taxonID	type	description	language	source
737287D7FFBEEC29FDB0FE24FB35F81A.taxon	diagnosis	Diagnosis (Condé 1955 a; Paclt 1957; Sendra et al. 2017 a) Since the only diagnostic character of Plusiocampinae (as suggested by Paclt 1957) is the higher number of macrosetae on the pronotum, this taxon seems to be paraphyletic. The pronotum never has less than 4 + 4 macrosetae. However, many of the genera can be considered monophyletic within Plusiocampinae. This is the case for Anisuracampa, Cestocampa, Condeicampa, Hystrichocampa, Patrizicampa, Plusiocampa, Plutocampa, Simlacampa and Vandelicampa, characterized by pretarsal lateral crests. Nonetheless, these lateral crests are absent in Silvestricampa and Turkmenocampa, clearly isolating them from the other genera.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFB1EC26FD04FEB3FAA2F99C.taxon	diagnosis	Diagnosis (Condé 1955 a; Sendra et al. 2012 a) Non-protruding frontal process with non-tubercular setae. Mesonotum with 1 + 1 medial anterior, 3 + 3 lateral anterior and 2 + 2 lateral posterior macrosetae; metanotum with 1 + 1 medial anterior, 2 + 2 lateral anterior and 2 + 2 lateral posterior macrosetae. With or without medial posterior notal macrosetae. One dorsal femoral macroseta. Two to three ventral tibial macrosetae. Subequal elbow-like claws, posterior claw equal (1.0) to subequal (1.1) to anterior claw. Laminar pretarsal lateral processes with long, rigid, ventral hook-tipped barbs. Urotergites I – II with 1 + 1 – 2 + 2 posterior macrosetae; urotergite III with 2 + 2 – 4 + 4 posterior macrosetae; urotergites VI – VII with 3 + 3 lateral anterior and 5 + 5 posterior macrosetae. Urosternite I with 7 + 7 (15 + 15 in C. iberica Sendra & Condé, 2012), urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae. Males without g 1 glandular setae but with a 2 glandular setae on first urosternite appendages. Females with subcylindrical appendages with a 1 glandular setae.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFB1EC26FD04FEB3FAA2F99C.taxon	distribution	Habitat and distribution Two of the four species of Cestocampa occupy the subterranean habitat in caves from the south of the Italian Peninsula (C. gasparoi Bareth, 1988) and the east of the Iberian Peninsula (C. iberica). The other two are soil-dwelling species (C. italica (Silvestri, 1912) and C. balcanica Condé, 1986) that have been reported from the Italian Peninsula and Dinaric Mountains (Fig. 210).	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFB1EC27FDC5F9DFFC04FDFE.taxon	diagnosis	Diagnosis (Condé 1955 a; Sendra et al. 2012 a) Non-troglomorphic species. With 25 antennomeres in a regenerated antenna; frontal process with nontubercular setae. Pronotum with 1 + 1 ma, 2 + 2 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp, 1 + 1 mp macrosetae; metanotum with 1 + 1 ma, 2 + 2 la, 2 + 2 lp, 1 + 1 mp macrosetae; long, barbed notal macrosetae; sparse smooth notal clothing setae. Two ventral tibial macrosetae; subequal small claws (1.03) with small lateral crests, posterior claw without a backward overhang. Urotergite I with 1 + 1 post macrosetae; urotergites II – III with 2 + 2 post macrosetae; urotergite IV with 1 + 1 la, 5 + 5 post macrosetae; urotergite V with 2 + 2 (2 + 1) la, 5 + 5 post macrosetae; urotergites VI – VII with 3 + 3 la, 5 + 5 post macrosetae; urotergite VIII with 6 + 6 post macrosetae; abdominal segment IX with 8 + 8 (8 + 7) post macrosetae; urosternite I with 7 + 1 + 7, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae. Male unknown; female subcylindrical appendages with a 1 glandular setae.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFB1EC27FDC5F9DFFC04FDFE.taxon	distribution	Habitat and distribution Soil-dwelling species found in two remote localities, the type locality in Rabitlje, south-eastern Dinaric Mountains, and on Monti Aurunci, Lazio, central Italian Peninsula.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFB0EC27FDCFFDF2FAE5FAF7.taxon	diagnosis	Diagnosis (Bareth 1988) Troglomorphic species. Antennae with 27 – 29 antennomeres (perhaps regenerated); 12 – 13 simple olfactory chemoreceptors in cupuliform organ; frontal process with non-tubercular setae. Pronotum with 1 + 1 ma, 3 + 3 – 4 + 4 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp macrosetae; metanotum with 1 + 1 ma, 2 + 2 la, 2 + 2 lp macrosetae; long, barbed notal macrosetae; notal setal clothing sparse, long, smooth or with two apical barbs. Two ventral tibial macrosetae; subequal small claws with small lateral crests, posterior claw without a backward overhang. Urotergite I with 1 + 1 post macrosetae; urotergite II with 1 + 2 post macrosetae; urotergite III with 2 + 2 post macrosetae; urotergite IV with 1 + 1 la, 5 + 5 post macrosetae; urotergite V with 2 + 2 la, 5 + 5 post macrosetae; urotergites VI – VII with 3 + 3 la, 5 + 5 post macrosetae; urotergite VIII with 6 + 6 post macrosetae; abdominal segment IX with 8 + 8 post macrosetae; urosternite I with 7 + 7, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae. Male without glandular g 1 setae, enlarged subcylindrical appendages with thin and long glandular a 2 setae. Spermatozoid fascicle wheels with 2 ½ turns of 350 µm long and 8 µm wide filament. Female unknown.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFB0EC27FDCFFDF2FAE5FAF7.taxon	distribution	Habitat and distribution Subterranean species found in Grotta Superiore di Sant’Angelo Calabria, south Italian Peninsula.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFB0EC24FD9FFAEBFAF6FEAA.taxon	diagnosis	Diagnosis (Sendra et al. 2012 a) Troglomorphic species. Antennae with 34 – 41 antennomeres; 8 – 10 complex olfactory chemoreceptors in cupuliform organ; non-protruding frontal process with non-tubercular setae. Pronotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp, 1 + 1 mp macrosetae; metanotum with 1 + 1 ma, 2 + 2 la, 2 + 2 lp, 1 + 1 mp macrosetae; middle-sized, barbed notal macrosetae; notal clothing setae sparse, smooth or with thin distal barbs. Two or three ventral tibial macrosetae; subequal claws (1.05 – 1.10) with large lateral crests, posterior claw without a backward overhang. Urotergite I with 1 + 1 post macrosetae; urotergite II with 2 + 2 post macrosetae; urotergite III with 1 + 1 la, 2 + 2 post macrosetae; urotergite IV with 1 + 1 la, 4 + 4 post macrosetae; urotergites V – VII with 3 + 3 la, 5 + 5 post macrosetae; urotergite VIII with 7 + 7 post macrosetae; abdominal segment IX with 9 + 9 post macrosetae; urosternite I with 15 + 15, urosternites II – VII with 5 + 5, urotergite VIII with 2 + 2 macrosetae. Male without glandular g 1 setae, enlarged subcylindrical appendages with glandular a 2 setae; female with subcylindrical appendages with glandular a 1 setae. Spermatozoid fascicle wheels with 1 ½ turns of 70 – 90 µm long and 4 – 5 µm wide filament.