taxonID	type	description	language	source
6414D960FFBFFF88FED755BB470AFCB7.taxon	materials_examined	Material examined Holotype: Leizou Bay, Guandong Province, 20 ◦ 10.20 ′ N, 110 ◦ 56.40 ′ E, 10 m, 15 April to 15 May 2007, male (sl 5.3 mm; No. 6), IOCAS-MBM 119736. Paratype: same data as holotype, one male (sl 7.1 mm; No. 7), CBM-ZC 10521. Description Shield (Figure 1 A) nearly subquadrate, almost as long as broad or slightly broader than long; anterior margin between rostrum and lateral projections slightly concave, with 3 – 6 tiny tubercles near each base of lateral projection; anterolateral margins sloping, with small spines at lateral angle; posterior margin truncate; dorsal surface with some transverse spinulose ridges laterally. Rostrum broadly rounded or terminally faintly bilobed; lateral projections each with small submarginal spine. Anterior part of branchiostegite with row of about 10 small spines on dorsal margin, no spine at anterior angle; lateral surface almost smooth, glabrous (Figure 1 B). Ocular peduncles (Figure 1 A) about 0.6 times as long as shield, moderately stout; cornea slightly inflated, corneal width about 0.3 of length of ocular peduncle. Ocular acicles with straight inner margins, slightly convex anterolaterally, with 2 or 3 prominent spines and 7 – 9 spinules decreasing in size laterally, not extending to entire length of terminal margin. Intercalary rostriform process reaching nearly to distal end of ocular acicles, slightly broadened basally, tapering to slender acute tip. Antennular peduncles (Figure 1 A) overreaching distal corneal margins by 0.8 – 0.9 length of ultimate segment and extending as far as antennal peduncles. Ultimate segment with some setae on dorsal surface and tufts of setae distally on both ultimate and penultimate segments; basal segment with row of minute spinules on distolateral margin. Antennal peduncles (Figure 1 A, B) overreaching distal corneal margins by 0.8 length of fifth segment. Fifth segment with two rows of moderately long to long stiff setae ventrally; fourth and third segments unarmed; second segment with strong spine at dorsolateral distal angle and small but distinct spine at dorsomesial distal angle, ventrodistal margin with row of spinules; first segment with row of spinules on distal margin laterally. Antennal acicle falling short of distolateral margin of fourth segment, terminating in simple or bifid spine, bearing seven or eight strong accessory spines on mesial margin and occasionally with one spine on lateral margin, and with some tufts of short stiff setae on both lateral and mesial margins. Antennal flagellum (Figure 1 C) less than twice of shield length, articles with paired long, setulose setae ventrally and few additional short setae, forming cast net-like structure. Third maxilliped (Figure 1 D) moderately slender. Carpus unarmed on dorsodistal margin. Ischium-basis fused segment with five strong, corneous-tipped spines (three on ischium and two on basis) on mesial surface (Figure 1 E). Exopod reaching distal margin of carpus. Left cheliped (Figure 2 A – E) large but not particularly elongate in male; not particularly setose. Chela strongly compressed, about 1.5 times longer than wide. Dactylus distinctly longer than palm, somewhat curved; three rows of spines on upper margin, spines in middle row distinctly larger than those in lateral or mesial rows; outer surface with scattered spines or tubercles increasing in size proximally and ventrally, shallowly sulcate along midline; cutting edge with single row of blunt but distinct calcareous teeth; inner surface with longitudinal row of small spines accompanied by distinct groove along midline and one additional row of tubercles adjacent to cutting edge. Palm slightly shorter than carpus; upper surface with small spines arranged in three irregular longitudinal rows; outer surface generally convex, but shallowly depressed at base of fixed finger, having covering of small spines or spinules increasing in size dorsally, but without pronounced crest or longitudinal row (s) of spines; proximal margin distinctly delimited by row of small spines; lower margin faintly sinuous in outer view (including fixed finger), with double row of small spines extending to tip of fixed finger and continuous to proximal margin; inner surface with scattered, numerous tiny tubercles. Fixed finger very slightly deflexed, not curved outward; outer surface not particularly elevated along midline, with scattered small spines increasing in size toward cutting edge, but without distinct row of tubercles or ridge; inner surface with scattered small tubercles