taxonID	type	description	language	source
6A6487C2FFAE7C74FF5BFBD66559F975.taxon	materials_examined	Type locality. Malaysia, Borneo, Sabah Ulu Segama Forest Reserve, Danum Valley conservation area, 05 ° 73 N, 117 ° 37.61 E. Type material. The species was hitherto known only from the male holotype (Solodovnikov 2008) which here was not examined. Additional males and female reported here fully match the original description. Additional material examined. 3 ♂, 1 ♀: “ SABAH, Danum, Valley, B. R. l. [Borneo Rainforest lodge], f. i. t. [flight intercept trap] 14. – 16. II. 2007, G. De Rougemont ” (GROG, JJRC).	en	Janák, Jiří (2017): Four new species and a new record of Anchocerus from the Oriental region (Coleoptera: Staphylinidae: Staphylininae: Staphylinini: Acylophorina). Zootaxa 4319 (3): 579-589, DOI: 10.11646/zootaxa.4319.3.9
6A6487C2FFAE7C74FF5BFBD66559F975.taxon	description	Description. Detailed description and illustration of the holotype was published by Solodovnikov (2008). Habitus of the specimen from Borneo Rainforest lodge is illustrated as in Figs. 1 – 2 and male and female characters of specimens from this locality are illustrated as follows: aedeagus as in Figs. 5 – 8, male sternite 9 as in Fig. 3, male tergite 10 as in Fig. 4; female tergite 10 as in Fig. 9, female valvifer and poorly chitinised sclerite of female genital segment as in Fig. 10.	en	Janák, Jiří (2017): Four new species and a new record of Anchocerus from the Oriental region (Coleoptera: Staphylinidae: Staphylininae: Staphylinini: Acylophorina). Zootaxa 4319 (3): 579-589, DOI: 10.11646/zootaxa.4319.3.9
6A6487C2FFAE7C74FF5BFBD66559F975.taxon	distribution	Distribution. The species is known only from the Danum valley area in Sabah, Borneo. Bionomics. The species was collected by a flight intercept trap in the rainforest.	en	Janák, Jiří (2017): Four new species and a new record of Anchocerus from the Oriental region (Coleoptera: Staphylinidae: Staphylininae: Staphylinini: Acylophorina). Zootaxa 4319 (3): 579-589, DOI: 10.11646/zootaxa.4319.3.9
6A6487C2FFAE7C70FF5BF96A62F1FE85.taxon	materials_examined	Type locality. Indonesia, Bali, Lake Buayan, ca. 8 ° 15 ' S, 115 ° 08 ' E. Type material. Holotype ♂: INDONESIA, BALI: “ BALI, L. Buayan, 27. VII. 1981, de Rougemont ” (GROC). Description. Body length 9.1 mm, forebody length 4.1 mm (Fig. 12). Body black, mouthparts, legs and antennae brown, antennal segments 1 – 3 and 11 reddish yellow. Head (Fig. 13) almost as long as wide (R = 0.94), straight behind eyes. Temples longer than eyes (R = 1.28). Dorsal macrosetae (one side only): one near antennal base; one medially between eyes; one between posterior margin of eye and neck constriction, markedly closer to neck constriction than to posterior margin of eye; one medially near neck constriction; two laterally on temples of which one near posterior margin of eye and one near neck constriction (Fig. 13). Surface with sparse and fine nonsetiferous punctation. Antennae long, fifth segment markedly longer than wide (R = 1.58), tenth segment markedly transverse (R = 0.67). Mandibles with wide tooth (Fig. 13). Pronotum (Fig. 13) almost as long as wide (R = 0.95), longer (R = 1.40) and wider (R = 1.39) than head, with punctation markedly finer and sparser than that on head. Dorsal macrosetae (one side only): one medial on disc and three laterally in anterior third. Elytra with dense and coarse punctation, shorter than wide (R = 0.92), longer (R = 1.10) and wider (R = 1.14) than pronotum. Wings well developed. Scutellum with punctation and setae similar to those on elytra. Abdomen with coarse and moderately dense punctation, all tergites with dense transverse microsculpture. Legs densely covered with brown setae, all tibiae with lateral spines. Male. Sternite 9 (Fig. 14) long, moderately narrowed apically, with numerous long setae at apicolateral portion; tergite 10 (Fig. 15) narrowed apically, with several long setae situated at apical and lateral margins. Aedeagus 1.25 mm long (Figs. 16 – 19); median lobe moderately slender, narrow apically; paramere moderately wide, distinctly shorter than median lobe; sensory peg setae absent. Female. Unknown.	