taxonID	type	description	language	source
6B0687C5A665FFF5ADD3F8AEE6829A86.taxon	materials_examined	Type locality. India: Andaman. Type depository. MNHN. Specimen examined. No specimens seen, description summarized from Tian & Deuve (2015).	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A665FFF5ADD3F8AEE6829A86.taxon	description	Description. Length 15.0 mm. Head, pronotum, scutellum and elytra brownish black; two reddish yellow spots on the frons; antennae with segments 1 – 4 light reddish brown, remaining segments dark brown; legs dark reddish yellow. Head with frons and vertex smooth, neck punctated and wrinkled; eyes only slightly protruding; antennae reaching the middle of elytra, with segment 3 longer than segments 1 and 2 combined. Pronotum smooth, disc convex, glabrous; basal and apical margins straight without sinuation, apical margin with anterior projection on lateral sides; disc laterally with anterior three-fourths convex and posterior fourth straight, hind angles almost forming a right angle, blunt, not projecting laterally; median groove shallow, deep and wide just before base; anterior and posterior sulcus deep, well impressed; lateral beads broad. Elytra with humerus obsolete, base narrowed, widest just before apex; apex obliquely truncate, rounded, not forming any re-entrant angle; elytra smooth with dense setae along the lateral and apical portion of disc, with central region of disc less pubescent.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A665FFF5ADD3F8AEE6829A86.taxon	distribution	Distribution. INDIA: Andaman Islands.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A665FFF5ADD3F8AEE6829A86.taxon	discussion	Remarks. This species is recorded only from the Andaman Islands (India), which are distant from the Indian mainland. It is different from all other Indian Brachinus with the presence of two reddish yellow spots on frons (head).	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A664FFF5ADD3FC8BE6919DAD.taxon	materials_examined	Type locality. India: Tamil Nadu: Tharangambadi, coll. Dr. Dohrn. Type depository. Type lost. Specimen examined. No specimens seen, description summarized from Andrewes (1923).	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A664FFF5ADD3FC8BE6919DAD.taxon	description	Description. Length 5.5 mm. Head, pronotum and narrow lateral margin of elytra (except the base) reddish yellow with head having a darker shade; first two segments of antennae yellow, rest brown; lateral margin of pronotum and elytra greyish black; scutellum dull brick-red; elytra with two reddish yellow spots. Head broad, frons and middle of vertex smooth, neck slightly wrinkled, with sparse pubescence, rest glabrous; shallow rounded foveae between eyes; antennae not reaching the middle of the elytra. Pronotum as long as broad, punctated, pubescent; lateral bead narrow; disc laterally with anterior two-thirds convex, posterior third straight, lateral portion of the apical margin projecting forwards. Elytra longer than broad, convex; weakly striated, intervals slightly convex, finely granulated, moderately pubescent; surface with two spots, the anterior spot oval, middle spot rounded, apical spot absent; apex brown.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A664FFF5ADD3FC8BE6919DAD.taxon	distribution	Distribution. INDIA: Tamil Nadu: Tharangambadi.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A664FFF5ADD3FC8BE6919DAD.taxon	discussion	Remarks. The type species, which was presumed to be in BMNH, is lost or misplaced (Hrdlička 2018 pers. comm.).	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A664FFF2ADD3F9E7E53799DD.taxon	materials_examined	Type locality. India: Himachal Pradesh: Shimla, coll. Dr. Bacon. Type depository. MNHN. Specimen examined. No specimens seen, description summarized from Chaudoir (1876).	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A664FFF2ADD3F9E7E53799DD.taxon	description	Description. Length 8.5 – 9.5 mm. Head, pronotum, antennae reddish brown; elytra black; legs pale yellowish red. Head stout, broad, glabrous, smooth; neck almost smooth. Eyes less protruding. Pronotum as long as wide, glabrous, not exceeding the width of head with eyes; laterally convex, moderately narrowed before the base, hind angles not pointed, not projecting laterally. Elytra convex, with faint, wide striations, having equally raised intervals; disc smooth with slight punctuations over the intervals.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A664FFF2ADD3F9E7E53799DD.taxon	distribution	Distribution. INDIA: Himachal Pradesh: Shimla.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A664FFF2ADD3F9E7E53799DD.taxon	discussion	Remarks. This species was described as Brachinus atripennis Chaudoir, 1876, but diagnosed as a homonym and renamed as Brachinus chaudoirianus by Jakobson (1908).	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A663FFF2ADD3FE77E5889DE8.taxon	description	Fig. 1 h	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A663FFF2ADD3FE77E5889DE8.taxon	materials_examined	Type locality. Bangladesh: Dhaka, coll. M. S. Stevens. Type depository. MNHN. Specimen examined (n = 1). Syntype labelled: “ ‘ Deccan’, coll. Stevens, det. Chaudoir ”.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A663FFF2ADD3FE77E5889DE8.taxon	description	Description. Length 6.7 mm. Head, pronotum, first two segments of antennae, palpi, scutellum, region around scutellum, sutural line, lateral line of elytra (except the base) and legs reddish yellow (with legs having a lighter shade); segments 3 – 4 of antennae reddish brown, rest dark brown; eyes and elytra black. Pronotum with median groove and lateral margins dark brown. Head as long as wide, shiny, vertex smooth, frons, neck and region around eyes wrinkled, sparsely pubescent; apical segment of palpi tapering, pointed; eyes protruding; frontal foveae moderately deep; antennae short, not reaching the middle of elytra, segment 3 shorter than segment 1. Pronotum flat, punctated, pubescent, fine wrinkles on each side of median groove; disc with apical margin sinuate, lateral region of apical margin not projecting forward, basal margin sinuate; disc laterally with anterior twothirds convex, posterior third slightly divergent, hind angles blunt, obtuse, very slightly projecting laterally; median groove shallow; lateral bead broad. Elytra with matte surface, punctated, pubescent, base narrowed, humerus straight, corners rounded; apex without re-entrant angle; scutellum narrow, pointed; striations weak, rounded, intervals broad; both striations and intervals covered by setae; sutural line broadest at base, gradually narrowing till the middle of elytra, extending till the apex.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A663FFF2ADD3FE77E5889DE8.taxon	distribution	Distribution. BANGLADESH: Dhaka. Pakistan (Hrdlička 2003, 2017).	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A663FFF2ADD3FE77E5889DE8.taxon	discussion	Remarks. Brachinus cinctellus Chaudoir, 1876 and B. limbellus Chaudoir, 1876 are very closely related species which, according to Chaudoir (1876) are “ … without any appreciable difference in the conformation of the external morphology, but different in size and colouring of the abdomen and the absence of spots on the antennae. ” We consider the variation in size and colouration of the abdomen as minor variations, and spots on the antennae of B. cinctellus are not visible, which may be due to decolouration over the years.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A663FFF3ADD3F9A5E30C9AD1.taxon	description	Fig. 1 a	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A663FFF3ADD3F9A5E30C9AD1.taxon	materials_examined	Type locality. India: Assam: Eastern Duars, coll. H. Stevens. Type depository. BMNH. Specimen examined (n = 1). Holotype labelled: “ Type, ‘ E. Dooars’, coll. H. Stevens, ‘ Brachinus stenomus ’ ‘ Brachinus dryas nom. nov. Andrewes 1936 p. 65 ’, det. H. E. Andrewes ”.