeys.1256.162609
CE7CFC045E7A562B97920ED659BA3E63.taxon	distribution	Distribution. At present only known from the type locality of Malinau, in North Kalimantan, Indonesia (Fig. 2).	en	Cumming, Royce T., Foley, Evelyn Marie, Hennemann, Frank H., Le Tirant, Stephane, Daawia,, Warikar, Evie Lilly, Yando, Heron, Suhartawan, Bambang, Henze, Katharina, Büscher, Thies H., Bank, Sarah (2025): A deeper look into the diversity of Phyllium leaf insects from Indonesia: seven new species and two unique egg morphologies (Phasmatodea, Phylliidae). ZooKeys 1256: 317-370, DOI: 10.3897/zookeys.1256.162609
ECA53CA0D5FC586F814588BAAB6AB0B6.taxon	description	Fig. 11	en	Cumming, Royce T., Foley, Evelyn Marie, Hennemann, Frank H., Le Tirant, Stephane, Daawia,, Warikar, Evie Lilly, Yando, Heron, Suhartawan, Bambang, Henze, Katharina, Büscher, Thies H., Bank, Sarah (2025): A deeper look into the diversity of Phyllium leaf insects from Indonesia: seven new species and two unique egg morphologies (Phasmatodea, Phylliidae). ZooKeys 1256: 317-370, DOI: 10.3897/zookeys.1256.162609
ECA53CA0D5FC586F814588BAAB6AB0B6.taxon	description	Description. Male. Coloration. Coloration based upon the dead, dried holotype (Fig. 11). Overall coloration pale green and yellow throughout with highlights of tan / orange. The antennae and compound eyes are the darkest areas on the specimen with burnt orange color. Abdominal segment V has a pair of slightly lighter eye spots that are not prominent. Ventral coxae coloration matches the surrounding tissue. Morphology. Head capsule slightly longer than wide, with a vertex that is rather smooth. The posteromedial tubercle is singularly pointed and distinctly raised above the head capsule (Fig. 11 E). Frontal convexities stout and bluntly pointed with sparse setae. Compound eyes large and bulbous, occupying ~ 2 / 5 of the head capsule lateral margins (Fig. 11 D). There are three well-developed ocelli distinctly raised above the capsule and located between the compound eyes. Antennae (including the scapus and pedicellus) consist of 25 segments, all segments except the scapus and pedicellus and terminal five segments are covered in dense setae where most are as long as the antenna segment is wide (Fig. 11 D). The terminal five segments are covered in dense short setae and the scapus and pedicellus are nearly completely bare with only a few sparse setae. Thorax. Pronotum with slightly concave anterior margin and straight lateral margins that converge to a slightly convex posterior margin that is ~ ½ the width of the anterior margin (Fig. 11 A). Anterior and lateral margins of the pronotum have distinct rims and the posterior margin has a weakly formed rim (Fig. 11 A). Face of the pronotum is marked by a distinct pit in the center, a sagittal furrow on the anterior ½, and slight perpendicular furrows originating from the central pit. The pronotum surface is slightly lumpy and lacking distinct granulation (Fig. 11 A). Prosternum surface is slightly granular. Mesosternum surface anterior 1 / 3 slightly granular, the remainder of the mesosternum surface is relatively smooth (Fig. 11 B). Metasternum surface finely wrinkled throughout. Mesoprescutum longer than wide, with lateral margins that slightly converge to the posterior margin which is ~ ¾ as wide as the anterior margin (Fig. 11 A). Lateral margins of the mesoprescutum with six or seven stout tubercles with the anterior three the most prominent (Fig. 11 A). Mesoprescutum surface slightly raised along the sagittal plane and is marked with three or four distinct but small spines and the remainder of the surface is relatively smooth (Fig. 11 A). Mesoprescutum anterior rim moderately formed with a distinct sagittal spine, and the remainder of the rim surface is relatively smooth (Fig. 11 E). Mesopleurae begin on the anterior mesoprescutum margin and diverge at a gradually increasing angle from the anterior to the posterior but are relatively narrow throughout their length (Fig. 11 A). Mesopleuron lateral margin with seven or eight moderately formed tubercles and a few small nodes interspersed throughout the length (Fig. 11 A). Mesopleuron face moderately wrinkled and marked by a distinct pit on the anterior 1 / 3 and a smaller pit on the posterior 1 / 3. Wings. Tegmina moderate length, extending ¼ of the way onto abdominal segment IV. Tegmina wing venation: the subcosta (Sc) is the first vein, is simple, and terminates ~ 1 / 3 of the way through the overall wing length. The radius (R) spans the entire length of the tegmina with the first radius (R 1) branching ~ 1 / 3 of the way through the wing length and terminating at the wing margin ~ ½ of the way through the wing length. There is an additional radius (R 2) branching near the midlength of the tegmina and the radial sector runs straight to the wing apex. The media (M) also spans the entire length of the tegmina and runs side by side along the radius / radial sector with the first media posterior (MP 1) branching off near the tegmina midlength, followed by a second media posterior (MP 2) near the distal 2 / 5, and the media anterior (MA) runs straight to the tegmina apex. The cubitus (Cu) cuts across the tegmina to the margin ~ 1 / 3 of the way through the length and runs along the edge of the wing where the first and second media posterior veins fuse with it and as the cubitus reaches the apex it fades. The first anal (1 A) vein terminates upon reaching the cubitus 1 / 3 of the way through the tegmina length. Alae well-developed in an oval fan configuration, long, reaching to the posterior of abdominal segments IX. Ala wing venation not visible in the holotype specimen. Abdomen. Lateral margins of abdominal segment II slightly converging, III diverging slightly, IV diverging to the widest point 2 / 3 of the way through the segment length and