identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
6F6ED021D62BFFCAFD18FF2B7E6CF93D.text	6F6ED021D62BFFCAFD18FF2B7E6CF93D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tomarus laticaudus Lopez-Garcia & Deloya 2019	<div><p>Tomarus laticaudus López-Garc´ıa and Deloya, new species</p> <p>Zoobank.org/ urn:lsid:zoobank.org:act: AD2C9C9C-89DE-4DE3-94EE-FD4F50CBCC7E (Figs. 2–5)</p> <p>Type Material. 56, 16♀. Holotype labeled “Corazal/ CZ / Pn / iv.27.11 / AugBusck ” (16, USNM). Paratypes: Same label data as holotype (263♀ USNM). “Corazal/ CZ / Pn / iv.7.12 / A. Busck / light” (16 USNM). “ Diablo Hts. / CZ / Pn / 3.v.1971 / W.E. Bivin ” (1♀ USNM). “Arc-light/ Ancon / C.Z. / Pan. / 12.v.11 / A.H. Jennings ” (16 UNSM, 4♀ USNM). “ Arc-light globe/ Ancon / C.Z. / Pan. / Apr.1911 / Kraft” (2♀ UNSM, 2♀ USNM). “ Arc-light globe/ Ancon / C.Z. / Pan. / 27. Apr.1911 / E.A. Schwarz ” (2♀ USNM). “Paraiso / C.Z. / iv.24.11 / August Busck ” (1♀ UNSM). “ Panama / 27.xii.1912 / W. Robinson” (1♀ USNM).</p> <p>Description. Holotype. Habitus as in Fig. 2. Length 22.2 mm; width across humeri 12.0 mm. Color brown to black, venter slightly lighter. Head: Frons coarsely rugose. Frontoclypeal region with 2 triangular tubercles separated by about 7 tubercle diameters. Clypeus narrowed towards apex, base 3 times as wide as apex. Clypeal teeth triangular, separated by 1 tooth diameter. Mandibles with 2 apical, acute teeth and a rounded basal lobe. Interocular distance equal to 3.3 times eye width. Antennal club short, subequal in length to antennomeres 2–7. Pronotum: Surface smooth on disc, with minute punctures on lateral margins. Apical tubercle small and rounded. Fovea shallow, striate, and about half as wide as interocular distance. Margin of apical angles homogeneously rounded, not emarginate. Elytra: First interval punctate, punctures smaller than on other intervals. Pygidium: Surface rugose on basal fourth and lateral angles, disc with minute punctures. Slightly convex in lateral view. Apex regularly rounded. Legs: Protibia tridentate with an additional small basal tooth. Protarsus subcylindrical, not enlarged; inner claw entire. Metatibia not narrowed before apex, sides nearly parallel. Apex of metatibia crenulate, with 11 spinules. Parameres: With 2 pairs of dorsal lateral teeth; basal teeth long, wide, and upwardly arched, apical teeth short and joined to the basal (Figs. 3–4).</p> <p>Variation. Male (4 paratypes): Length 18.8–23.3 mm; width across 9.9–12.1 mm. Female (16 paratypes): Length 19.9–24.4; width across 10.0– 12.2 mm; apex of pygidium dilated apically (Fig. 5); apex of metatibia with 10–12 spinules.</p> <p>Etymology. The epithet laticaudus comes from the Latin words lati (large) and cauda (tail), which refer to the enlarged pygidium of the females.</p> <p>Distribution. Tomarus laticaudus is known from four localities in the former Canal Zone in Panama (Fig. 23), which correspond to tropical moist forest areas at 10–80 m elevation in Panamá province.</p> <p>Temporal Distribution. April (14), May (6), December (1).</p> <p>Diagnosis. Tomarus laticaudus can be recognized by the nearly impunctate pronotum, small pronotal tubercle and fovea, and protibia with three teeth and an additional small basal tooth. It is similar to Tomarus laevicollis (Bates), Tomarus colombianus López-Garc´ıa and Gasca-Álvarez, and Tomarus maracaiboensis López-Garc´ıa and Deloya, new species, but it can be differentiated by the strongly dilated pygidium of the females (Fig. 5), which is homogeneously rounded in the other species (e.g., T. laevicollis (Fig. 6), and the wide basal teeth of the parameres (Figs. 3–4).</p> </div>	https://treatment.plazi.org/id/6F6ED021D62BFFCAFD18FF2B7E6CF93D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	López-García, Margarita M.;Deloya, Cuauhtémoc	López-García, Margarita M., Deloya, Cuauhtémoc (2019): Five New Species of the Dynastine Genus Tomarus Erichson (Coleoptera: Scarabaeidae), with an Illustrated Key to Species. The Coleopterists Bulletin 73 (1): 127-141, DOI: 10.1649/0010-065X-73.1.127, URL: http://dx.doi.org/10.1649/0010-065x-73.1.127