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A663FFF3ADD3F9A5E30C9AD1.taxon	description	Description. Length 6.5 – 7.5 mm. Colour reddish brown (rust-coloured); venter, elytra, margins of pronotum (base rather widely), mes- and metepisterna and apical joints black; legs brownish red. Microsculpture isodiametric, very clear on the elytra, weak on head and pronotum. Head with frontal foveae shallow, subrugose; neck sparsely punctuate, surface otherwise smooth; eyes moderately prominent; apical segment of palpi narrowed and subtruncate at apex; antennae reaching middle of elytra. Pronotum cordate, narrower than head, wider than long; extremities truncate, base as wide as apex, sides round- ed in front and rather strongly sinuate at about basal fourth, both front and hind angles fairly sharp, the latter slightly acute and slightly projecting laterally; median groove short, moderately impressed, punctate at extremities, basal foveae rather shallow, surface sparsely and finely punctate. Elytra convex, two and a quarter times as wide as pronotum, two-fifths longer than wide, contracted in front; humerus straight, corners rounded; elytra widest just before apex, apex with a re-entrant angle, the pale membranous margin very narrow, glabrous; striae weak, without any rows of punctures, intervals moderately raised but hardly costate, surface finely but not very closely punctate. Body covered with rather scanty grey pubescence, slightly thicker on the elytra.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A663FFF3ADD3F9A5E30C9AD1.taxon	distribution	Distribution. INDIA: Sikkim: Gopaldhara; Assam: Eastern Duars.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A663FFF3ADD3F9A5E30C9AD1.taxon	discussion	Remarks. Brachinus dryas is endemic to the Eastern Himalayas. It was described as Brachinus stenomus Andrewes, 1932, which was later found to be a homonym and thus changed to Brachinus dryas Andrewes, 1936.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A662FFF3ADD3FD73E4869DB1.taxon	description	Fig. 1 i	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A662FFF3ADD3FD73E4869DB1.taxon	materials_examined	Type locality. India: Uttarakhand: Dehra Dun, coll. J. C. M. Gardner and G. D. Bhasin. Type depository. BMNH. Specimen examined (n = 1). Holotype labelled “ Type, Dehra Dun, U. P., coll. J. C. M. Gardner, 20. III. 1933, ‘ Ex. Descented termite mound’, ‘ 2328 ’, det. H. E. Andrewes, Brit. Mus. 1937.548. ”	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A662FFF3ADD3FD73E4869DB1.taxon	description	Description. Length 18.0 – 19.0 mm. Head and pronotum brownish red; base and lateral margin of pronotum brownish black; palpi, antennae and legs reddish brown; elytra black with three reddish yellow spots. Head convex; wide, shallow frontal foveae; surface finely punctate; eyes prominent; antennae stout, almost reaching the middle of elytra. Pronotum convex, cordate, wider than head, wider than long; surface finely punctate, becoming coarse along base; base as wide as apex; lateral bead narrow; disc laterally with anterior two-thirds strongly convex, posterior third strongly sinuate, hind angles acute, sharp, projecting laterally; median groove fine, deeper near base; transverse impression present, shallow, clear. Elytra convex, dilated behind, longer than broad, humerus rounded; apex with a re-entrant angle and glabrous membranous border; intervals convex with fine scattered punctures; three reddish yellow spots present, first fairly large, placed near to humerus and rounded, second irregularly rounded and just behind mid region, third small, oblique, near apex.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A662FFF3ADD3FD73E4869DB1.taxon	distribution	Distribution. INDIA: Uttarakhand: Dehra Dun.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A662FFF0ADD3F99BE24799F9.taxon	description	Fig. 2 a	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A662FFF0ADD3F99BE24799F9.taxon	materials_examined	Type locality. India: Tamil Nadu: Coromondel Coast, coll. Guerin. Type depository. MNHN. Specimen examined (n = 1). Syntype labelled “ ‘ Coromandel’, coll. Guerin, det. Chaudoir ”.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A662FFF0ADD3F99BE24799F9.taxon	description	Description. Length 5.0 – 6.5 mm. Head, pronotum, antennae, scutellum and sutural line reddish brown; eyes black; elytra bluish black; legs pale reddish yellow. Head broad, wider than long, punctate, pubescent, neck rough, wrinkled; eyes protruding; antennae stout, short, not reaching the middle of elytra, first four segments of lighter colour, segment 3 equals the combined length of segments 1 and 2, apical segment elongate oval. Pronotum longer than broad, rough, wrinkled, pubescent; disc with apical and basal margin sinuate, with apical outwardly sinuate and basal inwardly sinuate, lateral edge of the apical margin not projecting forward; disc laterally with anterior two-thirds convex, posterior third divergent, hind angles blunt, acute, slightly projecting laterally; lateral bead narrow, lateral margin brownish black; median groove deep, deepest at basal and apical region; basal transverse foveae present, shallow. Elytra long, sub-parallel, glossy, punctated, pubescent; humerus prominent, narrow, rounded; apex without reentrant angle, having membranous border; very weak, broad, rounded striations, narrow intervals.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A662FFF0ADD3F99BE24799F9.taxon	distribution	Distribution. INDIA: Maharashtra: Nagpur; Tamil Nadu: Coromondel coast	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A661FFF0ADD3FE5CE7B19CFA.taxon	materials_examined	Type locality. India: Pune; coll. P. V. Joshi Type depository. BMNH Specimen examined. No specimens seen, description summarized from Hrdlička (2019).	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A661FFF0ADD3FE5CE7B19CFA.taxon	description	Description. Length: 9.8 mm. Head, pronotum and mandibles rusty red; antennomeres 1 – 4 yellow, 5 – 11 yellow brown; elytra black with four yellow spots; legs yellow. Head longer than wide, frons (anteriorly), neck (entirely) punctuate, sparsely pubescent. Pronotum longer than wide; apical margin straight with lateral region projecting forward, basal margin sinuate; disc convex, slightly matt, punctated pubescent, laterally with anterior three-fourths convex, posterior fourth divergent; median groove uniformly deep, reaching the apical and basal margins. Elytral disc matte, punctate, pubescent; base narrowed, apex wide, widest just before apex; humerus with corners rounded; apex without re-entrant angle.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A661FFF0ADD3FE5CE7B19CFA.taxon	distribution	Distribution. INDIA: Maharashtra: Pune.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A661FFF0ADD3FE5CE7B19CFA.taxon	discussion	Remarks. This species is close to B. illotus Chaudoir, 1876 among the described Indian Brachinus. Brachinus illotus differs by having a smaller form, elytra having lateral reddish yellow margins, elytral apex straightly truncate, reddish yellow spot near to elytral apex smaller, and flattened pronotal disc. For further description of the species see Hrdlička (2019).	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A661FFF1ADD3FB54E5F798E4.taxon	description	Fig. 2 b	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A661FFF1ADD3FB54E5F798E4.taxon	materials_examined	Type locality. India: Bengal: Darjeeling, coll. Maj. R. W. G. Hingston, 14. iii. 1924. Type depository. BMNH. Specimen examined (n = 1). Holotype labelled “ Type, N. India: Darjeeling, 5000 ft., 14. III. 1924, coll. Maj. R. W. G. Hingston, ‘ Everest Exp. Brit. Mus. 1924 - 386. ’, det. H. E. Andrewes.