proximally; cutting edge with row of blunt but conspicuous calcareous teeth (middle tooth strongest), subterminal concavity receiving tip of dactylus. Carpus wider than long, with single row of moderately large spines with curved tips on upper margin; outer surface convex, with scattered numerous small spines or spine-like tubercles decreasing in size ventrally, dorsodistal margin finely spinulose; lower angle with prominent spine; mesial surface with scattered small spines or spinules and median row of small spines in proximal half; ventral surface with few tiny tubercles. Merus about as long as high; dorsal surface narrowing proximally and forming blunt crest in proximal half, bearing small spines over entire length; tufts of long stiff plumose setae present on dorsal surface; dorsodistal margin with row of sharp spines; lateral surface with scattered tiny tubercles, distolateral margin minutely spinulose, ventrolateral margin with row of small spines; mesial surface nearly smooth, ventromesial margin with row of small spines and one prominent spine at proximal angle; ventral surface with scattered minute tubercles and tufts of short setae. Ischium with two low protuberances on ventral surface and row of spinules on ventrolateral distal margin. Right cheliped (Figure 3 A – D) moderately stout, setose; chela 1.8 times longer than wide, tips of fingers crossing. Dactylus only slightly curved, about 2.9 times longer than palm; dorsal surface with three irregular longitudinal rows of small spines and accompanying tufts of long stiff setae; mesial surface with median row of small spines extending beyond midlength and accompanying tufts setae and prominent groove inferior to midline in proximal half; ventral surface nearly smooth, with tufts of moderately long stiff setae laterally; cutting edge with row of small, acute, calcareous teeth over entire length. Palm about half length of carpus; dorsal surface with tufts of stiff setae and four irregular longitudinal rows of small spines, of them two rows extending onto fixed finger (spines strongest on fixed finger), lateral margin delimited by row of spinules extending onto fixed finger, dorsomesial face with numerous scattered spines; mesial surface with some small tubercles; ventral surface very slightly convex, with three rows of tiny tubercles laterally, extending onto fixed finger. Carpus with dorsolateral and dorsomesial rows of spines (spines of dorsomesial row stronger than those of dorsolateral row), dorsal surface with few spinules proximally; lateral surface with scattered tiny tubercles or spinules, distolateral margin with row of spinules; mesial surface with single row of spinules adjacent to dorsal margin and median row of tiny tubercles, distomesial margin with row of tiny tubercles extending ventrally. Merus with row of spines decreasing in size proximally and tufts of setae on dorsal margin, dorsodistal margin with six moderately small spines; lateral surface with scattered simple or bifid tiny tubercles and tufts of short setae, ventrolateral margin only delimited in distal part, with small spines or spinules; mesial surface nearly smooth, with tiny tubercles adjacent to ventromesial margin, ventromesial margin with row of spinules; ventral surface with scattered tiny tubercles and tufts of setae. Ischium unarmed on ventromesial margin; ventrolateral distal margin with row of tiny tubercles; some small protuberances present on ventral surface. Ambulatory legs (Figures 4 A, B, 5 A – C) moderately long and stout; left third having stronger armature than other legs. Dactyli about 1.5 – 1.6 times as long as propodi and 8.5 – 9.0 times longer than wide; dorsal surfaces each with row of sparse short stiff setae and row of spinules laterally (in proximal 0.3 – 0.4 in second, proximal 0.2 in right third, proximal half in left third) and with row of dense stiff setae mesially; lateral surfaces each with shallow median sulcus running over entire length and median row of short sparse setae; mesial surfaces each with short median sulcus and median row of setae proximally, otherwise nearly flat, flanked by rows of moderately long stiff setae; ventral surfaces rounded, each with row of short stiff setae, unarmed (second and right third) or armed with some minute tubercles proximally (left third). Propodi slightly curved, distinctly longer than carpi; dorsal surfaces each with two rows of small spines and tufts of short stiff setae; lateral surfaces each with one row of small spines adjacent to dorsal margin (second and right third) or with two rows of minute spinules dorsally and one row of tiny tubercles ventrally