taxonID	type	description	language	source
6945604EED55FF87FF60FF21FF74A0EA.taxon	diagnosis	Diagnosis. Adults generally blackish-brown with yellowish thoracic and abdominal markings, and sometimes with pale markings on head. Antennae and legs usually pale yellow or yellowish brown. Media anterior in the hindwing invisible. Male gonocoxite 9 dome-like, much wider than long in the I. crassicornis group, while much longer than wide in the I. fulvostigmata group; inner side with a stylus near subapical portion, and usually bearing some bristle tufts. Pseudostyli (basal parts of gonapophyses 9) feebly sclerotized, generally narrow and foliate, closely approximated to each other in the middle. Parameres (gonocoxite-gonapophyses-gonostyli complex 10) unpaired, proximal portion flattened, with a slender distal projection. Gonarcus (fused gonocoxites 11) generally shield-like, sometimes with projections from its posterior surface. Endophallus short, mostly with some bristles or bristle tufts. Female sternum 7 broad, posteriorly produced more or less, with posterior margin truncate or concave; subgenital plate (fused gonocoxites 8) reduced into a small ovoid sclerite or much more modified into a diversely shaped plate.	en	Liu, Xingyue, Aspöck, Horst, Zhang, Weiwei, Aspöck, Ulrike (2012): New species of the snakefly genus Inocellia Schneider, 1843 (Raphidioptera: Inocelliidae) from Yunnan, China. Zootaxa 3298: 43-52, DOI: 10.5281/zenodo.211415
6945604EED55FF87FF60FF21FF74A0EA.taxon	distribution	Distribution. Europe (Austria, Czech Republic, European part of Russia, Finland, Germany, Netherlands, Poland, Sweden); Asia (Afghanistan, Armenia, Bhutan, China, India, Japan, Mongolia, North Korea, Pakistan, Russia Far East, Thailand).	en	Liu, Xingyue, Aspöck, Horst, Zhang, Weiwei, Aspöck, Ulrike (2012): New species of the snakefly genus Inocellia Schneider, 1843 (Raphidioptera: Inocelliidae) from Yunnan, China. Zootaxa 3298: 43-52, DOI: 10.5281/zenodo.211415
6945604EED55FF87FF60FF21FF74A0EA.taxon	discussion	Comments The heuristic value, as well as the conflicts of subdividing the genus Inocellia into the I. crassicornis and I. fulvostigmata groups have been discussed elsewhere (Liu et al. 2010 a, 2010 b, U. Aspöck et al. 2011).	en	Liu, Xingyue, Aspöck, Horst, Zhang, Weiwei, Aspöck, Ulrike (2012): New species of the snakefly genus Inocellia Schneider, 1843 (Raphidioptera: Inocelliidae) from Yunnan, China. Zootaxa 3298: 43-52, DOI: 10.5281/zenodo.211415
6945604EED56FF85FF60FC6FFEE4A6ED.taxon	diagnosis	Diagnosis. The appearance of the species is conspicuously characterized by black elements including head, thorax, wing venation, legs, and abdomen. The male is characterized in the genitalia by the short gonocoxite 9 with a rather short and cone-like stylus, the sub-quadrate gonarcus (fused gonocoxites 11) with ventromedial protrusion, and the endophallus with only one pair of tufts with bristles on dorsal surface.	en	Liu, Xingyue, Aspöck, Horst, Zhang, Weiwei, Aspöck, Ulrike (2012): New species of the snakefly genus Inocellia Schneider, 1843 (Raphidioptera: Inocelliidae) from Yunnan, China. Zootaxa 3298: 43-52, DOI: 10.5281/zenodo.211415
6945604EED56FF85FF60FC6FFEE4A6ED.taxon	description	Description. Male. Body length 7.7 – 8.3 mm; forewing length 6.7 – 7.7 mm, hindwing length 5.6 – 6.6 mm. Head (Fig. 1) sub-quadrate, black throughout. Antennal sclerite (torulus) and antennae blackish brown. Mouthparts black, mandibles with distal half reddish-brown. Thorax black throughout. Legs (Fig. 2) blackish brown with brownish setae; coxae, trochanters, and bases of femora yellow. Wings (Fig. 1) hyaline, pterostigma blackish brown, veins blackish brown. Anterior branch of Rs with one forked vein and one simple vein running to wing margin. Abdomen blackish brown; tergum of each pregenital segment medially with a small yellow