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A661FFF1ADD3FB54E5F798E4.taxon	description	Description. Length 8.5 – 9.0 mm. Head, pronotum and elytra pitch brownish black; scutellum brownish black; epistome, first four segments of antennae and legs reddish yellow; eyes black. Head stout, wider than long, vertex smooth, glabrous, frons and neck punctated, pubescent; eyes protruding; antennae slender, long, reaching the middle of elytra, segment 3 length equals combined length of segments 1 and 2, apical segment elongate-oval. Pronotum broad, wider than long, shiny, wrinkled, pubescent; disc with apical margin straight, lateral edge projecting forward, basal margin inwardly sinuate; disc laterally with anterior half convex, posterior half divergent, hind angles blunt, acute; median groove moderately deep, complete, connecting the apical and basal margin; lateral bead narrow. Elytra convex, shiny, pubescent with dense yellow setae, base narrowed, humerus erased; apex without re-entrant angle, having narrow membranous border; scutellum pointed; striations clear, rounded, intervals narrow, layers of setae along the striations as well as intervals.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A661FFF1ADD3FB54E5F798E4.taxon	distribution	Distribution. INDIA: Sikkim: Gopaldhara; Bengal: Darjeeling.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A660FFF1ADD3FEAEE3779CB3.taxon	description	Fig. 1 c	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A660FFF1ADD3FEAEE3779CB3.taxon	materials_examined	Type locality. India: Bengal, coll. Dr. Bacon. Type depository. MNHN. Specimens examined (n = 7). Syntype labelled “ Ind. Or. Bor. ’, coll. Dr. Bacon, det. Chaudoir ”; 5 specimens (2 males, 3 females) labelled “ India: Tamil Nadu: Ettimadai, Hand Picking, 08. IX. 2015, coll. S. V. Akhil ”; 1 specimen labelled “ India: Gujarat: Anand, 17. XI. 2017, coll. Thangavel ”.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A660FFF1ADD3FEAEE3779CB3.taxon	description	Description. Length 8.5 – 9.5 mm. Head, pronotum and antennal segments 2 – 11 reddish brown; antennal segment one, legs and six spots on elytra yellowish red; pronotum with brownish black lateral margins; scutellum reddish brown; eyes and elytra black. Head short, stout, rough, closely punctated, sparsely pubescent; neck highly wrinkled; eyes protruding; last segment of palpi narrowed and sub-truncate at apex; antennae short and stout, not reaching the middle of the elytra; length of antennal segment 3 equals the combined length of segments 1 and 2. Pronotum broad, wider than long; disc laterally with anterior two-thirds convex, posterior third slightly divergent; hind angles pointed, sharp, but not protruding laterally; disc with surface highly punctated, wrinkled, pubescent with distantly placed setae; median groove shallow but visible in the mid region of disc, erased near to base and apex of pronotum. Elytra broad, longer than wide, widest just before apex; scutellum pointed; humerus straight, corners rounded; apex straight, without re-entrant angle; striations broad, weak, shallow, rounded, clear towards base but faint towards apex, intervals shallow, narrow; surface highly punctated, densely pubescent; three yellowish red spots on each elytron, larger square shaped spot just below the humerus extending between the fourth and eighth striations, smaller, rounded spot just below the middle of elytra, oval shaped spot along the apical edge of elytra, not covering the apical edge.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A660FFF1ADD3FEAEE3779CB3.taxon	distribution	Distribution. INDIA: Bengal; Gujarat: Anand; Tamil Nadu: Ettimadai; Arunachal Pradesh; Himachal Pradesh; Kashmir; Sikkim, Uttarpradesh. BHUTAN (Hrdlička 2003, 2017). PAKISTAN (Hrdlička 2003, 2017).	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A660FFFEADD3FA9DE4669915.taxon	description	Fig. 2 c	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A660FFFEADD3FA9DE4669915.taxon	materials_examined	Type locality. Bangladesh: Dhaka (‘ Deccan’), coll. M. S. Stevens. Type depository. MNHN. Specimen examined (n = 1). Syntype labelled: “ ‘ Deccan’, coll. Stevens, det. Chaudoir ”.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A660FFFEADD3FA9DE4669915.taxon	description	Description. Length 6.5 mm. Head and pronotum reddish yellow; legs, spots on elytra, lateral margin of elytra (except the basal portion) and first four segments of antennae pale reddish yellow; eyes black; elytra brownish black. Head slender, longer than broad, neck with fine setae, otherwise glabrous, frons, vertex smooth, shiny; frontal foveae shallow; eyes protruding; antennae long, slender, reaching the middle of elytra, segment 3 equals the combined length of segments 1 and 2. Pronotum narrower than head, wider than long, densely pubescent, smooth; disc with apical margin inwardly sinuate, without lateral sides projecting forward, basal margin straight; disc laterally with anterior two-thirds convex, posterior third divergent, hind angles blunt, acute, slight lateral projection; median groove shallow; lateral bead narrow, with dark brown shade. Elytra long, narrow, subparallel, base narrowed, rounded, humerus erased, apex with less evident re-entrant angle, having membranous border; disc densely pubescent, covered with yellow setae; striations shallow, faint, rounded, intervals narrow; two reddish yellow spots, first elongate oval, placed near base, extending between fifth stria and lateral margin, second rounded, placed just after the mid region, a wide reddish yellow line running from the basal spot to apex.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A660FFFEADD3FA9DE4669915.taxon	distribution	Distribution. INDIA: Madhya Pradesh: Dr. Ambedkar Nagar. BANGLADESH: Dhaka. CAMBODIA: Phnom Penh. VIETNAM: Cochinchina.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A66FFFFEADD3FE3FE4879D7B.taxon	description	Fig. 1 b	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A66FFFFEADD3FE3FE4879D7B.taxon	materials_examined	Type locality. India: Uttarakhand: Almora, coll. H. G. Champion. Type depository. BMNH. Specimens examined (n = 4). Holotype labelled “ Type, ‘ India: W. Almora Du, Almora U. P’, Mar. ’ 19, coll. H. G. Champion, B. M. 1932 - 27, det. H. E. Andrewes; 2 males, 1 female, labelled “ India: Karanataka: Kolar: Krishnarajapuram, 25. xii. 2001, coll. Dr. Raghunathan ”.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A66FFFFEADD3FE3FE4879D7B.taxon	description	Description. Length 7.0 – 8.0 mm. Elytra, lateral borders of pronotum, metepisterna, and apical joints of venter black; head, pronotum and legs brownish red with head darker. Microsculpture isodiametric, distinct on elytra, indistinct on pronotum, invisible on head. Head with frontal fovea subrugose and moderately deep; sides of neck punctated, middle and front of vertex smooth; eyes moderately prominent; apical segment of palpi bluntly pointed; antennae long, slender, extending beyond the middle of elytra. Pronotum cordate, narrower than head, wider than long, base narrower than apex; disc punctate with lateral sides convex anteriorly, diverging posteriorly; basal and apical angles fairly sharp, basal angle projecting laterally; median groove fine, shallow medially and deeper at extremities; basal fovea moderately deep. Metepisterna length twice the width. Elytra convex, widest just before apex, longer than wide, highly contracted to base, slightly contracted to apex; humerus straight, corners rounded; apex with a re-entrant angle; pale membranous margin without long seta, having minute pale hairs; striae wide, shallow, without any rows of punctures, intervals moderately convex; disc densely punctate.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A66FFFFEADD3FE3FE4879D7B.taxon	distribution	Distribution. INDIA: Uttarakhand: Tanakpur, Haldwani, Ranikhet, West Almora, Kumaon, Haridwar, River Sardah Gorge; Karnataka: Kolar: Krishnarajapuram.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A66FFFFFADD3FAD5E44199DD.taxon	description	Fig. 2 d	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A66FFFFFADD3FAD5E44199DD.taxon	materials_examined	Type locality. Bangladesh: Dhaka, coll. M. S. Stevens. Type depository. MNHN. Specimen examined (n = 1). Syntype labelled “ ‘ Deccan’, coll. Stevens, ‘ suturiger ’ Chaudoir, ‘ limbellus’ Chaudoir ”.