identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
6B7BE270BC1DAA23FF02FE72FB16FD04.text	6B7BE270BC1DAA23FF02FE72FB16FD04.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Radix dgebuadzei Aksenova, Vinarski, Bolotov & Kondakov 2019	<div><p>Radix dgebuadzei Aksenova, Vinarski, Bolotov &amp; Kondakov sp. nov.</p> <p>Figs. 1–4, Table 2</p> <p>https://zoobank.org/ urn:lsid:zoobank.org:pub: 0C283AE4-A94E-456D-9FD4-466A97D0BABB</p> <p>Type series. The holotype and paratype are deposited in ZIN (accession numbers: 1-515/2019 and 2- 515/2019, respectively), and 3 paratypes are deposited in RMBH (accession numbers: MLym-686/2, MLym- 686/3, MLym-686b). The reference DNA sequence numbers of the paratypes MLym-686/2 and MLym- 686/3 are presented in Table 2.</p> <p>*ZIN; **RMBH; n/a – not available</p> <p>Type locality. CHINA: Qinling Region, Gansu Province, Sun Jia He River (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.222&amp;materialsCitation.latitude=33.6673" title="Search Plazi for locations around (long 106.222/lat 33.6673)">Yangtze River Basin</a> ), in the vicinity of the <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.222&amp;materialsCitation.latitude=33.6673" title="Search Plazi for locations around (long 106.222/lat 33.6673)">Shangtan village</a>, lakelet on the shore, 33.6673°N, 106.2220°E, altitudes 1000 m above sea level, 09.ix.2017, V. S. Artamonova, A.A. Makhrov &amp; Y. Fang leg.</p> <p>Etymology. Species is named in honor of Dr. Yury Yu. Dgebuadze, an Academician of the Russian Academy of Sciences, leading hydrobiologist, well-known researcher of Central Asia, and the organizer of the Russian-Chinese scientific expeditions for freshwater biodiversity studies throughout the Qinling Mountains and Tibetan Plateau.</p> <p>Differential diagnosis. Externally, shell of Radix dgebuadzei sp. nov. is rather similar to the shells of Radix plicatula and R. euphratica (Mousson, 1874) (see Aksenova et al. 2018 for their morphological characteristics), but the new species can be distinguished from the latter taxa by a smaller size (shell size of Radix plicatula and R. euphratica may reach 20 mm or a bit more) and by spire height and aperture shape (Fig. 2G, J). Also, Radix dgebuadzei sp. nov. has a darker mantle pigmentation and clearer “freckles” (small black dots) on the foot (Fig. 2C, D) compared to the two other species (Fig. 2H, I, K).</p> <p>Molecular diagnosis. The most reliable method of identification of this species is molecular diagnostics. Phylogenetically, the new species is most closely related to Radix plicatula and R. euphratica (mean uncorrected COI p-distances 7.7% and 5.8%, respectively) (Fig. 3) but can be distinguished from all other Radix congeners by 3 fixed nucleotide substitutions in the COI gene as follows: 212 C, 314 T, 485 C. The nucleotide substitutions that distinguish the new species from the other are synonymous. Two previously unknown haplotypes were found within the studied sample (Fig. 4). The 28S gene fragment has not unique substitutions.</p> <p>Description. Shell morphology. Shell small (up to 10.5 mm in height) (Fig. 2A, B), relatively thick-walled, light brown colored, ovoid or drop-like in shape, with moderately enlarged body whorl and relatively high spire (spire constitutes approximately 1/3 of a total shell height). Whorl number 3.12–3.50. Tangential line almost straight. Whorls very weakly convex, separated by shallow suture. Shell surface sculpture represented by thin axial growth lines. Aperture ovoid, its basal margin evenly rounded. Columellar fold almost invisible. Columellar lip thin and narrow, umbilicus absent. The measurements of the type series are presented in Table 2. Soft body. Mantle black with white spots and “freckles” on the foot (Fig. 2C, D). Reproductive anatomy. The structure of the copulatory organ is typical for the radicines (Fig. 2E). Praeputium is oblong, light gray, its width is almost the same along the entire length. The penis sheath narrower, its distal end somewhat swollen. The praeputium length slightly shorter or equal to the length of the penis sheath. The spermathecal duct long (Fig. 2F). Prostate with a single fold inside (not shown).</p> <p>Habitat. The type specimens were collected from a fast-flowing mountain river with transparent water, rapids and runs, on the stones.</p> <p>Distribution. Known only from its type locality, the Sun Jia He River, Gansu Province, China (Fig. 1). It is likely that this species can also be found in surrounding areas of the Yangtze River basin.</p></div> 	https://treatment.plazi.org/id/6B7BE270BC1DAA23FF02FE72FB16FD04	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Aksenova, Olga V.;Vinarski, Maxim V.;Kondakov, Alexander V.;Tomilova, Alyona A.;Artamonova, Valentina S.;Makhrov, Alexander A.;Kononov, Oleg D.;Gofarov, Mikhail Yu.;Fang, Yun;Bolotov, Ivan N.	Aksenova, Olga V., Vinarski, Maxim V., Kondakov, Alexander V., Tomilova, Alyona A., Artamonova, Valentina S., Makhrov, Alexander A., Kononov, Oleg D., Gofarov, Mikhail Yu., Fang, Yun, Bolotov, Ivan N. (2019): A new Radix species from Qinling Mountains, China (Gastropoda: Lymnaeidae). Ecologica Montenegrina 26: 137-146, DOI: 10.37828/em.2019.26.10, URL: http://dx.doi.org/10.37828/em.2019.26.10