spot, sternum of each pregenital segment with posterior yellow margin. Genital segments entirely blackish brown. Gonocoxite 9 (Figs. 8 – 10) dome-like, nearly semicircular, much wider than long; distal portion of inner side with a rather short stylus, median portion of the stylus slightly prominent dorsally, with numerous bristles. Pseudostylus (basal part of gonapophysis 9) (Fig. 9) paired and foliate. Fused parameres (complex of fused gonocoxites, gonapophyses, and gonostyli 10) (Figs. 9 – 11) small, proximal portion flattened, expanded, subtrapezoidal and concave ventrad, distal projection curved dorsad. Gonarcus (fused gonocoxites 11) (Figs. 9 – 11) shield-like, with dorsal margin feebly produced posteriorly; in caudal view subquadrate, with ventromedial portion produced into a subtriangular lobe. Endophallus (Figs. 9 – 10) short, distal portion medially with a pair of bristle tufts directed dorsad. Ectoproct (Fig. 8) subtrapezoidal with rounded posterodorsal corner in lateral view. Hypandrium internum small, lobes narrow. Female. Unknown.	en	Liu, Xingyue, Aspöck, Horst, Zhang, Weiwei, Aspöck, Ulrike (2012): New species of the snakefly genus Inocellia Schneider, 1843 (Raphidioptera: Inocelliidae) from Yunnan, China. Zootaxa 3298: 43-52, DOI: 10.5281/zenodo.211415
6945604EED56FF85FF60FC6FFEE4A6ED.taxon	materials_examined	Type material. Holotype 3, CHINA: “ Yunnan, Baoshan, along the road from Wayao to Laoyingshan [ca. 25 ° 26 ʹN, 99 ° 15 ʹE, 1800 m], 12. V. 2009, Liang Ding ” (CAU). Paratype 13, CHINA: “ Yunnan, Gaoligong mts., 25.22 N, 98.49 E, 1500 – 2500 m, 17 - 24. V. 1995, Vít Kubáň (HUAC).	en	Liu, Xingyue, Aspöck, Horst, Zhang, Weiwei, Aspöck, Ulrike (2012): New species of the snakefly genus Inocellia Schneider, 1843 (Raphidioptera: Inocelliidae) from Yunnan, China. Zootaxa 3298: 43-52, DOI: 10.5281/zenodo.211415
6945604EED56FF85FF60FC6FFEE4A6ED.taxon	distribution	Distribution. The new species is currently known only from the southern part of the Gaoligongshan Range.	en	Liu, Xingyue, Aspöck, Horst, Zhang, Weiwei, Aspöck, Ulrike (2012): New species of the snakefly genus Inocellia Schneider, 1843 (Raphidioptera: Inocelliidae) from Yunnan, China. Zootaxa 3298: 43-52, DOI: 10.5281/zenodo.211415
6945604EED56FF85FF60FC6FFEE4A6ED.taxon	etymology	Etymology. The specific epithet ‘ nigra’ refers to the predominately blackish coloration including head, thorax, wing venation, legs and abdomen. It is an adjective in the feminine nominative case from Latin niger, - a, - um = black.	en	Liu, Xingyue, Aspöck, Horst, Zhang, Weiwei, Aspöck, Ulrike (2012): New species of the snakefly genus Inocellia Schneider, 1843 (Raphidioptera: Inocelliidae) from Yunnan, China. Zootaxa 3298: 43-52, DOI: 10.5281/zenodo.211415
6945604EED56FF85FF60FC6FFEE4A6ED.taxon	discussion	Remarks. The new species belongs to the Inocellia crassicornis group and appears to be closely related to I. crassicornis by having a tiny stylus near the posterior margin of the inner side of male gonocoxite 9, however, it is distinguished from I. crassicornis in appearance by the dark legs. In I. crassicornis as well as the other Inocellia species, the legs are generally yellow. Considering the male genitalia, the new species differs from I. crassicornis by the subquadrate gonarcus in caudal view and the endophallus with only one pair of tufts. In I. crassicornis the gonarcus is subtrapezoidal in caudal view, and the endophallus has two pairs of tufts of bristles on both dorsal and ventral surfaces.	en	Liu, Xingyue, Aspöck, Horst, Zhang, Weiwei, Aspöck, Ulrike (2012): New species of the snakefly genus Inocellia Schneider, 1843 (Raphidioptera: Inocelliidae) from Yunnan, China. Zootaxa 3298: 43-52, DOI: 10.5281/zenodo.211415