6F6ED021D62CFFCBFD13FF2B7DA8FB68.text	6F6ED021D62CFFCBFD13FF2B7DA8FB68.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tomarus maracaiboensis Lopez-Garcia & Deloya 2019	<div><p>Tomarus maracaiboensis López-Garc´ıa and Deloya, new species</p> <p>Zoobank.org/ urn:lsid:zoobank.org:act: 10BCDC71-EA26-4D32-9CAC-BABC1468FE9F (Figs. 7–9)</p> <p>Type Material. 36. Holotype labeled “Maracaibo” (16 USNM). Paratypes: Same label data as holotype (16 USNM, 16 UNSM).</p> <p>Description. Holotype. Habitus as in Fig. 7. Length 23.3 mm; width across humeri 2.2 mm. Color brown to black, venter slightly lighter. Head: Frons coarsely rugose. Frontoclypeal region with 2 triangular tubercles separated by about 7 tubercle diameters. Clypeus narrowed towards apex, base 3 times as wide as apex. Clypeal teeth triangular, separated by 1 tooth diameter. Mandibles with 2 apical, acute teeth and a rounded basal lobe. Interocular distance equal to 3.3 times eye width. Antennal club short, subequal in length to antennomeres 2–7. Pronotum: Surface smooth on disc, with minute punctures on lateral margins. Apical tubercle small and rounded. Fovea shallow, striate, and about half as wide as interocular distance. Elytra: First interval punctate, punctures smaller than on other intervals. Pygidium: Surface rugose on basal fourth and lateral angles, disc with minute punctures. Slightly convex in lateral view. Apex regularly rounded. Legs: Protibia tridentate with an additional small basal tooth. Protarsus subcylindrical, not enlarged; inner claw entire. Metatibia not narrowed before apex, sides nearly parallel. Apex of metatibia crenulate, with 8 spinules. Parameres: With 2 pairs of dorsal lateral teeth, both long and upwardly arched (Figs. 8–9).</p> <p>Variation. Male (2 paratypes): Length 20.9–21.1 mm; width across humeri 10.0– 12.3 mm. Apex of metatibia with 8–9 spinules.</p> <p>Etymology. The name maracaiboensis refers to the type locality.</p> <p>Distribution. Tomarus maracaiboensis is known only from Maracaibo, a municipality in northwestern Venezuela (Fig. 23).</p> <p>Diagnosis. This species can be recognized by the nearly impunctate pronotum, small pronotal tubercle and fovea, and tridentate protibia with an additional fourth basal tooth. It is similar to T. laevicollis, T. colombianus, and T. laticaudus but can be differentiated by the parameres with a very short apex and the apical teeth similar in shape and as long as the basal tooth.</p> <p>Comments. Tomarus laticaudus, T. colombianus, and T. maracaiboensis are found at elevations below 100 m. In particular, T. colombianus and T. maracaiboensis, which are the most similar in external appearance, are from coastal areas in South America. However, they occur in very distinct regions. Tomarus colombianus lives in moist forest in the Biogeographic Chocó with extremely high levels of precipitation, whereas T. maracaiboensis is from dry forest in northern Venezuela.</p> </div>	https://treatment.plazi.org/id/6F6ED021D62CFFCBFD13FF2B7DA8FB68	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	López-García, Margarita M.;Deloya, Cuauhtémoc	López-García, Margarita M., Deloya, Cuauhtémoc (2019): Five New Species of the Dynastine Genus Tomarus Erichson (Coleoptera: Scarabaeidae), with an Illustrated Key to Species. The Coleopterists Bulletin 73 (1): 127-141, DOI: 10.1649/0010-065X-73.1.127, URL: http://dx.doi.org/10.1649/0010-065x-73.1.127
