identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
6F64DB1B5E43FFA7FC7CF914FDD0FE80.text	6F64DB1B5E43FFA7FC7CF914FDD0FE80.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heraeus Stal 1862	<div><p>GENUS HERAEUS STÅL, 1862</p><p>Type species: Lygaeus (Plociomerus) triguttatus Guérin-Méneville, 1857, by monotypy.</p><p>Heraeus Stål, 1862: 314 –315; Stål, 1874: 147; Distant, 1882: 204; Provancher, 1886: 80; Berg, 1892: 162; Lethierry &amp; Severin, 1894: 191; Van Duzee, 1917: 178–179; Barber, 1918: 75; Blatchley, 1926: 389; Barber, 1928: 175; Barber, 1939: 349, 354; Froeschner, 1944: 640; Torre Bueno, 1946: 64, 69–70; Slater &amp; Hurlbutt, 1957: 73; Scudder, 1957: 155; Sweet &amp; Slater, 1961: 339; Slater, 1964: 1081–1082; Slater, 1972: 149; Slater, 1974: 167, 170; Sweet, 1967: 225; Brailovsky, 1979: 549; Harrington, 1980: 108; Brailovsky, 1981: 219; Froeschner, 1981: 43; Froeschner, 1985: 22; Harrington, 1987: 81; Ashlock &amp; Slater, 1988: 227; Slater &amp; Baranowski, 1990: 130; Baranowski &amp; Slater, 1998: 79; Froeschner, 1999: 244; Baranowski &amp; Slater, 2005: 136; Dellapé 2014: 428, 431; Henry et al. 2015: 488, 499.</p><p>Abbreviations: APLL, anterior pronotal lobe length; APLW, anterior pronotal lobe width; BFL, basiflagellomere length; CL, collar length; DFL, distiflagellomere length; DFBL, distiflagellomere band length; HL, head length; HW, head width; IOcW, interocular width; IOclW, interocellar width; PL, pedicellus length; POcL, postocular length; PPLL, posterior pronotal lobe length; PPLW, posterior pronotal lobe width; R1, rostral segment-I length; R2, rostral segment-II length; R3, rostral segment-III length; R4, rostral segment-IV length; SCL, scapus length; TL, total length.</p><p>Diagnosis</p><p>The species included in Heraeus are recognized by the anteriorly extended collar on the ventral region (Fig. 4 A), which is a synapomorphy for the genus. All species have: an elongate head with a postocular region longer than the interocular space; the dorsally narrowed and ventrally broadened collar, extending forwards beneath the head (Figs 1, 4 A); and the aedeagus with a bilobed vesica and a simple conjunctiva, showing relatively small spines in most of the species.</p><p>Redescription</p><p>Head elongate, postocular distance longer than interocular distance. Buccular juncture V-shaped. Juga rounded. Collar narrow dorsally and broadening ventrally, extending forwards beneath the head. Procoxa with a spine. Profemur with two rows of spines ventrally. Mesofemur with or without spines. Protibia without spines or with small denticles or spines. Tibiae with spiniform setae. Corial margin smooth. Clavus with more than three rows of punctures. Mesepimeron emergent. Evaporative area extensive, not restricted to a small portion of metapleuron surrounding the auricle. Male genitalia: aedeagus without spines or with relatively small spines, with two lobes on vesica lateral to the ejaculatory reservoir.</p><p>Distribution</p><p>The genus is widely distributed from Canada to Argentina, including the West Indies and the Galapagos Islands. Most of its 39 species are distributed on continental areas. Heraeus plebejus, H. guttatus, and H. triguttatus are also found on at least one island of the West Indies, and some species are known only from the West Indies. Heraeus dominicanus sp. nov. is only known from the Dominican Republic, Heraeus steineri sp. nov. is known from Turks and Caicos Islands, and the islands of the Bahamas, H. caliginosus is known from the Dominican Republic and Haiti, H. hollyae is known from the Dominican Republic, Haiti, and Antigua Island; H. pulchellus is recorded from the Bahamas, Cuba, and the Cayman Islands, and H. concolor is recorded from the Dominican Republic, Haiti, Jamaica, Antigua, Saba, and the Nevis islands. Heraeus pacificus is known only from the Galápagos islands.</p><p>INCLUDED SPECIES GROUPED ACCORDING TO EXTERNAL APPEARANCE AND PHYLOGENETIC</p><p>RELATIONSHIPS</p><p>The coquilletti group</p><p>Heraeus cinnamomeus Barber, 1948</p><p>Heraeus coquilletti Barber, 1914</p><p>Heraeus costalis sp. nov.</p><p>Heraeus itzelae sp. nov.</p><p>Heraeus setosus sp. nov.</p><p>The caliginosus group</p><p>Heraeus caliginosus Slater &amp; Baranowski, 1994</p><p>Heraeus dominicanus sp. nov.</p><p>The guttatus group</p><p>Heraeus guttatus (Dallas, 1852)</p><p>Heraeus hollyae Baranowski, 2005</p><p>Heraeus steineri sp. nov.</p><p>Heraeus triguttatus (Guérin-Méneville, 1857)</p><p>The illitus group</p><p>Heraeus annulatus sp. nov.</p><p>Heraeus antennalis sp. nov.</p><p>Heraeus chamamecinus sp. nov.</p><p>Heraeus illitus Distant, 1882</p><p>Heraeus inca sp. nov.</p><p>Heraeus panamaensis sp. nov.</p><p>Heraeus similis sp. nov.</p><p>Heraeus spinosus sp. nov.</p><p>Heraeus splendens sp. nov.</p><p>Heraeus tiputini sp. nov.</p><p>The plebejus group</p><p>Heraeus alvarengai sp. nov.</p><p>Heraeus apicalis sp. nov.</p><p>Heraeus bahiensis sp. nov.</p><p>Heraeus baranowskii sp. nov.</p><p>Heraeus bolivianus sp. nov.</p><p>Heraeus brevirostris sp. nov.</p><p>Heraeus concolor Slater &amp; Baranowski, 1994</p><p>Heraeus ecuatorianus sp. nov.</p><p>Heraeus loja sp. nov.</p><p>Heraeus mesoamericanus sp. nov.</p><p>Heraeus mexicanus sp. nov.</p><p>Heraeus morganae sp. nov.</p><p>Heraeus nicaraguensis sp. nov.</p><p>Heraeus pacificus Barber, 1925</p><p>Heraeus pallidinervis sp. nov.</p><p>Heraeus penai sp. nov.</p><p>Heraeus plebejus Stål, 1874</p><p>Heraeus pulchellus Barber, 1954</p></div>	https://treatment.plazi.org/id/6F64DB1B5E43FFA7FC7CF914FDD0FE80	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pablo M. Dellapé;María C. Melo;Thomas J. Henry	Pablo M. Dellapé, María C. Melo, Thomas J. Henry (2016): A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species. Zoological Journal of the Linnean Society 177 (177): 29-134, DOI: 10.1111/zoj.12362
6F64DB1B5E48FFA7FE8FFEBEFF30FC06.text	6F64DB1B5E48FFA7FE8FFEBEFF30FC06.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heraeus coquilletti	<div><p>THE COQUILLETTI GROUP</p><p>Included species: Heraeus cinnamomeus, H. coquilletti, H. costalis sp. nov., H. itzelae sp. nov., and H. setosus sp. nov.</p><p>Diagnosis</p><p>Sparsely pilose species, with head smooth and shiny (except H. setosus sp. nov.); ocelli located posteriorly to an imaginary line passing the posterior border of eyes; posterior pronotal lobe and clavus uniformly coloured; corium without pale spots, at most with a small spot restricted to costal margin; evaporative area short, distance from dorsal margin of auricle to dorsal margin of evaporative area shorter than distance from dorsal margin of evaporative area to dorsal margin of metapleura; males with strongly incrassate profemora with one ventral spine much larger than the others and protibiae with a row of small tubercles on inner surface.</p></div>	https://treatment.plazi.org/id/6F64DB1B5E48FFA7FE8FFEBEFF30FC06	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pablo M. Dellapé;María C. Melo;Thomas J. Henry	Pablo M. Dellapé, María C. Melo, Thomas J. Henry (2016): A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species. Zoological Journal of the Linnean Society 177 (177): 29-134, DOI: 10.1111/zoj.12362
6F64DB1B5E48FFA4FF5BFC34FC32FC0E.text	6F64DB1B5E48FFA4FF5BFC34FC32FC0E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heraeus cinnamomeus Barber 1948	<div><p>HERAEUS CINNAMOMEUS BARBER, 1948</p><p>(FIGS 3 A, 4A–C, 5A, 6A–D, 7)</p><p>Heraeus coquilletti Barber, 1914a: 165 –166 (in part); Van Duzee, 1917: 179 (in part); Blatchley, 1934: 8 (in part); Torre Bueno, 1946: 70 (in part).</p><p>Heraeus cinnamomeus Barber, 1948: 67; Slater, 1964: 1082; Schaefer, 1972: 812; Harrington, 1980: 108; Ashlock &amp; Slater, 1988: 227.</p><p>Diagnosis</p><p>Head shiny, contrasting with a dull pronotum and hemelytra. Pronotum and scutellum without erect setae. Distiflagellomere and hemelytra uniformly coloured cinnamon brown. Profemora pale yellowish brown.</p><p>Heraeus cinnamomeus and H. costalis sp. nov. are the only two species of the coquilletti group that have a shiny head, contrasting with a dull pronotum and hemelytra, and the scutellum lacking erect setae. The general colouration of H. cinnamomeus is cinnamon brown, with distiflagellomeres and hemelytra uniformly coloured, whereas H. costalis sp. nov. is dark brown with the distiflagellomere mostly white and with the apex darker.</p><p>Redescription (Fig. 3 A)</p><p>Paratype ♂</p><p>Head: Strongly convex dorsally (Fig. 4 A). Colour orange brown, shiny and smooth, with short recumbent and erect setae dorsally. Eyes slightly protruding, not surpassing the dorsal margin of head in lateral view. Ocelli posterior to an imaginary line passing the posterior border of eyes. Labium pale brown with erect setae, extending to mesocoxae. Antennae pale brown, distiflagellomere slightly paler; with short recumbent, semi-erect, and erect setae.</p><p>Thorax: Pronotum dull brown, anterior lobe darker; pruinose, punctate (Fig. 4 B); collar punctate, delimit- ed posteriorly by a sulcus and a row of punctures, anterior pronotal lobe with punctures more conspicuous posteriorly, posterior pronotal lobe coarsely punctate; anterior and posterior lobes with minute setae. Pleurae brown. Evaporative area short. Scutellum brown, pruinose (Fig. 4 C), punctate, with minute recumbent setae. Hemelytra pale brown, pruinose (Fig. 4 C), with minute recumbent setae. Corium with a diffuse subapical pale spot; lateral margins slightly concave. Membrane pale brown, veins concolorous. Legs pale brown, coxae and apical third of metafemur, and apex of tibiae, darker (Fig. 5 A); with sparse, short, recumbent and erect setae. Tibiae with spiniform setae. Profemur conspicuously incrassate in males; spines on profemur small, except for one much larger spine. Protibia with small tubercles bearing spiniform setae in a row along inner surface.</p><p>Abdomen: Brown, with abundant short recumbent setae. Male genitalia: Pygophore (Fig. 6 A, B) rounded, anterior margin of dorsal aperture slightly rounded, inner projections projecting posteriorly. Parameres: Figure 6 (C, D). Aedeagus: Conjunctiva unspined, vesica with small spines, lobes of vesica strongly sclerotized at base; processus gonopori long and slender.</p><p>Distribution</p><p>Mexico (NEW RECORD) and USA (Fig. 7).</p><p>Type material examined</p><p>Holotype: ♂, USA, Texas, Brownsville, Los Borregos, 5-VI-[19]04, type #58494 (USNM).</p><p>Paratypes: 1♂, 1♀, Tex.[as], Brownsville, 15-V- [19]36, J.N. Knull (OSUC); 1♂, Tex., Brownsville, 8-V-[19]36, J.N. Knull (OSUC); 3♀, Los Borregos, Brownsville, 5-VI-[19]04, H.S. Barber (USNM); 1♀, Brownsville, 29-II-[19]36, P.A. Glick (USNM); 1♂, 8–25-V-[19]35, J.N. Knull (USNM); 1♂, 15-V- [19]35, J.N. Knull (USNM); 1♀, 30-V-[19]04, H.S. Barber (USNM); 1♂, 2♀ (USNM); 1♀, Esper Ranch, Brownsville, Brooklyn Museum coll. (USNM); 1♂, 20-V-1030, J.C. Gaines (USNM).</p><p>Additional material studied</p><p>MEXICO: 1♀, Mexico, intercepted at Br. Tex. in plane, #69209, 1-27-50-2072 (USNM); 1♀, Mexico, intercept- ed at Laredo, Tx., 30-I-[19]43, with tomatoes, #31333 (USNM); 1♀, Mexico, 7-VII-[19]44, tomato fruit, inter-</p><p>cepted at Brownsville #59250 (USNM); 1♂, Mexico, in plane, 28-X-[19]41, intercepted at Brownsville, Tx (USNM). Chihuahua: 1♀ Salaices, 5200 ft, 20-VIII- 1947, G.M. Bradt (AMNH); San Luis Potosi: 1♂, Tamazunchales, intercepted at Laredo, Tx., 10-X- 1957, with orchid plants (USNM); 2♂, El Salto Falls, 2000–2500 ft, 22-IV-[19]65, in blacklight trap, H.V. Weems (USNM); Sinaloa: 1♀, 43 mi. N Mazatlan, 27-VII-1952, J. Lattin (AMNH); Tamaulipas, 1♂, paratype [this is not a paratype], C[iudad] Victoria, 31-III-1951, J. Lattin (AMNH); 1♀, Guemes, 28-VI- 1965, collected at blacklight, P.J. Spangler (USNM); 1♂, El Salto Falls, 26 mi. W Antiguo Morelos, 2000 ft, 11/ 14-VII-1963, Duckworth &amp; Davis (USNM); 1♀, Victoria, intercepted at Br. TX #68098, 14-VI-[19]49, avocado budwood (USNM); Nuevo Leon, 35♂, 31♀, two without abdomen, anegade arroya, 16 mi. S Linares, N.L., 1250 ft, 9-VII-1963, Duckworth &amp; Davis (USNM); 1♀, Linares, Rio Camacho, 21/22-VI-1965, O.S. Flint (USNM); 5♂, 4♀, 31 mi. E Galeana, 5000 ft, 7/9-VIII- 1963, Duckworth &amp; Davis (USNM); Jalisco: 1♀, Chamela, Station UNAM, 6/7-IX-1984, D. &amp; B. Sigwalt (MNHN).</p><p>USA: Texas: 9♂, 11♀, Brownsville, X-[19]42, E.S. Ross</p><p>(CAS); 4♂, 7♀, X-[19]42, E.S. Ross, at light (CAS);</p><p>6♂, 14♀, 16-IX-[19]42, T.M. Burns (CAS); 1♀, Hidalgo</p><p>Co., 24-III-[19]54, D.J. &amp; J.N. Knull (OSUC); 3♂,</p><p>2♀, Weslaco, 11-X-1930, S.W. Clark (CNC); 1♂, 1♀,</p><p>Brazos Co., N. Banks (AMNH); 1♂, Brownsville, 11/</p><p>16-VI-[19]33, Darlington (AMNH); 1♀, SE Hidalgo</p><p>Co., 28-IV/4-V-[19]46, at light, G.B. Vogt (USNM); 1♀,</p><p>26/31-VII-[19]46 (USNM); 1♀, 29-VIII-[19]47 (USNM);</p><p>1♀, 1/5-X-[19]47 (USNM); 1♀, 6-IV-[19]46, beating</p><p>flwrs and foliage of Prosopis juliflora Swartz de Candolle (USNM). 1♂, Canal Z. [PANAMA] or GUA- TEMALA, 7-IX-[19]42, intercepted at Brownsville, Tx. (USNM).</p></div>	https://treatment.plazi.org/id/6F64DB1B5E48FFA4FF5BFC34FC32FC0E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pablo M. Dellapé;María C. Melo;Thomas J. Henry	Pablo M. Dellapé, María C. Melo, Thomas J. Henry (2016): A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species. Zoological Journal of the Linnean Society 177 (177): 29-134, DOI: 10.1111/zoj.12362
6F64DB1B5E49FFA4FDE4FF05FA4AFC95.text	6F64DB1B5E49FFA4FDE4FF05FA4AFC95.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heraeus	<div><p>KEY TO THE SPECIES OF HERAEUS</p><p>1. Hemelytra nearly uniformly coloured, without a large pale spot at apex of corium (or with a small, pale subapical corial spot, with apex darker; Fig. 3 A–E); head smooth ( coquilletti group) ................................................... 2</p><p>– Hemelytra more heterogeneously coloured, often with a large pale spot at apex of corium (Fig. 3 F, K); head rugose. ......................................................................................................................................................6</p><p>2. Pronotum and scutellum with long erect setae ....................................................................................... 3</p><p>– Pronotum and scutellum without long erect setae ................................................................................... 4</p><p>3. Distiflagellomere pale brown, with a pale band; hemelytra with only short, recumbent setae (Mexico; Fig. 3 D) ... ..................................................................................................................... Heraeus itzelae sp. nov.</p><p>– Distiflagellomere yellowish brown, without a pale band; hemelytra with long, erect setae (Honduras and Mexico; Fig. 3 E).......................................................................................................... Heraeus setosus sp. nov.</p><p>4. Head, pronotum, and hemelytra shiny (Mexico and USA; Fig. 3 B).................... Heraeus coquilletti Barber, 1914</p><p>– Head shiny, contrasting with a dull pronotum and hemelytra ................................................................... 5</p><p>5. General colouration dark brown; distiflagellomere mostly white, with the apex dark; profemur mostly dark brown (Costa Rica, Honduras, and Nicaragua; Fig. 3 C) ................................................... Heraeus costalis sp. nov.</p><p>– General colouration cinnamon brown; distiflagellomere uniformly brown; profemur pale yellowish brown (Mexico and USA; Fig. 3 A)............................................................................... Heraeus cinnamomeus Barber, 1948</p><p>6. Hemelytron with a distinct, white, subapical spot on corium and a white mark on apex of membrane; species usually glabrous ( guttatus and caliginosus groups; Fig. 3 F–K)..............................................................................7</p><p>– Hemelytron with different combination of spots; if subapical corial spot present, shape is irregular and the colour pattern is more complex; species often setose (Figs 15, 25, 36).................................................................12</p><p>7. Labium extending beyond metacoxae; posterior pronotal lobe with four small pale spots strongly contrasting with dark background (Dominican Republic and Haiti; Fig. 3 J).......... Heraeus caliginosus Slater &amp; Baranowski, 1994</p><p>– Labium extending only to mesocoxae, at most reaching metacoxae; posterior pronotal lobe without four distinct spots...............................................................................................................................................8</p><p>8. Apical third of metafemur contrastingly dark brown (Fig. 11 A, B)..............................................................9</p><p>– Metafemur uniformly pale or brown, if paler basally, not strongly contrasting............................................10</p><p>9. Posterior pronotal lobe brown, mottled with paler areas, sometimes forming irregular lines; profemur dark brown with apex usually paler (Central America and Colombia; Figs 3 F, 11A) .............. Heraeus guttatus (Dallas, 1852)</p><p>– Posterior pronotal lobe uniformly dark, with two short yellow lines; profemur uniformly dark brown (Antigua, Dominican Republic, and Haiti; Figs 3 G, 11B)............................................... Heraeus hollyae Baranowski, 2005</p><p>10.Metafemur reddish brown; head smooth; distiflagellomere uniformly brown; males without small spines on protibia and mesofemur (Dominican Republic; Fig. 3 K).............................................. Heraeus dominicanus sp. nov.</p><p>– Metafemur pale; head rugose; distiflagellomere with pale band; males with small spines on protibia and mesofemur. .................................................................................................................................................... 11</p><p>11. Head, pronotum, and most of hemelytra anterior to distal white spot reddish or cinnamon brown; legs brownish yellow (USA and West Indies; Fig. 3 H) ....................................... Heraeus triguttatus (Guérin-Méneville, 1857)</p><p>– Head and anterior pronotal lobe dark brown, posterior pronotal lobe and hemelytra brown anterior to distal spot, mottled with yellow; legs uniformly pale, almost white (Dominican Republic; Fig. 3 I)...................................... .................................................................................................................... Heraeus steineri sp. nov.</p><p>12.Profemur pale brown with dark-brown spots, if predominantly dark brown, the dorsal region clearly pale ( illitus group; Figs 16, 21) ........................................................................................................................... 13</p><p>– Profemur uniformly dark brown, some species with apex paler ( plebejus group; Figs 26, 33, 39) .................... 22</p><p>13.Labium long, extending to metacoxae or beyond....................................................................................14</p><p>– Labium shorter, not extending beyond mesocoxae..................................................................................15</p><p>14.Strongly setose; metatibiae densely setose, with setae two times the diameter of the segment; labium long, surpassing metacoxae, usually extending to abdominal segment II (Central and South America; Fig. 15 D).............. ................................................................................................................. Heraeus illitus Distant, 1882</p><p>– Less setose; metatibial setae less abundant and shorter; labium shorter, usually extending only to posterior margin of metasternum (Argentina, Bolivia, Brazil, French Guiana, and Peru; Fig. 15 G)......... Heraeus similis sp. nov.</p><p>15.Basiflagellomere thickened distally, diameter subequal to or greater than diameter of scape.........................16</p><p>– Basiflagellomere slender, diameter less than diameter of scape................................................................ 17</p><p>16.Pronotum with a pale inverted V-shaped mark; outer margin of clavus pale (Panama; Fig. 15 I)........................ ................................................................................................................ Heraeus splendens sp. nov.</p><p>– Pronotum and hemelytra uniformly pale brown, with evenly scattered brown-stained punctures, except for distal white marks (Ecuador; Fig. 15 B)................................................................... Heraeus antennalis sp. nov.</p><p>17.Distiflagellomere uniformly brown (Argentina and Brazil; Fig. 15 C) .............. Heraeus chamamecinus sp. nov.</p><p>– Distiflagellomere with a distinct white band sub-basally.........................................................................18</p><p>18.Profemur with a distinct subapical brown band (Figs 16 A, 21B)...............................................................19</p><p>– Profemur without a subapical brown band ...........................................................................................20</p><p>19.Brown dark band on profemur narrow and contrasting (Fig. 16 A); apex of pygophore (Fig. 17 A–D) rounded, without a small dorsally directed protuberance; aedeagus with lobes sclerotized with a few spines distally (Central America, from Mexico to Panama; Fig. 15 A).................................................................. Heraeus annulatus sp. nov.</p><p>– Brown band on profemur broader and diffuse (Fig. 21 B); pygophore with a small dorsally directed protuberance near apex (Fig. 20 N); aedeagus with spines on lobes and laterally to the ejaculatory reservoir (Brazil and Ecuador; Fig. 15 H)...................................................................................................... Heraeus spinosus sp. nov.</p><p>20.Profemur irregularly pigmented with brown, metafemoral band subapical, processus gonopori long and slender (Panama; Figs 15 F, 16F) ........................................................................................ Heraeus panamaensis sp. nov.</p><p>– Profemur paler dorsally, metafemoral band apical (a few specimens of H. inca sp. nov. show a dark band on metafemur instead of being darker distally), processus gonopori long, widening towards apex (Fig. 21 D).........................21</p><p>21.Pronotum usually without contrasting dark areas laterally; dorsal anterior margin of aperture of pygophore rounded (Fig. 23 E); aedeagus with spines restricted to unsclerotized lobes (Fig. 8 I) (Ecuador; Fig. 15 J).......................... ................................................................................................................... Heraeus tiputini sp. nov.</p><p>– Pronotum usually with contrasting dark areas laterally; dorsal anterior margin of aperture of pygophore subquadrangular (Fig. 20 A); aedeagus with two pairs of lobes, the anterior lobe short, not sclerotized, and with spines; the posterior lobe large, strongly sclerotized posteriorly, with a row of spines (Peru; Fig. 15 E)............................................ ........................................................................................................................ Heraeus inca sp. nov.</p><p>22.Distiflagellomere without a pale band..................................................................................................23</p><p>– Distiflagellomere with a pale band......................................................................................................25</p><p>23.Antenna uniformly brown; dorsal margin of pygophore in lateral view almost straight and slightly declivent posteriorly (Fig. 40 F) (Galapagos Islands; Fig. 36 E).............................................. Heraeus pacificus Barber, 1925</p><p>– Antenna brown with apex of basiflagellomere and distiflagellomere darkened; dorsal margin of pygophore in lateral view sinuate and more declivent posteriorly (Fig. 41 B, F) ....................................................................... 24</p><p>24.Small species, less than 5.00 mm; subapical pale spot on corium well defined (West Indies; Fig. 36 I)................. ............................................................................................................ Heraeus pulchellus Barber, 1954</p><p>– Larger species, over 5.25 mm; subapical pale spot irregular, not well defined (USA and Central America; Fig. 36 H). Heraeus plebejus Stål, 1874</p><p>25.Labial segment IV extending only to mesocoxae .................................................................................... 26</p><p>– Labial segment IV extending to at least metacoxae................................................................................32</p><p>26.Metafemoral band apical (Figs 25 B, 26B)............................................................ Heraeus apicalis sp. nov.</p><p>– Metafemoral band subapical .............................................................................................................. 27</p><p>27.Pale band on distiflagellomere short, about one-quarter of segment length (Argentina and Bolivia; Fig. 25 F) ...... .............................................................................................................. Heraeus brevirostris sp. nov.</p><p>– Pale band on distiflagellomere longer, occupying at least one-third of segment length..................................28</p><p>28.Membranae with whitish apical spot...................................................................................................29</p><p>– Membranae without whitish apical spot...............................................................................................31</p><p>29.Anterior margin of dorsal aperture of pygophore rounded (Fig. 34 I) (Costa Rica and Mexico; Fig. 36 B) .............. ............................................................................................................... Heraeus mexicanus sp. nov.</p><p>– Anterior margin of dorsal aperture of pygophore almost straight (Figs 27 I, 40A).........................................30</p><p>30.Dark species; head blackish; posterior lobe of pronotum dark brown, with four longitudinal pale stripes (Nicaragua and Costa Rica; Fig. 36 D).................................................................. Heraeus nicaraguensis sp. nov.</p><p>– Pale species; head dark reddish brown; posterior lobe of pronotum pale brown with four difuse longitudinal pale stripes (Brazil; Fig. 25 C)................................................................................ Heraeus bahiensis sp. nov.</p><p>31.Apex of pygophore broadly rounded posteriorly in dorsal view (Fig. 32 A); outer projection of paramere not conspicuous (Fig. 32 C, D) (Bolivia; Fig. 25 E)......................................................... Heraeus bolivianus sp. nov.</p><p>– Apex of pygophore acutely rounded posteriorly in dorsal view (Fig. 34 E); outer projection of paramere conspicuous (Fig. 34 G, H) (Central America; Fig. 36 A)............................................... Heraeus mesoamericanus sp. nov.</p><p>32.Ocelli located anterior to an imaginary line passing through posterior margin of eyes; pygophore with a conspicuous protuberance at posterior margin pointed dorsally (Fig. 27 B) (Brazil; Fig. 25 A)......................................... ............................................................................................................... Heraeus alvarengai sp. nov.</p><p>– Ocelli located at level of an imaginary line passing through the posterior margin of eyes; pygophore without a dorsally directed protuberance at posterior margin, but if present, smaller and indistinct.............................33</p><p>33.Pale band on distiflagellomere narrow, less than one-third length of segment.............................................34</p><p>– Pale band on distiflagellomere broader, more than one-third length of segment .......................................... 36</p><p>34.Metafemur with a broad, subapical, dark band (Fig. 39 C); pygophore in lateral view abruptly declivent posteriorly (Fig. 40 J) (Bolivia and Brazil; Fig. 36 F)....................................................... Heraeus pallidinervis sp. nov.</p><p>– Metafemur with a narrower subapical and well-defined dark band (Figs 33C, 39D); pygophore in lateral view slightly declivent posteriorly (Figs 34 B, 40N)................................................................................................35</p><p>35.Inner projections of pygophore subquadrangular and slightly projecting posteriorly (Fig. 40 M) (Argentina, Bolivia, and Paraguay; Fig. 36 G)...................................................................................... Heraeus penai sp. nov.</p><p>– Inner projections of pygophore narrow, elongate, and strongly projecting posteriorly (Fig. 34 A) (Ecuador; Fig. 25 I).............................................................................................................. Heraeus loja sp. nov.</p><p>36.Metafemur with a narrow subapical dark band (Fig. 26 D); pygophore projecting posteriorly and dorsally directed (Fig. 27 N) (Belize and Mexico; Fig. 25 D)........................................................ Heraeus baranowski sp. nov.</p><p>– Metafemur with a broader subapical dark band (Fig. 33 A, B, F); pygophore not projecting posteriorly nor pointed dorsally (Figs 32 J, N, 34N)................................................................................................................37</p><p>37.Small species, less than 5 mm long; pygophore with inner projections of dorsal aperture subquadrangular (Fig. 32 I) (Dominican Republic and Haiti; Fig. 25 G) ................................... Heraeus concolor Slater &amp; Baranowski, 1994</p><p>– Larger species, over 5 mm long; pygophore with inner projections of dorsal aperture elongate, not subquadrangular (Figs 32 M, 34M) .............................................................................................................................. 38</p><p>38.Pygophore rounded; with inner projections of dorsal aperture subtriangular, not well delimited anteriorly (Fig. 32 M); parameres with inner projection narrow (Fig. 32 O, P) (Ecuador; Fig. 25 H) ........ Heraeus ecuatorianus sp. nov.</p><p>– Pygophore declivent and truncate posteriorly in lateral view (Fig. 34 N); inner projections of dorsal aperture elongate, well delimited anteriorly (Fig. 34 M); parameres with inner projection broad (Fig. 34 O, P) (Panama; Fig. 36 C).................................................................................................... Heraeus morganae sp. nov.</p></div>	https://treatment.plazi.org/id/6F64DB1B5E49FFA4FDE4FF05FA4AFC95	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pablo M. Dellapé;María C. Melo;Thomas J. Henry	Pablo M. Dellapé, María C. Melo, Thomas J. Henry (2016): A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species. Zoological Journal of the Linnean Society 177 (177): 29-134, DOI: 10.1111/zoj.12362
6F64DB1B5E4BFFBEFC2EFC27FE22F99F.text	6F64DB1B5E4BFFBEFC2EFC27FE22F99F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heraeus coquilletti Barber 1914	<div><p>HERAEUS COQUILLETTI BARBER, 1914</p><p>(FIGS 3 B, 4D–E, 5B, 6E–H, 7, 8A)</p><p>Heraeus coquilletti Barber, 1914a: 165 –166 (in part); Van Duzee, 1916: 21 (in part); Van Duzee, 1917: 179 (in part); Blatchley, 1934: 8 (in part); Torre Bueno, 1946: 70 (in part); Slater, 1964: 1082; Harrington, 1980: 108; Ashlock &amp; Slater, 1988: 227–228; Slater &amp; Brailovsky, 2000: 332.</p><p>Heraeus nitens Van Duzee, 1914: 8 –9 (synonymized by Van Duzee, 1916).</p><p>Diagnosis</p><p>Heraeus coquilletti can be easily distinguished from all other species of the genus by the shiny head, pronotum, and hemelytra with minute setae. In addition, the distiflagellomeres and hemelytra are uniformly brown and the aedeagus has minute spines on conjunctiva and vesica.</p><p>Redescription</p><p>Body shiny, with minute setae on head, pronotum, scutellum and hemelytra (Fig. 3 B).</p><p>Head: Strongly convex dorsally, orange–brown, smooth, with sparse short recumbent setae. Head transversely rugose ventrally. Eyes slightly protruding, small (larger in specimens from Mexico), not surpassing the dorsal margin of head in lateral view. Ocelli posteri- or to an imaginary line passing through posterior border of eyes. Labium pale brown with short erect setae, extending to mesocoxae. Antennal tubercles divergent. Antennae pale brown, with short recumbent and scattered erect setae, basiflagellomere and distiflagellomere darker, with more abundant setae.</p><p>Thorax: Pronotum dark orange–brown, smooth (Fig. 4 D); posterior lobe paler than anterior lobe; punctate, punctures more conspicuous and abundant on posterior lobe; anterior lobe with sparse, minute, erect setae.</p><p>Pleurae orange–brown. Evaporative area short. Scutellum orange–brown, punctate. Hemelytra shiny, smooth (Fig. 4 E), pale brown, with abundant punctures. Membrane smoky pale brown. Legs: Coxae orange–brown, femora, tibiae, and tarsi pale brown, except apex of each tibia darker, meso- and metafemur slightly darkened apically (Fig. 5 B). Male profemur strongly enlarged with two rows of spines along ventral region, four spines from the apex on posterior row longer and curved anteriorly, with short and sparse setae. Protibia with numerous small tubercles, metatibia with spiniform setae.</p><p>Abdomen: Orange–brown with short sparse recumbent setae. Male genitalia: Pygophore (Fig. 6 E, F) rounded, anterior margin of dorsal aperture rounded, inner projections subrectangular. Parameres: Figure 6 (G, H). Aedeagus (Fig. 8 A): conjunctiva with a few minute spines laterally; lobes of vesica slightly sclerotized, with a few minute spines laterally and distally; processus gonopori long and slender.</p><p>Distribution</p><p>Mexico and USA (Fig. 7).</p><p>Remarks</p><p>Barber (1914a) based his original description on two males and two females from Standford University, California (Nathan Banks Collection), and a female from Brownsville, Texas, USA, that was referred to as H. cinnamomeus by Barber (1948).</p><p>Type material examined</p><p>To ensure nomenclatural stability, we select a male syntype in the USNM collection with the following labels as the lectotype of H. coquilletti: Label 1, ‘Stan[ford] U[.], Cal[.], 24 Dec 1909’; 2 (red) ‘Cotype’; 3, ‘H.G. Barber Collection’; 4 (handwritten), ‘ Heraeus coquilletti Barber [,]♂[,] Cotype’; 5 (red, here added), Lectotype: ♂, Heraeus coquilletti Barber, desig. by Dellapé, Melo, &amp; Henry’. A female labelled as co-type, also in the USNM collection, and a male (missing the head) in the AMNH collection with the same locality data are considered paralectotypes.</p><p>Additional material studied</p><p>MEXICO: Baja California: 1♀, Santo Tomás, 8-VII- 1953, W.J. &amp; J.W. Gertsch (AMNH); 1♀, Mexico, Brassica sp., 18-II-2003, San Diego 0 30359, Miami Port, March 2007 (USNM); Sinaloa: 1♂, 40 mi. N Mazatlan, 27-VII-1952, at light, J.D. Lattin (AMNH); 8♂, 10♀, 26 mi. N Pericos, 13-VIII-1960, P.H. Arnaud Jr, E.S. Ross &amp; D.C. Reutz (CAS); 1♂, 1♀, Mazatlan, 27-VI-1918, Venedio, J.A. Kusche, pres. by B. Preston Clark (CAS); 1♂, 2♀, 1-VI-1918, J.A. Kusche, pres. by B. Preston Clark (CAS); 1♂, 5-VI-1918, J.A. Kusche, pres. by B. Preston Clark (CAS); 2♂, Presidio Riv., 26-IX- [19]18, J.A. Kusche (USNM).</p><p>USA: Arizona: one without abdomen, Phoenix (USNM); California: 2♂, Claremont, Baker (CAS); 1♀, San Joaquin Co., 9 mi. E Stockton, 10-VIII-1971, Chenopodium, D. Shepard, (AMNH); 1♂, 2♀, Oakland, Alameda Co., 25-II-1906, van Dyke (CAS); 1♀, Alameda Co., van Dyke (CAS); 1♀, van Dyke (CAS); 1♂, Burbank, 22-II-1930, C.H. Hicks, (USNM); 1♀, 24-VIII-1930 (USNM); 1♀, Leona Hgts, Alameda, Aug., J.C. Bradley (CAS); 1♂, Alameda, Nov., Koebele (CAS); 1♂, 1♀, S Sonoma Co., 18-II-1911, van Dyke (CAS); 1♀, Sonoma Co., VI-1919, J.R. de la Torre Bueno (CAS); 1♂, 1♀, Sonoma Co., C. Olsen (AMNH); 2♂, Berkeley, 15-I-1922, pres. by E.C. van Dyke (CAS); 1♀, Redwood City, 1-I-1943, P.H. Arnaud (CAS); 1♂, 1♀, San Mateo Co., 8-I-1944, P.H. Arnaud, 20– 161 (CAS); 1♂, 8-I-1944, P.H. Arnaud, 20–161 (USNM); 1♀, Oroville, 13-VII-1926, H.H. Kelfer (CAS); 1♀, Fairfax, 9-III-1919, E.P. Van Duzee (CAS); 1♂, Redding, Shasta Co., 31-VII-1947, elev. 500, H.P. Chandler (CAS); 6♂, 3♀, San Luis Obispo, 28- IV-1919, E.P. van Duzee (CAS); 1♂, Morro beach, 6-IV- 1928, pres. by E.R. Lesch (CAS); 1♂, 2♀, Morro, S.L. Ob. Co., 20-XII-1928, pres. by E.C. van Dyke (CAS); 2♀, Romero Canyon, San Ysidro, 3-I-[19]11, W.M. Wheeler &amp; H.M. Parshley (CAS); 1♀, 3-I- [19]11, W.M. Wheeler &amp; H.M. Parshley, V.D. no. 529 (CAS); 1♂, Ojai, Ventura Co., 16-VI-1957, W.E. Simonds, (AMNH); 1♀ Madera, VI-22-[19]59, C.A. Toschi (AMNH); 2♂, 1♀ Sonoma Co., C.E. Olsen (AMNH); 1♂, H.G. Barber (USNM); 1♀, Vacaville, 6-IX-[19]47, A.T. McClay (USNM); 1♀ Monterrey Co., 4.5 mi. N Parkfield, el. abt 2000 ft, IV-2-1977, K.W. Brown, R.E. Somerby (AMNH); 4♂, 5♀, Los Angeles, Coquillett (USNM); 3♂, Los Angeles, P.R. Uhler (USNM); 3♂, 5♀, San Diego, 30-I, Hubbard (USNM); 1♀, 11-III-[19]14, E.P. Van Duzee (USNM); New Mexico: 1♂, Sandoval Co., San Ysidero, IX- 1967, Wilton Ivie (AMNH).</p></div>	https://treatment.plazi.org/id/6F64DB1B5E4BFFBEFC2EFC27FE22F99F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pablo M. Dellapé;María C. Melo;Thomas J. Henry	Pablo M. Dellapé, María C. Melo, Thomas J. Henry (2016): A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species. Zoological Journal of the Linnean Society 177 (177): 29-134, DOI: 10.1111/zoj.12362