in addition to dorsal row of small spines (left third); mesial surfaces each with two rows of small spinules (in third, ventral spinules reduced to minute tubercles); ventral margins smooth (right second and right third) or with row of small spines (left third) or spinules (left second). Carpi each with row of small spines increasing in size distally and tufts of setae on dorsal margin; lateral surfaces each with median row of minute tubercles accompanied by tufts of short setae and additional row of small tubercles (second) or small low protuberances ventral to median row (third), and vertical short row of spinules along distal margin; mesial surfaces each with median row of tufts of short setae. Meri with dorsal and ventral tufts of long setae; dorsal margins each with one or two rows of small spines or spinules, distalmost spine largest; lateral surfaces with spinules dorsally and ventrally, otherwise smooth; mesial surfaces each with row of long plumose setae adjacent to dorsal margin; ventral surfaces each with two rows of spinules (spinules on lateral row often reduced to tiny protuberances). Ischia each with one or few rounded protuberances on ventral surface. Coxae of chelipeds and ambulatory legs with large blister-like protuberances. Fourth pereopods chelate (Figure 1 F). Propodi each with small spine on dorsodistal margin; rasp consisting of more than 10 rows of corneous scales. Carpi each with small dorsodistal spine. Anterior lobe of sixth thoracic sternite (of third pereopods) (Figure 1 G) distinctly bilobed, each lobule with 1 spinule or tiny tubercle anterolaterally. Telson (Figure 1 H) with small median cleft, terminal margin slightly oblique on left, nearly transverse on right, both with small, unequal spines; lateral margin of left posterior lobe minutely denticulate, with some submarginal spines (posteriormost one or two spines prominent). Coloration Based on formalin preserved specimen: Shield generally light brown. Ocular peduncles and antennular peduncles pale brown; corneas black. Antenna with second segment and antennal acicle brown, fifth segment and flagellum white. Chelipeds and ambulatory legs generally light brick red. Distribution Known only from the type locality, Leizou Bay, Guandong Province; 10 m. Remarks The present new species is referred to the Diogenes edwardsii (De Haan, 1849) species group by the intercalary rostriform process being marginally smooth and not reaching beyond the ocular acicles, the antennal peduncle distinctly overreaching the distal corneal margin, and the antennal flagellum having long paired setae on the distal margin of each article ventrally (Asakura and Tachikawa 2010). Diogenes acanthochela sp. nov. is readily distinguished from other species in this group by a combination of the following characters: left chela with spinose upper margin and outer surface, but without distinct crest or row (s) of spines; merus of left cheliped with one prominent spine on ventromesial margin proximally; cutting edges of fingers of right chela armed each with row of small acute teeth over entire length; dorsal margin of dactylus of left third pereopod with row of spinules; propodi and carpi of ambulatory legs serrated with row (s) of spines, propodus of left third with additional rows of spinules on lateral surface. Diogenes spinulimanus Miers, 1880 may be similar to D. acanthochela sp. nov. in the armature of the left chela (Miers 1880), but the former differs from the latter in the non-denticulate ocular acicles and the smooth dactyli of the ambulatory legs. Etymology The specific name is a combination of the Greek words “ acanthus ” (= spiny) and “ chele ” (= claw), in reference to the strongly spinose left chela of this new species.	en	Komai, Tomoyuki, Liang, Jingzhen, Yang, Tingbao (2012): Records of four species of the shallow water hermit crab genus Diogenes (Crustacea: Decapoda: Anomura: Diogenidae) from southern China, with description of a new species. Journal of Natural History 46 (19 - 20): 1219-1248, DOI: 10.1080/00222933.2011.654279, URL: http://dx.doi.org/10.1080/00222933.2011.654279
6414D960FFB7FF90FEDE552F4235FC46.taxon	description	(Figures 6 – 9)	en	Komai, Tomoyuki, Liang, Jingzhen, Yang, Tingbao (2012): Records of four species of the shallow water hermit crab genus Diogenes (Crustacea: Decapoda: Anomura: Diogenidae) from southern China, with description of a new species. Journal of Natural History 46 (19 - 20): 1219-1248, DOI: 10.1080/00222933.2011.654279, URL: http://dx.doi.org/10.1080/00222933.2011.654279