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A66FFFFFADD3FAD5E44199DD.taxon	description	Description. Length 4.5 – 5.0 mm. Head, pronotum, sutural line, base of elytra and antennal segments 3 – 11 reddish brown; antennal segments 1 – 2, palpi and legs reddish yellow; elytra brownish black. Head stout, as long as wide, smooth, glabrous; frontal foveae clear, moderately deep; eyes protruding; antennae short, stout, not reaching the middle of the elytra, antennal apical segment longest, segment 3 shorter than segment 1 and 2 combined. Pronotum as wide as long, punctated, pubescent; disc with apical and basal margins straight, lateral portion of apical margin slightly projecting forward; disc laterally with anterior 2 / 3 rd convex, posterior 1 / 3 rd divergent, hind angle blunt, obtuse, projecting laterally; median groove faint, superficial; lateral bead very narrow, dark brown. Elytra convex, base narrow, humerus straight, corners rounded, narrow, pubescent with closely placed setae; apex with a very faint re-entrant angle, having a membranous border; scutellum very small; lateral margin of disc lined with a flat, reddish yellow bead which increases in width towards apex; very weak striations; sutural colour extending to the apex, broadest at base, extending between first and third striae, gradually narrowed towards apex.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A66FFFFFADD3FAD5E44199DD.taxon	distribution	Distribution. INDIA: Assam; Bengal: Kolkata; Odisha: Puri; Maharashtra: Bandra; Tamil Nadu: Thiruchirapalli. SRI LANKA: Murunkan. BANGLADESH: Dhaka. MYANMAR: Rangoon, Senmigion, Palon, Magwe. PAKISTAN (Hrdlička 2003, 2017).	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A66EFFFFADD3FE77E5179CB3.taxon	description	Fig. 2 e	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A66EFFFFADD3FE77E5179CB3.taxon	materials_examined	Type locality. Bangladesh: Dhaka, coll. M. S. Stevens. Type depository. MNHN. Specimen examined (n = 1). Syntype labelled “ ‘ Deccan’, coll. Stevens, det. Chaudoir ”.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A66EFFFFADD3FE77E5179CB3.taxon	description	Description. Length 9.0 mm. Head, mid region of pronotum reddish brown; palpi, antennal segments 1 – 4, and legs reddish yellow; lateral sides of pronotum brownish black; eyes black; scutellum brownish black; elytra black with light brown shade. Head longer than broad, frons rough, vertex smooth, fine pubescence around the eyes, otherwise glabrous; frontal foveae shallow; eyes protruding; antennae long, reaching the middle of the elytra, antennal segment 3 equals the combined length of segments 1 and 2; last segment of palpi pointed. Pronotum as wide as long, punctated, densely pubescent, with fine yellow setae; disc with apical margin inwardly sinuate, basal margin straight, apical margin without lateral region projecting forward; disc laterally with anterior two-thirds convex, posterior third divergent, hind angle blunt, acute; median groove shallow; lateral bead narrow. Elytra sub-parallel, long, densely pubescent with yellow setae; humerus narrow, straight, corners rounded; apex with very faint re-entrant angle, having narrow membranous border; scutellum small, pointed; striations weak, intervals shallow.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A66EFFFFADD3FE77E5179CB3.taxon	distribution	Distribution. INDIA: Bihar: Pusa. BANGLADESH: Dhaka.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A66EFFFCADD3FA9DE4CA9A4D.taxon	materials_examined	Type locality. India: Bengal, coll. Dejean, Dr. Bacon. Type depository. ZMUC (Hrdlička 2019 pers. comm.) Specimen examined (n = 1). 1 ex. labelled “ longipalpis wied. ‘ Bengal Western’ ”.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A66EFFFCADD3FA9DE4CA9A4D.taxon	description	Description. Length 5.5 – 7.0 mm. Head, pronotum, first four segments of antennae, legs, markings on the elytra, scutellum, and region around scutellum reddish yellow, with head darker; eyes grey; elytra black; antennal segments 5 – 11 dark brown. Head stout, wider than long, smooth, shiny, only a small patch of setae around the eyes, otherwise glabrous; frontal foveae shallow; eyes protruding; antennae long, slender, reaching the middle of the elytra, segment 3 the longest, longer than segments 1 and 2 combined; last segment of palpi oval, pointed. Pronotum as long as wide, smooth, shiny, without any setae; disc with apical margin straight, lateral region of apical margin projecting forward, basal margin outwardly bi-sinuate; disc laterally with anterior half wide, posterior half parallel, hind angles pointed, forming right angles, not projecting laterally; median groove deep, reaching both apical and basal margin. Elytra convex, symmetrically rounded, flat, moderately pubescent, with yellow setae; humerus rounded; apex rounded without re-entrant angle; scutellum small, narrow, pointed; striations weak, broad, rounded, intervals broad, flatter but rounded, rows of setae covering both striations and intervals; two reddish yellow markings, first a transverse band, in the middle of elytra, beginning from the lateral margin, extending up to first striation, second a triangular one, with base of the triangle over the apex of elytra, a narrow reddish yellow line running laterally, starting just above the median band and extending till the tip of apex of elytra.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A66EFFFCADD3FA9DE4CA9A4D.taxon	distribution	Distribution. INDIA: Sikkim: Teesta Valley; Uttarakhand: Almora, Dehra Dun, Tanakpur, Haldwani, Garhwal: Jumnagwar, Boxar, Dikhwala; Bengal. MYANMAR: Senmigion, Myenkyan, Rangoon. AFGHANISTAN (Hrdlička 2003, 2017). PAKISTAN (Hrdlička 2003, 2017).	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A66EFFFCADD3FA9DE4CA9A4D.taxon	discussion	Remarks. A species with unique markings on elytra, having a reddish yellow median transverse band similar to the related genus Pheropsophus Solier, 1833.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A66DFFFCADD3FDC7E33C9E3A.taxon	description	Fig. 2 f	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A66DFFFCADD3FDC7E33C9E3A.taxon	materials_examined	Type locality. India: Bengal: Kolkata. Type depository. NMPC. Specimen examined (n = 1). Type labelled: “ Bengal ”.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A66DFFFCADD3FDC7E33C9E3A.taxon	description	Description. Length 5.2 mm. Head, pronotum, legs, sutural line, antennal segments, palpi, scutellum and region around scutellum reddish brown; eyes grey; elytra brownish black. Head stout, broad, wider than long, punctated, pubescent; vertex smooth, glabrous, frons, region around eyes and neck punctated, pubescent; eyes protruding; frontal foveae deep; antennal segments stout, short, not reaching the middle of the elytra, segment 3 shorter than segment 1. Pronotum slightly broader than long, punctated, densely pubescent; disc with apical margin straight, lateral region of apical margin not projecting forward, basal margin slightly sinuated outward; disc laterally with anterior three-fourths convex, posterior fourth divergent, hind angle obtuse, blunt, projecting laterally; median groove deep reaching both apical and basal margin; lateral bead narrow, dark brown. Elytra subparallel, punctated, densely pubescent, with golden brown setae; base narrow, rounded, humerus erased; apex without a re-entrant angle; scutellum very small, pointed; striations weak, rounded, intervals narrow, setae covering both striations and intervals; sutural colour extending till middle of elytra, widest near base where it extends between 1 st and 3 rd striae, narrowing gradually towards apex.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A66DFFFCADD3FDC7E33C9E3A.taxon	distribution	Distribution. INDIA: Bengal: Kolkata. MYANMAR: Tharrawaddy. Prome. Toungoo. Rangoon. Moulmein. VIETNAM: Cochinchina. CHINA: Fujiyan (Hrdlička, 2017).	