6F64DB1B5E51FFBDFE88F942FB27FC3B.text	6F64DB1B5E51FFBDFE88F942FB27FC3B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heraeus costalis	<div><p>HERAEUS COSTALIS SP. NOV.</p><p>(FIGS 3 C, 5C, 6I–L, 7, 8B)</p><p>Diagnosis</p><p>Head shiny, contrasting with dull pronotum and hemelytra. Distiflagellomeres mostly white, with apex darker. Scutellum without erect setae. Profemur mostly dark brown.</p><p>Heraeus costalis sp. nov. and H. cinnamomeus are the only species of the coquilletti group that have a shiny head, contrasting with a dull pronotum and hemelytra, and the scutellum lacking erect setae. The general colouration of H. costalis sp. nov. is dark brown, with the distiflagellomere mostly white with the apex darker, whereas the general colouration of H. cinnamomeus is cinnamon brown, with distiflagellomere and hemelytra uniformly coloured.</p><p>Description</p><p>Total length 6.27 (Fig. 3 C).</p><p>Head: Convex dorsally, dark reddish brown, shiny, smooth, with short recumbent and long, erect setae.</p><p>Head length 1.34, width 0.91. Postocular length 0.48. Eyes with setae between ommatidea; not surpassing the dorsal margin of head in lateral view. Ocelli posterior to an imaginary line passing the posterior border of eyes. Interocular width 0.46, interocellar width 0.27. Labium pale brown with erect setae, extending to mesocoxae. Labial segment lengths: I 0.93, II 0.96, III 0.88, and IV 0.37. Antennae brown, except scapus darker and distiflagellomere paler with a broad diffuse subbasal yellowish band, and abundant short recumbent and sparse erect setae. Antennal lengths: scape 0.62, pedicel 1.26, basiflagellomere 0.94, and distiflagellomere 1.30. Length of pale band on distiflagellomere 0.72.</p><p>Thorax: Pronotum pruinose, with sparse, short, recumbent, and erect setae. Pronotum dark brown; anterior lobe darker; humeral angles with a small pale spot. Pronotum more conspicuously punctate on posterior lobe. Collar length 0.07, anterior lobe length 0.86, posteri- or lobe length 0.48; anterior lobe width 0.98, posteri- or lobe width 1.50. Pleurae dark brown, acetabular areas reddish brown; punctate, with short recumbent setae. Evaporative area short. Scutellum dark brown, pruinose, punctate, with short recumbent setae. Hemelytra pruinose, with short recumbent setae. Hemelytron brown, claval commisure and about apical half of inner margin of clavus, base of corium, anterior threequarters of lateral margins, and a small subapical spot yellowish; apex of corium and adjacent area of membrane pale, membrane pale brown with a diffuse pale apical spot, veins concolorous. Legs: Coxae and protrochanters reddish brown; profemur reddish brown, except base and apex paler; meso- and metafemur bicoloured, base yellowish and apex brown; tibiae and tarsi pale brown, apex of tibiae darker (Fig. 5 C). With sparse, short, semi-erect setae. Profemur conspicuously incrassate, spines on profemur small except for one spine contrastingly larger. Protibia with small tubercles bearing spiniform setae on inner surface.</p><p>Abdomen: Reddish brown; with abundant, short, recumbent setae. Male genitalia: Pygophore (Fig. 6 I, J) rounded, anterior margin of dorsal aperture slightly rounded, inner projections subquadrangular; declivent posteriorly in lateral view. Parameres: Figure 6 (K, L). Aedeagus (Fig. 8 B): conjunctiva with spines laterally beneath the ejaculatory reservoir; vesica with a few minute spines laterally and two sclerotized lobes with a few minute spines distally; processus gonopori long and slender.</p><p>Distribution</p><p>Costa Rica, Guatemala, Honduras, and Nicaragua (Fig. 7).</p><p>Etymology</p><p>This species is named ‘ costalis ’ to denote the pale costal margin of the hemelytra.</p><p>Type material</p><p>Holotype: ♂, COSTA RICA, Las Canas, 13-VII- 1965, P.J. Spangler (USNM).</p><p>Paratypes: 10♂, 20♀, same data as for holotype (USNM); 1♂, 1♀, same data as for holotype (MLP); 1♀, same data as for holotype (UKANS); 1♀, 9 mi. NW Esparta, 22-VII-1965, P.J. Spangler (USNM). GUATEMALA: 1♂, Los Amates, J.S. Hine (OSUC). HONDURAS: 1♀, intercepted at Atlanta (Georgia) from Honduras, 5-III-1999, on Cucurbita moschata (USNM); 1♂, intercepted at Miami (Florida) from Honduras, 3-I-2005, on Cucumis sativus (USNM); 1♀, intercepted at West Palm Beach (Florida) from Honduras, 19-IV-1993, on Cucurbita sp. (USNM); Comayagua: 2♀, 6-VI-[19]66, J.M. Matta (USNM); 1♂, 13-IV-[19]66, mosquito trap (USNM); 1♂, Tegucigalpa, M. Mojica, 62–6225 (USNM). NICARA- GUA: 1♂, intercepted at Ft. Lauderdale (Florida) from Nicaragua, 21-II-1996 on Citrullus lanatus (USNM); one without abdomen, intercepted at Ft. Lauderdale (Florida) from Nicaragua, 28-III- 1994 on Allium cepa (USNM); 1♀, Altamira, Managua, IX-1970, E. Moore (USNM).</p></div>	https://treatment.plazi.org/id/6F64DB1B5E51FFBDFE88F942FB27FC3B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pablo M. Dellapé;María C. Melo;Thomas J. Henry	Pablo M. Dellapé, María C. Melo, Thomas J. Henry (2016): A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species. Zoological Journal of the Linnean Society 177 (177): 29-134, DOI: 10.1111/zoj.12362
6F64DB1B5E52FFBCFC07FC05FAB2FE80.text	6F64DB1B5E52FFBCFC07FC05FAB2FE80.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heraeus itzelae	<div><p>HERAEUS ITZELAE SP. NOV.</p><p>(FIGS 3 D, 5D, 6M–P, 7)</p><p>Diagnosis</p><p>Antenna with a pale band on distiflagellomere, pronotum with long, erect setae on anterior lobe, and hemelytra with only short recumbent setae.</p><p>Heraeus itzelae sp. nov. and H. setosus sp. nov. have long erect setae on pronotum and scutellum. The distiflagellomere of H. itzelae sp. nov. has a pale band, and the hemelytra have only short recumbent setae; the distiflagellomere of H. setosus sp. nov. lacks a pale band and the hemelytra possess erect setae similar to those on the pronotum and scutellum.</p><p>Description</p><p>Total length 7.03 (Fig. 3 D).</p><p>Head: Strongly convex dorsally, orange brown, shiny, smooth, with long erect and short recumbent setae. Head length 1.03, width 0.96. Postocular length 0.38. Eyes slightly protruding, not surpassing the dorsal margin of head in lateral view. Ocelli posterior to an imaginary line passing the posterior border of eyes. Interocular width 0.51, interocellar width 0.30. Labium pale brown with short erect setae, extending to mesocoxae. Labial segment lengths: I 0.98, II 1.04, III 0.88, and IV 0.40. Antennal tubercles slightly divergent. Antenna pale brown, except for darker scapus, with abundant semi-erect and sparse erect setae; distiflagellomere with a wide pale band. Antennal lengths: scape 0.67, pedicel 1.33, basiflagellomere 1.04, and distiflagellomere 1.28. Length of pale band on distiflagellomere 0.53.</p><p>Thorax: Pronotum pruinose, brown, punctate, posterior lobe with more abundant and larger punctures, and sparse short, recumbent setae; anterior lobe with long erect setae. Collar length 0.13, anteri- or lobe length 0.75, posterior lobe length 0.58; anterior lobe width 1.07, posterior lobe width 1.70. Pleurae dark brown. Evaporative area short. Scutellum brown, pruinose, punctate with long erect and short recumbent setae. Hemelytra pruinose, with short recumbent setae. Colour uniformly pale brown, costal margin on proximal three-quarters, small inner corial spot, and a small subapical spot hardly paler; membrane greyish brown, veins concolorous. Legs: Coxae brown, shiny, remainder of legs pale brown except protrochanter, meso- and metafemur darker distally, tibiae darker apically (Fig. 5 D), setose, with short semi-erect setae, longest on profemur. Profemur with two rows of small spines on ventral surface, one spine larger. Protibia with small tubercles bearing spiniform setae on inner surface.</p><p>Abdomen: Brown with abundant, short, recumbent setae. Male genitalia: Pygophore (Fig. 6 M, N) rounded, anterior margin of dorsal aperture rounded, inner projections quadrangular. Parameres: Figure 6 (O, P). Aedeagus: Conjunctiva not spined, vesica with a few minute spines laterally and on sclerotized lobes; processus gonopori long and slender.</p><p>Variability observed in paratypes</p><p>Similar to holotype in all aspects. Hemelytra varying from almost uniformly pigmented to having a welldifferentiated pattern.</p><p>Distribution</p><p>Mexico (Fig. 7).</p><p>Etymology</p><p>This species is named after our friend and colleague Sara Itzel Montemayor (Museo de La Plata).</p><p>Type material</p><p>Holotype: ♂, MEXICO, intercepted at Nogales Arizona, 28-XI-2011, on Brassica sp. APHIS port no. 0 35394 (USNM).</p><p>Paratypes: 1♀, same data as for holotype, 23-XI- 2011; 1♀, 30-I-2011, APHIS port no. 035412; 1♂, intercepted at Nogales Arizona, from Mexico, 1-XII- 2011, on Brassica chinensis; 1♀, intercepted at Nogales (Arizona) from Mexico, 18-X-2011, with Vigna unguiculata, Aphis Port, no. 0 35304 (USNM); 1♀, intercepted at Nogales (Arizona) from Mexico, 16- XI-2011, with Brassica oleracea, Aphis Port, no. 0 35376 (MLP); 1♂, 3♀, Jalisco, Tepatitlan, 20 mi. SW, 5500 ft, 19-VIII-1954, J.G. Chillcott (CNC); 1♀, Lagos de Moreno, 6400 m a.s.l., 19-VIII-1954, J.G. Chillcott (CNC); Sinaloa: 1♂, Mazatlan, 27-VI-1918, Venedio, J.A. Kusche, pres. by B. Preston Clark (CAS).</p></div>	https://treatment.plazi.org/id/6F64DB1B5E52FFBCFC07FC05FAB2FE80	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pablo M. Dellapé;María C. Melo;Thomas J. Henry	Pablo M. Dellapé, María C. Melo, Thomas J. Henry (2016): A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species. Zoological Journal of the Linnean Society 177 (177): 29-134, DOI: 10.1111/zoj.12362
6F64DB1B5E53FFBAFC77FE58FDD0F96E.text	6F64DB1B5E53FFBAFC77FE58FDD0F96E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heraeus setosus	<div><p>HERAEUS SETOSUS SP. NOV.</p><p>(FIGS 3 E, 5E, 7, 8C, 9A–D)</p><p>Diagnosis</p><p>Antennae uniformly coloured; pronotum, scutellum, and hemelytra with erect setae; labium extending to or slightly beyond metacoxae.</p><p>Heraeus setosus sp. nov. and H. itzelae sp. nov. have long erect setae on pronotum and scutellum. In H. setosus sp. nov., the hemelytra also have erect setae, the distiflagellomere lacks a pale band, and the labium extends to the metacoxae, whereas in H. itzelae sp. nov., the distiflagellomere has a pale band, the hemelytra have only scattered, short, recumbent setae, and the labium is shorter, extending to mesocoxae.</p><p>Description</p><p>Total length 6.46 (Fig. 3 E). Strongly setose species.</p><p>Head: Strongly convex dorsally, brown, coriaceous with abundant recumbent and erect setae. Head length 1.27, width 0.88. Postocular length 0.41. Eyes slightly protruding, not surpassing dorsal margin of head in lateral view. Ocelli posterior to an imaginary line passing the posterior border of eyes. Interocular width 0.45, interocellar width 0.30. Labium light brown with erect setae, extending to metacoxae. Labial segment lengths: I 0.90, II 0.99, III 0.90, and IV 0.37. Antennal colouration yellowish brown. Pedicel with abundant erect setae, equal to or longer than diameter of segment. Antennal lengths: scape 0.62, pedicel 1.60, basiflagellomere, and distiflagellomere missing.</p><p>Thorax: Pronotum pruinose; punctate, those of anterior lobe shallow; dark brown, posterior lobe and humeral angles paler; pruinose, with abundant, long, erect setae. Collar length 0.08, anterior lobe length 0.82, posterior lobe length 0.54; anterior lobe width 1.14, posterior lobe width 1.68. Pleurae brown. Evaporative area short. Scutellum brown, pruinose, punctate, with abundant long erect setae. Hemelytra pruinose, brown, with abundant erect setae shorter than those on scutellum; clavus unicolorous; costal margin pale on proximal three-quarters. Inner corial spot well defined. Membrane brown, veins paler. Legs: Coxae and protrochanter brown; profemur brown with apex paler, rest of legs pale brown; meso- and metafemur slightly darkened distally, tibiae darker at apices (Fig. 5 E). Strongly setose, with long erect and semi-erect setae. Protibia with small tubercles bearing spiniform setae on inner surface.</p><p>Abdomen: Brown with abundant short recumbent setae. Male genitalia: Pygophore rounded, anterior margin of dorsal aperture slightly rounded (Fig. 9 A, B). Parameres: Figure (9C, D). Aedeagus with minute spines; vesical lobes sclerotized; processus gonopori long and slender (Fig. 8 C).</p><p>Variability observed in paratypes</p><p>All antennal segments yellowish brown.</p><p>Distribution</p><p>Honduras and Mexico (Fig. 7).</p><p>Etymology</p><p>This species is named ‘ setosus ’, Latin for bristly, to denote the long, erect dorsal setae.</p><p>Type material</p><p>Holotype: ♂, HONDURAS, Dept Cortes, Lk. Yooa, Agua Azul, U.F. Co., 1-VI-[19]64, F.S. Blanton, A.B. Brose &amp; R.E. Woodruff, blacklight trap (USNM).</p><p>Paratypes: MEXICO: 2♀, Mexico Aircraft, 26-IX-2003 (USNM); Nayarit, 1♀, Nayarit: 18 km SW Compostela, 20-VII-1974, blacklight, M.E. &amp; P.D. Perkins (USNM); 1♀, 20-VII-1974, blacklight, M.E. &amp; P.D. Perkins (MLP); Chiapas: 1♀, 25 mi. E of Tuxtla Gutiérrez, 22-VII- 1964, P.J. Spangler (USNM).</p></div>	https://treatment.plazi.org/id/6F64DB1B5E53FFBAFC77FE58FDD0F96E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pablo M. Dellapé;María C. Melo;Thomas J. Henry	Pablo M. Dellapé, María C. Melo, Thomas J. Henry (2016): A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species. Zoological Journal of the Linnean Society 177 (177): 29-134, DOI: 10.1111/zoj.12362
6F64DB1B5E55FFBAFEFCF8C1FB21F9BC.text	6F64DB1B5E55FFBAFEFCF8C1FB21F9BC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heraeus caliginosus	<div><p>THE CALIGINOSUS GROUP</p><p>Diagnosis</p><p>Included species: Heraeus caliginosus and</p><p>General colour dark brown, with four small pale spots on posterior pronotal lobe, basal half of meso- and metafemur whitish; labium long, usually extending to abdominal sternum II.</p><p>H. dominicanus sp. nov.</p><p>Diagnosis</p><p>Evaporative area short, distance from dorsal margin of auricle to dorsal margin of evaporative area shorter than distance from dorsal margin of evaporative area to dorsal margin of metapleura; posterior half of hemelytra darkened, with a subquadrangular whitish spot on apical region of corium and apex of membrane. Labium long, extending to or surpassing metacoxae.</p></div>	https://treatment.plazi.org/id/6F64DB1B5E55FFBAFEFCF8C1FB21F9BC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pablo M. Dellapé;María C. Melo;Thomas J. Henry	Pablo M. Dellapé, María C. Melo, Thomas J. Henry (2016): A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species. Zoological Journal of the Linnean Society 177 (177): 29-134, DOI: 10.1111/zoj.12362
6F64DB1B5E56FFB8FC21FDB3FD36F9CD.text	6F64DB1B5E56FFB8FC21FDB3FD36F9CD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heraeus dominicanus	<div><p>HERAEUS DOMINICANUS SP. NOV.</p><p>(FIGS 3 K, 5G, 9I–L, 10)</p><p>Diagnosis</p><p>Posterior pronotal lobe uniformly brown; meso- and metafemora reddish, pale basally; labium long, extending to metacoxae.</p><p>The uniformly brown posterior pronotal lobe easily distinguishes H. dominicanus sp. nov. from H. caliginosus . Also, H. dominicanus sp. nov. has a shorter rostrum that extends only to the metacoxae.</p><p>Description</p><p>Total length 6.64 (Fig. 3 K).</p><p>Head: Strongly convex dorsally, brown, smooth, and shiny, with short recumbent and long erect setae dorsally. Head length 1.55, width 0.99. Postocular length 0.59. Eyes slightly protruding, not surpassing the dorsal margin of head in lateral view. Ocelli placed at level of an imaginary line passing posterior border of eyes. Interocular width 0.50, interocellar width 0.26. Labium light brown with erect setae, extending to meso- or metacoxae (pinned through posterior pronotal lobe). Labial segment lengths: I 0.94, II 1.04, III 0.80, and IV 0.24. Antenna light brown, scapus darker, basiflagellomere reddish distally; with abundant short recumbent and sparse, short, erect setae. Antennal lengths: scape 0.67, pedicel 1.44, basiflagellomere 1.28, and distiflagellomere missing.</p><p>Thorax: Pronotum dull, pruinose, uniformly brown, punctate, punctures more conspicuous on posterior lobe; setae short and recumbent, anterior pronotal lobe with long erect setae. Collar length 0.11, anterior lobe length 0.78, posterior lobe length 0.56; anterior lobe width 0.99, posterior lobe width 1.55. Pleurae brown. Evaporative area short. Scutellum brown, punctate, pruinose, with long erect setae. Hemelytra pruinose, with short recumbent setae; general colour brown, darkened on posterior half, with a large whitish subapical corial spot. Membrane brown, with an elongate pale apical spot, veins concolorous. Legs: Femora reddish brown, meso- and metafemur paler basally, tibiae pale reddish brown, tarsi pale brown (Fig. 5 G). Legs with semi-erect and erect setae; protibia with small spines on inner surface.</p><p>Abdomen: Brown, shiny, with abundant short recumbent setae. Male genitalia: Pygophore (Fig. 9 I, J) rounded. Anterior margin of dorsal aperture slightly rounded. Parameres: Figure 9 (K, L). Aedeagus with minute spines, vesical lobes sclerotized; processus gonopori long and slender.</p><p>Variability observed in a female paratype Distiflagellomeres uniformly light brown, without a pale band. Spines of profemur small and restricted to apical half.</p><p>Distribution</p><p>Dominican Republic (Fig. 10).</p><p>Etymology</p><p>The specific epithet ‘ dominicanus ’ is an adjective referring to the country where all of the specimens of this species were collected.</p><p>Type material</p><p>Holotype: ♂, DOMINICAN REPUBLIC: S. Frncsco Mts. [sic], St. Domingo, W.I., IX-[20]05, Aug. Busck (USNM).</p><p>Paratype: 1♀, same data as for holotype (USNM).</p></div>	https://treatment.plazi.org/id/6F64DB1B5E56FFB8FC21FDB3FD36F9CD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pablo M. Dellapé;María C. Melo;Thomas J. Henry	Pablo M. Dellapé, María C. Melo, Thomas J. Henry (2016): A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species. Zoological Journal of the Linnean Society 177 (177): 29-134, DOI: 10.1111/zoj.12362
6F64DB1B5E57FFB8FEE3F970FA78FC06.text	6F64DB1B5E57FFB8FEE3F970FA78FC06.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heraeus guttatus	<div><p>THE GUTTATUS GROUP</p><p>Included species: Heraeus guttatus, H. hollyae, H. steineri sp. nov., H. triguttatus .</p><p>Diagnosis</p><p>Ocelli located posteriorly; posterior half of hemelytra darkened, with a subquadrangular whitish spot on apical region of corium and apex of membrane. Male generally with a larger and more globose anterior pronotal lobe, and with small spines on protibia and mesofemur.</p></div>	https://treatment.plazi.org/id/6F64DB1B5E57FFB8FEE3F970FA78FC06	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pablo M. Dellapé;María C. Melo;Thomas J. Henry	Pablo M. Dellapé, María C. Melo, Thomas J. Henry (2016): A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species. Zoological Journal of the Linnean Society 177 (177): 29-134, DOI: 10.1111/zoj.12362
6F64DB1B5E57FFB5FC1DFC34FACEFE67.text	6F64DB1B5E57FFB5FC1DFC34FACEFE67.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heraeus guttatus	<div><p>HERAEUS GUTTATUS DALLAS, 1852</p><p>(FIGS 3 F, 8E, 11A, 12A–D, 14)</p><p>Orthaea ? guttata Dallas, 1852: 580–581; Dohrn, 1859: 34; Walker, 1872: 121; Stål, 1874: 166.</p><p>Heraeus guttatus: Distant, 1882: 205; Lethierry &amp; Severin, 1894: 191; Distant, 1903: 254, 255 (synonym of Nabis elegans Walker, 1873); Barber, 1939: 354; Ramos, 1946: 27; Wolcott, 1948: 205; Anonymous, 1958: 906; Scudder, 1967: 267–268 (lectotype designation); Slater, 1964: 1083; Scudder, 1970: 100 (raised from synonymy with N. elegans); Slater &amp; O’Donnell 1995: 147; Harrington 1980: 109; Slater 1988: 50; Froeschner 1999: 244; Baranowski, 2005: 139 .</p><p>Diagnosis</p><p>This species can be separated from the other species of the group by the conspicuous pale band on distiflagellomere, the contrastingly darkened apical third of the metafemur, and the pale spots on the posterior pronotal lobe.</p><p>Redescription (Fig. 3 F)</p><p>♀ lectotype.</p><p>Head: Convex dorsally. Colour dark brown, shiny, coriaceous, with short recumbent setae. Eyes protruding, not surpassing the dorsal margin of head in lateral view. Ocelli posterior to an imaginary line passing the posterior border of eyes. Labium pale brown with short erect setae, extending to mesocoxae. Antenna (with only scape of right antenna) pale brown with short recumbent setae.</p><p>Thorax: Pronotum dark brown, dull, posterior lobe paler, with five incomplete longitudinal pale stripes and a small pale spot on humeral angles; collar delimited posteriorly by a sulcus; anterior and posterior lobes with short recumbent setae. Pronotum punctate, punctures more conspicuous on posterior lobe. Pleurae brown. Scutellum dark brown, dull, punctate, with short recumbent setae. Hemelytra (without corium and membrane of right hemelytron) dull, with short recumbent setae. Clavus brown, anterior one-third between median and external rows of punctures pale brown. Corium with lateral margins slightly concave, general colour brown, with a subapical pale brown spot, costal margin pale on proximal three-quarters. Membrane fuscous, veins paler, with a round pale spot at apex. Posterior half of hemelytra dark, with the subapical corial spot and apical spot on membrane conspicuously pale. Legs (only coxa and anterior right leg): Profemur with sparse, short, erect setae, and with spines restricted to apical half. Protibia with short erect setae and small tubercles bearing spiniform setae on apical half.</p><p>Abdomen: Brown, with abundant, short, recumbent setae.</p><p>Variability observed in other material studied</p><p>Pedicellus and basiflagellomere pale brown, basiflagellomere reddish brown at apex; distiflagellomere brown with a pale band; all segments with abundant, short, recumbent and scattered erect setae. Legs: Brown, meso- and metafemur with basal two-thirds yellowish and apical third brown, meso- and metatrochanter, tibiae, and tarsi paler (Fig. 11 A). Males generally with a strongly globose anterior pronotal lobe. Male mesofemur with a row of small spines. Male genitalia: Pygophore (Fig. 12 A, B) rounded, anterior margin of dorsal aperture rounded, posterior lobe more protuberant. Parameres as shown in Figure 12 (C, D). Aedeagus (Fig. 8 E): conjunctiva and vesica lacking spines, lobes of vesica weakly sclerotized; processus gonopori long and slender.</p><p>Distribution</p><p>Belize, Colombia, Dominican Republic, Haiti, Honduras, Jamaica, Mexico (NEW RECORD), Panama, and Puerto Rico (Fig. 14).</p><p>Comments</p><p>Dallas described this species from female(s?) from Jamaica. Distant (1882) reported this species from Guatemala and Jamaica. According to Scudder (1967), the Guatemalan specimens mentioned by Distant are not conspecific with the type, but he did not give additional clarification, and we were unable to examine these specimens.</p><p>Type material examined</p><p>Lectotype: ♀, JAM[AICA], Hope, Saunders, 66-13 (BMNH).</p><p>Additional material studied</p><p>BELIZE: 1♀, British Honduras, San Antonio, VI- 1931, J.J. White (USNM); 1♀, J.J. White, III-1931 (USNM); 1♀, IV-1934, J.J. White (AMNH); 8♂, 8♀, Orange Walk Dist., Chan Chich Lodge, 1-VI-[20]09, BLT, E. Flota (USNM); 1♀, 25-VII-[20]04, BLT, E. Flota (USNM); 1♀, 19-X-[20]03, BLT, H.V. &amp; R.M. Baranowski (USNM); 1♀, 22-X-[20]03, BLT, H.V. &amp; R.M. Baranowski (USNM); 1♀, 25-X-[20]03, BLT, H.V. &amp; R.M. Baranowski (USNM); 1♀, 28-X-[20]03, BLT, H.V. &amp; R.M. Baranowski (USNM); 1♀, 26-VI-[20]08, BLT, H.V. &amp; R.M. Baranowski (USNM); 1♂, 27-VI-[20]08, BLT, H.V. &amp; R.M. Baranowski (USNM); 1♀, 2-VII- [20]09, BLT, H.V. &amp; R.M. Baranowski (USNM); 2♀, 3-VIII-[20]09, BLT, H.V. &amp; R.M. Baranowski (USNM); 1♀, 5-VII-[20]09, BLT, H.V. &amp; R.M. Baranowski (USNM); 2♀, 7-IX-[20]06, BLT, C. Sanabria (USNM); 3♂, 1♀, 13-XI-[20]04, BLT, C. Sanabria (USNM); 2♀, Orange Walk Dist., Gallion Jug, BLT, 31-I-[20]04, B. Miller (USNM); 1♂, 17-VII-[20]04, BLT, B. Miller (USNM); 1♂, 13-IV-[20]05, BLT, B. Miller (USNM); 4♂, 19-VI- [20]05, BLT, B. Miller (USNM); 2♂, 11-X-[20]05, BLT, B. Miller (USNM); 1♀, 9-XI-[20]05, BLT, B. Miller (USNM); 1♀, Cayo District, Benque Viejo del Carmen, riverside N, Mopan River resort, BLT, 27-VI-[20]06, A. Trujillo (USNM); 2♀, 14-V-[20]07, BLT, P. Picon (USNM); 1♂, 12-I-[20]08, BLT, M. Chrysler (USNM); 1♀, 17-X-2009, M. Chrysler (USNM); 1♀, 22-IX- 2009, M. Chrysler (USNM); 1♀, 23-IX-2009, M. Chrysler (USNM).</p><p>COLOMBIA: 1♂, Isla de Providencia, Santa Isabel, 10/27-VIII-[19]69, C.R. Gilbert (USNM).</p><p>DOMINICAN REPUBLIC: Pedernales: 1♂, Cabo Rojo, Alcoa (Cafetería), 9/13-IV-2000, black light, R.E. Woodruff &amp; T.J. Henry (USNM); Altagracia: 1♂, Nisibón, 9-VI-1976, R.E. Woodruff, blacklight trap (USNM); 1♀, R.E. Woodruff, blacklight trap, 8-VI-1976, Baranowski (USNM); 2♂, 2♀, La Romana Prov., La Romana, 13-IX-[19]76, blacklight trap, E. Folch (USNM); 2♂, 18-IX-[19]76, blacklight trap, E. Folch (USNM); 2♂, 1♀, 22-VIII-[19]77, blacklight trap, R.E. Woodruff col. (USNM); 2♀, La Romana Prov., Higueral, 15-VIII-[19]77, blacklight trap, R.E. Woodruff &amp; E. Folch (USNM); 2♀, 17-VIII-[19]77, blacklight trap, R.E. Woodruff &amp; E. Folch (USNM); 1♀, 18-VII- [19]77, blacklight trap, R.E. Woodruff &amp; E. Folch (USNM).</p><p>HAITI: 1♂, 1♀, Trou Caiman, 4-IX-1934, M. Bates (AMNH).</p><p>HONDURAS: 1♀, Dept. Cortes, Puerto Cortes beach, 3-VI-[19]64, BL trap, Blanton &amp; Broce (USNM).</p><p>JAMAICA: 1♀, Kingston Institute, XI-1956, ex. electric light bulb shade, T.H. Farr (USNM); one without abdomen, St. Andrew, Beverly Hills, 5-VIII-1961, R.P. Bengry (USNM); 1♀, Parish of St. Ann, Kellits Mason R. Field Sta. 22-VIII-[19]69, R.E. Woodruff (USNM); 1♀, Parish of St. Ann, 3 mi. W Ocho Rios, 4-VII-1971, under Ficus, Slater, Baranowski &amp; Harrington (USNM); 1♀, Parish of St. Andrew, Hardwar Gap, Hollywell Cabins, 21-V-[19]69, blacklight trap, R.E. Woodruff, (USNM); 1♀, 22-V- [19]69, at light, R.E. Woodruff &amp; P.C. Drummond (USNM); 1♂, St. Andrew, Hardwar Gap, 19-IV- 1959, T.H. Farr (USNM); 1♂, 1♀, Parish of St Andrew, Kingston, Mona, 17-X-1971, blacklight trap, R.M. Baranowski (USNM); 2♂, blacklight trap, R.M. Baranowski, 19-X-1971 (USNM); 1♀, 16-X- 1971, blacklight trap, R.M. Baranowski (USNM); 1♂, 1♀, Hardwar Gap, 4000 ft, 8-VII-1966, Howden &amp; Becker (CNC); 3♂, Pt. Antonio, A.E. Wigte (AMNH); 1♂, Parish of St. Andrew, Irishtown, 7-VII-1971, black light trap., J.A. Slater, R.M. Baranowski &amp; J.E. Harrington, H. gutattus det. J. Slater 1978 comp. type (AMNH); one without abdomen, 5-VII-1971, black light trap., J.A. Slater, R.M. Baranowski &amp; J.E. Harrington (USNM); 1♀, Parish of St. Catherine, Worthy Park, 11-V-[19]69, blacklight trap, R.E. Woodruff, compared with broken lectotype of H. guttatus (Dallas) T. Henry det. (USNM); 2♂, blacklight trap, R.E. Woodruff (USNM); 1♀, 17-V- [19]69, blacklight trap, R.E. Woodruff (USNM); 1♀, Parish of St. Catherine, Worthy Park, 10-VI-[19]75, blacklight trap, R.E. Woodruff, compared with lectotype of H. guttatus (Dallas) T. Henry det. (USNM); 3♂, 1♀, 24-VIII-[19]69, blacklight trap, R.E. Woodruff (USNM); 1♂, 1♀, 9-V-[19]69, blacklight trap, R.E. Woodruff (USNM); 5♂, 2♀, blacklight trap, R.E. Woodruff (USNM); 4♂, 1♀, 11-V-[19]69, blacklight trap, R.E. Woodruff (USNM); 1♂, 2♀, 13- V-[19]69, blacklight trap, R.E. Woodruff (USNM); 1♂, Parish Manchester De Carteret College Mandeville, 12-V-[19]69, blacklight trap, K. Stanton (USNM); 1♂, 18-V-[19]69, blacklight trap, K. Stanton (USNM); 11♂, 4♀, Parish of St. Andrew, Bamboo Lodge, near Irishtown, 23-VI-1972, blacklight trap, 2500 ft, R.M. Baranowski (USNM); 3♂, 3♀, 19-VI-1972, blacklight trap, 2500 ft, R.M. Baranowski (USNM); 1♂, 1♀, VII-1972, R.M. Baranowski (USNM); 1♂, Balaclava, 15-IV-1909, A.E. Wigte (AMNH); 1♂, 1♀ one without abdomen, 24-IV-1909, A.E. Wigte (AMNH); 1♀, 20-IV-1909, A.E. Wigte (AMNH); 2♂, 1♀, Baron Hill, Trelawney, 16/30-III-[19]31, E.L. Bell (USNM); 1♀, one nymph, Trelawny, Windsor Estate, 12-II- 1960, T.H. Farr (USNM); 1♂, Mandeville, Oct., Cockerell (USNM).</p><p>MEXICO: 1♀, Cozumel Island, Celerain Pt., Sta. 20, 21-IV-1960, J.F.G. Clarke (USNM); 4♀, one without abdomen, Camp. 16 mi. N Champoton, 9-VIII- 1974, C.W. &amp; L.B. O’Brien &amp; G.B. Marshall (USNM).</p><p>PANAMA: 2♂, 3♀, C[anal] Z[one], Coco Solo Hospital, 14-V-1975, D. Engleman (AMNH); 1♀, D. Engleman, 13-VI-[19]72, LT (USNM).</p></div>	https://treatment.plazi.org/id/6F64DB1B5E57FFB5FC1DFC34FACEFE67	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pablo M. Dellapé;María C. Melo;Thomas J. Henry	Pablo M. Dellapé, María C. Melo, Thomas J. Henry (2016): A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species. Zoological Journal of the Linnean Society 177 (177): 29-134, DOI: 10.1111/zoj.12362
6F64DB1B5E5AFFB4FCC7FE20FD89F9E8.text	6F64DB1B5E5AFFB4FCC7FE20FD89F9E8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heraeus hollyae Baranowski 2005	<div><p>HERAEUS HOLLYAE BARANOWSKI, 2005</p><p>(FIGS 3 G, 11B, 12E–H, 13)</p><p>Heraeus hollyae Baranowski, 2005: 141 (in Baranowski &amp; Slater, 2005) .</p><p>Diagnosis</p><p>Posterior pronotal lobe nearly uniformly dark, with two short yellow lines. Meso- and metafemur bicoloured, with basal two-thirds yellowish and apical third brown.</p><p>Specimens of H. hollyae share with H. guttatus the contrastingly darkened apical third of metafemur, but differ in having the posterior pronotal lobe nearly uniformly dark, with only two short yellowish lines.</p><p>Description of characters used in phylogenetic</p><p>analysis (Fig. 3 G)</p><p>Head: Coriaceous. Eyes not surpassing the dorsal margin of head in lateral view. Ocelli posterior to an imaginary line passing the posterior border of eyes. Labium extending to mesocoxae. Distiflagellomere with a pale band.</p><p>Thorax: Anterior and posterior pronotal lobe without long setae. Posterior pronotal lobe nearly uniformly dark, with two short yellow lines. Males generally with a strongly globose anterior pronotal lobe. Scutellum without erect setae. Inner corial spot present. Costal margin pale on proximal three-quarters, with a subapical corial spot. Veins of membrane paler and with a pale apical spot. Posterior half of hemelytra dark with the subapical corial spot and apical spot of membrane conspicuously pale. Legs: Profemur dark with apices paler, apical half of metafemur darker (Fig. 11 B). Profemur with sparse short setae. Mesofemur and protibia spinose.</p><p>Abdomen, male genitalia: Pygophore (Fig. 12 E, F) rounded. Anterior margin of dorsal aperture rounded. Parameres: Figure 12 (G, H). Aedeagus: conjunctiva and vesica unspined, lobes of vesica weakly sclerotized, without spines; processus gonopori long and slender.</p><p>Distribution</p><p>Antigua, Dominican Republic (NEW RECORD), and Haiti (NEW RECORD) (Fig. 13).</p><p>Type material examined</p><p>Holotype: ♂, ANTIGUA B., W.I., Collins, 20-VIII- [19]95 (under Bucida sp.), R.M. &amp; H.V. Baranowski, J.A. Slater (FSCA).</p><p>Paratypes: 7♂, 7♀ same data as for holotype (2♂, 4♀, AMNH; 5♂, 3♀ USNM).</p><p>Additional material studied</p><p>HAITI: 1♂, 1♀, Trou Cayman, 4-IX-1934, M. Bates (AMNH).</p><p>DOMINICAN REPUBLIC: Pedernales: 1♀, Cabo Rojo, Alcoa (cafetería), 9/13-IV-2000, blacklight, R.E. Woodruff &amp; T.J. Henry (USNM).</p></div>	https://treatment.plazi.org/id/6F64DB1B5E5AFFB4FCC7FE20FD89F9E8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pablo M. Dellapé;María C. Melo;Thomas J. Henry	Pablo M. Dellapé, María C. Melo, Thomas J. Henry (2016): A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species. Zoological Journal of the Linnean Society 177 (177): 29-134, DOI: 10.1111/zoj.12362