6414D960FFB7FF90FEDE552F4235FC46.taxon	materials_examined	Material examined Leizhou Bay, Guandong Province, 20 ◦ 10.20 ′ N, 110 ◦ 56.40 ′ E, 10 m, 15 April to 15 May 2007, trawl, 1 male (sl 4.1 mm; No. 8, figured), IOCAS-MBM 119737; Hailing Bay, Guandong Province, 21 ◦ 34.48 ′ N, 111 ◦ 43.36 ′ E, 10 m, July to August 2007, trawl, 1 male (sl 4.3 mm; No. 9), CBM-ZC 10522. Redescription Shield (Figure 6 A) slightly longer than broad, almost subquadrate; anterior margin between rostrum and lateral projections slightly concave, with four to six tiny tubercles near each base of lateral projection; anterolateral margins sloping, with few tiny tubercles near lateral angle; posterior margin truncate; dorsal surface with several transverse spinulose ridges laterally. Rostrum broadly rounded; lateral projections each with moderately strong submarginal spine. Anterior part of branchiostegite with nine to 10 small spines on dorsal margin, anterior angle with one small spine; lateral surface with scattered tufts of short to moderately long setae (Figure 6 B). Ocular peduncles (Figure 6 A) about 0.6 times as long as shield, moderately stout; cornea not dilated, corneal width about 0.3 of peduncular length; ocular acicles with straight inner margins, nearly straight anterolaterally, four or five small spines decreasing in size laterally, not extending to entire length of terminal margin. Intercalary rostriform process reaching nearly to distal end of ocular acicles, broad basally, tapering to slender acute tip. Antennular peduncles (Figure 6 A) overreaching distal corneal margins by about 0.8 length of ultimate segment and reaching or slightly overreaching antennal peduncles. Ultimate segment with some setae on dorsal surface and tufts distally on both ultimate and penultimate segments; basal segment unarmed. Antennal peduncles (Figure 6 A, B) overreaching distal corneal margins by about 0.7 of fifth segment. Fifth segment with rows of numerous moderately long to long stiff setae ventrolaterally; fourth segment with few scattered tufts of short setae, unarmed; second segment with strong spine at dorsolateral distal angle and small but prominent spine at dorsomesial distal angle, ventrodistal margin with row of spinules; first segment with row of spinules on distal margin laterally. Antennal acicle reaching to distolateral margin of fourth segment of peduncle, terminating in simple or bifid spine, bearing five or six strong accessory spines on mesial margin, and with some tufts of short stiff setae on both lateral and mesial margins. Antennal flagellum (Figure 6 C) more than twice of shield length, articles with paired long, setulose setae and few additional short setae, forming cast net-like structure. Third maxilliped (Figure 6 D) moderately slender. Carpus with minute spine on dorsodistal margin. Ischium-basis fused segment with three moderately strong, corneous-tipped spines (two on ischium and one on basis) on mesial surface (Figure 6 E). Exopod overreaching distal margin of carpus. Left cheliped (Figures 7 A – E; 8 A – C) elongate. Dactylus 1.1 – 1.2 times longer than palm; double row of moderately small to moderately large tubercles on upper margin (those tubercles sometimes spine-like); outer surface weakly elevated along midline, with short row of tubercles proximally adjacent to upper margin and scattered small tubercles proximally; cutting edge with single row of small rounded calcareous teeth; inner surface nearly smooth with short row of small tubercles in midline proximally. Palm distinctly longer than wide, and shorter than carpus; outer surface with two rows of tubercles, upper row extending from proximal to or beyond midlength of palm, middle row extending from proximal to at most midlength of palm, remaining outer surface with numerous small blunt tubercles; proximo-lower margin not markedly carinate or crested; lower margin nearly straight in outer view, with row of low, rounded tubercles; inner surface with numerous low, sometimes squami-form tubercles increasing in size ventrally. Fixed finger scarcely or slightly deflexed, somewhat curved outward; outer surface with row of tubercles on elevated midline, otherwise nearly smooth; inner surface nearly smooth; cutting edge with row of rounded calcareous teeth, strongest at proximal 0.3 and subterminal. Carpus about twice longer than wide, with double row of moderately small spines on upper margin; outer surface convex, with numerous small, simple to multifid tubercles, dorsodistal margin with row of spinules; lateral surface with numerous low, rounded tubercles (simple to multifid); ventral surface with small tubercles. Merus slightly longer than high; dorsal surface rounded, with tufts of stiff