the remaining 1 / 3 of the segment is parallel, V parallel sided or slightly subparallel (converging slightly), VI through X converging gradually with smooth margins, giving the abdomen a spade-shaped appearance. Genitalia. Poculum broad and ends in an apex that slightly passes the anterior margin of the abdominal segment X with a margin that is broad and straight (Fig. 11 F). Cerci long and slender, with slightly> ½ of their length extending from under abdominal segment X, nearly flat, covered in a granulose surface and several short setae with those along the margins slightly longer (Fig. 11 F). Vomer broad and stout with straight sides evenly converging until near the apex where the margins converge more sharply to the apical hook which is thick and has a singular point (Fig. 11 F). Legs. The profemoral exterior lobe is narrow, approx. the same width as the profemoral shaft at its widest. The profemoral exterior lobe margin is relatively smooth (Fig. 11 D). The profemoral interior lobe is obtusely triangular and at its greatest width it is ~ 2 × the greatest width of the profemoral shaft. The profemoral interior lobe is ornamented with four serrate teeth arranged in a three-one pattern with looping gaps between them, where the third and fourth tooth has a notable wider gap between them (Fig. 11 D). The central two teeth are notably larger than the first and fourth teeth (Fig. 11 D). Mesofemoral exterior lobe arcs end to end but is slightly wider on the distal 1 / 3 which is marked with two teeth, while the proximal 2 / 3 of the lobe is thinner and lacks teeth. Mesofemoral interior lobe and mesofemoral shaft are approximately the same width, and the exterior lobe is slightly thicker. The mesofemoral interior lobe is slightly broader on the distal end and the distal ½ is ornamented with six small serrate teeth while the proximal portion of the lobe is thin and lacks teeth. Metafemoral exterior lobe has a straight margin along the metafemoral shaft and is mostly unornamented but does have two small teeth on the distal 1 / 3. Metafemoral interior lobe arcs end to end with nine or ten sharply serrate teeth on the distal ½, which is slightly wider than the smooth proximal portion of the lobe. Protibia lacking exterior lobe; interior lobe reaching end to end as a rounded triangle with the widest portion slightly distal to the midlength with a maximum width of ~ 1.5 × as wide as the protibial shaft width (Fig. 11 D). Meso- and metatibiae simple, lacking lobes completely. Measurements of holotype male [mm]. Length of body (including cerci and head, excluding antennae) 63.8, length / width of head 4.4 / 3.8, antennae 28.4, pronotum 3.4, mesonotum 4.9, length of tegmina 25.0, greatest width of abdomen 16.3, profemora 10.9, mesofemora 10.3, metafemora 12.8, protibiae 6.5, mesotibiae 6.6, metatibiae 9.8.	en	Cumming, Royce T., Foley, Evelyn Marie, Hennemann, Frank H., Le Tirant, Stephane, Daawia,, Warikar, Evie Lilly, Yando, Heron, Suhartawan, Bambang, Henze, Katharina, Büscher, Thies H., Bank, Sarah (2025): A deeper look into the diversity of Phyllium leaf insects from Indonesia: seven new species and two unique egg morphologies (Phasmatodea, Phylliidae). ZooKeys 1256: 317-370, DOI: 10.3897/zookeys.1256.162609
ECA53CA0D5FC586F814588BAAB6AB0B6.taxon	etymology	Etymology. Latin in origin, adjective. Meaning drunk or intoxicated. For anyone who has seen living phylliids, when in the wild on a bush or in a tree, they blend completely with the leaves swaying in the breeze. However, most people rarely see them in the wild, more often they are seen handled as pets indoors or in terrariums which lack a gusty breeze. This results in the leaf insect swaying wildly, as is their natural predisposition (Leigh 1912; Steiniger 1933), but looking very much like a drunken leaf stumbling home after a wild evening of too much merriment. Hence, the binomial meaning “ drunken leaf ”.	en	Cumming, Royce T., Foley, Evelyn Marie, Hennemann, Frank H., Le Tirant, Stephane, Daawia,, Warikar, Evie Lilly, Yando, Heron, Suhartawan, Bambang, Henze, Katharina, Büscher, Thies H., Bank, Sarah (2025): A deeper look into the diversity of Phyllium leaf insects from Indonesia: seven new species and two unique egg morphologies (Phasmatodea, Phylliidae). ZooKeys 1256: 317-370, DOI: 10.3897/zookeys.1256.162609
ECA53CA0D5FC586F814588BAAB6AB0B6.taxon	distribution	Distribution. At present known from the type locality; Mount Bawang in West Kalimantan, Indonesia (Fig. 2).	en	Cumming, Royce T., Foley, Evelyn Marie, Hennemann, Frank H., Le Tirant, Stephane, Daawia,, Warikar, Evie Lilly, Yando, Heron, Suhartawan, Bambang, Henze, Katharina, Büscher, Thies H., Bank, Sarah (2025): A deeper look into the diversity of Phyllium leaf insects from Indonesia: seven new species and two unique egg morphologies (Phasmatodea, Phylliidae). ZooKeys 1256: 317-370, DOI: 10.3897/zookeys.1256.162609
371848E37B985710B15D86D1AEBA6D41.taxon	description	Figs 12, 13, 14	en	Cumming, Royce T., Foley, Evelyn Marie, Hennemann, Frank H., Le Tirant, Stephane, Daawia,, Warikar, Evie Lilly, Yando, Heron, Suhartawan, Bambang, Henze, Katharina, Büscher, Thies H., Bank, Sarah (2025): A deeper look into the diversity of Phyllium leaf insects from Indonesia: seven new species and two unique egg morphologies (Phasmatodea, Phylliidae). ZooKeys 1256: 317-370, DOI: 10.3897/zookeys.1256.162609