6F64DB1B5E5BFFB3FE88F97DFB28FBA2.text	6F64DB1B5E5BFFB3FE88F97DFB28FBA2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heraeus steineri	<div><p>HERAEUS STEINERI SP. NOV.</p><p>(FIGS 3 I, 11C, 12I–L, 13)</p><p>Diagnosis</p><p>Legs whitish, coxa pale brown; posterior pronotal lobe mottled with yellow; hemelytra brown, mottled with yellow at base.</p><p>The uniformly whitish legs easily distinguish this species from the others in the group.</p><p>Description</p><p>Total length 6.56 (Fig. 3 I).</p><p>Head: Slightly convex dorsally, brown, shiny, coriaceous, with short recumbent and scattered erect setae. Head length 1.30, width 0.93. Postocular length 0.46. Eyes not surpassing the dorsal margin of head in lateral view. Ocelli placed posterior to an imaginary line passing the posterior border of eyes. Interocular width 0.34, interocellar width 0.22. Labium pale brown with short erect setae, nearly extending to mesocoxae. Labial segment lengths: I 0.80, II 0.88, III 0.64, and IV 0.35. Antennae pale brown, except apices of pedicel and basiflagellomere slightly darker, with short recumbent and scattered erect setae, distiflagellomere brown, with a medial pale band.</p><p>Antennal lengths: scape 0.48, pedicel 1.31, basiflagellomere 1.18, and distiflagellomere 1.28. Length of pale band on distiflagellomere 0.38.</p><p>Thorax: Pronotum brown, posterior lobe paler with four incomplete longitudinal pale stripes; anterior and posterior lobe with short recumbent setae. Pronotum pruinose, anterior lobe impunctate, posterior lobe punctate. Collar length 0.08, anterior lobe length 0.77, posterior lobe length 0.51; anterior lobe width 1.04, posterior lobe width 1.52. Pleurae brown, acetabular areas reddish brown; punctate, with short recumbent setae. Scutellum brown, except two medial pale spots and apex yellowish; pruinose, punctate, with short recumbent setae. Hemelytra pruinose, with short recumbent setae; posterior half dark with the subapical corial spot and apical spot of membrane conspicuously pale. Clavus brown, with anterior one-third between median and external rows of punctures pale brown. Corium brown with a subapical whitish spot, costal margin pale on proximal three-quarters. Membrane fuscous, veins paler with an apical rounded whitish spot. Legs: Coxae pale brown, remainder of legs whitish (Fig. 11 C), with sparse short erect setae. Profemur with spines on apical twothirds. Protibia slightly curved at base, with small tubercles bearing spiniform setae on apical half. Mesofemur with three or four spines.</p><p>Abdomen: Brown, with abundant short recumbent setae. Male genitalia: Pygophore (Fig. 12 I–J) rounded, anterior margin of dorsal aperture rounded, inner projections elongate, more produced posteriorly; slightly declivent posteriorly in lateral view. Parameres: (Fig. 12 K, L). Aedeagus without spines, and with two sclerotized lobes; processus gonopori long and slender.</p><p>Variability observed in paratypes</p><p>The posterior pronotal lobe varies from having four longitudinal pale stripes to only irregular pale areas, but it is never homogeneously or evenly coloured.</p><p>Distribution</p><p>Turks and Caicos Islands and the Bahamas (Fig. 13).</p><p>Etymology</p><p>This species is named in honour of Warren E. Steiner, one of the collectors of this species.</p><p>Type material</p><p>Holotype: ♂, TURKS AND CAICOS ISLANDS,</p><p>Providenciales, King’s Town, 21°48 ′ N, 72°14 ′ W, 30-I-</p><p>1998, at black light in sandy scrub forest, W.E. Steiner</p><p>&amp; J.M. Swearingen (USNM).</p><p>Paratypes: 1♀, same data as for holotype (USNM); 1♂, 4♀, Providenciales Grace Bay Hills, 21°48 ′ N, 72°13 ′ W, 27-I-1998, at blacklight in sandy scrub forest, W.E. Steiner &amp; J.M. Swearingen (USNM); 1♀, at blacklight in sandy scrub forest, W.E. Steiner &amp; J.M. Swearingen (MLP); 1♂, 5♀, Providenciales Isl., 23-XI-[19]94, R.M. &amp; H.V. Baranowski (USNM); 1♀, 22-XI-[19]94, R.M. &amp; H.V. Baranowski (USNM); 1♀, Providenciales Isl., Grace Bay, 20-X-93, R.M. &amp; H.V. Baranowski (USNM); 2♀, North Caicos Island, 20-VII-[19]93, B.L.T., A. Swan (USNM); 1♂, 3♀, Whitby, North Caicos Island, 23-X-[19]93, B.L.T., R.M. &amp; H.V. Baranowski (USNM); 1♂, 26-XI-[9]94, B.L.T., R.M. &amp; H.V. Baranowski (USNM); 1♂, 21-X-[9]93, B.L.T., R.M. &amp; H.V. Baranowski (USNM); 1♀, North Caicos Island, Pelican Beach Hotel, 31-V-[19]91, BL Trap, H.V. &amp; R.M. Baranowski (USNM); 3♂, 10♀, Bambarra, North Caicos Island, 12-XII-[19]93, B.L.T., B.M. Riggs (USNM); 2♂, 2♀, B.L.T., B.M. Riggs (MLP). BAHAMAS ISLANDS: 1♂, Mayaguana Isl. 1-VIII-1963, blacklight trap, C. Murvosh (USNM); 1♀, 25-VIII-1967, blacklight trap, C. Murvosh (USNM); 1♀, 26-VIII-1963, blacklight trap, C. Murvosh (USNM); 1♂, 3♀, 3-VIII-[19]63, blacklight trap, C. Murvosh (USNM); 1♀, 20-VIII-[19]63, blacklight trap, C. Murvosh (USNM); 1♂, 2♀, 24-VIII-[19]63, blacklight trap, C. Murvosh (USNM); 1♀, 25-VIII- [19]63, blacklight trap, C. Murvosh (USNM); 1♂, 5♀, 26-VIII-[19]63, blacklight trap, C. Murvosh (USNM); 1♂, 4♀, 27-VIII-[19]63, blacklight trap, C. Murvosh (USNM); 3♂, 1♀, 28-VIII-[19]63, blacklight trap, C. Murvosh (USNM); 1♂, 1♀, 30-VIII-[19]63, blacklight trap, C. Murvosh (USNM).</p></div>	https://treatment.plazi.org/id/6F64DB1B5E5BFFB3FE88F97DFB28FBA2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pablo M. Dellapé;María C. Melo;Thomas J. Henry	Pablo M. Dellapé, María C. Melo, Thomas J. Henry (2016): A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species. Zoological Journal of the Linnean Society 177 (177): 29-134, DOI: 10.1111/zoj.12362
6F64DB1B5E5CFFB0FC82FB97FE1CFB27.text	6F64DB1B5E5CFFB0FC82FB97FE1CFB27.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heraeus triguttatus	<div><p>HERAEUS TRIGUTTATUS GUÉRIN-MÉNEVILLE, 1857</p><p>(FIGS 3 H, 11D, 12M–P, 14)</p><p>Lygaeus (Plociomerus) triguttatus Guérin-Méneville, 1857: 400 .</p><p>Heraeus triguttata: Stål, 1862: 315; Stål, 1874: 147; Uhler, 1886: 15; Lethierry &amp; Severin, 1894: 191.</p><p>Plociomerus triguttatus: Walker, 1873b: 52 .</p><p>Heracus [sic] trigutatus [sic]: Valdés Ragués, 1914: 98.</p><p>Heraeus triguttatus: Barber, 1914b: 514; Van Duzee, 1916: 21; Van Duzee, 1917: 179; Blatchley, 1926: 389– 390; Torre Bueno, 1946: 70; Barber, 1954a: 4; Barber, 1954b: 342; Scudder, 1958: 146–147; Barber &amp; Ashlock, 1960: 121; Frost, 1964: 135; Slater, 1964: 1084–1085; Harrington, 1980: 109; Slater, 1988: 40– 41, 52, 59; Ashlock &amp; Slater, 1988: 228; Slater &amp; Baranowski, 1990: 130, 132–134; Baranowski &amp; Slater, 1998: 79, 87; Baranowski, 2005: 144; Cervantes Peredo, 2005: 371, 381.</p><p>Diagnosis</p><p>Distiflagellomere with a diffuse pale band, the legs yellowish brown, with the distal third of meso- and metafemur slightly darkened, the posterior pronotal lobe uniformly coloured, and the hemelytra reddish brown, with the apex of corium and most of the membrane darker.</p><p>Heraeus triguttatus is most closely related to H. steineri sp. nov., whereas this last species has uniformly whitish legs, the posterior pronotal lobe is mottled with yellow, and the hemelytra are mottled brown with yellow basally.</p><p>Redescription (Fig. 3 H)</p><p>Head: Convex dorsally, postocular region elongate; brown, shiny, coriaceous, with short recumbent and erect setae dorsally. Eyes not surpassing the dorsal margin of head in lateral view. Ocelli placed posterior to an imaginary line passing the posterior border of eyes. Labium pale brown, with erect setae, extending to mesocoxae. Antennae pale brown, with abundant recumbent and scattered short erect setae. Distiflagellomere with a pale band, diffuse in some specimens.</p><p>Thorax: Pronotum brown, posterior pronotal lobe paler; dull, with scattered short recumbent and long erect setae, shorter on posterior lobe. Males generally with a strongly globose anterior pronotal lobe. Scutellum brown, pruinose, with long erect setae. Pleurae brown. Hemelytra pruinose, dull, with short recumbent setae; posterior half of hemelytra dark with the subapical corial spot and apical spot of membrane conspicuously pale. Membrane fuscous, veins concolorous. Legs: Coxae brown, profemur pale brown, rest of legs yellowish brown, meso- and metafemur slightly darkened distally (Fig. 11 D). Profemur with sparse short setae. Protibia and male mesofemur spinose.</p><p>Abdomen: Pale reddish brown, with abundant short recumbent setae. Male genitalia: Pygophore (Fig. 12 M, N) rounded, anterior margin of dorsal aperture rounded, inner projections elongate, more pronounced posteriorly. Parameres with a tubercle at base of blade on outer surface (Fig. 12 O, P). Aedeagus without spines; vesical lobes sclerotized; processus gonopori long and slender.</p><p>Distribution</p><p>Cuba, Dominican Republic, Jamaica, USA, and the West Indies (Bahamas, South Bimini Island, Grand Cayman Island, and Turks &amp; Caicos Islands; Fig. 14).</p><p>Type material</p><p>According to the original description, the type series consists of three specimens from Cuba. After an unsuccessful search of the Muséum National d’Histoire Naturelle in Paris by Dr Eric Guilbert, we have concluded that the specimens of Lygaeus (Plociomerus) triguttatus are lost. To ensure nomenclatural stability, we are designating the following female neotype (in good condition with only the right antenna missing) deposited in the USNM collection: label 1, ‘Baraguá, Cuba, X-24-26’; label 2, ‘T P R F Ent. No. 188’; label 3, ‘at light’; label 4, ‘C.F. Stahl collector’; label 5 (red, here added), ‘Neotype ♀: Lygaeus triguttatus Guérin- Méneville, desig. by Dellapé, Melo, &amp; Henry’. There is also a male (in good condition, but with both distiflagellomeres missing) in the USNM with the same label data, except for the date X-19-25.</p><p>Additional material studied</p><p>CUBA: 1♀ (abdomen on separate label) (CAS); 1♂, (CAS); 1♂, Baraguá, XII-13, 29, Grasses, L.D. Christenson (AMNH); 1♂, Baraguá, X-24-[19]26, TPRF ent. N°188, at light, C.F. Stahl (AMNH); 1♀, Baraguá, 5-X-[19]25, TPRF ent. N°188, at light, C.F. Stahl (AMNH); 1♀, one without abdomen (USNM); 2♂, one without abdomen, Cuba (USNM); 1♂, at light, TPRF Ent. #188, 10-X-[19]25, C.F. Stahl (USNM); 1♀, Cayamas, 5–6 m, E.A. Schwarz (USNM); 1♂, Colon, Mat., 15-XI-1922, A. Comas (USNM); 1♀, Guantamano Bay, 14-XI-[19]72, blacklight trap, S. Calhoun (USNM); 1♂, blacklight trap, 16-XI-[19]72, S. Calhoun (USNM); 1♀, Navy Base, Caravella Pt., blacklight trap, 17-XI- [19]72, S. Calhoun (USNM); one without abdomen, blacklight trap, 10-I-[19]73, S. Calhoun (USNM); 1♂, 1♀, Guantanamo, La Fa, Imiaz, VII-[19]90, a la luz, L.F. Farmas (USNM); 1♂, Cayo Sabina, Pinares de Mayari, 645 m a.s.l., VII-[19]90, a la luz, L.F. Armas &amp; V. Becker (USNM); 1♀, Holguin, Pinares de Mayari, Loma de la Mensura, 780 m a.s.l., 7-VII-[19]90, L.F. Armas &amp; V. Becker (USNM).</p><p>BAHAMAS: 14♂, 13♀, Great Abaco, Pinewoods Nursery, Marsh Harbor, 17-X-[19]90, BL trap, R. Keys (USNM); 2♂, 2♀, 22-XII-[19]90, BL trap, R. Keys (USNM); 1♂, 11-X-[19]90, BL trap, R. Keys (USNM); 2♂, Long Island, Stella Maris Resort, 30-VI- [19]94, B.L.T., R.M. &amp; H.V. Baranowski (USNM); 1♀, Eleuthera Rainbow Bay, 18-XII-[19]75, blacklight trap, J. Wiley (USNM); 1♀, N. Eleuthera, Current Club, 10-I-[19]77, blacklight trap, R.M. Baranowski (USNM); 2♂, 3♀, New Providence, Isl. Nassau, 5-IV-1953, Van Voast–AMNH Bahama Islands Exped., E.B. Hayden (AMNH); 2♀, Abaco Cays/Allans Cay, 9-V-1953, Van Voast–AMNH Bahama Islands Exped., E.B. Hayden &amp; G.B. Rabb (AMNH); 1♂, Crooked Island, Landrail Point, 5-III-1953, Van Voast–AMNH Bahama Islands Exped., E.B. Hayden &amp; L. Giovannoli (AMNH); 2♀, South Bimini Island, B.W.I., V-1951, Cazier &amp; Gertsch (AMNH); 1♂, South Bimini Island, 18-VIII-1951, C. &amp; P. Vaurie (USNM); 1♀, Mangrove Cay, Andros Island, V/VI-1917, Wm. M. Mann (AMNH).</p><p>GRAND CAYMAN ISLAND: 1♂, 22-I-[19]89, BL trap, P. Fitzgerald (USNM); 1♂, 17-I-[19]89, BL trap, P. Fitzgerald (USNM); 1♀, 30-X-[19]89, BL trap, P. Fitzgerald (USNM); 1♂, 28-V-[19]89, BL trap, P. Fitzgerald (USNM); 1♂, 10-VI-[19]90, BL trap, P. Fitzgerald (USNM); 1♀, 23-I-[19]93, BL trap, P. Fitzgerald (USNM); 1♀, 31-I-[19]93, BL trap, P. Fitzgerald (USNM); 1♂, 19-I-[19]92, BL trap, P. Fitzgerald (USNM); 2♂, 23-I-[19]88, BL trap, P. Fitzgerald (USNM); 1♀, 13-XII-[19]87, BL trap, P. Fitzgerald (USNM); 1♂, 5-VIII-[19]88, BL trap, P. Fitzgerald (USNM); 1♂, 12-III-[19]88, BL trap, P. Fitzgerald (USNM); 1♂, 23-V-[19]88, BL trap, P. Fitzgerald (USNM); 2♂, 1♀, South Sound, 2-VIII- [19]86, BL trap, F. Fitzgerald (USNM); 2♀, 12-II- [19]86, BL trap, F. Fitzgerald (USNM); 1♀, 16-XI- [19]86, BL trap, F. Fitzgerald (USNM); 1♀, Savannah, 12-XII-[19]89, R.M. Baranowski (USNM); 1♀, Pine Villa Hotel, 1-XI-[19]96, BLT, H.V. &amp; R.M. Baranowski (USNM); 2♂, 2-XI-[19]96, BLT, H.V. &amp; R.M. Baranowski (USNM).</p><p>JAMAICA: 1♂, St. James, Montego Bay, 23-IV- 1960, Baranowski (USNM).</p><p>TURKS &amp; CAICOS ISLANDS: 1♀, Cays, 3.5 m SW of North Caicos Island, 28-II-1953, Van Voast– AMNH Bahama Island Exped., E.B. Hayden &amp; L. Giovannoli (AMNH).</p><p>USA: Florida: 1♂, Archbold Biol. Stn., Highlands Co., 27-I-[19]59, S.W. Frost (PSUC); 1♂, Isla Morada, upper Matecumba Key, 15-IV-1963, J.G. &amp; B.L. Rozen (AMNH); 1♂, Naples, 19-X-1979, Museum Paris 1994 coll., G. Perrault (MNHN); 1♂, 1♀, Paradise Key, III-[19]19, H. Barber (USNM); 1♀, 3-III-[19]19, Schwartz &amp; Barber (USNM); 2♂, 3-III-[19]19, H.G. Barber (USNM); 1♀, Naranja, 24-IX-1960, R.M. Baranowski (USNM); 1♂, Dade Co., Ross &amp; Castello Ham., 4-X-[19]70, R.M. Baranowski (USNM); 1♂, Prinston, 5-I-[19]70, black light trap, R.M. Baranowski (USNM); 1♂, Miami Dade Co., Black Point Marina Buttonwood, 25°32.4 ′ N, 80°19.7 ′ W, 26-IX-2006, T. Dobbs (USNM); 1♀, T. Dobbs (USNM); 3♀, Dade Co., Agr. Res. &amp; Ed. Ctr., Homestead, 21- X-[19]76, blacklight trap, R.M. Baranowski (USNM); one without abdomen, blacklight trap, R.M. Baranowski, 21-IV-[19]69 (USNM); 2♂, 2♀, Homestead, 26-XI-[19]69, R.M. Baranowski (USNM); 7♂, 4♀, 7-X-[19]69, R.M. Baranowski (USNM); 2♀, 2-IV-[19]70, R.M. Baranowski (USNM); 1♂, 1♀, 11- IV-[19]69, R.M. Baranowski (USNM); 2♂, 2♀, 18- XI-[19]69, R.M. Baranowski (USNM); 1♀, 15-X- [19]69, R.M. Baranowski (USNM); 1♂, 18-XI- [19]68, R.M. Baranowski (USNM); 1♀, Dade Co., 11 km SW Florida city, 30-IV-1986, W. Steiner &amp; D. Bogar (USNM); 2♂, 1♀, Goulds, 24-I-[19]41, in light trap, O.D. Link (USNM); 7♂, 7♀, Monroe Co., Everglades National Park, Flamingo Praire, 8-IV- [19]72, blacklight trap, R.M. Baranowski (USNM); 11♂, 6♀, 23-III-[19]70, blacklight trap, R.M. Baranowski, (USNM); 2♂, 1♀, 4-V-[19]70, blacklight trap, R.M. Baranowski (USNM); 9♂, 22♀, 28-III-[19]70, blacklight trap, R.M. Baranowski (USNM); 2♀, 27-II[19]72, blacklight trap, R.M. Baranowski (USNM); 1♂, 5-IV-[19]58, R.E. Woodruff (USNM); 2♂, 2♀, Monroe Co., Tavernier, Indian Mound Trail, MM89, 30-Xi- [19]87, BL Trap, S. Jewel (USNM); 12♂, 10♀, Monroe Co., North Key Largo Key, 5-III-[19]82, J.A. Slater &amp; R.M. Baranowski (USNM); 1♀, Monroe Co., Stock Island, 23-XII-[19]65, blacklight trap, F.A. Buchanan (USNM); 1♂, 1♀, 28-V-[19]68, blacklight trap, F.A. Buchanan (USNM); one without abdomen, 5-VIII- [19]62, blacklight trap, F.A. Buchanan (USNM); 1♂, 25-X-[19]84, blacklight trap, F.A. Buchanan, (USNM); 2♂, 1♀, Mosq. Control Hdqrs., 10-XI-[19]71, W.H. Pierce (USNM); 21♂, 10♀, Monroe Co., Big Pine Key, Key Deer Refuge, 5-XI-[19]82, blacklight trap, S. Klett (USNM); 13♂, 11♀, 15-XI-[19]82, blacklight trap, S. Klett (USNM); 1♂, 10-XI-[19]82, blacklight trap, S. Klett (USNM).</p></div>	https://treatment.plazi.org/id/6F64DB1B5E5CFFB0FC82FB97FE1CFB27	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pablo M. Dellapé;María C. Melo;Thomas J. Henry	Pablo M. Dellapé, María C. Melo, Thomas J. Henry (2016): A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species. Zoological Journal of the Linnean Society 177 (177): 29-134, DOI: 10.1111/zoj.12362
6F64DB1B5E5FFFB0FEDDFB0AFF29F8D9.text	6F64DB1B5E5FFFB0FEDDFB0AFF29F8D9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heraeus illitus	<div><p>THE ILLITUS GROUP</p><p>Included species: Heraeus annulatus sp. nov., H. antennalis sp. nov., H. chamamecinus sp. nov., H. illitus, H. inca sp. nov., H. panamaensis sp. nov., H. similis sp. nov., H. spinosus sp. nov., H. splendens sp. nov., and H. tiputini sp. nov.</p><p>Diagnosis</p><p>Often setose species; head coriaceous, setose; collar without a well-developed furrow posteriorly (except H. annulatus sp. nov.); evaporative area extended, distance from dorsal margin of auricle to dorsal margin of evaporative area subequal to or longer than distance from dorsal margin of evaporative area to dorsal margin of metapleura; hemelytra with heterogeneous colouration pattern; profemur pale brown with dark-brown spots, if predominantly dark, the dorsal region clearly pale; male mesofemur without spines; meso- and metatibia generally with a subproximal dark band; aedeagus with well-developed spines.</p></div>	https://treatment.plazi.org/id/6F64DB1B5E5FFFB0FEDDFB0AFF29F8D9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pablo M. Dellapé;María C. Melo;Thomas J. Henry	Pablo M. Dellapé, María C. Melo, Thomas J. Henry (2016): A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species. Zoological Journal of the Linnean Society 177 (177): 29-134, DOI: 10.1111/zoj.12362
6F64DB1B5E5FFF8CFC00FF17FB8BFA75.text	6F64DB1B5E5FFF8CFC00FF17FB8BFA75.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heraeus annulatus	<div><p>HERAEUS ANNULATUS SP. NOV.</p><p>(FIGS 15 A, 16A, 17A–D, 18)</p><p>Diagnosis</p><p>Labium extending to mesocoxae. Pedicel with abundant erect setae, equal to or longer than diameter of segment; distiflagellomere with a pale band subbasally. Collar delimited by a furrow; with long erect setae only on anterior pronotal lobe; profemur mottled, with the base and a subapical band darker; metatibia yellowish. The combination of these last four characters allows us to separate H. annulatus sp. nov. from the other species of the genus.</p><p>Description (Fig. 15 A)</p><p>Total length 5.80. Strongly setose species.</p><p>Head: Flattened dorsally, dark brown clypeus paler, shiny, with abundant short recumbent and long erect setae. Head length 1.15, width 0.93. Postocular length 0.41. Eyes not surpassing the dorsal margin of head in lateral view; with long erect setae between ommatidea. Ocelli placed at level of an imaginary line passing the posterior border of eyes. Interocular width 0.45, interocellar width 0.27. Labium pale brown with short erect setae, almost extending to mesocoxae. Labial segment lengths: I 0.53, II 0.72, III 0.61, and IV 0.35. Antennae pale brown except apex of basiflagellomere and distiflagellomere darker, distiflagellomere with a pale band sub-basally; setae short recumbent and semi-erect, pedicel with abundant erect setae, equal to or longer than diameter of segment; basiflagellomere slightly clavate. Antennal lengths: scape 0.48, pedicel 1.04, basiflagellomere 0.86, distiflagellomere 1.04. Length of pale band on distiflagellomere 0.40.</p><p>Thorax: Anterior pronotal lobe dark brown, posteri- or lobe reddish brown with irregular pale areas, humeral angles yellowish; pruinose, with short recumbent and long erect setae, longer on anterior lobe. Pronotum punctate, punctures more conspicuous on posterior lobe. Collar delimited by a furrow, length 0.08, anterior lobe length 0.53, posterior lobe length 0.50; anterior lobe width 0.91, posterior lobe width 1.41. Pleurae brown, acetabular areas reddish brown; punctate, with abundant, short, recumbent setae. Evaporative area extended. Scutellum dark brown, pruinose, punctate, with short recumbent and long erect setae, as on the anterior lobe. Hemelytra pruinose, with short semierect setae. General colour pale brown, costal margin pale on proximal three-quarters, corium darker distally with a subapical whitish spot and a rounded pale spot internally. Membrane fuscous, with an apical pale spot, veins paler (Fig. 15 A). Legs: Coxae and protrochanter brown, rest of leg whitish, profemur mottled, with base and a subapical band brown; meso- and metafemur with a subapical brown band; tibiae brown apically (Fig. 16 A); setae long and erect, longer on profemur. Profemur with spines restricted to apical two-thirds.</p><p>Abdomen: Brown, with abundant, short, recumbent setae. Male genitalia: Pygophore (Fig. 17 A, B) rounded, anterior margin of dorsal aperture rounded, inner projections elongate, more produced posteriorly; slightly declivent posteriorly in lateral view. Parameres: Figure 17 (C, D). Aedeagus: conjunctiva with spines laterally, beneath the ejaculatory reservoir; two lobes sclerotized with a few spines distally; processus gonopori long and slender.</p><p>Distribution</p><p>Belize, Costa Rica, Honduras, Mexico, and Panama (Fig. 18).</p><p>Etymology</p><p>The specific ephitet ‘ annulatus ’ is a Latin adjective meaning ringed, and refers to the presence of a subapical band on the profemur.</p><p>Type material</p><p>Holotype: ♂, BELIZE: Cayo District, Benque Viejo, riverside N, Mopan R. Resort, 21-VI-[20]08, BLT, R.M. &amp; H.V. Baranowski (USNM).</p><p>Paratypes: 3♂, 2♀, same data as for holotype (USNM); 1♂, 1♀, same data as for holotype, 25-VI-[20]08 (MLP); 2♀, San Antonio, VI-1931, J.J. White (USNM); 1♂, Orange Walk, Dist. Gallion Jug, BLT, 17-VII-[20]04, B. Miller (USNM); 1♂, 18-XII-[20]04, BLT, B. Miller (USNM); 2♂, 2♀, 19-VI-[20]05, BLT, B. Miller (USNM); 1♂, 3-V-[20]05, BLT, B. Miller (USNM); 1♀, 27-VI- [20]08, BLT, B. Miller (USNM); 1♂, Orange Walk, Rio Bravo, Cons. Area, well Trail, 9-VII-1996, C.W. &amp; L.B. O’Brien (USNM); 1♂, C.W. &amp; L.B. O’Brien, Lagunitas Trail, 15-VII-1996 (USNM); 1♂, Texas Camp., 21-VII-1996, C.W. &amp; L.B. O’Brien (USNM); 8♂, 10♀, Orange Walk Dist., Chan Chich Lodge, 1-VI-[20]09, BLT, E. Flota (USNM); 2♂, 2♀, BLT, E. Flota (MLP); 1♂, 18-X-[20]09, BLT, E. Flota (USNM); 1♀, 30-X-[20]09, BLT, E. Flota (USNM); 1♂, 1♀, 1-VII- [20]09, BLT, E. Flota (USNM); 1♀, 30-IV-[20]05, C. Sanabria (USNM); 3♀, Cayo District, Benque Viejo, riverside N, Mopan R. Resort, 22-VI-[20]08, BLT, R.M. &amp; H.V. Baranowski (USNM); 3♂, one without abdomen, 23-VI-[20]08, BLT, R.M. &amp; H.V. Baranowski (USNM); 1♀, 24-VI-[20]08, BLT, R.M. &amp; H.V. Baranowski (USNM); 1♂, 22-IX-[20]09, M. Chrysler (USNM); 1♂, 8-X-2009, M. Chrysler (USNM); 1♀, 10-X-[20]09, M. Chrysler (USNM); 1♂, 12-X-2009, M. Chrysler (USNM); 1♂, 16-X-2009, M. Chrysler (USNM); 1♂, 1♀, 25-X-2009, M. Chrysler (USNM); 1♂, 22-X-[20]06, A. Trujillo (USNM); 1♀, 25-X-[20]06, A. Trujillo (USNM); 1♂, 16-I-[20]07, P. Picon (USNM).</p><p>Additional material studied</p><p>COSTA RICA: 1♀, intercepted at Dover (Delaware, USA) from Costa Rica, 17-I-2000, on Musa paradisiaca (USNM); 1♀, La Selva OTS Station, 19-V-1988, J.E. O’Donnell, UVL (UCMS); 1♀, 22/23-V-1988, blacklight, J.E. O’Donnell (UCMS); 1♂, Guanacaste Prov., Palo Verde OTS Station, 22/23-VI-[1]988, blacklight, J.E. O’Donnell (UCMS).</p><p>HONDURAS: 1♂, 1♀, S.C.Y., Lago Yojoa, 21-VII- 1974, C.W. &amp; L. O’Brien &amp; Marshall (AMNH); 1♂, Dept. Cortes, Puerto Cortes, 3-VI-[19]69, blacklight trap, Blanton et al. (USNM); 2♀, Com., 5 km NW Taulabe, Rio Jaitique, 2-VIII-[19]77, O’Brien &amp; Marshall (USNM).</p><p>MEXICO: Veracruz: 1♀, UNAM, Los Tuxtlas Stac. Biol., N. Catemaco, 16/19-IX-1989, E. Barrera, T.J. Henry &amp; I.M. Kerzhner (USNM).</p><p>PANAMA: C[anal] Z[one]: 1♀, Gatun, III-1930, Tres Rios Plantation, T.O. Zschokke (CAS); 1♂, Coco Solo Hosp., 14-V-1975, D. Engleman, light trap (AMNH); 1♂, Barro Colorado, XI-1934, M. Bates (AMNH); 1♀, Barro Colorado Island, 30-IV-1962, H. Ruckes (AMNH); 3♀, 24/28-V-[19]64, W.D. &amp; S.S. Duckworth (USNM); 1♂, IV-1941, J. Zetek, lots 41–7231 (USNM); 1♀, Cano Saddle, Gatun L., 3-V-[19]23, R.C. Shannon (USNM); 1♂, 12-V-[19]23, R.C. Shannon, close’s Pltn (USNM); Chiriqui: 1♀, Bugaba, Champion, from coll. Biol. C. Am. (USNM).</p></div>	https://treatment.plazi.org/id/6F64DB1B5E5FFF8CFC00FF17FB8BFA75	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pablo M. Dellapé;María C. Melo;Thomas J. Henry	Pablo M. Dellapé, María C. Melo, Thomas J. Henry (2016): A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species. Zoological Journal of the Linnean Society 177 (177): 29-134, DOI: 10.1111/zoj.12362
6F64DB1B5E63FF8BFC0CFA25FB56FE24.text	6F64DB1B5E63FF8BFC0CFA25FB56FE24.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heraeus antennalis	<div><p>HERAEUS ANTENNALIS SP. NOV.</p><p>(FIGS 15 B, 16B, 17E–H, 18)</p><p>Diagnosis</p><p>Strongly setose. Labium extending to mesocoxae. Basiflagellomere clavate. Anterior pronotal lobe dark brown, posterior lobe paler with humeral angles concolorous. Profemur pale brown, mottled with brown. Hemelytra uniformly pale brown, with a white subapical corial spot. Aedeagus spinose, with spines forming an arc on sclerotized vesical lobes.</p><p>Heraeus antennalis sp. nov. and H. splendens sp. nov. are the only two species in the genus with the basiflagellomere clavate. Heraeus antennalis sp. nov. is strongly setose, with mottled profemora, and the posterior pronotal lobe uniformly pale brown, whereas H. splendens sp. nov. is less setose, with a pale invert- ed V-shaped mark on the posterior pronotal lobe, the profemora are dark but paler dorsally, and the outer margin of clavus is pale.</p><p>Description (Fig. 15 B)</p><p>Total length 6.36. Strongly setose.</p><p>Head: Reddish brown, with an elongate neck and short recumbent and long erect setae. Head length 1.46, width 0.99. Postocular length 0.62. Eyes not surpassing the dorsal margin of head in lateral view. Ocelli placed at level of an imaginary line passing the posterior border of eyes. Interocular width 0.53, interocellar width 0.27. Labium brown, with erect setae, extending to mesocoxae. Labial segment lengths: I 0.66, II 0.80, III 0.62, and IV 0.32. Antenna brown, pedicel and base of basiflagellomere paler, with short recumbent and sparse short erect setae, distiflagellomere with a narrow sub-basal whitish band. Basiflagellomere clavate. Antennal lengths: scape 0.43, pedicel 1.23, basiflagellomere 0.96, and distiflagellomere 1.06. Length of pale band on distiflagellomere 0.32.</p><p>Thorax: Anterior pronotal lobe dark brown, posteri- or lobe paler, humeral angles concolorous; collar impunctate, delimited posteriorly by a sulcus; anteri- or and posterior lobes with long erect setae, posterior lobe with short recumbent setae. Pronotum pruinose, posterior lobe punctate. Collar length 0.10, anterior lobe length 0.55, posterior lobe length 0.55; anterior lobe width 0.94, posterior lobe width 1.47. Pleurae dark brown, acetabular areas paler, punctate, with short recumbent setae. Evaporative area extended. Scutellum brown darker at base and along a longitudinal line to the apex, with long erect setae. Hemelytra pale brown, pattern as in Figure 15 (B); inner corial spot diffuse, subapical corial spot present. Membrane brown, with veins paler and with a pale apical spot. Legs: Pale brown except mottled profemur, metafemur with a subapical dark band; and meso- and metatibia with a subproximal dark band, apex of tibiae darker (Fig. 16 B). Profemur with long erect setae.</p><p>Abdomen: Dark brown to reddish brown, with abundant short recumbent setae. Male genitalia: Pygophore (Fig. 17 E, F) rounded, anterior margin of dorsal aperture rounded. Parameres: Figure 17 (G, H). Aedeagus spinose, with spines forming an arc on sclerotized vesical lobes; processus gonopori long and slender.</p><p>Distribution</p><p>Ecuador (Fig. 18).</p><p>Etymology</p><p>The specific ephitet is a noun and refers to the clavate basiflagellomere of the antenna.</p><p>Type material</p><p>Holotype: ♂, ECUADOR: Yanayacu, 400 m a.s.l., IX/ X-1977, L.E. Peña G. (USNM).</p></div>	https://treatment.plazi.org/id/6F64DB1B5E63FF8BFC0CFA25FB56FE24	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pablo M. Dellapé;María C. Melo;Thomas J. Henry	Pablo M. Dellapé, María C. Melo, Thomas J. Henry (2016): A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species. Zoological Journal of the Linnean Society 177 (177): 29-134, DOI: 10.1111/zoj.12362
6F64DB1B5E64FF89FC34FE1AFE03FCB5.text	6F64DB1B5E64FF89FC34FE1AFE03FCB5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heraeus chamamecinus	<div><p>HERAEUS CHAMAMECINUS SP. NOV.</p><p>(FIGS 8 F–G, 15C, 16C, 17I–L, 19)</p><p>Diagnosis</p><p>Strongly setose. Ocelli located posteriorly. Distiflagellomere without or with only a weakly developed pale band. Profemur irregularly pigmented. Metafemur with a conspicuous dark apical band. Posterior margin of pygophore in dorsal view weakly bilobed; aedeagus with spines laterally to the ejaculatory reservoir and on the sclerotized vesical lobes forming an arc.</p><p>Heraeus chamamecinus sp. nov. shares with H. illitus and H. similis sp. nov. a highly setose body, and the presence of a slightly bilobed posterior margin of pygophore, but it can be distinguished by the absence of a well-defined pale band on distiflagellomere, and the short labium not surpassing the mesocoxae.</p><p>Description (Fig. 15 C)</p><p>Total length 6.18. Strongly setose.</p><p>Head: Convex dorsally, dark brown, shiny, coriaceus, with short recumbent and long erect setae. Head length 1.25, width 1.01, postocular length 0.46. Eyes protruding, not surpassing dorsal margin of head in lateral view. Ocelli placed posterior to an imaginary line passing posterior border of eyes. Interocular width 0.51, interocellar width 0.34. Labium pale brown, apex of article IV darker; with erect setae, extending to mesocoxae. Labial segment lengths: I 0.80, II 0.88, III 0.78, and IV 0.32. Antennal tubercles parallel. Antennae pale brown, except distal third of basiflagellomere and distiflagellomere darker; distiflagellomere without pale band; all segments with short recumbent and long erect setae, pedicel with abundant erect setae, equal to or longer than diameter of segment. Antennal lengths: scape 0.48, pedicel 1.18, basiflagellomere 0.96, distiflagellomere 1.02.</p><p>Thorax: Pronotum pruinose; dark brown, posterior lobe with irregular pale areas, and humeral angles dark brown with a small pale spot. Surface punctate, more conspicuous on posterior lobe. Anterior lobe with long erect and short recumbent setae, posterior lobe with erect setae shorter than setae of anterior lobe. Collar length 0.08, anterior lobe length 0.62, posteri- or lobe length 0.54; anterior lobe width 0.99, posteri- or lobe width 1.57. Pleurae brown. Evaporative area extended. Scutellum brown, pruinose, punctate, setae similar to setae on anterior lobe. Hemelytra pruinose, with erect and semi-erect setae. Colour pale brown, costal margin pale on proximal three-quarters, with a subapical corial spot, dark pattern as in Figure 15 (C); membrane with a pale, ovoid, apical spot, veins pale. Legs: Pale brown, coxae, profemur, and pretarsus darker; profemur irregularly pigmented; metafemur with a conspicuous dark band apically, less evident on mesofemur. Meso- and metatibia with a subproximal dark band, apex of tibiae darker (Fig. 16 C). Legs with abundant erect setae, longest on profemur. Profemur with spines restricted to apical two-thirds. Protibia slightly curved.</p><p>Abdomen: Brown with abundant short recumbent setae. Male genitalia: Pygophore (Fig. 17 I, J) subquadrangular, with posterior margin slightly bilobed and depressed towards aperture, anterior margin of dorsal aperture rounded, inner projections subquadrangular. Parameres: Figure 17 (K, L). Aedeagus (Fig. 8 F, G): conjunctiva with an anterior lobe basal to ejaculatory reservoir; vesica with spines lateral to ejaculatory reservoir, forming an arc on the sclerotized vesical lobes; processus gonopori long and slender.</p><p>Variability observed in paratypes</p><p>Some specimens show a weakly developed pale band on distiflagellomere.</p><p>Distribution</p><p>Argentina and Brazil (Fig. 19).</p><p>Etymology</p><p>The specific epithet is a noun in aposition and refers to the typical music style (Chamamé) from the Argentinian province of Corrientes.</p><p>Type material</p><p>Holotype: ♂, ARGENTINA, Corrientes: Ituzaingo, Reserva Santa María, 30-X-2003, T. luz, M.C. Melo (MLP).</p><p>Paratypes: 22♂, 14♀, same data as for holotype (MLP); 6♂, 8♀, same data as for holotype (USNM); 1♀, same data as for holotype, P.M. Dellapé (MLP); 2♂, 4♀, same data as for holotype, M. Chayle (MLP); 1♂, 31-X-2003, M.C. Coscarón col. (MLP); 2♂, 3♀, 29- X-2003, P.M. Dellapé col. (AMNH); 2♂, 1♀, 29-X- 2003, P. Dellapé col. (USNM); 6♂, 7♀, 29-X-2003, P.M. Dellapé col. (MLP); 1♂, 1♀, 27-IV-2003, M.C. Melo col. (MLP); 3♂, P.M. Dellapé col. (MLP); Tucumán: 1♂, Ciudad Universitaria, 800 m a.s.l., 18- II-[19]59, J.F.G. Clarke (USNM); 1♀, Depto Burruyacú, Rio Salas, 12-II-1982, R. Golbach (IFML); 1♀, Horco Molle, 1/18-IX-1966, L. Stange (IFML); Entre Rios: 5♂, X-1994 (USNM); 1♀, X-[19]93, E. Foerster (USNM); 1♂, 1♀, X-[19]93, Peña &amp; Ugarte (USNM). BRAZIL: 1♀, Banhado Purane, 14- II-[19]02, E.G. &amp; E.A. Monroe (CNC); Paraná: 1♂, 58 km S Curitiba, Sao Jose dos Pinhais, 24-XI- 1985, T.J. Henry (USNM).</p></div>	https://treatment.plazi.org/id/6F64DB1B5E64FF89FC34FE1AFE03FCB5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pablo M. Dellapé;María C. Melo;Thomas J. Henry	Pablo M. Dellapé, María C. Melo, Thomas J. Henry (2016): A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species. Zoological Journal of the Linnean Society 177 (177): 29-134, DOI: 10.1111/zoj.12362
6F64DB1B5E66FF88FEBCFCECFBD1FABD.text	6F64DB1B5E66FF88FEBCFCECFBD1FABD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heraeus illitus Distant 1882	<div><p>HERAEUS ILLITUS DISTANT, 1882</p><p>(FIGS 8 H, 15D, 16D, 17M–P, 19)</p><p>Heraeus illitus Distant, 1882: 205; Distant, 1893: 397; Lethierry &amp; Severin, 1894: 191; Scudder, 1967: 268 (lectotype designation); Slater, 1964: 1083; Slater &amp; O’Donnell, 1995: 147; Harrington, 1980: 109; Froeschner, 1999: 244; Slater &amp; Brailovsky, 2000: 332; Baranowski &amp; Slater, 2005: 142.</p><p>Diagnosis</p><p>Strongly setose. Ocelli located posteriorly. Distiflagellomere with a pale band. Labium extending to abdominal sternum II. Profemur irregularly pigmented; metafemur with a darker subapical band. Posterior margin of pygophore in dorsal view slightly bilobed; aedeagus with conjunctiva and vesica with a few spines laterally, and lobes sclerotized posteriorly, with spines forming an arc.</p><p>Heraeus illitus shares with H. chamamecinus sp. nov. and H. similis sp. nov. a strongly setose body and a slightly bilobed posterior margin of pygophore. Heraeus illitus and H. caliginosus are the only species of the genus that have a long labium exceeding the metacoxae.</p><p>Redescription (Fig. 15 D)</p><p>Strongly setose.</p><p>Head: Convex dorsally, dark brown, clypeus paler, shiny, with short recumbent setae. Eyes protruding, not surpassing the dorsal margin of head in lateral view. Ocelli placed posteriorly to an imaginary line passing the posterior border of eyes. Labium pale brown, extending to base of abdomen, with short erect setae. Antennal tubercles parallel. Antennae pale brown except apex of pedicel, basiflagellomere and distiflagellomere darker, distiflagellomere with a pale band subbasally; all segments with short recumbent and semierect setae, pedicel with abundant erect setae, equal to or longer than diameter of segment.</p><p>Thorax: Pronotum brown, pruinose, posterior lobe with irregular pale areas and a pale spot on humeral angles, anterior and posterior lobes with short recumbent and long erect setae. Pronotum punctate, punctures more conspicuous on posterior lobe. Pleurae brown, with semi-erect setae. Evaporative area extended. Scutellum dark brown, pruinose, punctate, with short recumbent, and long erect setae. Hemelytra pruinose, with abundant erect setae. General colour pale brown, costal margin pale on proximal three-quarters. Corium darker distally, with a subapical whitish spot and a rounded pale spot internally. Membrane testaceous, with an apical rounded pale spot, veins paler. Legs pale brown, coxae darker, profemur darker and irregularly pigmented; metafemur with a subapical darker band, only slightly evident on mesofemur. Meso- and metatibiae with a subproximal dark band, apices darker (Fig. 16 D), with long erect setae, longest on profemur. Profemur with spines restricted to apical threequarters.</p><p>Abdomen: Brown, with abundant, short, recumbent, and longer erect setae. Male genitalia: Pygophore subquadrangular (Fig. 17 M, N), posterior margin slightly bilobed; anterior margin of dorsal aperture rounded, inner projections large and subrectangular. Parameres as shown in Figure 17 (O, P). Aedeagus (Fig. 8 H): conjunctiva and vesica with spines laterally; lobes sclerotized posteriorly, with spines forming an arc; processus gonopori long.</p><p>Distribution</p><p>Guatemala, Mexico, Panama, Trinidad; Argentina, Belize, Bolivia, Brazil, Colombia, Costa Rica, French Guiana, Guatemala, Honduras, Nicaragua, Peru, Suriname, and Venezuela (NEW RECORDS) (Fig. 19).</p><p>Type material examined</p><p>The type series of this species consists of specimens from Guatemala and Panama; Scudder (1967) designated a male from San Juan, Verapaz, Guatemala, as the lectotype. Lectotype ♂, San Juan, Vera Paz, Champion (BMNH).</p><p>Additional material studied</p><p>BELIZE: 1♂, Cayo District, Benque Viejo, riverside N, Mopan R. Resort, 16-I-[20]08, BLT, R.M. &amp; H.V. Baranowski (USNM); 1♂, 21-VI-[20]08, BLT, R.M. &amp; H.V. Baranowski (USNM); 3♂, 22-VI-[20]08, BLT, R.M. &amp; H.V. Baranowski (USNM); 7♂, 6♀, 22-VI- [20]06, BLT, R.M. &amp; H.V. Baranowski (USNM); 1♂, 3♀, 23-VI-[20]08, BLT, R.M. &amp; H.V. Baranowski (USNM); 1♂, 24-VI-[20]08, BLT, R.M. &amp; H.V. Baranowski (USNM); 1♀, 1-X-2009, M. Chrysler (USNM); 1♂, 4-X-</p><p>2009, M. Chrysler (USNM); 1♀, 7-X-2009, M. Chrysler</p><p>(USNM); 1♀, Orange Walk Dist., Chan Chich Lodge,</p><p>12-IV-[20]04, BLT, R.M. &amp; H.V. Baranowski (USNM);</p><p>1♀, 13-XI-[20]04, C. Sanabria (USNM); 1♂, 21-IV-</p><p>[20]04, E. Flota (USNM); 2♀, 1-VI-[20]09, E. Flota</p><p>(USNM); 1♂, Orange Walk Dist., Gallon Jung., BLT,</p><p>7-IV-[20]04, B. Miller (USNM).</p><p>BOLIVIA: Cochabamba: 1♀, Chapare Prov., Cristal- Mayu, 14-X-1950, L.E. Pena (USNM); La Paz: 2♂, 1♀, Rio Coroico, 1200 m a.s.l., 24/26-XI-[19]84, L.E. Pena (USNM); 2♀, Mapiri, N La Paz, 10/16- VIII-[19]89, L.E. Peña (USNM); 1♂, 1♀, Yungas, Pte. Mururata, 1200–1600 m a.s.l., 24/26-XII-1984, L.E. Pena (USNM); Santa Cruz: 1♂, San Esteban Prov., Muyurina, 49 km N Santa Cruz, 1120 ft, 12- X-[19]59, blacklight trap, R.B. Cumming (USNM); 1♂, 2♀, 5-X-[19]59, blacklight trap, R.B. Cumming (USNM); 2♀, 26-X-[19]59, blacklight trap, R.B. Cumming (USNM).</p><p>BRAZIL: 1♀, Monat. IV-1935, P. Sandig (USNM); 1♀, DF, 35 km N Brasilia, nr Planaltina, 15°35 ′ S, 47°42 ′ W, 1000 m a.s.l., 17-XI-1997, T.J. Henry (USNM); Mato Grosso: 1♀, Corumba, H.G. Barber (USNM); Rondonia: 1♂, 62 km S Ariquemes, Faz. Rancho Grande, 10°32 ′ S, 62°48 ′ W, 11/22-XI-1991, B.C. Ratcliffe (USNM); 1♂, 30-III/10-IV-1992, collected with mercury vapour &amp; blacklight, J.E. Eger (USNM); 20♂, 23♀, 62 km SW Ariquemes, nr Fzda. Rancho Grande, 5/17- X-1993, blacklight trap, J.E. Eger (USNM).</p><p>COLOMBIA: 1♀, Apiay, Meta, 14-VII-1965, J.A. Ramos (USNM).</p><p>COSTA RICA: 1♀, intercepted on fruit of from Costa Rica.</p><p>ECUADOR: 1♀, Guay., Olon, 29-II-1976, blacklight, J. Cohen, Ecuador Peace Corps, Smithsonian Institution Aquatic Insect Survey (USNM); 1♀, Catacocha, Macara, 400–650 m a.s.l., 14/15-VIII-1977, L.E. Pena G. (USNM).</p><p>GUATEMALA: 1♂, Cerro Zunil, 4000 ft, Champion, BCA, Hem. 1, H. illitus (BMNH).</p><p>HONDURAS: 1♂, Com., 21 km NW Siquatepeque Ch., 5 Rd, 1-VIII-1977, O’Brians &amp; Marshall (AMNH).</p><p>MEXICO: Tabasco: 1♂, Teapa, Feb., H.H.S., BCA, Hem. 1, H. illitus . (BMNH); Campeche: 1♀, 19 mi. SW Esparcedga, 4-VIII-1974, C.W. &amp; L.B. O’Briens &amp; G.B. Marshall (AMNH); 1♀, Esparcedga, 25-IV- 1962, trap light, F. Islas S. (USNM); Chiapas: 1♂, Rio Tulija, 48 km S Palenque, 17-V-1981, C.M. &amp; O.S. Flint Jr. (USNM).</p><p>NICARAGUA: 1♀, Puerto Cabezas, VII-1971, J. Maldonado C. (USNM).</p><p>PANAMA: 1♂, Panama aircraft, 6-II-[20]02 (USNM); 3♂, 1♀, V. de Chiriqui, 25–4000 ft Champion, BCA, Hem. 1, H. illitus (BMNH); Chiriqui: 1♀, Rovira, 5-VII-[19]64, 2500 ft, mosquito light trap, A. Bruse (USNM); 1♂, Bugaba, Champion (USNM); Canal Zone: 1♀, Madden Dam, 7-VI-[19]73, light trap, D. Engleman (USNM); 1♂, Coco Solo Hospital, 9°21 ′ N, 79°51 ′ W, 23-V-[19]72, D. Engleman (USNM); 1♀, Barro Colorado Island, IV-1941, at light, J. Zetek (USNM); 1♀, Puerto Bello, 19-II-[19]12, A. Busck (USNM); 1♂, Trinidad Riv., 4-V-[19]11, A. Busck (USNM).</p><p>PERU: 1♂, 2♀, Cuzco, Pilcopata, 600 m a.s.l., 8/10- XII-1979, premontane moist forest, J.B. Heppner (USNM).</p><p>SURINAME: 1♀, 40 km E Paramaribo, 22/30-VII- [19]75, D. Engleman (AMNH); 1♀, 22/30-VII- [19]75, D. Engleman (USNM).</p><p>TRINIDAD: 1♂, Curepé, Santa Margarita circular Rd, 7-VII-1971, blacklight trap, F.D. Bennett (USNM); 1♂, 6-VII-[19]71, black light trap, F.D. Bennett (USNM); 1♀, 23-VII-1971, blacklight trap, F.D. Bennett (USNM); 2♀, 9-VI-[19]75, blacklight trap, F.D. Bennett (USNM); 1♂, 15-XI-[19]71, black light trap, F.D. Bennett (USNM); 1♂, 1♀, 28-VII-[19]75, blacklight trap, F.D. Bennett (USNM); 1♀, 29-VI- [19]75, blacklight trap, F.D. Bennett (USNM); 1♂, 1♀, Maracas Valley, 1 mi. N St Joseph, 3-IV-[19]79, blacklight trap, L. Du Bruijn (USNM); 1♀, 8-IV- [19]79, blacklight trap, L. Du Bruijn (USNM); 1♂, 1♀, B.W.I., Arima Valley, 3-V-1952 (AMNH), 1♂, Curepé, Santa Margarita Circular Rd., 23-VII- 1971, blacklight trap, F.D. Bennett (AMNH); 1♀, 15- VI-[19]75, blacklight trap, F.D. Bennett (AMNH); 3♂, 2♀, Simla Biol. Sta. Trinidad, at light, M. Emsley (AMNH); 1♀, Port of Spain, R.J. Crew (USNM); 3♀, Trinidad, IV/V-1976, J. Carayon (MNHN).</p><p>VENEZUELA: 1♀, Acarigua Est. Portuguesa, VI- [19]81 (USNM).</p></div>	https://treatment.plazi.org/id/6F64DB1B5E66FF88FEBCFCECFBD1FABD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pablo M. Dellapé;María C. Melo;Thomas J. Henry	Pablo M. Dellapé, María C. Melo, Thomas J. Henry (2016): A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species. Zoological Journal of the Linnean Society 177 (177): 29-134, DOI: 10.1111/zoj.12362