setae and short transverse rows of small spinules or spinulose tubercles, distalmost row adjacent to distal margin longest, extending onto lateral and mesial faces; lateral surface with scattered small, simple to multifid tubercles or granules increasing in size ventrally, ventrolateral margin with row of small rounded tubercles; mesial surface with distally spinulose tubercles dorsally and ventrally (dorsal tubercles with tufts of short setae), ventromesial margin weakly tuberculate; ventral surface with small, low tubercles and tufts of short setae. Ischium with row of small granules on laterodistal margin ventrally and proximal margin; ventromesial margin tuberculate. Right cheliped (Figure 8 D – G) moderately stout. Dactylus strongly curved, about 2.6 times longer than palm; dorsal margin with short row of small spines proximal on midline and one additional short row of spines just lateral to median row; two rows of tufts of stiff setae extending entire length on dorsal surface; mesial surface with one short row of moderately small spines on midline proximally and single row of tufts of setae adjacent to ventral margin; cutting edge without conspicuous teeth. Palm about 0.6 times as long as carpus; dorsal surface with tufts of long stiff setae and four or five irregular longitudinal rows of small tubercles or spines (becoming larger and sharper distally), lateral rows extending onto fixed finger, dorsomesial margin with row of moderately small spines, dorsolateral surface sloping ventrally; mesial surface with some small low protuberances; ventral surface very slightly convex, with single row of small spines laterally, extending onto fixed finger. Carpus with row of spines on dorsal margin and row of tubercles on dorsolateral surface, dorsodistal margin with row of tiny spinulose tubercles; mesial surface with row of low, distally spinulose protuberances adjacent to dorsal margin, otherwise with few very low protuberances, distomesial margin spinulose. Merus with short transverse spinulose ridges in distal half and tufts of long setae over entire length of dorsal margin, dorsodistal margin spinose; lateral surface with scattered tiny tubercles or low protuberances and tufts of short setae, ventrolateral margin with two small spines or tubercles subdistally; mesial surface nearly smooth, with small tubercles adjacent to ventral margin, ventromesial margin with row of tiny to small tubercles; ventral surface with small, low tubercles and tufts of setae. Ischium with row of spinulose tubercles on ventromesial margin. Ambulatory legs (Figure 9 A – D) similar, but third pair slightly longer than second pair. Dactyli 1.2 – 1.3 times as long as propodi and 8.2 – 8.9 times longer than wide; dorsal margins unarmed, each with row of moderately short stiff dense setae; lateral surfaces each with distinct median sulcus over entire length; mesial surfaces each with short median sulcus and median row of setae proximally, otherwise nearly flat, and with dorsal and ventral row of long stiff dense setae; ventral surfaces each with row of short sparse setae. Propodi slightly curved, distinctly longer than carpi; dorsal surfaces each with irregular two rows of spinules and two rows of tufts of stiff setae (second) or nearly smooth but with two irregular rows of stiff setae (third); lateral surfaces each with row of tufts of setae adjacent to dorsal margin and sparse row of tufts of very short setae ventrally; mesial surfaces each with row of tufts of moderately short setae on either side of midline; ventral surfaces with sparse setae. Carpi each with two rows of small spinules and tufts of setae on dorsal margin (spines sometimes fairly reduced on third); lateral surfaces each with row of tufts of short setae dorsally; mesial surfaces each with longitudinal row of tufts of setae dorsal to midline. Meri with dorsal and ventral tufts of setae; dorsal margins each with row of spinules (second) or row of low protuberances (third); lateral surfaces with row of tufts of short setae adjacent to ventral margin; ventral margins each without conspicuous armature. Ischia unarmed, with tufts of moderately long setae. Fourth pereopods chelate (Figure 6 F). Propodi each with dorsodistal margin not particularly produced, unarmed; rasp consisting of five or six rows of corneous scales. Carpi without dorsodistal spine. Anterior lobe of sixth thoracic sternite (of third pereopods) (Figure 6 G) distinctly bilobed, each lobule with spinule at anterolateral angle. Telson (Figure 6 H) with small median cleft, terminal margins of both lobes nearly transverse, with long, curved spines interspersed