371848E37B985710B15D86D1AEBA6D41.taxon	description	Description. Female. Coloration. Coloration description is based upon the type material, which is dead and dried reasonably well (not too many dark rotten areas; Fig. 13). The general coloration is pale green throughout (although some areas have faded to yellow, likely due to the drying process). There are several areas, commonly occurring on other Phyllium species, which appear to be more variably marked with muddled brown / tan coloration; these areas are: the protibiae, profemoral interior lobe, mesofemoral lobes, and the margins of abdominal segments VII and VIII. Meso- and metacoxae ventrally marked with a dark spot. Morphology. Head capsule approximately as long as wide, with a vertex that is smooth, lacking granulation (Fig. 13 B). The posteromedial tubercle is present, singularly lobed, but not very prominent (Fig. 13 B). Frontal convexity broad and ending in a blunted point; there are several short setae across the surface. Compound eyes slightly protruding from the head capsule, not bulbous, taking up ~ ¼ of the head capsule lateral margins (Fig. 13 B). Ocelli absent. Antennal fields slightly wider than the first antennomere width. Antennae consist of ten segments, with the terminal segment the narrowest and approx. the same length as the preceding four segments’ lengths combined (Fig. 13 B). Antennomeres VIII and IX appear derived from a single segment as they are tightly situated, but have a prominent suture separating them into two, and each segment has a distinct texture, with segment IX a rough, fuzzy texture (like segment X) and segment VIII smoother with sparse setae (like on segment VII; Fig. 13 B). Antennomeres I – VIII are smooth, and sparsely marked with short setae, the terminal two antennomeres are covered in short, dense setae, giving these segments a fuzzy appearance (Fig. 13 B). Thorax. Pronotum with slightly concave anterior margin and lateral margins that anteriorly start wide, angle inward strongly, then for the middle portion are angled more gently, followed by a strong incurve to the posterior margin (Fig. 13 F). The posterior margin is ~ ½ the width of the anterior margin (Fig. 13 F). The pronotum anterior margin has a prominent rim, while the lateral and posterior margins are less prominent. The pronotum surface is relatively smooth, with only a prominent pit in the center and a few furrows (Fig. 13 F). Prosternum, mesosternum, and metanotum are covered throughout by moderately spaced granulation (Fig. 13 D). Mesoprescutum slightly longer than wide, lateral rims with eight or nine nodes (none particularly prominent; Fig. 13 F). Mesoprescutum anterior rim prominently raised into a raised, broad sagittal spine (Fig. 13 F). Mesoprescutum surface smooth except for the slightly raised mesoprescutum sagittal crest marked with variably granulation throughout the length, but only along the sagittal crest; areas lateral to the sagittal crest smooth (Fig. 13 F). Mesopleurae begin to diverge ~ 1 / 3 of the way along the mesoprescutum, angle prominently away with straight margins (Fig. 13 F). Mesopleurae lateral margins with six or seven nodes with interspersed granulation throughout, giving the margin a rough textured appearance (Fig. 13 F). Face of the mesopleura slightly wrinkled, with two notable divots, one on the anterior margin and one near the middle (Fig. 13 F). Wings. Tegmina long, reaching onto abdominal segment VIII. Tegmina venation; the subcosta (Sc) is the first vein in the forewing, running parallel with the margin for the first ½, and then bending and running towards the margin. The subcosta runs for ~ 1 / 3 of the tegmina length. The radius (R) spans the central portion of the forewing with two subparallel branched veins; the first radius (R 1) branches ~ ¼ of the way through the wing length and terminates slightly proximal to the midline; the radial sector (Rs) branches ~ 1 / 3 of the way through the wing length and terminates near the distal 1 / 3 of the wing length. There is a weak continuation of the radius following the prominent Rs branching which continues on as a short but distinct R – M crossvein that weakly connects the two veins. The media (M) is bifurcate with both the media anterior (MA) and media posterior (MP) terminating near to the posterior of the tegmina. There is a weak continuation of the media following the prominent media posterior (MP) branching which continues on as a somewhat long M – Cu crossvein that fades before fully connecting the two veins. The cubitus (Cu) is also bifurcate, branching near the posterior ¼ of the wing into the cubitus anterior (CuA) and cubitus posterior (CuP) which both terminate near the wing apex. The first anal vein (1 A) is simple and fuses with the cubitus ~ 1 / 3 of the way through the tegmina length. Alae vestigial, with their apex only just reaching abdominal segment I (~ 6.0 mm long as measured in a paratype). Abdomen. Abdominal segments II through the anterior 2 / 3 of IV gradually diverging. The posterior 1 / 3 of segment IV through the anterior 2 / 3 of segment VII are gradually and uniformly converging. The posterior 1 / 3 of segment VII is rounded inwards towards segment VIII which like VII starts converging gradually and then rounds inward to segment IX. Segments IX – X have straight, converging margins ending in a broad rounded apex (Fig. 13 H). Genitalia. Subgenital plate starts at the anterior margin of tergum VIII, is moderately broad, and extends most of the way onto tergum X. The shape is approximately teardrop-shaped, with the apex a fine point (Fig. 3 E). Gonapophyses VIII are long and moderately broad, reaching the apex of abdominal tergum X; gonapophyses IX are obstructed from view (Fig. 13 H). Cerci flat, slightly broadening to the apical ¼, with a slightly granular surface (Fig. 13 H). Legs. Profemoral exterior lobe broad, rounded, and obtusely angled, slightly narrower than the width of the interior lobe (Fig. 13 C). Distal margin of the profemora with three small, finely pointed teeth (Fig. 13 C). Profemoral interior lobe ~ 3 × as wide as the greatest width of the profemoral shaft, approximately right angled, and marked with three large teeth with looping gaps between them, each gap with a singular smaller tooth (Fig. 13 C). Mesofemoral lobes significantly broadened on the distal 1 / 3, with the interior lobe greatest width ~ 2 × wider than the mesofemoral shaft width, and the exterior lobe greatest width ~ 1.5 × wider than the mesofemoral shaft width. Mesofemoral exterior lobe with three or four small, distally pointing teeth on the distal 1 / 3 of the lobe. Mesofemoral interior lobe with five small, distally pointing teeth on the distal 1 / 3 of the lobe. Metafemoral interior lobe arcs end to end, with the distal ½ slightly wider than the proximal ½ and marked with five or six serrate teeth on the distal ½ of the lobe. Metafemoral exterior lobe lacks dentation and has a width similar to the metafemoral shaft width. Protibiae exterior has a thin expansion near the middle, less than the width of the protibial shaft (Fig. 13 C). Protibiae interior lobe spans the entire length of the protibiae and is ~ 2.5 × the width of the protibiae shaft itself. The lobe is roundly triangular with the widest portion on the distal ½. Mesotibiae and metatibiae simple, lacking exterior and interior lobes. Measurements of holotype female [mm]. Length of body (including cerci and head, excluding antennae) 88.0, antennae 5.8, pronotum 5.9, mesonotum 10.5, length of tegmina 54.7, greatest width of abdomen 32.0, profemora 16.8, mesofemora 15.0, metafemora 18.5, protibiae 11.2, mesotibiae 10.3, metatibiae 15.0. Measurements of paratype females [mm]. Length of body (including cerci and head, excluding antennae) 73.0 – 77.0, antennae 5.0 – 5.5, pronotum 5.2 – 5.5, mesonotum 9.0 – 9.5, metanotum 9.0, length of tegmina 46.0 – 50.0, length of alae 6.0, greatest width of abdomen 28.5 – 29.0, profemora 14.5 – 15.0, mesofemora 13.0 – 14.0, metafemora 15.0 – 16.0, protibiae 9.0 – 10.5, mesotibiae 9.0 – 9.5, metatibiae 13.0 – 14.5. Eggs (Figs 12, 14). The lateral flaps and the fused capitular frill are brown; the feather-like pinnae are tan; and the exposed, smooth sides of the egg capsule under the lateral flaps are yellowish / tan in color. The actual egg capsule is notably smaller than the habitus due to the lateral flaps covering the entire lateral surfaces. These lateral flaps are connected to the capsule by a small, stiff section on the posterior of the egg. These lateral flaps are slightly convex and have an irregular, somewhat lumpy surface that lacks pinnae, instead the raised lumpy areas are slightly lighter in color but are smooth in texture. On the microscopical level fine hair-like protrusions cover the surface of the flap (Fig. 14 D). These lateral flaps are held slightly away from the actual egg capsule, with the anterior margin simple and open, while the lateral flaps lateral margins are curled under slightly, but this rim does not rest on the egg capsule. A thin membrane runs along the inner side of the flap sealing the space between flap and capsule (Fig. 14 H). The actual egg capsule lateral surface is slightly convex, with a smooth, only slightly lumpy surface (Fig. 14 B). The dorsal surface has a thin micropylar plate which is ~ ½ the capsule length, but it is situated on the posterior 1 / 2 of the capsule (Fig. 12 A). Running along each side of the micropylar plate is a continuous line of short, feather-like pinnae. The micropylar plate is thin, really only a slit with the widest portion around the small micropylar cup which is on the posterior ¼ of the capsule (Fig. 12 A). These feather-like pinnae run fully around the sagittal plane of the egg, so when it is viewed laterally, the pinnae fully surround the capsule, with only the fused capitular frill projecting above the anterior pinnae (Fig. 12 B). The operculum is ovular, and the outer margin has a fused capitular frill fully surrounding the opercular margin (Fig. 14 C). This frill is fully fused around the entire margin and is prominently projecting above the flat, smooth surface of the cap, leaving only a narrow, long opening along the top of the egg (Fig. 12 E). The fused capitular frill is roughly textured, appearing slightly fuzzy. The ventral surface of the egg capsule has the continuous encircling feather-like pinnae coming down from each side from the anterior, these two lines of pinnae gradually draw together until they fuse into one line near the posterior 1 / 3 of the capsule, where they continue on to the posterior (Fig. 12 C). On the posterior, the encircling feather-like pinnae split around a central pinnae stalk with a feather-like apex (Fig. 12 D).	en	Cumming, Royce T., Foley, Evelyn Marie, Hennemann, Frank H., Le Tirant, Stephane, Daawia,, Warikar, Evie Lilly, Yando, Heron, Suhartawan, Bambang, Henze, Katharina, Büscher, Thies H., Bank, Sarah (2025): A deeper look into the diversity of Phyllium leaf insects from Indonesia: seven new species and two unique egg morphologies (Phasmatodea, Phylliidae). ZooKeys 1256: 317-370, DOI: 10.3897/zookeys.1256.162609
371848E37B985710B15D86D1AEBA6D41.taxon	description	Measurements including the extended pinnae [mm]. Length (including operculum expansion): 7.6 – 7.7; maximum width of capsule when viewed from lateral aspect 4.8 – 4.9; length of micropylar plate 3.0 – 3.1.	en	Cumming, Royce T., Foley, Evelyn Marie, Hennemann, Frank H., Le Tirant, Stephane, Daawia,, Warikar, Evie Lilly, Yando, Heron, Suhartawan, Bambang, Henze, Katharina, Büscher, Thies H., Bank, Sarah (2025): A deeper look into the diversity of Phyllium leaf insects from Indonesia: seven new species and two unique egg morphologies (Phasmatodea, Phylliidae). ZooKeys 1256: 317-370, DOI: 10.3897/zookeys.1256.162609