en	Janák, Jiří (2017): Four new species and a new record of Anchocerus from the Oriental region (Coleoptera: Staphylinidae: Staphylininae: Staphylinini: Acylophorina). Zootaxa 4319 (3): 579-589, DOI: 10.11646/zootaxa.4319.3.9
6A6487C2FFAE7C70FF5BF96A62F1FE85.taxon	diagnosis	Differential diagnosis. Anchocerus balianus sp. nov. can be inserted in a key (Solodovnikov 2008) near A. aparamerus and differs from this species by longer and finely punctured pronotum and by the different shape of the aedeagus with very wide paramere.	en	Janák, Jiří (2017): Four new species and a new record of Anchocerus from the Oriental region (Coleoptera: Staphylinidae: Staphylininae: Staphylinini: Acylophorina). Zootaxa 4319 (3): 579-589, DOI: 10.11646/zootaxa.4319.3.9
6A6487C2FFAE7C70FF5BF96A62F1FE85.taxon	etymology	Etymology. The specific epithet refers to the type locality.	en	Janák, Jiří (2017): Four new species and a new record of Anchocerus from the Oriental region (Coleoptera: Staphylinidae: Staphylininae: Staphylinini: Acylophorina). Zootaxa 4319 (3): 579-589, DOI: 10.11646/zootaxa.4319.3.9
6A6487C2FFAE7C70FF5BF96A62F1FE85.taxon	distribution	Distribution. The species is known only from Buayan Lake on Bali island. Bionomics. Nothing is known about the bionomics of the new species.	en	Janák, Jiří (2017): Four new species and a new record of Anchocerus from the Oriental region (Coleoptera: Staphylinidae: Staphylininae: Staphylinini: Acylophorina). Zootaxa 4319 (3): 579-589, DOI: 10.11646/zootaxa.4319.3.9
6A6487C2FFAA7C70FF5BFE006559F9AF.taxon	materials_examined	Type locality. Indonesia, Borneo, Sabah, Danum Valley, ca. 5 ° 01 ' 16 '' N, 117 ° 44 ' 48 '' E. Type material. Holotype ♂: INDONESIA, BORNEO: “ SABAH, Danum, Valley, B. R. l. [Borneo Rainforest lodge], f. i. t. [flight intercept trap] 14. – 16. II. 2007, G. De Rougemont ”, “ Holotypus Anchocerus differens sp. nov., J. Janák det. 2017 ” (GROG).	en	Janák, Jiří (2017): Four new species and a new record of Anchocerus from the Oriental region (Coleoptera: Staphylinidae: Staphylininae: Staphylinini: Acylophorina). Zootaxa 4319 (3): 579-589, DOI: 10.11646/zootaxa.4319.3.9
6A6487C2FFAA7C70FF5BFE006559F9AF.taxon	description	Description. Body length 8.6 mm, forebody length 3.4 mm (Fig. 20). Body black, mouthparts, legs and antennae brown, antennal segments 4 – 11 reddish yellow. Head (Fig. 21) markedly shorter than wide (R = 0.77), markedly narrowed behind eyes. Temples shorter than eyes (R = 0.72). Dorsal macrosetae (one side only): one near antennal base; one medially between eyes, markedly before maximal width of head; one postero-medially, near neck constriction; one posterolaterally near neck constriction; two laterally on temple, of which one near posterior margin of eye, one between posterior margin of eye and neck constriction. Surface with sparse and fine nonsetiferous punctation. Antennae moderately short, fifth segment about as long as wide (L / W = 0.96), tenth segment markedly transverse (R = 0.62). Mandibles very wide in basal half, then gradually narrowed apically, basal tooth only very slightly protruding, almost invisible (Fig. 21). Pronotum (Fig. 21) wider than long (R = 0.89), longer (R = 1.66) and wider (R = 1.45) than head, with punctation only slightly finer than those on head. Dorsal macrosetae (one side only): one medial and two lateral. Elytra with dense and coarse punctation, shorter