taxonID	type	description	language	source
6909C857F200FF895390BB89FA2C3FD2.taxon	diagnosis	Diagnosis. Fore tarsus with tarsomeres 3 – 5 slightly flattened; first mid tarsomere with strong basal spine on ventral surface; hind coxa bare.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F200FF895390BB89FA2C3FD2.taxon	description	Description. Holotype male (Fig. 1 A) Head: eye densely setulose. Frontal vitta velvety black, pollinose. 18 fr. 1 pair of oc, proclinate and developed, surrounded by 5 less developed setulae. Fronto – orbital plate black, greyish pollinose. Parafacial bare, wide ventrally, greyish pollinose. Fronto – genal suture with brownish hairs and protruding anteriorly. Gena brownish pollinose and setulose. Scape black, 2 short dorsal setae reaching pedicel. Pedicel black pollinose, dorsal setae developed, one of this stronger. Postpedicel black, golden pilose, 2.5 times the length of pedicel. Arista black, with short pubescence, rays shorter than width of base of arista. Palpus black, filiform. Thorax: black; scutum without vittae. Acr without a pattern, irregular rows. Dc 2: 3. Pprn 2, developed, setulose area. Pra 2, seta closest to suture longest. Proepimeron setulose. Katespisternum setulose on ventral surface. Wing: with superior third, between C vein and superior half of R 2 + 3 vein darkened. Calypter brownish with margin darkened. Haltere dark brown. Legs: black with tarsi black and pulvilli brownish. Fore femur setulose on posterior surface with 1 row of long pv and d; ventral surface bare. Fore tibia with 1 strong pre – apical d. Fore tarsomeres 3 – 5 slightly flattened (Fig. 2 A). Mid femur slightly constricted on pre – apical ventral surface; 1 row of short ad; 1 row of strong av, irregular length; 1 row of strong pv and 1 row of p. Mid tibia on ventral surface slightly constricted on basal half area, densely setulose on apical half area; 1 developed pre – apical ad; 1 developed median a; 2 developed apical av and 1 v; 2 apical strong pv; 1 median p and 1 apical p. First mid tarsomere with strong basal spine on ventral surface (Fig. 2 B). Hind coxa bare on posterior margin. Hind femur on ventral surface without protuberance (Fig. 2 C); 1 row of ad, 4 stronger setae running towards dorsal surface (Fig. 2 C); 1 row of strong av, 2 longer and strong setae on apical half; posterior surface with a small tuft of setae on apical half. Hind tibia with 1 developed median d and 1 pre – apical; 1 long and developed sub – median ad; 2 median av, 1 apical (Fig. 2 C). Abdomen: elongate, brownish with long and developed setae. Sternite 1 densely setulose. Sternite 5 (Fig. 5 A). Terminalia (Fig. 6 A): epandrium wider than long, with strong setae on basal half; cercal plate elongate and pointed apically, with short weak setae along the plate, fused to mid – basal half; surstylus slightly articulated with epandrium, wide on base and curved apically, shaped as a comma in frontal and lateral views; bacilliform process spiral and simple. Female: unknown.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F200FF895390BB89FA2C3FD2.taxon	biology_ecology	Biology. Label information indicates that this species was found in decomposing matter. This species occurs in the Páramos in the Central Cordillera in Colombia. Nothing else is known about its habits or biology.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F200FF895390BB89FA2C3FD2.taxon	etymology	Etymology. Name in apposition. The species epithet refers to the Awá indigenous people, that inhabit the regions of northern Ecuador (provinces of Carchi and Sucumbios) and southern Colombia (particularly the departments of Nariño and Putumayo).	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F200FF895390BB89FA2C3FD2.taxon	materials_examined	Type material. Holotype male (CEUA). COLOMBIA: Antioquia. San José de la montaña. El Congo / 10 – 15 \ set \ 2011 / 3000 m. / L. Rios / VSR pez. Trampa 5. Paratypes. COLOMBIA. Antioquia: San José de la montaña. El Congo / 10 – 15 \ set \ 2011 / 3000 m. / L. Rios / T. VSR pez. Bosque laguna (1 male, CEUA. Voucher code DG 017); Caldas: Colombia, Manizales. Reserva Rio / Blanco. 2995 m. VSR pescado / Abril \ 2005 / Seminario de entomologia / CEUA (1 male, CEUA). The holotype is not completely in good condition because it was used to extract DNA. It was not macerated, but rather immersed in Proteinase K. The specimen has 1 mid leg, 2 posterior legs and the rest of the legs glued to a paper triangle. The code of the voucher is DG 017.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F200FF895390BB89FA2C3FD2.taxon	distribution	Distribution. Colombia: departments of Antioquia and Caldas (Central Cordillera, Andean region) (Fig. 8 A).	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F203FF895390BB08FABD3970.taxon	description	(Figs. 1 B, 2 D – E, 5 B, 6 B, 8 B)	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F203FF895390BB08FABD3970.taxon	diagnosis	Diagnosis. Fore tarsomeres 2 – 5 slightly flattened; hind femur on ventral surface without protuberance with 3 strong av on mid half and 2 pv on mid half; hind coxa setulose on posterior margin	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F203FF895390BB08FABD3970.taxon	description	Description. Holotype male (Fig. 1 B) Head: eye densely setulose. Frontal vitta velvety black, pollinose. 16 fr. 1 pair of oc, proclinate and developed, surrounded by 3 less developed setulae. Fronto – orbital plate black, greyish pollinose. Parafacial bare, wide ventrally, greyish pollinose. Gena brownish and setulose. Scape black, 1 short dorsal setae reaching pedicel. Pedicel black pollinose, dorsal setae developed, one is stronger. Postpedicel black, golden pilose, 2 times the length of pedicel. Arista black, with short pubescence, rays shorter than width of base of arista. Palpus black, filiform. Thorax: black; scutum without vittae. Acr 2 – 3: 3 – 4, irregular rows. Dc 2: 3. Pprn 2, developed, setulose area. Pra 2, seta closest to suture longest. Proepimeron setulose. Katespisternum setulose on ventral surface. Wing: brownish, with superior third, between C vein and superior half of R 2 + 3 vein darkened. Calypter whitish. Haltere brown and lightened base. Legs: black with tarsi black and pulvilli brownish. Fore femur setulose on posterior surface; ventral surface bare. Fore tibia with 1 strong pre – apical d, 1 sub – median and 1 apical p, 1 v. Fore tarsomeres 2 – 5 slightly flattened (Fig. 2 D). Mid femur lost. Hind coxa with 2 setulae on posterior margin. Hind femur on ventral surface without protuberance (Fig. 2 E); 1 row of ad, setae running towards dorsal surface (Fig. 2 E); 3 strong av on mid half; 2 pv on mid half. Hind tibia with 1 developed sub-median d and 1 pre – apical; 1 long and developed sub – median ad; 1 median av (Fig. 2 E). Abdomen: elongate, brown with long and developed setae. Sternite 1 densely setulose. Sternite 5 (Fig. 5 B). Terminalia (Fig. 6 B): epandrium almost as long as wide, projected downwards and covering part of the surstylus, short and weak setae on basal half, in the middle part of the epandrium and downwards with bifid projection next to the cercal plate; surstylus widened at the base and pointed at the apex, strongly merged with epandrium; cercal plate elongated and pointed apically, with short weak setae along the plate, fused from de second third; bacilliform absent. Female: unknown.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F203FF895390BB08FABD3970.taxon	biology_ecology	Biology. Label information indicates that it was found near decomposing matter, in the Páramo in the West Cordillera in Colombia. Nothing else is known about its habits or biology.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F203FF895390BB08FABD3970.taxon	etymology	Etymology. Name in apposition. The species epithet refers to the Barí indigenous people, sometimes called Motilones, that inhabit the forest of the Catatumbo river, between Colombia and Venezuela.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F203FF895390BB08FABD3970.taxon	materials_examined	Type material. Holotype male (CEUA). COLOMBIA: Antioquia. San José de la montaña. El Congo / 10 – 15 \ set \ 2011 / 3000 m. / L. Rios / VSR pez. Trampa 5 / Voucher DG 002. The holotype is not in perfect condition because it was used to extract DNA. It was not macerated, but rather immersed in Proteinase K. The individual has 1 foreleg and 1 posterior leg (without tarsi). The voucher code is DG 002.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F203FF895390BB08FABD3970.taxon	distribution	Distribution. Colombia: department of Antioquia (Central Cordillera, Andean region) (Fig. 8 B).	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F20DFF855390BFD0FBDA3FD2.taxon	description	(Figs. 1 C, 2 F – I, 5 C, 6 C, 8 C)	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F20DFF855390BFD0FBDA3FD2.taxon	diagnosis	Diagnosis: male dichoptic; fifth fore tarsomere flatten and darkened, tarsomeres 1 – 4 white – yellowish, second fore tarsomere with one short ventral spine; katepisternum with strong ventral spine; hind coxa with strong and long hook setae on ventral surface.