6F64DB1B5E67FF86FC5FFA8CFC60F912.text	6F64DB1B5E67FF86FC5FFA8CFC60F912.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heraeus inca	<div><p>HERAEUS INCA SP. NOV.</p><p>(FIGS 15 E, 16E, 20A–D, 22)</p><p>Diagnosis</p><p>Ocelli posterior. Postocular region subequal to twice the length of an eye. Labium extending to mesocoxae. Distiflagellomere with a pale band. Pronotum with long erect setae only on anterior pronotal lobe. Profemur darker on distal half, paler dorsally; meso- and metafemur with a subapical dark band; meso- and metatibia with a narrow band on basal region. Aedeagus with two pairs of lobes on vesica, with the posterior lobe large and strongly sclerotized posteriorly. Processus gonopori long, widening towards apex.</p><p>Heraeus inca sp. nov. is similar to H. tiputini sp. nov. in having the ocelli placed posteriorly, the profemur paler dorsally, the metafemur darker apically (with a dark band instead of darker distally in a few specimens of H. inca sp. nov.), and the processus gonopori long and widening towards apex. Heraeus inca sp. nov. shows a narrow band on basal region of metatibia that is absent in H. tiputini sp. nov., in which the metatibia is uniformly yellowish. In H. inca sp. nov., the anteri- or margin of dorsal aperture of pygophore is slightly rounded with lateral regions angulate, and the posterior vesical lobe is large and strongly sclerotized posteriorly, whereas in H. tiputini sp. nov., the anterior margin of dorsal aperture of pygophore is rounded, and the vesical lobes are not sclerotized.</p><p>Description (Fig. 15 E)</p><p>Total length 6.18. Setose species.</p><p>Head: Convex dorsally, dark brown, dull, with short recumbent and long erect setae. Head length 1.02, width 0.93, postocular length 0.56. Eyes protruding, not surpassing the dorsal margin of head in lateral view. Ocelli posterior to an imaginary line passing posterior border of eyes. Interocular width 0.46, interocellar width 0.26. Labium pale brown, segments III and IV darker, with short erect setae; nearly extending to mesocoxae. Labial segment lengths: I 0.80, II 0.83, III 0.64, and IV 0.37. Antennal tubercles parallel. Antennae pale brown, except for dark-brown scape and distal half of basiflagellomere; distiflagellomere with a pale band; setae short and recumbent. Antennal lengths: scape 0.50, pedicel 1.38, basiflagellomere 1.14, and distiflagellomere missing.</p><p>Thorax: Pronotum pruinose, anterior lobe dark brown, posterior lobe brown with two longitudinal incomplete stripes medially and two diffuse lateral pale stripes. Anterior lobe of pronotum with long erect and short recumbent setae, posterior lobe with short recumbent and semi-erect setae, longest on lateral margins. Collar length 0.08, anterior lobe length 0.62, posterior lobe length 0.56; anterior lobe width 0.93, posterior lobe width 1.42. Pleurae dark brown. Evaporative area extended. Scutellum dark brown, pruinose, punctate, with long erect and short recumbent setae. Hemelytra pruinose, with erect and semi-erect setae; pale brown, costal margin pale on proximal threequarters, with a subapical corial spot; colour pattern as in Figure 15 E. Membrane fuscous with an apical pale spot, veins paler. Legs: Legs pale brown, profemur darker on distal half and paler dorsally; meso- and metafemur with an apical dark band; meso- and metatibia with a thin dark band on basal region, apex of tibiae darker (Fig. 16 E) tarsus I missing. Profemur elongate, slightly incrassate, with spines restricted to distal two-thirds. Legs with erect setae, longest on profemora.</p><p>Abdomen: Brown, with abundant, short, recumbent setae. Male genitalia: Pygophore (Fig. 20 A, B) rounded, anterior margin of dorsal aperture slightly rounded with lateral regions angulate, inner projections more pronounced posteriorly. Parameres:</p><p>Variability observed in paratypes</p><p>Most of the paratypes have a dark apical band on the metafemur, but in a few specimens the band is reduced and appears subapical; even the apical bands sometimes have a narrowly pale apex. Both apical and subapical metafemoral bands occur in specimens from the same series.</p><p>Distribution</p><p>Bolivia, Brazil, Ecuador, and Peru (Fig. 22).</p><p>Etymology</p><p>The specific epithet is a noun in aposition and refers to the Inca people from Ecuador and Peru.</p><p>Type material</p><p>Holotype: ♂, PERU: Avispas, X-1952, L.E. Peña (USNM).</p><p>Paratypes: BOLIVIA: La Paz: 1♂, 3♀, Guanay, X/XI- 1993, L.E. Peña (USNM); 2♂, 4♀, Guanay, Tres Esteros, 19/25-VIII-[19]89, L.E. Peña (USNM); 1♂, 1♀, L.E. Peña (MLP); 1♂, 1♀, Guanay, Uyapi, X-1994, G. Castillo (USNM); 1♂, 1♀, G. Castillo (MLP). BRAZIL: Mato Grosso: 1♀, NE Caceres, 21-VII- [19]88 (USNM). ECUADOR: Napo: 1♂, Limoncocha, 15-VI-1977, collected in Malaise trap, P.J. Spangler &amp; D.R. Givens (USNM); Zamora-Chinchipe: 1♂, Yanzaza, 16-VI-1976, A. Langley et al., Ecuador Peace Corps, Smithsonian Institution Aquatic Insect Survey (USNM); Tungurahua: 1♂, ∼ 29 km N of Puyo, 22- I-1974, 3800 ft, R.M. King, on Austroeupatorium inulaefolium, compositae at USNM coll. (USNM); 1♂, 2♀ 1, Gualaquiza, 1000 m a.s.l., 25/28-VIII-1977, L.E. Pena G. (USNM). PERU: 3♀, Madre de Dios, Avispas, 10/30-IX-1962, L.E. Peña (AMNH); 1♀, L.E. Peña (MLP); 1♂, 2♀, Sinchono, VII-[19]28, J.G. Sanders (OSUC); 2♀, Cueva de los Pavos, 15 km S Tingo María, 15-VII-1968, C.W. &amp; L. O’Brien (AMNH); 1♀, Bella Durmiente near Tingo María, 12- VII-1968, C.W. &amp; L. O’Brien (AMNH); 1♂, 24 km W Aguaytia between Pucallpa y Tingo María, C.W. &amp; L. O’Brien (AMNH); 1♀, Satipo, 10-VIII-1941, P. Paprzycki (USNM); 1♀, 17-VIII-1941, P. Paprzycki (USNM).</p></div>	https://treatment.plazi.org/id/6F64DB1B5E67FF86FC5FFA8CFC60F912	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pablo M. Dellapé;María C. Melo;Thomas J. Henry	Pablo M. Dellapé, María C. Melo, Thomas J. Henry (2016): A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species. Zoological Journal of the Linnean Society 177 (177): 29-134, DOI: 10.1111/zoj.12362
6F64DB1B5E69FF85FC0BF91FFB7EFD53.text	6F64DB1B5E69FF85FC0BF91FFB7EFD53.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heraeus panamaensis	<div><p>HERAEUS PANAMAENSIS SP. NOV.</p><p>(FIGS 15 F, 16F, 18, 20E–H)</p><p>Diagnosis</p><p>Strongly setose. Ocelli placed at level of an imaginary line passing posterior border of eyes. Labium extending to mesocoxae. Distiflagellomere with a pale band sub-basally. Profemur irregularly pigmented with brown; meso- and metatibia with a sub-basal brown band. Vesica with two sclerotized lobes bearing a few spines anteriorly near apices.</p><p>Description (Fig. 15 F)</p><p>Total length 5.65. Strongly setose.</p><p>Head: Brown, clypeus paler, shiny, coriaceous, with abundant, short, recumbent, and long erect setae. Head length 1.10, width 0.90. Postocular length 0.43. Eyes not surpassing the dorsal margin of head in lateral view, with long erect setae between ommatidea. Ocelli at level of an imaginary line passing posterior border of eyes. Interocular width 0.46, interocellar width 0.27. Labium pale brown, with short erect setae, extending to mesocoxae. Labial segment lengths: I 0.64, II 0.72, III 0.54, and IV 0.38. Antennae pale brown, except basiflagellomere distally and distiflagellomere darker, distiflagellomere with a pale band sub-basally; all segments with short, recumbent, and erect setae, pedicel with abundant erect setae, equal to or longer than diameter of segment; basiflagellomere weakly clavate. Antennal lengths: scape 0.34, pedicel 1.10, basiflagellomere 0.96, distiflagellomere 1.09. Length of pale band on distiflagellomere 0.32.</p><p>Thorax: Pronotum with anterior lobe brown, collar and posterior lobe paler; posterior lobe with irregular pale areas, humeral angle with a whitish spot surrounded by a darker area; pruinose; with short recumbent and long erect setae, longest on anterior lobe. Collar delimited posteriorly by a punctate sulcus. Pronotum punctate, punctures larger on posterior lobe. Collar length 0.06, anterior lobe length 0.53, posteri- or lobe length 0.51; anterior lobe width 0.85, posteri- or lobe width 1.34. Pleurae brown, punctate, pruinose, with semi-erect setae. Evaporative area extended. Scutellum brown, pruinose, punctate, with short recumbent and long erect setae, as on anterior lobe. Hemelytra pruinose, with abundant, short, semi-erect setae; pale brown, costal margin pale on proximal three-quarters, corium darker distally, with a subapical whitish spot and a rounded pale spot internally. Membrane dark brown, with an apical whitish spot, veins paler (Fig. 15 F). Lateral margins slightly concave. Legs: Coxae brown, remainder of legs whitish, except profemur irregularly pigmented with brown; femora and tibiae yellowish brown, with subapical band on meso- and metafemur, and sub-basal band on meso- and metatibia darker, tibiae and tarsi apically and pretarsi darker brown (Fig. 16 F); setae long and erect, longest on profemur. Profemur with spines restricted to apical half; protibia straight, with minute tubercles. Protibiae and mesofemur without spines.</p><p>Abdomen: Brown, with abundant short recumbent setae. Male genitalia: Pygophore (Fig. 20 E, F) rounded, elongate, anterior margin of dorsal aperture rounded, inner projections elongate, more produced posteriorly; declivent posteriorly in lateral view. Parameres: Figure 20 (G, H). Aedeagus: vesica with two sclerotized lobes and a few spines anteriorly near apices; processus gonopori long and slender.</p><p>Distribution</p><p>Panama (Fig. 18).</p><p>Etymology</p><p>The specific epithet ‘ panamaensis ’ is an adjective referring to the country where the holotype was collected.</p><p>Type material</p><p>Holotype: ♂, PANAMA: Cerro Campana, Dist. Chame, 2-V-1976, D. Engleman (AMNH).</p></div>	https://treatment.plazi.org/id/6F64DB1B5E69FF85FC0BF91FFB7EFD53	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pablo M. Dellapé;María C. Melo;Thomas J. Henry	Pablo M. Dellapé, María C. Melo, Thomas J. Henry (2016): A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species. Zoological Journal of the Linnean Society 177 (177): 29-134, DOI: 10.1111/zoj.12362
6F64DB1B5E6AFF83FC7AFCCBFA57FA9E.text	6F64DB1B5E6AFF83FC7AFCCBFA57FA9E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heraeus similis	<div><p>HERAEUS SIMILIS SP. NOV.</p><p>(FIGS 15 G, 20I–L, 21A, 22)</p><p>Diagnosis</p><p>Ocelli located posteriorly. Distiflagellomere with a subbasal pale band. Labium extending to metacoxae. Profemur irregularly pigmented; metafemur with a subapical dark band. Posterior margin of pygophore slightly bilobed.</p><p>Heraeus similis sp. nov. shares with H. chamamecinus sp. nov. and H. illitus a highly setose body and the presence of a weakly bilobed posterior margin of pygophore, but it can be distinguished by the length of the labium extending to the metacoxae (whereas it attains abdominal segment II in H. illitus and extends to the mesocoxae in H. chamamecinus sp. nov.).</p><p>Description (Fig. 15 G)</p><p>Total length 6.56. Strongly setose.</p><p>Head: Convex dorsally, dark brown, shiny, with short recumbent and long erect setae. Head length 1.44, width 0.96. Postocular length 0.54. Eyes protruding, not surpassing the dorsal margin of head in lateral view. Ocelli placed posterior to an imaginary line passing the posterior border of eyes. Interocular width 0.48, interocellar width 0.29. Labium pale brown with long erect setae, extending to metacoxae. Labial segment lengths: I 1.01, II 1.12, III 1.12, and IV 0.40. Antennal tubercles parallel. Antennae: Scape brown, pedicel and basiflagellomere light brown, darkened apically, distiflagellomere brown with a sub-basal pale brown band; all segments with short semi-erect and erect setae, pedicel with abundant erect setae, equal to or longer than diameter of segment. Antennal lengths: scape 0.53, pedicel 1.36, basiflagellomere 1.15, distiflagellomere 1.28. Length of pale band on distiflagellomere 0.38.</p><p>Thorax: Pronotum pruinose, anterior lobe dark brown, posterior lobe brown with pale areas, each humeral angle with a small pale spot. Anterior and posterior lobes with short recumbent and long erect setae, posterior lobe with shorter setae. Pronotum punctate, punctures more conspicuous on posterior lobe. Collar length 0.10, anterior lobe length 0.69, posterior lobe length 0.54; anterior lobe width 1.01, posterior lobe width 1.52. Pleurae dark brown. Evaporative area extended. Scutellum dark brown, pruinose, punctate, with setae similar to pronotal setae. Hemelytra pruinose, with erect setae, costal margin pale on proximal three-quarters, with a subapical corial spot, and an inner corial spot, colour pattern as in Figure 15 (G). Membrane dark, veins paler, with a pale apical spot. Legs: Coxae and protrochanter reddish brown, remainder of legs pale brown, profemur spotted with brown and with a subapical brown band; meso- and metafemur with a sub-basal brown band, meso- and metatibia with a pale-brown band subbasally, apex of tibiae, distal half of tarsus I, and pretarsus darker (Fig. 21 A). Setae long and erect, longer on profemur.</p><p>Abdomen: Brown with abundant, short, recumbent setae. Male genitalia: Pygophore (Fig. 20 I, J) subquadrangular, with posterior margin weakly bilobed, anterior margin of dorsal aperture rounded. Parameres: Figure 20 (K, L). Aedeagus: conjunctiva spined; lobes strongly sclerotized and with many spines; processus gonopori long and slender.</p><p>Distribution</p><p>Argentina, Bolivia, Brazil, French Guiana, and Peru (Fig. 22).</p><p>Etymology</p><p>The specific epithet ‘ similis ’, Latin for like, refers to the strongly setose body that is similar to and shared with H. chamamecinus sp. nov. and H. illitus .</p><p>Type material</p><p>Holotype: ♂, BRAZIL: Rondonia, 62 km SW Ariquemes nr. F[a]zenda, Rancho Grande, 20-X-1994, BLT, U. Schmitz (USNM).</p><p>Paratypes: 16♂, 22♀, same data as for holotype (USNM); 3♂, 2♀, same data as for holotype (MLP).</p><p>Additional material studied</p><p>ARGENTINA: Formosa: 2♂, Ea. La Marcela, 35 km</p><p>E El Colorado, VIII-2003, light trap, J. Williams col.</p><p>(MLP); Tucumán: one without abdomen, Tafi del Valle,</p><p>Quebrada de la Angostura, 1800 m a.s.l., 26-II-</p><p>[19]53, T. Ramirez (USNM).</p><p>BOLIVIA: Beni: 1♀, Romansos, 1 km. N junction Rio Itenez and Rio Paragua, 30-VII-1964 (AMNH); Santa Cruz: 1♀, Prov. San Esteban Muyurina, 49 km N Santa Cruz, 1120 ft, 26-X-[19]59, blacklight trap, R.B. Cumming (USNM); 1♀, Saavedra Res. Sta., 27- III-1978, UV trap, C.R. Ward &amp; C.W. O’Brien (USNM).</p><p>BRAZIL: Bahia: 3♂, Encruzilhada, 960 m a.s.l., XI- 1972, M. Alvarenga (AMNH); Espirito Santo: 1♀, Linhares, XI-1967, M. Alvarenga (AMNH); Estado do Rio: 1♀, Guanabara, Represa Rio Grande, III- IV-1972, F.M. Oliveira (AMNH); Mato Grosso: 1♀, Barra do Tapirape, 14-XII-1964, B. Malkin (MZSP); 1♂, Campo Grande, 1941, A. Maller, F. Johnson (AMNH); Minas Gerais: 2♀, Pedra Azul, 900 m a.s.l., XI-1972, M. Alvarenga (AMNH); Para: 1♀, Jacareacanga, XII-1968, M. Alvarenga (AMNH); Sao Paulo: 1♂, Ilha dos Buzios, 16-X/4-XI-[1]963, Exp. Dep. Zool. (MZSP); 3♂, 2♀, Piracicaba, 12-X-1965, C.A. Triplehorn, black light (AMNH); Rondonia: 3♂, 6♀, 62 km SW Ariquemes nr Fzenda, Rancho Grande, 25-X-1994, BLT, U. Schmitz (USNM); 2♀, 28-X- 1994, BLT, U. Schmitz (USNM); 2♀, 16-18-X-1996, BLT, U. Schmitz (USNM); 1♂, 1♀, 7-X-1994, BLT, U. Schmitz (USNM); 1♀, 12/26-IV-1992, BLT, U. Schmitz (USNM); 5♂, 9♀, 8/20-XI-1994, J.E. Eger (USNM); 2♂, 6♀, 4–16-XI-1997, J.E. Eger (USNM); 2♂, 5♀, 5/17-X-1993, J.E. Eger (USNM); 1♂, 1♀, 10-XI-1994, C. O’Brien (USNM); 1♂, 14-XI-1994, C. O’Brien (USNM); 1♀, 9-XI-1994, UV and mercury vapor light, C.W. &amp; L.B. O’Brien (USNM); 1♂, 1♀, 13-XI-1994, UV and merc. vap light, C.W. &amp; L.B. O’Brien (USNM); 1♀, 15-XI-1994, UV and merc. vap light, C.W. &amp; L.B. O’Brien (USNM); 1♀, 16-XI- 1994, UV and merc. vap light, C.W. &amp; L.B. O’Brien (USNM); 1♀, 18-XI-1994, UV and mercury vapor light, C.W. &amp; L.B. O’Brien (USNM).</p><p>FRENCH GUIANA: 1♂, 2♀, Grand Santi, 23-IX- 2000, A. Matocq (MNHN).</p><p>PERU: Cuzco: 1♀, Pilcopata, 600 m a.s.l., 8/10-VII- 1979, premontane moist forest, J.B. Heppner (USNM).</p></div>	https://treatment.plazi.org/id/6F64DB1B5E6AFF83FC7AFCCBFA57FA9E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pablo M. Dellapé;María C. Melo;Thomas J. Henry	Pablo M. Dellapé, María C. Melo, Thomas J. Henry (2016): A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species. Zoological Journal of the Linnean Society 177 (177): 29-134, DOI: 10.1111/zoj.12362
6F64DB1B5E6CFF82FC09FA8CFC60FE24.text	6F64DB1B5E6CFF82FC09FA8CFC60FE24.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heraeus spinosus	<div><p>HERAEUS SPINOSUS SP. NOV.</p><p>(FIGS 15 H, 20M–P, 21B, 24)</p><p>Diagnosis</p><p>Distiflagellomere with a wide sub-basal whitish band. Labium extending to mesocoxae. Profemur spotted, darker subapically, forming a band. Pygophore rounded, apex with a dorsally directed protuberance.</p><p>Description (Fig. 15 H)</p><p>Total length 6.27.</p><p>Head: Dark brown, shiny; with abundant short recumbent and long erect setae. Head length 1.34, width 0.96. Postocular length 0.48. Eyes not surpassing the dorsal margin of head in lateral view. Ocelli at level of an imaginary line passing the posterior border of eyes. Interocular width 0.48, interocellar width 0.24. Labium pale brown with long erect setae, almost extending to mesocoxae. Labial segment lengths: I 0.67, II 0.77, III 0.58, and IV 0.35. Antennae pale brown, except distal half of basiflagellomere and distiflagellomere darker, distiflagellomere with a wide sub-basal whitish band; all segments with abundant, short, semi-erect, and sparse erect setae. Antennal lengths: scape 0.55, pedicel 1.15, basiflagellomere 0.98, and distiflagellomere 1.13. Length of pale band on distiflagellomere 0.48.</p><p>Thorax: Anterior pronotal lobe dark brown, posteri- or lobe paler with irregular pale areas (Fig. 15 H), each humeral angle with a small whitish spot; collar delimited posteriorly by a punctate sulcus; anterior and posterior lobes with short recumbent and long erect setae, longest on anterior lobe. Pronotum pruinose, punctate, punctures larger on posterior lobe. Collar length 0.10, anterior lobe length 0.60, posterior lobe length 0.55; anterior lobe width 0.94, posterior lobe width 1.51. Pleurae brown, acetabular areas paler, punctate, with short recumbent setae. Evaporative area extended. Scutellum basally and a longitudinal median stripe dark brown, remainder paler brown, pruinose, punctate, with short recumbent and long erect setae, as on anterior pronotal lobe. Hemelytra brown, costal margin pale on proximal three-quarters, apical half of corium darker, with a subapical whitish spot and a small pale rounded spot internally (Fig. 15 H); setae short, semi-erect; membrane brown with a diffuse subapical pale spot, veins paler. Legs: Pale brown; profemur darker, mottled laterally and darker subapically forming a band, apex paler; coxae, protrochanter, a subapical band on meso- and metafemur, apex of tibiae, apex of tarsi, and pretarsi brown (Fig. 21 B); setae abundant and erect, longest on profemur. Profemur with spines restricted to apical two-thirds.</p><p>Abdomen: Brown, with abundant, short, recumbent setae. Male genitalia: Pygophore (Fig. 20 M, N) rounded, apex with a dorsally directed protuberance; slightly declivent posteriorly in lateral view. Anterior margin of dorsal aperture slightly rounded, inner projections quadrangular. Parameres: Figure 20 (O, P). Aedeagus: conjunctiva with spines laterally beneath the ejaculatory reservoir; vesica with a few spines laterally and two sclerotized lobes with a few spines distally; processus gonopori long and slender.</p><p>Distribution</p><p>Brazil and Ecuador (Fig. 24).</p><p>Etymology</p><p>The specific epithet ‘ spinosus ’, Latin for spiny, refers to the spined aedeagus.</p><p>Type material</p><p>Holotype: ♂, BRAZIL: Rondonia, 62 km SW Ariquenes, near Fazenda Rancho Grande, 16/18-III-1996, BLT, U. Schmitz (USNM).</p><p>Paratypes: 1♂, same locality as for holotype, 5–17-X- 1993, J.E. Eger, BLT (USNM). BRAZIL: 1♂, Bahia: Encruzilhada, 960 m a.s.l., XI-1972, M. Alvarenga (AMNH). ECUADOR: Napo: 1♀, vic. Puerto Misahuali, 1°2 ′ 4.2 ″ S, 77°39 ′ 49.2 ″ W, 1650–1900 ft, 6/19- IX-1998, mercury vapor and ultraviolet light, J.E. Eger (USNM).</p></div>	https://treatment.plazi.org/id/6F64DB1B5E6CFF82FC09FA8CFC60FE24	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pablo M. Dellapé;María C. Melo;Thomas J. Henry	Pablo M. Dellapé, María C. Melo, Thomas J. Henry (2016): A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species. Zoological Journal of the Linnean Society 177 (177): 29-134, DOI: 10.1111/zoj.12362
6F64DB1B5E6DFF81FC07FE05FA57FB5E.text	6F64DB1B5E6DFF81FC07FE05FA57FB5E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heraeus splendens	<div><p>HERAEUS SPLENDENS SP. NOV.</p><p>(FIGS 15 I, 21C, 23A–D, 24)</p><p>Diagnosis</p><p>Basiflagellomere clavate. Labium extending to mesocoxae. Anterior pronotal lobe with long erect setae; posterior pronotal lobe and hemelytra with short setae. Posterior pronotal lobe with a pale inverted V-shaped mark. Outer margin of clavus pale. Profemur brown, paler dorsally.</p><p>Heraeus splendens sp. nov. and H. antennalis sp. nov. are the only two species in the genus with the basiflagellomere clavate; whereas H. antennalis sp. nov. is strongly setose, with the profemora mottled, and the posterior pronotal lobe and clavus are uniformly pale brown.</p><p>Description (Fig. 15 I)</p><p>Total length 6.65.</p><p>Head: Reddish brown, shiny, coriaceous, with short and erect setae. Head length 1.42, width 1.01. Postocular length 0.65. Eyes not surpassing the dorsal margin of head in lateral view. Ocelli placed at level of an imaginary line passing the posterior border of eyes. Interocular width 0.53, interocellar width 0.24. Labium pale brown with short erect setae, extending to mesocoxae. Labial segment lengths: I 0.88, II 0.96, III 0.69, IV 0.35. Antennae reddish brown, distiflagellomere with a pale reddish-brown band. Basiflagellomere clavate. Antennal lengths: scape 0.53, pedicel 1.39, basiflagellomere 1.23, and distiflagellomere 1.28. Length of pale band on distiflagellomere 0.38.</p><p>Thorax: Pronotum pruinose, punctate; dark brown, posterior pronotal lobe brown with a pale inverted V-shaped mark and a pale submarginal line along lateral margin. Anterior pronotal lobe with long erect setae, posterior pronotal lobe with shorter setae. Collar length 0.10, anterior lobe length 0.69, posterior lobe length 0.60; anterior lobe width 1.01, posterior lobe width 1.60. Pleurae brown. Evaporative area extended. Scutellum brown with erect setae. Hemelytra: costal margin pale on proximal three-quarters, with a subapical corial spot. Membrane dark brown, with veins paler and a slender, pale apical spot. Legs: coxae, protrochanter, and profemur reddish brown, profemur paler dorsally; meso- and metafemur with a subapical dark band; meso- and metatibia with a sub-basal dark band, tibiae darkened apically (Fig. 21 C). Legs with semi-erect and erect setae, longest on profemora.</p><p>Abdomen: Brown, with abundant, short, recumbent setae. Male genitalia: Pygophore (Fig. 23 A, B) rounded, anterior margin of dorsal aperture rounded. Parameres: Figure 23 (C, D). Aedeagus: vesical lobes sclerotized posteriorly, with spines distally; processus gonopori long and slender.</p><p>Distribution</p><p>Panama (Fig. 24).</p><p>Etymology</p><p>The specific epithet ‘ splendens ’, Latin for beauty, refers to the attractive dorsal markings and colouration.</p><p>Type material</p><p>Holotype: ♂, PANAMA: Cerro Campana, 800 m a.s.l., Dist. Chame, 2-V-1976, D. Engleman (AMNH).</p><p>Paratypes: PANAMA: Panama: 1♀, Cerro Campana, 850 m a.s.l., 8°40 ′ N, 79°56 ′ W, 24-XII-[19]73, Stockwell (USNM); Chiriqui: 1♀, Res. For. La Fortuna, cont. divide Tr., 20-VII-1995, C.W. &amp; L.B. O’Brien (USNM).</p></div>	https://treatment.plazi.org/id/6F64DB1B5E6DFF81FC07FE05FA57FB5E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pablo M. Dellapé;María C. Melo;Thomas J. Henry	Pablo M. Dellapé, María C. Melo, Thomas J. Henry (2016): A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species. Zoological Journal of the Linnean Society 177 (177): 29-134, DOI: 10.1111/zoj.12362
6F64DB1B5E6EFF9FFC03FAF3FC0DFEAF.text	6F64DB1B5E6EFF9FFC03FAF3FC0DFEAF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heraeus tiputini	<div><p>HERAEUS TIPUTINI SP. NOV.</p><p>(FIGS 8 I, 15J, 21D, 23E–H, 24)</p><p>Diagnosis</p><p>Ocelli posteriorly. Distiflagellomere with a pale band sub-basally. Labium extending to mesocoxae. Profemur brown, paler dorsally; metafemur pale brown, darker apically; metatibia yellowish. Aedeagus: vesica with two unsclerotized lobes; processus gonopori long, widening towards apex.</p><p>Heraeus tiputini sp. nov. is similar to H. inca sp. nov. in having: the ocelli placed posteriorly behind the eyes; the profemur paler dorsally; the metafemur darker apically (a few specimens of H. inca sp. nov. show a dark band with a narrowly pale apex on the metafemur instead of being entirely dark distally); and the processus gonopori long, widening towards apex. In H. tiputini sp. nov. the metatibia is yellowish; the anterior margin of dorsal aperture of pygophore is rounded; and the aedeagus has a vesica with two unsclerotized lobes. Heraeus inca sp. nov. has a thin band on the basal region of the metatibia; the anterior margin of dorsal aperture of pygophore is slightly rounded with the lateral regions angulate; and the vesica has two pairs of lobes, the posterior lobe large and strongly sclerotized posteriorly.</p><p>Description (Fig. 15 J)</p><p>Total length 6.27.</p><p>Head: Elongate, postocular region longer than preocular region; flattened dorsally; shiny, dark brown, clypeus paler; with short recumbent and long erect setae. Head length 1.46, width 0.93. Postocular length 0.60. Eyes with setae between ommatidea; not surpassing dorsal margin of head in lateral view. Ocelli posteri- or of an imaginary line passing the posterior border of eyes. Interocular width 0.48, interocellar width 0.30. Labium pale brown with erect setae, extending to mesocoxae. Labial segment lengths: I 0.75, II 0.85, III 0.75, IV 0.32. Antennae brown; scapus, apical region of pedicel, and basiflagellomere darker, distiflagellomere brown, with a pale band sub-basally; all segments with abundant, short, semi-erect setae and sparse erect setae, more abundant on distiflagellomere. Basiflagellomere slightly incrassate apically. Antennal lengths: scape 0.45, pedicel 1.22, basiflagellomere 1.09, distiflagellomere 1.12. Length of pale band on distiflagellomere 0.48.</p><p>Thorax: Pronotum pruinose; anterior lobe dark brown, posterior lobe brown with two pale longitudinal stripes, with short recumbent and long erect setae, longest on anterior lobe. Collar delimited posteriorly by a row of punctures. Pronotum punctate, punctures more conspicuous on posterior lobe. Collar length 0.08, anterior lobe length 0.59, posterior lobe length 0.54; anterior lobe width 0.93, posterior lobe width 1.41. Pleurae dark brown, punctate, with short recumbent setae. Evaporative area extended. Scutellum dark brown, pruinose, punctate, with short recumbent and long erect setae. Hemelytra pruinose, with short semi-erect setae. General colour brown, costal margin pale on proximal threequarters, corium darker distally, with a subapical whitish spot and a small rounded pale spot internally; membrane brown, with veins paler and an apical pale spot (Fig. 15 J). Legs: Coxae brown; protrochanter, profemur laterally, and meso- and metafemur apically brown, remainder of legs pale brown; apex of tibiae, tarsi, and pretarsi darker (Fig. 21 D). Legs with long semi-erect setae, longest on profemur. Profemur with two rows of spines on apical two-thirds.</p><p>Abdomen: Brown, with abundant short recumbent setae. Male genitalia: Pygophore (Fig. 23 E, F) rounded, anterior margin of dorsal aperture rounded, inner projections subquadrangular; slightly declivent posteriorly in lateral view. Parameres: Figure 23 (G, H); shank with inner and outer projections indistinguishable. Aedeagus (Fig. 8 I): unsclerotized lobes of vesica with a group of strong spines near apex; processus gonopori long, widening towards apex.</p><p>Distribution</p><p>Ecuador and Peru (Fig. 24).</p><p>Etymology</p><p>The specific epithet ‘ tiputini ’ is a noun in aposition referring to ‘ Tiputini Biodiversity Station’, in Orellana Province, Ecuador, where most of the material was collected.</p><p>Type material</p><p>Holotype: ♂, ECUADOR: Orellana Prov. [Napo Prov. on label in error], Tiputini Biodiversity Station, 216 m a.s.l., 0°37 ′ 55 ′ S, 76°8 ′ 39 ′ W, 5-II-1999, T.L. Erwin et al., insecticidal fogging of mostly bare green leaves, some with covering of lichenous or bryophytic plants, lot 2082, transect T-9 (USNM).</p><p>Paratypes: 3♂, 3♀, same data as for holotype (USNM); 1♂, same data as for holotype, 22-X-1998, lot 1978, transect T-8 (MLP); 1♀, same data as for holotype, 22-X-1998, lot 1973, transect T-8 (MLP); 1♀, same data as for holotype, 6-II-1999, lot 2082, transect T-9 (USNM); 1♂, same data as for holotype, 8-II-1999, lot 2026, transect T-3 (USNM); 1♂, 1♀, same data as for holotype, 22-X-1998, lot 1972, transect T-8 (USNM); 1♀, same data as for holotype, lot 1962, transect T-7 (USNM); 1♂, same data as for holotype, 26-X-1998, lot 1941, transect T-5 (USNM); 3♂, same data as for holotype, 26-X-1998, lot 1942, transect T-5 (USNM); 1♂, 1♀, Reserva Ethnica Waorani, 1 km S Onkone Gare Camp., Trans. Ent., 4-X-1996, 220 m a.s.l., 0°39 ′ 10 ″ S, 76°26 ′ 0 ″ W, T.L. Erwin et al., insecticidal fogging of mostly bare green leaves, some with covering of lichenous or bryophytic plants in terra firme forest, lot 1744, transect T-9 (USNM); 2♂, 2♀, lot 1743, transect T-9 (USNM); 1♀, 25-VI- 1994, lot 731 (USNM); 1♀, 29-VI-1994, at 6 x-trans, 81 m mark Proj. Maxus, lot 759 (USNM); 1♀, Reserva Ethnica Waorani, Onkone Gare Camp., 38 km S of Pompeya, 220 m a.s.l., 0°39 ′ 10 ″ S, 76°26 ′ W, 2–14-II- 1996, T.J. Henry (USNM). PERU: Ucayali: 1♀, Kirigueti, 11°38 ′ 13 ″ S, 73°7 ′ 8 ″ W, luz, VII-[20]04, J. Williams (MLP).</p></div>	https://treatment.plazi.org/id/6F64DB1B5E6EFF9FFC03FAF3FC0DFEAF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pablo M. Dellapé;María C. Melo;Thomas J. Henry	Pablo M. Dellapé, María C. Melo, Thomas J. Henry (2016): A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species. Zoological Journal of the Linnean Society 177 (177): 29-134, DOI: 10.1111/zoj.12362
6F64DB1B5E70FF9FFC67FE82FBC2FBDA.text	6F64DB1B5E70FF9FFC67FE82FBC2FBDA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heraeus plebejus	<div><p>THE PLEBEJUS GROUP</p><p>Included species: Heraeus alvarengai sp. nov., H. apicalis sp. nov., H. bahiensis sp. nov., H. baranowskii sp. nov., H. bolivianus sp. nov., H. brevirostris sp. nov., H. concolor, H. ecuatorianus sp. nov., H. loja sp. nov., H. mesoamericanus sp. nov., H. mexicanus sp. nov., H. morganae sp. nov., H. nicaraguensis sp. nov., H. pacificus, H. pallidinervis sp. nov., H. penai sp. nov., H. plebejus, and H. pulchellus .</p><p>Diagnosis</p><p>Head rugose, setose; collar with a well-developed posterior furrow; evaporative area extended, distance from dorsal margin of auricle to dorsal margin of evaporative area subequal to or longer than distance from dorsal margin of evaporative area to dorsal margin of metapleura (except H. alvarengai sp. nov. and H. mesoamericanus sp. nov.); hemelytra with a heterogeneous colouration pattern; profemur uniformly dark brown, although in some species the apex is paler; male mesofemora without spines; and aedeagus spined.</p><p>Although this group is paraphyletic in our analysis, we are keeping it as a group for identification purposes because the relationships among the species are not well supported and additional characters are needed for better resolution.</p></div>	https://treatment.plazi.org/id/6F64DB1B5E70FF9FFC67FE82FBC2FBDA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pablo M. Dellapé;María C. Melo;Thomas J. Henry	Pablo M. Dellapé, María C. Melo, Thomas J. Henry (2016): A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species. Zoological Journal of the Linnean Society 177 (177): 29-134, DOI: 10.1111/zoj.12362