than wide (R = 0.90), longer (R = 1.12) and wider (R = 1.10) than pronotum. Wings well developed. Scutellum with punctation and setae similar to those on elytra. Abdomen with coarse and moderately dense punctation, all tergites with dense transverse microsculpture. Legs densely covered with brown setae, all tibiae with lateral spines. Male. Sternite 9 unknown (damaged in holotype); tergite 10 (Fig. 22) narrowed apically, with several long setae situated at apical and lateral margins. Aedeagus 1.2 mm long (Figs. 23 – 26); median lobe moderately slender, narrowed apically; paramere moderately narrow, distinctly shorter than median lobe; sensory peg setae absent. Female. Unknown.	en	Janák, Jiří (2017): Four new species and a new record of Anchocerus from the Oriental region (Coleoptera: Staphylinidae: Staphylininae: Staphylinini: Acylophorina). Zootaxa 4319 (3): 579-589, DOI: 10.11646/zootaxa.4319.3.9
6A6487C2FFAA7C70FF5BFE006559F9AF.taxon	diagnosis	Differential diagnosis. Anchocerus differens sp. nov. differs from other Anchocerus species by the very wide basal part of mandibles, posteriorly narrowed temples and slender paramere lacking sensory peg setae. The new species can be inserted in a key (Solodovnikov 2008) near Anchocerus birmanus Fauvel, 1905.	en	Janák, Jiří (2017): Four new species and a new record of Anchocerus from the Oriental region (Coleoptera: Staphylinidae: Staphylininae: Staphylinini: Acylophorina). Zootaxa 4319 (3): 579-589, DOI: 10.11646/zootaxa.4319.3.9
6A6487C2FFAA7C70FF5BFE006559F9AF.taxon	etymology	Etymology. The specific epithet refers to the different shape of the mandibles and temples compared to other species of the genus.	en	Janák, Jiří (2017): Four new species and a new record of Anchocerus from the Oriental region (Coleoptera: Staphylinidae: Staphylininae: Staphylinini: Acylophorina). Zootaxa 4319 (3): 579-589, DOI: 10.11646/zootaxa.4319.3.9
6A6487C2FFAA7C70FF5BFE006559F9AF.taxon	distribution	Distribution. The species is distributed in the Danum valley, Sabah, Borneo. Bionomics. The species was collected by a flight intercept trap in the rainforest.	en	Janák, Jiří (2017): Four new species and a new record of Anchocerus from the Oriental region (Coleoptera: Staphylinidae: Staphylininae: Staphylinini: Acylophorina). Zootaxa 4319 (3): 579-589, DOI: 10.11646/zootaxa.4319.3.9
6A6487C2FFAA7C7DFF5BF9156559FC28.taxon	materials_examined	Type locality. Indonesia, Borneo, Sabah, Danum Valley, ca. 5 ° 01 ' 16 '' N, 117 ° 44 ' 48 '' E. Type material. Holotype ♂: INDONESIA, BORNEO: “ SABAH, Danum, Valley, B. R. l. [Borneo Rainforest lodge], f. i. t. [flight intercept trap] 14. – 16. II. 2007, G. De Rougemont ”, “ Holotypus Anchocerus rougemonti sp. nov., J. Janák det. 2017 ” (GROG). Paratype ♂: same data as the holotype, “ Paratypus Anchocerus rougemonti sp. nov., J. Janák det. 2017 ” (JJRC).	en	Janák, Jiří (2017): Four new species and a new record of Anchocerus from the Oriental region (Coleoptera: Staphylinidae: Staphylininae: Staphylinini: Acylophorina). Zootaxa 4319 (3): 579-589, DOI: 10.11646/zootaxa.4319.3.9
6A6487C2FFAA7C7DFF5BF9156559FC28.taxon	description	Description. Body length 10.3 (HT) – 10.7 (PT) mm, forebody length 4.3 (HT) – 4.5 (PT) mm (Fig. 27). Body black, mouthparts, legs and apical part of abdominal segments 7 and 8 reddish brown, tarsi and antennal segments 8 – 11 reddish yellow. Head (Fig. 28) markedly shorter than wide (R PT = 0.85, HT = 0.87), slightly widened behind eyes. Temples markedly longer than eyes (R HT = 1.15, PT = 1.18). Dorsal macrosetae (one side only): one near antennal base; one medially between eyes; one postero-medially, closer to neck constriction than to posterior margin of eye; one medially near neck constriction; two laterally on temple, of which one