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F20DFF855390BFD0FBDA3FD2.taxon	description	Description. Holotype male (Fig. 1 C) Head: dichoptic. Eye with sparse setulae. Frontal vitta velvety dark. 6 fr, posterior setae not developed. A single pair of oc proclinate and developed, surrounded by 4 less developed setulae. Fronto – orbital plate, parafacial and gena dark with golden pollinosity; 2 orb, 1 reclinate upper and 1 lateroclinate lower seta. Frontal plate with 2 series of irregular setulae on anterior area and 2 setae between orb. Parafacial bare. Scape dark – brown, 2 setae reaching pedicel. Pedicel dark, weak golden pollinosity, 1 dorsal strong seta and 6 less developed setae. Postpedicel with golden pollinosity, 3 times the length of pedicel. Arista dark with short pubescence. Palpus dark, spatulate. Thorax: dark; scutum without vittae. Acr 3: 3, irregular rows. Dc 2: 3. Pprn 2, developed and 1 weak. Pra 2, strong. Katerpisternum with tuft of right and strong ventral spine; proepimeron setulose. Subapical scutellar setae 1 pair. Discal scutellar setae 2. Wing: yellowish. Calypters brownish. Haltere brown with stem and base lighter in colour. Legs: dark with pulvilli yellowish; anterior leg brownish with the apex and tibia, fore tarsus whitish and yellowish pollinosity, ventral surface translucent, apical tarsomere darkened (Fig. 2 G). Fore femur with 1 series of short av setae; 1 series of long pv; 2 series of p, less developed than pv; 1 series of long pd. Fore tibia with 1 preapical d; 1 p and 1 pv, short. Fore tarsi bare on v surface, except for 1 short spine on the second tarsomere (Fig. 2 F); apical tarsomere with 2 long and curved d. Mid coxa with 6 long d and 1 v. Mid femur constricted on ventral preapical surface; 1 series of long ad (Fig. 2 H); 7 short and right av on apical third; 1 irregular series of long v, diminishing in length on apical half (Fig. 2 H); 1 series of short pv forming an apical ctenidium; 1 series of long and hook pd, setae on apical half developed with right apex. Mid tibia constricted weakly on basal v surface; 1 preapical ad; 1 a on apical third, 2 apical a; 1 apical av; 2 strong apical v; 1 median p and 2 apical. Hind coxa with 4 p, strong and long hook setae on ventral surface. Hind femur strongly curved (Fig. 2 I); preapical strong protuberance on v surface, conspicuous tuft of hooked av, v and pv setae (Fig. 2 I); 1 series of long ad towards d surface; ventral surface on basal half bare; 1 series of pv on basal half; 1 series of hooked p on basal half (Fig. 2 I). Hind tibia with 1 median d and 1 preapical d; 9 – 13 of strong a on apical half, 1 seta on apical third and 1 apical seta (Fig. 2 I); 2 – 3 median av and 1 apical; 2 apical v. Abdomen: trimaculate. Syntergite 1 + 2 with short lateral setae, setulose on apex. Sternite 1 bare. Sternite 5 (Fig. 5 C). Terminalia (Fig. 6 C): epandrium wider than long, with sparse and weak setae; cercal plate V – shaped with long setae on external margin; surstylus strongly fused with epandrium without visible articulation, long and right, inner basal region setulose with short setae, acute on apex, with lateral – apical projections or process with short setae; bacilliform process absent. Female: dichoptic, similar to male, except: Frontal vitta black velvety. 8 – 10 fr. Upper orb and lower orb reclinate. Row of short and irregular fro. Fronto – orbital plate shining black, without pollinosity. Parafacial greyish pollinose until arista, with short setae almost same length of fro and in the same line. Pedicel with 1 short seta. Postpedicel 3 times the length of pedicel. Mesonotum reddish brown. Intra – alar area, postsutural supra – alar area and lateral area black. Calypter yellowish. Haltere whitish with base yellowish. Legs with tibia orange on apex. Fore femur with short pv, increasing in length to apex; 1 row of pd. Fore tibia with 1 row of av. Mid femur with 1 row of short av diminishing in length on apical half; 1 row of short pv diminishing in length to apex. Mid tibia without constriction on ventral base; 1 apical v; 2 p on apical third. Hind femur with small constriction and flattened on anterior and posterior surfaces; 1 row of short ad, increasing in length on apical half; developed av on apical half; short p on apical base; ventral face bare. Hind tibia with 1 median seta. Abdomen not trimaculate reddish brown; fourth tergite with 1 pair of lateral setae. Terminalia: cercus longer than hypoproct, with long and sinuous setae on apex; hypoproct round and setulose, mainly on apical area, setae shorter than cercus setae; sternite 8 reduced to 2 separated plates with long setae, developed and sinuous; sternite 7 wider than long with setae on apical half, longer apical setae; sternite 6 bigger than sternite 7, with apical disorder setae. Spermathecae 2, semicircular and flattened apically.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F20DFF855390BFD0FBDA3FD2.taxon	biology_ecology	Biology. Label information indicates that this species occurs in highland forests in Costa Rica, above 2800 m. Nothing else is known about its habits or biology.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F20DFF855390BFD0FBDA3FD2.taxon	discussion	Comments. This species is similar to Fannia gilvitarsis Chillcott, 1961 because of the yellow fore tarsus and the distinct broad frontal vitta. Similar to Fannia arcuata Chillcott, 1961 because of broad frontal vitta. Differs in the combination of the remaining diagnostic characters. F. gilvitarsis and F. arcuata occur in Central America, in Mexico, 2300 – 3000 m (Chillcott 1961).	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F20DFF855390BFD0FBDA3FD2.taxon	etymology	Etymology. Name in apposition. The species epithet refers to the Boruca indigenous people, who once inhabited Guanacaste province in Costa Rica.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F20DFF855390BFD0FBDA3FD2.taxon	materials_examined	Type material. Holotype male (INBio). COSTA RICA: San José. Est. Cuerici, Send. El Carbon, 5 km / Este de Villa Mills, San josé, Costa / Rica 2800 m. 19 ENE 1996. B. / Gamboa, de Luz / L _ S 390300 _ 500300 # 6785 / Costa Rica INBio CRI 002 378780. Paratypes. COSTA RICA. San José: Est. Cuerici, Send. El Carbon, 5 km / Este de Villa Mills, San José, Costa / Rica 2800 m. 19 ENE 1996. B. / Gamboa, de Luz / L _ S 390300 _ 500300 # 6785 / Costa Rica INBio CRI 002 378781 (1 male, INBio); “ Idem ” / Costa Rica INBio CRI 002 378789 (1 male, DZUP); “ idem ” / Costa Rica INBio CRI 002 378782 (1 female, INBio); “ idem ” / 20 ENE 1996. A. Picado / “ idem ” # 6806 / Costa Rica INBio CRI 002 392809 (1 male, INBio); “ idem ” / Costa Rica INBio CRI 002 392820 (1 male, DZUP). The holotype is in good condition.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F20DFF855390BFD0FBDA3FD2.taxon	distribution	Distribution. Costa Rica: Guanacaste province (Guanacaste Cordillera) (Fig. 8 C).	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F20FFF845390BB08FED73F66.taxon	description	(Figs. 1 D, 2 J – K, 5 D, 6 D, 8 D)	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F20FFF845390BB08FED73F66.taxon	diagnosis	Diagnosis. Tiny palpi, as short as the basal third of the proboscidae; eye with short and sparse setulae; mid tibia with long pubescence on ventral surface, almost the same length as the tibia; cercal plate completely fused.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F20FFF845390BB08FED73F66.taxon	description	Description. Holotype male (Fig. 1 D) Head: eye with short and sparse setulae. Frontal vitta dark brown. 14 fr. 1 pair of oc, proclinate and developed and 2 posterior setae. Parafacial with silver pollinosity, gena thin and black. Parafacial bare. Scape black, 1 dorsal seta. Pedicel black with 1 strong seta and some short setae. Postpedicel dark brown, silver hairy, 2 times the length of pedicel. Arista dark brown, with short pubescence, rays shorter than width of base of arista. Tiny palpus, dark brown and spatulate (Fig. 2 J). Thorax: dark brown; scutum without vittae. Acr 2: 2, irregular rows. Dc 2: 3. Pprn 3, developed and surrounded by setulae. Pra 1. Proepimeron setulose, 1 conspicuous setae and setulose area. Katespisternum setulose. Wing: yellowish. Calypters and haltere brownish. Legs: coxae, femora and tibia brown; tarsi brown, pulvilli yellowish. Fore femur 1 row of pd and pv, p surface setulose, with thin and long setae. Fore tibia with 1 pre – apical d; 1 pv; 1 v. Mid femur with 1 row of p, 3 stronger on the apex; 2 rows of pv; 1 row of ad and av; short av setae forming a tuft on apex. Mid tibia slightly setulose on v apical half, 1 strong and long apical seta; 1 long apical a; 1 d on apical half; 3 ad on apical half; 4 p. Hind coxa with setula on posterior margin. Hind femur (Fig. 2 K) without pre – apical swelling on v surface; 3 apical ad, towards d surface; 1 row of av, longer on the apex; setulose on posterior surface. Hind tibia with 1 median and 1 pre – apical d; 3 ad; 2 av and 1 apical. Abdomen: dark brown, elongated. Sternite 1 setulose. Sternite 5 (Fig. 5 D). Terminalia (Fig. 6 D): epandrium square, on basal half with two developed setae; surstylus broad and concave internally, articulated with epandrium; cercal plate broad and concave, with fine and long projections from the middle of the plate, longer that surtylus. Bacilliform process present, width on base half and acute on apical half, shaped as a comma. Female: unknown.