6F64DB1B5E70FF9BFC1EFB76FF23FAF0.text	6F64DB1B5E70FF9BFC1EFB76FF23FAF0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heraeus alvarengai	<div><p>HERAEUS ALVARENGAI SP. NOV.</p><p>(FIGS 25 A, 26A, 27A–D, 28)</p><p>Diagnosis</p><p>Ocelli placed anterior to an imaginary line passing the posterior border of eyes. Labium extending to metacoxae. Evaporative area short, distance from dorsal margin of auricle to dorsal margin of evaporative area shorter than distance from dorsal margin of evaporative area to dorsal margin of metapleura. Membrane with a diffuse, apical whitish spot. Meso- and metafemur with a subapical brown band. Apex of pygophore pointed dorsally in lateral view.</p><p>This is the only species with the ocelli placed anterior to an imaginary line passing the posterior border of eyes.</p><p>Description (Fig. 25 A)</p><p>Total length 4.75.</p><p>Head: Brown, shiny, with short recumbent and scattered erect setae. Head length 1.06, width 0.82. Postocular length 0.37. Eyes not surpassing dorsal margin of head in lateral view. Ocelli placed anterior to an imaginary line passing the posterior border of eyes. Interocular width 0.42, interocellar width 0.26. Labium pale brown, with short erect setae; extending to metacoxae. Labial segment lengths: I 0.72, II 0.83, III 0.72, and IV 0.34. Antennae (right antenna with three segments, last segment darker with a diffuse pale band) pale brown, apex of basiflagellomere darker; distiflagellomere absent; with abundant, short, recumbent setae. Antennal lengths: scape 0.42, pedicel 0.93, and basiflagellomere 0.78.</p><p>Thorax: Pronotum brown, posterior lobe paler, with two irregular pale spots, each humeral angle with a pale spot; anterior and posterior lobes with short recumbent setae, with long scattered setae on anterior lobe. Pronotum pruinose, punctate, punctures larger on posterior lobe. Collar length 0.08, anterior lobe length 0.61, posterior lobe length 0.37; anterior lobe width 0.80, posterior lobe width 1.14. Pleurae brown, punctate, with short recumbent setae. Evaporative area short. Scutellum brown, with a longitudinal, dark, median stripe, pruinose, punctate, with short recumbent and sparse long erect setae. Hemelytra with short recumbent setae. Costal margin pale on proximal threequarters, corium darker distally, with a subapical whitish spot and a rounded pale spot internally (Fig. 25 A); membrane dark brown, veins paler, with a diffuse apical whitish spot. Legs: Coxae, protrochanter, and profemur brown, remainder of legs whitish, except for a darker subapical band on meso- and metafemur, tibiae darker apically (Fig. 26 A); setae short and semierect. Profemur with spines on apical two-thirds. Protibia straight.</p><p>Abdomen: Brown, with abundant short recumbent setae. Male genitalia: Pygophore (Fig. 27 A, B) rounded, anterior margin of dorsal aperture slightly rounded, apex with a dorsally directed protuberance; declivent posteriorly in lateral view. Inner projections elongate, more produced posteriorly. Parameres: Figure 27 (C, D). Aedeagus: vesica with a few spines laterally and with two sclerotized lobes bearing spines anteriorly; processus gonopori long and slender.</p><p>Distribution</p><p>Brazil (Fig. 28).</p><p>Etymology</p><p>This species is named after the late Brazilian entomologist Moacyr Alvarenga, who collected the holotype.</p><p>Type material</p><p>Holotype: ♂, BRAZIL, [Río Grande do Sul], Guanabara, Represa Rio Grande, VI-1972, M. Alvarenga (AMNH).</p></div>	https://treatment.plazi.org/id/6F64DB1B5E70FF9BFC1EFB76FF23FAF0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pablo M. Dellapé;María C. Melo;Thomas J. Henry	Pablo M. Dellapé, María C. Melo, Thomas J. Henry (2016): A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species. Zoological Journal of the Linnean Society 177 (177): 29-134, DOI: 10.1111/zoj.12362
6F64DB1B5E74FF99FE9AFAAEFD55FEE2.text	6F64DB1B5E74FF99FE9AFAAEFD55FEE2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heraeus apicalis	<div><p>HERAEUS APICALIS SP. NOV.</p><p>(FIGS 25 B, 26, 27E–H, 29)</p><p>Diagnosis</p><p>Labium extending to mesocoxae. Distiflagellomere with a sub-basal yellowish band. Membrane with a subapical pale spot. Metafemur darker on apical one-quarter. Processus gonopori widening towards apex.</p><p>Description (Fig. 25 B)</p><p>Total length 5.51.</p><p>Head: Dark reddish brown, clypeus paler, shiny; with abundant short recumbent and long erect setae dorsally. Head length 1.22, width 0.82. Postocular length 0.38. Eyes not surpassing the dorsal margin of head in lateral view. Ocelli placed at level of an imaginary line passing the posterior border of eyes. Interocular width 0.38, interocellar width 0.17. Labium pale brown, with erect setae; extending to mesocoxae. Labial segment lengths: I 0.72, II 0.91, III 0.72, and IV 0.24. Antennae: Scapus brown, pedicel and base of basiflagellomere pale brown, basiflagellomere distally reddish brown, distiflagellomere brown with a sub-basal yellowish band; all segments with abundant short recumbent setae and short semi-erect setae; basiflagellomere weakly clavate. Antennal lengths: scape 0.43, pedicel 1.03, basiflagellomere 0.96, and distiflagellomere 1.10. Length of pale band on distiflagellomere 0.34.</p><p>Thorax: Pronotum: pruinose, punctate, punctures larger on posterior lobe; anterior pronotal lobe reddish brown; collar paler, delimited posteriorly by a punctate sulcus, posterior lobe brown with irregular paler areas, each humeral angle with a pale spot; anterior and posterior lobes with short recumbent and erect setae, and long erect setae on anterior lobe. Collar length 0.08, anterior lobe length 0.51, posterior lobe length 0.48; anterior lobe width 0.83, posterior lobe width 1.30. Pleurae reddish brown, punctate, with short recumbent setae. Evaporative area extended. Scutellum reddish brown, pruinose, punctate, with short recumbent and long erect setae, as on anterior pronotal lobe. Hemelytra brown, with short recumbent setae, costal margin pale on proximal three-quarters, corium darker distally, with a subapical whitish spot and a pale spot internally (Fig. 25 B); membrane brown, veins paler, with a small subapical pale spot. Legs: Coxae, protrochanter, profemur, and meso- and metafemur distally brown, remainder of legs pale brown, tibiae darker apically (Fig. 26 B); with short recumbent and semi-erect setae, longer on profemur. Profemur with spines restricted to apical two-thirds. Protibia with minute denticles. Protibiae and mesofemur without spines.</p><p>Abdomen: Brown, with abundant short recumbent setae. Male genitalia: Pygophore (Fig. 27 E, F) rounded, elongate, with a straight posterior margin; slightly declivent posteriorly in lateral view; anterior margin of dorsal aperture rounded, inner projections subquadrangular. Parameres (Fig. 27 G, H) with outer projection not differentiated. Aedeagus: sclerotized lobes of vesica with a few spines; processus gonopori widening towards apex.</p><p>Distribution</p><p>Brazil, Ecuador, and Panama (Fig. 29).</p><p>Etymology</p><p>The specific ephitet ‘ apicalis ’, from the Latin apex, is an adjective, and refers to the distinct apical band on the metafemur.</p><p>Type material</p><p>Holotype: ♂, PANAMA: Distrito Chame, Cerro Campana, 800 m a.s.l., 22-II-[19]75, H.D. Engleman (USNM).</p><p>Paratypes: 1♂, ECUADOR, Tung.[urahua]: ∼ 29 km N Puyo, 22-I-1974, 3800 ft, on Austroeupatorium inulaefolium, R.M. King (USNM). BRAZIL: Rondonia: 1♀, 62 km SW Ariquemes, nr Fza. Rancho Grande, 4/16-XI-1997, blacklight trap, J.E. Eger (USNM); 1♂, 5–17-X-1993, blacklight trap, J.E. Eger (MLP); 1♂, 18-IX-199?, BL trap, U. Schmitz (USNM).</p></div>	https://treatment.plazi.org/id/6F64DB1B5E74FF99FE9AFAAEFD55FEE2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pablo M. Dellapé;María C. Melo;Thomas J. Henry	Pablo M. Dellapé, María C. Melo, Thomas J. Henry (2016): A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species. Zoological Journal of the Linnean Society 177 (177): 29-134, DOI: 10.1111/zoj.12362
6F64DB1B5E76FF99FEACFE58FB86FB51.text	6F64DB1B5E76FF99FEACFE58FB86FB51.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heraeus bahiensis	<div><p>HERAEUS BAHIENSIS SP. NOV.</p><p>(FIGS 25 C, 26C, 27I–L, 29, 30A)</p><p>Diagnosis</p><p>Distiflagellomere with a sub-basal whitish band. Labium extending to mesocoxae. Membrane brown with an apical pale spot. Meso- and metafemur with a subapical brown band. Pygophore with the apex pointed dorsally in lateral view.</p><p>The diagnostic characters relate this new species to H. nicaraguensis sp. nov., but they can be distinguished by the paler general colouration of H. bahiensis sp. nov., with the head dark reddish brown and the posterior pronotal lobe brown with diffuse longitudinal stripes, whereas in H. nicaraguensis sp. nov. the head is blackish and the posterior pronotal lobe is dark brown with four longitudinal pale stripes.</p><p>Description (Fig. 25 C)</p><p>Total length 5.61.</p><p>Head: Brown, shiny, with short recumbent and long erect setae dorsally. Head length 1.12, width 0.88. Postocular length 0.37. Eyes not surpassing the dorsal margin of head in lateral view. Ocelli at level of an imaginary line passing the posterior border of eyes. Interocular width 0.42, interocellar width 0.24. Labium pale brown, with erect setae; extending to mesocoxae. Labial segment lengths: I 0.80, II 0.85, III 0.66, and IV 0.35. Antennae pale brown, scape externally, basiflagellomere distally, and distiflagellomere darker, distiflagellomere with a sub-basal whitish band occupying about one-third of segment length; all segments with abundant short recumbent and sparse short erect setae. Antennal lengths: scape 0.43, pedicel 1.04, basiflagellomere 0.91, and distiflagellomere 0.96. Length of pale band on distiflagellomere 0.37.</p><p>Thorax: Anterior pronotal lobe brown, posterior lobe paler with mottled pale areas, humeral angles yellowish; collar punctate, delimited posteriorly by a punctate sulcus; anterior and posterior lobes with short recumbent setae, with longer erect setae on anterior lobe. Pronotum pruinose, punctate, punctures larger on posterior lobe. Collar length 0.08, anterior lobe length 0.48, posterior lobe length 0.48; anterior lobe width 0.90, posterior lobe width 1.44. Pleurae brown, acetabular areas paler, punctate, with short recumbent setae. Evaporative area extended. Scutellum brown, with a longitudinal median dark stripe, apex paler, pruinose, punctate, with short recumbent and long erect setae, as on anterior pronotal lobe. Hemelytra pale brown, irregularly pigmented, with short recumbent setae. Costal margin pale on proximal three-quarters, apex of corium darker with a subapical whitish spot and an inner small pale spot; membrane brown, veins paler with an apical pale spot (Fig. 25 C). Legs: Coxae, protrochanter, profemur, except distally, and a subapical band on meso- and metafemur brown; remainder of legs pale brown, apex of tibiae, tarsi, and pretarsi darker (Fig. 26 C); with semi-erect setae longer on profemur. Profemur with spines restricted to apical two-thirds.</p><p>Abdomen: Brown, with abundant short recumbent setae. Male genitalia: Pygophore (Fig. 27 I, J) rounded, anterior margin of dorsal aperture slightly rounded, apex with a small protuberance; declivent posteriorly in lateral view. Inner projections elongate, more pronounced posteriorly. Parameres: Figure 27 (K, L). Aedeagus (Fig. 30 A): vesica with a row of spines laterally and with two sclerotizated spined lobes; processus gonopori long and slender.</p><p>Distribution</p><p>Brazil (Fig. 29).</p><p>Etymology</p><p>The specific epithet ‘ bahiensis ’ is an adjective that refers to the state of Bahia, Brazil, where all specimens were collected.</p><p>Type material</p><p>Holotype: ♂, BRAZIL, Bahia, Encruzilhada, 960 m a.s.l., XI-1972, M. Alvarenga (AMNH).</p><p>Paratypes: 1♂, 2♀, same data as for holotype (AMNH); 1♂, same data as holotype (MLP); 1♀, same data as for holotype (USNM).</p></div>	https://treatment.plazi.org/id/6F64DB1B5E76FF99FEACFE58FB86FB51	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pablo M. Dellapé;María C. Melo;Thomas J. Henry	Pablo M. Dellapé, María C. Melo, Thomas J. Henry (2016): A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species. Zoological Journal of the Linnean Society 177 (177): 29-134, DOI: 10.1111/zoj.12362
6F64DB1B5E76FF96FC27FACCFE2CFA01.text	6F64DB1B5E76FF96FC27FACCFE2CFA01.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heraeus baranowskii	<div><p>HERAEUS BARANOWSKII SP. NOV.</p><p>(FIGS 25 D, 26D, 27M–P, 31)</p><p>Diagnosis</p><p>Labium extending to metacoxae. Distiflagellomere with a sub-basal whitish band. Membrane with an elongate apical pale spot. Mesofemur with a diffuse band and metafemur with a narrow subapical brown band. Pygophore apex pointed dorsally in lateral view.</p><p>Description (Fig. 25 D)</p><p>Total length 4.66.</p><p>Head: Brown, shiny, coriaceus; with short recumbent and long erect setae dorsally. Head length 1.01, width 0.77. Postocular length 0.34. Eyes not surpassing the dorsal margin of head in lateral view. Ocelli at level of an imaginary line passing the posterior border of eyes. Interocular width 0.35, interocellar width 0.24. Labium pale brown, with erect setae, extending to metacoxae. Labial segment lengths: I 0.59, II 72.</p><p>Antennae pale brown, basiflagellomere distally and distiflagellomere darker, distiflagellomere with a subbasal whitish band; all segments with abundant short recumbent and sparse short erect setae. Antennal lengths: scape 0.34, pedicel 0.84, basiflagellomere 0.72, and distiflagellomere 0.90. Length of pale band on distiflagellomere 0.30.</p><p>Thorax: Pronotum brown, anterior pronotal lobe darker, posterior lobe with irregular pale areas, humeral angles with a whitish spot; collar with scattered punctures delimited posteriorly by a punctate sulcus; anterior and posterior lobes with short recumbent and long erect setae, longer on anterior lobe. Pronotum pruinose, punctate, punctures larger on posterior lobe. Collar length 0.06, anterior lobe length 0.54, posterior lobe length 0.51; anterior lobe width 0.74, posterior lobe width 1.50. Pleurae brown, acetabular areas paler, punctate, with short recumbent setae. Evaporative area extended. Scutellum brown, apex paler; pruinose, punctate, with short recumbent and long erect setae, as on anterior pronotal lobe. Hemelytra pale brown, irregularly pigmented, with short recumbent setae. Costal margin pale on proximal twothirds, apex of corium darker with a subapical whitish spot and an inner small pale spot; membrane brown, veins paler, with an elongate apical pale spot and diffuse paler spots between veins (Fig. 25 D). Legs: Coxae, protrochanter, profemur, except distally, a diffuse subapical band on mesofemur, and a subapical narrow band on metafemur brown, remainder of legs pale brown (Fig. 26 D); with semierect setae, longest on profemur. Profemur with spines restricted to apical half. Protibia slightly curved.</p><p>Abdomen: Brown, with abundant short recumbent setae. Male genitalia: Pygophore (Fig. 27 M, N) rounded, with pointed posterior margin; apex with a dorsally directed protuberance; declivent posteriorly in lateral view; anterior margin of dorsal aperture weakly rounded, inner projections elongate, more pronounced posteriorly. Parameres: Figure 27 (O, P). Aedeagus: vesica with minute spines laterally and two sclerotizated lobes with minute spines distally; processus gonopori long and slender.</p><p>Distribution</p><p>Belize and Mexico (Fig. 31).</p><p>Etymology</p><p>This species is named in honour of our friend and colleague Richard M. Baranowski for his many contributions, and who collected most of the specimens of this new species.</p><p>Type material</p><p>Holotype: ♂, BELIZE, Cayo Dist., Benque Viejo, Riverside N, Mopan R. Resort, 22-VI-[20]06, BLT, R.M. &amp; H.V. Baranowski (USNM).</p><p>Paratypes: 2♂, same data as for holotype, 23-VI- [20]06 (USNM); 1♂, same data as for holotype, 24- VI-[20]06 (USNM); 1♂, same data as for holotype, 25-VI-[20]06 (USNM); 2♀, BLT, 1-X-2009, M. Chrysler (USNM); 1♀, 12-X-2009, M. Chrysler (USNM); 1♂, 7-XI-2009, M. Chrysler (USNM); 1♂, 23-XI-2009, M. Chrysler (MLP); 1♂, 25-X-[20]06, BLT, A. Trujillo (MLP); 1♂, Orange Walk District, Gallon Jug, 11- X-[20]05, BLT, B. Miller (USNM); 1♂, 19-VI- [20]05, BLT, B. Miller (USNM); 2♂, 31-IV-[20]05, BLT, B. Miller (USNM); one without abdomen, 14-I- [20]05, BLT, B. Miller (USNM); 1♂, Orange Walk Distr., Chan Chich Ldge., 25-X-[20]03, BLT, R.M. &amp; H.V. Baranowski (USNM); 1♂, 18-IV-[20]04, BL trap, E. Flota (USNM); 2♂, 1♀, 13-XI-[20]04, BL trap, C. Sanabria (USNM); 1♂, 1-IX-[20]05, BL trap, C. Sanabria (USNM); 1♂, O.W. Dist., Rio Bravo Cons. Area, Mahogany Tr, 10-VII-1996, C.W. &amp; L.B. O’Brien (USNM).</p><p>Additional material studied</p><p>MEXICO: Yucatan: 1♂, Kiuick, 13-XI-1974, J. Reddell (USNM); Quintana Roo: 1♀, Miguel Cozumel, VII- 1959, N.L.H. Kraus (USNM).</p></div>	https://treatment.plazi.org/id/6F64DB1B5E76FF96FC27FACCFE2CFA01	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pablo M. Dellapé;María C. Melo;Thomas J. Henry	Pablo M. Dellapé, María C. Melo, Thomas J. Henry (2016): A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species. Zoological Journal of the Linnean Society 177 (177): 29-134, DOI: 10.1111/zoj.12362
6F64DB1B5E79FF94FE98FA23FE06FEAF.text	6F64DB1B5E79FF94FE98FA23FE06FEAF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heraeus bolivianus	<div><p>HERAEUS BOLIVIANUS SP. NOV.</p><p>(FIGS 25 E, 26E, 29, 30B, 32A–D)</p><p>Diagnosis</p><p>Labium extending to mesocoxae. Distiflagellomere with a sub-basal whitish band. Membrane brown, with veins paler. Meso- and metafemur with a subapical brown band. Pygophore rounded, anterior margin of dorsal aperture rounded, inner projections elongate, more pronounced posteriorly.</p><p>Description (Fig. 25 E)</p><p>Total length 5.22.</p><p>Head: Brown, clypeus paler, shiny, coriaceous, with short recumbent and long erect setae dorsally. Head length 1.08, width 0.85. Postocular length 0.43. Eyes not surpassing the dorsal margin of head in lateral view. Ocelli at level of an imaginary line passing the posterior border of eyes. Interocular width 0.40, interocellar width 0.26. Labium pale brown, extending to mesocoxae, with erect setae. Labial segment lengths: I 0.77, II 0.77, III 0.62, and IV 0.29. Antennae pale brown, basiflagellomere distally and distiflagellomere darker, distiflagellomere with a sub-basal whitish band; all segments with abundant short recumbent and sparse short erect setae. Antennal lengths: scape 0.48, pedicel 1.01, basiflagellomere 0.84, and distiflagellomere 1.03. Length of pale band on distiflagellomere 0.22.</p><p>Thorax: Anterior pronotal lobe brown, collar and posterior lobe paler, with irregular paler areas, humeral angles with a small whitish spot; collar with punctures delimited posteriorly by a punctate sulcus; anterior and posterior lobes with short recumbent setae and long erect setae on anterior lobe. Pronotum pruinose, punctate, punctures larger on posterior lobe. Collar length 0.08, anterior lobe length 0.51, posteri- or lobe length 0.48; anterior lobe width 0.86, posteri- or lobe width 1.34. Pleurae brown, acetabular areas paler, punctate, with short recumbent setae. Evaporative area extended. Scutellum brown, apex paler, pruinose, punctate, with short recumbent and long erect setae, as on anterior pronotal lobe. Hemelytra pale brown, irregularly pigmented. Costal margin pale on proximal three-quarters, apex of corium darker with a subapical whitish spot and an inner small pale spot; with short recumbent setae; membrane brown with small paler spots between veins, veins paler (Fig. 25 E). Legs (left profemur, tibia, and tarsi missing): Coxae, protrochanter, profemur, except distally, and a subapical band on meso- and metafemur brown, remainder of legs pale brown, apex of tibiae, tarsi, and pretarsi darker (Fig. 26 E); semi-erect setae longer on profemur. Profemur with spines restricted to apical two-thirds.</p><p>Abdomen (glued on card): Brown, with abundant short recumbent setae. Male genitalia: Pygophore (Fig. 32 A, B) rounded, anterior margin of dorsal aperture rounded, inner projections elongate, more pronounced posteriorly; slightly declivent posteriorly in lateral view. Parameres: Figure 32 (C, D). Aedeagus (Fig. 30 B): vesica with a patch of spines laterally and with two sclerotizated lobes; processus gonopori long and slender.</p><p>Distribution</p><p>Bolivia (Fig. 29).</p><p>Etymology</p><p>The specific epithet ‘ bolivianus ’ is an adjective, and refers to Bolivia, the country where the only known specimen was collected.</p><p>Type material</p><p>Holotype: ♂, BOLIVIA, N La Paz, Mapiri, 10/16-VIII- [19]89, L.E. Peña (USNM).</p></div>	https://treatment.plazi.org/id/6F64DB1B5E79FF94FE98FA23FE06FEAF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pablo M. Dellapé;María C. Melo;Thomas J. Henry	Pablo M. Dellapé, María C. Melo, Thomas J. Henry (2016): A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species. Zoological Journal of the Linnean Society 177 (177): 29-134, DOI: 10.1111/zoj.12362
6F64DB1B5E7BFF94FEAEFE9FFBE6FA2F.text	6F64DB1B5E7BFF94FEAEFE9FFBE6FA2F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heraeus brevirostris	<div><p>HERAEUS BREVIROSTRIS SP. NOV.</p><p>(FIGS 25 F, 26F, 28, 30C, 32E–H)</p><p>Diagnosis</p><p>Labium extending to mesocoxae. Distiflagellomere with a narrow sub-basal pale band. Membrane with an apical pale spot. Meso- and metafemur with a narrow subapical brown band. Apex of pygophore with a small protuberance pointed dorsally.</p><p>Heraeus brevirostris sp. nov., H. pallidinervis sp. nov., H. penai sp. nov., and H. loja sp. nov. have a narrow pale band on the distiflagellomere, occupying about onequarter of the segment length; H. brevirostris sp. nov., however, can be distinguished by the shorter labium extending only to mesocoxae, whereas the other species have a longer labium, extending to metacoxae.</p><p>Description (Fig. 25 F)</p><p>Total length 5.61.</p><p>Head: Dark brown, shiny, coriaceous; with short recumbent setae and erect setae dorsally. Head length 1.15, width 0.85. Postocular length 0.42. Eyes not surpassing the dorsal margin of head in lateral view. Ocelli placed at level of an imaginary line passing the posterior border of eyes. Interocular width 0.45, interocellar width 0.29. Labium pale brown with short erect setae, extending to mesocoxae. Labial segment lengths: I 0.78, II 0.88, III 0.72, and IV 0.35. Antennae pale brown, apex of basiflagellomere and distiflagellomere darker, distiflagellomere with a narrow sub-basal pale band; all segments with abundant short recumbent and sparse erect setae. Antennal lengths: scape 0.45, pedicel 1.04, basiflagellomere 0.93, and distiflagellomere 1.02. Length of pale band on distiflagellomere 0.19.</p><p>Thorax: Pronotum dark brown, collar and posterior lobe paler, with irregular pale spots, humeral angles with a pale spot; pruinose, punctate, punctures larger on posterior lobe; anterior and posterior lobes with short recumbent and erect setae, longer on anterior lobe. Collar punctate, delimited by a punctate sulcus. Collar length 0.08, anterior lobe length 0.51, posterior lobe length 0.51; anterior lobe width 0.91, posterior lobe width 1.41. Pleurae dark reddish brown, punctate, with short recumbent setae. Evaporative area extended. Scutellum dark brown with a longitudinal darker median stripe, pruinose, punctate, with short recumbent and long erect setae, as on anterior pronotal lobe. Hemelytra: costal margin pale on proximal twothirds, with a subapical corial spot, membrane brown with veins paler and a pale apical spot, colouration pattern complex (Fig. 25 F); with short recumbent setae.</p><p>Legs: Coxae, protrochanter, and profemur dark brown, except apex of profemur and remainder of legs yellowish, with a subapical narrow brown band on meso- and metafemur (Fig. 26 F); setae short and semierect, longer on profemur. Profemur with spines on apical two-thirds. Protibia slightly curved.</p><p>Abdomen: Brown, with abundant short recumbent setae. Male genitalia: Pygophore (Fig. 32 E, F) rounded, anterior margin of dorsal aperture rounded, apex with a small protuberance pointed dorsally; declivent posteriorly in lateral view; inner projections elongate, more produced posteriorly. Parameres: Figure 32 (G, H). Aedeagus (Fig. 30 C) with strong spines, conjunctiva with a few spines laterally; vesica with two spined sclerotized lobes and small unsclerotized lobes anteriorly with a patch of spines; processus gonopori long and slender.</p><p>Distribution</p><p>Argentina and Bolivia (Fig. 28).</p><p>Etymology</p><p>The specific epithet ‘ brevirostris ’ is an adjective meaning short-beaked, and refers to the relatively short labium that extends only to the mesocoxae.</p><p>Type material</p><p>Holotype: ♂, ARGENTINA, Salta, Orán, Abra Grande, 10-I/1-X-[19]67, R. Golbach (IFML).</p><p>Paratypes: 2♂, 1♀, same data as for holotype (IFML); Salta: 1♂, 1♀, XII-2012, L. Alvarez col. (MLP); Tucumán: 1♂, (MLP). BOLIVIA: La Paz: 1♂, Yungas, Inquisivi, 5-XII-1984, L.E. Peña (USNM); 1♂, Guanay, Tres Esteros, 19/25-VIII-[19]89, L.E. Peña (USNM). Cochabamba: 2♂, Chapare Prov., Villa Tunari, Hotel de Selva El Puente, 340 m a.s.l., 16°59.04 ′ S 65°24.53 ′ W, 10-12-XII-2013, T.J. Henry (USNM); 1♀, Chapare Prov., Highway 4 at Rio Antahuacana bridge, 38 km SW of Villa Tunari, 850 m a.s.l., 17°05.35 ′ S, 65°40.62 ′ W, 12-XII-2013, T.J. Henry (USNM).</p></div>	https://treatment.plazi.org/id/6F64DB1B5E7BFF94FEAEFE9FFBE6FA2F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pablo M. Dellapé;María C. Melo;Thomas J. Henry	Pablo M. Dellapé, María C. Melo, Thomas J. Henry (2016): A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species. Zoological Journal of the Linnean Society 177 (177): 29-134, DOI: 10.1111/zoj.12362
6F64DB1B5E7BFF92FCF3FA1CFB59FEC3.text	6F64DB1B5E7BFF92FCF3FA1CFB59FEC3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heraeus concolor Slater & Baranowski 1994	<div><p>HERAEUS CONCOLOR SLATER &amp; BARANOWSKI, 1994</p><p>(FIGS 25 G, 30D, 31, 32I–L, 33A)</p><p>Heraeus concolor Slater &amp; Baranowski, 1994: 493 –494; Slater &amp; O’Donnell, 1995: 147; Baranowski &amp; Slater, 2005: 137.</p><p>Diagnosis</p><p>Small species, total length less than 5 mm. Pedicel with abundant erect setae, equal to or longer than diameter of segment. Distiflagellomere with a pale band. Labium extending to metacoxae. Membrane with an apical pale spot. Metafemur darker on apical one-quarter.</p><p>Heraeus concolor is similar in external appearance to H. pulchellus . Both are small species with a long labium extending to or near metacoxae; H. concolor can be differentiated by the dark-brown distiflagellomere with a conspicuous pale sub-basal band, whereas in H. pulchellus it is uniformly dark brown. These two species are known from the West Indies, and probably derived from ancestral stock related to H. plebejus that reached the islands from Florida (USA).</p><p>Description (Fig. 25 G)</p><p>Head: Coriaceous. Eyes not surpassing the dorsal margin of head in lateral view. Ocelli placed at level of an imaginary line passing the posterior border of eyes. Labium extending to metacoxae. Distiflagellomere with a pale band. Pedicel with abundant erect setae, equal to or longer than diameter of segment.</p><p>Thorax: Posterior pronotal lobe with pale spots and a pale spot on humeral angles. Anterior pronotal lobe with long erect setae. Evaporative area extended. Scutellum with erect setae. Inner corial spot diffuse. Costal margin pale on proximal three-quarters. Corium with a subapical corial spot. Membrane with an apical pale spot, veins pale. Profemur dark, at most with apex paler, meso- and metafemur darker apically (Fig. 33 A); setae on profemur long and erect; protibia and mesofemur without spines.</p><p>Abdomen, male genitalia: Pygophore (Fig. 32 I, J) rounded, anterior margin of dorsal aperture rounded. Parameres: Figure 32 (K, L). Aedeagus (Fig. 30 D): conjunctiva with a few minute spines laterally, vesica with lobes weakly sclerotized and spined apically; processus gonopori long and slender.</p><p>Distribution</p><p>Antigua, Dominican Republic, Jamaica, Nevis, and Saba; Haiti (NEW RECORD) (Fig. 31).</p><p>Type material examined</p><p>Holotype ♀, DOMINICAN REPUBLIC, Bayaguana, 4-IX-[19]91, BL Trap, D. Brown, Holotype Heraeus concolor J. A. Slater and R. M. Baranowski (USNM).</p><p>Additional material studied</p><p>ANTIGUA: 1♀, Christian Valley, 14-X-1961, blacklight trap, FAO Insect Survey, Baranowski (USNM).</p><p>DOMINICAN REPUBLIC: 1♂, Finca Goya, 5-IV- [19]89, Bl trap, G. Anzerro (USNM); Azua: 1♂, 8 km NE Padre Las Casas, Rio Las Cuevas, 18°46 ′ N, 70°53 ′ W, 580 m a.s.l., 7-VIII-1990, J. Rawlins &amp; S. Thompson (AMNH); Barahona: 1♂, 5 km NW Barahona, Agr. Exp. Sta., B.L.T., 29/30-IV-[19]78, Woodruff, Fairchild &amp; Mercado (USNM); El Seibo: 5♂, 5♀, Pedro Sanchez, 10-VI-6, blacklight trap at shallow stream, R.E. Woodruff; 2♀, 7 km N Pedro Sanchez, Loma de Chivo, 25-VI-[19]98, blacklight trap, R.M. Baranowski &amp; R.E. Woodruff (USNM); 1♀, 20- VI-[19]98, blacklight trap, R.M. Baranowski &amp; R.E. Woodruff (USNM); 1♀, 5000 ft, R.E. Woodruff &amp; P.H. Freytag (USNM); La Altagracia: 2♂, 7♀, one without abdomen, Nisibon, finca ‘Papagallo’, 17-VI- 1999, blacklight trap, abandoned building, R.E. Woodruff &amp; R.M. Baranowski (USNM); 2♂, 2♀, Nisibon, Papagallo, 26-VI-[19]98, blacklight trap, R.M. Baranowski &amp; R.E. Woodruff (USNM); 1♂, 4♀, Nisibon, 5 mi. W Las Lagunas, 16-VI-1999, blacklight trap, R.E. Woodruff &amp; R.M. Baranowski (USNM); 1♀, 19-VI-1999, abandoned house (USNM); 1♂, 18-VI- 1999 (USNM); 3♂, 1♀, Rio Maimon, near La Guama, 18-VI-[19]98, black light trap, P.H. Freytag &amp; R.E. Woodruff (USNM); 1♀, Rio Maimon, La Laguna, Nisibon, 18-Vi-[19]98, black light trap, R.E. Woodruff (USNM); La Romana: 1♀, Higueral, 15-VIII- [19]77, blacklight trap, R.E. Woodruff &amp; E. Folch (USNM); 1♂, 17-VIII-[19]77, blacklight trap, R.E. Woodruff &amp; E. Folch (USNM); Monseñor Novel: 1♀, Bonao, Jacaranda Hotel, 28-VI-[19]98, blacklight trap, R.M. &amp; R.E. Woodruff (USNM); Monte Plata: 1♀, NE Sierra de la Agua, 1-V-[19]78, blacklight trap, R.E. Woodruff, G. Fairchild &amp; E. Mercado (USNM); Pedernales: 1♀, 19-VI-[19]76, blacklight trap, R.E. Woodruff (AMNH); 1♀, 24 km N Cabo Rojo, 11- VI-[19]98, blacklight trap, P.H. Freytag, B.K. Dezier &amp; R.E. Woodruff (USNM); 1♀, 26 km N Cabo Rojo, 10-VI-[19]98, blacklight trap, P.H. Freytag, B.K. Dezier &amp; R.E. Woodruff (USNM); San Juan: 1♀, 6 km W San Juan, 7-VIII-1979, UV light, O’Brien &amp; Marshall (USNM); San Pedro de Macoris: 1♂, 1♀, 0.8 mi. W Juan Dolio, 22-VI-1998, black light trap, R.M. Baranowski &amp; R.E. Woodruff (USNM).</p><p>HAITI: 1♂, III-[19]28, A.J. Poole (USNM).</p><p>JAMAICA: 1♂, Parish of St Andrew, 4000 ft, Holywell For. Camp, 18-X-1971, R.M. Baranowski (USNM); 1♀, Parish of St James, 4 mi. E of Montego Bay, Ironshore, 30-V-[19]70, U.V. light trap, E.G. Farnworth (USNM).</p><p>NEVIS ISLAND: 1♂, 1♀, Hurricane Cove, 22-VII- [19]91, BL trap, H.V. &amp; R.M. Baranowski (USNM); 1♀, 18-IX-[19]92, B. Brandy (USNM); 3♂, 8-IX- [19]92, B. Brandy (USNM); 1♂, 1♀, Butler Village, 29-IX-[19]92, B.L. trap, B. Brandy (USNM); 1♀, 22- IX-[19]92, B.L. trap, B. Brandy (USNM).</p><p>SABA ISLAND: 2♂, 3♀, Mt Scenery, 26-IX-[19]90, H.V. &amp; R.M. Baranowski (USNM); 4♂, 1♀, 19-VIII- [19]92, H.V. &amp; R.M. Baranowski (USNM); 1♂, 20- VII-[19]91, H.V. &amp; R.M. Baranowski (USNM); 1♂, N.A. Booby Hill, 19-IX-[19]93, B.L.T., T. van Oosteren, Baranowski coll. (USNM).</p></div>	https://treatment.plazi.org/id/6F64DB1B5E7BFF92FCF3FA1CFB59FEC3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pablo M. Dellapé;María C. Melo;Thomas J. Henry	Pablo M. Dellapé, María C. Melo, Thomas J. Henry (2016): A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species. Zoological Journal of the Linnean Society 177 (177): 29-134, DOI: 10.1111/zoj.12362
6F64DB1B5E7DFF91FC1CFE7BFDF7FB5E.text	6F64DB1B5E7DFF91FC1CFE7BFDF7FB5E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heraeus ecuatorianus	<div><p>HERAEUS ECUATORIANUS SP. NOV.</p><p>(FIGS 25 H, 30E, 31, 32M–P, 33B)</p><p>Diagnosis</p><p>Labium extending to metacoxae. Distiflagellomere with a sub-basal whitish band. Membrane with an apical pale spot. Meso- and metafemur with a subapical brown band. Pygophore rounded, inner projections elongate, more pronounced posteriorly; declivent posteriorly in lateral view. Parameres with inner projections narrow and elongate.</p><p>Description (Fig. 25 H)</p><p>Total length 5.70.</p><p>Head: Brown, shiny, coriaceus; with abundant short recumbent and sparse long erect setae dorsally. Head length 1.22, width 0.83. Postocular length 0.43. Eyes not surpassing the dorsal margin of head in lateral view. Ocelli at level of an imaginary line passing the posterior border of eyes. Interocular width 0.42, interocellar width 0.26. Labium pale brown with erect setae, extending to metacoxae. Labial segment lengths: I 0.82, II 0.86, III 0.82, and IV 0.34. Antennae pale brown, scape externally, pedicel, basiflagellomere distally, and distiflagellomere darker, distiflagellomere with a sub-basal whitish band; with abundant, short recumbent and sparse semi-erect setae more abundant on distiflagellomere. Antennal lengths: scape 0.48, pedicel 1.08, basiflagellomere 1.01, and distiflagellomere 1.06. Length of pale band on distiflagellomere 0.29.</p><p>Thorax: Pronotum: anterior pronotal lobe brown, collar and posterior lobe paler, with irregular pale areas, each humeral angle with a whitish spot; collar punctate, delimited posteriorly by a punctate sulcus; anterior and posterior lobes with short recumbent setae and sparse long erect setae on anterior lobe. Pronotum pruinose, punctate, punctures larger on posterior lobe. Collar length 0.08, anterior lobe length 0.56, posterior lobe length 0.51; anterior lobe width 0.89, posterior lobe width 1.39. Pleurae brown, acetabular areas paler, punctate, with short recumbent setae. Evaporative area extended. Scutellum brown, darker basally, with a darker longitudinal median stripe, apex paler, pruinose, punctate, with short recumbent and long erect setae, as on anterior pronotal lobe. Hemelytra brown, irregularly pigmented, costal margin pale on proximal threequarters; apex of corium darker, with a subapical whitish spot and an inner pale spot; setae short and semi-erect; membrane brown, with an apical pale spot, veins paler (Fig. 25 H). Legs: Coxae, protrochanter, profemur, except apex, and a subapical band on meso- and metafemur brown, remainder of legs pale brown, apex of tibiae, apex of tarsi, and pretarsi darker (Fig. 33 B); setae semi-erect, longer on profemur. Profemur with spines restricted to apical half.</p><p>Abdomen: Brown, with abundant short recumbent setae. Male genitalia: Pygophore (Fig. 32 M, N) rounded, anterior margin of dorsal aperture rounded, inner projections elongate, more pronounced posteriorly; declivent posteriorly in lateral view. Parameres: Figure 32 (O, P). Inner projections narrow and elongate. Aedeagus (Fig. 30 E): conjunctiva with spines laterally beneath the ejaculatory reservoir; vesica with two pairs of lobes, the anterior lobe short, not sclerotized, and with spines; the posterior lobe larger, strongly sclerotized, with a few spines distally; processus gonopori long and slender.</p><p>Distribution</p><p>Ecuador (Fig. 31).</p><p>Etymology</p><p>The specific epithet ‘ ecuatorianus ’ is an adjective and refers to Ecuador, the country where the known specimens were collected.</p><p>Type material</p><p>Holotype: ♂, ECUADOR, Guay, Olon, 29-II-1976, blacklight, J. Cohen (USNM).</p><p>Paratypes: 2♂, 1♀, same data as for holotype (USNM); 1♂, 2♀, ECUADOR, Victoria, Arenillas, 150 m a.s.l., 18/19-VIII-1977, L. Peña G. (USNM); 1♂, 1♀, 18/ 19-VIII-1977, L. Peña G. (MLP); 1♂, 1♀, Pilalo (W), 9/10-X-1977, L. Peña G. (USNM).</p></div>	https://treatment.plazi.org/id/6F64DB1B5E7DFF91FC1CFE7BFDF7FB5E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pablo M. Dellapé;María C. Melo;Thomas J. Henry	Pablo M. Dellapé, María C. Melo, Thomas J. Henry (2016): A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species. Zoological Journal of the Linnean Society 177 (177): 29-134, DOI: 10.1111/zoj.12362