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFB0EC24FD9FFAEBFAF6FEAA.taxon	distribution	Habitat and distribution Subterranean species found in 11 caves around the Castilian-Valencian Branch of the Iberian Mountain Range and its border with the northwest of the Pre-Baetic Mountains, eastern Iberian Peninsula.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFB3EC24FDC2FE1EFE31FC0B.taxon	diagnosis	Diagnosis (Silvestri 1912; Condé 1955 a) Non-troglomorphic species. Antennae with 22 antennomeres. Pronotum with 1 + 1 ma, 4 + 4 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp macrosetae; metanotum with 1 + 1 ma, 2 + 2 la, 2 + 2 lp macrosetae. Two ventral tibial macrosetae; subequal small claws with small lateral crests, posterior claw without a backward overhang. Urotergites I – II with 2 + 2 post macrosetae; urotergite III with 4 + 4 post macrosetae; urotergite IV with 2 + 2 la, 5 + 5 post macrosetae; urotergites V – VII with 3 + 3 la, 5 + 5 post macrosetae; urosternite I with 7 + 7, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae. Male without glandular g 1 setae and slightly enlarged appendages.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFB3EC24FDC2FE1EFE31FC0B.taxon	distribution	Habitat and distribution Soil-dwelling species known from the type locality in Strongoli, Calabria, southern Italian Peninsula, and northern Macedonia.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFB3EC24FDEEFBBEFE6DFAE0.taxon	diagnosis	Diagnosis Mesonotum and metanotum with medial posterior macrosetae. Five dorsal femoral macrosetae; one or two ventral and two dorsal tibial macrosetae. Unequal, elbow-like claws. Thick, subcylindrical, setiform, pubescent lateral process of pretarsus. Urotergites with up to 3 + 3 lateral anterior and 5 + 5 posterior macrosetae.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFB3EC25FDA9FAD2FDF9FEAB.taxon	diagnosis	Diagnosis (Condé 1948 a, 1955 a; Bareth 1968, 2006) Troglomorphic species. Antennae with 31 – 36 antennomeres; 8 to 10 olfactory chemoreceptors in cupuliform organ; protruding frontal process with tubercular setae. Pronotum with 1 + 1 ma, 4 + 4 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 2 + 2 la, 2 + 2 lp, 1 + 1 mp macrosetae; metanotum with 1 + 1 ma, 1 + 1 la, 2 + 2 lp, 1 + 1 mp macrosetae; long, barbed notal macrosetae; sparse long, thin clothing setae. Unequal, elbow-like claws (1.2) with large lateral crests, posterior claw with a backward overhang. Urotergites I – II with 1 + 1 post macrosetae; urotergite III with 1 + 1 la, 1 + 1 post macrosetae; urotergite IV with 1 + 1 la, 4 + 4 post macrosetae; urotergite V with 2 + 2 la, 5 + 5 post macrosetae; urotergites VIVII with 3 + 3 la, 5 + 5 post macrosetae; urotergite VIII with 6 + 6 post macrosetae; urosternite I with 7 + 7, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae. Male with glandular g 1 setae, enlarged subcylindrical appendages with glandular a 1 setae; female subcylindrical appendages with glandular a 1 setae. Spermatozoid fascicles 57 – 80 µm in diameter and 7 – 9 µm in width; wheels with 1 ½ turns of 150 – 200 µm long and 7 – 8 µm thick filament.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFB3EC25FDA9FAD2FDF9FEAB.taxon	distribution	Habitat and distribution Subterranean species widespread in the Jura karst mountains between France and Switzerland, where it is found in 18 caves (Fig. 210).	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFB2EC25FDFDFE26FC46FD10.taxon	diagnosis	Diagnosis (Condé 1962) Without dorsal femoral macrosetae or ventral tibial macrosetae; abundant macrosetae on all urosternites, including 3 + 3 macrosetae on urosternite VIII. Its apomorphic feature, the shape of the laminar pretarsal lateral process, is unique among diplurans. It starts with a large laminar lateral process, well barbed on the ventral side and ending in a thin, smooth, pointed process.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFB2EC22FDC9FD53FBDBFBC9.taxon	description	Figs 1 – 2	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFB2EC22FDC9FD53FBDBFBC9.taxon	diagnosis	Diagnosis (Condé 1962) Troglomorphic species. Cupuliform organ with 10 – 14 complex olfactory chemoreceptors. Protruding frontal process with non-tubercular setae. Pronotum with 1 + 1 ma, 4 + 4 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp macrosetae; metanotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp macrosetae; long, barbed notal macrosetae; long clothing setae covered with thin barbs. Subequal elbow-like claws (1.1) with large lateral crests, posterior claw with a backward overhang. Urotergite III with 1 + 1 la, 0 + 0 – 1 + 1 post macrosetae; urotergite IV with 2 + 2 la, 1 + 1 – 3 + 3 post macrosetae; urotergite V with 2 + 2 – 3 + 3 la, 2 + 2 – 4 + 4 post macrosetae; urotergites VI – VII with 3 + 3 la, 4 + 4 post macrosetae; urotergite VIII with 6 + 6 post macrosetae; abdominal segment IX with 8 + 8 – 8 + 9 post macrosetae. Urosternite I with 15 + 15 – 16 + 16, urosternites II – VII with 7 + 7 – 8 + 8, urosternite VIII with 3 + 3 macrosetae. Male without glandular g 1 setae, slightly subtrapezoidal appendages with glandular a 1 and a 2 setae; females with subcylindrical appendages and an apical area with glandular a 1 setae.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFB2EC22FDC9FD53FBDBFBC9.taxon	discussion	Remarks (Figs 1 – 2) The study of many specimens collected in 6 new caves (see Table S 2 in Supplementary Material) allowed us to complete Condé’s diagnosis, as well as to supplement it with some minor taxonomic details. Medial antennomeres with 10 – 12 gouge sensilla 40 – 50 µm long. End of metathoracic leg reaches end of abdomen; calcars covered with barbs; subapical setae with thin barbs almost from base up to twothirds of length. Stylus with an apical seta with a few thin barbs and a large, thick basal tooth, medial and subapical setae covered with thin barbs. Male appendages with up to 60 glandular a 2 setae and 20 a 1 glandular setae (Fig. 1); female appendages with up to 18 a 1 glandular setae. Spermatozoid fascicle wheels with 2 turns of 140 – 160 µm long and up to 4 µm wide filament (Fig. 2). Habitat and distribution Subterranean species found in eight caves in eastern Sardinia (Fig. 210).	