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F20FFF845390BB08FED73F66.taxon	biology_ecology	Biology. Label information indicates that this species occurs in forests in Costa Rica, 1000 – 2050 m. Nothing else is known about its habits or biology.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F20FFF845390BB08FED73F66.taxon	discussion	Comments. This species possibly belongs to the carbonaria group, subgroup minutipalpis (Chillcott 1961), because of their tiny palpi and cercal plate with ventral long process. Fannia minutipalpis (Stein), F. brevipalpis Chillcott, 1961, F. neopolychaeta Chillcott, 1961 and F. trogonifera Chillcott, 1961 also belong to this group and are found in the Holarctic region.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F20FFF845390BB08FED73F66.taxon	etymology	Etymology. Name in apposition. The species epithet refers to the Bribri indigenous people, who once inhabited the Puntarenas province in Costa Rica.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F20FFF845390BB08FED73F66.taxon	materials_examined	Type material. Holotype male (INBio). COSTA RICA: Send. ac. Pittier, Prov. Punta, Costa / Rica. 1800 – 2050 m. 16 – 17 ENE 1995 / M. A. Zumbado. L _ S _ 331800 _ 577400 # 6248 / Costa Rica INBIO CRI 002 354938. Paratypes. COSTA RICA. Puntarenas: La Amistad, sector Altamira, Cerro Biolley / A. C. Amistad, Prov. Punta, Costa Rica / 1800 m. Ene 1994, R. Delgado, L _ S 572400 _ 332700 # 2547 / Costa Rica INBIO CRI 001 854095 (1 male, INBio); San Luis, Monteverde, Prov. Punta / Costa Rica. 1000 – 1350 m. Mar 1994. Z. / Fuentes, L N 449250 _ 250850 # 2771 / Costa Rica INBIO CRI 001 756422 (1 male, INBio); Send ac. Pittier, Prov. Punt, Costa / Rica. 1800 – 2050 m. 16 – 17 ENE 1995 / M. Chinchilla, L _ S _ 331800 _ 577400 / # 5325 / Costa Rica INBIO CRI 002 147608 (1 male, DZUP); “ idem ” / # 6248 / Costa Rica INBIO CRI 002 354935 (1 male, DZUP); “ idem ” / # 6248 / Costa Rica INBIO CRI 002 354936 (1 male, INBio); Costa Rica, Prov. Puntarenas / Finca Cafrosa, Embalce, 800 m NO. / de Tigra, 1280 m. 9 – 10 FEB 1997 / E. Navarro. L _ S _ 317800 _ 596200 / # 45444 / Costa Rica INBIO CRI 002 490364 (1 male, INBio)	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F20FFF845390BB08FED73F66.taxon	distribution	Distribution. Costa Rica: Province Puntarenas (Talamanca Cordillera – Southern Costa Rica and Western Panama) (Fig. 8 D).	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F20EFF845390B89CFC973991.taxon	description	(Figs. 1 E, 3 A – E, 5 E, 6 E, 8 E)	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F20EFF845390B89CFC973991.taxon	diagnosis	Diagnosis. Vibrissa and subvibrissal setulae hook – like; fore coxa with strong spine on posterior margin; fore tarsomeres 2 – 3 flattened; mid coxa with spine on ventral surface; hind tibia without setae on antero – ventral surface; facial margin with strong anterior projection.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F20EFF845390B89CFC973991.taxon	description	Description. Holotype male (Fig. 1 E) Head: eye setulose. Frontal vitta dark brown. 18 fr. 1 pair of oc, proclinate and developed and 1 posterior seta. Parafacial golden pollinose, gena black with brownish pollinosity. Parafacial bare. Scape black, 2 dorsal setae. Pedicel black with strong and short hook setae. Postpedicel dark brown, golden hairy, 1.5 times the length of pedicel. Arista dark brown, with short pubescence, rays shorter than width of base of arista. Palpus dark brown, flat and filiform. Vibrissa and subvibrissal setulae hook – like (Fig. 3 A). Thorax: dark brown; scutum without vittae. Acr without a pattern, setulose area, irregular rows. Dc 2: 3. Pprn 3, well – developed and surrounded setulae. Pra 2, anterior one stronger. Proepimeron setulose. Katespisternum setulose. Wing: brownish. Calypters and haltere brownish. Legs: coxae, femora and tibia brown; tarsi brown, pulvilli yellowish. Fore coxa with a strong apical spine on posterior surface (Fig. 3 B). Fore femur 1 row of pd, p and pv surface setulose, with thin and long setae. Fore tibia with 1 pre – apical d; 2 short p and 1 av. Fore tarsomeres 2 – 3 flattened (Fig. 3 C). Mid coxa with 1 row of strong a and 1 strong spine on v surface. Mid femur setulose on p surface, short setae; 1 row of decreasing pv, long on basal half and shorter on apical half; a few short v. Mid tibia without constrictions on v surface on basal half, setulose on apical half, 1 strong and long apical v; 1 apical a; 1 ad; 1 d; 2 apical av; 2 d. Hind coxa with setulae on posterior margin. Hind femur without pre – apical swelling on v surface (Fig. 3 D); 1 row of ad, apical setae towards d surface (Fig. 3 D); 1 row of av (Fig. 3 D). Hind tibia with 1 median and 1 pre – apical d; 4 ad; 1 row of 8 av and 1 apical; 1 row of pv. Hind tarsus with conspicuous short setae (Fig. 3 E). Abdomen: dark brown, elongated. Sternite 1 setulose. Sternite 5 (Fig. 5 E). Terminalia (Fig. 6 E): epandrium wider than long with weakly setae on through; surstylus long width in the base and the same width until the apex, fusioned with epandrium, with short setae; cercal plate rounded and globose, plate fusioned until the middle half. Bacilliform process present and spiraled. Female: unknown.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F20EFF845390B89CFC973991.taxon	biology_ecology	Biology. Label information indicates that this species occurs in highland forests of Costa Rica, 2600 – 3000 m. Nothing else is known about its habits or biology.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F20EFF845390B89CFC973991.taxon	discussion	Comments. This species possibly belongs to the scalaris group and maybe is close to F. ciliatisima Chillcott, 1961 and Fannia columbiana Chillcott, 1961 because of the strong setae on fore and mid coxa. These species occur in the Holarctic region.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F20EFF845390B89CFC973991.taxon	etymology	Etymology. Name in apposition. The species epithet refers to the Huetares indigenous people (also known as Pacacuas or Guetares), who once inhabited the San Jose province in Costa Rica.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F20EFF845390B89CFC973991.taxon	materials_examined	Type material. Holotype male (INBio). COSTA RICA. Villa Mills, 3000 m. Prov. / San Jose, Costa Rica / Curso Tachinidae y / Syrphidae, 24 feb 1992 / L _ S 390000 _ 495000 / Costa Rica INBio CRI 000 408379. Paratype. COSTA RICA: San José: Est. Cuerici, Sendero al Mirador, 4.6 / km al E. de Villa Mills, San José / Costa Rica. 2640 – 2700 m. 25 NOV / 1995. A. Picado, de Luz / L _ S _ 389700 _ 499600 # 6795 / Costa Rica INBio CRI 002 391462 (1 male, INBio).	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F20EFF845390B89CFC973991.taxon	distribution	Distribution. Costa Rica: San Jose Province (Fig. 8 E).	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F208FF825390B9BCFBB83B99.taxon	description	(Figs. 1 F, 3 F, 5 F, 6 F, 8 F)	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F208FF825390B9BCFBB83B99.taxon	diagnosis	Diagnosis: Hyaline wing, calypters and haltere yellowish, hind coxa bare on posterior margin, hind femur with weakly protuberance on apical half, without tuft of setae, just 6 av more differentiated on apical third.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F208FF825390B9BCFBB83B99.taxon	description	Description. Holotype male (Fig. 1 F) Head: eye with sparse setulae. Frontal vitta velvety brownish. 15 fr. 1 pair of oc proclinate and developed, surrounded by 5 setulae. Fronto – orbital plate and parafacial width, with silver pollinosity, gena brown. Parafacial bare. Scape dark – brown, 2 setae. Pedicel dark with dorsal short setae. Postpedicel with golden pollinosity, 1.5 times the length of pedicel. Arista dark with some part yellowish, very short pubescence. Palpus dark, spatulate. Thorax: dark with silver pollinosity; scutum with 3 silvery vittae, along acr and dc. Acr irregular rows. Dc 2: 3. Pprn 4 developed. Pra 2, setae near to suture a little longer. Proepimeron setulose. Discal scutellar setulose. Wing: hyaline. Calypters and haltere yellowish. Legs: brownish with pulvilli brownish. Fore femur with 1 series of short av; pd, p and pv surface setulose; 2 series of p, less developed than pv; 1 series of long pd. Fore tibia with 1 pre – apical d; 1 pd and 1 v. Mid femur constricted on ventral preapical surface; 1 series of ad; short and right av on apical third; pv surface setulose with series of long pv, on apical third with short setae; 1 series of long p, setae on apical third more developed. Mid tibia constricted weakly on basal mid v surface; 1 mid a and 1 apical; 1 apical av; 1 pre – apical d, 1 v. Hind coxa bare on posterior margin. Hind femur with weakly protuberance on apical half, without tuft of setae, just 6 av more differentiated on apical third (Fig. 3 F); 1 series of ad towards to d surface; apical third on p surface bare. Hind tibia with 1 median d and 1 pre – apical d; 2 av, 1 mid ad (Fig. 3 F). Abdomen: Sternite 1 bare. Sternite 5 (Fig. 5 F). Terminalia (Fig. 6 F): epandrium wider than long, with sparse and weak setae; cercal plate width throws with a hollow on base and short setae around; surstylus strong and width, articulated with epandrium with visible articulation, acute on apex; bacilliform process strong, sclerotized and the same length of the surtstylus, curved on the middle third, comma – shape. Female: unknown.