near posterior margin of eye, one near neck constriction. Surface with sparse and fine nonsetiferous punctation. Antennae slender, fifth segment about one and half as long as wide (R PT = 1.62, HT = 1.64), tenth segment markedly transverse (R PT = 0.74, HT = 0,76). Pronotum (Fig. 28) shorter than wide (R HT, PT = 0.89), longer (R HT = 1.22, PT 1.25) and wider (R HT = 1.19, PT = 1.20) than head, with punctation slightly finer than that on head. Dorsal macrosetae (one side only): one medial and two lateral. Elytra with dense and coarse punctation, shorter than wide (R PT = 0.88, HT = 0.92), longer (R HT = 1.10, PT = 1.11) and wider (R HT = 1.07, PT = 1.12) than pronotum. Wings well developed. Scutellum with punctation finer and setae shorter than those on elytra. Abdomen with coarse and moderately dense punctation, all tergites with dense transverse microsculpture. Legs densely covered with brown setae, all tibiae with lateral spines. Male. Sternite 9 (Fig. 29) long, moderately narrowed apically, with numerous long setae at apicolateral portion; tergite 10 (Fig. 30) narrowed apically, with several long setae situated at apical and lateral margins. Aedeagus 1.36 (PT) – 1.43 (HT) mm long (Figs. 31 – 36); median lobe wide, markedly narrowed apically; paramere long and relatively narrow, distinctly shorter than median lobe, with large elongate triangular orifice; with a few fine light tubercles visible in lateral view in holotype. Female. Unknown.	en	Janák, Jiří (2017): Four new species and a new record of Anchocerus from the Oriental region (Coleoptera: Staphylinidae: Staphylininae: Staphylinini: Acylophorina). Zootaxa 4319 (3): 579-589, DOI: 10.11646/zootaxa.4319.3.9
6A6487C2FFAA7C7DFF5BF9156559FC28.taxon	diagnosis	Differential diagnosis. Anchocerus rougemonti sp. nov. differs from other Anchocerus species by long and narrow teeth on mandibles. The new species can be inserted in a key (Solodovnikov 2008) near A. aparamerus and differs from this species by more coarse punctures on head and pronotum and a different shape of the paramere.	en	Janák, Jiří (2017): Four new species and a new record of Anchocerus from the Oriental region (Coleoptera: Staphylinidae: Staphylininae: Staphylinini: Acylophorina). Zootaxa 4319 (3): 579-589, DOI: 10.11646/zootaxa.4319.3.9
6A6487C2FFAA7C7DFF5BF9156559FC28.taxon	etymology	Etymology. This species is dedicated to Guillaume de Rougemont, collector of the holotype.	en	Janák, Jiří (2017): Four new species and a new record of Anchocerus from the Oriental region (Coleoptera: Staphylinidae: Staphylininae: Staphylinini: Acylophorina). Zootaxa 4319 (3): 579-589, DOI: 10.11646/zootaxa.4319.3.9
6A6487C2FFAA7C7DFF5BF9156559FC28.taxon	distribution	Distribution. The species is distributed in the Danum valley, Sabah, Borneo. Bionomics. The species was collected by a flight intercept trap in the rainforest.	en	Janák, Jiří (2017): Four new species and a new record of Anchocerus from the Oriental region (Coleoptera: Staphylinidae: Staphylininae: Staphylinini: Acylophorina). Zootaxa 4319 (3): 579-589, DOI: 10.11646/zootaxa.4319.3.9
6A6487C2FFA77C7FFF5BFBAB62F1FF18.taxon	materials_examined	Type locality. Thailand, Doi Inthanon, ca. 18 ° 35 ' N, 98 ° 29 ' E. Type material. Holotype ♂: THAILAND: “ THAILAND, Doi Inthanon, 15. III. 1982, G. de Rougemont ”, “ Anchocerus sp., det. 1988 G. de Rougemont ”, “ Holotypus Anchocerus thai sp. nov., J. Janák det. 2017 ” (GROG).	en	Janák, Jiří (2017): Four new species and a new record of Anchocerus from the Oriental region (Coleoptera: Staphylinidae: Staphylininae: Staphylinini: Acylophorina). Zootaxa 4319 (3): 579-589, DOI: 10.11646/zootaxa.4319.3.9