6F64DB1B5E7EFF91FE83FAF3FAB4F8D9.text	6F64DB1B5E7EFF91FE83FAF3FAB4F8D9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heraeus loja	<div><p>HERAEUS LOJA SP. NOV.</p><p>(FIGS 25 I, 33C, 34A–D, 35)</p><p>Diagnosis</p><p>Labium extending to metacoxae. Distiflagellomere with a narrow sub-basal pale band. Membrane brown, with veins paler. Metafemur with a subapical dark band. Heraeus loja sp. nov. shares with H. pallidinervis sp. nov. and H. penai sp. nov. a narrow pale band on the distiflagellomere, occupying at most onequarter of the segment length, and a long labium extending to the metacoxae. These three species, however, are readily distinguished by the male genitalia, with H. loja sp. nov. having a slightly declivent pygophore with narrow, elongate inner projections.</p><p>Description (Fig. 25 I)</p><p>Total length 5.89.</p><p>Head: Dark brown, shiny; setae long and erect dorsally. Head length 1.20, width 0.93. Postocular length 0.40. Eyes not surpassing the dorsal margin of head in lateral view. Ocelli placed at level of an imaginary line passing the posterior border of eyes. Interocular width 0.48, interocellar width 0.29. Labium pale brown with erect setae, extending to at least metacoxae. Labial segment lengths: I 0.85, II 0.96, III 0.83, and IV 0.35. Antennae pale brown, distiflagellomere with a narrow sub-basal pale band; setae abundant, short, and semierect. Antennal lengths: scape 0.46, pedicel 1.09, basiflagellomere 0.90, and distiflagellomere 0.96. Length of pale band on distiflagellomere 0.13.</p><p>Thorax: Anterior pronotal lobe and scutellum dark brown, posterior pronotal lobe paler, with pale spots and with a pale spot on humeral angles. Anterior pronotal lobe with long erect setae, posterior lobe with short erect setae. Collar with scattered punctures, delimited posteriorly by a punctate sulcus. Collar length 0.08, anterior lobe length 0.56, posterior lobe length 0.45; anterior lobe width 1.02, posterior lobe width 1.52. Pleurae dark brown, except acetabular areas reddish brown; punctate, with short recumbent setae. Evaporative area extended. Scutellum dark brown, with a median darker longitudinal stripe, apex paler, pruinose, punctate; with erect setae. Hemelytra mostly pale brown, with irregular darker spots, pattern complex (Fig. 25 I). Inner corial spot diffuse; costal margin pale on proximal three-quarters; with a subapical corial spot; membrane brown with veins paler and pale spots between them. Legs: Profemur dark, at most, with apices paler, remainder of legs paler, except metafemur with a subapical dark band (Fig. 33 C); setae on profemur long and erect. Protibia curved, protibia and mesofemur without spines.</p><p>Abdomen: Dark brown, with abundant short recumbent setae. Male genitalia: Posterior margin of pygophore flattened in dorsal view (Fig. 34 A, B), anterior margin of dorsal aperture slightly rounded, with inner projections narrow, elongate, strongly projecting posteriorly. Parameres: Figure 34 (C, D). Aedeagus: spinose; conjunctiva with spines laterally beneath the ejaculatory reservoir; vesica with sclerotized lobes and spines distally and anteriorly; processus gonopori long and slender.</p><p>Distribution</p><p>Ecuador (Fig. 35).</p><p>Etymology</p><p>The specific epithet is a noun in aposition and refers to Loja Province in Ecuador, where the holotype was collected.</p><p>Type material</p><p>Holotype: ♂, ECUADOR, Loja, 26 km W of Loja, along road to Catamayo, 2350 m a.s.l., 18-II-2002, 3°59 ′ 71 ″ S, 79°18 ′ 41 ″ W, T.J. Henry &amp; P.S.F. Ferriera (USNM).</p></div>	https://treatment.plazi.org/id/6F64DB1B5E7EFF91FE83FAF3FAB4F8D9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pablo M. Dellapé;María C. Melo;Thomas J. Henry	Pablo M. Dellapé, María C. Melo, Thomas J. Henry (2016): A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species. Zoological Journal of the Linnean Society 177 (177): 29-134, DOI: 10.1111/zoj.12362
6F64DB1B5E00FFECFF51FB07FE30F8D9.text	6F64DB1B5E00FFECFF51FB07FE30F8D9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heraeus mesoamericanus	<div><p>HERAEUS MESOAMERICANUS SP. NOV.</p><p>(FIGS 30 F, 33D, 34E–H, 36A, 37)</p><p>Diagnosis</p><p>Labium extending to mesocoxae. Distiflagellomere with a sub-basal whitish band. Evaporative area short, distance from dorsal margin of auricle to dorsal margin of evaporative area shorter than distance from dorsal margin of evaporative area to dorsal margin of metapleura. Membrane brown, veins paler, without an apical pale spot. Meso- and metafemur with a subapical narrow brown band. Pygophore apex pointed dorsally in lateral view.</p><p>Description (Fig. 36 A)</p><p>Total length 5.61.</p><p>Head: Brown, shiny, coriaceous, with short recumbent and long erect setae dorsally. Head length 1.18, width 0.85, postocular length 0.43. Eyes not surpassing dorsal margin of head in lateral view. Ocelli posterior of an imaginary line passing the posterior border of eyes. Interocular width 0.42, interocellar width 0.22. Labium pale brown with abundant erect setae, extending to mesocoxae. Labial segment lengths: I 0.72, II 0.82, III 0.60, and IV 0.24. Antennae pale brown, apical half of basiflagellomere and distiflagellomere darker, distiflagellomere with a subbasal whitish band; all segments with abundant short recumbent and sparse short erect setae. Antennal lengths: scape 0.46, pedicel 1.03, basiflagellomere 0.91, and distiflagellomere 1.03. Length of pale band on distiflagellomere 0.24.</p><p>Thorax: Anterior pronotal lobe brown, collar and posterior lobe paler, with four longitudinal pale stripes, humeral angles yellowish; collar delimited posteriorly by a punctate sulcus; anterior and posterior lobes with short recumbent and long erect setae, longer on anterior lobe. Pronotum pruinose, punctate, punctures larger on posterior lobe. Collar length 0.08, anterior lobe length 0.51, posterior lobe length 0.54; anterior lobe width 0.89, posterior lobe width 1.42. Pleurae brown, acetabular areas paler, punctate, with short recumbent setae. Evaporative area short. Scutellum brown, with a longitudinal median dark stripe, apex paler; pruinose, punctate; with short recumbent and long erect setae, as on anterior pronotal lobe. Hemelytra pale brown, irregularly pigmented. Costal margin pale on proximal three-quarters, apex of corium darker with a subapical whitish spot and an small pale inner spot; setae short and recumbent; membrane brown with veins paler (Fig. 36 A). Legs: Coxae, protrochanter, profemur, except distally, and a subapical narrow band on meso- and metafemur brown, remainder of legs pale brown, apex of tibiae, tarsi, and pretarsi darker (Fig. 33 D); setae semi-erect, longest on profemur. Profemur with spines restricted to apical half.</p><p>Abdomen: Brown, with abundant short recumbent setae. Male genitalia: Pygophore (Fig. 34 E, F) rounded, elongate, with pointed posterior margin, apex with a dorsally directed protuberance, declivent posteriorly in lateral view; anterior margin of dorsal aperture rounded. Inner projections subquadrangular. Parameres: Figure 34 (G, H). Aedeagus: conjunctiva and vesica with a few sparse spines laterally, vesica with two sclerotized lobes; processus gonopori long and slender (Fig. 30 F).</p><p>Variability observed in other material studied In the specimens from Belize, the labium is slightly longer, barely extending to metacoxae.</p><p>Distribution</p><p>Belize, Costa Rica, Ecuador, Honduras, and Panama (Fig. 37).</p><p>Etymology</p><p>The specific epithet ‘ mesoamericanus ’ is an adjective meaning Central American, and refers to the known geographic distribution of the species.</p><p>Type material</p><p>Holotype: ♂, COSTA RICA, San Jose, San Isidro de Coronado, 550 ft, 11/12-VI-[19]67, BL trap, E.B. Fogar (USNM).</p><p>Paratypes: 5♂, 11♀, same data as for holotype (USNM); 1♂, 2♀, San Isidro de Coronado, Finca Joseph Smids, 5500 ft, 31-V/2-VI-[19]67, blacklight trap, E.B. Fagan (USNM); 1♂, 1♀, 31-V/2-VI-[19]67, blacklight trap, E.B. Fagan (MLP); Puntarenas: 7♂, 7♀, finca Las Cruces, 6 km S San Vito de Java, 4200 ft, 28-IX/2- X-1986, J.E. Eger (USNM); 1♂, 1♀, 28-IX/2-X- 1986, J.E. Eger (MLP).</p><p>Additional material studied</p><p>BELIZE: 2♂, 3♀, Orange Walk Dist., Chan Chich</p><p>Lodge, 1-IV-[20]09, BLT, E. Flota (USNM); 1♂, Cayo</p><p>Dist., Chiquibul Forest Res., 27-III-[20]08, BL Trap,</p><p>27-III-[20]08, B. Miller (USNM); 1♂, Cayo Dist., Benque</p><p>Viejo del Carmen, Riverside N, Mopan</p><p>River Resort, 20-XI-[20]07, BL Trap, M. Chrysler</p><p>(USNM); 1♂, 13-XII-[20]07, BL Trap, M. Chrysler</p><p>(USNM).</p><p>COSTA RICA: 1♂, Torrazu, 20-VIII-[19]36, on Coffee arabica, E. Marin (USNM); Cartago: 1♀, Turrialba, Int. Amer. Inst. Agr. Sci., blacklight trap, 18/20-VIII- [19]64, R.E. Woodruff (USNM); Puntarenas: 1♂, Turrialba, 27-I-1965, J.A. Slater &amp; N.T. Davis (AMNH); 1♀, on Chrysanthemum, V-[19]87 (USNM).</p><p>ECUADOR: Tungurahua: 2♂, Baños, 6200 ft, 24-I- 1976, Spangler et al. (USNM); 2♂, 3♀, Finca Las Cruces, 6 km S San Vito de Java, 4200 ft, 28-IX/2- X-1986, J.E. Eger (USNM).</p><p>HONDURAS: 1♂, 1♀, Comayagua, Siguatepeque, 25- VIII-[19]66, mosquito trap, R.E. Woodruff (USNM); 1♂, 21 km NW Siguatepeque, Ch. 5 rd., 1-VIII- 1977, O’Brien &amp; Marshall (AMNH); 1♀, Sta. B., 13 km SE El Mochito, 22-VII-1977, C.W. &amp; L. O’Brien &amp; Marshall (AMNH).</p><p>PANAMA: Chiriqui, 3♂, 15♀, Dist. Renacimiento, Santa Clara, 4000 ft, 28/29-V-[19]76, at light, Engleman &amp; Thurman (AMNH); 1♂, 1♀, 28/29-V- [19]76, at light, Engleman &amp; Thurman (MLP); 2♂, 1♀, 20/22-V-[19]77, Engleman (AMNH); 1♂, 3♀, 29- V-[19]76, D. Engleman (AMNH); 1♂, Rovira, Elvira Farm, 15-VIII-[19]64, 220 ft, blacklight trap, A.B. Broce (USNM); 3♂, Rovira, 5-VII-[19]64, 2500 ft, mosquito light trap, A. Bruse (USNM); 1♂, 9-V-[19]64, mosquito light trap, A. Bruse (USNM); 1♀, Rovira, 4-VII- [19]64, blacklight trap, A.B. Brose (USNM); 1♂, 1♀, Volcan Chiriqui, 3500 ft, 4-XI-[19]73, D. Engleman (AMNH); 1♂, 1♀, Volcan Aria, 4500 ft, 25-V- [19]73, lagos, D. Engleman (USNM); 1♂, 11-IV- [19]73, D. Engleman (USNM); 1♀, 2.5 mi. W El Volcan, 4400 ft, Las Lagunas, 9-VII-1974, O’Briens &amp; Marshall (AMNH); 1♂, 2♀, Res. For. La Fortuna, El Vivero, 20-VII-1995, UV-HG lights, L.B. O’Brien (USNM); Panama, 1♂, Cerro Campana, 29-VII- 1995, C.W. &amp; L.B. O’Brien (USNM); 1♀, Interamerican Hwy. 3 km E of Ipeli, 3-V-1992, UV and HG vapour light, A. Gillogly (USNM).</p></div>	https://treatment.plazi.org/id/6F64DB1B5E00FFECFF51FB07FE30F8D9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pablo M. Dellapé;María C. Melo;Thomas J. Henry	Pablo M. Dellapé, María C. Melo, Thomas J. Henry (2016): A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species. Zoological Journal of the Linnean Society 177 (177): 29-134, DOI: 10.1111/zoj.12362
6F64DB1B5E03FFEBFC06FF17FC79F992.text	6F64DB1B5E03FFEBFC06FF17FC79F992.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heraeus mexicanus	<div><p>HERAEUS MEXICANUS SP. NOV.</p><p>(FIGS 30 G, 33E, 34I–L, 36B, 38)</p><p>Diagnosis</p><p>Labium extending to mesocoxae. Distiflagellomere with a pale band. Membrane with an apical pale spot. Metafemur with a subapical dark band. Apex of pygophore pointed posteriorly in lateral view.</p><p>Description (Fig. 36 B)</p><p>Total length 5.32.</p><p>Head: Brown, shiny, coriaceous; with short recumbent and long erect setae dorsally. Head length 1.10, width 0.82. Postocular length 0.38. Eyes not surpassing the dorsal margin of head in lateral view. Ocelli placed at level of an imaginary line passing the posterior border of eyes. Interocular width 0.40, interocellar width 0.22. Labium pale brown, with short erect setae; extending to mesocoxae. Labial segment lengths: I 0.72, II 0.78, III 0.64, and IV 0.32. Antennae pale brown, apex of basiflagellomere and distiflagellomere darker, distiflagellomere with a sub-basal whitish band; all segments with abundant short recumbent and sparse short erect setae. Antennal lengths: scape 0.43, pedicel 1.01, basiflagellomere 0.80, and distiflagellomere 0.88. Length of pale band on distiflagellomere 0.24.</p><p>Thorax: Pronotum pruinose, punctate, punctures larger on posterior lobe. Anterior pronotal lobe brown, collar and posterior lobe paler, with four longitudinal pale stripes, humeral angles yellowish; collar delimited posteriorly by a punctate sulcus; anterior and posterior lobes with short recumbent setae and long erect setae, longer on anterior lobe. Collar length 0.06, anterior lobe length 0.53, posterior lobe length 0.43; anterior lobe width 0.85, posterior lobe width 1.31. Evaporative area extended. Scutellum brown, with a longitudinal median dark stripe, apex paler; pruinose, punctate; with short recumbent and long erect setae, as on anterior pronotal lobe. Hemelytra pale brown, irregularly pigmented. Costal margin pale on proximal three-quarters, apex of corium darker with a subapical whitish spot and an inner small pale spot; setae short and recumbent; membrane irregularly brown, veins paler, with a pale apical spot. (Fig. 36 A). Legs: Coxae, protrochanter, and profemur brown, at most with apices paler, meso- and metafemur with a subapical narrow dark band; remainder of legs pale brown, apex of tibiae, tarsi, and pretarsi darker (Fig. 33 E); setae long and erect, longer on profemur.</p><p>Abdomen: Brown, with abundant short recumbent setae. Male genitalia: Pygophore (Fig. 34 I, J) rounded, with a pointed posterior margin; apex with a small protuberance pointing posteriorly in lateral view; anterior margin of dorsal aperture rounded. Parameres: Figure 34 (K, L). Aedeagus (Fig. 30 G) with minute spines on conjunctiva and sclerotized vesical lobes; processus gonopori long and slender.</p><p>Distribution</p><p>Costa Rica and Mexico (Fig. 38).</p><p>Etymology</p><p>The specific epithet ‘ mexicanus ’ is an adjective referring to Mexico, the country where most of the type series was collected.</p><p>Type material</p><p>Holotype: ♂, MEXICO, N[uevo] L[eon], Anegade Arroya, 16 mi. S Linares, 1250 ft, 9-VII-1963, Duckworth &amp; Davis (USNM).</p><p>Paratypes: 1♂, 2♀, same data as for holotype (USNM); MEXICO: Nuevo Leon: 1♂, 3♀, 3 mi. E Galeana, 5000 ft, 7/9-VIII-1963, Duckworth &amp; Davis (USNM); 1♀, 7/9-VIII-1963, Duckworth &amp; Davis (UNAM); 1♂, 7/9-VIII-1963, Duckworth &amp; Davis (MLP); Campeche: 1♀, Escarcega, 3-VI-1962, light trap, F. Islas S. (USNM); 1♂, 28-IV-1962, light trap, F. Islas S. (USNM); Veracruz: 1♂, 25 mi. S Acayucan, 4-VII- 1965, collected at blacklight trap, P.J. Spangler (USNM); COSTA RICA: 1♂, Puntarenas, San Vito, Darryl Coles Farm, Loma Linda, 30-VII-1986, D.V. Bennett (UCMS); 1♀, Puntarenas, nr Monteverde, 31-V-1988, blacklight, J.E. O’Donnell (UCMS).</p></div>	https://treatment.plazi.org/id/6F64DB1B5E03FFEBFC06FF17FC79F992	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pablo M. Dellapé;María C. Melo;Thomas J. Henry	Pablo M. Dellapé, María C. Melo, Thomas J. Henry (2016): A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species. Zoological Journal of the Linnean Society 177 (177): 29-134, DOI: 10.1111/zoj.12362
6F64DB1B5E04FFEAFC10F9A0FA7BFDF7.text	6F64DB1B5E04FFEAFC10F9A0FA7BFDF7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heraeus morganae	<div><p>HERAEUS MORGANAE SP. NOV.</p><p>(FIGS 31, 33 F, 34M–P, 36C)</p><p>Diagnosis</p><p>Labium extending to metacoxae. Distiflagellomere with a wide sub-basal yellowish band. Membrane with an apical pale spot. Meso- and metafemur with a subapical brown band. Pygophore rounded, declivent, and truncate posteriorly in lateral view; anterior margin of dorsal aperture subquadrangular, inner projections elongate, more pronounced posteriorly. Inner projections of parameres broad.</p><p>Description (Fig. 36 C)</p><p>Total length 5.51.</p><p>Head: Brown, shiny, coriaceous, with short recumbent and long erect setae dorsally. Head length 1.18, width 0.83. Postocular length 0.43. Eyes not surpassing the dorsal margin of head in lateral view. Ocelli placed at level of an imaginary line passing the posterior border of eyes. Interocular width 0.40, interocellar width 0.24. Labium pale brown with erect setae, extending to metacoxae. Labial segment lengths: I 0.77, II 0.74, III 0.72, and IV 0.34. Antenna pale brown, distiflagellomere with a wide sub-basal yellowish band; setae abundant, short, and recumbent. Antennal lengths: scape 0.46, pedicel 1.08, basiflagellomere 0.96, and distiflagellomere 1.01. Length of pale band on distiflagellomere 0.34.</p><p>Thorax: Pronotum brown, anterior pronotal lobe darker, posterior lobe with irregular longitudinal pale stripes, each humeral angle with a whitish spot; collar with scattered punctures, delimited posteriorly by a punctate sulcus, punctures larger on posterior lobe; anterior and posterior lobes with short recumbent setae, and anterior lobe with long erect setae. Collar length 0.08, anterior lobe length 0.53, posterior lobe length 0.48; anterior lobe width 0.86, posterior lobe width 1.34. Pleurae brown, acetabular areas paler, punctate, with short recumbent setae. Evaporative area extended. Scutellum brown, apex paler, pruinose, punctate, with short recumbent and long erect setae, as on anterior pronotal lobe. Hemelytra pale brown, costal margin pale on proximal three-quarters, apex of corium darker, with a subapical whitish spot and an inner pale spot; setae short and recumbent; membrane irregularly brown, with an apical pale spot, veins paler (Fig. 36 C). Legs: Coxae, protrochanter, profemur, except distally, and a subapical band on meso- and metafemur brown, remainder of legs pale brown, apex of tibiae, tarsi, and pretarsi darker (Fig. 33 F); setae semi-erect, longer on profemur. Profemur with spines restricted to apical two-thirds. Protibia slightly curved.</p><p>Abdomen: Brown, with abundant short recumbent setae. Male genitalia: Pygophore (Fig. 34 M, N) rounded, declivent and truncate posteriorly in lateral view; anterior margin of dorsal aperture subquadrangular, inner projections elongate, more pronounced posteriorly. Parameres: inner projections broad (Fig. 34 O, P). Aedeagus: vesica with spines laterally, and with two sclerotized lobes with many spines anteriorly; processus gonopori long and slender.</p><p>Distribution</p><p>Panama (Fig. 31).</p><p>Etymology</p><p>Named after our friend and colleague Cecilia C. Morgan (MLP).</p><p>Type material</p><p>Holotype: ♂, PANAMA, Cano Saddle, Gatun Lake, close’s Plta, 12-V-[19]23, B.C. Shannon (USNM).</p><p>Paratypes: 2♂, 3♀, same data as for holotype (USNM); PANAMA, 1♂, 4♀, Cano Saddle, Gatun Lake, 12- V-[19]23, B.C. Shannon (USNM); 1♂, 12-V-[19]23, B.C. Shannon (MLP); 1♀, 27-IV-[19]23, B.C. Shannon (MLP); 1♂, CC, 28-IV-1973, H. Hespenheide (AMNH).</p></div>	https://treatment.plazi.org/id/6F64DB1B5E04FFEAFC10F9A0FA7BFDF7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pablo M. Dellapé;María C. Melo;Thomas J. Henry	Pablo M. Dellapé, María C. Melo, Thomas J. Henry (2016): A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species. Zoological Journal of the Linnean Society 177 (177): 29-134, DOI: 10.1111/zoj.12362
6F64DB1B5E05FFE9FC19FDA8FAB8F99A.text	6F64DB1B5E05FFE9FC19FDA8FAB8F99A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heraeus nicaraguensis	<div><p>HERAEUS NICARAGUENSIS SP. NOV.</p><p>(FIGS 36 D, 38, 39A, 40A–D)</p><p>Diagnosis</p><p>Labium extending to mesocoxae. Distiflagellomere with a pale band. Membrane with an apical pale spot. Metafemur with a subapical dark band. Apex of pygophore pointed dorsally in lateral view.</p><p>The diagnostic characters relate H. nicaraguensis sp. nov. to H. bahiensis sp. nov. They can be distinguished by the darker general colouration of H. nicaraguensis sp. nov., with a blackish head and a dark posterior pronotal lobe with four longitudinal pale stripes, whereas in H. bahiensis sp. nov., the head is dark reddish brown and the posterior pronotal lobe is brown, with diffuse longitudinal stripes.</p><p>Description (Fig. 36 D)</p><p>Total length 5.32. Dark brown species.</p><p>Head: Dark brown, shiny, coriaceous, with short recumbent and long erect setae dorsally. Head length 1.10, width 0.83, postocular length 0.43. Eyes not surpassing the dorsal margin of head in lateral view. Ocelli at level of an imaginary line passing the posterior border of eyes. Interocular width 0.40, interocellar width 0.24. Labium pale brown, with erect setae; extending to mesocoxae. Labial segment lengths: I 0.70, II 0.75, III 0.48, and IV 0.29. Antennae brown, basal half of basiflagellomere and base of distiflagellomere darkened, distiflagellomere with a wide sub-basal yellowish band; setae abundant, short, and recumbent. Antennal lengths: scape 0.42, pedicel 0.99, basiflagellomere 0.80, and distiflagellomere 1.04. Length of pale band on distiflagellomere 0.40.</p><p>Thorax: Pronotum dark brown, posterior pronotal lobe slightly paler, with four longitudinal pale bands and a pale spot on each humeral angle. Anterior and posterior lobes with short recumbent setae and long erect setae, longer on anterior lobe. Collar length 0.08, anterior lobe length 0.54, posterior lobe length 0.46; anterior lobe width 0.86, posterior lobe width 1.28. Scutellum dark brown, apex paler, pruinose, punctate, with short recumbent and erect setae, as on anterior lobe. Pleurae dark brown, punctate, with short recumbent setae. Evaporative area extended. Hemelytra dark brown, with a pale inner corial spot and a subapical corial spot; costal margin pale on proximal three-quarters; membrane brown, with veins paler and an elongate apical pale spot (Fig. 36 D). Legs: Coxae, protrochanter, and profemur dark brown, at most with apices paler, meso- and metafemur with a subapical dark band; remain- der of legs pale brown, apex of tibiae, tarsi, and pretarsi darker (Fig. 39 A); setae long and erect, longest on profemur.</p><p>Abdomen: Dark brown, with abundant short recumbent setae. Male genitalia: Pygophore (Fig. 40 A, B) rounded, apex with a dorsally directed protuberance, anterior margin of dorsal aperture slightly rounded. Parameres: Figure 40 (C, D). Aedeagus: spinose; conjunctiva with spines laterally beneath the ejaculatory reservoir; vesica with spines laterally and sclerotized lobes with spines distally; processus gonopori long and slender.</p><p>Distribution</p><p>Costa Rica and Nicaragua (Fig. 38).</p><p>Etymology</p><p>The specific epithet ‘ nicaraguensis ’ is an adjective referring to Nicaragua, the Central American country where the holotype was collected.</p><p>Type material</p><p>Holotype: ♂, NICARAGUA, Puerto Cabezas, VII- 1971, J. Maldonado C. (USNM).</p><p>Paratypes: 1♂, 1♀, same data as for holotype (USNM); COSTA RICA: Heredia: 1♀, La Selva OTS Station, 22/23-V-1988, blacklight, J. O’Donnell (UCMS); 1♀, 24/25-V-1988, blacklight, J. O’Donnell (UCMS).</p></div>	https://treatment.plazi.org/id/6F64DB1B5E05FFE9FC19FDA8FAB8F99A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pablo M. Dellapé;María C. Melo;Thomas J. Henry	Pablo M. Dellapé, María C. Melo, Thomas J. Henry (2016): A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species. Zoological Journal of the Linnean Society 177 (177): 29-134, DOI: 10.1111/zoj.12362
6F64DB1B5E06FFE6FC29F9B3FED1FB83.text	6F64DB1B5E06FFE6FC29F9B3FED1FB83.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heraeus pacificus Barber 1925	<div><p>HERAEUS PACIFICUS BARBER, 1925</p><p>(FIGS 36 E, 38, 39B, 40E–H)</p><p>Heraeus pacificus Barber, 1925: 244 –246; Barber, 1934: 286; Slater, 1964: 1083; Linsley &amp; Usinger, 1966: 134; Ashlock, 1972: 101; Parkin et al., 1972: 102; Linsley, 1977: 11; Harrington, 1980: 109; Schaefer et al., 1980: 46; Froeschner, 1981: 43; Froeschner, 1985: 22; Peck, 2001: 258.</p><p>Diagnosis</p><p>Ocelli posterior to an imaginary line passing the posterior border of eyes. Labium extending to or slightly surpassing mesocoxae. Antennae uniformly dark brown. Pronotum with long erect setae only on anterior pronotal lobe. Membrane brown, without an apical corial spot. Metafemur with a dark subapical band. Anterior margin of dorsal aperture of pygophore slightly rounded with lateral regions angulate.</p><p>Heraeus pacificus together with H. pulchellus and H. plebejus are the only species in the group that lack a pale band on the distiflagellomere (although in some specimens of H. pulchellus a pale band is slightly evident). Heraeus pacificus can be distinguished by the uniformly coloured dark-brown antenna and the ocelli placed posterior of an imaginary line passing the posterior border of eyes; H. pulchellus and H. plebejus have the antennae pale brown to brown with the apex of basiflagellomere and distiflagellomere darker, and the ocelli are placed at the level of an imaginary line passing the posterior border of eyes.</p><p>Redescription (Fig. 36 E)</p><p>Head: Brown, shiny, coriaceous, with short recumbent and long erect setae dorsally. Eyes not surpassing the dorsal margin of head in lateral view. Ocelli posterior of an imaginary line passing the posterior border of eyes. Labium pale brown, with short erect setae; extending to or slightly surpassing mesocoxae. Antenna uniformly dark brown, with short recumbent and long erect setae.</p><p>Thorax: Pronotum brown, pruinose, punctate. Posterior pronotal lobe with pale spots and with a pale spot on each humeral angle. Anterior and posterior lobes with short recumbent setae, and erect longer setae on anterior lobe. Scutellum brown, with long erect setae. Evaporative area extended. Hemelytra: irregularly brown, costal margin pale on proximal three-quarters; inner corial spot well developed; membrane brown, with veins paler; setae short and recumbent. Legs: coxae, protrochanter, and profemur dark, at most with apices paler, meso- and metafemur with a narrow subapical dark band; apex of tibiae, tarsi, and pretarsi darker (Fig. 39 B); setae long and erect, longest on profemur.</p><p>Abdomen: Brown, with abundant short recumbent setae. Male genitalia: Pygophore (Fig. 40 E, F) rounded, with posterior margin straight, anterior margin of dorsal aperture almost straight, inner projections long, slightly bilobed. Parameres: Figure 40 (G, H). Aedeagus: conjunctiva unspined, with an anterior lobe at base of the ejaculatory reservoir; vesica with minute spines lateral to the ejaculatory reservoir and on the lobes; processus gonopori long and slender.</p><p>Variability observed</p><p>We have examined specimens from Santa Cruz Island that differ from Barber’s (1925) original description.</p><p>These specimens are paler; the head is slightly narrower than the anterior lobe of the pronotum; all antennal segments have abundant setae; the labium is shorter, extending to mesocoxae, and in some specimens the labium surpasses the mesocoxae, extending to the anterior region of the metacoxae; and the hemelytra have short recumbent setae. In addition, we have observed some characters not mentioned in the original description: the scutellum has setae longer than on the anterior lobe of pronotum; the legs have short semi-erect setae; and the anterior tibiae bear small denticles ventrally that are more conspicuous in the males.</p><p>Distribution</p><p>Galapagos: Santa María Island, San Salvador Island, and Santa Cruz Island; Fernandina Island, Floreana Island, Santiago Island (NEW RECORDS) (Fig. 38).</p><p>Type material</p><p>This species was described based on a male holotype, and two female and one male paratypes from James Island, deposited in AMNH.</p><p>Material studied</p><p>GALAPAGOS: Santa Cruz Island: 2♀, Indefatigable Island, Puerto Azora, XII-1971/III-[19]72, at light, J. Vagvolgyi (CAS); 23♂, 32♀, Hornemann Ranch, 8-II- 1964, I.L. Wiggins (CAS); 1♂, 1♀, Hornemann Farm, 220 m a.s.l. 5-IV-1964, D.Q. Cavagnaro (CAS); 1♂, 1♀, D.Q. Cavagnaro (MLP); 1♀, 7-V-1964, D.Q. Cavagnaro (CAS); 1♂, grassland, 750 m a.s.l., 10-IV-1964, D.Q. Cavagnaro (CAS); 1♂, 4♀, Academy Bay, Darwin Research Station, 8-II-1964, R.O. Schuster (CAS); 1♀, 12-II-1964, D.Q. Cavagnaro &amp; R.O. Schuster (CAS); 1♂, 1♀, vic. Media Luna, 4-VI-[19]70, 600 m a.s.l., at light, Miconia ‘Belt’, R. Silverglied &amp; T.J. de Vries (USNM); 2♂, vic. Media Luna, 4-VI-[19]70, 600 m a.s.l., at light, Miconia ‘Belt’, R. Silverglied (USNM); 2♂, 2♀, Cerro Crocker, 800 m a.s.l., 19-VI-[19]85, volcano crater, ferns &amp; Psychotria litter, S. &amp; J. Peck (USNM); 9♂, 19♀, Los Gemelos, 31 km N Sta. Ro[sa], 15-VII-[19]85, Scalesia forest, 570 m a.s.l., FIT &amp; Malaise, S. &amp;. J. Peck (USNM); 17♂, 34♀, 13-VI/15-VII-[19]85, FIT &amp; Malaise, S. &amp;. J. Peck (USNM); 1♂, 2♀, Scalesia forest litter, FIT &amp; Malaise, S. &amp;. J. Peck (USNM); 1♀, 9 km N Los Gemelos, 330 m a.s.l., 17-V-[19]91, up. trans. for UV light, J. Heraty &amp; S. Peck (USNM); 1♂, El Granillo, 9 km N Los Gemelos, 15-VI-[19]91, trans. for UV light, S. Peck (USNM); 8♂, 12♀, 4 km NE St Rosa, 28-VI- [19]85, Scalesia forest litter N of Cerro Banderas, S. &amp; J. Peck (USNM); 1♂, 1♀, 4 km N Bellavista Medialuna, 620 m a.s.l., 14-V/13-VII-[19]85, Miconia zone, FIT, S. &amp; J. Peck (USNM); 1♀, Media Luna, 600 m a.s.l., 29-V-[19]91, wet Miconia litter, S. Peck (USNM); 1♂, 35 km W Bellavista, Finca Vilema, 210 m a.s.l., Scalesia pasture, FIT, 1/30-V-[19]92, S. Peck (USNM); Isabella Island: 7♂, 12♀, Alcedo, 20/24-VI- [19]91, Crater rim, UV light, 1100 m, S. Peck (USNM); 1♀, Volcan Alcedo crater rim, 1100 m a.s.l., 21/23-VI- 1991, CVALC-09–017, C. Vogel (USNM); 1♀, Volcan Alcedo, 21/26-VI-[19]91, C. Vogel (USNM); 1♂, 3♀, NE rim Alcedo, 1100 m a.s.l., 21/25-VI-[19]91, pit &amp; bottle tps under shrubs, S. Peck (USNM); 1♀, Alcedo SE crater rim, 23-VI-[19]91, 1150 m a.s.l., Bermoss &amp; epiphytes on trees, S. Peck (USNM); 1♂, Alcedo, 10 km SW NE Playa, 1100 m a.s.l., 24-VI-[19]91, Heraty, scrub for. H91-114 (USNM); 1♀, Cerro Azul, 7 km NE Caleta Iguana, 700 m a.s.l., pampas, 23-V-[19]91, leaf litter sifting, S. Peck (USNM); 1♀, Cerro Azul, 2 km NE Caleta Iguana, FIT, 19/25-V-[19]91, 155 m a.s.l., S. Peck (USNM); 1♀, Cerro Azul, 300 m a.s.l., 18/26-C-1991, C. Vogel (USNM); 3♀, 350 m a.s.l., 22/25-V-1991, C. Vogel (USNM); 1♀, Cerro Azul, 8 km W Cal. Iguana, 850 m a.s.l., 22-V-[19]91, crater/grass, J. Heraty (USNM); Fernandina Island: 1♀, 10 km NE Cabo Hammond, 400 m a.s.l., 6/9-V-[19]91, open meadow, UV, S. &amp; J. Peck (USNM); Santiago Island: 1♂, 1♀, Pla. Espumilla, 2 m, arid zone opening, 3/10-VI-[19]91, UV light, S. Peck (USNM); 1♂, 1♀, Aguacate Camp., 550 m a.s.l., 4-7/ 8-VI-[19]91, humid forest, UV light, S. Peck (USNM); Floreana Island: 1♀, base Cerro Pajas, 300 m a.s.l., 28- III/23-IV-[19]96, Scalesia forest, Malaise, S. Peck (USNM).</p></div>	https://treatment.plazi.org/id/6F64DB1B5E06FFE6FC29F9B3FED1FB83	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pablo M. Dellapé;María C. Melo;Thomas J. Henry	Pablo M. Dellapé, María C. Melo, Thomas J. Henry (2016): A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species. Zoological Journal of the Linnean Society 177 (177): 29-134, DOI: 10.1111/zoj.12362
6F64DB1B5E09FFE5FEA9FBBBFD49FF4D.text	6F64DB1B5E09FFE5FEA9FBBBFD49FF4D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heraeus pallidinervis	<div><p>HERAEUS PALLIDINERVIS SP. NOV.</p><p>(FIGS 35, 36 F, 39C, 40I–L)</p><p>Diagnosis</p><p>Labium extending to metacoxae. Distiflagellomere with a narrow pale band sub-basally. Membrane brown, with veins paler and without an apical corial spot. Meso- and metafemur with a wide subapical brown band. In addition to Heraeus pallidinervis sp. nov., H. penai sp. nov. and H. loja sp. nov. have a narrow pale band on the distiflagellomere, occupying about onequarter of the segment length, and a long labium extending to the metacoxae; but H. pallidinervis sp. nov. has a broader pale band on the metafemur than the other two species. The male genitalic characters also differentiate these species.</p><p>Description (Fig. 36 F)</p><p>Total length 4.80.</p><p>Head: Convex dorsally, dark brown, shiny, coriaceous, with short recumbent and long erect setae. Head length 1.15, width 0.77, postocular length 0.40. Eyes with setae between ommatidea; not surpassing dorsal margin of head in lateral view. Ocelli small, at level of an imaginary line passing the posterior border of eyes. Interocular width 0.45, interocellar width 0.24. Labium pale brown, with erect setae; extending to metacoxae. Labial segment lengths: I 0.77, II 0.80, III 0.72, and IV 0.34. Antennae pale brown, scapus, apical region of pedicel and basiflagellomere darker, distiflagellomere brown with a narrow pale band sub-basally; all segments with abundant short recumbent and sparse semierect setae. Antennal lengths: scape 0.43, pedicel 0.88, basiflagellomere 0.75, and distiflagellomere 0.99. Length of pale band on distiflagellomere 0.27.</p><p>Thorax: Pronotum pruinose; dark brown, except collar and four longitudinal stripes on posterior lobe paler, humeral angles with a pale spot; punctate, more conspicuous on posterior lobe; anterior lobe with short recumbent and long erect setae. Collar length 0.08, anterior lobe length 0.48, posterior lobe length 0.32; anterior lobe width 0.80, posterior lobe width 1.18. Pleurae dark brown, punctate, with short recumbent setae. Evaporative area extended. Scutellum dark brown, pruinose, punctate, with short recumbent and long erect setae. Hemelytra pruinose, with short recumbent setae, costal margin pale on proximal three-quarters, with a subapical corial spot, colouration pattern as Figure 36 (F). Membrane brown with pale spots between veins, veins pale. Legs: Coxae and protrochanter brown; profemur brown, except apical region paler; meso- and metafemur yellowish with a wide subapical brown band; tibiae and tarsi pale brown, tibiae darker at apex (Fig. 39 C); setae short and semi-erect. Profemur with two rows of spines on apical two-thirds.</p><p>Abdomen: Dark brown, with abundant short recumbent setae. Pygophore (Fig. 40 I, J) rounded, slightly quadrangular, anterior margin of dorsal aperture rounded, inner projections elongate, more produced posteriorly; strongly declivent posteriorly in lateral view. Parameres: Figure 40 (K, L). Aedeagus: spinose, spines relatively larger than in most species; conjunctiva with spines laterally beneath the ejaculatory reservoir; vesica with spines laterally and two pairs of lobes, the posterior pair sclerotized and without spines, and the anterior pair with spines; processus gonopori long and slender.</p><p>Distribution</p><p>Bolivia and Brazil (Fig. 35).</p><p>Etymology</p><p>The specific epithet ‘ pallidinervis ’ is an adjective referring to the contrasting pale colouration of the veins on a darker brown membrane.</p><p>Type material</p><p>Holotype: ♂, BOLIVIA, La Paz, (SE) Coroico, 1800– 2100 m a.s.l., 30-XI/2-XII-[19]84, L.E. Peña G. (USNM).</p><p>Paratypes: BOLIVIA: La Paz: 3♂, 2♀, Yungas, Inquisivi, 5-XII-1984, L.E. Peña (USNM); 1♂, 5-XII- 1984, L.E. Peña (MLP); BRAZIL: Minas Gerais: 1♀, Varginha, II-1972, M. Alvarenga (AMNH).</p></div>	https://treatment.plazi.org/id/6F64DB1B5E09FFE5FEA9FBBBFD49FF4D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pablo M. Dellapé;María C. Melo;Thomas J. Henry	Pablo M. Dellapé, María C. Melo, Thomas J. Henry (2016): A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species. Zoological Journal of the Linnean Society 177 (177): 29-134, DOI: 10.1111/zoj.12362