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F208FF825390B9BCFBB83B99.taxon	biology_ecology	Biology. Label information indicates that this species occurs above 4000 m in the Andes. Nothing else is known about its habits or biology.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F208FF825390B9BCFBB83B99.taxon	etymology	Etymology. Name in apposition. The species epithet refers to the Humahuaca indigenous people (also known as Omaguaca) that once inhabited the Andean highlands in northwestern Argentina, in the province of Jujuy.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F208FF825390B9BCFBB83B99.taxon	materials_examined	Type material. Holotype male (DZUP). ARGENTINA: Jujuy / El Quemado. Ruta 52 km 38 / S 23.69390 ° / W 0 65.64913. 4150 m. / 17. i. 09 / L. R. Faria Jr. Leg. The holotype is not in perfect conditions because it was used in molecular study. It was not macerated, but rather immersed in Proteinase K. The main characters are visible, and the genitalia is in good condition. The voucher code is DG 026.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F208FF825390B9BCFBB83B99.taxon	distribution	Distribution. Argentina: province of Jujuy (Andean highlands) (Fig. 8 F).	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F208FF815390BCC5FC6C3DEE.taxon	description	(Figs. 1 G, 3 G – I, 5 G, 6 G, 8 G)	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F208FF815390BCC5FC6C3DEE.taxon	diagnosis	Diagnosis. Fore tibia with modified apical ventral seta; fore tarsomeres 3 – 5 slightly flattened; mid tibia with one modified posteroventral seta; hind coxa bare on posterior margin; hind femur strongly curved on ventral surface.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F208FF815390BCC5FC6C3DEE.taxon	description	Description. Holotype male (Fig. 1 G) Head: eye setulose. Frontal vitta dark brown. 18 fr. 1 pair of oc, proclinate and developed, surrounded by 5 setulae. Parafacial bare, silver and golden pollinosity, gena black with brownish polinosity. Scape black, 2 dorsal setae. Pedicel black with strong and short setae. Postpedicel dark brown, golden hairy, 2 times the length of pedicel. Arista dark brown, with short pubescence, rays shorter than width of base of arista. Palpus dark brown, right. Thorax: dark brown; scutum without vittae. Acr 2 – 3: 3, irregular rows. Dc 2: 3. Pprn 3, developed and surrounded by setulae. Pra 2. Proepimeron setulose. Katespisternum setulose ventrally, long setulae. Wing: brownish, with superior third, between C vein and superior half of vein R 2 + 3 darkened. Calypter whitish, outer margin yellowish. Haltere brownish. Legs: coxae, femora and tibia brown; tarsi brown, pulvilli yellowish. Fore femur 1 row of pd, p and pv surface setulose, with thin and long setae. Fore tibia with 1 apical and modified v (Fig. 3 G) and 1 short; 2 short pd on basal half. Fore tarsus with tarsomeres slightly flattened (Fig. 3 H). Mid femur setulose on p surface, short setae; 1 row of unequal av, long on basal half and shorter on apical half; 2 rows of long pv and v, a few short apical hooked setae on v surface. Mid tibia constricted on v surface on basal half and highly setulose on apical half, 1 strong and long apical setae; 1 long sub – median a; 2 apical av; 1 pre – apical ad; 1 sub – median and 1 pre – apical p; 1 modified (truncated) pv. Hind coxa without setulae on posterior margin. Hind femur curved with pre – apical swelling on v surface, tuft of long and curved setae on the swelling (Fig. 3 I); conspicuous swelling on p surface with tuft of straight setae shorter than on v surface. Hind tibia with 1 median and 1 pre – apical d; 1 sub – median ad; 1 row of 5 av and 1 apical; 1 apical v. Abdomen: dark brown, elongated. Sternite 1 setulose. Sternite 5 (Fig. 5 G). Terminalia (Fig. 6 G): epandrium longer than wide, on basal half with long but fine setae; surstylus strong, broad on base and the apex strongly hooked, articulated with epandrium and with short setae; cercal plate pointed on apex, plate fusioned until basal third, setulose apically; baciliform process small, spiral and simple. Female: unknown.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F208FF815390BCC5FC6C3DEE.taxon	biology_ecology	Biology. Label information indicates that this species occurs in highland forests of Costa Rica, above 2600 m. Nothing else is known about its habits or biology.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F208FF815390BCC5FC6C3DEE.taxon	etymology	Etymology. Name in apposition. The species epithet refers to the Kabekwa indigenous people, who once inhabited the provinces of Cartago and Limón.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F208FF815390BCC5FC6C3DEE.taxon	materials_examined	Type material. Holotype male (INBio). COSTA RICA: Prov. Limón: Cerros / Tararia. 2659 m. 23 – 25. ABR. 1997 \ R. Villalobos \ L _ S 344600 _ 576700. # 46221 / INBIO CRI 002 504535. Paratype. COSTA RICA. Cartago: Rio Macho, La \ Esperanza del Guarco. 3 Km carret. \ Interam (KM 61). Camino ICE 2750 m \ 1 – 2 JUN 1998. M. A. Zumbado \ LN _ 185000 _ 550700 # 50734 (1 male, INBio).	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F208FF815390BCC5FC6C3DEE.taxon	distribution	Distribution. Costa Rica: Provinces of Limón and Cartago (Fig. 8 G).	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F20BFF805390BD14FE7D3A12.taxon	description	(Figs. 1 H, 3 J – L, 5 H, 6 H, 8 H)	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F20BFF805390BD14FE7D3A12.taxon	diagnosis	Diagnosis. Fore tarsomere 1 with 1 broad leaf – like spine at the tip of posterior surface and another tiny leaf – like spine; tarsomeres 3 – 5 flattened; tarsomere 1 and 2 yellowish on apical half.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F20BFF805390BD14FE7D3A12.taxon	description	Description. Holotype male (Fig. 1 H). Head: eye densely setulose. Frontal vitta velvety black, pollinose. 15 fr. Ocellar triangle black; oc proclinate and developed, 5 less developed setulae on ocellar area. Fronto – orbital plate black and pollinose. Parafacial bare, narrowing ventrally, golden pollinose. Fronto – genal suture and gena golden pollinose, gena with long setae. Scape black, 3 setae reaching pedicel. Pedicel black pollinose, dorsal setae developed, one much longer than the others. Postpedicel black, golden pilose, 2 times the length of pedicel. Arista black, with short pubescence, rays shorter than width of base of arista. Palpus black and slightly spatulate. Thorax: black. Scutum without vitae. Acr 2: 3 – 4, not arranged in rows. Dc 2: 3, long, posterior – most postsutural pair developed. Pprn 2, developed. Pra 2. Pro – epimeron setulose. Wing: brownish, with upper third between vein C and half of vein R 2 + 3, and cells br, bm and cup darker. Calypters whitish. Haltere brown with stem and base lighter in colour. Legs: black, pulvilli brownish and tarsi black, except for fore tarsomere 1 and 2 with apical half yellowish. Fore femur setulose on posterior surface with 1 row of long pv and d; ventral surface bare. Fore tibia with 1 strong pre – apical d; 1 apical pv; 1 apical v. Fore tarsomere 1 with 1 broad leaf – like spine at the tip of posterior surface and another tiny leaf – like spine (Fig. 3 J); tarsomeres 3 – 5 flattened (Fig. 3 K). Mid femur with 1 row of decreasing av from base to apex; 2 rows of long pv with small tuft of setae with hooked apex on apical third. Mid tibia with 2 pronounced constrictions on ventral surface on basal half area, densely setulose on entire surface, 1 v; 1 developed pre – apical ad; 1 median a and 1 apical; 1 apical av; 1 apical modified pv; 2 pd; (Fig. 3 L). Hind coxa with setae on posterior margin. Hind femur on ventral surface with strong pre – apical protuberance and tuft of long setae on the protuberance; 1 row of ad, 4 stronger setae running towards dorsal surface; posterior surface with strong protuberance on median third (Fig. 3 M – N). Hind tibia with 1 median d and 1 pre – apical; 1 median ad; 3 av on apical half and 1 apical; anterior surface with short setae and 1 short apical; 2 strong median av, 1 apical (Fig. 3 M – N). Abdomen: elongate, black, weakly greyish pollinose with developed setae. Sternite 1 densely setulose. Sternite 5 (Fig. 5 H). Terminalia (Fig. 6 H): epandrium wider than long, strong setae on basal half, short apical internal expansions; cercal plate elongate, globose, apically wide, short weak setae on the plate; surstylus articulates with epandrium, curved apically, shaped as a comma in lateral view, basal half with internal basal expansion; bacilliform process spiral and simple. Female: unknown.