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A66DFFFCADD3FDC7E33C9E3A.taxon	discussion	Remarks. Comparison of the type specimens of B. modestus Schmidt-Göbel, 1846, B. peltastes Andrewes, 1931 and B. suturellus Chaudoir, 1876 shows that there is no significant variation between the three species other than the colour (elytra black in B. suturellus, pitch black in B. peltastes and brownish black in B. modestus) and size (5.2 mm in B. modestus, 6.0 mm in B. peltastes and 8.5 mm in B. suturellus). We have not synonymised Brachinus suturellus and B. peltastes with B. modestus as we are not certain how far the colour of the types have faded.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A66DFFFDADD3F914E7489AFD.taxon	materials_examined	Type locality. India: Himachal Pradesh: Poo, coll. Fleischer. Type depository. Type in collection of A. Fleischer (Andrewes 1928)	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A66DFFFDADD3F914E7489AFD.taxon	description	Description. Length 11.0 mm. First four segments of antennae, palpi and legs pale yellow; antennal segments 5 – 11, head and pronotum yellowish red; scutellum brownish red; elytra bluish black; eyes black. Head narrow, longer than broad; surface smooth, glabrous; eyes large; last segment of palpi narrowed and sub-truncate at apex; antennae long, slender, reaching beyond the middle of elytra, with segment three the longest, segment 3 longer than segments 1 and 2 combined, segment 2 shortest. Pronotum narrow, longer than broad, glabrous, wrinkled; disc laterally convex with anterior half rounded and posterior half divergent; lateral bead present, having a darker brownish black shade; hind angles sharp, projecting laterally; median groove fine, shallow medially, deep towards base and apex. Elytra broad, slightly rounded, widest just before apex, apex and base narrow, apex rounded, without re-entrant angle; humerus straight, rounded at corners; striations weak, broad, rounded, striae 1 to 6 (counting suture as 1) clear, well impressed, intervals narrow; surface rough, wrinkled, highly pubescent.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A66DFFFDADD3F914E7489AFD.taxon	distribution	Distribution. INDIA: Jammu & Kashmir: Gurais valley; Himachal Pradesh: Pooh, Bashahr, Kangra; Uttarakhand: Almora, Dehra Dun, Tanakpur, River Sarda gorge. PAKISTAN: Gilgit-Baltistan, Khyber Pakhtunkhwa.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A66DFFFDADD3F914E7489AFD.taxon	discussion	Remarks. Brachinus pallidipes was misidentified by Andrewes (1924 a) as B. stenoderus Bates, 1873, an entirely different species with a Japanese distribution (Andrewes 1928). The species with a Himalayan distribution is B. pallidipes.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A66CFFFDADD3FD56E51C9E27.taxon	description	Fig. 2 g	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A66CFFFDADD3FD56E51C9E27.taxon	materials_examined	Type locality. India: Dehra Dun: Dobhalwala, coll. H. E. Andrewes. Type depository. BMNH. Specimen examined (n = 1). Holotype labelled “ Dehra Dun: Dhobalwala, ‘ 2,304 ft. ’, H. E. Andrewes Coll. B. M. 1945 - 97, ‘ bought from Staudinger & Bang-Haas, 1930, det. H. E. Andrewes ”.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A66CFFFDADD3FD56E51C9E27.taxon	description	Description. Length 6.0 mm. Head, pronotum, scutellum, region around scutellum, sutural line reddish brown; first four segments of antennae, palpi and legs pale reddish yellow; eyes grey; elytra black. Head stout, wider than long, shiny, glabrous, vertex smooth, frons, region around eyes and neck wrinkled; eyes protruding; last segment of palpi pointed; antennae stout, just reaching the middle of elytra; 3 rd segment of antennae shorter than 1 st and 2 nd combined. Pronotum broad, wider than long, punctated, pubescent, shiny; disc with apical margin inwardly sinuate, lateral portion of apical margin not projecting forward, basal margin slightly outwardly sinuate; disc laterally with anterior three-fourths convex, posterior fourth parallel, hind angle blunt, slightly obtuse, not projecting laterally; median groove deep, reaching the apical and basal margins; apical transverse furrow visible, shallow; lateral bead moderately wide, lateral margin dark brown. Elytra shiny, pubescent, covered with short yellow setae; base narrow, rounded, humerus obliterate; apex straight with less evident re-entrant angle, having a membranous border; suture not closed completely towards apex; scutellum small, pointed; striations weak, narrow, keel shaped, intervals broad setae covering both striations and intervals; sutural colour extending till middle of elytra, widest near base extending between 1 st and 4 th striae, narrowing gradually towards apex.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A66CFFFDADD3FD56E51C9E27.taxon	distribution	Distribution. INDIA: Uttarakhand: Dehra Dun: Dobhalwala.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A66CFFFAADD3F969E5329C65.taxon	description	Fig. 1 g	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A66CFFFAADD3F969E5329C65.taxon	materials_examined	Type locality. India: Maharashtra: Pune, Sykes collection. Type depository. Type lost. Specimens examined (n = 3). 2 exs., (1 male, 1 female) labelled “ India: Andhra Pradesh: Cudappah dt, ‘ 181 m Alt’, 28. xi. 04, coll. Dr. Prabhakar ”, det. S. V. Akhil; 1 ex.; (male) labelled “ India: Kerala: Chinnar, x. 2011, coll. K. V. Vinod & S. T. Raj, det. S. V. Akhil ”.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A66CFFFAADD3F969E5329C65.taxon	description	Description. Length 12.4 mm. Head, pronotum, basic colour of elytra, scutellum, legs and venter reddish yellow. Eyes, three markings on elytra, abdominal lateral margins and posterior segments black; antennae reddish brown with first four segments pale red. Pronotum with median groove and lateral margins reddish brown. Head stout, as long as broad, glabrous, smooth, a patch of setae around the eyes; eyes prominent, protruding; neck smooth. Pronotum broad, as long as wide, smooth, few setae near to base and apex, otherwise glabrous, symmetrically wrinkled on both sides of median groove; lateral beads broad, prominent, disc with posterior two-thirds convex, basal third parallel, without projecting laterally; apical and basal margins straight with lateral edge of apical margin projecting forward; median groove deep, deepest at both ends, not reaching the apical and basal margins. Elytra longer than broad, slightly convex, pubescent, humerus straight, prominent, projecting; apex with a membranous lining, having a re-entrant angle; scutellum pointed; striations 1 and 2 (counting suture as 1) faint, 3 – 5 prominent, keel shaped, rest normal, intervals deep, pubescent along the intervals; surface with a black rectangular longitudinal band starting from the sides of scutellum covering the area between 1 st and 3 rd striations but ending just before the mid region of elytra, two comma shaped black spots near the lateral margin in the anterior half with a narrow projection towards the humerus, a black transverse band just before the apex with its lateral portions large and rounded.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A66CFFFAADD3F969E5329C65.taxon	distribution	Distribution. INDIA: Siwaliks; Delhi; Bengal; Haryana: Kalka; Jharkhand: Medininagar; Maharashtra: Pune, Nagpur; Andhra Pradesh: Cudappah; Karnataka: Belgavi, Bengaluru; Tamil Nadu: Chennai; Kerala: Thrissur, Chinnar. SRI LANKA: Hambantota; PAKISTAN: Lyallpur.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A66CFFFAADD3F969E5329C65.taxon	discussion	Remarks. Unique Indian Brachinus with basic colour of elytra reddish yellow with black spots, whereas all other species have black, bluish black or brownish black elytra. The type, which was in the Hope collection of BMNH, is presumed to be lost or misplaced (Andrewes 1919).	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A66BFFFAADD3FB2DE50F9F3A.taxon	materials_examined	Type locality. Nepal: Pokhara. Type depository: coll. D. W. Wrase, Berlin. Specimen examined. No specimens seen, description summarized from Kirschenhofer (2012).	