6F64DB1B5E0AFFE5FE89FEFDFB29F9C7.text	6F64DB1B5E0AFFE5FE89FEFDFB29F9C7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heraeus penai	<div><p>HERAEUS PENAI SP. NOV.</p><p>(FIGS 29, 30 H, 36G, 39D, 40M–P)</p><p>Diagnosis</p><p>Labium almost extending to metacoxae. Distiflagellomere with a narrow pale band sub-basally. Membrane brown, with veins paler. Metafemur with a narrow subapical brown band.</p><p>In addition to Heraeus penai sp. nov., only H. pallidinervis sp. nov. and H. loja sp. nov. have a narrow pale band on distiflagellomere, occupying about one-quarter of segment length, and a long labium extending to metacoxae. The male genitalic characters, however, will differentiate these species.</p><p>Description (Fig. 36 G)</p><p>Total length 5.61.</p><p>Head: Brown, shiny, coriaceous, with abundant short recumbent and sparse long erect setae. Head length 1.22, width 0.84. Postocular length 0.43. Eyes not surpassing the dorsal margin of head in lateral view. Ocelli at level of an imaginary line passing the posterior border of eyes. Interocular width 0.43, interocellar width 0.26. Labium pale brown with erect setae, almost extending to metacoxae. Labial segment lengths: I 0.83, II 0.96, III 0.80, and IV 0.32. Antennae pale brown, except apex of scapus, pedicel, and basiflagellomere; distiflagellomere darker, with a narrow pale band subbasally; all segments with abundant short semi-erect and sparse longer erect setae. Antennal lengths: scape 0.46, pedicel 0.96, basiflagellomere 0.86, and distiflagellomere 0.98. Length of pale band on distiflagellomere 0.24.</p><p>Thorax: Anterior pronotal lobe brown, collar and posterior lobe paler, with irregular paler areas, humeral angles with a whitish spot; collar delimited posteriorly by a punctuate sulcus; punctate, punctures larger on posterior lobe; anterior and posterior lobes with short recumbent and erect setae, longer erect setae on anterior lobe. Collar length 0.07, anterior lobe length 0.53, posterior lobe length 0.50; anterior lobe width 0.90, posterior lobe width 1.39. Pleurae brown, acetabular areas paler, punctate, with short recumbent setae. Evaporative area extended. Scutellum brown, with a darker longitudinal median stripe, pruinose, punctate, with short recumbent and long erect setae, as for anterior pronotal lobe. Hemelytra brown, costal margin pale on proximal three-quarters, apical half of corium darker, with a subapical whitish spot and a small pale rounded spot internally (Fig. 36 G); setae short and semierect; membrane brown with pale areas, veins paler. Legs: Coxae, protrochanter, profemur, except distally, and a subapical narrow band on metafemur brown, remainder of legs pale brown; mesofemur with a narrow band subapically and apex of tibiae darker (Fig. 39 D); setae abundant and semi-erect, longest on profemur. Profemur with spines restricted to apical two-thirds.</p><p>Abdomen: Brown, with abundant short recumbent setae. Male genitalia: Pygophore (Fig. 40 M, N) rounded, anterior margin of dorsal aperture slightly rounded, inner projections subquadrangular; declivent posteriorly in lateral view. Outer projection of parameres not developed (Fig. 40 O, P). Aedeagus (Fig. 30 H): conjunctiva with spines laterally beneath the ejaculatory reservoir; vesica with two pairs of lobes, the anterior lobe short, unsclerotized, and with many spines, and the posterior lobe larger and sclerotized, with a few spines distally; processus gonopori long and slender.</p><p>Distribution</p><p>Argentina, Bolivia, and Paraguay (Fig. 29).</p><p>Etymology</p><p>This species is named after the late Luiz E. Peña, who collected the holotype and some of the paratypes.</p><p>Type material</p><p>Holotype: ♂, BOLIVIA, Mataral (N), V. Grande, 1800– 2000 m a.s.l., 15/17-XII-1984, L.E. Peña (USNM).</p><p>Paratypes: 2♂, 1♀, same data as for holotype (USNM); 2♂, 1♀, Chuquisaca, Monteagudo, 24-XII-[19]84, L.E. Peña (USNM); 1♂, 1♀, 24-XII-[19]84, L.E. Peña (MLP); 4♂, 1♀, (E) Muyupampa, 1600 m a.s.l., 21/ 25-XII-[19]84, L.E. Peña G. (USNM); La Paz: 3♀, Rio Coroico, 1200 m a.s.l., 24/26-XI-[19]84, L.E. Peña (USNM); 7♀, Tres Esteros, Guanay, 19/25-VIII- [19]89, L.E. Peña (USNM); 1♀, Yungas, Coripata, 1700 m a.s.l., 1-XII-1984, L.E. Peña (USNM); 1♀, N La Paz, Mapiri, 10/16-VIII-[19]89, L.E. Peña (USNM); 2♀, Yungas, Pte. Mururata, 1200–1600 m a.s.l., 24/ 26-XII-1984, L.E. Peña (USNM); Santa Cruz: 1♀, Pto. Camacho (S), 20-XII-[19]84, L.E. Peña (USNM). AR- GENTINA: Salta: 1♂, Orán, Abra Grande, 10-I/1- III-[19]67, R. Golbach (IFML); 1♀, Orán, 5 R.N., 12- VII-1971, Porter-Stange (IFML); 1♀, 15 km S Orán, 13-VI-[19]95, Flores &amp; Roig (IADIZA); Catamarca: 4♂, 2♀, El Arenal, 29°4 ′ 48.95 ″ S, 65°30 ′ 51.9 ″ W, 1106 m a.s.l., XII-[20]03, S. Roig (IADIZA). Paraguay: 1♂, Capiata, XI-1993 (USNM).</p></div>	https://treatment.plazi.org/id/6F64DB1B5E0AFFE5FE89FEFDFB29F9C7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pablo M. Dellapé;María C. Melo;Thomas J. Henry	Pablo M. Dellapé, María C. Melo, Thomas J. Henry (2016): A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species. Zoological Journal of the Linnean Society 177 (177): 29-134, DOI: 10.1111/zoj.12362
6F64DB1B5E0AFFFFFC18F974FA57FC40.text	6F64DB1B5E0AFFFFFC18F974FA57FC40.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heraeus plebejus Stal 1874	<div><p>HERAEUS PLEBEJUS STÅL, 1874</p><p>(FIGS 36 H, 39E, 41A–D, 42)</p><p>Heraeus plebejus Stål, 1874: 147; Uhler, 1886: 15; Van Duzee, 1894: 174; Lethierry &amp; Severin, 1894: 191; Torre Bueno, 1908: 231; Banks, 1910: 65; Torre Bueno, 1910: 30; Barber, 1914b: 514; Osborn &amp; Drake, 1915a: 506; Weiss, 1916: 10; Van Duzee, 1916: 21; Van Duzee, 1917: 179; Hussey, 1922: 21; Barber, 1923: 725; Torre Bueno, 1924: 50; Torre Bueno, 1925: 70; Blatchley, 1926: 390; Torre Bueno, 1930: 101; Glick, 1939: 23, 134; Froeschner, 1944: 643, 662; Torre Bueno, 1946: 69; Slater, 1952: 532, Barber, 1954a: 4; Ashlock, 1957: 422; Sweet &amp; Slater, 1961: 339; Frost, 1964: 135; Slater, 1964: 1083–1084; Schaefer, 1972: 812; Slater, 1974: 158; Scudder, 1977: 34; Harrington, 1980: 109; Ashlock &amp; Slater, 1988: 228; Slater &amp; Baranowski, 1990: 130–132; Slater &amp; O’Donnell, 1995: 147; Lago &amp; Testa, 2000: 192; Slater &amp; Brailovsky, 2000: 332; Baranowski &amp; Slater, 2005: 142.</p><p>Diagnosis</p><p>Apex of basiflagellomere and distiflagellomere darker, the latter without a pale band. Pedicel with abundant erect setae, equal to or longer than diameter of segment. Labium extending to mesocoxae. Membrane without an apical corial spot. Metafemur with a subapical dark band. Pygophore apex pointed dorsally in lateral view.</p><p>Heraeus plebejus, together with H. pulchellus and H. pacificus, are the only species in the group that lack a pale band on distiflagellomere (although in some specimens of H. pulchellus a narrow pale band is apparent). Heraeus plebejus and H. pulchellus have pale brown to brown antennae, with the apex of basiflagellomere and distiflagellomere darker, and the ocelli are placed at the level of an imaginary line passing the posteri- or border of the eyes, whereas in H. pacificus the antenna is uniformly coloured dark brown and the ocelli are placed posterior of an imaginary line passing the posterior border of the eyes. Heraeus plebejus can be distinguished from H. pulchellus based mainly on body size ( H. plebejus over 5.25 mm long and H. pulchellus less than 5 mm long) and characters of the male genitalia.</p><p>Redescription (Fig. 36 H)</p><p>Head: Coriaceous. Eyes not surpassing the dorsal margin of head in lateral view. Ocelli at level of an imaginary line passing the posterior border of eyes. Labium extending to mesocoxae. Antennae pale brown to brown, with the apex of basiflagellomere and distiflagellomere darker. Distiflagellomere without a pale band. Pedicel with abundant erect setae, equal to or longer than diameter of segment.</p><p>Thorax: Posterior pronotal lobe with pale spots and with a pale spot on humeral angles. Anterior pronotal lobe with long erect setae. Scutellum with erect setae. Evaporative area extended. Inner corial spot diffuse. Costal margin pale on proximal three-quarters, with a subapical corial spot. Membrane brown, with veins paler. Legs: Profemur dark, at most with apices paler and long erect setae; metafemur with a subapical dark band (Fig. 39 E). Protibiae and mesofemur without spines.</p><p>Abdomen: Brown, with abundant short recumbent setae. Male genitalia: Pygophore (Fig. 41 A, B) rounded, apex with a protuberance pointed dorsally; anterior margin of dorsal aperture slightly rounded, inner projections more pronounced posteriorly. Parameres: Figure 41 (C, D). Aedeagus: conjunctiva with some minute spines laterally, lobes of vesica slightly sclerotized, with a few minute spines distally; processus gonopori long and slender.</p><p>Variability observed</p><p>The specimens from Tamaulipas and Nueva León (Mexico) differ from the other specimens examined by the short pale band on the basal region of distiflagellomere.</p><p>Distribution</p><p>Canada, Mexico, and the USA; the Bahamas, South Bimini Island, Haiti, Belize, and Honduras (NEW RECORDS) (Fig. 42).</p><p>This species has a broad distribution in North and Central America, probably originating from an ancestral stock that reached the West Indies and Galapagos Islands where it differentiated into other species: H. pacificus in the Galapagos Islands and H. pulchellus and H. concolor in Cuba, the Cayman Islands, and the Dominican Republic.</p><p>Type material</p><p>The type series is deposited in SMNH and is from Texas and New Jersey (USA); Scudder (1977) designated a female lectotype and paralectotypes.</p><p>Material studied</p><p>BELIZE: 3♂, 2♀, Orange Walk Dist., Chan Chich Lodge, 14-IV-[20]04, BLT, R.M. &amp; H.V. Baranowski (USNM); 4♂, 2♀, 30-X-[20]09, BLT, R.M. &amp; H.V. Baranowski (USNM); 1♂, 1♀, 1-VI-[20]09, E. Flota (MLP); 1♂, 15-X-[20]03, E. Flota (USNM); 3♀, 12-IV- [20]04, E. Flota (USNM); 2♂, 4♀, 13-IV-[20]04, E. Flota (USNM); 1♂, 5-VII-[20]09, E. Flota (USNM); 1♂, 21- IV-[20]04, BLT, E. Flota (USNM); 1♂, 1♀, 4-V- [20]04, BLT, E. Flota (USNM); 16♂, 20♀, 1-VI- [20]09, BLT, E. Flota (USNM); 1♀, 18-X-[20]09, BLT, E. Flota (USNM); 1♂, 1♀, 23-X-[20]09, BLT, E. Flota (USNM); 1♂, 24-X-[20]09, BLT, E. Flota (USNM); 2♂, 3♀, 25-X-[20]09, BLT, E. Flota (USNM); 1♂, 27-X- [20]09, BLT, E. Flota (USNM); 2♂, 1♀, 28-X-[20]09, BLT, E. Flota (USNM); 2♂, 2♀, 29-X-[20]09, BLT, E. Flota (USNM); 3♂, 1♀, 31-X-[20]09, BLT, E. Flota (USNM); 2♂, 3♀, 1-XI-[20]09, BLT, E. Flota (USNM); 2♂, 2-XI-[20]09, BLT, E. Flota (USNM); 1♀, 13-XI- [20]04, BL trap, C. Sanabria (USNM); 3♂, 2-I-[20]05, BL trap, C. Sanabria (USNM); 1♀, 27-I- [20]05, BL trap, C. Sanabria (USNM); 2♂, 30-IV- [20]05, BL trap, C. Sanabria (USNM); 2♂, 7-XII- [20]04, BL trap, C. Sanabria (USNM); 1♂, 4♀, 2-VII-[20]04, BL trap, C. Sanabria (USNM); 1♂, Cayo Dist., Benque del Carmen, Riverside N., Mopan River Resort, 25-V-[20]07, blacklight trap, P. Picon (USNM); 1♂, 30-V-[20]07, blacklight trap, P. Picon (USNM); 1♂, 1♀, Mopan R. Resort, Riverside N., Benque Viejo del Carmen, Cayo, BLT, 27-VI-[20]06, A. Trujillo (USNM); 1♂, 2♀, Cayo District, Benque Viejo, Riverside N. Mopan R. Resort, 22-IX-[20]09, BLT, M. Chrysler (USNM); 2♂, 29-IX-[20]09, BLT, M. Chrysler (USNM); 1♂, 4-X-[20]09, BLT, M. Chrysler (USNM); 1♂, 7-X-[20]09, BLT, M. Chrysler (USNM); 1♀, 15-X- [20]09, BLT, M. Chrysler (USNM); 1♀, 6-XI-[20]09, BLT, M. Chrysler (USNM); 2♂, Orange Walk District, Gallon Jug, 7-IV-[20]04, BLT, B. Miller (USNM); 1♂, 13-IV-[20]05, BLT, B. Miller (USNM); 1♀, 11-X- [20]05, BLT, B. Miller (USNM); 1♀, Cayo District, Central farms, mile 67, Western Highway, 29-VI- [19]68, blacklight trap, W.L. Hasse (USNM); 2♂, O.[range] W.[alk] Dist., Rio Bravo, Area hdqtrs., 11- VII-1996, UV trap, P. Shuey (USNM); 2♂, 13-VII- 1996, UV trap, P. Shuey (USNM); 5♂, 8-VII-1996, UV trap, P. Shuey (USNM); 3♂, Rio Bravo Cons. Area, Mahogany Trail, 10-VII-1996, Uv &amp; Hg vapor light, C.W. &amp; L.B. O’Brien (USNM).</p><p>HONDURAS: 1♂, intercepted at Miami, 21-V-[19]54, A.S. Mills (USNM).</p><p>MEXICO: Coahuila: 1♀, 10 mi. S Allende, 11-VIII- 1958, H.F. Howden (CNC); Durango: 1♂, 23 mi. W Durango, 7500 ft, 13-VI-1964, L.A. Kelton (CNC); Oaxaca: 1♀, Rio Jatepec, Isthmus of Tehuantepec, blacklight trap, 21-V-[20]09, Blanton et al. (USNM); Tamaulipas: 1♀, Tamps, Galeana Canyon, 19-VI- 1975, L.E. Watrous (OSUC); 1♀, Antiguo Morelos 5 mi. W, 1400 ft, 20-VII-1954, J.G. Chillcott (CNC); 1♂, CD, Mante, 27-VII-[19]60, at light, H. Howden (CNC); San Luis Potosí: 1♀, Tamazunchale, 19-IV-1963, W.J. Gertsch &amp; W. Ivie (AMNH). Veracruz: 1♂, 5 mi. S Veracruz, 30-VI-1975, at light, C.A., W.E., B.W. Triplehorn (OSUC).</p><p>USA: Alabama: 1♀, C.F. Parker (USNM); Arizona: one without abdomen, Phoenix, 4-VI-1928, A.A. Nichol (USNM); 2♀, one without abdomen, #90 (USNM); 1♂, 2♀, Salt Riv. Valley, 16-II-1933, rubbish nr. cotton field, C.J. King (USNM); 1♀, Thatcher, 20-VI-1951, E.J. Taylor (USNM); Arkansas: 1♀, Hope, 11-VI- 1954, J.W. Green (CAS); 1♂, 1-VII-1931, light trap, L. Knobel (USNM); 1♀, 14-VI-[19]25, light trap, L. Knobel (USNM); 1♀, 5-X-[19]25, light trap, L. Knobel (USNM); 1♂, 23-VI-[19]25 (USNM); one without abdomen, 25-V-[19]25, light trap, L. Knobel (USNM); 1♂, Queen Wilhelmina State Park, 4-IV- 1987, G.F. &amp; J.F. Hevel (USNM); Connecticut: 1♂, Storrs, 29-VII-1954, F.B. Levi (USNM); 1♀, 26-VIII- 1954, J.A. Slater (USNM); District of Columbia: 1♀, Washington D.C., 15-V-[18]92 (CUIC); 1♂, 8-VI- [19]03, Heidemann (CUIC); 1♂, V-[19]08, Van Duzee (CAS); 1♂, 2♀, Washington, 13-III-[19]10, B.M. (USNM); 5♂, 1♀, 22-II-[19]06, D.H. Clemons (USNM); 1♂, 4-VII-[19]14, H.H.T. Jackson (USNM); 1♂, 1♀, 24-VII-[19]07, W.L. McAtee (USNM); 1♂, 15-VII- [19]14, W.L. McAtee (USNM); 1♀, 6-V-[19]13, at light, W.L. McAtee (USNM); 1♂, J.D. Hood (USNM); 1♀, 6-VII-[19]07, J.D. Hood (USNM); 1♀, DC, 2-XII- [19]88 (USNM); 1♂, DC, 3-XI-[19]38, C.V. Riley (USNM); 1♀, Woodbridge, 4-VII-1913, A. Wetmore (USNM); 1♂, Receiving Reservoir, 22-II-1913, W.D. Appel (USNM); 1♂, 13-IV-1913, W.D. Appel (USNM); Florida: 3♂, 2♀, Lake Alfred, 24-VI- [19]63, on Eremochloa ophiuroides, C.D. Risk (USNM); 1♂, 1♀, Volusia Co., 4-VIII-[19]56, at light, H.A. Denmark (USNM); 2♀, 6-VIII-[19]56, at light, H.A. Denmark (USNM); 1♂, Cleveland, IV-[19]25, D.M. DeLong (PSUC); 2♂, 1♀, Archbold Biological Station, Highlands Co., 17-III-[19]62, S.W. Frost (PSUC); 1♂, 1♀, one without abdomen, 24-IV- [19]67, D.E. Bright (CNC); 1♂, 3♀, 23-IV-[19]67, D.E. Bright (CNC); 1♀, 23-IV-[19]67, D.E. Bright (PSUC); 1♀, 7-IV-[19]62, S.W. Frost (PSUC); 1♀, IX- 1975, W. Rosenberg (USNM); 1♀, 13 km S lake Placid, 4-V-1986, W. Steiner (USNM); 1♀, Lake Placid, Archbold Biological Sta., 4-V-1969 (CNC); 1♂, 9-V- 1964, R.W. Hodges (USNM); 1♂, 1♀, 18-V-1964, R.W. Hodges (USNM); 1♂, Highlands Co., 2-V- 1983, UV light trap, M. Deyrup (USNM); 1♂, 1♀, 6-IV-1983, UV light trap, M. Deyrup (USNM); 1♀, Putnam Co., 2 mi. NW Orange Springs, 1/3-VIII- [19]75, D. Bowman (USNM); 3♂, 1♀, Washington Co., 5 mi. E Chipley, 31-VIII-[19]60, blacklight trap, W.C. Rhodes (USNM); 1♀, Hendry Co., LaBelle, 10/ 11-III-1974, UV in oak groove, J. Reynolds (AMNH); 1♀, Clewiston, 30-IV-[19]61, L.A. Kelton (CNC); one without abdomen, 6-II-[19]41, M. Wrigth (USNM); 1♀, Gainesville, 24-IV-1952, O. Peck (CNC); 1♀, Alachua Co., Gainesville, 29°36.916 ′ N, 82°23.102 ′ W, 12-IV-2007, at light, C.L. Staines (USNM); 2♂, 3♀, Doyle Connor Bldg, 2/4-V-[19]75, BLT, F.W. Mead (USNM); 3♂, 1♀, 23/25-V-[19]75, BLT, F.W. Mead (USNM); 1♂, 1♀, 2/4-V-[19]75, BLT, F.W. Mead (USNM); 2♂, 12-V-[19]75, BLT, F.W. Mead (USNM); 1♀, Punta Gorda, 11-IV-[19]52, J.R. Vockeroth (CNC); 3♂, 3♀, Dade Co., Homestead, 11-IV-[19]69, blacklight trap, R.M. Baranowski (USNM); 2♀, Homestead, 1-IV-1962, at light, S.S. Waley (CNC); 2♂, 3♀, Dade Co., Agr. Res. &amp; Ed. Ctr., Homestead, 12-IV-[19]69, blacklight trap, R.M. Baranowski (USNM); 5♂, 4♀, Dade Co., Grossman Hammock Sta. Pk., 24-V-[19]72, blacklight trap, R.M. Baranowski (USNM); 2♀, 2-V-[19]72, blacklight trap, R.M. Baranowski (USNM); 1♂, Elfers, 14-IV-1952, O. Peck (CNC); 3♂, 51–55, Stuart, 25- VI-1951, O. Bryant (CAS); 3♀, Dunedin, 4-II- [19]26, W.S. Blatchley (CAS); 1♀, Monroe Co., Everglades Natl Park, Flamingo Praire, 4-XII- 1970, sea level, P.H. &amp; M. Arnaud (CAS); 7♂, 19♀, 28-III-[19]70, R.M. Baranowski (USNM); 33♂, 36♀, 4-V-[19]70, R.M. Baranowski (USNM); 1♂, St Augustine, 7-III-[19]40, S. VanDyke (CAS); 1♂, Sanford, 5-VI-[20]08 (CAS); 1♂, 5-V-[19]28, E.D. Ball (USNM); 2♂, 15-V-[19]26, E.D. Ball. (USNM); 1♂, Palmdale, Fisheating Cr., 10-V-1964, R.W. Hodges (USNM); 3♂, 2♀, Boynton Beach, 12-VI-1968, collected at blacklight, F.E. Wood &amp; J.A. Davidson (USNM); 4♂, 9♀, Marion Co., Sharpes Ferry Fld. Sta., 30-IV-[19]75, blacklight trap, N. Holler (USNM); 2♂, 5♀, 15-V-[19]75, blacklight trap, N. Holler (USNM); 3♂, 1♀, 30-X-[19]75, blacklight trap, N. Holler (USNM); 1♂, Marion Co., Lake Eaton, 8-IV- [19]75, blacklight trap, P.C. Drummond (USNM); 1♀, 22-V-[18]75, blacklight trap, P.C. Drummond (USNM); 2♂, 1♀, Marion Co., 14 mi. SW Ocala, Ross Praire, 12-VI-[19]75, blacklight trap, P.C. Drummond (USNM); 3♀, Taylor Co., Williams Landing, 21/30-VI-1967, R. Smith (USNM); 1♂, Miami, 7-VII-1920 (USNM); 1♀, Tampa, 5-II-1946, R.C. Froeschner (USNM); 1♀, Paradise Key, 3-III-[19]19, H. Barber (USNM); 2♀, 28-II-[19]19, H. Barber (USNM); 1♂, VI-1919, C.A. Mosier (USNM); 1♂, Ft. Lauderdale, 24-IV- 1928, D.M. Bates (USNM); 1♀, 26-III-1928, D.M. Bates (USNM); 1♀, Ducal Co. (USNM); 3♂, 2♀, Baker Co., Olustee, 30-V-[19]63, blacklight trap, E.P. Merkel (USNM); 6♂, 3♀, 12-VII-[19]63, blacklight trap, E.P. Merkel (USNM); 1♂, 5♀, 2/4-VII-[19]66, blacklight trap, E.P. Merkel (USNM); 1♀, Alva, 3-IV- 1919, A.A. Hardisty (USNM); 2♂, 1♀, Edgewater, 28-II-[19]39, C.A. Frost (USNM); 1♀, Royal Palm Pk., 20-VII-1948, E.L. Todd (USNM); 1♀, 22-VII-1948, E.L. Todd (USNM); 1♀, Old Town, 11-VII-1939, Oman (USNM); 1♂, Orange Co., 2-VII-[19]29, Florida Fruit Fly Surv., E.T. Bates (USNM); 1♂, 2♀, Wakulla Co., Surf Panacea, 5/15-VII-1967, C. Hilfiker (USNM); Georgia: 1♀, P.R. Uhler (USNM); 1♂, Atlanta, 12- VIII-1973 (CUIC); 1♂, Pine Mountain, Rabun Co., 1400 ft, 14-V-1957, J.R. Vockeroth (CNC); 1♂, Grady Co., Beachton, 18/24-V-1967, E.V. Komarek Sr. (USNM); 2♀, 19/26-VI-1967, E.V. Komarek Sr. (USNM); 1♂, Thomasville, 13-III-[19]03, H.G. Barber (USNM); 1♂, Demorest, 14-VIII-[19]39, Valentine (USNM); 4♂, 2♀, Peach Co., 18-V-[19]43, light trap, Turner (USNM); Illinois: 1♂, P.R. Uhler (USNM); Indiana: 1♂, Oxford, VIII-[19]13, Mrs. W.L. McAtee (USNM); 1♀, Marion Co., W.S.B., 6-X-[19]21, H.G. Barber (USNM); 1♂, 20-VI-[19]20, H.G. Barber (USNM); 1♂, Jennings Co., 15-X-1912, H.O. Deay (USNM); Kansas: 1♂, Orange, E.P. Van Duzee (CAS); 1♀, Bourbon Co., 1 mi. N Redfield, 13-VII-1966, J. &amp; W. Ivie (AMNH); 1♂, 1♀, Douglas Co., Breidenthal Reserve, 15 mi. SE Lawrence, 29-VI-1979, R.J. McGinley (USNM); 1♂, Independence, 25-VIII- 1950, R.I. Sailer (USNM); 1♀, Topeka, 8-VII (USNM); Kentucky: 3♀, Louisville, 23-VII-[19]40, D.R. Young (USNM); Louisiana: 1♀, 30-XII-[19]31, J.H. Moreland (USNM); Maine: 4♀, Augusta, 28-III-[19]62, in pkg. Chicory, A.E. Brower (USNM); Massachusetts: 1♀, Chicopee, 29-X-[18]96, C.F. Baker (USNM); 1♀, Holliston, 27-IX, N. Banks (USNM); 1♀, Framingham, 1-XII-1932, C.A. Frost (USNM); 1♂, 21-VI-[19]49, C.A. Frost (USNM); 1♂, 19-X-[19]35, C.A. Frost (USNM); 1♀, Natick, 27-IV-[19]35, C.A. Frost (USNM); 1♀, Sherbon, thru sifting, 7-X-[19]33, C.A. Frost (USNM); Maryland: 1♂, P.R. Uhler (USNM); 1♂, Forest Gln., 9-X-[19]25, electric light, Heidemann (USNM); 1♀, KabJohn Br., April 1, Heidemann (CUIC); 1♀, Montgomery Co., Plummers Island, 38°58 ′ N, 77°10 ′ W, 27-VI/11-VII-2005, Malaise trap, lower trap, D. Smith &amp; J. Brown (USNM); 16♂, 26♀, Plummers, 30-III-1913, in drift, W.L. McAtee (USNM); 1♂, 18-VI-[19]14, on mullen, W.L. McAtee (USNM); 2♂, 1♀, 12-IV-[19]14, R.C. Shannon (USNM); 1♀, 7-IX-[19]12 (USNM); 3♀, 8-II-[19]05, D.H. Clemons (USNM); 2♂, 28-X-[19]13, R.C. Shannon (USNM); 1♀, 13-VIII-[19]14, at light, R.C. Shannon (USNM); 1♂, at light, R.C. Shannon 8-VIII-[19]14 (USNM); 1♀, 4-VII-[19]14, W.L. McAtee (USNM); 1♂, 23-IX- [19]15, W.L. McAtee (USNM); 1♂, 28-IX-[19]14, W.L. McAtee (USNM); 1♂, 25-I-[19]14, W.L. McAtee (USNM); 1♀, 18-I-[19]14, W.L. McAtee (USNM); 1♂, 30-VIII-[19]14, W.L. McAtee (USNM); 1♂, 24-V- [19]14, W.L. McAtee (USNM); 1♀, 1-V-[19]14, W.L. McAtee (USNM); 1♂, 7-VI-[19]14, W.L. McAtee (USNM); 1♀, 4-X-[19]14, W.L. McAtee (USNM); 2♂, 5-X-[19]13, W.L. McAtee (USNM); 1♂, 16-VIII- [19]12, W.L. McAtee (USNM); 1♂, 13-X-[19]07, W.L. McAtee (USNM); 1♀, 17-V-[19]08, W.L. McAtee (USNM); 1♀, 11-IV-[19]11, H.G. Barber (USNM); 1♂, 29-V-[19]09, Brooklyn Museum (USNM); 4♂, 5♀, Maryland near Plummers Island, 14-II-1915, W.L. McAtee (USNM); 1♂, 28-III-1915, W.L. McAtee (USNM); 1♂, Prince George Co., Patuxent Res. Refuge, Central tract nr Jct. River Road &amp; Beech Tree Road, 39°2 ′ 57.06 ″ N, 76°47 ′ 54.38 ″ W, 5/8-V- 2006, Malaise trap, S.J. Scheffer (USNM); 1♂, 1♀, Pr. Geo. Co., Cheverly, 38°56 ′ N, 76°55 ′ W, 10-VIII- 1993, W.E. Steiner &amp; J.M. Swearingen (USNM); 1♂, A. Arun. Co., 6 km S Edgewater (SERC), 38°53 ′ N, 76°33 ′ W, 15-VI-1976, J.H. Falk (USNM); 1♂, 3 mi. S Beltsville, 23-VI-1969, Druckenbrod (USNM); 1♂, Beltsville, 17-V-[19]14, W.L. McAtee (USNM); 1♂, College Park, VII-1941, at light, L.P. Ditman (USNM); 1♀, Breton Bay, Potomac Riv., R. Budlong, 13- VII[19]23, H.S. Barber (USNM); 1♂, 1♀, Wicomico Co., Bivalve, 8-VI-1990, M.J. Rothschild (USNM); 2♂, 4♀, 18-VII-1990, M.J. Rothschild (USNM); 1♀, 11- IX-1990, M.J. Rothschild (USNM); 1♀, 13-IX-1990, M.J. Rothschild (USNM); 1♀, 17-VII-1990, M.J. Rothschild (USNM); 1♀, Cabin John, 16-X- [19]61, R.I. Sailer (USNM); 1♂, 1♀, 8-III-[19]08, H.O. Marsh (USNM); 1♂, C.H. Popenoe (USNM); 1♂, Minnie L., nr Cabin John, 5-VI-[19]31, E.E. Myers (USNM); 1♂, Annapolis, 8-VII-[19]32, mosquito trap, F.C. Bishop (USNM); 1♂, Chestertown, 8-IX-[19]32, mosquito trap, F.C. Bishop (USNM); 1♂, Princess Anne, 28-VII-[19]32, mosquito trap, F.C. Bishop (USNM); 3♂, 1♀, one nymph, Glen Echo, summer 1922, J.C. Bridwell (USNM); 1♀, 10-VI-[19]23, J.R. Malloch (USNM); 1♂, Plum Point, 20-VI-[19]14, W.L. McAtee (USNM); 1♂, 1♀, nr Bladesburg, 23- III-1913, W.L. McAtee (USNM); 1♂, Paint Branch, 2 mi. W of Beltsville, 31-X-[19]31, H.G. Barber (USNM); 1♂, Jacksons Id., 22-V-[19]13, Shannon &amp; Barber (USNM); 1♂, Contee, 7-V-[19]14, A. Wetmore (USNM); 1♂, Hagerstown, 2-IX-[19]14, H.L. Parker (USNM); 1♂, 6-V-[19]15, H.L. Parker (USNM); 1♂, 12-VII-[19]15, H.L. Parker (USNM); 1♀, Ch. Chase Lk., 24-X-[19]16, B.P. Currie (USNM); Mississippi: 1♀, Gulfport, 10/30-VI-[19]38, R.E. Blackwelder (USNM); Missouri: 1♀, McDonald Co., Elk River at Hwy. 59, 3 mi. N Noel, 8-VIII-1967, at light combined blacklight and Coleman lantern, H.B. Leech (CAS); 1♂, Columbia, 27-VIII-1968, Malaise, F.D. Parker (USNM); 1♂, 1-VII-1941, R. Froeschner (USNM); 1♂, Lanagan, 18-IX-1942, R. Froeschner (USNM); 1♂, Howard Bend, 6-IX-1938, R. Froeschner (USNM); 1♂, Sarcoxie, 25-VII-1943, R.C. Froeschner (USNM); 1♂, Kimaswick, 17-X-1943, R.C. Froeschner (USNM); 1♂, Newmelle, 28-XI-1946, R.C. Froeschner (USNM); 1♂, 2 km W St Louis, 23-V-[19]04, W.V. Warner (USNM); 1♂, Webster Groves, 21-VII- [19]20, at light, V.R. Stones (USNM); New Hampshire: 1♀, Hampton, S. Alberto Shaw, 3-IV-1932, A.E. Brower (USNM); New Jersey: 1♂, Parshley (CAS); 1♀, Arlington, Parshley (CAS); 1♂, 9-III- 1902, Joutell (USNM; 1♂, Orange, elect. light, Chittenden (USNM); 2♂, Roselle Pk., V-[19]25, collected by sifting, H.G. Barber (USNM); 1♂, 30-II- [19]07, collected by sifting, H.G. Barber (USNM); 1♂, 1♀, 16-II-[19]07, collected by sifting, H.G. Barber (USNM); 1♂, 28-II-[19]09, H.G. Barber (USNM); 2♂, 1♀, Madison, 29-VII-[18]97, P.R. Uhler (USNM); 1♂, 15-X-[18]97, P.R. Uhler (USNM); 1♀, 15-VIII- [18]98, P.R. Uhler (USNM); 1♂, 10-IV-[18]98, P.R. Uhler (USNM); 1♀, Cape May Co., Cold Springs, 12-IV-[19]11, under dead leaves, H.G. Barber (USNM); 1♂, 1♀, 12-IV-[19]11, H.G. Barber, under board (USNM); 1♀, Cape May Co., nr. Town Bank, 10-IV- [19]11, under board, H.G. Barber (USNM); 1♂, Lakehurst, collected by sifting, H.G. Barber (USNM); 1♂, 3♀, IX-[19]26, H.G. Barber (USNM); 1♀, 1-IV- 1916, beneath board, H.G. Barber (USNM); New York: 1♂, Pinelawn, 4-V-[19]11, G.P. Engelhardt (CUIC); 1♂, Hamburg, 31-VI-[19]02, Van Duzee (CAS); 1♂, 12-IV-[19]01, Van Duzee (CAS); 1♀, White Plains, 28-V-[19]09, Parshley (CAS); 1♀, 4-IV-[19]09, Parshley (CAS); 1♀, 11-IV-[19]09, Parshley (CAS); 2♀, 2-IV- [19]25, H.G. Barber (USNM); 1♀, 29-IX-[19]25, H.G. Barber (USNM); 1♀, Rochester, R.M. Moore (CAS); 1♀, Ithaca, 10-XI-[19]17, E.C. Van Dyke (CAS); 1♀, East Aurora, 30-X-[19]21, M.C. Van Duzee (CAS); 1♀, Batavia, 26-VIII-1914, H.H. Knight (USNM); 1♂, Staten Island, 6-XI-[19]09, salt meadow, collected by sifting on edge, H.G. Barber (USNM); 1♀, 2–22, collected by sifting, H.G. Barber (USNM); 2♂, 3♀, Long Island, Cold Spring Harbor, 25-VIII-1900, H.G. Barber (USNM); 1♀, 27-VIII-1900, H.G. Barber (USNM); 1♂, L.[ong] I.[island], near Brighton, 14-IX-[19]09, collected by sifting, H.G. Barber (USNM); 1♀, L.[ong] I.[island], Flatbush, 14-X-1892, Jlzabriskic (USNM); North Carolina: 1♀, Rocky Mt, Edgecombe Co., 4-VII- 1973, R. Schrammel (AMNH); 1♀, Black Mts, 13- III-1912, Beutenmuller (CNC); 2♀, X-1966, J. &amp; W. Irvie (AMNH); 1♀, Wilmington, 7-IV-[19]14, H.G. Barber (USNM); 1♀, Asheville, 12-V-[19]44, W.E. Hoffman (USNM); South Carolina: 1♂, Charleston, 26-III-[19]32, Adams &amp; Gay (USNM); Ohio: 1♂, Ada, 6–41, C.R. Neiswander (OSUC); 1♀, McGuffey, 7–41, C.R. Neiswander (OSUC); 1♀, W. Unity., 6–41, C.R. Neiswander (OSUC); 1♀, Hocking Co., 5–20, D.J.</p><p>&amp; J.N. Knull (OSUC); 1♀, Scioto Co., 10-VI-[19]44, D.J. &amp; J.N. Knull (OSUC); 7♂, 10♀, Columbus, 7-VII- 1930, C.H. Hicks (USNM); 1♀, Athens, City Park, 3-II-[19]50, P.J. Spangler (USNM); Pennsylvania: 3♂, Philadelphia, VIII-[19]37, J.O. Pepper (PSUC); 2♀, Martinsburg, 12-VII-[19]48, S.W. Frost (PSUC); 1♂, 13-VII-[19]48, S.W. Frost (PSUC); 1♂, 6-VII-[19]48, S.W. Frost (PSUC); one without abdomen, 6-VII- [19]48, S.W. Frost (PSUC); 7♂, 1♀, Phila., Chestnut Hill, 25-III-[19]39, E. Endy (USNM); 1♀, Phila.[delphia], 24-VII-1928, J.C. Lutz (USNM); 1♀, Lancaster, 6-I-[19]48, B.F. Coon (PSUC); 2♂, 1♀, 14- I-[19]48, B.F. Coon (PSUC); 1♀, Kennett sq., 26- VIII-[19]47, C.A. Thomas (PSUC); 1♀, State College Center Co., 29-VII-[19]83, Verda Haas (PSUC); 1♂, 1♀, Bucks Co., NE Jamison, Horseshoe Bend, Neshaminy Creek, I/II-1954, W. Ivie (AMNH); 1♂, Bucks Co., Neshaminy Creek, E of Jamison, 3-VII- 1966, J. &amp; W. Ivie (AMNH); 1♀, Bucks Co., Neshaminy Creek, 2 mi. E of Jamison, VI-29– 1966, J. &amp; W. Ivie (AMNH); Tennessee: 1♀, Gr. Smoky Mt., Nat. Pk., 2-VII-1957, J.R. Vockeroth (CNC); 1♂, P.R. Uhler (USNM); 1♀, Clarksville, 5-III-[19]09, hibernating Tob, field, C. Morgan (USNM); 1♂, 1♀, hiberbating in leaves, C. Morgan (USNM); 1♂, 15- IV-[19]09, C. Morgan (USNM); 1♂, 3♀, Hamilton Co., 15-VII-[19]42, collected at light, peach orchard, Turner (USNM); 6♂, 6♀, 24-VI-[19]43, light trap edge peach orchard (USNM); 1♂, 3♀, 23-VIII-[19]43 (USNM); one without abdomen, 12-VIII-[19]42 (USNM); 1♂, Knoxville, 15-V-[19]17, sweep from sedges, Geo. G. Ainslie (USNM); Texas, 1♀, P.R. Uhler (USNM); 1♂, 1♀, C.V. Riley (USNM); 2♂, 2♀, P.R. Uhler (USNM); 1♂, Belfrage, C.V. Riley (USNM); 1♂, 2♀, Sommerset, 27-XII-1942, E.S. Ross (CAS); 1♀, Austin, 20-VI-[19]30, G. Linsley (CAS); 1♂, San Antonio, 5-X- [19]47, E.S. Ross (CAS); 2♀, Brownsville, X-[19]42, E.S. Ross (CAS); 1♂, at light (CAS); 6♂, 6♀, 16- IX-[19]42, T.M. Burnst, (CAS); 5♀, L.R. Joyce (USNM); 1♀, on weeds deputy, 11-V-[19]38 (USNM); 1♀, 23- I-[19]43, from pink bollworm, I. Shiller (USNM); 1♀, Hidalgo Co., Santa Ana Refuge, 15/16-II-1971, A. &amp; M.E. Blanchard (USNM); 1♀, Hidalgo Co., 29-VII- [19]47, at light, G.B. Vogt (USNM); 1♂, 12/20-IX- [19]47, at light, G.B. Vogt (USNM); 3♀, Plano, VII- [19]07, taken at trap light, E.S. Tucker (USNM); 2♂, one without abdomen, July at night, E.S. Tucker (USNM); 1♂, Houston, Harris Co., 1/4-VI-1967, A. &amp; M.E. Blanchard (USNM); 1♀, Sinton, Welder Wildlife Foundation, 3-V-1967, A. &amp; M.E. Blanchard (USNM); 1♂, 1♀, Victoria, Sept[ember] (USNM); 1♀, Dallas, 25-IV-[19]07, F.C. Pratt (USNM); 2♂, 1♀, Kerrville, 31-V-[19]06, at light, F.C. Pratt (USNM); 1♀, College Station, 29-V-1933, J.R. Reinhard (USNM); 1♀, Tyler, 15-II-[19]39 (USNM); Virginia: 1♂, 1♀, Fairfax Co., Turkey Run (West), 38°57.9 ′ N- 77°09.4 ′ W, 29-VI/12-VII-2007, Malaise trap, D. Smith (USNM); 2♂, 7♀, 3/17-VII-2008, Malaise trap, D. Smith (USNM); 1♀, 5/19-VI-2008, Malaise trap, D. Smith (USNM); 1♂, 14/28-VI-2007, Malaise trap, D. Smith (USNM); 1♂, 2♀, Fairfax Co., 1 km E Fairfax city, 26/27-VII-2005, blacklight, J. Brown (USNM); 1♂, Louisa Co., 4 mi. S Cuckoo, 1-VI-1985, Malaise trap, J. Kloke &amp; D.R. Smith (USNM); 1♀, Giles Co., Mtn. L. Scenic Area, 3800 ft, 14-VII-1971, A.B. Gurney (USNM); 1♂, Clarke Co., U. Va. Blandy Exp. Farm, 2 mi. S Boyce, 39°5 ′ N, 78°10 ′ W, 19-IX/25-X-1993, Malaise trap, D.R. Smith (USNM); 1♂, Scotts Run, Stubblefield Falls, 23-X-1921, on Pinus virginiana, J.R. Malloch (USNM; 1♀, Falls Ch., 7-VI-[19]14, R.C. Shannon (USNM); 1♂, 29-XII, H. Barber (USNM); 1♀, 21-IX, H.G. Barber (USNM); 1♀, Vienna, 21-V-[19]32, H.G. Barber (USNM); 1♂, 1♀, 7-VI- [19]28, H.G. Barber (USNM); 1♂, 3-VI-[19]28, H.G. Barber (USNM); 1♂, 5-VI-[19]28, H.G. Barber (USNM); 1♀, VIII-[19]18, H.G. Barber (USNM); 2♀, 31-VIII-[19]32, J.C. Bridwell (USNM); 2♂, 22-X- [19]32, at light, J.C. Bridwell (USNM); 2♀, 1-IX- 1932, at light, J.C. Bridwell (USNM); 1♀, 22-IX- 1932, at light, J.C. Bridwell (USNM); one without abdomen, 31-V-[19]32, at light, J.C. Bridwell (USNM); 2♂, 1♀, Norfolk, 3-VII-[19]63, B/L Ser. x22 (USNM); 1♂, Four Mile Run, 23-V-[19]14, A. Wetmore (USNM).</p></div>	https://treatment.plazi.org/id/6F64DB1B5E0AFFFFFC18F974FA57FC40	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pablo M. Dellapé;María C. Melo;Thomas J. Henry	Pablo M. Dellapé, María C. Melo, Thomas J. Henry (2016): A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species. Zoological Journal of the Linnean Society 177 (177): 29-134, DOI: 10.1111/zoj.12362
6F64DB1B5E10FFFEFC39FBE5FC60FE67.text	6F64DB1B5E10FFFEFC39FBE5FC60FE67.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heraeus pulchellus Barber 1954	<div><p>HERAEUS PULCHELLUS BARBER, 1954</p><p>(FIGS 36 I, 39F, 41E–H, 42)</p><p>Heraeus pulchellus Barber, 1954b: 342 –343; Slater, 1964: 1084; Harrington, 1980: 109; Slater, 1988: 50, 59; Baranowski &amp; Slater, 1998: 79, 87; Baranowski &amp; Slater, 2005: 143.</p><p>Diagnosis</p><p>Small species, total length less than 5 mm. Labium extending to mesocoxae or metacoxae. Antennae pale brown to brown, with the apex of basiflagellomere and distiflagellomere darker. Distiflagellomere without a pale band. Membrane with an apical pale spot. Metafemur with a subapical dark band. Anterior margin of dorsal aperture of pygophore straight. Pygophore apex pointed dorsally in lateral view. Aedeagus unspined.</p><p>Heraeus pulchellus, together with H. plebejus and H. pacificus, are the only species in the group that lack a pale band on distiflagellomere (although in some specimens of H. pulchellus a band is vaguely apparent). In H. plebejus and H. pulchellus, the antenna is pale brown to brown, with the apex of basiflagellomere and distiflagellomere darker, and the ocelli are placed at the level of an imaginary line passing the posterior border of the eyes, whereas in Heraeus pacificus the antenna is uniformly dark brown and the ocelli are placed posterior of an imaginary line passing the posterior border of the eyes. Heraeus pulchellus can be distinguished from H. plebejus based mainly on body size ( H. plebejus over 5.25 mm long and H. pulchellus less than 5 mm long) and the male genitalia.</p><p>Redescription (Fig. 36 I)</p><p>Head: Eyes not surpassing the dorsal margin of head in lateral view. Ocelli at level of an imaginary line passing the posterior border of eyes. Labium extending to mesocoxae or metacoxae. Antennae pale brown to brown, with the apex of basiflagellomere and distiflagellomere darker. Distiflagellomere without a pale band.</p><p>Thorax: Posterior pronotal lobe with pale spots and a pale spot on each humeral angle. Anterior pronotal lobe with long erect setae. Evaporative area extend- ed. Scutellum with erect setae. Hemelytra: inner corial spot diffuse and with a subapical corial spot; costal margin pale on proximal three-quarters. Membrane brown, with veins paler and an apical pale spot. Legs: Profemur dark, at most with apices paler, metafemur with a subapical dark band (Fig. 39 F). Profemur with long erect setae. Protibiae and mesofemur without spines.</p><p>Abdomen: Brown, with abundant short recumbent setae. Male genitalia: Pygophore (Fig. 41 E, F) rounded, apex with a protuberance pointed dorsally; anterior margin of dorsal aperture rounded, inner projections quadrangular, more pronounced posteriorly. Parameres: Figure 41 (G, H). Aedeagus with minute spines on conjunctiva and sclerotized vesical lobes; processus gonopori long and slender.</p><p>Distribution</p><p>Bahamas, Cayman Island, Cuba, and Saba Island (Fig. 42).</p><p>Type material examined</p><p>Holotype: ♂, CUBA, ‘Capitolio’, Rio Yara, May 15–19 [19]48, Elev. 1150 ft appr., J. Ferrás, type #61922 (USNM).</p><p>Additional material studied</p><p>BAHAMAS: 1♂, Mayaguana Isl., 28-VIII-[19]63,</p><p>blacklight trap, C. Murvosh (AMNH); 1♀, 26-VIII-</p><p>[19]63, blacklight trap, C. Murvosh (USNM).</p><p>CUBA: 1♀, Soledad, Santa Clara Prov., 24/30-V- 1939, C.T. Parsons (AMNH); 1♀, Central Soledad, 1-VII-1932, B.B. Leavitt (AMNH); 1♂, Guantanamo, Imias, 10 m.s.n.m., 16-VII-[19]90, a la luz, L.F. Armas &amp; V. Becker (USNM).</p><p>GRAND CAYMAN ISLAND: 1♂, 1-XI-[19]92, BLT, P. Fitzgerald (USNM); 1♂, South Sound, 23-VIII- 1987, BL trap, P. Fitzgerald (USNM); 1♀, 2-VIII-1987, BL trap, P. Fitzgerald (USNM); 1♀, 30- XI-1987, BL trap, P. Fitzgerald (USNM); 1♀, Grand Cayman, 17-V-[19]92, B.L.T., P. Fitzgerald (USNM); 1♀, 8-V-[19]92, B.L.T., P. Fitzgerald (USNM); 1♂, 1♀, VI-[19]86, R.M. Baranowski (USNM); 1♀, Savannah, 12/13-XI[19]85, R.M. Baranowski (USNM).</p><p>CAYMAN BRAC ISLAND: One without abdomen, The Creek, 5-VI-[19]96, blacklight trap, E.A. Dilbert (USNM); 1♀, 8-V-[19]97, blacklight trap, E.A. Dilbert (USNM).</p></div>	https://treatment.plazi.org/id/6F64DB1B5E10FFFEFC39FBE5FC60FE67	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pablo M. Dellapé;María C. Melo;Thomas J. Henry	Pablo M. Dellapé, María C. Melo, Thomas J. Henry (2016): A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species. Zoological Journal of the Linnean Society 177 (177): 29-134, DOI: 10.1111/zoj.12362