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F20BFF805390BD14FE7D3A12.taxon	biology_ecology	Biology. Label information indicates that this species occurs in the Andean highlands, above 3000 m, from southern Ecuador to the Central Cordillera in Colombia. Nothing else is known about its habits or biology.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F20BFF805390BD14FE7D3A12.taxon	discussion	Comments. Fannia palta sp. nov. is morphologically similar to Fannia albitarsis Stein, 1911, in the first fore tarsomere having 1 broad leaf – like spine at the tip of posterior surface but differ in the fore tarsi that are not strong, flattened and yellowish. Fannia palta sp. nov. has fore tarsomeres 1 and 2 with apical half yellowish and the remainder brownish, tarsomeres 3 – 5 slightly flattened.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F20BFF805390BD14FE7D3A12.taxon	etymology	Etymology. Name in apposition. The species epithet refers to the Palta indigenous people that once inhabited the Province of Loja in the pre-Incan era.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F20BFF805390BD14FE7D3A12.taxon	materials_examined	Type material. Holotype male (UTPL). ECUADOR: 13155 – (EC) Loja: Cajanuma / Podocarpus National Park, trail Oso / de Anteojos, 3000 m, 27 / 02 / 2009 – 5 / 03 / 2009 (YPT), leg. Marc Pollet & / Anja De Braekeleer, sorted by Marc Pollet – sample code: EC / 2009 – 37 / MP & ACB – 011. Paratype. COLOMBIA. Antioquia: San José de la montaña / El Congo. 10 – 15 \ set \ 2011 / 3000 m. L. Rios / VSR pez trampa 5 (1 male, CEUA). Voucher code DG 013. The abdomen of holotype was taken to dissect the terminalia and it was fixed with the holotype. The paratype was used as voucher in a molecular study of Fanniidae (Voucher code DG 013).	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F20BFF805390BD14FE7D3A12.taxon	distribution	Distribution. Colombia: Antioquia department (Central Cordillera, Andean region); Ecuador: Loja province (Andean region) (Fig. 8 H).	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F20AFF9F5390BD48FC003B56.taxon	description	(Figs. 1 I, 3 O – P, 5 I, 7 A, 8 I)	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F20AFF9F5390BD48FC003B56.taxon	diagnosis	Diagnosis. Wing brownish, with upper third darker, fore tarsomeres 3 – 5 flattened, hind coxa bare on posterior margin, hind femur strongly curved, ventral surface with strong pre – apical protuberance extending over anterior and posterior surfaces.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F20AFF9F5390BD48FC003B56.taxon	description	Description. Holotype male (Fig. 1 I) Head: eye densely setulose. Frontal vitta velvety black, brownish pollinose. 15 fr. 1 pair of oc proclinate and developed, 5 less developed setulae on ocellar area. Fronto – orbital plate black, greyish pollinose. Parafacial bare, narrowing ventrally, greyish pollinose. Oral margin projecting anteriorly. Fronto – genal suture with brownish hairs. Gena black, yellowish pollinose. Scape black, 3 – 4 short setae reaching pedicel. Pedicel black pollinose, dorsal setae developed. Postpedicel black, brownish pilose, 2 times the length of pedicel. Arista black, with short pubescence, rays shorter than width of base of arista. Palpus black and filiform. Thorax: black. Scutum without vittae. Acr 2: 3 – 4, not arranged in rows. Dc 2: 3, long, posterior most postsutural pair developed. Pprn 4, 3 developed and 1 weak. Pra 2, strong. Pro – epimeron setulose. Pre – basal scutellar area with ground setulae. Wing: brownish, with upper third between C vein and half of R 2 + 3 vein, and cells br, bm and cup darker. Legs: black with tarsi black and pulvilli brownish. Fore femur with 1 row of long d; a and av inconspicuous; ventral surface bare; 1 row of long pv with straight apices; posterior surface setulose, with long and weak setae with curved apices. Fore tibia with 1 strong pre – apical d; 2 short ad; 1 apical pv; 1 median p and 1 apical; 1 apical v. Fore tarsus with 1 strong and thin v on first tarsomere; tarsomeres 3 – 5 flattened (Fig. 3 O). Mid femur constricted on pre – apical ventral surface; 1 row of short ad; 1 row of strong av that decrease in length towards apex; 1 row of long pv with hooked apices, setae decrease in length towards apex; 1 row of weak p with 6 longer on apical third. Mid tibia on ventral surface with pronounced constriction on basal half area, densely setulose on apical half, 1 long and strong apical v; 1 developed pre – apical ad; 1 developed a on mid apical half and 1 apical; 1 developed apical av; 1 apical pv; 2 p. Hind coxa bare on posterior margin. Hind femur strongly curved; ventral surface with strong pre – apical protuberance extending over anterior and posterior surfaces (Fig. 3 P); 1 row of ad, 4 setae running towards dorsal surface (Fig. 3 P); anteroventral surface setulose, with setae that increase in length to pre – apical protuberance; protuberance with tuft of long setae with hooked apices; posteroventral and posterior surfaces weakly setulose. Hind tibia with 1 developed median d and 1 pre – apical; 1 long and developed median ad at the same level of d; 1 short apical a; 1 median av, 1 apical. Abdomen: elongate, black, weakly greyish pollinose and long and with developed setae. Sternite 1 densely setulose. Sternite 5 (Fig. 5 I). Terminalia (Fig. 7 A): epandrium wider than long, with short apical internal projections and a few fine setae on apical half; cercal plate elongates apically, with short weak setae mainly on the basal third; surstylus articulated with epandrium, curved apically, shaped as a comma in frontal and lateral views, basal half broad; bacilliform process spiral and simple. Female: unknown.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F20AFF9F5390BD48FC003B56.taxon	biology_ecology	Biology. Label information indicates that this species occurs in the Andes in southern Ecuador, above 3000 m. Nothing else is known about its habits or biology.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F20AFF9F5390BD48FC003B56.taxon	discussion	Comments. Fannia polleti sp. nov. is morphologically similar to Fannia chingaza Grisales, Wolff & Carvalho, 2012 from Colombia. Fannia polleti sp. nov differs on the hind femur (with a tuft of more developed setae on preapical protuberance in F. chingaza) and the fore tarsomere (less flat in F. chingaz a) and the shape of the tip of the cercal plate (square in F. chingaz a).	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F20AFF9F5390BD48FC003B56.taxon	etymology	Etymology. Name in apposition. The species epithet is in homage to Marc Pollet, a researcher who collected a lot of interesting Diptera material in and near Podocarpus National Park (southern Andes, Ecuador) in 2009.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F20AFF9F5390BD48FC003B56.taxon	materials_examined	Type material. Holotype male (UTPL). ECUADOR: 13115 – (EC) [Ecuador] Loja: Cajanuma \ Podocarpus National Park, trail Oso \ de anteojos, 3000 m, 20 / 02 / 2009 – 27 / 02 / 2009 (YPT), leg. Marc Pollet & \ Anja De Braekeleer, sorted by Marc \ Pollet – Sample code: EC / 2009 – 34 MP & ADB – 0 12. Paratypes. ECUADOR. Loja: 13114 – (EC) [Ecuador] Loja: Cajanuma \ Podocarpus National Park, trail Oso \ de anteojos, 3000 m, 20 / 02 / 2009 – 27 / 02 / 2009 (YPT), leg. Marc Pollet & \ Anja De Braekeleer, sorted by Marc \ Pollet – Sample code: EC / 2009 – 34 / MP & ADB – 0 11 (1 male, UTPL); 13170 / “ idem ” / 5 / 03 / 2009 (MT) / “ idem ” / EC / 2009 – 37 / MP & ADB – 0 13 (1 male, DZUP); 13065 / “ idem ” / 20 / 02 / 2009 (YTP) / EC / 2009 – \ 31 / MP & ADB – 0 0 3 (1 male, UTPL); 13145 / “ idem ” / 5 / 03 / 2009 (MT) / “ idem ” / EC / 2009 – 37 / MP & ADB – 0 0 1 (3 males, UTPL); 13074 / “ idem ” / 20 / 02 / 2009 (YTP) / “ idem ” / EC / 2009 – 31 / MP & ADB – 0 12 (1 male, UTPL); 13112 / “ idem ” / 27 / 02 / 2009 (BPT) / “ idem ” / EC / 2009 – 34 / MP & ADB – 0 0 9 (1 male, UTPL); 13146 / “ idem ” / 27 / 02 / 2009 – 5 / 03 / 2009 (YPT) / “ idem ” / EC / 2009 – 37 / MP & ADB – 0 0 2 (2 males, DZUP); 13064 / “ idem ” / 16 / 02 / 2009 – 20 / 02 / 2009 (YPT) / “ idem ” / EC / 2009 – 31 / MP & ADB – 0 0 2 (2 males, DZUP). The holotype is in good condition and has all structures.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F20AFF9F5390BD48FC003B56.taxon	distribution	Distribution. Ecuador: province of Loja (Andean region) (Fig. 8 I).	