6F6ED021D62DFFCBFF11FB577DAAFAC2.text	6F6ED021D62DFFCBFF11FB577DAAFAC2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tomarus moroni Lopez-Garcia & Deloya 2019	<div><p>Tomarus moroni López-Garc´ ıa and Deloya, new species</p> <p>Zoobank.org/ urn:lsid:zoobank.org:act: 66D8087E-B9F5-43F0-BC8C-3A38FDDAEB06 (Figs. 10–13)</p></div> 	https://treatment.plazi.org/id/6F6ED021D62DFFCBFF11FB577DAAFAC2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	López-García, Margarita M.;Deloya, Cuauhtémoc	López-García, Margarita M., Deloya, Cuauhtémoc (2019): Five New Species of the Dynastine Genus Tomarus Erichson (Coleoptera: Scarabaeidae), with an Illustrated Key to Species. The Coleopterists Bulletin 73 (1): 127-141, DOI: 10.1649/0010-065X-73.1.127, URL: http://dx.doi.org/10.1649/0010-065x-73.1.127
6F6ED021D62FFFC9FF7DFEF47F4BFC43.text	6F6ED021D62FFFC9FF7DFEF47F4BFC43.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tomarus paranaensis Lopez-Garcia & Deloya 2019	<div><p>Tomarus paranaensis López-Garc´ ıa and Deloya, new species</p> <p>Zoobank.org/ urn:lsid:zoobank.org:act: 5FF67FC3-A050-475F-9541-D42E2CCFFA26 (Figs. 17–19)</p> <p>Type Material. 16. Holotype labeled “ Ponta Grossa / Paraná / 3.ii 1965 /Moses / 26438 // Ligyrus gibbosus DeGeer // Halik det. 1965 // BRASIL Halik / 1966 / Collection” (16 USNM).</p> <p>Description. Holotype. Habitus as in Fig. 17. Length 15.5 mm; width across humeri 8.7 mm. Color reddish brown. Head: Frons coarsely punctate, area between eyes nearly impunctate. Frontoclypeal region with 2 cariniform tubercles separated by less than 1 tubercle diameter. Clypeus slightly narrowed towards apex, base 2 times as wide as apex. Clypeal teeth triangular, separated by a tooth diameter. Mandibles with 2 apical, acute teeth and a widely rounded basal lobe. Interocular distance equal to 5 times eye width. Antennal club long, 1.5 times longer than antennomeres 2–7. Pronotum: Surface with small, dense punctures. Apical tubercle and fovea absent. Margin of apical angles entire. Elytra: First interval punctate, punctures similar to those on other intervals. Pygidium: Surface with small, evenly distributed punctures; not rugose Punctures small, evenly distributed. Strongly convex in lateral view. Apex regularly rounded. Legs: Protibia tridentate without an additional small basal tooth. Protarsus subcylindrical, not enlarged; inner claw entire. Metatibia strongly narrowed before apex. Apex of metatibia entire, with 19 spinules. Parameres: With a medial tooth on each side; apex widened but ending in an acute, triangular point (Figs. 18–19).</p> <p>Etymology. The species’ name refers to the type locality in the state of Paraná, Brazil.</p> <p>Distribution. The holotype was collected in Ponta Grossa (Fig. 23), a municipality located in the Second Plateau of the state of Paraná (Brazil) within the Atlantic Forest biome, represented by the Araucaria Forest ecosystem (Mielke et al. 2012).</p> <p>Temporal Distribution. February (1).</p> <p>Diagnosis. Tomarus paranaensis and T. burmeisteri are similar in their external morphology. However, the metatibiae are strongly narrowed before the apex and have 19 spinules on the apical margin in T. paranaensis, whereas they have the sides nearly parallel and the apical margin with 25–27 spinules in T. burmeisteri. The parameres are apically acute in T. paranaensis (Figs. 18–19) but rounded to subquadrate in T. burmeisteri (Fig. 49; see also fig. 18 in Neita-Moreno and Ratcliffe 2017).</p> </div>	https://treatment.plazi.org/id/6F6ED021D62FFFC9FF7DFEF47F4BFC43	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	López-García, Margarita M.;Deloya, Cuauhtémoc	López-García, Margarita M., Deloya, Cuauhtémoc (2019): Five New Species of the Dynastine Genus Tomarus Erichson (Coleoptera: Scarabaeidae), with an Illustrated Key to Species. The Coleopterists Bulletin 73 (1): 127-141, DOI: 10.1649/0010-065X-73.1.127, URL: http://dx.doi.org/10.1649/0010-065x-73.1.127