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFB5EC3DFDF4FB84FAA2F963.taxon	diagnosis	Diagnosis (Silvestri 1912; Condé 1955 a; Sendra & Weber 2018) Apparently smooth epicuticle, usually reticulated at high magnification, rosetta pores absent. Head with a frontal process with or without tuberculate setae. In non-troglomorphic species, cupuliform organ with four or five spheroidal olfactory chemoreceptors with polygonal net, with pore surface made by at least one cup-shaped fold and a central structure ending in a terminal pore. In troglobiomorphic species, spheroidal olfactory chemoreceptor with more folds in spiral, radial or other complex shape and a more visible polygonal net with pore surface. Sensillum of third antennomere in ventral position. Meso- and metathorax with a few macrosetae but frequently with medial anterior, lateral anterior, lateral posterior and medial posterior (exceptionally medial intermediate and lateral intermediate) macrosetae. Femur with one to five dorsal macrosetae; tibia with one to three ventral macrosetae. Elbow-like claws with usually large lateral crests and setiform lateral processes, rarely with a few barbs on proximal part. Abdomen with lateral anterior and posterior macrosetae and never with medial anterior macrosetae. Sternal macrosetae: sternite I with 6 + 6 up to 60 macrosetae; sternites II – VII with 4 + 4 to up to 14 + 14 macrosetae; sternite VIII with 2 + 2 up to 4 + 4 macrosetae. Secondary sexual differences in shape of first urosternite appendages and number of glandular setae.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFB5EC3DFDF4FB84FAA2F963.taxon	distribution	Habitat and distribution This is the most diverse genus of Plusiocampinae, with a clear preference for the subterranean ecosystems (Condé 1955 a; Sendra & Weber 2018). It has 72 species (including the seven new ones described here) and 10 subspecies, distributed around the Mediterranean Basin (Figs 210 – 211). The exceptions are three Chinese species considered representatives of Plusiocampa by Silvestri (1931 a), Condé (1993 a) and Sendra et al. (2012 a): P. sinensis Silvestri, 1931, P. lipsae Condé, 1993 and P. kashiensis Chou & Tong, 1980. Nevertheless, these species must be removed from Plusiocampa due to the macrosetal formula on urosternites II to VIII.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFAAEC3AFDC6F95EFD08FE00.taxon	diagnosis	Diagnosis (Paclt 1957; Condé 1996) Dydimocampa shares with Plusiocampa s. str. the same taxonomic features, with the exception of the two dorsal femoral macrosetae. This fact, together with their distribution patterns, suggests that Dydimocampa is an artificial subgenus based on a single apomorphic feature that has probably appeared several times in Plusiocampa s. str.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFAAEC3AFDC6F95EFD08FE00.taxon	distribution	Habitat and distribution The three species treated here are cave-dwellers with a subterranean life-style and they are distributed in three disjunct karst areas in Europe (Fig. 211).	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFADEC3AFE23FDBBFBF8FAD4.taxon	diagnosis	Diagnosis (Condé & Sendra 1989) Troglomorphic species. Antennae with 35 – 40 antennomeres; up to 17 digitiform complex olfactory chemoreceptors on cupuliform organ; slightly protruding frontal process with tubercular setae. Pronotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 2 + 2 la, 2 + 2 lp, 1 + 1 mp macrosetae; metanotum with 1 + 1 ma, 1 + 1 la, 2 + 2 lp, 1 + 1 mp macrosetae; long, barbed notal macrosetae with thin barbs; sparse long, barbed notal clothing setae. Two to four ventral tibial macrosetae, femur / tibia ratio 0.69; subequal, elbow-like claws with large lateral crests, posterior claw with a small backward overhang; long lateral process. Urotergites I – II with 1 + 1 post macrosetae; urotergite III with 0 + 0 – 1 + 1 la, 1 + 1 post macrosetae; urotergite IV with 1 + 1 la, 2 + 2 – 4 + 4 post macrosetae; urotergite V with 1 + 1 la, 4 + 4 – 5 + 5 post macrosetae; urotergites VI – VII with 1 + 1 la, 5 + 5 post macrosetae; urotergite VIII with 6 + 6 post macrosetae; abdominal segment IX with up to 8 + 8 macrosetae; urosternite I with 7 + 7, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae. Male with an area of glandular g 1 setae, subcylindrical appendages with glandular a 1 setae; female appendages like male appendages. Spermatozoid fascicle wheels with 1 ½ turns of 100 – 140 µm long and 2 µm wide filament. Habitat and distribution Troglomorphic species found in four caves around the Sub-Baetic Interior Mountain Range, from Antequera (Málaga) to Sierra María (Almería), southern Iberian Peninsula.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFADEC3AFE6DFA97FBB9F817.taxon	diagnosis	Diagnosis (Condé 1996) Troglomorphic species. Antennae with 44 – 45 antennomeres; eight to ten complex olfactory chemoreceptors in cupuliform organ; slightly protruding frontal process with a few anterior tubercular setae. Pronotum with 1 + 1 ma, 1 + 1 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 2 + 2 la, 2 + 2 lp macrosetae; metanotum with 1 + 1 ma, 1 + 2 la, 2 + 2 lp macrosetae; long, barbed notal macrosetae; sparse short, smooth notal clothing setae. Three ventral tibial macrosetae; unequal claws (1.25) with large lateral crests, posterior claw with a small backward overhang; short lateral processes. Urotergites I – III with 1 + 1 post macrosetae; urotergite IV with 1 + 1 la, 4 + 4 post macrosetae; urotergite V with 2 + 2 la, 4 + 4 post macrosetae; urotergites VI – VII with 2 + 2 la, 5 + 4 post macrosetae; urotergite VIII with 6 + 6 post macrosetae; abdominal segment IX with 9 + 9 post macrosetae; urosternite I with 8 + 7, urosternites II – VII with 6 + 6, urosternite VIII with 2 + 2 macrosetae. Male without glandular g 1 setae, enlarged subcylindrical appendages with a large area of glandular a 1 setae; adult female unknown.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFADEC3AFE6DFA97FBB9F817.taxon	distribution	Habitat and distribution Subterranean species inhabiting the Movile Cave, Southern Dubrudja, Romania.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFACEC38FE3BFD13FED5FE53.taxon	description	Figs 3 – 8; Table 1	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFACEC38FE3BFD13FED5FE53.taxon	diagnosis	Diagnosis (Silvestri 1949; Condé 1996) Troglomorphic species. Antennae with 33 antennomeres. Protruding frontal process with tubercular setae. Pronotum with 1 + 1 ma, 2 + 2 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 1 + 1 la, 2 + 2 lp macrosetae; metanotum with 1 + 1 ma, 2 + 2 lp macrosetae; long, barbed macrosetae. Two ventral tibial macrosetae; unequal claws with large lateral crests, posterior claw with a backward overhang; short lateral anterior process and long lateral posterior process. Urotergites I – III with 1 + 1 post macrosetae; urotergites IV – VI with 1 + 1 la, 2 + 2 post macrosetae; urotergite VII with 1 + 1 la, 5 + 5 post macrosetae. Urosternite I with 7 + 7, urosternites II – VII with 6 + 6, urosternite VIII with 2 + 2 macrosetae. Male unknown; female with subcylindrical appendages with glandular a 1 setae.