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F215FF9D5390BCB7FC093AA1.taxon	description	(Figs. 1 J, 4 A – C, 5 J, 7 B, 8 J)	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F215FF9D5390BCB7FC093AA1.taxon	diagnosis	Diagnosis. Fore tarsomeres 2 – 5 flattened and tarsomeres 2 – 3 laterally expanded; fore tarsomere 1 yellowish, 2 – 3 partially yellowish; fronto – orbital plate, frontal vitta and facial ridge strongly protruding; gena with a shallow groove under the eye.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F215FF9D5390BCB7FC093AA1.taxon	description	Description. Holotype male (Fig. 1 J) Head: eye densely setulose. Frontal vitta velvety black, pollinose. 14 – 15 fr. 1 pair of oc proclinate and developed, 5 less developed setulae on ocellar area. Fronto – orbital plate silver and pollinose. Parafacial bare, golden pollinose and wide, almost the same width along. Fronto – orbital plate, frontal vitta and facial ridge strongly protruding (Fig. 4 A). Fronto – genal suture and gena golden pollinose, gena with a shallow groove under the eye (Fig. 4 B). Scape black, 3 – 4 setae reaching pedicel. Pedicel black pollinose with a dorsal seta developed and surrounded minor setae. Postpedicel black, golden pilose, 2 times the length of pedicel. Arista black, with short pubescence, rays shorter than width of base of arista. Palpus black and slightly spatulate. Prementum long and sclerotized. Thorax: black. Scutum without vittae. Acr 3: 3 not arranged in rows. Dc 2: 3, long. Pprn 4, developed. Pra 2. Pro – epimeron setulose. Wing: brownish, with upper third between C vein and half of R 2 + 3 vein slightly darker. Calypters brownish with border darker. Haltere brown with stem and base lighter in colour. Legs: black, pulvilli yellowish and tarsi black, except for first fore tarsomere completely yellowish and tarsomeres 2 and 3 partially yellowish (Fig. 1 J). Fore femur setulose on posterior surface with 1 row of long pv and d. Fore tibia with 2 sub – median av and 1 stronger sub – apical; 1 apical p; 1 pd. Fore tarsomere 1 short with a little tuft of short basal v; tarsomeres 2 – 5 flattened and tarsomeres 2 – 3 expanded laterally. Mid femur with 2 rows of long pv, strong setae on apical third and 1 row of av. Mid tibia on ventral surface with pronounced constriction on basal half area, densely setulose on apical half, 2 apical v; 3 median a; 1 ad; 4 median p and 1 apical; 1 apical pv. Hind coxa bare setae on posterior margin. Hind femur on ventral surface with a strong almost median protuberance, with tuft of setae and some of them stronger and hook – like, apical half setulose; av, a and ad surfaces with strong sparse setae (Fig. 4 C); pv surface setulose. Hind tibia with 1 median d; 2 median ad; 1 a; 2 median av and 1 apical (Fig. 4 C); p surface slightly setulose on apical half. Abdomen: elongate, black, weakly greyish pollinose with developed setae. Sternite 1 densely setulose. Sternite 5 (Fig. 5 J). Terminalia (Fig. 7 B): epandrium longer as wide with strong and stout setae; cercal plate square apically, fused on the middle portion; surstylus strong and articulated with epandrium, wide apically; bacilliform process spiral. Female: unknown.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F215FF9D5390BCB7FC093AA1.taxon	biology_ecology	Biology. Label information indicates that this species occurs in southern Ecuador, in the Andes above 2200 m. Nothing else is known about its habits or biology.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F215FF9D5390BCB7FC093AA1.taxon	discussion	Comments. Fannia quillacingas sp. nov. resembles the anthracina species group of Fannia, mainly in the characters of the fore tarsomeres (expanded, flattened and yellowish). This new species differs in the extent of the yellowish color in these tarsomeres and in the lateral expansion.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F215FF9D5390BCB7FC093AA1.taxon	etymology	Etymology. Name in apposition. The species epithet refers to the Quillacingas indigenous people, who once inhabited the higher elevations between Colombia and Ecuador.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F215FF9D5390BCB7FC093AA1.taxon	materials_examined	Type material. Holotype male (MZSP). ECUADOR: Girón, Portete de / Tarqui, 2200 – 2800 m. / Ecuador, XII. 1970 / L. E. Peña col. / MZSP. The abdomen is absent in the holotype because it was taken to dissect the terminalia and was fixed under the holotype.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F215FF9D5390BCB7FC093AA1.taxon	distribution	Distribution. Ecuador: Azuay province (Andean region) (Fig. 8 J).	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F217FF9C5390BDDDFC883852.taxon	description	(Figs. 1 K, 4 D – F, 5 K, 7 C, 8 K)	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F217FF9C5390BDDDFC883852.taxon	diagnosis	Diagnosis. Partially white yellowish fore tarsomeres, except for the fifth tarsomere darkened and flattened; fore second tarsomere on ventral surface with a tuft of short whitish setae, forming a comb and almost in entire tarsomere; ventral katepisternal spine; hind coxa without setulae on posterior margin, and 2 long setae on anterior surface.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F217FF9C5390BDDDFC883852.taxon	description	Description. Male holotype (Fig. 1 K) Head: eye with sparse setulae. Frontal vitta dark brown. 16 fr. 1 pair of oc, proclinate and developed, surrounded by 5 setulae. Fronto – orbital plate and parafacial with silver pollinosity, gena brown. Parafacial bare. Scape dark brown with 2 dorsal setae. Pedicel black with golden yellow pollinosity, 1 long dorsal seta and surrounded by short setulae. Postpedicel dark brown, 2.5 times the length of pedicel. Arista dark brown, with short pubescence, rays shorter than width of base of arista. Palpus dark brown, claviform. Thorax: dark brown; scutum without vittae. Acr 2 – 3: 3, irregular rows. Dc 2: 3. Pprn 3, developed and surrounded setulae. Pra 2. Proepimeron setulose. Katespisternum with long setulae and ventrally with a tuft of straight short setae. Wing: brownish, with superior third, between C vein and superior half of R 2 + 3 vein darkened. Calypter whitish. Haltere yellowish. Legs: coxae, femora and tibia brown; tarsi brown, except for the fore 1 – 4 tarsomeres white yellowish, pulvilli brownish. Fore femur with a few av on basal third, apical half almost bare; 1 row of d; pd and p surface setulose, with thin and long hooked setae; 1 row of long pv. Fore tibia with 1 pre – apical d; 1 apical pv and 1 v. Fore second tarsomere on ventral surface with a tuft of short whitish setae (Fig. 4 D). Mid femur with 1 row of unequal av, long on basal half and shorter on apical half; 1 row of ad; 1 row p; 1 row of pv; 1 row of shorter v until apical third, near pv row, 5 apical hooked setae. Mid tibia strongly constricted on ventral surface on basal half and highly setulose on apical half (Fig. 4 E), 1 strong and long apical v; 1 sub – median and 1 apical a; 1 apical av; 1 pre – apical ad; 1 submedian and 1 pre – apical pd, both short; 1 pre – apical p. Hind coxa without setulae on posterior margin, 2 long a. Hind femur curved and pre – apically enlarged, with 5 long strong v on the swelling (Fig. 4 F); 4 differentiated ad on apical third; short straight a, ad and av, apical third almost bare; setulose ventral surface until the apical third, 6 long and strong v on the swelling and 1 apical (Fig. 4 F); 1 pre – apical d; 1 pre – apical pd; pv surface setulose; posterior surface bare, except on basal third. Hind tibia with 1 median and 1 pre – apical d; 1 sub – median ad; 2 submedian and 1 apical av. Abdomen: dark brown. Sternite 1 setulose. Sternite 5 (Fig. 5 K). Terminalia (Fig. 7 C): epandrium wider than long, on basal half with short setae and two developed setae, two long acute apical internal projections, externally covering a basal part of surstylus; surstylus fusioned with epandrium, curved apically, shaped as a comma in frontal and in lateral view, basal half with internal basal expansion and setae; cercal plate much smaller than epandrium, slightly pointed and and fused to the basal third, weakly setulose; baciliform process long and simple, almost as long as half of the surstylus. Female: unknown.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F217FF9C5390BDDDFC883852.taxon	biology_ecology	Biology. Label information indicates that this species occurs in highland forests of Costa Rica, above 2000 m. Nothing else is known about its habits or biology.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F217FF9C5390BDDDFC883852.taxon	discussion	Comments. This species is similar to Fannia terraba sp. nov. but differs in that the katepisternal ventral setae are less developed, calypter whitish, haltere yellowish, fore second tarsomere on ventral surface with a tuft of short whitish setae on almost the entire tarsomere and hind coxa without setulae on posterior margin, and 2 long setae on anterior surface. Also similar to Fannia arcuata Chillcott, 1961 (pretiosa group) but differs due to dark legs. Similar to Fannia gilvitarsis Chillcott, 1961 (pretiosa group) from which it differs due to the pale yellow fore tarsus, apex of fifth is darkened.