6A6487C2FFA77C7FFF5BFBAB62F1FF18.taxon	description	Description. Body length 12.4 mm, forebody length 5.4 mm (Fig. 37). Body black, basal and apical part of first segment of antenna, tarsi, knees and apical part of abdominal segment 7 and 8 reddish brown. Head (Fig. 38) markedly shorter than wide (R = 0.86), distinctly widened behind eyes. Temples longer than eyes (R = 1.24). Dorsal macrosetae (one side only): one near antennal base; one medially between eyes; one posteromedially, much closer to neck constriction than to posterior margin of eye; one medially near neck constriction; two laterally on temple, of which one near posterior margin of eye, one near neck constriction. Surface with sparse and fine nonsetiferous punctation. Antennae very slender, fifth segment about twice as long as wide (R = 2.16), tenth segment slightly transverse (R = 0.90). Pronotum (Fig. 38) shorter than wide (R = 0.88), longer (R = 1.41) and wider (R = 1.37) than head, with punctation slightly finer than those on head. Dorsal macrosetae (one side only): one medial and two or three lateral. Elytra with dense and coarse punctation, shorter than wide (R = 0.84), longer (R = 1.09) and wider (R = 1.15) than pronotum. Wings well developed. Scutellum with punctation and setae similar to those on elytra. Abdomen with coarse and moderately dense punctation, all tergites with dense transverse microsculpture. Legs densely covered with brown setae, all tibiae with lateral spines. Male. Sternite 9 (Fig. 39) long, moderately narrowed apically, with numerous long setae at apicolateral portion; tergite 10 (Fig. 40) narrowed apically, with several long setae situated at apical and lateral margins. Aedeagus 1.7 mm long (Figs 41 – 45); median lobe slender, gradually narrowed apically; paramere very wide, distinctly shorter than median lobe; with a few fine setae at apex, sensory peg setae absent. Female. Unknown.	en	Janák, Jiří (2017): Four new species and a new record of Anchocerus from the Oriental region (Coleoptera: Staphylinidae: Staphylininae: Staphylinini: Acylophorina). Zootaxa 4319 (3): 579-589, DOI: 10.11646/zootaxa.4319.3.9
6A6487C2FFA77C7FFF5BFBAB62F1FF18.taxon	diagnosis	Differential diagnosis. Anchocerus thai sp. nov. differs from other Anchocerus species by the wide paramere without sensory peg setae in combination with a markedly transverse head. The new species can be inserted in a key (Solodovnikov 2008) near Anchocerus birmanus Fauvel, 1905. From this species and from the geographically close Chinese species from Yunnan with similar body size and distinct punctation on head and pronotum (A. giganteus Hu & Li 2010, A. pengzhongi Hu & Li 2012, A. yunnanensis Hu & Li 2010) it differs by the wide paramere without peg setae and orifice. Anchocerus thai sp. nov. differs from A. thailandicus Solodovnikov 2008, also described from Thailand, by the distinct punctation on the head and pronotum and different shape of the paramere.	en	Janák, Jiří (2017): Four new species and a new record of Anchocerus from the Oriental region (Coleoptera: Staphylinidae: Staphylininae: Staphylinini: Acylophorina). Zootaxa 4319 (3): 579-589, DOI: 10.11646/zootaxa.4319.3.9
6A6487C2FFA77C7FFF5BFBAB62F1FF18.taxon	etymology	Etymology. The specific epithet refers to the type locality.	en	Janák, Jiří (2017): Four new species and a new record of Anchocerus from the Oriental region (Coleoptera: Staphylinidae: Staphylininae: Staphylinini: Acylophorina). Zootaxa 4319 (3): 579-589, DOI: 10.11646/zootaxa.4319.3.9
6A6487C2FFA77C7FFF5BFBAB62F1FF18.taxon	distribution	Distribution. The species is distributed in Thailand. Bionomics. Nothing is known about the bionomics of the new species.	en	Janák, Jiří (2017): Four new species and a new record of Anchocerus from the Oriental region (Coleoptera: Staphylinidae: Staphylininae: Staphylinini: Acylophorina). Zootaxa 4319 (3): 579-589, DOI: 10.11646/zootaxa.4319.3.9