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A66BFFFAADD3FB2DE50F9F3A.taxon	description	Description. Length 6.1 – 6.8 mm. Head, pronotum, scutellum and region around scutellum, suture and lateral border of elytra dark reddish yellow; first four segments of antennae, legs and palpi pale reddish yellow; eyes grey; elytra black with yellowish setae. Head smooth, shiny, glabrous (except few setae on the frons); eyes protruding; antennae long, slender, reaching beyond the middle of elytra, with 3 rd segment longest, longer than segments 1 and 2 combined; apical segment of palpi pointed. Pronotum as long as broad, smooth, sparsely punctated pubescent; apex sinuate, base straight, apical margin with slight anterior projection on lateral sides; disc laterally with anterior 3 / 4 th moderately convex, posterior 1 / 4 th divergent, hind angles acute; median groove deep in the middle of the disc and shallow towards base and apex. Elytra long, convex, widest after the mid-region; humerus obsolete, rounded; apex straight, truncate without re-entrant angle; striations weak, broad, shallow, intervals rounded; disc densely pubescent.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A66BFFFAADD3FB2DE50F9F3A.taxon	distribution	Distribution. INDIA: Uttarakhand (Anichtchenko, 2019). NEPAL (Hrdlicka, 2017).	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A66BFFFAADD3FB2DE50F9F3A.taxon	discussion	Remarks. Similar to B. stevensi Andrewes, 1924 (Andrews 1924 a but) differing in having the base of the pronotum straight without sinuation and dark black colour of elytra.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A66AFFFBADD3FC7AE7FC9F35.taxon	description	Fig. 1 d	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A66AFFFBADD3FC7AE7FC9F35.taxon	materials_examined	Type locality. India: India oriental, coll. Dejean. Type depository. MNHN. Specimens examined (n = 2). Syntype labelled “ ‘ Indes orient’, ‘ 6 - maculatus’, coll. Dejean ”; 1 ex., (1 female), labelled “ India: Tamil Nadu: Rajapalayam, ‘ Light’, 15. X. 2017, coll. & det. S. V. Akhil ”.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A66AFFFBADD3FC7AE7FC9F35.taxon	description	Description. Length 9.0 – 11.0 mm. Head, pronotum, antennal segments and scutellum reddish brown; legs and spots on elytra reddish yellow; margin around eyes and elytra black. Head short, punctated, pubescent, neck glabrous; antennae short, stout, not reaching the middle of the elytra; eyes prominent. Pronotum elongated, longer than broad, glabrous, shiny, faint wrinkles in middle of the disc, highly wrinkled in basal and apical portion of disc; lateral sides of disc convex with anterior two-thirds highly convex, posterior third divergent; apical margin convex with lateral portion of apical margin projecting forwards, basal margin sinuous, hind angles acute, pointed; median groove deep, reaching the anterior and posterior margins. Elytra subparallel, long, narrow, humerus obliterate, rounded; apex rounded without re-entrant angle; scutellum pointed; striation clear, visible, broad, intervals weak, smooth, shallow; three reddish yellow spots over each elytron, first and third longitudinal, along the corners of base and apex respectively, second rounded, just below the middle of elytra, not reaching either the suture or the lateral margin, a reddish yellow line running from the first spot to the apex forming a apical yellow band; elytral sutures not closed apically.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A66AFFFBADD3FC7AE7FC9F35.taxon	distribution	Distribution. INDIA: Bengal: Kolkata; Himachal Pradesh; Sikkim; Arunachal Pradesh; Uttar Pradesh; Tamil Nadu: Rajapalayam. SRI LANKA. MYANMAR: Tharrawaddy. BHUTAN (Hrdlička 2003, 2017). PAKISTAN (Hrdlička 2003, 2017).	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A66AFFFBADD3FF33E34B9B29.taxon	description	Fig. 2 h	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A66AFFFBADD3FF33E34B9B29.taxon	materials_examined	Type locality. India: Odisha: Ganjam, coll. R. P. J. Rey. Type depository. BMNH. Specimen examined (n = 1). Holotype labelled: “ ‘ Madras: Surada’, coll. R. P. J. Rey, ex. coll. Babault, ‘ figured specimen’, ‘ H. E. Andrewes Coll. B. M. 1945 - 97 ’, det. H. E. Andrewes. ”	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A66AFFFBADD3FF33E34B9B29.taxon	description	Description. Length 11.0 – 13.0 mm. Head, disc of pronotum (mid region) and first three segments of antennae brownish red; three spots on elytra and legs pale reddish yellow; segments 4 – 11 of antennae dark brown; apical, basal and lateral margins of pronotum black, elytra black. Head convex, shiny, rough, vertex punctated; frontal foveae shallow; eyes protruding; antennae long, reaching the middle of elytra, segment 1 very short. Pronotum longer than wide, surface rough, wrinkled, glabrous; disc laterally with anterior 2 / 3 rd convex, posterior 1 / 3 rd divergent, hind angles slightly protruding laterally, angles acute, pointed; median groove shallow, deepest at the extremities. Elytra convex, humerus squared; striations deep, distinctly punctated, intervals carinate; three spots on each elytron, anterior spot placed towards humerus, broad, rounded, covering intervals 4 – 8, smaller middle spot covering intervals 2 – 5, very small posterior spot close to angle outside the truncation.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A66AFFFBADD3FF33E34B9B29.taxon	distribution	Distribution. INDIA: Odisha: Ganjam, Surada; Maharashtra: Igatpuri, Bandra; Karanataka: Kanara.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A677FFE6ADD3FF33E2459C65.taxon	description	Fig. 2 i	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A677FFE6ADD3FF33E2459C65.taxon	materials_examined	Type locality. India: Tamil Nadu: Uduparani, coll. N. C. Chatterjee. Type depository. BMNH. Specimen examined (n = 1). Syntype labelled: “ Uduparani, North Salem?, coll. N C. Chatterjee, 28. I. 1930, ‘ India. Dehra Dun Coll. B. M. 1932 - 26 ’. det. H. E. Andrewes ”.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A677FFE6ADD3FF33E2459C65.taxon	description	Description. Length: 10.0 mm. Head, pronotum, first four segments of antennae and legs reddish yellow; scutellum dark reddish yellow; eyes and elytra black. Head stout, broad, wider than long, shiny, sparsely punctated, surface slightly rough, frontal foveae moderately deep; eyes protruding; antennae long, reaching middle of elytra, antennal segment 3 shorter than segment 1 and 2 combined. Pronotum broad, wider than long, surface rough, densely punctated, pubescent; disc with apical margin inwardly sinuate, lateral portion not projecting forward, basal margin straight; disc laterally with anterior three-fourths convex, posterior fourth divergent, hind angles blunt, obtuse, not projecting laterally; median groove shallow, medially brown, not reaching the apical and basal margin; lateral bead narrow, dark brown. Elytra convex, densely punctated, pubescent, with short yellow setae; base narrowed, humerus rounded; apex with slight but distinct re-entrant angle, having a membranous border; scutellum small, pointed; striations very weak, broad, rounded, intervals narrow.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A677FFE6ADD3FF33E2459C65.taxon	distribution	Distribution. INDIA: Salem, Uduparani?; Uttarakhand: Dehra Dun.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A677FFE6ADD3FF33E2459C65.taxon	discussion	Remarks. Andrewes (1933) remarked that both the specimens considered as types have the locality of Uttarakhand: Dehra Dun, and that the Madras province label was a typographical error. Hence, distribution data provided on the labels of the type species are misleading and the locality of the species is Uttarakhand in the foothills of the Himalayas and not the southern Indian locality (Tamil Nadu, Salem, Uduparani).	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A677FFE7ADD3FB2EE2C19881.taxon	description	Fig. 3 a	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A677FFE7ADD3FB2EE2C19881.taxon	materials_examined	Type locality. India: Gielle Khola, Teesta Valley, coll. H. G. Stevens, 28. xi – 3. xii. 1919. Type depository. BMNH. Specimen examined (n = 1). Holotype labelled “ Giellle Khola, Tista Valley, Darjiling. 