with minute denticles (Figure 6 I), extending onto lateral margin of left only. Coloration Based on formalin preserved specimen: generally tan; non conspicuous markings on ocular peduncle, antennular peduncle and antenna; cornea black. Left cheliped mottled with brown. Ambulatory legs with brownish markings on middle parts of propodi and meri; carpi with longitudinal brown spots on lateral surfaces. Distribution Recorded from the Chinese coast from Bohai to Hainan Island; intertidal to 30 m (Wang 2008). Remarks	en	Komai, Tomoyuki, Liang, Jingzhen, Yang, Tingbao (2012): Records of four species of the shallow water hermit crab genus Diogenes (Crustacea: Decapoda: Anomura: Diogenidae) from southern China, with description of a new species. Journal of Natural History 46 (19 - 20): 1219-1248, DOI: 10.1080/00222933.2011.654279, URL: http://dx.doi.org/10.1080/00222933.2011.654279
6414D960FFAFFF9FFED155BF4137FEC5.taxon	description	(Figures 10 – 14)	en	Komai, Tomoyuki, Liang, Jingzhen, Yang, Tingbao (2012): Records of four species of the shallow water hermit crab genus Diogenes (Crustacea: Decapoda: Anomura: Diogenidae) from southern China, with description of a new species. Journal of Natural History 46 (19 - 20): 1219-1248, DOI: 10.1080/00222933.2011.654279, URL: http://dx.doi.org/10.1080/00222933.2011.654279
6414D960FFAFFF9FFED155BF4137FEC5.taxon	materials_examined	Material examined Dapeng Bay, near Hong Kong, Guandong Province, 22 ◦ 33.59 ′ N, 114 ◦ 25.09 ′ E, 18 m, mud, 15 April to 15 May 2007, trawl, 1 male (sl 6.8 mm; No. 1), CBM-ZC 10518; same data, 1 male (sl 7.4 mm; No. 2, figured), IOCAS-MBM 119738; Hailing Bay, Guandong Province, 21 ◦ 34.48 ′ N, 111 ◦ 43.36 ′ E, 10 m, 7 November 2007, 1 male (sl 4.9 mm; No. 3), IOCAS-MBM 119739; same data, 1 male (sl 5.3 mm; No. 4), CBM-ZC 10519; same data (sl 4.9 mm; No. 5), CBM-ZC 10520. Redescription Shield (Figure 10 A) slightly longer than broad, almost subquadrate; anterior margin between rostrum and lateral projections slightly concave, with four to six tiny tubercles near each base of lateral projection; anterolateral margins sloping, with few tiny tubercles or spines near or at lateral angle; posterior margin truncate; dorsal surface with some transverse spinulose ridges laterally. Rostrum broadly rounded; lateral projections each with small, simple or bifid submarginal spine. Anterior part of branchiostegite with row of about 10 small spines on dorsal margin, anterior angle with one small spine; lateral surface with scattered small tubercles bearing tuft of setae (Figure 10 B). Ocular peduncles (Figure 10 A) about 0.6 times as long as shield, moderately stout; cornea not dilated, corneal width about 0.3 of peduncular length; ocular acicles with straight or slightly concave inner margins, nearly straight or slightly convex anterolaterally, with about 10 small spines decreasing in size laterally, not extending to entire length of terminal margin. Intercalary rostriform process reaching nearly to distal end of ocular acicles, slightly broadened basally, tapering to slender acute tip. Antennular peduncles (Figure 10 A) overreaching distal corneal margins by 0.5 – 0.6 length of ultimate segment and extending as far as antennal peduncles. Ultimate segment with some setae on dorsal surface and tufts distally on both ultimate and penultimate segments; basal segment with row of minute spinules on distolateral margin. Antennal peduncles (Figure 10 A, B) overreaching distal corneal margins by 0.8 – 0.9 length of fifth segment. Fifth segment with rows of moderately long to long stiff setae ventrolaterally; fourth and third segments unarmed; second segment with strong spine at dorsolateral distal angle and small but prominent spine at dorsomesial distal angle, ventrodistal margin with row of spinules; first segment with row of spinules on distal margin laterally. Antennal acicle slightly falling short of distolateral margin of fourth segment, terminating in simple or bifid spine, bearing six or seven strong accessory spines on mesial margin, and with some tufts of short stiff setae on both lateral and mesial margins. Antennal flagellum (Figure 10 C) more than twice shield length, articles with paired long, setulose setae and few additional short setae, forming cast net-like structure. Third maxilliped (Figure 10 D) moderately slender. Carpus unarmed on dorsodistal margin. Ischium-basis fused segment with five strong, corneous-tipped spines (three on ischium and two on basis) on mesial surface (Figure 10 E). Exopod slightly overreaching distal margin of merus. Left cheliped (Figure 11 A – F) large but not particularly elongate