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFACEC38FE3BFD13FED5FE53.taxon	discussion	Remarks The study of four specimens, a juvenile and 3 females, collected in three caves (see Table S 2 in Supplementary Material) allowed a comparison with the species description, but new topotype material is needed to clarify the taxonomic status of this species (Table 1). The epicuticle of dorsal sclerites is reticulate when observed in high magnification (Fig. 6). Antennae with 34 and 35 antennomeres in two adults, and 40 in the juvenile; adult medial antennomeres 1.6 × as long as wide, apical antennomere 2.5 × as long as wide; one distal whorl with 20 – 22 gouge sensilla in each medial and distal antennomere, 36 – 38 µm in length (Fig. 4). Cupuliform organ occupies 1 / 5 of total length of last antennomere, with more than ten spheroidal olfactory chemoreceptors with a central column unfolded in complex expansions with a visible polygonal net with porous surface (Fig. 3). Protruding frontal process with abundant tubercular setae (Fig. 5). Pronotum with 3 + 3 la macrosetae (2 + 2 la in original description). Unequal claws (1.4 – 1.5) with large lateral crests (Fig, 7), posterior claw with a large backward overhang; short lateral anterior pretarsal processes, less than half length of claw, and long lateral posterior pretarsal anterior process overpassing end of claw, as described in type form. Calcars with two rows of long barbs almost from base; smooth subapical tarsal setae. Urotergites with important differences from type form: 2 + 2 (3 + 2) la and 4 + 4 – 5 + 5 post on urotergites VI – VII; 6 + 6 post on urotergite VII; 9 + 9 post macrosetae on abdominal segment IX (undescribed in type form). Stylar setae with a few long, thin barbs. Urosternite I with 7 + 7, urosternites II – VII with 6 + 6, urosternite VIII with 2 + 2 macrosetae (Fig. 8). Cerci 1.9 × as long as body, with only 6 articles plus base.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFACEC38FE3BFD13FED5FE53.taxon	distribution	Habitat and distribution Subterranean species found in four caves, including the type locality in the I-El Koba (= Kizil-Koba, Krasnaya) Сave in the Crimean Mountains. The occurrence of such a distribution for P. (D.) evallonychia in the currently unrelated isolated karst regions of the Crimean Mountains is probably associated with the recent colonization of the caves by the edaphic form of this species in the Würm glacial stage. Such a recent isolation of the populations of this species might explain the absence of differences between them.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFAEEC39FE03FD99FB63FADF.taxon	diagnosis	Diagnosis (Sendra & Weber 2018) Troglomorphic species. Sparse thin and long clothing notal setae with thin barbs. Antennae in adults with 32 – 33 antennomeres. Cupuliform organ with 12 – 14 complex olfactory chemoreceptors. Frontal process developed, with slightly tubercular setae and macrosetae. Pronotum with 1 + 1 ma, 4 + 4 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp, 1 + 1 mp macrosetae; metanotum with 1 + 1 ma, 1 + 1 la, 2 + 2 lp, 1 + 1 mp macrosetae; long, thin macrosetae, with very thin barbs; subequal to slightly unequal claws (posterior claw 1.05 – 1.15) with large lateral crests; posterior claw with backward overhang; pretarsal lateral process long, smooth and setiform with several short proximal barbs. Urotergites I – II with 1 + 1 post 1 macrosetae; urotergite III with 1 + 1 la, 1 + 1 post macrosetae; urotergite IV with 1 + 1 la, 2 + 2 (3 + 2, 3 + 3) post macrosetae; urotergites V – VII with 1 + 1 la, 5 + 5 post macrosetae; urotergite VIII with 6 + 6 (5 + 6, 5 + 5) post macrosetae; abdominal segment IX with 8 + 8 (8 + 7) post macrosetae. Urosternite I with 7 + 7 (7 + 6), urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae. Male with a narrow area with glandular g 1 setae, enlarged subcylindrical appendages with glandular a 1 setae; female with glandular g 1 setae with subcylindrical appendages, thinner than in male. Habitat and distribution Monotypic subgenus known only from a single cave in southwest Germany (Fig. 211).	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFAEEC39FE67FEB3FC7EFE26.taxon	diagnosis	Diagnosis (Sendra & Weber 2018) Mesothoracic and metathoracic femora with five dorsal macrosetae, up to three on prothoracic femur; presence of a narrow area of glandular g 1 setae in adult females.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFAEEC39FD95FA92FA83F8E2.taxon	diagnosis	Diagnosis (Silvestri 1912; Condé 1955 a; Sendra & Weber 2018) Lateral crests on claws from small to large, with subequal to very unequal, elbow-like claws; one dorsal femoral; from one to four ventral tibial macrosetae; setifom and smooth (in several species with a few proximal barbs) pretarsal lateral processes. Sternite I with 6 + 6 – 9 + 9, sternites II – VII with 4 + 4 – 7 + 7 and sternite VIII with 2 + 2 macrosetae (with the exception of P. (P.) dargilani). Cerci elongated, with long articles covered in long, thin macrosetae with thin barbs and thin, smooth clothing setae. Habitat and distribution Within humid soils and MSS, but with a preference for the deep subterranean spaces found in caves, including the deepest caves in the world. The 60 species and 9 subspecies of this subgenus are wellspread around the Euro-Mediterranean region, from Southern Europe, including a small karst area in North Africa (Kabylian Mountains, Tell Atlas), throughout south of Central to Southeastern Europe and also reaching some spots in the Anatolian and Crimean peninsulas and Western Caucasus (Fig. 211).	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFAEEC36FE71F8DDFAC9FD02.taxon	diagnosis	Diagnosis (Condé 1947 a) Non-troglomorphic species. Antennae with 23 – 26 antennomeres. Pronotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp macrosetae; metanotum with 1 + 1 ma, 1 + 1 la, 2 + 2 lp macrosetae; long, barbed notal macrosetae; smooth or distally barbed notal clothing setae. Two ventral tibial macrosetae; unequal claws (1.30) with large lateral crests, posterior claw with backward overhang. Urotergites I – II with 1 + 1 post macrosetae; urotergite III with 1 + 1 la, 1 + 1 post macrosetae; urotergite IV with 1 + 1 la, 2 + 2 post macrosetae; urotergites V – VII with 1 + 1 la, 4 + 4 post macrosetae; urosternite I with 7 + 7, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae. Male with an area of glandular g 1 setae, subcylindrical appendages with glandular a 1 setae; female appendages like male appendages. Habitat and distribution This species is only known from Županska pećina, Lubnice, Berane Municipality, Montenegro.