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F217FF9C5390BDDDFC883852.taxon	etymology	Etymology. Name in apposition. The species epithet is in homage to the Teribi indigenous people (also called Naso), that once inhabited northwestern Panama to northeastern Costa Rica.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F217FF9C5390BDDDFC883852.taxon	materials_examined	Type material. Holotype male (INBio). COSTA RICA: Prov. San Jose, Est. \ Cuerici, 4.6 Km al E. de Villa Mills. \ 2600 – 2640 m, 15 JUL 1996. A. \ Picado. L _ S _ 389400 _ 499600 # 7723 / INBIO CRI 002 467259. Paratypes. COSTA RICA. San José: Est. Cuerici, 4.6 Km al E. de Villa / Mills. Alrededores de la estación / Prov. San José. Costa Rica / 2600 m, 17 SET 1995. A. Picado / L _ S _ 389400 _ 499600 # 6303 / INBIO CRI 002 568225 (1 male, INBio); Est. Cuerici, sendero al Mirador / 4.6 km al E de Villa Mills, San / Jose, Costa Rica. 2700 m. 24 MAR / 1996. A. Picado / L _ S _ 389400 _ 499600 # 7027 / INBIO CRI 002 495063 (1 male, INBio); “ idem ” / 2640 m. 16 FEB 1996. B. / Gamboa, L _ S _ 389700 _ 499600 # 7034 / INBIO CRI 002 369699 (1 male, INBio); Est. Cuerici, sendero al Carbon 5 km / al E de Villa Mills, Prov. San Jose / Costa Rica. 2600 m. 26 OCT 1995 / B. Gamboa / L _ S _ 390100 _ 500100 # 6328 / INBIO CRI 002 468203 (1 male, DZUP); F. Cementerio de la Máquina / 2100 – 2500 m, P. N. Chirripó, Prov. / S. José, Costa Rica. 2 mar 1993 / M. A. Zumbado / L _ S _ 378700, 512500 / INBIO CRI 001 305770 (1 male, INBio); San Gerardo de Dota, 2000 / – 2500 m, Prov. San Jose / Costa Rica, Curso / Tachinidae y Syrphidae / 22 – 26 feb 1992 / L _ S _ 387400 _ 482700 / INBIO CRI 000 407258 (1 male, INBio); “ ídem ” / INBIO CRI 000 407349 (1 male, DZUP). The holotype is in good condition and has all structures.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F217FF9C5390BDDDFC883852.taxon	distribution	Distribution. Costa Rica: San José province (Fig. 8 K).	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F216FF9B5390BF88FCFF3929.taxon	description	(Figs. 1 L, 4 G – I, 5 L, 7 D, 8 L)	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F216FF9B5390BF88FCFF3929.taxon	diagnosis	Diagnosis. Fore tarsomeres 1 – 3 partially whitish – yellow, fifth fore tarsomere flattened, second fore tarsomere with setae forming a comb. Ventral katepisternal spine strong.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F216FF9B5390BF88FCFF3929.taxon	description	Description. Male holotype (Fig. 1 L) Head: eye setulose. Frontal vitta dark brown. 18 fr. 1 pair of oc, proclinate and developed, surrounded by long setulae. Fronto – orbital plate and parafacial with silver pollinosity, gena brown. Parafacial bare. Scape dark brown, 2 dorsal setae reaching pedicel. Pedicel black with golden yellow pollinosity, 1 long dorsal seta and surrounded by short setulae. Postpedicel dark brown, 2.5 times the length of pedicel. Arista dark brown, with short pubescence, rays shorter than width of base of arista. Palpus dark brown, claviform. Thorax: dark brown; scutum without vittae. Acr 3: 3, irregular rows. Dc 2: 3. Pprn 3, developed, surrounded setulae. Pra 2, seta closest to suture longer. Proepimeron setulose. Katespisternum with long setulae and tuft of straight short setae ventrally. Wing: brownish, with superior third, between C vein and superior half of R 2 + 3 vein darkened. Calypter whitish. Haltere brown. Legs: coxae, femora and tibia brown; tarsi brown, except the 1 – 3 fore tarsomeres white yellowish, apex of the third tarsomere and pulvilli brownish. Fore femur on basal half with 1 row of av, apical half almost bare; 1 row of d; pd and p surface setulose, with thin and long hooked setae; 1 series of long pv. Fore tibia with 1 pre – apical d; 1 apical pv and 1 v. Fore second tarsomere on ventral surface with a tuft of short whitish setae (Fig. 4 G). Mid femur with 1 row of av, long on basal half and shorter on apical half; 1 row of ad on basal half; 1 row of unordered p, with 8 pre – apical differentiated setae; 1 row of pv with 2 basal longer setae; 1 row of short v, near pv row. Mid tibia strongly constricted on v surface on basal half and highly setulose on apical half (Fig. 4 H), 1 strong and long apical setae; 1 av apical; 2 ad on apical half; 1 sub – median and 1 pre – apical pd; 1 short pre – apical p; 1 pre – apical pv. Hind coxa with 1 seta on posterior margin and a tuft of long setae on anterior surface. Hind femur strongly curved and preapically enlarged, with long strong v and pv on the swelling (Fig. 4 I); 1 row of ad, 4 setae strongly differentiated on apical half; a long and strong, apical third bare; 2 av on the swelling, separated between them; tuft of long and strong v on the swelling, short setae on the median and basal third; 1 pre – apical d; pd surface with a short pre – apical seta; pv surface with a row of short setae; p surface bare on apical third. Hind tibia with 1 median and 1 pre – apical d; 1 sub – median ad; 2 sub – median and 1 apical av. Abdomen: dark brown. Sternite 1 setulose. Sternite 5 (Fig. 5 L). Terminalia (Fig. 7 D): epandrium longer than wide, strong setae on basal half with two developed setae, two long and pointed apical internal projections; cercal plate much smaller than epandrium, with short and weak setae along the plate; surstylus fused with epandrium, curved, shaped as a comma in apical half, easily visible in frontal and lateral view, basal half with internal basal setae; bacilliform process absent. Female: unknown.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F216FF9B5390BF88FCFF3929.taxon	biology_ecology	Biology. Label information indicates that this species occurs in the highland forests of Costa Rica, 2100 – 3050 m. Nothing else is known about its habits or biology.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F216FF9B5390BF88FCFF3929.taxon	discussion	Comments. This species is similar to Fannia arcuata Chillcott, 1961 (pretiosa group) except it has dark legs. The new species is similar to Fannia gilvitarsis Chillcott, 1961 (pretiosa group) from which it differs in the paleyellow fore tarsus and apex of fifth tarsomere darkened.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F216FF9B5390BF88FCFF3929.taxon	etymology	Etymology. Name in apposition. The species epithet refers to the Terraba indigenous people, who once inhabited Puntarenas province in Costa Rica.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F216FF9B5390BF88FCFF3929.taxon	materials_examined	Type material. Holotype male (INBio). COSTA RICA. Prov. Puntarenas, P. Int. / La Amistad, de Cerro Becón a Tres / Colinas, 2124 m. 17 ABR 2002, M. / Alfaro, Libre, L _ S _ 343750 _ 565700 / # 69143 / INBIO 0003479937 INBIOCRI COSTA RICA. Paratypes. COSTA RICA. San José: Villa Mills, 3000 m. Prov. / San Jose, Costa Rica / Curso Tachinidae y / Syrphidae. 24 feb 1992 / L _ S 390000, 495000 / INBIO CRI 000 408439 (1 male, INBio); “ ídem ” / INBIO CRI 000 408387 (1 male, DZUP); “ ídem ” / INBIO CRI 000 408381 (1 male, INBio); “ ídem ” / INBIO CRI 000 408382 (1 male, INBio); Estac. Barva. Braulio / Carrillo, N. P. 2500 m / Heredia COSTA RICA / Feb 1990. G Rivera / 233400 – 523200 / INBIO CRI 000 202055 (1 male, INBio); “ idem ” / Mar 1990. A Fernandez / INBIOCRI 000 169464 (1 male, DZUP); Est. Cuerici, Sendero al Mirador, 4.6 / km al E. de Villa Mills, San Jose / Costa Rica 2640 – 2700 m. 20 – 22 ENE / 1996. A Picado, de Luz / L _ S 389700 _ 499600 # 6807 / INBIO CRI 002 392887 (1 male, DZUP); Costa Rica, Prov. San Jose / Sendero a Cerro Cuerici, Limite del / P. N. Chirripo. 3050 m. 5 JUL 1996. A. / Picado. L _ S _ 391300 _ 500100 # 7718 / INBIO CRI 002 444089 (1 male, INBio) The holotype is in good conditions.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F216FF9B5390BF88FCFF3929.taxon	distribution	Distribution. Costa Rica: Puntarenas province (Fig. 8 L).	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F211FF9A5390BE58FEDB3851.taxon	description	(Figs. 1 M, 4 J – K, 5 M, 7 E, 8 M)	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F211FF9A5390BE58FEDB3851.taxon	diagnosis	Diagnosis. Fore tarsus with tarsomeres 3 – 5 flattened, tarsomere 1 and 2 yellowish on part of v surface.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F211FF9A5390BE58FEDB3851.taxon	description	Description. Holotype male (Fig. 1 M) Head: eye slightly setulose. Frontal vitta velvety black, pollinose. 