in male, shape and armature rather variable; not particularly setose. Dactylus distinctly longer than palm; three irregular longitudinal rows of moderately small, spine-like tubercles on upper margin; outer surface very slightly concave, with scattered tiny tubercles or granules in lower part; cutting edge with two irregular rows of small tubercles; inner surface slightly elevated along midline with row of small tubercles, upper half nearly smooth, and lower half with scattered tiny tubercles. Palm slightly shorter than carpus; upper surface with three irregular rows of small, spine-like tubercles; outer surface with two rows of small tubercles, upper row extending from proximal to or beyond midlength of palm, middle row extending from proximal to at most midlength of palm, latter forming weak crest extending from carpal articulation, remaining outer surface with numerous microscopically small tubercles or granules (greater part apparently smooth to naked eye); proximal margin carinate; lower margin slightly sinuous in outer view (including fixed finger), with row of small rounded tubercles extending to base of fixed finger and continuous to proximal margin; inner surface with scattered, numerous tiny tubercles. Fixed finger slightly deflexed, not curved outward; outer surface elevated along midline, with scattered small granules, but without distinct row of tubercles or ridge; inner surface with scattered small tubercles proximally; cutting edge with row of rounded calcareous teeth in proximal half and two irregular rows of small tubercles in distal half. Carpus about as long as wide, with single row of moderately large spines on upper margin; outer surface convex, with scattered numerous small spines or spine-like tubercles, dorsodistal margin finely denticulate; mesial surface with row of small spines adjacent to upper margin and with scattered spinules; ventral surface also with scattered spine-like tubercles. Merus slightly longer than high; dorsal surface rounded, with tufts of stiff setae and short irregular transverse rows of small spines, distalmost row adjacent to distal margin longest, extending onto lateral and mesial surfaces; dorsodistal margin with row of spines; lateral surface with scattered tiny, simple or bifid tubercles or granules increasing in size ventrally, distolateral margin finely denticulate, ventrolateral margin with row of spines decreasing in size proximally; mesial surface nearly smooth except for dorsal part bearing tiny spines or spinules and longitudinal row of spines adjacent to ventromesial margin, distomesial margin with row of spines increasing in size ventrally and extending onto ventromesial margin; spines on ventromesial margin large; ventral surface with scattered minute spinules. Ischium with row of small tubercles on laterodistal margin ventrally and proximal margin; ventromesial margin with row of small tubercles. Right cheliped (Figure 12 A – D) moderately stout, setose. Dactylus fairly arched, about 2.6 times longer than palm; dorsal surface with two irregular longitudinal rows of small spines and accompanying tufts of stiff setae extending beyond midlength; mesial surface with median row of small tubercles and accompanying tufts of setae and prominent groove inferior to midline in proximal half; ventral surface with row of tiny tubercles and tufts of stiff setae on midline and with few small tubercles proximomesially; cutting edge with row of low, tiny tubercles. Palm about twice length of carpus; dorsal surface with tufts of stiff setae and four or five irregular longitudinal rows of small tubercles or spines (spines strongest around base of fixed finger), lateral rows extending onto fixed finger, dorsomesial face with numerous spines; mesial surface with some small tubercles dorsally, otherwise nearly smooth; ventral surface very slightly convex, with two rows of small spines laterally, extending onto fixed finger. Carpus with row of spines on dorsal margin and on dorsolateral surface, dorsodistal margin spinose; mesial surface with row of small spines or spine-like tubercles adjacent to dorsal margin and median row of small spinulose tubercles, distomesial margin spinulose. Merus with short transverse spinulose ridges or row of spines and tufts of long setae over entire length of dorsal margin, dorsodistal margin spinose; lateral surface with scattered tiny spinulose tubercles and tufts of short setae, ventrolateral margin only delimited in distal part, with two small spines or tubercles subdistally; mesial surface nearly smooth, with small spine-like tubercles adjacent to ventral margin, ventromesial margin