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFA1EC35FEF2FCBCFBC1FE95.taxon	description	Figs 9 – 11; Table 2	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFA1EC35FEF2FCBCFBC1FE95.taxon	etymology	Etymology The specific name ʻapolloʼ is proposed since the Parnassos Mountain, its type locality, is the home of the divinity ʻApolloʼ.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFA1EC35FEF2FCBCFBC1FE95.taxon	materials_examined	Material examined Holotype GREECE • 1 ♀; Viotìa, O. Parnassós, crevices; 1660 m a. s. l.; 20 Jun. 2004 – 9 Jun. 2005; Pier Mauro Giachino and Dante Vailati leg.; MSS habitat; pitfall trap; MZB 2019 - 1032. Paratypes GREECE • 4 ♀♀; same collection data as for holotype; Coll. AS • 9 ♂♂, 18 ♀♀; Fokída, Ori Vardoússia; 1575 m a. s. l.; 10 Jun. 2005 – 8 Jun. 2006; Pier Mauro Giachino and Dante Vailati leg.; MSS habitat; pitfall trap; Coll. AS.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFA1EC35FEF2FCBCFBC1FE95.taxon	description	Description BODY. Body length 4.5 to 6.1 mm (females) and 5.0 to 6.4 mm (males). Epicuticle smooth under optical microscope; body with thin, middle-sized clothing, covered or not by a few thin barbs. HEAD. Two intact antennae in a 5.6 mm long female paratype with 28 antennomeres and a 5.1 mm long female paratype with 32 antennomeres, in both shorter than body length (Table 2). Small subcylindrical sensillum of third antennomere located in ventral position between c and d macrosetae. Central antennomeres 1.8 × as long as wide, apical antennomere 2.0 × as long as wide. Cupuliform organ occupying ¼ of total length of apical antennomere, with 8 – 9 complex olfactory chemoreceptors. Gouge sensilla 30 – 38 µm long, in a single distal whorl of 16 – 18 sensilla on each medial and distal antennomere. Frontal process plain, with non-tubercular setae and macrosetae with a few barbs along distal half; macrosetae along line of insertion of antennomere and x setae longer than other macrosetae (a / i / p / x with relative lengths of 40 / 48 / 37 / 68 in holotype). Suboval labial palps with a small, coniform lateroexternal sensillum, with two guard setae, up to 9 setae on anterior border and up to 190 neuroglandular setae. THORAX. Thoracic macrosetal distribution (Fig. 9): pronotum with 1 + 1 ma, 2 + 2 la 3,4 and 3 + 3 sla, 2 + 2 lp 2,3 (additionally, 1 + 1 extra s ma appear near sagittal plane); mesonotum with 1 + 1 ma, 3 + 3 la 1,2,3, 2 + 2 lp 2,3, 1 + 1 mp; metanotum with 1 + 1 ma, 2 + 2 lp 2,3, 1 + 1 mp. All notal macrosetae long and robust, covered by thin barbs almost from their base; submacrosetae sla and sma thinner and shorter than notal macrosetae, but with same large base; marginal setae longer than clothing setae, covered by thin barbs along distal half. Legs elongated, pretarsus of metathoracic legs slightly overpassing end of abdomen (Table 2). Femur I without dorsal macrosetae, but with one long, barbed ventral macroseta; femora II – III with one long, barbed dorsal and one long, barbed ventral macroseta. Tibia I with one short, barbed ventral macroseta; tibiae II – III with two short, barbed ventral macrosetae. Calcars with 2 – 3 rows of long barbs from base to tip, smooth on one narrow side along calcar. Dorsal and lateral subapical tarsal setae smooth. Subequal claws (posterior claw 1.1 × as long as anterior one); posterior claw without backward overhang; lateral crests well developed. Pretarsal process long and setiform, reaching extremities of claws. ABDOMEN. Distribution of abdominal macrosetae on tergites (Fig. 10): 1 + 1 post 1 on I – III; 1 + 1 la, 3 + 3 (also 2 + 2, but less frequent) post 1,2,3 on IV; 1 + 1 la, 6 + 6 post 1 – 5 on V – VII; 8 + 8 post on VIII; 10 + 10 post on abdominal segment IX. All post urotergal macrosetae thick, long and covered by thin barbs along distal four-fifths; la urotergal macrosetae shorter than post macrosetae, covered by barbs along distal half. Urosternite I with 7 + 7 macrosetae; urosternites II – VII with 5 + 5 macrosetae; urosternite VIII with 2 + 2 macrosetae; all urosternal macrosetae robust and large, covered by long barbs along distal threefourths to four-fifths. Apical, subapical and ventromedial setae with abundant long barbs. SECONDARY SEX CHARACTERS. Male urosternite I (Fig. 11) with up to 180 glandular g 1 setae arranged in up to five rows, with enlarged subcylindrical appendages each bearing up to 52 glandular a 1 setae. Female appendages slightly thinner, with up to 38 glandular a 1 setae. Phyletic affinities, habitat and distribution The modest subequal claws, including the posterior claw lacking a backward overhang, and the absence of lateral anterior macrosetae on the metanotum relate P. (P.) apollo sp. nov. to P. (P.) festae. Nonetheless the more abundant posterior urotergal macrosetae, with 6 + 6 post on urotergites IV – VII, as well as 8 + 8 and 10 + 10 post on abdominal segments VIII – IX in P. (P.) apollo sp. nov., rather than 5 + 5 on urotergites IV – VII, and 7 + 7 and 9 + 9 on abdominal segments VIII – IX in P. (P.) festae (Silvestri 1933 a), separate the two species. Interestingly, the new species was collected in traps placed in MSS over the forest line (type locality) or in Abies Mill. forest (all the other sites), at altitudes from 1075 to 1660 m a. s. l. This large range of hypogean habitats also occurs in P. (P.) festae (see below).	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFA2EC35FE17FED4FB53FA42.taxon	diagnosis	Diagnosis (Bareth & Condé 2001) Non-troglomorphic species. Antennae with 23 and 26 antennomeres; five simple olfactory chemoreceptors in cupuliform organ; slightly protruding frontal process with slightly tubercular setae. Pronotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp macrosetae; metanotum with 1 + 1 ma, 1 + 1 (2 + 1) la, 2 + 2 lp macrosetae; long, barbed notal macrosetae; abundant barbed notal clothing setae. Two ventral tibial macrosetae; subequal claws (1.05 – 1.1) with large lateral crests, posterior claw with a small backward overhang. Urotergites I – II with 1 + 1 post macrosetae; urotergite III with 2 + 2 post macrosetae; urotergites IV – V with 1 + 1 la, 5 + 5 post macrosetae; urotergites VI – VII with 2 + 2 (2 + 1) la, 5 + 5 post macrosetae; urotergite VIII with 6 + 6 post macrosetae; abdominal segment IX with 8 + 8 post macrosetae; urosternite I with 7 + 7, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae. Male with glandular g 1 setae, subcylindrical appendages with glandular a 1 setae; female appendages similar to those of male.