6945604EED57FF8FFF60F9E3FD24A51B.taxon	diagnosis	Diagnosis. In appearance, the species is entirely black on head and thorax, and dorsally blackish brown with transverse yellowish bands on pregenital segments of the abdomen. The male is characterized in the genitalia by the elongated gonocoxite 9 with a conspicuous unguiform stylus and a subtriangular lobe posteroventrally to stylus. The female is characterized by the tergum 8 posteroventrally with a short rounded protrusion.	en	Liu, Xingyue, Aspöck, Horst, Zhang, Weiwei, Aspöck, Ulrike (2012): New species of the snakefly genus Inocellia Schneider, 1843 (Raphidioptera: Inocelliidae) from Yunnan, China. Zootaxa 3298: 43-52, DOI: 10.5281/zenodo.211415
6945604EED57FF8FFF60F9E3FD24A51B.taxon	description	Description. Male. Body length 10.0 mm; forewing length 8.2 mm, hindwing length 6.6 mm. Head (Fig. 3) subquadrate, black throughout. Antennal sclerite (torulus) and antennae blackish brown. Mouthparts black, mandibles with distal half reddish brown. Thorax black throughout. Legs yellow with yellowish setae. Wings (Fig. 3) hyaline, pterostigma blackish brown, veins blackish brown. Anterior branch of Rs with one forked vein and one simple vein running to wing margin. Abdomen blackish brown with venter much paler; joints of each pregenital segment transversely yellow. Genital segments entirely yellow. Tergum 9 (Figs. 12 – 13) approximately 1.5 times as long as sternum, anterior margin feebly incised medially, posterior margin nearly truncate. Sternum 9 (Figs. 12 – 14) arcuate, with anterior margin slightly prominent. Gonocoxite 9 (Figs. 12 – 16) broadly dome-like, much longer than width of its proximal portion, with apex acutely pointed in lateral view; stylus (gonostylus 9) produced from medial portion of inner side, unguiform and feebly curved ventrad; a rounded lobe posteroventral to the stylus; a bristle tuft anterior to stylus. Pseudostylus (basal part of gonapophysis 9) (Figs. 14 – 15) paired, feebly sclerotized, narrowly foliate. Fused parameres (complex of fused gonocoxites, gonapophyses, gonostyli 10) (Figs. 16, 18) flattened and subtrapezoidal on proximal portion, ventrally bearing a slender distal projection, which is slightly curved dorsad. Gonarcus (fused gonocoxites 11) (Figs. 15 – 17) shield-like, nearly trapezoidal in caudal view, medially prominent posteriad. Endophallus (Figs. 14 – 16) short, dorsally with a pair of short bristle tufts on proximal portion and a pair of long bristle tufts on feebly sclerotized areas of distal portion, ventrally with a short bristle tuft medially. A scabrous membranous structure is present near the proximodorsal portion of endophallus. Ectoproct (Fig. 12) in lateral view subquadrate. Hypandrium internum (Fig. 19) small with lateral lobes foliate and slightly sinuate posteriorly. Female. Forewing length 13.3 mm, hindwing length 11.5 mm. Sternum 7 (Figs. 20 – 21, 24) nearly trapezoidal in lateral view, posterior margin slightly produced. Tergum 8 (Figs. 20, 22, 24) posteroventrally with a short and round protrusion. Subgenital plate (fused gonocoxites 8) (Figs. 20 – 21, 24) separated into a broadly subquadrate sclerite anteriorly and a small ovoid sclerite posteriorly, which are connected by a grooved membrane. Atrium bursae (Figs. 20, 23) subtriangular, slightly sclerotized ventrad; sacculus bursae and receptaculum seminis damaged in female paratype, but well preserved in the female specimen from Mt. Meili (Fig. 23).	en	Liu, Xingyue, Aspöck, Horst, Zhang, Weiwei, Aspöck, Ulrike (2012): New species of the snakefly genus Inocellia Schneider, 1843 (Raphidioptera: Inocelliidae) from Yunnan, China. Zootaxa 3298: 43-52, DOI: 10.5281/zenodo.211415