28. xi- 3. xii. 19, 600 ft., coll. H. Stevens, ‘ H. E. Andrewes Coll. B. M. 1945 - 97 ’, det. H. E. Andrewes ”.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A677FFE7ADD3FB2EE2C19881.taxon	description	Description. Length 5.0 – 6.5 mm. Head, pronotum, scutellum, region around scutellum, sutural line and base of elytra dark reddish yellow; first four segments of antennae, lateral margin of elytra and legs reddish yellow; elytra brownish black, eyes pitch black. Head convex, almost smooth, shiny, frontal foveae shallow, vertex smooth, region around the eyes, frons, neck punctated, pubescent; antennae long, reaching the middle of elytra, segment 3 longest, longer than segments 1 and 2 combined. Pronotum narrower than head, longer than broad, disc rough, with wrinkles, sparsely pubescent; apical margin of disc straight, with lateral region of apical margin projecting forward, posterior margin bisinuate; disc laterally with anterior two-thirds convex, posterior third divergent, hind angles pointed, acute, projecting laterally; median groove shallow, not reaching the apical and basal margin. Elytra subparallel, punctated, pubescent, with golden brown setae; base narrowed, rounded, humerus obliterate; surface matt; apex straightly truncate without re-entrant angle, having broad membranous border; scutellum small, blunt; striations weak; sutural colour broadest at basal quarter covering the base, narrows down till basal fourth of the elytra, then straight with uniform thickness to the apex; pale yellow lateral margin beginning just below the humerus, widening to apex, forming a pale reddish yellow apical border.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A677FFE7ADD3FB2EE2C19881.taxon	distribution	Distribution. INDIA: Assam: Kobo, Dihing river; Sikkim: Teesta Valley; Bengal: Kalimpong; Himachal Pradesh: Kangra; Uttarakhand: Garhwal: Dhikala, River Sarda Gorge, Ranikhet, Dehra Dun.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A676FFE7ADD3FE83E5C79CB6.taxon	description	Fig. 3 b	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A676FFE7ADD3FE83E5C79CB6.taxon	materials_examined	Type locality. India: North India, coll. Dr. Bacon and Captain Boys. Type depository. MNHN. Specimen examined (n = 1). Syntype labelled “ North India, coll. Dr. Bacon and Captain Boys ”.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A676FFE7ADD3FE83E5C79CB6.taxon	description	Description. Length 8.5 mm. Head, pronotum, elytral sutural line, scutellum, region around scutellum, femur, first two antennal segments dark reddish yellow; tibia and tarsi pale reddish yellow; antennal segments 3 – 11 dark reddish brown; eyes grey; elytra brownish black. Head stout, wider than long, vertex smooth, shiny, glabrous; frons, region around eyes and neck punctated, pubescent; eyes protruding; antennae stout, segments wide, short, not reaching the middle of the elytra; segment 2 very short, segment 3 shorter than segment 1. Pronotum shiny, punctated, densely pubescent; disc with apical margin outwardly sinuate, without lateral portion of apical margin projecting forward, basal margin slightly outwardly sinuate; disc laterally with anterior half convex, posterior half slightly divergent, hind angles pointed, at right angles, not projecting laterally; median groove shallow, not reaching the apical and basal margin; anterior transverse foveae moderately deep; lateral bead broad, dark brown. Elytra subparallel, shiny, punctated, pubescent, with short golden brown setae; base narrowed, humerus present, rounded; apex without re-entrant angle; scutellum small, wider, blunt; striations weak, keel shaped, intervals broad, carinate; sutural line extending from region around scutellum to apex, gradually narrowing from base to apex, broadest at base; having narrow reddish yellow apical band.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A676FFE7ADD3FE83E5C79CB6.taxon	distribution	Distribution. INDIA: Uttarakhand: Almora; Uttar Pradesh: Lucknow; Himachal Pradesh: Narpur (Kangra); Bengal: Bowali; Jharkhand: Barway; Madhya Pradesh: Dr. Ambedkar Nagar (Mhow); Karnataka: Belgavi, Bengaluru. MYANMAR: Tharawaddy, Rangoon. VIETNAM: Cochinchina.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A676FFE4ADD3FA98E33D98AD.taxon	description	Fig. 1 e	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A676FFE4ADD3FA98E33D98AD.taxon	materials_examined	Type locality. Bangladesh: Dhaka, coll. M. S. Stevens. Type depository. MNHN. Specimen examined (n = 1). Syntype labelled. “ ‘ Deccan’, coll. M. S. Stevens, det. Chaudoir ”.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A676FFE4ADD3FA98E33D98AD.taxon	description	Description. Length 16.0 mm. Head, pronotum, antennae and scutellum dark reddish brown; palpi and legs light reddish brown; lateral margin of pronotum and elytra brownish black. Head stout, as wide as long, smooth, glabrous, a patch of setae around the eyes; eyes prominent, protruding; frontal impression deep, clear; neck smooth; antennae short, stout, not reaching the middle of elytra. Pronotum broad, wider than long, smooth, glabrous, wrinkles very fine; disc laterally convex, anterior twothirds rounded, posterior third divergent, hind angle sharp, acute, slightly projecting laterally; apical and basal margin straight, lateral edge of apical margin projecting forward; median groove shallow; anterior transverse furrow clear, deep. Elytra subparallel, broad, glabrous; humerus straight, squared, corners rounded; apex rounded, without reentrant angle; scutellum pointed; striations clear, keel shaped, pointed, intervals flat, broad, smooth; three reddish yellow spots on each elytron, first transverse, extending between fourth and eight striations, second rounded, just below the middle portion, third very narrow along the apical region not reaching the suture; a reddish yellow line starting as a very faint line from the humerus running along the lateral edge joining the first and the third spot, apical portion of first and second striae including the interval brownish black.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A676FFE4ADD3FA98E33D98AD.taxon	distribution	Distribution. INDIA: Bengal: Asansol. BANGLADESH: Dhaka. SRI LANKA: Vavuniya, Hambantota.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A675FFE4ADD3FEE7E7309CCB.taxon	description	Fig. 1 f	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A675FFE4ADD3FEE7E7309CCB.taxon	materials_examined	Type locality. India: Bengal, coll. Dr. Bacon. Type depository. MNHN. Specimen examined (n = 1). Syntype labelled “ ‘ Ind. or. bor. ’, coll. Dr. Bacon, det. Chaudoir ”.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A675FFE4ADD3FEE7E7309CCB.taxon	description	Description. Length 8.5 – 9.0 mm. Head, pronotum, palpi and first four segments of antennae dark reddish yellow; legs and two spots on each elytron pale reddish yellow; antennal segments 5 – 11 dark brown; eyes grey; scutellum brownish black; median groove of pronotum dark reddish brown, lateral margin dark brown; elytra black. Head stout, wider than long, smooth, glabrous, matt; eyes, protruding; apical segment of palpi sub-cylindrical, truncate; antennae, short, stout, not reaching the middle of the elytra, segment 3 longest, equals the combined length of segments 1 and 2. Pronotum narrow, longer than broad, matt, rough, punctated, pubescent; disc with apical margin straight, lateral region of apical margin slightly projecting forward, basal margin sinuate, convex; anterior two-thirds of lateral margin convex, posterior third divergent, hind angles pointed, not projecting laterally; median groove moderately deep; lateral bead narrow. Elytra subparallel, flat, matt, punctated, pubescent; humerus rounded; apex with a well-defined re-entrant angle, having a membranous border; scutellum very small, pointed; striations moderately deep, carinate, intervals wide, both striations and intervals covered by setae; two spots on each elytron, first extended oval with transverse elongation, extending between striations 4 and 8, in the middle of elytra, second at the apex, square shaped.