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFA2EC35FE17FED4FB53FA42.taxon	discussion	Remarks The studied material allowed us to complete the above description. The complete antennae of a female from Jama 1 v Kanjaducah show 23 antennomeres with five simple olfactory chemoreceptors within the cupuliform organ; it has a distal whorl of 8 – 11 gouge sensilla that are 25 µm long. All specimens show a simple frontal process with non-tubercular setae; smooth subapical tarsal setae; abdominal segment IX with 8 + 8 post macrosetae; urosternite I of male with a narrow area of up to 80 glandular g 1 setae arranged in two or three rows; subcylindrical appendages with up to 12 glandular a 1 setae; females with subcylindrical appendages as in male, with up to 9 glandular a 1 setae. Nevertheless, the large distance from the type locality casts serious doubts regarding the correct identification of this rare species. Habitat and distribution Hypogean species found in two geographically distant karst regions, the Lyashkata Peshtera Cave, Balkan Mountains in Bulgaria, and two caves in the Slovenian Dinaric Mountains.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFA2EC30FE76FA79FD04FEAA.taxon	description	Figs 12 – 23	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFA2EC30FE76FA79FD04FEAA.taxon	diagnosis	Diagnosis (Sendra et al. 2004) Troglomorphic species. Antennae with 39 antennomeres; 12 very complex olfactory chemoreceptors in a large cupuliform organ; protruding frontal process with tubercular setae. Pronotum with 1 + 1 ma, 2 + 2 la, 2 + 2 lp macrosetae; mesonotum 1 + 1 with ma, 2 + 2 lp macrosetae; metanotum without macrosetae; short, barbed notal macrosetae except pronotal lp; sparse smooth clothing setae. Two ventral tibial macrosetae; slightly unequal claws (1.16) with large lateral crests, posterior claw with a short backward overhang. Urotergites V – VI with 1 + 1 la, 1 + 1 post macrosetae; urotergite VII with 1 + 1 la, 2 + 2 – 3 + 3 post macrosetae; urotergite VIII with 5 + 5 – 6 + 6 post macrosetae; abdominal segment IX with 8 + 8 post macrosetae; urosternite I with 8 + 8, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae. Male with a narrow area of glandular g 1 setae, with subcylindrical appendages similar to those of female, with glandular a 1 setae.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFA2EC30FE76FA79FD04FEAA.taxon	discussion	Remarks SEM observations have shown several taxonomical features to complete the original description (Sendra et al. 2004). Each olfactory chemoreceptor shows a spiral fold with a central cylindrical structure, all with a multiporous surface, all tightly open to the exterior through a small opening (Figs 12, 14 – 15), surrounded by tiny coniform sensilla (Figs 12 – 13) that are present in the whorl of thin gouge sensilla on each medial and distal antennomere (Figs 16 – 17). Tibia longer than femur and tarsus (Fig. 18). Pretarsus with subequal claws with large lateral crests, the claws carved by longitudinal ridges and a micro-tuberculated surface (Figs 20, 22 – 23). Lateral process smooth, with several thin basal barbs (Figs 22 – 23). Stylar setae with abundant barbs (Fig. 21). Male with subcylindrical appendages with only glandular a 1 setae and a narrow area of glandular g 1 seate (Fig. 19).	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFA2EC30FE76FA79FD04FEAA.taxon	distribution	Habitat and distribution Subterranean species known only from Nerja Cave, a touristic cave in the internal zone of the Baetic Mountain Range, southern Iberian Peninsula.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFA7EC30FE74FE26FBFFFB40.taxon	diagnosis	Diagnosis (Condé 1947 b, 1948 b; Bareth 2006) Troglomorphic species. Antennae with 30 – 40 antennomeres; 12 complex olfactory chemoreceptors in cupuliform organ; protruding frontal process with tubercular setae. Pronotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 2 + 2 la, 2 + 2 lp macrosetae; metanotum 1 + 1 ma, 1 + 1 – 2 + 2 la, 2 + 2 lp macrosetae; large, long, barbed notal macrosetae; scarce barbed clothing setae. One or two ventral tibial macrosetae; unequal claws with large lateral crests, posterior claw with a backward overhang. Urotergites I – II with 1 + 1 post macrosetae; urotergite III with 0 + 0 – 1 + 1 la, 1 + 1 post macrosetae; urotergite IV with 1 + 1 la, 2 + 2 post macrosetae; urotergites V – VII with 1 + 1 la, 4 + 4 post macrosetae; urotergite VIII with 6 + 6 post macrosetae; abdominal segment IX with 8 + 8 post macrosetae; urosternite I with 7 + 7, urosternites II – VII with 6 + 6, urosternite VIII with 2 + 2 macrosetae. Male with a narrow area of glandular g 1 setae, with subtrapezoidal appendages with glandular a 2 setae; female with subcylindrical appendages with glandular a 1 setae. Spermatozoid fascicles with a diameter of 55 – 60 µm and a width of 15 µm, its filament wheels with 2 ½ turns of 460 µm long and 12 – 13 µm diameter filament. Habitat and distribution Subterranean species found in 49 caves around the Central Massif, France.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFA7EC30FE3BFB7AFE14F838.taxon	diagnosis	Diagnosis (Bareth & Condé 2001) Slightly troglomorphic species. Antennae with 26 – 29 antennomeres; 5 – 6 complex olfactory chemoreceptors on cupuliform organ; non-protruding frontal process with non-tubercular setae. Pronotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp, 1 + 1 mp macrosetae; metanotum with 1 + 1 ma, 1 + 1 (2 + 1) la, 2 + 2 lp, 1 + 1 mp macrosetae; long, barbed notal macrosetae; abundant notal clothing setae. Two ventral tibial macrosetae; unequal claws (1.30) with large lateral crests, posterior claw with a backward overhang. Urotergites I – II with 1 + 1 post macrosetae; urotergite III with 2 + 2 post macrosetae (1 + 1 la, 1 + 1 post on one specimen from Novata Peştera, western Rhodope); urotergite IV with 1 + 1 la, 4 + 4 – 5 + 5 post macrosetae; urotergite V with 1 + 1 – 2 + 2 la, 5 + 5 post macrosetae; urotergites VI – VII with 2 + 2 – 3 + 3 la, 5 + 5 post macrosetae; urotergite VIII with 6 + 6 post macrosetae; abdominal segment IX with 8 + 8 post macrosetae; urosternite I with 7 + 7, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae. Male with an area of glandular g 1 setae, subcylindrical appendages with glandular a 1 setae (adults from Gruffre Padez, Sofia with short, thick appendages with a large area of glandular a 1 setae); female with subcylindrical appendages with glandular a 1 setae. Spermatozoid fascicle wheels with 2 turns of 50 µm long and 5 µm diameter filament. Habitat and distribution Subterranean species found in six caves in different karst regions of the Bulgarian Balkan Mountains (= Stara Planina Mts).	