371848E37B985710B15D86D1AEBA6D41.taxon	etymology	Etymology. Patronym; named to honor Frank Hennemann (Germany). At the time of this writing, Frank has named more than 350 phasmid species, reflecting decades of his dedicated work. Specifically within the phylliids, Frank’s 2009 publication (Hennemann et al. 2009) was instrumental in sparking the first author’s passion for leaf insects. At the time of its publication, Frank’s work was a major step towards revising the family and served as the foundation for many subsequent works on the group.	en	Cumming, Royce T., Foley, Evelyn Marie, Hennemann, Frank H., Le Tirant, Stephane, Daawia,, Warikar, Evie Lilly, Yando, Heron, Suhartawan, Bambang, Henze, Katharina, Büscher, Thies H., Bank, Sarah (2025): A deeper look into the diversity of Phyllium leaf insects from Indonesia: seven new species and two unique egg morphologies (Phasmatodea, Phylliidae). ZooKeys 1256: 317-370, DOI: 10.3897/zookeys.1256.162609
371848E37B985710B15D86D1AEBA6D41.taxon	distribution	Distribution. At present known from two provinces on Sulawesi, Indonesia (Fig. 2). The holotype locality of Bungadidi (South Sulawesi Province), and the paratype locality of Tiulapolu [= Tipulu] (Southeast Sulawesi Province).	en	Cumming, Royce T., Foley, Evelyn Marie, Hennemann, Frank H., Le Tirant, Stephane, Daawia,, Warikar, Evie Lilly, Yando, Heron, Suhartawan, Bambang, Henze, Katharina, Büscher, Thies H., Bank, Sarah (2025): A deeper look into the diversity of Phyllium leaf insects from Indonesia: seven new species and two unique egg morphologies (Phasmatodea, Phylliidae). ZooKeys 1256: 317-370, DOI: 10.3897/zookeys.1256.162609
0EA356D5712E588686BB4E39CB33E1EE.taxon	type_taxon	Type species. Phyllium siccifolium (Linnæus, 1758); type locality: ‘ Indies’.	en	Cumming, Royce T., Foley, Evelyn Marie, Hennemann, Frank H., Le Tirant, Stephane, Daawia,, Warikar, Evie Lilly, Yando, Heron, Suhartawan, Bambang, Henze, Katharina, Büscher, Thies H., Bank, Sarah (2025): A deeper look into the diversity of Phyllium leaf insects from Indonesia: seven new species and two unique egg morphologies (Phasmatodea, Phylliidae). ZooKeys 1256: 317-370, DOI: 10.3897/zookeys.1256.162609
AD89A2DC707A58F38F867AD3EC1594B5.taxon	description	Fig. 15	en	Cumming, Royce T., Foley, Evelyn Marie, Hennemann, Frank H., Le Tirant, Stephane, Daawia,, Warikar, Evie Lilly, Yando, Heron, Suhartawan, Bambang, Henze, Katharina, Büscher, Thies H., Bank, Sarah (2025): A deeper look into the diversity of Phyllium leaf insects from Indonesia: seven new species and two unique egg morphologies (Phasmatodea, Phylliidae). ZooKeys 1256: 317-370, DOI: 10.3897/zookeys.1256.162609
AD89A2DC707A58F38F867AD3EC1594B5.taxon	etymology	Etymology. The species epithet illusorium is the singular neuter form of the Latin illusorius, meaning mocking or ironical. While many leaf insects are colloquially described as “ walking leaves, ” Phyllium illusorium sp. nov. offers a subtle twist: it is not merely a leaf in motion, but rather a master of deception, a “ mocking leaf ” that plays with perception itself. Its intricate mimicry not only camouflages the insect among the foliage but also teases the observer, blurring the boundary between flora and fauna. The name celebrates this playful deceit, emphasizing the species’ role as a living illusionist within its arboreal habitat.	en	Cumming, Royce T., Foley, Evelyn Marie, Hennemann, Frank H., Le Tirant, Stephane, Daawia,, Warikar, Evie Lilly, Yando, Heron, Suhartawan, Bambang, Henze, Katharina, Büscher, Thies H., Bank, Sarah (2025): A deeper look into the diversity of Phyllium leaf insects from Indonesia: seven new species and two unique egg morphologies (Phasmatodea, Phylliidae). ZooKeys 1256: 317-370, DOI: 10.3897/zookeys.1256.162609
AD89A2DC707A58F38F867AD3EC1594B5.taxon	distribution	Distribution. At present only known from the type locality; Buton Island, Southeast Sulawesi Province, Indonesia (Fig. 2).	en	Cumming, Royce T., Foley, Evelyn Marie, Hennemann, Frank H., Le Tirant, Stephane, Daawia,, Warikar, Evie Lilly, Yando, Heron, Suhartawan, Bambang, Henze, Katharina, Büscher, Thies H., Bank, Sarah (2025): A deeper look into the diversity of Phyllium leaf insects from Indonesia: seven new species and two unique egg morphologies (Phasmatodea, Phylliidae). ZooKeys 1256: 317-370, DOI: 10.3897/zookeys.1256.162609
3425B9B3481E53218DB06ECFBF22F0CC.taxon	description	Fig. 16	en	Cumming, Royce T., Foley, Evelyn Marie, Hennemann, Frank H., Le Tirant, Stephane, Daawia,, Warikar, Evie Lilly, Yando, Heron, Suhartawan, Bambang, Henze, Katharina, Büscher, Thies H., Bank, Sarah (2025): A deeper look into the diversity of Phyllium leaf insects from Indonesia: seven new species and two unique egg morphologies (Phasmatodea, Phylliidae). ZooKeys 1256: 317-370, DOI: 10.3897/zookeys.1256.162609
3425B9B3481E53218DB06ECFBF22F0CC.taxon	description	Description. Female. Coloration. Coloration description is based upon photographs of the live holotype specimen before it was preserved and dried (which resulted in most of the color being lost). The general coloration was lime green throughout, with the antennae brown, the larger veins of the tegmina dark orange / brown, and a few tan / brown patches present on the profemoral interior lobes and the mesofemoral lobes on the distal ends. Morphology. Head capsule slightly longer than wide, with a vertex that is slightly lumpy, not perfectly smooth (Fig. 16 C). The posteromedial tubercle is present, singularly lobed, but not very prominent (Fig. 16 C). Frontal convexity not very prominent and ending in a blunted point with several short setae across the surface. Compound eyes slightly protruding from the head capsule, not bulbous, taking up slightly <1 / 3 of the head capsule lateral margins (Fig. 16 C). Ocelli absent. Antennal fields slightly wider than the first antennomere width. Antennae consist of nine segments, with segment VIII slightly wider than segments VII or IX. The terminal antennomere is not particularly long, only slightly longer than the preceding segment. Antennomeres I – VII are smooth, and sparsely