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A675FFE4ADD3FEE7E7309CCB.taxon	distribution	Distribution. INDIA: Bengal; Bihar: Pusa; Maharashtra: Nagpur, Bandra; Odisha: Surada; Karnataka: Belgavi; Kerala: Malabar.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A675FFE4ADD3FB45E2F49FAE.taxon	description	Fig. 3 c	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A675FFE4ADD3FB45E2F49FAE.taxon	materials_examined	Type locality. India: North India, coll. Dr. Bacon. Type depository. MNHN. Specimen examined (n = 1). Syntype labelled “ ‘ Ind. or. bor’, coll. Dr. Bacon, det. Chaudoir.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A675FFE4ADD3FB45E2F49FAE.taxon	description	Description. Length 7.5 mm. Head, pronotum, first two antennal segments, scutellum, region around scutellum and sutural line dark reddish yellow; antennal segments 3 – 11 dark reddish brown; legs pale reddish yellow; eyes greyish black; lateral margin of pronotum dark brown; elytra brownish black. Head stout, wider than long, rough, punctated, pubescent; eyes moderately protruding; antennae short, segments stout, not reaching the middle of elytra, antennal segment 3 shorter than segment 1. Pronotum broad, wider than long, rough, matt, punctated, pubescent; disc with apical margin outwardly sinuate, lateral region of apical margin projecting slightly forward, basal margin straight; disc laterally with anterior threefourths convex, posterior fourth divergent, hind angle blunt, obtuse, projecting laterally; median groove shallow, not reaching the apical and basal margins; lateral bead broad. Elytra convex, matt, punctated, densely pubescent, with golden brown short setae; base narrowed, humerus rounded; apex without re-entrant angle; scutellum long, pointed; striations moderate, shallow, narrow, intervals broad, both striations and intervals covered by setae; sutural line narrow, widest at base, narrowing suddenly after the tip of scutellum, extending till apex.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A675FFE4ADD3FB45E2F49FAE.taxon	distribution	Distribution. INDIA: Jharkhand: Tetara; Maharashtra: Nagpur; Tamil Nadu: Chennai.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A674FFE0ADD3FC6FE4099949.taxon	description	Fig. 4	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A674FFE0ADD3FC6FE4099949.taxon	materials_examined	Type material. Holotype (male) labelled: “ Male; India: Kerala: Pattambi (KAU Campus, 10 ° 48 ’ 42.9 ” N 76 ° 11 ’ 25.7 ” E), ‘ Light’, 14. XI. 2016, coll. and det. S. V. Akhil ”, deposited in ZSIC; Paratypes labelled “ Female, India: Kerala: Pattambi (KAU Campus, 10 ° 48 ’ 42.9 ” N 76 ° 11 ’ 25.7 ” E), ‘ Light’, 14. XI. 2016, coll. and det. S. V. Akhil ”, deposited in ZSIC; labelled: “ Male; India: Kerala: Peruvannamoozhi (IISR station, 11 ° 36 ’ 26.0 ” N 75 ° 49 ’ 24.9 ” E), 02. ii. 2017, coll. V. A. Jithmon, det. S. V. Akhil ” deposited in ZSIC.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A674FFE0ADD3FC6FE4099949.taxon	description	Description: Length (TLA) 5.8 – 6.1 mm. Head, pronotum, scutellum and region around scutellum dark reddish brown; first two segments of antennae, palpi and legs pale reddish yellow (apical segments of palpi darker); femur, apex of tibiae and tarsi dark brown; antennal segments 3 – 4 dark reddish yellow, rest of the antennal segments reddish brown; lateral margin of pronotum and elytra black; eyes pitch black; genae, gula and prosternum dark reddish yellow; mesosternum, metasternum and abdominal ventrites brownish yellow. Head broad, wider than long, sparsely pubescent, smooth, with very faint wrinkles; frons with frontal foveae setose, shallow, densely punctated (mid region finely punctated, with short setae), smooth; vertex very faintly wrinkled, glabrous; neck rough, densely punctated, pubescent; labrum rectangular, transverse, anterior margin straight, apical angles almost at right angle, having six setae at anterior margin with two long setae at the corners, four (2 long, 2 intermediate) setae in the middle, rest glabrous. Clypeus sub-rectangular, anterior angles obtuse, with one long setae on each lateral edge, minute setae in the anterior margin; fronto-clypeal suture well impressed, deep, intended. Mentum transverse with two triangular, large lateral lobes which are toothed laterally, a single long seta present at the apex of lateral tooth, rest of the mentum glabrous; lateral lobes pointed apically; mentum without median tooth. Submentum very narrow, glabrous. Antennae long, slender, reaching beyond the middle of elytra; segment 1 longest, segment 2 elongate oval, all the segments densely pubescent with a ring of long setae in the apex of the segments. Palpi with segment 2 longest, wide, rectangular; the entire segment pubescent, penultimate segment long, slender, in the shape of an inverted cone, apical segment conical, tapering towards apex. Mandibles stout, scrobe plurisetose, tip pointed, arcuate, not crossing. Eyes highly protruding, prominent. Genae evidently wrinkled except at the base, a few setae present below eyes, rest of the genae glabrous. Gula large, widening towards pronotum, gular suture divergent, well impressed. Pronotum shiny, as long as wide, disc punctated, sparsely pubescent, smooth without wrinkles; disc apically sinuate without forward projection on lateral region of apical margin, disc straight at base, hind angles rounded, blunt, right angled; disc broad, laterally with anterior two-thirds convex and posterior third narrowed, with parallel sides; surface of disc convex; median groove deep, reaching both apical and basal margin; lateral bead narrow, widest near apical region. Elytra subparallel, slightly narrowed towards base, shiny, densely punctated and densely pubescent with short yellow setae; humerus prominent, corners rounded; apex obliquely truncate without any re-entrant angle; scutellum long, narrow, pointed, glabrous; striations well marked, deep, carinate; intervals broad, convex / carinate; suture not completely closed apically; setae present on striations as well as on intervals. Hind wings well developed. Legs strong, densely pubescent; protibial comb rounded with spur at the basal end of the comb. Two tibial spurs pointing inwards. Tarsi with article 1 longest, article 4 shortest. Ventrally, pronotum smooth, pubescent, punctate. Prosternal process prominent, pubescent, extending beyond procoxae, apex pointing downwards. Ventral region densely punctate, pubescent with golden brown setae; hind coxae contiguous; abdomen rounded till apex, apex pointed; mesepisternum prominent, long and broad. Sexual dimorphism. Symmetrically and diagonally arranged adhesive pads on male pro-tarsomeres 1, 2 and 3. Male genitalia. Median lobe (in lateral view) straight in the basal half, sharply curving downwards towards the apex, with apex blunt, pointing straight rather than downwards. Measurements. Holotype (male), TLA = 6.08 mm, TLB = 5.74 mm, PL = 1.10 mm, PW = 1.16 mm, EL = 3.60 mm, EW = 2.39 mm; Paratype (female), TLA = 5.81 mm, TLB = 5.74 mm, PL = 1.10 mm, PW = 1.18 mm, EL = 3.52 mm, EW = 2.48 mm.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A674FFE0ADD3FC6FE4099949.taxon	distribution	Distribution. INDIA: Kerala: Pattambi, Peruvannamoozhi. Collecting circumstances. Light-attracted, collected using low-intensity UV light trap.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A674FFE0ADD3FC6FE4099949.taxon	etymology	Etymology. Named after the host institution of the authors.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
6B0687C5A674FFE0ADD3FC6FE4099949.taxon	discussion	Remarks. This new species is similar to B. dryas but differs in having rounded, obliquely truncate elytral apex without re-entrant angle, deeply impressed elytral striations, hind angle of pronotum right angled, blunt, not projecting laterally and strongly protruding eyes.	en	Akhil, S. V., Divya, M., Sabu, K. Thomas (2020): Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India. Zootaxa 4816 (4): 576-600, DOI: 10.11646/zootaxa.4816.4.7