6F64DB1B5E11FFFDFC7CFDD4FE4EFB1E.text	6F64DB1B5E11FFFDFC7CFDD4FE4EFB1E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Baranowskiobius	<div><p>BARANOWSKIOBIUS GEN. NOV.</p><p>Type species: Nabis elegans Walker, 1873a: 144 . HERE DESIGNATED.</p><p>Diagnosis</p><p>Postocular region elongate, longer than interocellar distance. Jugal ridge present. Mesepimeron emergent. Evaporative area extensive. Clavus with three complete rows of punctures and additional incomplete rows between inner and median row. Procoxa with a spine. Protrochanters without spines. Aedeagus unspined, vesica with two lobes weakly sclerotized laterally; ejaculatory ductus wide; processus gonopori broadened towards apex.</p><p>Description</p><p>Relatively large and elongate. Pruinose dorsally.</p><p>Head: Elongate, vertex rounded. Postocular region elongate, longer than interocellar distance, slightly narrowing, forming a neck. Eyes surpassing dorsal margin of head. Jugal ridge present. Buccular juncture V-shaped, near insertion of labium.</p><p>Thorax: Pronotum punctate, more conspicuous on collar and posterior lobe. Mesepimeron emergent. Evaporative area extensive, covering a thin fringe on mesopleura and all of metapleural area, except on dorsal quarter. Clavus with three complete rows of punctures and filled with two additional incomplete rows on distal three-quarters between inner and median rows. Procoxa with a small spine in both sexes. Profemur incrassate, elongate, with spines on apical half arranged in two rows. Protibia straight, with spiniferous tubercles, with or without spines in males. Protrochanter and mesofemur without spines in both sexes. Meso- and metatibiae with spiniform setae.</p><p>Abdomen, male genitalia: Aedeagus unspined, vesica with two lobes weakly sclerotized laterally, ejaculatory ductus wide; processus gonopori broadened towards apex.</p><p>Etymology</p><p>This new genus is named in honor of our friend and colleague Dr Richard Baranowski, who has published numerous important papers on the Lygaeoidea, including the ‘ Lygaeidae of Florida’ (Slater &amp;</p><p>© 2016 The Linnean Society of London, Zoological Journal of the Linnean Society, 2016, 177, 29–134</p><p>Baranowski, 1990) and the ‘ Lygaeidae of the West Indies’ ( Baranowski &amp; Slater, 2005). The gender is masculine.</p><p>This new genus is created to accommodate Heraeus elegans (Walker, 1873) and two new species. These are the largest species that previously would have been included in the genus Heraeus, but they lack the characteristic anteriorly protruding collar found on all members of that genus. Their general appearance and the male genitalia, with an unspined aedeagus with two basally sclerotized lobes, resemble species of the Neotropical genus Paisana Dellapé, 2008, including five species distributed in southern South America (Argentina and Brazil), but the species included in Baranowskiobius gen. nov. are larger and more slender, the juga is angulate, the aedeagus has a wide ejaculatory ductus, and the processus gonopori is broadened towards the apex. As in Paisana, the presence or absence of tubercles or spines on the male protibia and the serrated corial margin are considered intrageneric variation.</p></div>	https://treatment.plazi.org/id/6F64DB1B5E11FFFDFC7CFDD4FE4EFB1E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pablo M. Dellapé;María C. Melo;Thomas J. Henry	Pablo M. Dellapé, María C. Melo, Thomas J. Henry (2016): A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species. Zoological Journal of the Linnean Society 177 (177): 29-134, DOI: 10.1111/zoj.12362
6F64DB1B5E12FFF8FF03FB03FCD1F9AA.text	6F64DB1B5E12FFF8FF03FB03FCD1F9AA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Baranowskiobius elegans (WALKER 1873) Walker 1873	<div><p>BARANOWSKIOBIUS ELEGANS (WALKER, 1873)</p><p>COMB. NOV.</p><p>(FIGS 43 A, 44A, 45A–D, 46A, 47)</p><p>Nabis elegans Walker, 1873a: 144 .</p><p>Heraeus cincticornis Stål, 1874: 147; Berg, 1892: 162; Lethierry &amp; Severin, 1894: 191; Osborn, 1904: 201; Osborn &amp; Drake, 1915b: 537; Pennington, 1921: 19; Slater, 1964: 1082; Scudder, 1977: 30; Harrington, 1980: 108; Melo et al., 2004: 67 (misidentification); Dellape, 2014: 438. (NEW SYNONYMY).</p><p>Heraeus guttatus: Distant, 1903: 254, 255; Slater, 1964: 1083.</p><p>Heraeus elegans: Scudder, 1967: 264; Scudder, 1970: 100 (raised from syn with H. guttatus); Harrington, 1980: 108; Slater &amp; O’Donnell, 1995: 147.</p><p>Redescription (Fig. 43 A)</p><p>Lectotype: ♂. Total length 7.79.</p><p>Head: Porrect, convex dorsally, brown, shiny, coriaceous, with abundant short recumbent and long semi-erect and erect setae dorsally. Postocular region longer than preocular length. Head length 1.44, width 1.09. Postocular length 0.42. Eyes with setae between ommatidea. Ocelli large, placed just behind an imaginary line across the posterior margin of eyes. Interocular width 0.53, interocellar width 0.35. Labium pale brown, surpassing procoxae, almost extending to mesocoxae, with sparse short erect setae. Labial segment lengths: I 0.69, II 0.99, III 0.83, and IV 0.50. Antennal tubercles slightly divergent. Antennae pale brown, apical region of basiflagellomere and distiflagellomere darker, distiflagellomere with a wide pale stripe sub-basally; setae abundant, short, and recumbent. Antennal lengths: scape 0.80, pedicel 1.68, basiflagellomere 1.63, and distiflagellomere 1.49. Length of pale band on distiflagellomere 0.58.</p><p>Thorax: Pronotum with short recumbent adpressed setae, anterior lobe with sparse long erect setae; anterior pronotal lobe brown, posterior pronotal lobe paler, with irregular pale spots. Maximum width of anterior pronotal lobe behind middle. Collar length 0.14, anterior lobe length 0.74, posterior lobe length 0.64; anterior lobe width 1.09, posterior lobe width 1.68. Pleurae brown, with acetabular areas paler; setae short and recumbent. Scutellum brown, punctate, with setae similar to those on anterior lobe of pronotum. Hemelytra with short recumbent setae, general colour pale brown, except for anterior three-quarters of corial margins and a small subapical spot paler (Fig. 43 A). Corial margins slightly concave and serrate. Membrane brown, veins paler. Legs: Coxa yellowish brown, remainder of legs pale brown, except apical two-thirds of profemur, about distal half of meso- and metafemora, subproximal dark band of meso- and metatibia, apex of tibia and tarsus, and pretarsus darker. Darker region of profemora weakly mottled (Fig. 44 A). Femora with short semi-erect setae. Protibia with small denticles and four spiniferous tubercles, each bearing a spiniform seta on distal half.</p><p>Abdomen: Brown, with abundant short recumbent setae. Male genitalia: Pygophore (Fig. 45 A, B) rounded, elongate, anterior margin of dorsal aperture rounded, inner projections produced posteriorly. Parameres: Figure 45 (C, D). Aedeagus (Fig. 46 A) unspined, vesica with two lobes weakly sclerotized laterally; ejaculatory ductus wide; processus gonopori slightly broadened towards apex.</p><p>Variability observed in females</p><p>Similar to males in all respects, except labium extending to mesocoxae, and without or barely apparent spiniform tubercles on protibia; some specimens with all tibiae having a sub-basal dark band.</p><p>Variability observed in other material studied General colour darker and more contrasting than in lectotype; hemelytra with a small pale spot on middle inner corial margin.</p><p>Distribution</p><p>Argentina, Bolivia, Brazil, Colombia, Guatemala; and Ecuador, Paraguay, Peru, Uruguay, Venezuela (NEW RECORDS) (Fig. 47).</p><p>In his world catalogue, Slater (1964) erroneously credited Osborn (1904) with a record of this species from British Guiana and Berg (1892) with a record from Uruguay. Osborn (1904) actually mentioned only Cochabamba, Bolivia, and Berg (1892) recorded this species only from Chacabuco, Buenos Aires, Argentina. All previous records of this species (as Heraeus cincticornis) from Argentina are erroneous, and here we record B. elegans only from Misiones Province. Berg’s (1892) record from Buenos Aires is uncertain; his description is vague, so we are not convinced the record is accurate. Melo et al. (2004) recorded H. cincticornis from Corrientes (Argentina), but we have concluded that the specimens they examined were misidentified and should be referred to H. chamamecinus sp. nov.</p><p>Taxonomy</p><p>Distant (1903) synonymized H. elegans under H. guttatus . Scudder (1967) designated a lectotype for Nabis elegans Walker, 1873 and determined the correct generic combination for this species as: ‘ Heraeus elegans (Walker) ’. Slater &amp; O’Donnell (1995) misinterpreted Scudder’s treatment as raising H. elegans from synonymy under H. guttatus; Scudder (1970) later formally made this systematic change.</p><p>According to Stål’s original description of H. cincticornis, the specimen (or specimens) examined should be at the Naturhistoriska Riksmuseet, Stockholm (NHRS), as indicated after the geographic distribution ‘(Mus. Holm.)’. Scudder (1977) mentioned that the type material is not present at NHRS. We also confirm that Stål’s type material is apparently lost. Also because Stål did not give the number of specimens he examined when describing species, it is not certain how many specimens he used to describe H. cincticornis (he reported the sex: a female, and single measurements). But the listing of a single sex, single measurements, and a single museum does not necessarily mean he had only one specimen.</p><p>Most of Stål’s collection was deposited in the NHRS, but he also described some species from Signoret’s collection deposited in the International Research Institute of Entomology, Natural History Museum Vienna, Austria (NHMW), and a few from Dohrn’s collection, which should now be in the Museum and Institute of Zoology, Polish Academy of Sciences, Warsaw, Poland (ZMPA). We checked with the curators of these two museums, and in addition with the Museum für Naturkunde Humboldt-Universität zu Berlin, Germany (ZMHB), without success, indicating that the type material of H. cincticornis is lost. Thus, Scudder’s (1967) lectotype and interpretation of H. elegans is followed.</p><p>Type material examined</p><p>Lectotype: ♂, BRAZIL, Petropolis, Feb. 1857, J. Gray, 31. Nabis elegans, Walker’s catal. Nabis elegans Distant, 1873, G.G.E. Scudder, 1965 (BMNH).</p><p>Additional material studied</p><p>ARGENTINA: Misiones: 1♂, PN Iguazú, X-1980, luz,</p><p>D.J. Carpintero (MACN); 1♀, Puerto Iguazú, 12/13-</p><p>II-2007, E. Lestani (MLP); 3♂, 2♀, Apartado, XI-</p><p>[19]80 (USNM); 1♂, without head, Iguazú, XI-</p><p>[19]88, Drake (USNM); 3♂, 2♀, XI-[19]86, Drake</p><p>(USNM).</p><p>BOLIVIA: Cochabamba: 1♀, Cochabamba, VIII/IX- [18]99 (OSUC); La Paz: 3♂, Rio Zongo, 1400 m a.s.l., 24/30-X-[19]84, L.E. Peña (USNM).</p><p>BRAZIL: 1♂, Monat, IV-1935, P. Sandig (USNM); Minas Gerais: 1♂, Carmo do R. Claro, 15-VI- [19]43, Carvalho (USNM); 1♂, Viçosa, 25-IV- [19]33; E.J. Hambleton (USNM); 1♂, 1♀, 13-X/1- XI-1985, T.J. Henry &amp; S.P. Fiuza F. (USNM); 2♀, Viçosa, Corrego da Paraiso (Mata do Prefeitura), 10- III-1993, T.J. Henry (USNM); 6♂, 6♀, 26/27-III- 1993, T.J. Henry (USNM); Santa Catarina: 1♂, 2♀, Corupa, Hansa Humbolt, I-1945, A. Maller, Frank Johnson donor (AMNH); 1♀, XI-1944, A. Maller, Frank Johnson donor (AMNH); 1♀, XII-1944 (AMNH); Sao Paulo: 1♀, Santos, S.A., 23-V-1919 (AMNH); Roraima: 1♂, Mt Roraina, alt. 6000 ft, Glycon Swamp, 10-XI- 1927 (AMNH); Estado do Rio: 1♂, Mendés, 92 km Rio de Janeiro, De Moldt (MNHN); Espirito Santo: 1♀, Espirito Santo (MNHN).</p><p>COLOMBIA: Cundinamarca: 1♀, La Mesa, 14-VIII- 1965, J.A. Ramos (USNM); Magdalena: 1♀, Socorpa Mission, Sierra de Perija, 5/25-VIII-1968, B. Malkin (AMNH).</p><p>ECUADOR: Loja: 1♀, Celica, 2200 m a.s.l., 16/18- VIII-1977, L.E. Peña G. (USNM); Napo: 1♀, Sierrazul, 2200 m a.s.l., SW of Baeza, 0°40 ′ S, 77°5 ′ W, 2/30-I- 1996, T.J. Henry (USNM); 1♀, Yanayacu, 400 m a.s.l., IX/X-1977, L.E. Peña G. (USNM); Tungurahua: 1♂, 1♀, Baños, 30 km E, 25-I-1976, 4200 ft, blacklight, Spangler et al. Ecuador Peace Corps, Smithsonian Institution Aquatic Insect Survey (USNM); 2♂, (one without head), Baños, 12 km E, 1570 m a.s.l., 1°24 ′ S, 78°20 ′ W, seepage, 15-IX-1990, P.J. Spangler, #22 (USNM).</p><p>PARAGUAY: Alto Paraná: 1♂, 2♀, Itabó Res., 19- VI-1984, st. 105, L. Baert &amp; J.P. Maelfait (IRSN); 1♀, Itabó Res., 19-VI-1984, st. 105, L. Baert &amp; J.P. Maelfait (IRSN); Caaguazú: 1♂, Pastoreo, 3/5-I-1972, L.E. Peña (USNM).</p><p>PERU: Huánuco: 1♀, Tingo Maria, Turista Hotel, 2500 ft, 11/17-IV-1987, J.E. Eger (USNM); 1♀, Avispas, X-1962, L.A. Peña (USNM); Cuzco, 2♂, 1♀, Santa Isabel, valley of river Cosñipata, 16-XI-1951, F. Woytkowski (USNM); 1♀, 30-XI-1951, F. Woytkowski (USNM); 1♂, 22-XI-1951, F. Woytkowski (USNM).</p><p>VENEZUELA: Bolivar: 1♀, 26 km N Rio Yuruani, Gn. Sabana, 29-VI/10-VIII-[19]87, forest grassland edge, S. &amp; J. Peck (AMNH); Lara: 1♀, Yacambu National Park, 13 km SE Sanare, 4800 ft, 4/7-III- 1978, blacklight, cloud forest, J.B. Heppner (USNM); Portuguesa: 1♀, Acarigua, VI-[19]81, Drake (USNM).</p></div>	https://treatment.plazi.org/id/6F64DB1B5E12FFF8FF03FB03FCD1F9AA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pablo M. Dellapé;María C. Melo;Thomas J. Henry	Pablo M. Dellapé, María C. Melo, Thomas J. Henry (2016): A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species. Zoological Journal of the Linnean Society 177 (177): 29-134, DOI: 10.1111/zoj.12362
6F64DB1B5E17FFF5FF5BF981FB5FFE80.text	6F64DB1B5E17FFF5FF5BF981FB5FFE80.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Baranowskiobius bimaculatus	<div><p>BARANOWSKIOBIUS BIMACULATUS SP. NOV.</p><p>(FIGS 43 B, 44B, 45E–H, 48)</p><p>Diagnosis</p><p>Without or with a few long erect setae on pronotum and scutellum. Hemelytra with the corial margin serrate and a large whitish subapical spot. Male protibia with five spiniferous tubercles, each bearing a seta on the distal half, the proximal seta large, forming a strong spine.</p><p>Description (Fig. 43 B)</p><p>Holotype: ♂. Total length 7.60.</p><p>Head: Convex dorsally, coriaceous, dark brown, shiny, with abundant short recumbent semi-erect and erect setae dorsally. Head length 1.51, width 1.12. Postocular length 0.67. Ocelli large, placed posterior of an imaginary line passing the posterior border of eyes. Interocular width 0.48, interocellar width 0.24. Labium brown, segment II paler, with short erect setae; surpassing procoxae, almost extending to mesocoxae. Labial segment lengths: I 0.80, II 0.99, III 0.72, and IV 0.45. Antennal tubercles slightly divergent. Antenna pale brown, apical region of basiflagellomere and distiflagellomere darker, distiflagellomere with a wide pale band sub-basally; all segments with abundant short recumbent setae. Antennal lengths: scape 0.83, pedicel 1.62, basiflagellomere 1.49, and distiflagellomere 1.52. Length of pale band on distiflagellomere 0.40.</p><p>Thorax: Pronotum with very short adpressed setae; anterior pronotal lobe dark brown, collar and posteri- or pronotal lobe paler, posterior lobe with four irregular longitudinal pale stripes. Maximum width of anterior pronotal lobe behind middle. Collar length 0.13, anterior lobe length 0.88, posterior lobe length 0.58; anterior lobe width 1.01, posterior lobe width 1.55. Pleurae dark brown, acetabular areas paler, with short recumbent adpressed setae. Scutellum dark brown, punctate, with setae as on anterior lobe of pronotum. Hemelytra with short recumbent setae. General colour dark brown (Fig. 43 B); costal margin on proximal threequarters and a subapical inner corial spot pale. Membrane brown, with veins and an apical spot paler. Corial margin serrated. Legs: Coxae yellowish brown; femora creamy basally, apical two-thirds of profemur and distal half (approximately) of meso- and metafemur brown; tibiae and tarsi pale brown, except apex of tibiae and pretarsus darker (Fig. 44 B); darker region of profemur weakly mottled. Femora with sparse short erect setae. Protibia with five spiniferous tubercles, each bearing a spiniform seta on distal half, the proximal seta large, forming a strong spine.</p><p>Abdomen: Dark brown, with abundant short recumbent setae. Male genitalia: Pygophore (Fig. 45 E, F) rounded; anterior margin of dorsal aperture rounded. Parameres: Figure 45 (G, H). Aedeagus unspined, vesica with two slightly sclerotized lobes basally and laterally; ejaculatory ductus thickened before coil; processus gonopori long, slightly broadened towards apex.</p><p>Variability observed in other material studied</p><p>Some specimens have the apical region of basiflagellomere reddish brown. In addition, the colour pattern of the posterior pronotal lobe and hemelytra varies in some details. Some specimens, such as the holotype, have only very short setae on the anterior pronotal lobe and scutellum, whereas other specimens in both sexes have a few long erect setae on the anterior pronotal lobe and scutellum. It is probable that these long setae are very fragile and are broken in the specimens appearing not to have them. In general, females have a shorter anterior pronotal lobe and, as a result, the labium reaches the mesocoxae, and lack spiniform tubercles on protibia.</p><p>Distribution</p><p>Argentina, Bolivia, Brazil, Paraguay, and Uruguay (Fig. 48).</p><p>This species is common within its range and is frequently taken at lights.</p><p>Etymology</p><p>The specific epithet ‘ bimaculatus ’ is an adjective referring to the two distinct white spots on the apical third of the corium.</p><p>Type material</p><p>Holotype: ♂, ARGENTINA, P.N. Chaco, 26°48 ′ 25 ″ S, 59°36 ′ 26 ″ W, 26-IX-2009, Trampa de luz, P.M. Dellapé &amp; M.C. Melo (MLP).</p><p>Paratypes: ARGENTINA, Chaco: 3♂, 5♀, same data as for holotype (MLP); 1♂, 4♀, P.N. Chaco, entrance, 26°48 ′ 34,1 ″ S, 59°36 ′ 21,9 ″ W, 27-IX-2009, P. Dellapé &amp; M.C. Melo (MLP); 1♀, P.N. Chaco, 26°48 ′ 25 ″ S, 59°26 ′ 36.5 ″ W, 17/28-X-2009, Trampa de luz, R. Pfoh (MLP); 4♂, 1♀, Resistencia, III-1988, Mitre (USNM); Corrientes: 2♂, Ituzaingó, Reserva Santa María, 30-X-2003, light, P.M. Dellapé (MLP); Santiago del Estero: 2♂, 4♀, Quimili, 12-III- [19]92, L.E. Peña (USNM). BOLIVIA: Santa Cruz: 2♂, Saavedra, Res. Sta., 22-III-1978, UVtrap, C.R. Ward &amp; C.W. O’Brien (USNM). Brazil, Sao Paulo, 1♀, Cosmópolis, 22-I-1974, J.G. Rozen, F.C. Thomson &amp; J.S. Moure (AMNH); 1♂, 25-I-1974, J.G. Rozen, F.C. Thomson &amp; J.S. Moure (AMNH); 2♂, 2♀, Linhares, IX-1972, M. Alvarenga (AMNH). PARA- GUAY: Central: 1♂, 2♀, Cord. San Bernardino, Lago Ypacarai, 6/7-XI-1987, black light trap, J. Kochalka (USNM); 4♂, 4♀, 7/8-XI-[19]87, black light trap, J. Kochalka (USNM).</p><p>Additional material studied</p><p>ARGENTINA: Buenos Aires: 1♀, La Lucila del Mar,</p><p>I-1999, luz, D.L. Carpintero (MLP); 1♂, Lanús, I-1979, D.L. Carpintero (MACN); Chaco: 1♀, Reserva del Loro Hablador, XI-2006, light, G. Marti (MLP); 1♂, Resistencia, XI-[19]52 (MLP); 2♀, R.S. Peña, VII- 1971 (MLP); 1♀, Río de Oro, IX-1962 (MLP); 4♂, 8♀, Fontana (MLP); 1♀, 40 km Tres Estacas, 27°4 ′ 58.8 ″ S- 61°31 ′ 38.4 ″ W, 20/30-XI-2008, Marti &amp; Pelliza (MLP); Córdoba: 1♀, Depto. Tulumba, Cerro Colorado, XI- 1998, A. Cicchino (MACN); 1♀, Cruz del Eje, 5-II- 1999, J.E. Barriga (MACN); Corrientes: 1♂, Manantiales (MLP); 1♂, Manantiales, Colonia Inés Apostol, 1960 (MLP); 1♂, Paso de La Patria, 28-VIII-1963 (MLP); 1♂, San Roque, 2003, luz, M.C. Melo (MLP); 1♂, I-1921, De Carlo (MLP); 5♂, 5♀, Ituzaingó, Reserva Santa María, 30-X-2003, light, P.M. Dellapé (MLP); 1♀, Reserva Santa María, 27-IV-2003, luz, P.M. Dellapé (MLP); 1♀, 29-IV-2003, light, P.M. Dellapé (MLP); 1♀, Pellegrini, 6-XII-2001, light, P.M. Dellapé (MLP); Entre Ríos: 1♀, Liebig, I-2003, light L. Caire (MLP); 2♂, Brazo Largo, II-[19]39 (MLP); 1♀, X-[19]93, Peña &amp; Ugarte (USNM); Formosa: 7♂, 5♀, Ea. (Estancia) ‘La Marcela’, 35 km E de El Colorado, VIII-2003, light, J. Williams (MLP); 1♂, VIII-2003, light, F. Brusa (MLP); Misiones: 1♂, Departamento Concepción, Santa María, XII- 1947, M. Viana (MLP); 1 ♂, Posadas, D.L. Carpintero (MACN); Neuquén: 1♂, Catandil, 25-II-1942, M. Biraben (MLP); Rio Negro: 1♀, Rio Limay, 25-XI-[19]90, L.E. Peña (USNM); Salta: 1♂, Orán (MLP); 1♂, 1♀, Rio Sauzal, 16-X-[19]33 (MLP); Santa Fe: 1♂, Rafaela, 8-XI-[19]32 (MLP); 1♂, 2♀, Santa Fe (MLP); Santiago del Estero: 10♀, Añatuya, II-1999, light, D.L. Carpintero (MACN); 4♂, 5♀, XII-1998, light, D.L. Carpintero (MACN); 1♂, Cuestas del Salado, desvío 511, 4-XII-1928 (MLP); 1♂, Beltrán, 28-XII-[19]92, L.E. Peña (USNM); 1♂, 2♀, Telares, 3-III-[19]92, L.E. Peña (USNM); Tucumán: 1♀, Tucumán, 450 m a.s.l. (USNM).</p><p>BOLIVIA: 1♀, (MLP); 1♂, 1♀, 1962 (MLP); Cochabamba: 1♂, Chapare, Cristal Mayu, 1300 m a.s.l., 9/10-XII-[19]84, L.E. Peña (USNM); Chuquisaca: 1♀, Monteagudo, 24-XII-[19]84, L.E. Peña (USNM); 1♂, aircraft, 31-III-[20]02, Miami 230415 (USNM); La Paz, 1♀, La Paz, X-1956, L.E. Peña (USNM); Santa Cruz: 2♂, 1♀, Prov. Sara, 2 km NW Santa Rosa, 21-II-[19]69, black light trap, A. Martinez &amp; R.E. Woodruff (USNM); 1♀, San Esteban, Mushurina, 40 km N Santa Cruz, 1120 ft (USNM); 1♂, 1♀, Prov. Sara, Steinbach (AMNH); 1♀, 25-III- 1978, UV trap, O’Brien &amp; Serrate (USNM); 3♂, Mataral (N), V. Grande, 1800–2000 m a.s.l., 15/17- XII-1984, L.E. Peña (USNM).</p><p>BRAZIL: Sao Paulo: one without abdomen, Nova Europa, Faz. Itaquere, XII-1964, K. Lenko (MZSP); 4♂, 3♀ one without abdomen, Ilha Dos Buzios, 16- X/4-XI [1]963, Exp. Dep. Zool. (MZSP); 1♂, Ulatula, 22-X-1962, J. Malik (USNM); 1♀, Piracicaba, 12-X- 1965, blacklight, C.A. Triplehorn (AMNH); 1♀, 3-II- 1966, blacklight, C.A. Triplehorn (AMNH); 1♂, 20- I-1966, blacklight, C.A. Triplehorn (AMNH); 1♀, ESALQ, 5-III-[19]89, BL Trap, R.M. Baranowski (USNM); 1♀, Cosmopolis, 22-I-1974, J.G. Rozen, F.C. Thomson &amp; J.S. Moure (AMNH); 1♂, 23-I- 1974, J.G. Rozen, F.C. Thomson &amp; J.S. Moure (AMNH); 2♂, 2♀, 25-I-1974, J.G. Rozen, F.C. Thomson &amp; J.S. Moure (AMNH); 1♂, 1♀, 26-I-1974, J.G. Rozen, F.C. Thomson &amp; J.S. Moure (AMNH); 1♀, Gramadinha, 6-II-[19]89, R.M. Baranowski (USNM); Espirito Santo: 1♂, Concenção da Barra, X-1972, M. Alvarenga (AMNH); 11♂, 8♀, Linhares, IX- 1972, M. Alvarenga (AMNH); Pernambuco: 1♀, Bnito Prov. (USNM); Minas Gerais: 1♀, Viçosa, 13-X/1- XII-1985, T.J. Henry &amp; S.P. Fiuza Ferreira (USNM); 1♂, Pedra Azul, 900 m a.s.l., XI-1972, M. Alvarenga (AMNH).</p><p>PARAGUAY: 1♀, Paraguay, XI-1989, Drake (USNM); Alto Paraguay: 1♂, 1♀, Caacupé, 15/19-XI-[19]90, G. Arriagada (USNM); Alto Paraná: 1♂, Alto Paraná, 4/12-XII-[19]90, G. Arriagada (USNM); 2♂, 1♀, 12/ 16-XI-[19]90, G. Arriagada (USNM); Boquerón: 2♀, Boquerón, 30-XI-[19]92, Drake (USNM); 1♀ one without abdomen, Boq. New Land, 14-X-90, Drake (USNM); Caaguazú: 1♂, Estancia Primera, 3-I- 1932, taken at light, R.F. Hussey (USNM); 1♂, Villa Rica, II-1934, F. Schade (USNM); 1♂, 19-VII-1937, F. Schade (USNM); 1♂, 1♀, IX-1934, F. Schade (USNM); Caazapá: 1♀, Caazapá, 19-XI-[19]91, Arriagada (USNM); Central: 1♀, Capiatá, II-1995, Drake (USNM); 1♀, XI-1993, Drake (USNM); 1♂, 1♀, 2-X-[19]93, Drake (USNM); 2♂, Capiata, II- [19]95, L.E. Peña (USNM); Concepcion: 1♀, Horqueta, 23°24 ′ N, 57°10 ′ W, 44 km E Paraguay river, 27-I- 1936, A. Schulze (USNM); 1♀, 8-XI-1934, A. Schulze (USNM); 1♂, 1♀, Horqueta, 1938, A. Schulze (USNM); 1♂, 17-XI-1933, A. Schulze (USNM); 1♀, 9-VI-1934, A. Schulze (USNM); 1♂, 1♀, 14-VII- 1935, A. Schulze (USNM); Cordillera: 1♀, Caacupé, Inst. Agr. Nac., 12-XII-[19]80, R.D. Cave (USNM); 1♀, 9-XII-[19]80, R.D. Cave (USNM); 1♂, 2-XII-[19]80, R.D. Cave (USNM); Distrito Capital: 2♀, Cerro Lambare, 10-XI-[19]90, Arriagada (USNM); 3♂, 2♀, Tati, 14-XI-[19]90, Arriagada (USNM); 1♀, Asunción, XI/XII-1944, A. Schulze (USNM); 1♂, Asunción, J. Botánico, XI-[19]90, G. Arriagada (USNM); 1♂, Asunción, 16-III-[19]32 (MLP); San Pedro: 1♂, 1♀, Cororó, 23-II-[19]93, L.E. Peña (USNM); Presidente Hayes: 1♂, Monte Lindo, 13-XI-[19]93 (USNM); 1♂, Orquidaceae, 15-V-[20]02 (USNM).</p><p>URUGUAY: 1♀, Casupá, Fda., 9-III-1961, C.S. Carbonel, A. Mesa &amp; L. Zollesi (MLP).</p></div>	https://treatment.plazi.org/id/6F64DB1B5E17FFF5FF5BF981FB5FFE80	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pablo M. Dellapé;María C. Melo;Thomas J. Henry	Pablo M. Dellapé, María C. Melo, Thomas J. Henry (2016): A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species. Zoological Journal of the Linnean Society 177 (177): 29-134, DOI: 10.1111/zoj.12362
6F64DB1B5E1AFFF4FCC4FEBEFE10FCB9.text	6F64DB1B5E1AFFF4FCC4FEBEFE10FCB9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Baranowskiobius muticus	<div><p>BARANOWSKIOBIUS MUTICUS SP. NOV.</p><p>(FIGS 43 C, 44C, 45I–L, 47)</p><p>Diagnosis</p><p>Highly setose species, with long erect setae on ante-</p><p>rior and posterior pronotal lobes. Hemelytra with the</p><p>corial margin smooth; clavus unicolorous, without a</p><p>large subapical corial spot. Male protibia without spines.</p><p>Description (Fig. 43 C)</p><p>Holotype: ♂. Total length 6.84. Strongly setose.</p><p>Head: Brown, shiny, coriaceous, with abundant short recumbent, semi-erect, and erect setae. Eyes with many long setae between ommatidea. Head length 1.39, width 1.09, postocular length 0.41. Ocelli at level of an imaginary line passing the posterior border of eyes. Interocular width 0.51, interocellar width 0.27. Labium pale brown, with long erect setae; extending to mesocoxae. Labial segment lengths: I 0.72, II 0.82, III 0.64, and IV 0.45. Antennal tubercles slightly divergent. Antennae with abundant semi-erect and erect setae; pedicel with erect setae, equal or longer than diameter of segment; antenna pale brown, distiflagellomere without a pale band. Antennal lengths: scape 0.59, pedicel 1.36, basiflagellomere 1.18, and distiflagellomere 1.39.</p><p>Thorax: Anterior and posterior pronotal lobes with short and long erect setae. Anterior pronotal lobe brown; posterior pronotal lobe pale brown, with paler areas and with humeral angles pale. Maximum width of anterior pronotal lobe behind middle. Collar length 0.11, anterior lobe length 0.75, posterior lobe length 0.53; anterior lobe width 0.96, posterior lobe width 1.46. Pleurae brown, with short recumbent setae. Scutellum brown, punctate, with setae as on anterior lobe of pronotum. Hemelytra with abundant short erect setae; general colour pale brown, except anterior threequarters of corial margins and a small subapical corial spot paler, clavus unicolorous (Fig. 43 C). Corial margin smooth. Membrane pale brown, veins concolorous. Legs: Coxae yellowish brown, profemur except base, onethird of meso- and metafemur, apex of tibiae, tarsus, and pretarsus brown, rest of legs pale brown (Fig. 44 C); legs setose, with abundant long erect setae. Protibiae without spines.</p><p>Abdomen: Brown, with abundant short recumbent setae. Male genitalia: Pygophore (Fig. 45 I, J) rounded; anterior margin of dorsal aperture rounded; inner projections subquadrangular, with an anterior digitiform projection. Parameres: Figure 45 (K, L). Aedeagus: unspined, vesica with two lobes weakly sclerotized laterally, ejaculatory ductus thickened before coil; processus gonopori broadened towards apex.</p><p>Distribution</p><p>Bolivia and Brazil (Fig. 47).</p><p>Etymology</p><p>The specific epithet ‘ muticus ’ is an adjective referring to the lack of spines on the protibiae.</p><p>Type material</p><p>Holotype: ♂, BOLIVIA, Santa Cruz: Prov. Sara, 2 km NW Santa Rosa, 21-II-1969, blacklight trap, A. Martinez &amp; R.E. Woodruff (USNM).</p><p>Paratypes: BOLIVIA, 1♀, Saavedra, Agr. Exp. Sta., 27-XII-1959, blacklight trap, R.B. Cuming (USNM); BRAZIL, 1♂, Espiritu Santo, Linhares, IX-1972, M. Alvarenga (AMNH).</p></div>	https://treatment.plazi.org/id/6F64DB1B5E1AFFF4FCC4FEBEFE10FCB9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pablo M. Dellapé;María C. Melo;Thomas J. Henry	Pablo M. Dellapé, María C. Melo, Thomas J. Henry (2016): A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species. Zoological Journal of the Linnean Society 177 (177): 29-134, DOI: 10.1111/zoj.12362
6F64DB1B5E1BFFF4FEFFFC9EFC0EFBC0.text	6F64DB1B5E1BFFF4FEFFFC9EFC0EFBC0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Paraheraeus	<div><p>PARAHERAEUS GEN. NOV.</p><p>Type species</p><p>Heraeus eximius Distant, 1882 . HERE DESIGNAT- ED.</p><p>Diagnosis</p><p>Postocular region elongate, longer than interocellar distance, subequal to interocular distance. Juga rounded. Mesepimeron emergent. Evaporative area extensive. Clavus with three complete rows of punctures and a fourth incomplete row on distal two-thirds between median and internal row. Procoxa with a spine. Protrochanters without spines. Aedeagus unspined, vesica with two slightly sclerotized lobes, and processus gonopori long and slender.</p><p>Description</p><p>Relatively large and elongate.</p><p>Head: Elongate, vertex convex, preocular region longer than postocular. Postocular region elongate, longer than interocellar distance, subequal to interocular distance. Juga rounded. Buccular juncture V-shaped, near insertion of labium. Eyes relatively small, not surpassing the dorsal margin of head in lateral view.</p><p>Thorax: Anterior pronotal lobe almost impunctate, except on collar delimited posteriorly by a sulcus and with a row of punctures; sulci and posterior pronotal lobe punctate. Mesepimeron emergent. Evaporative area extensive, covering a thin fringe on mesopleura and approximately half of metapleural area. Clavus with three complete rows of punctures and a fourth incomplete row on distal two-thirds between median and internal row. Procoxa with a spine in both sexes. Protibia straight and without spines. Profemur incrassate, elongate, with two rows of spines. Protrochanters without spines. Mesofemur without spines.</p><p>Abdomen, male genitalia: Aedeagus unspined, vesica with two slightly sclerotized lobes; processus gonopori long and slender.</p><p>Etymology</p><p>This new generic name is based on the Greek prefix ‘para’, meaning near, and the generic name Heraeus, to denote the general similarity of these two taxa. The gender is masculine.</p><p>This new genus is created to accommodate Heraeus eximius Distant, 1882, a species that does not fit into any other known myodochine genus. The relative large size, shiny head, with postocular length subequal to the interocular width, a rounded vertex and bucculae, and the subshiny anterior pronotal lobe, together with the characters from the diagnosis, distinguish Paraheraeus from all other genera.</p><p>Paraheraeus eximius runs to the couplet 33 in Harrington’s (1980) key to the Myodochini of the world, and the genera Catenes Distant, 1893 and Heraeus Stål, 1862 . The species of Catenes have a distinctive flattened head and an aedeagus with large spines on conjunctiva (Dellapé &amp; Melo, 2008). Species of Heraeus have an elongated head, similar to that of P. eximius; however, the diagnostic pronotal collar in Heraeus species, which narrows dorsally and broadens ventrally to extend forwards beneath the head, is absent in P. eximius .</p></div>	https://treatment.plazi.org/id/6F64DB1B5E1BFFF4FEFFFC9EFC0EFBC0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pablo M. Dellapé;María C. Melo;Thomas J. Henry	Pablo M. Dellapé, María C. Melo, Thomas J. Henry (2016): A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species. Zoological Journal of the Linnean Society 177 (177): 29-134, DOI: 10.1111/zoj.12362
6F64DB1B5E1BFFF2FC86FB66FB4AFD72.text	6F64DB1B5E1BFFF2FC86FB66FB4AFD72.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Paraheraeus eximius (DISTANT 1882) Distant 1882	<div><p>PARAHERAEUS EXIMIUS (DISTANT, 1882) COMB. NOV.</p><p>(FIGS 43 D, 44D, 45M–P, 46B, 49)</p><p>Heraeus eximius Distant, 1882: 204 –205; Lethierry &amp; Severin, 1894: 191; Van Duzee, 1916: 21; Van Duzee, 1917: 179; Torre Bueno, 1946: 69–70; Scudder, 1967: 265 (lectotype designation); Slater, 1964: 1082– 1083; Schaefer, 1972: 812; Harrington, 1980: 109; Ashlock &amp; Slater, 1988: 228; Slater &amp; O’Donnell, 1995: 147; Slater &amp; Brailovsky, 2000: 332; Cervantes Peredo &amp; Brailovsky, 2004: 97.</p><p>Redescription (Fig. 43 D)</p><p>Paralectotype ♂.</p><p>Head: Porrect, brown, shiny, coriaceous, with short recumbent, long semi-erect, and erect setae dorsally; elongate, convex dorsally, preocular length longer than postocular length. Eyes relatively small, not surpassing the dorsal margin of head in lateral view. Ocelli small, posterior to an imaginary line passing the posterior border of eyes. Labium pale brown, with short erect setae; almost extending to mesocoxae. Antennae brown, apex of basiflagellomere reddish brown, distiflagellomere with a wide pale band sub-basally; all segments with abundant short recumbent setae.</p><p>Thorax: Pronotum pale brown to brown, pruinose, anterior lobe subshiny, collar well delimited posteriorly by a sulcus and with a row of punctures, anterior and posterior lobes with short recumbent and long erect setae. Posterior pronotal lobe and sulci punctate. Maximum width of anterior pronotal lobe behind middle. Pleurae brown. Scutellum brown, dull, basal half darker, punctate, with short recumbent and long erect setae. Hemelytra: clavus brown with a pale stripe between outer and inner row of punctures on basal half; corium with lateral margins slightly concave, smooth, general colour brown, costal margin pale on proximal three-quarters, with a large subapical pale brown spot (Fig. 43 D); with short recumbent and semierect setae. Membrane brown, veins concolourous, with a diffuse apical pale spot. Legs: Brown, meso- and metatrochanter and meso- and metafemur pale brown basally (Fig. 44 D); with abundant erect setae, longer on profemur. Mesofemur with long erect setae ventrally only; metafemur with long erect setae ventrally and restricted to apical third dorsally. Profemur with two rows of spines on apical half. Protibia without spines. Tibiae with spiniform setae.</p><p>Abdomen: Brown, with abundant short recumbent setae. Male genitalia: Pygophore (Fig. 45 M, N) rounded, anterior margin of dorsal aperture almost straight, inner projections subrectangular, more produced posteriorly. Parameres with outer projection bearing long setae and a prominence basally on inner projection (Fig. 45 O, P). Aedeagus (Fig. 46 B) unspined, vesica with two weakly sclerotized lobes, wings of the ejaculatory reservoir large and bent apically, apex directed backwards; processus gonopori long and slender.</p><p>Distribution</p><p>Guatemala, Mexico, and USA; Honduras (NEW RECORD) (Fig. 49).</p><p>Variability observed</p><p>Distant described this species based on specimens from San Geronimo, Capetillo, and Las Mercedes in Guatemala; Scudder (1967) designated a female from Las Merdeces as the lectotype. Some specimens differ from the paralectotype by the smaller size, the longer labium extending to mesocoxae, the presence of long erect setae on posterior pronotal lobe, the generally darker colouration, and the more contrasting colour pattern of the hemelytra. In some specimens, the darker basal half of the scutellum is not quite evident and males generally have a more strongly globose anterior pronotal lobe than females.</p><p>Type material examined</p><p>Paralectotype: ♂, [GUATEMALA], Las Mercedes, 3000 ft, Champion, Distant coll. 1911, 383 (BMNH).</p><p>Additional material studied</p><p>GUATEMALA: Escuintla: one without abdomen, Palin,</p><p>V-1924, W.M. Mann (USNM); Chimaltenango: 1♀, Agua</p><p>Caliente, Kelerman (AMNH).</p><p>HONDURAS: 1♂, 2♀, Dept. Comayagua, Rancho Chiquito, km 62, 29-V-[19]64, blacklight trap, F.S. Blanton, A.B. Broce &amp; R.E. Woodruff (USNM); 1♂, Dept. Atlantida, Lancetilla, U.F. Co., 4-VI- [19]64, blacklight trap, F.S. Blanton, A.B. Broce (USNM); 1♀, on Cucurbita sp., 15-V-2005, Miami, 283412, compared with lectotype female of H. eximius T.J. Henry 2008 (USNM).</p><p>MEXICO: 1♂, intercepted at Nogales (Arizona) from Mexico, 3-XI-2010, on Physalis philadelphica (USNM); Baja California: 2♂, 1♀, 5 mi. S Miraflores, 10-VII- [19]38, Michelbacher &amp; Ross (CAS); 1♀, Baja California Sur, 12 mi. S Loreto, Rancho de Parras, 29- I-[19]65, V. Roth (AMNH); Chiapas: 1♂, 17 mi. SE Teopisca, Rt. 24, 3/4-VI-1969, J.E.H. Martin (CNC); Jalisco: 1♀, Estación de Biología, Chamela, noct., 7-II-[19]75, H. Brailovsky (CAS); 1♂, Plan de Barrancas, SE Ixtlan, 8-IX-1966, J. &amp; W. Ivie (AMNH); Michoacan: 2♂, 12 mi. S Tzitzio, on Ehuetamo rd., 1050 m a.s.l., 19°20 ′ N, 100°50 ′ W, 10-VII-1947, 91, T.H. Hubbell (UMMZ); Morelos: 1♂, Tepoztlan, 5-V- 1963, W.J. Gertsch &amp; W. Ivie (AMNH); 1♂, Cuernavaca, 1959, N. Krauss (USNM); Nayarit: 1♀, Puerta de La Lima, 10-XI-1950, R.F. Smith (AMNH); one without abdomen, Nayarit, 15 mi. E of San Blas, CL1026, 21-IV-1964, J.T. &amp; M.S. Polhemus (AMNH); Oaxaca: 1♂, 5 mi. W El Camarón, 20-V-1969, J.E.H. Martin (CNC); one without abdomen, Juquilla, III-[19]64, W. Miller (CNC); Puebla: 1♀, 8 mi. S Izucar de Matamoros, 10-XII-[19]48, E.S. Ross (CAS); Sonora: 1♀, Alamos, 27-II-1963, P.H. Arnaud Jr. (CAS); 1♀, Bakachaka, Rio Mayo, 4-VII-[19]35, L.G. Gentner (CAS); Veracruz: 1♂, 1♀ Cotaxtla Exp. Sta., Cotaxtla, 26-VI-1962, D.H. Janzen (AMNH); 1♀, Palma Sola, 15-VII-1972, P. Reyes (MLP).</p><p>USA: Arizona: 1♂, Parker Ranch, Sixshooter Cn. Globe, Gila Co., 22-VIII-[19]52, H.B. Leech &amp; J.W. Green (CAS); 1♀, Pima Co., 1 mi. SE Catalina, VII-10- 1977, blacklight sheet, S. Kuba (AMNH); 1♂, Arizona, C.U. lot 34., P.R. Uhler (USNM); 1♀, C.F. Baker, C.U. lot 34., P.R. Uhler (USNM).</p></div>	https://treatment.plazi.org/id/6F64DB1B5E1BFFF2FC86FB66FB4AFD72	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pablo M. Dellapé;María C. Melo;Thomas J. Henry	Pablo M. Dellapé, María C. Melo, Thomas J. Henry (2016): A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species. Zoological Journal of the Linnean Society 177 (177): 29-134, DOI: 10.1111/zoj.12362