marked with short setae, the terminal two antennomeres are covered in short, dense setae, giving these segments a fuzzy appearance (Fig. 16 A). Thorax. Pronotum with slightly concave anterior margin and lateral margins that converge only slightly, to the posterior margin which is slightly more than ½ the width of the anterior margin (Fig. 16 A). The pronotum anterior margin has a prominent rim, while the lateral and posterior margins are less prominent. The pronotum surface is relatively smooth, with a prominent sagittal slit in the center and a few furrows lateral to this slit, and a prominent sagittal slit near the anterior margin (Fig. 16 A). Prosternum and the anterior 1 / 3 of the mesosternum are marked by sparse, but prominent nodes, while the rest of the ventral thorax surfaces are relatively smooth. Mesoprescutum approximately as long as wide, lateral rims with five prominent tubercles (Fig. 16 C). Mesoprescutum anterior rim prominently raised into a raised, broad sagittal spine (Fig. 16 B). Mesoprescutum surface smooth except for the slightly raised mesoprescutum sagittal crest which is marked with at least two distinctly raised nodes; areas lateral to the sagittal crest smooth or slightly wrinkled (Fig. 16 C). Mesopleurae begin on the anterior margin and diverge with nearly straight margins gradually (Fig. 16 C). Mesopleurae lateral margins with five or six distinct tubercles which are relatively evenly spaced, and some of these have a node between them (Fig. 16 C). Face of the mesopleura slightly wrinkled, with two notable divots, one on the anterior 1 / 3 and one near the posterior 1 / 3 (Fig. 16 C). Wings. Tegmina long, reaching onto abdominal segment VIII. Tegmina venation; the subcosta (Sc) is the first vein in the forewing, running parallel with the margin for the first ½, and then bending and running towards the margin. The subcosta runs for ~ 1 / 5 of the tegmina length. The radius (R) spans the anterior ½ of the forewing with two subparallel branched veins; the first radius (R 1) branches ~ 1 / 5 of the way through the wing length and terminates ~ 1 / 3 of the way through the tegmina length; the radial sector (Rs) branches ~ 1 / 3 of the way through the wing length and terminates near the distal 2 / 5 of the wing length. There is a continuation of the radius following the prominent Rs branching which continues on as a short but distinct R – M crossvein that connects the two veins. The media (M) is bifurcate with the media anterior (MA) terminating near the distal 1 / 5 of the tegmina and media posterior (MP) terminating near to the apex of the tegmina. There is a weak continuation of the media following the prominent media posterior (MP) branching which continues on as a somewhat long M – Cu crossvein that weakly connects the two veins. The cubitus (Cu) is also bifurcate, branching near the apex of the tegmina into the cubitus anterior (CuA) and cubitus posterior (CuP) which both terminate near the wing apex. The first anal vein (1 A) is simple and fuses with the cubitus ~ ¼ of the way through the tegmina length. Alae vestigial. Abdomen. Abdominal segments II through the anterior 2 / 3 of IV gradually diverging. The posterior 1 / 3 of segment IV through the anterior 2 / 3 of segment VII are slightly and uniformly converging. The posterior 1 / 3 of segment VII is rounded inwards towards segment VIII which like VII starts converging gradually and then rounds inward to segment IX. Segments IX and X have straight, converging margins ending in a broad rounded apex (Fig. 16 F). Genitalia. Subgenital plate starts at the anterior margin of tergum VIII, is broad and triangular, with straight margins, and extends ~ 1 / 2 of the way onto tergum X (Fig. 16 F). Gonapophyses VIII are long and moderately broad, reaching the apex of the abdominal tergum X; gonapophyses IX are obstructed from view (Fig. 16 F). Cerci flat, slightly broadening to the apical 1 / 3, with a slightly granular surface (Fig. 16 F). Legs. Profemoral exterior lobe thin and arching from end to end gently, with a maximum width only ~ 1.5 × the greatest width of the profemoral shaft (Fig. 16 C). Margin of the profemoral exterior lobe smooth or with slight granulation. Profemoral interior lobe ~ 2 × as wide as the greatest width of the profemoral shaft, and marked with six or seven variably sized teeth with looping gaps between them of varying widths (Fig. 16 C). Mesofemoral interior lobe slightly thicker on the distal end, with the greatest width similar in wider to the mesofemoral shaft. The mesofemoral exterior lobes greatest width is also approx. as wide as the mesofemoral shaft width, but the weighting is towards the center, with the proximal and distal ends thin. The mesofemoral exterior lobe has one or two small, distally pointing teeth on the distal 1 / 3 of the lobe. The mesofemoral interior lobe has eight or nine small, distally pointing teeth on the distal ½ of the lobe. Metafemoral interior lobe arcs end to end, with the distal ½ slightly wider than the proximal ½ and marked with 12 or 13 small, serrate teeth on the distal ½ of the lobe. Metafemoral exterior lobe with only two or three very small teeth on the distal 1 / 3 and has a width similar to the metafemoral shaft width. Protibiae exterior simple, lacking a lobe. Protibiae interior lobe spans the entire length of the protibiae and is only slightly wider than the width of the protibiae shaft itself. The lobe is roundly triangular with the widest portion slightly situated on the distal ½. Mesotibiae and metatibiae simple, lacking exterior and interior lobes. Measurements (mm). Holotype, female: body length (including cerci and head, excluding antennae): 79.6, length / width of head: 7.7 / 6.0, antennae: 5.2, pronotum: 5.6, mesonotum: 6.5, length of tegmina: 51.0, greatest width of abdomen: 37.3, profemora: 15.6, mesofemora: 14.7, metafemora: 18.1, protibia: 9.6, mesotibia: 9.8, metatibia: 16.0.	en	Cumming, Royce T., Foley, Evelyn Marie, Hennemann, Frank H., Le Tirant, Stephane, Daawia,, Warikar, Evie Lilly, Yando, Heron, Suhartawan, Bambang, Henze, Katharina, Büscher, Thies H., Bank, Sarah (2025): A deeper look into the diversity of Phyllium leaf insects from Indonesia: seven new species and two unique egg morphologies (Phasmatodea, Phylliidae). ZooKeys 1256: 317-370, DOI: 10.3897/zookeys.1256.162609