12 fr. Ocellar triangle black; oc proclinate and developed, 3 less developed setulae on ocellar area. Fronto – orbital plate silver and pollinose. Parafacial bare, silver pollinose. Fronto – genal suture and gena silver pollinose, gena with long setae. Scape black, 2 setae reaching pedicel. Pedicel black pollinose with a dorsal seta developed and surrounded minor setae. Postpedicel black, golden silver pilose, 2 times the length of pedicel. Arista black, with short pubescence, rays shorter than width of base of arista. Palpus black and filiform. Thorax: black. Scutum without vittae. Acr 2: 2 – 3 arranged in rows. Dc 2: 3, long, posterior most postsutural pair developed. Pprn 4, developed. Pra 2. Pro – epimeron setulose. Wing: brownish, with upper third between vein C and half of cell r 2 + 3 darker. Calypters whitish. Haltere brown with stem and base lighter in color. Legs: black, pulvilli brownish and tarsi black, except for fore tarsomere 1 and 2 with part of them yellowish on v surface. Fore femur setulose on posterior surface with 1 row of long pv and d. Fore tibia with 1 strong pre – apical d; 1 apical pv; 1 apical p; 1 apical v. Fore tarsomere 1 and 2 yellowish on part of v surface; tarsomeres 3 – 5 flattened (Fig. 4 J). Mid femur with 2 rows of long pv, hooked apices on apical third and 1 row of developed av decreasing to the apex. Mid tibia on ventral surface with pronounced constriction on basal half area, densely setulose on apical half, 1 v; 2 median ad; 1 d; 2 p; 1 apical pv; 1 pd. Hind coxa bare on posterior margin. Hind femur on ventral surface with pre – apical protuberance; av, v and p surfaces setuloses, av setae longer and curved, increasing in size from base to apex (Fig. 4 K). Hind tibia with 1 median d and 1 pre – apical; 1 median a; 1 median av and 1 apical (Fig. 4 K). Abdomen: elongate, black, weakly greyish pollinose with developed setae. Sternite 1 densely setulose. Sternite 5 (Fig. 5 M). Terminalia (Fig. 7 E): epandrium wider than long, with apical hook extensions; cercal plate pointed apically, base fused almost completely; surstylus strongly articulated with epandrium, pointed apically; bacilliform process spiral and short near epandrium extensions. Female: unknown.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F211FF9A5390BE58FEDB3851.taxon	biology_ecology	Biology. Label information indicates that this species occurs in the Andean highlands in Ecuador, from 2500 m in the western foothills. Nothing else is known about its habits or biology.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F211FF9A5390BE58FEDB3851.taxon	discussion	Comments. Fannia tsachilas sp. nov. is morphologically similar to Fannia iguaque, F. chingaza and F. sutagao described by Grisales, Wolff and Carvalho (2012), due to the fore tarsomeres 3 – 5 flattened, but differs in the hind femur and the terminalia.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F211FF9A5390BE58FEDB3851.taxon	etymology	Etymology. Name in apposition. The species epithet refers to the Tsachilás indigenous people, from Ecuador.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F211FF9A5390BE58FEDB3851.taxon	materials_examined	Type material. Holotype male (MZSP). ECUADOR: N. Latacunga / Ecuador, 3300 m. / XII. 1970 / L. E. Peña Leg / MZSP. Paratypes. ECUADOR. Latacunga: N. Latacunga / Ecuador, 3300 m. / XII. 1970 / L. E. Peña Leg / MZSP (6 males MZSP, 5 males DZUP); Saraguro: 2500 m. Ecuador / XI. 1970 / L. E. Peña col. / MZSP (5 males MZSP, 5 males DZUP); Quito: Ascázubi, Pifo / N. E. Quito, Ecuador / 2500 – 2800 m. XII – 1970 / L. E. Peña col. / MZSP (1 male, DZUP); Santo. Domingo de los Colorados: Ecuador / 2600 m. XII – 1970 / L. E. Peña col. / MZSP (1 male, MZSP); Otavalo: N. Perucho / Otavalo 2000 m. / Ecuador. I. 1971 / L. E. Peña col. / MZSP (1 male, MZSP). The holotype has no abdomen because it was taken to dissect the terminalia and it was fixed under the holotype.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F211FF9A5390BE58FEDB3851.taxon	distribution	Distribution. Ecuador: provinces of Cotopaxi, Loja, Santo Domingo de los Tsáchilas and Imbabúra (Andean region) (Fig. 8 M).	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F210FF995390BF8DFEBE389A.taxon	description	(Figs. 1 N, 4 L – N, 5 N, 7 F, 8 N)	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F210FF995390BF8DFEBE389A.taxon	diagnosis	Diagnosis. Wing brownish, without upper third darker, fore tarsomeres 3 – 5 flattened, mid tibia with apical modified setae (leaf – like) on pv surface, hind coxa setulose on posterior margin, hind femur slightly curved, anterior surface slightly setulose and posterior surface with a tuft of short setae on the posterior protuberance.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F210FF995390BF8DFEBE389A.taxon	description	Description. Holotype male (Fig. 1 N) Head: eye densely setulose. Frontal vitta velvety black, pollinose. 15 fr. 1 pair of oc, proclinate and developed, surrounded by 5 less developed setulae. Fronto – orbital plate black, greyish pollinose. Parafacial bare, narrowing ventrally, greyish pollinose. Fronto – genal suture with brownish hairs. Gena brownish pollinose. Scape black, 3 – 4 short dorsal setae reaching pedicel. Pedicel black pollinose, dorsal setae developed, one of which is stronger. Postpedicel black, brownish pilose, 2 times the length of pedicel. Arista black, with short pubescence, rays shorter than width of base of arista. Palpus black, filiform. Thorax: black; scutum without vittae. Acr 2: 3 – 4, irregular rows. Dc 2: 3. Pprn 4, developed. Pra 2, strong. Proepimeron setulose. Wing: slightly brownish without upper third darker. Calypter whitish. Haltere brown with stem and base lighter in colour. Legs: black with tarsi black and pulvilli brownish. Fore femur setulose on posterior surface with 1 row of long pv and d; ventral surface bare. Fore tibia with 1 pre – apical d; 1 apical pv; 1 apical v. Fore tarsus with 1 thin v on first tarsomere; tarsomeres 3 – 5 flattened (Fig. 4 L). Mid femur slightly constricted on pre – apical ventral surface; 1 row of short ad with 4 developed setae on the apex; 1 row of strong av that decrease in length towards apex; 1 row of long pv with hooked apex, setae decrease in length towards apex; 1 row of weak p with 6 longer on apical third. Mid tibia on ventral surface with 2 pronounced constrictions on basal half area, densely setulose on through the surface; 1 developed pre – apical ad; 2 developed a on mid apical half and 1 apical; 1 developed apical av; 1 apical modified pv (leaflike) (Fig. 4 M); 2 p. Hind coxa setulose on posterior margin. Hind femur on ventral surface with a slightly pre – apical protuberance; 1 row of ad, 4 stronger setae running towards dorsal surface (Fig. 4 N); 1 row of longer curved av, that does not form a tuft on the pre – apical protuberance; anterior surface slightly setulose (Fig. 4 N); posterior surface with a tuft of short setae on the posterior protuberance. Hind tibia with 1 developed median and 1 pre – apical d; 1 long and developed median ad; anterior surface with short setae along and 1 short apical; 2 strong median av, 1 apical. Abdomen: elongate, black, greyish pollinose, developed setae. Sternite 1 densely setulose. Sternite 5 (Fig. 5 N). Terminalia (Fig. 7 F): epandrium wider than long; cercal plate elongated, with short weak setae along the plate, fusioned on basal third; surstylus strongly articulated with epandrium, broad at the base and thin at the apex, shaped like a foot, bacilliform process spiral and simple. Female: unknown.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F210FF995390BF8DFEBE389A.taxon	biology_ecology	Biology. Label information indicates that this species is found on or near decomposing flesh (carcasses). It occurs from the Oriental Cordillera in Colombia to the Andean highlands of Ecuador (2300 – 2900 m). Nothing else is known about its habits or biology.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F210FF995390BF8DFEBE389A.taxon	discussion	Comments. Fannia yukpa sp. nov. is similar to Fannia pijao Grisales, Wolff & Carvalho, 2012 and Fannia quimbaya Grisales, Wolff & Carvalho, 2012 from Colombia. Fannia yukpa sp. nov. differs due to its mid tibia with apically modified setae (leaflike) on pv surface.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F210FF995390BF8DFEBE389A.taxon	etymology	Etymology. Name in apposition. The species epithet refers to the Yukpa indigenous people, of the Serranía del Perija, between Colombia and Venezuela.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F210FF995390BF8DFEBE389A.taxon	materials_examined	Type material. Holotype male (ICN). COLOMBIA: Cundinamarca. Bogotá \ Sierras del Chicó \ T 4 T 5 Mayo 15 / 2007 \ I. Morales \ ICN. Paratypes. COLOMBIA. Norte de Santander: Pamplona / Universidad de Pamplona. 2360 msnm \ En cerdo (forense). Jul – Ago / 2006 \ Leidy Ardila \ CEUA F 14449 \ Local \ Alt. _ \ Hosp: CPE \ Fech: 3 – 08 – 06 \ Col: Leidy Ardila (1 male, CEUA). ECUADOR. Quito: Baeza / E. Papallacta / 2900 m. I – 71 / L. E. Peña col. (2 males, MZSP); Saraguro: Sur de Saraguro / 2900 m. / XI – 1970 / L. E. Peña leg. (1 male, MZSP). The holotype is in good condition and has all structures.	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
6909C857F210FF995390BF8DFEBE389A.taxon	distribution	Distribution. Colombia: departments of Cundinamarca and Norte de Santander (Oriental Cordillera, Andean region), Ecuador: province of Napo (Andean region), province of Loja (Saraguro Canton, Ecuadorian highlands) (Fig. 8 N).	en	Grisales, Diana, De Carvalho, Claudio J. B. (2019): Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America. Zootaxa 4551 (3): 330-360, DOI: 10.11646/zootaxa.4551.3.4