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFA6EC31FE1FFEB3FAF8FCBA.taxon	diagnosis	Diagnosis (Condé 1948 c, 1949, 1950; Condé & Poivre 1982; Bareth 2006) Troglomorphic species. Antennae with 31 – 37 antennomeres; 7 – 8 complex olfactory chemoreceptors in cupuliform organ. Pronotum with 1 + 1 ma, 4 + 4 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp, 1 + 1 mp macrosetae; metanotum with 1 + 1 ma, 1 + 1 la, 2 + 2 lp, 1 + 1 mp macrosetae; long, barbed notal macrosetae; sparse notal clothing seate with a few distal barbs. Two or three ventral tibial macrosetae; subequal claws (1.05) with large lateral crests, posterior claw with a small backward overhang. Urotergites I – III with 1 + 1 post macrosetae; urotergites IV – VII with 1 + 1 la, 5 + 5 post macrosetae; urotergite VIII with 6 + 6 post macrosetae; abdominal segment IX with 8 + 8 post macrosetae; urosternite I with 7 + 7, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae. Male with abundant glandular g 1 setae, subcylindrical appendages like female appendages, with glandular a 1 setae. Habitat and distribution Subterranean species found in 11 caves and at one edaphic site in the Île Saint-Honorat caves around the Western Alps, in the mountain reliefs near the Mediterranean coast in the Provence Alps, France.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFA6EC31FE33FC34FD71FACF.taxon	diagnosis	Diagnosis differential against the type form (Condé 1961; Condé & Poivre 1982) Troglomorphic subspecies. Antennae with 27 – 33 antennomeres. Urotergite III with 2 + 2 post macrosetae. No adult males have been observed, and the young males from Valais do not have glandular g 1 setae, without glandular a 2 setae. Habitat and distribution Subterranean subspecies found in a cave of Valais (Switzerland) as well as in two caves in Brescia and Toscana (Italy), around the Western Alps.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFA6EC31FEFEFA82FC36F9A7.taxon	diagnosis	Differential diagnosis against the type form (Condé & Poivre 1982) Troglomorphic subspecies. Antennae with 31 – 36 antennomeres; urosternite VIII with 7 + 7 post macrosetae. Habitat and distribution Subterranean subspecies found in 13 caves around the Western Alps.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFA6EC0FFE39F91AFAA3FE2F.taxon	description	Figs 24 – 29	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFA6EC0FFE39F91AFAA3FE2F.taxon	diagnosis	Diagnosis (Condé 1948 c; Sendra 1985; Bareth & Condé 1996; Sendra & Escolà 2004; Bareth 2006) Troglomorphic species. Antennae with 28 – 40 antennomeres; nine to ten complex olfactory chemoreceptors in cupuliform organ; protruding frontal process with tubercular setae. Pronotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp macrosetae; metanotum with 1 + 1 ma, 2 + 2 la, 2 + 2 lp macrosetae; long, barbed notal macrosetae; abundant notal clothing setae with distal barbs. Two ventral tibial macrosetae; slightly unequal claws (1.1 – 1.2) with large lateral crests, posterior claw with a backward overhang. Urotergites I – III with 1 + 1 post macrosetae; urotergite IV with 1 + 1 la, 5 + 5 post macrosetae; urotergite V with 1 + 1 – 2 + 2 la, 5 + 5 post macrosetae; urotergites VI – VII with 2 + 2 la, 5 + 5 post macrosetae; urotergite VIII with 6 + 6 post macrosetae; abdominal segment IX with 8 + 8 post macrosetae; urosternite I with 7 + 7, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae. Male with a large area of glandular g 1 setae, subtrapezoidal appendages with glandular a 2 setae; female with subcylindrical appendages with glandular a 1 setae. Habitat and distribution Subterranean species found in three caves in the southern reliefs of France, west of the Rhône River, and in 34 caves distributed from the Central Pyrenees to the Pre-Pyrenees without reaching the Ebro Basin.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF98EC0FFE0EFDA3FD92FA3B.taxon	description	Figs 30 – 41	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF98EC0FFE0EFDA3FD92FA3B.taxon	diagnosis	Differential diagnosis against the type form (Sendra 1985; Sendra & Escolà 2004) Antennae with 30 – 43 antennomeres; eight to nine complex olfactory chemoreceptors in cupuliform organ; protruding frontal process with slightly tubercular setae. Spermatozoid fascicle wheels with 2 – 2 ½ turns of 300 – 400 µm long and 7 µm diameter filament.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF98EC0FFE0EFDA3FD92FA3B.taxon	discussion	Remarks (Figs 24 – 35) One male of P. (P.) bonneti bonneti from Cova Mora (San Juan de la Peña, Huesca, Spain) and six specimens of P. (P.) bonneti condei from Cova Toll (Moià, Barcelona, Spain) (see Table S 2 in Supplementary Material) were used for SEM to remark on some features that are difficult to observe under the optical microscope. In P. (P.) bonneti bonneti, the first urosternite shows very long subtrapezoidal appendages with a large area of glandular a 2 setae and a large area of glandular g 1 setae, with the surface covered by barbed clothing setae and well-barbed macrosetae (Figs 24 – 27). In P. (P.) bonneti condei, the epicuticle varies from completely smooth to gently reticulate (Fig. 35); antennomeres with a whorl of gouge sensilla, 16 – 18 µm long (Figs 31, 33). Olfactory chemoreceptors with a central column unfolded in several layers with a reticulate surface covered by pores (Fig. 30). Protruding frontal process with slightly tubercular macrosetae and setae (Figs 32, 34). Claws with large lateral crest with ventral longitudinal grooves separated by reticulate surfaces (Figs 36 – 39). Stylar setae totally covered by short, thin barbs (Fig. 40). Large eversible vesicles (Fig. 41). Habitat and distribution Subterranean subspecies found in 35 caves in the Eastern Pyrenees and the northern part of the Catalan Mediterranean Mountain System.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF98EC0FFEE9F9B6FB1AF83F.taxon	diagnosis	Differential diagnosis against the type form (Bareth & Condé 1996) Antennae with 34 – 38 antennomeres; six to seven simple olfactory chemoreceptors in cupuliform organ; slight protruding frontal process with a few anterior tubercular setae; abundant barbed notal clothing setae. One ventral tibial macroseta. Unequal claws with large lateral crests, posterior claw with a backwards overhang. Urotergites IV – VII with 1 + 1 la, 5 + 5 post macrosetae. Male with an area of glandular g 1 setae; subtriangular appendages with glandular a 2 setae. Spermatozoid fascicle wheels with 1 ½ – 2 turns of 8 µm diameter filament. Habitat and distribution Subterranean subspecies found in two caves in the northern part of the Eastern Pyrenees.