3425B9B3481E53218DB06ECFBF22F0CC.taxon	etymology	Etymology. Eponym; named to honor Morgan Brock-Smith (USA), recent wife to the first author. None of the hundreds of hours of research that has been focused on the phylliids over the years by the first author would have been possible without her support and love. Morgan is the stalwart partner that makes the adventures of life exciting and the challenges of life possible to overcome.	en	Cumming, Royce T., Foley, Evelyn Marie, Hennemann, Frank H., Le Tirant, Stephane, Daawia,, Warikar, Evie Lilly, Yando, Heron, Suhartawan, Bambang, Henze, Katharina, Büscher, Thies H., Bank, Sarah (2025): A deeper look into the diversity of Phyllium leaf insects from Indonesia: seven new species and two unique egg morphologies (Phasmatodea, Phylliidae). ZooKeys 1256: 317-370, DOI: 10.3897/zookeys.1256.162609
3425B9B3481E53218DB06ECFBF22F0CC.taxon	distribution	Distribution. At present only known from the type locality of Yapen Island, Papua Province, Indonesia (Fig. 2).	en	Cumming, Royce T., Foley, Evelyn Marie, Hennemann, Frank H., Le Tirant, Stephane, Daawia,, Warikar, Evie Lilly, Yando, Heron, Suhartawan, Bambang, Henze, Katharina, Büscher, Thies H., Bank, Sarah (2025): A deeper look into the diversity of Phyllium leaf insects from Indonesia: seven new species and two unique egg morphologies (Phasmatodea, Phylliidae). ZooKeys 1256: 317-370, DOI: 10.3897/zookeys.1256.162609
95185840172C59A6AE6CCDDDBBC7B753.taxon	description	Fig. 17	en	Cumming, Royce T., Foley, Evelyn Marie, Hennemann, Frank H., Le Tirant, Stephane, Daawia,, Warikar, Evie Lilly, Yando, Heron, Suhartawan, Bambang, Henze, Katharina, Büscher, Thies H., Bank, Sarah (2025): A deeper look into the diversity of Phyllium leaf insects from Indonesia: seven new species and two unique egg morphologies (Phasmatodea, Phylliidae). ZooKeys 1256: 317-370, DOI: 10.3897/zookeys.1256.162609
95185840172C59A6AE6CCDDDBBC7B753.taxon	description	Description. Female. Coloration. Coloration description is based upon the dead, dried holotype specimen (Fig. 17). Unfortunately, the holotype appears to have not been dried very well, and has many dark, discolored areas. Some coloration that can be seen is that the mesocoxae and metacoxae differ in color, with the mesocoxae a similar color to the surrounding tissue, while the metacoxae appear to have a dark spot, similar to Phyllium tobeloense but not quite as crisply edged as in that species. Additionally, the profemoral interior lobes and protibiae interior lobes appear to have some orange striping, a feature which is known from several Phyllium species, but is often variable in its intensity or completely absent in some individuals. Morphology. Head capsule is longer than wide, with a vertex that is relatively smooth except for a singularly pointed posteromedial tubercle which is not very large (Fig. 17 F). Frontal convexity broad and ending in a relatively fine point; surfaces smooth except for a few short setae. Compound eyes only slightly protruding from the head capsule, not overly large, taking up ~ ¼ of the head capsule lateral margins (Fig. 17 F). Ocelli absent. Antennal field slightly wider than the first antennomere. Antennae consist of nine segments, with the terminal segment approx. the same length as the preceding two segments’ lengths combined (Fig. 17 B). Antennomeres I – VII smooth but sparsely marked with small transparent setae, the terminal two antennomeres are covered in dense, short setae giving them a fuzzy appearance (Fig. 17 B). Thorax. Pronotum with slight concave anterior margin and straight lateral margins, which converge to the posterior margin that is ½ the width of the anterior margin (Fig. 17 F). The pronotum surface is relatively smooth, with only a prominent pit in the center and a sagittal furrow on the anterior 1 / 2 (Fig. 17 F). The pronotum has a distinctly formed anterior rim, moderately formed lateral rims, and a weakly formed posterior rim (Fig. 17 F). Prosternum, mesosternum, and metanotum have nodes running along the sagittal plane, with the lateral margins relatively smooth, lacking nodes. Mesoprescutum approx. as long as wide, lateral rims with four tubercles and one or two smaller nodes interspersed (Fig. 17 F). The tegmina membranous attachment areas are large and encroach on the mesoprescutum lateral margins, shortening them and compressing the tubercles somewhat (Fig. 17 F). Mesoprescutum anterior rim prominent and raised into a blunted sagittal spine; the mesoprescutum rim surface is slightly lumpy (Fig. 17 D). Mesoprescutum raised slightly along the sagittal crest, which is marked with three small nodes, while the remainder of the surface is smooth (Fig. 17 F). Mesopleurae begin near the anterior rim, have straight margins, and run uniformly diverging throughout their lengths (Fig. 17 F). Mesopleurae lateral margins with five or six moderately sized tubercles with minimal granulation between a few of the larger ones (Fig. 17 F). Face of the mesopleura smooth or slightly wrinkled, with two notable divots, one on the anterior 1 / 3 and one less prominent one slightly posterior to the middle (Fig. 17 F). Wings. Tegmina long, reaching the anterior margin of abdominal segment IX. Tegmina venation; the subcosta (Sc) is the first vein in the forewing, running parallel