6945604EED57FF8FFF60F9E3FD24A51B.taxon	materials_examined	Type material. Holotype 3, “ CHINA / Yunnan, Puer city, Jingdong county, Wuliangshan Nature Reserve, Gibbon Observation Station [1787 m], N 24 ° 21 ʹ 0 7 ʺ / E 100 ° 42 ʹ 12 ʺ, April, 2.2010, ZHANG Weiwei leg. ” (CAU). Paratype 1 ♀, same data as holotype (CAU). Other material. 1 ♀, CHINA: “ YUNNAN, 5. - 8. vii. 1996, 28 ° 06 ʹN 98 ° 54 ʹE, 2700 m, Hengduan mts-part MEILI, Vít Kubáň leg. ” (HUAC); 1 ♀, CHINA: Yunnan, Tengchong, Datang [25 ° 33 ʹN, 98 ° 53 ʹE, 1118 m] (CAU).	en	Liu, Xingyue, Aspöck, Horst, Zhang, Weiwei, Aspöck, Ulrike (2012): New species of the snakefly genus Inocellia Schneider, 1843 (Raphidioptera: Inocelliidae) from Yunnan, China. Zootaxa 3298: 43-52, DOI: 10.5281/zenodo.211415
6945604EED57FF8FFF60F9E3FD24A51B.taxon	distribution	Distribution. The new species is found to be distributed in three mountain ranges, i. e. Wuliangshan Range in central Yunnan, Gaoligongshan Range in western Yunnan, and Meili in northwestern Yunnan, all of which belong to the Hengduan orographic uplift.	en	Liu, Xingyue, Aspöck, Horst, Zhang, Weiwei, Aspöck, Ulrike (2012): New species of the snakefly genus Inocellia Schneider, 1843 (Raphidioptera: Inocelliidae) from Yunnan, China. Zootaxa 3298: 43-52, DOI: 10.5281/zenodo.211415
6945604EED57FF8FFF60F9E3FD24A51B.taxon	etymology	Etymology. The specific epithet ‘ yunnanica’ refers to the endemic distribution in Yunnan Province. It is an adjective in the feminine nominative case from Latin Yunnanicus, - a, - um = Yunnanian.	en	Liu, Xingyue, Aspöck, Horst, Zhang, Weiwei, Aspöck, Ulrike (2012): New species of the snakefly genus Inocellia Schneider, 1843 (Raphidioptera: Inocelliidae) from Yunnan, China. Zootaxa 3298: 43-52, DOI: 10.5281/zenodo.211415
6945604EED57FF8FFF60F9E3FD24A51B.taxon	discussion	Remarks. The new species belongs to the Inocellia fulvostigmata group and appears to be closely related to Inocellia longispina from Thailand by having similar male gonocoxite 9 with a conspicuous stylus and a subtriangular lobe. However, the new species differs from I. longispina by the elongated male gonocoxite 9, the unguiform stylus, and the endophallus with five tufts, while in I. longispina the male gonocoxite 9 is much shorter with digitiform stylus and the endophallus possesses only one pair of tufts on the dorsal surface. The female of the new species can be easily distinguished from the other Inocellia species by the tergum 8 with a posteroventral protrusion. Since we did not find male specimens of the new species from the Gaoligongshan and Meili ranges, the current identification of the two female specimens from these two localities was made tentatively; these specimens are not placed in the type series. The following biological notes pertain to the type specimens. These specimens were collected from a naturally dead tree (species unidentified) at the top of a mountain, where some large trees were cut down, leaving a relatively open area. The male was collected as a pupa sitting in a small hole when removing the tree bark in the afternoon (ca. 4: 00 pm). The female was collected as a freshly emerged adult on the bark of the same tree in the evening (ca. 9: 00 pm). Discussion Until a few years ago it was assumed that the east of Asia (where the southern boundary of the occurrence of Raphidioptera lies) harbors only one or two species of snakeflies (H. Aspöck et al. 1991). In recent years this view has been entirely changed. Surprisingly, several new species belonging to the family Inocelliidae had been discovered in the south of China and in the north of Thailand, respectively (Liu et al. 2010 a, b; U. Aspöck et al. 2011). Altogether nine species of Raphidioptera, all belonging to the family Inocelliidae, can be attributed to the Yunnanian centre: Inocellia digitiformis Liu, H. Aspöck, Yang & U. Aspöck, 2010 (China: Sichuan) Inocellia nigra sp. nov. (China: Yunnan)	en	Liu, Xingyue, Aspöck, Horst, Zhang, Weiwei, Aspöck, Ulrike (2012): New species of the snakefly genus Inocellia Schneider, 1843 (Raphidioptera: Inocelliidae) from Yunnan, China. Zootaxa 3298: 43-52, DOI: 10.5281/zenodo.211415