6F6ED021D620FFC6FF64FF2B7830FC25.text	6F6ED021D620FFC6FF64FF2B7830FC25.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tomarus pilcopataensis Lopez-Garcia & Deloya 2019	<div><p>Tomarus pilcopataensis López-Garc´ıa and Deloya, new species</p> <p>Zoobank.org/ urn:lsid:zoobank.org:act: AC215467-CC35-4350-8574-00D3BA0652B2 (Figs. 20–22)</p> <p>Type Material. 16. Holotype labeled “ PERU: / Cuzco; / Pilcopata, / 600 meters / 11-14 xii 1979 / J.B. Heppner / premontane moist forest” (16 USNM).</p> <p>Description. Holotype. Habitus as in Fig. 20. Length 27.7 mm; width across humeri 14.3 mm. Head: Frons coarsely rugose. Frontoclypeal region with 2 transverse tubercles separated by about 3 tubercle diameters. Clypeus narrowed towards apex, base width 3.8 times as apex. Clypeal teeth triangular, separated by less than a tooth diameter. Mandibles with 2 apical, acute teeth and a rounded basal lobe. Interocular distance equal to 4.1 times eye width. Antennal club short, subequal in length to antennomeres 2–7. Pronotum: Surface with dense, large punctures. Apical tubercle small, rounded. Fovea shallow, striate, and about 1/5 as wide as interocular distance. Margin of apical angles entire. Elytra: First interval punctate, punctures similar in size to those on other intervals. Pygidium: Surface coarsely rugose. Slightly convex in lateral view. Apex regularly rounded. Legs: Protibia tridentate with an additional inconspicuous, basal convexity. Protarsus subcylindrical, not enlarged; inner claw entire. Metatibia not narrowed before apex, sides nearly parallel; apex slightly crenulate, with 33 spinules. Parameres: Lateral teeth short, widely triangular; apical fourth narrowed, apices directed outwards (Figs. 21–22).</p> <p>Etymology. The name of the species refers to the type locality, Pilcopata.</p> <p>Distribution. Tomarus pilcopataensis is known only from Pilcopata, a locality in the premontane moist forest of Peru at an elevation of 600 m (Fig. 23).</p> <p>Temporal Distribution. December (1).</p> <p>Diagnosis. Tomarus pilcopataensis is similar to T. pumilus and T. roigjunenti Neita and Ratcliffe, but it can be differentiate by the completely rugose surface of the pygidium (rugose on the basal third in the other two species). The general shape of the parameres of T. pumilus (Fig. 67) is similar that in the new species, but in T. pilcopataensis the apical fourth is strongly narrowed and the lateral medial teeth are wider and not directed backwards (Figs. 21–22).</p> <p>Comments. There are specimens of Tomarus maternus Prell and Tomarus gyas Erichson with the same collection and locality data (deposited at USNM), which indicates that these three species have sympatric populations in the western Amazonian region. Unfortunately, we were not able to find additional specimens of T. pilcopataensis in three entomological collections in Peru nor during a visit to the type locality in Manu National Park (Cusco).</p> </div>	https://treatment.plazi.org/id/6F6ED021D620FFC6FF64FF2B7830FC25	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	López-García, Margarita M.;Deloya, Cuauhtémoc	López-García, Margarita M., Deloya, Cuauhtémoc (2019): Five New Species of the Dynastine Genus Tomarus Erichson (Coleoptera: Scarabaeidae), with an Illustrated Key to Species. The Coleopterists Bulletin 73 (1): 127-141, DOI: 10.1649/0010-065X-73.1.127, URL: http://dx.doi.org/10.1649/0010-065x-73.1.127