with the margin for the first 1 / 2, and then bending and running towards the margin where it terminates 1 / 3 of the way through the wing length. The radius (R) spans the central portion of the forewing with two subparallel branched veins; the first radius (R 1) branches ~ 1 / 5 of the way through the wing length and terminates slightly proximal to the midline, and the radial sector (Rs) branches ~ 1 / 3 of the way through the wing length and terminates near the distal 1 / 3 of the wing length. There is a weak continuation of the radius following the prominent Rs branching which continues on as a short and thin R – M crossvein that does not appear to actually connect the two veins. The media (M) is trifurcate with a media anterior (MA; originating near the middle of the tegmina length and terminating on the distal 1 / 5), first media posterior (MP 1; originating on the distal 1 / 3 of the tegmina length and terminating near the apex), and a second media posterior (MP 2; which is small, originates near the distal 1 / 5 of the tegmina length and terminates near the tegmina apex). The cubitus (Cu) is also bifurcate, branching near the posterior 1 ⁄ 10 of the wing into the cubitus anterior (CuA) and cubitus posterior (CuP) which both terminate at the wing apex. The first anal vein (1 A) is simple and fuses with the cubitus early on, ~ ¼ of the way through the tegmina length. Alae vestigial. Abdomen. Abdominal segments II through the anterior 2 / 3 of IV uniformly diverging. The posterior 1 / 3 of segment IV through segment VII are converging, with the degree of convergence slightly increasing from IV through VII. The posterior 1 / 3 of segment VIII ends in a slightly rounded lobe. Segments IX and X are notably narrower than the previous segments, and have straight, converging margins to the broad rounded apex. Genitalia. Subgenital plate starts at the anterior margin of tergum VIII, is moderately broad, and extends halfway onto tergum X with straight margins ending in a fine point (Fig. 17 G). Gonapophyses VIII are long and moderately broad, reaching the apex of abdominal tergum X; gonapophyses IX are shorter and narrower, hidden below gonapophyses VIII (Fig. 17 G). Cerci flat, somewhat rectangular, with a slightly rough textured surface (Fig. 17 G). Legs. Profemoral exterior lobe smoothly arching from end to end, ~ 3 × wider than the width of the profemoral shaft greatest width (Fig. 17 C). Edge of the profemoral exterior lobe smooth and unadorned (Fig. 17 C). Profemoral interior lobe as wide as the exterior lobe, approximately right angled, and marked with six teeth (four large and two small) arranged in a small-large-large-small-large-large pattern, with looping gaps between the teeth (Fig. 17 C). Mesofemoral exterior lobe roughly a narrow, rounded triangle with the greatest width only slightly wider than the mesofemoral shaft width, and the greatest width situated on the distal 1 / 3 of the mesofemora. Just distal to the mesofemoral exterior lobe greatest width is a singular, small tooth. Mesofemoral interior lobe is approx. the same width as the mesofemoral shaft, and is similar to the exterior lobe with the greatest width on the distal 1 / 3, but the interior lobe is slightly more rounded, not as straight edged as the exterior lobe. Mesofemoral interior lobe distal 1 / 2 is also marked by six or seven small, distally pointing, serrate teeth. Metafemoral interior lobe arcs end to end, with the distal 1 / 2 slightly wider than the proximal 1 / 2 and marked with nine serrated teeth on the distal 1 / 2 of the lobe. Metafemoral exterior lobe is thin and smooth, hugging the metafemoral shaft and lacks dentation. Protibia lacking an exterior lobe (Fig. 17 C). Protibia interior lobe spans the entire length of the protibia and is ~ 2 × the width of the protibia shaft itself. The lobe is roundly triangular with the widest portion on the distal 1 / 2. Mesotibiae and metatibiae simple, lacking exterior and interior lobes. Measurements (mm). Holotype, female: body length (including cerci and head, excluding antennae): 93.5, length / width of head: 10.5 / 8.1, antennae: 5.3, pronotum: 6.7, mesonotum: 7.8, length of tegmina: 62.6, greatest width of abdomen: 37.0, profemora: 18.8, mesofemora: 16.5, metafemora: 22.7, protibia: 11.3, mesotibia: 11.7, metatibia: 17.8.	en	Cumming, Royce T., Foley, Evelyn Marie, Hennemann, Frank H., Le Tirant, Stephane, Daawia,, Warikar, Evie Lilly, Yando, Heron, Suhartawan, Bambang, Henze, Katharina, Büscher, Thies H., Bank, Sarah (2025): A deeper look into the diversity of Phyllium leaf insects from Indonesia: seven new species and two unique egg morphologies (Phasmatodea, Phylliidae). ZooKeys 1256: 317-370, DOI: 10.3897/zookeys.1256.162609
95185840172C59A6AE6CCDDDBBC7B753.taxon	etymology	Etymology. Patronym, named after Pierre-Olivier Ouellet, who donated the specimen to the IMQC.	en	Cumming, Royce T., Foley, Evelyn Marie, Hennemann, Frank H., Le Tirant, Stephane, Daawia,, Warikar, Evie Lilly, Yando, Heron, Suhartawan, Bambang, Henze, Katharina, Büscher, Thies H., Bank, Sarah (2025): A deeper look into the diversity of Phyllium leaf insects from Indonesia: seven new species and two unique egg morphologies (Phasmatodea, Phylliidae). ZooKeys 1256: 317-370, DOI: 10.3897/zookeys.1256.162609
95185840172C59A6AE6CCDDDBBC7B753.taxon	distribution	Distribution. At present only known from Obi Island, Indonesia (Fig. 2).	en	Cumming, Royce T., Foley, Evelyn Marie, Hennemann, Frank H., Le Tirant, Stephane, Daawia,, Warikar, Evie Lilly, Yando, Heron, Suhartawan, Bambang, Henze, Katharina, Büscher, Thies H., Bank, Sarah (2025): A deeper look into the diversity of Phyllium leaf insects from Indonesia: seven new species and two unique egg morphologies (Phasmatodea, Phylliidae). ZooKeys 1256: 317-370, DOI: 10.3897/zookeys.1256.162609