with row of tiny to small spines; ventral surface with tiny spines and tufts of setae. Ischium with row of small tubercles on ventromesial margin; some small tubercles present on ventral surface. Ambulatory legs (Figures 13 A, B, 14 A, B) similar, but third pair slightly longer than second pair. Dactyli about 1.5 times as long as propodi and 7.5 – 8.0 times longer than wide; dorsal margins unarmed (second and right third) or armed with row of minute spinules (left third) (Figure 14 C), and each with row of moderately short stiff dense setae; lateral surfaces each with shallow median sulcus running over entire length; mesial surfaces each with short median sulcus and median row of setae proximally, otherwise nearly flat, and with dorsal and ventral rows of long stiff dense setae; ventral surfaces each with row of short sparse setae. Propodi slightly curved, distinctly longer than carpi; dorsal surfaces each with two rows of spinules and tufts of short stiff setae; lateral surface each with one row of small spinules and tufts of short stiff setae dorsally, otherwise nearly smooth; mesial surfaces each with row of small spinules dorsally (second) or unarmed (third), and with few tufts of short setae; ventral margins nearly smooth (second) or with row of small, low protuberances (third). Carpi each with row of small spines and tufts of setae on dorsal margin (spines smaller in third than in second; distalmost spine strongest); lateral surfaces each with median row of minute tubercles (second) or row of tiny, low protuberances (third); mesial surfaces each with one or more additional spine near base of dorsodistal spine (second) or unarmed (third). Meri with dorsal and ventral tufts of setae; dorsal margins each with row of spinules; lateral surfaces with minute spinulose tubercles dorsally and ventrally; mesial surfaces each with row of tufts of moderately long setae adjacent to dorsal margin; ventral surfaces each with two rows of spinules and row of tufts of setae. Ischia with rounded tubercles on ventral surfaces, dorsal margins bearing small spine (second) or unarmed (third). Fourth pereopods chelate (Figure 10 F). Dactyli reaching distal margins of propodi. Propodi each with distinctly produced dorsodistal margin sometimes bearing one small spine; rasp consisting of about 10 rows of corneous scales. Carpi each with or without dorsodistal spine. Anterior lobe of sixth thoracic sternite (of third pereopods) (Figure 10 G) distinctly bilobed, each lobe with two small spines laterally and anteriorly. Telson (Figure 10 H) with small median cleft, terminal margin oblique on left, nearly transverse on right, both with row of spinules or small spines (row of spinules extending onto left lateral margin), left with three prominent spines at lateral angle. Coloration Based on formalin preserved specimen: generally tan; non conspicuous markings on ocular peduncle, antennular peduncle and antenna; cornea black. Left chela with sparse brown dots on outer surface; carpus and merus with few brown dots. Ambulatory legs with trace of browning markings on middle parts of propodi and carpi. Distribution Known only from Chinese coast from Bohai to Zhejiang; intertidal to 30 m. Remarks	en	Komai, Tomoyuki, Liang, Jingzhen, Yang, Tingbao (2012): Records of four species of the shallow water hermit crab genus Diogenes (Crustacea: Decapoda: Anomura: Diogenidae) from southern China, with description of a new species. Journal of Natural History 46 (19 - 20): 1219-1248, DOI: 10.1080/00222933.2011.654279, URL: http://dx.doi.org/10.1080/00222933.2011.654279
6414D960FFA0FF9FFE4D573D4045FA11.taxon	materials_examined	Material examined Haling Bay, Guandong Province, 20 ◦ 10.20 ′ N, 110 ◦ 56.40 ′ E, 10 m, sand, November 2007, one male (sl 5.5 mm; No. 10), CBM-ZC 10523; Shantou Bay, 23 ◦ 26.28 ′ N, 116 ◦ 53.41 ′ E, 10 m, mud, November 2007, one female (sl 5.5 mm; No. 11), CBM- ZC 10524; Leizhou Bay, Guandong Province, 21 ◦ 02.48 ′ N, 110 ◦ 39.54 ′ E, 10 m, sand, December 2006, one male (sl 5.7 mm), IOCAS-MBM 119740; Shantou Bay, 23 ◦ 21.26 ′ N, 116 ◦ 50.25 ′ E, 10 m, mud, one male (sl 5.3 mm), IOCAS-MBM 119741. Distribution Gulf of Aden, India, Sri Lanka, Thailand, Torres Straight, Australia, China, Taiwan; intertidal to 36 m. Remarks	en	Komai, Tomoyuki, Liang, Jingzhen, Yang, Tingbao (2012): Records of four species of the shallow water hermit crab genus Diogenes (Crustacea: Decapoda: Anomura: Diogenidae) from southern China, with description of a new species. Journal of Natural History 46 (19 - 20): 1219-1248, DOI: 10.1080/00222933.2011.654279, URL: http://dx.doi.org/10.1080/00222933.2011.654279