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF9BEC0AFE43FEB3FDFCFEAB.taxon	diagnosis	Diagnosis (Loksa 1960) Non-troglomorphic species. Antennae with 19 – 20 doliiform antennomeres. Pronotum with 1 + 1 ma, 2 + 2 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp macrosetae; metanotum with 1 + 1 ma, 1 + 1 la, 2 + 2 lp macrosetae; thin notal macrosetae; sparse notal clothing setae. Two ventral tibial macrosetae. Urotergites I – II with 1 + 1 post macrosetae; urotergite III with 3 + 3 post macrosetae; urotergites IV – VII with 4 + 4 post macrosetae; la macrosetae not mentioned in original description. Habitat and distribution Hypogean species, only found in Lóczy Cave, Balatonfüred, in the Balaton Uplands, southern Transdanubian Mountains, Hungary.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF9DEC0AFE58FBB8FDFEFA9C.taxon	diagnosis	Differential diagnosis against the type form (Sendra et al. 2004) Frontal process with tubercular setae and 1 + 1 la macrosetae on urotergites VI – VII. Habitat and distribution Subterranean species inhabiting two caves in Eivissa and four caves in Formentera, two of the Balearic Islands. Plusiocampa (P.) aff. breuili is only known from a cave in the eastern Baetic Mountains, southeastern Iberian Peninsula.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF9DEC0AFE70FE1DFC3FFC0E.taxon	diagnosis	Diagnosis (Condé 1955 b; Condé & Sendra 1989; Sendra et al. 2004) Troglomorphic species. Antennae with 30 and 32 antennomeres; 21 complex olfactory chemoreceptors in cupuliform organ; protruding frontal process with non-tubercular setae. Pronotum with 1 + 1 ma, 2 + 2 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp, 1 + 1 mp macrosetae; metanotum with 1 + 1 ma, 1 + 1 la, 2 + 2 lp, 1 + 1 mp macrosetae; long, barbed notal macrosetae, sparse notal clothing setae. 3 or 4 ventral tibial macrosetae (ratio of femur / tibia 0.66); subequal claws (1.1) with large lateral crests, posterior claw with a small backward overhang. Urotergites I – II with 1 + 1 post macrosetae; urotergite III with 0 + 0 – 1 + 1 la, 1 + 1 post macrosetae; urotergite IV with 2 + 2 (2 + 1) la, 5 + 5 post macrosetae; urotergites V – VII with 2 + 2 la, 5 + 5 post macrosetae; urotergite VIII with 6 + 6 post macrosetae; abdominal segment IX with 8 + 8 post macrosetae; urosternite I with 8 + 8, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae. Male with an area of glandular g 1 setae, subcylindrical appendages with glandular a 1 setae; female with subcylindrical appendages with glandular a 1 setae.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF9DEC0BFE5FFAD7FB07FDD6.taxon	description	Figs 42 – 43	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF9DEC0BFE5FFAD7FB07FDD6.taxon	diagnosis	Diagnosis (Silvestri 1931 b; Bareth & Condé 2001) Troglomorphic species. Antennae with 24 – 30 antennomeres; 10 complex olfactory chemoreceptors in cupuliform organ; protruding frontal process with tubercular setae. Pronotum with 1 + 1 ma, 4 + 4 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp, 1 + 1 mp macrosetae; metanotum with 1 + 1 ma, 2 + 2 la, 2 + 2 lp, 1 + 1 mp macrosetae; long, barbed notal macrosetae; a few long, distally barbed notal clothing setae. Two or three ventral tibial macrosetae; unequal claws (1.30) with large lateral crests, posterior claw with a backward overhang. Urotergites I – II with 1 + 1 post macrosetae; urotergite III with 0 + 0 – 1 + 1 la, 1 + 1 – 2 + 2 post macrosetae; urotergite IV with 1 + 1 la, 5 + 5 post macrosetae; urotergite V with 2 + 2 la, 5 + 5 post macrosetae; urotergites VI – VII with 3 + 3 la, 5 + 5 post macrosetae; urotergite VIII with 6 + 6 post macrosetae; abdominal segment IX with 8 + 8 post macrosetae; urosternite I with 7 + 7, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae. Male with a large area of glandular g 1 setae and slightly enlarged appendages with glandular a 1 setae; female with subcylindrical appendages with glandular a 1 setae.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF9DEC0BFE5FFAD7FB07FDD6.taxon	discussion	Remarks The study of 38 specimens collected from two new caves (see Table S 2 in Supplementary Material) allowed the addition of information to the diagnosis, including the observation of the pretarsal structures (Figs 42 – 43). Habitat and distribution Subterranean species found in 16 caves in the Balkan and Rhodope Mountains, Bulgaria.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF9CEC08FE71FD89FC35FEAA.taxon	diagnosis	Diagnosis (Condé 1950, 1954 a; Christian et al. 1996) Slightly troglomorphic species. Antennae with 28 – 30 antennomeres. Pronotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp, 1 + 1 mp macrosetae; metanotum with 1 + 1 ma, 1 + 1 la, 2 + 2 lp, 1 + 1 mp macrosetae; long, barbed notal macrosetae; abundant notal clothing setae, either smooth or with a few thin barbs. Three ventral tibial macrosetae; unequal claws with large lateral crests, posterior claw with a backward overhang. Urotergites I – II with 1 + 1 post macrosetae; urotergite III with 2 + 2 post macrosetae; urotergite IV with 1 + 1 la, 5 + 5 (4 + 4) post macrosetae; urotergite V with 2 + 2 (1 + 1) la, 5 + 5 post macrosetae; urotergites VI – VII with 2 + 2 la, 5 + 5 post macrosetae; urotergite VIII with 6 + 6 post macrosetae; abdominal segment IX with 8 + 8 post macrosetae. Urosternite I with 7 + 7, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae. Male without glandular g 1 setae, slightly enlarged subcylindrical appendages with a 1 glandular setae; female with subcylindrical appendages with a 1 glandular setae.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF9CEC08FE71FD89FC35FEAA.taxon	discussion	Remarks One male and one female of P. (P.) caprai from Babja jama (Slovenia) have been studied. However, two females collected from an MSS habitat in Corinthia (Peloponnese Peninsula) (see Table S 2 in Supplementary Material) appear to be very closely related to P. (P.) caprai, although the substantial geographical distance between these two areas and the observation of only females cast doubts on this specific attribution. Furthermore, all antennae and cerci are broken. The frontal process shows a protrusion and bears tubercular setae. Habitat and distribution Hypogean species inhabiting subterranean and soil habitats, found in six caves, one mine and two soil and MSS habitats in the Austrian, Italian, Slovenian and Swiss Alps.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF9FEC06FE0BFE15FC4DF80A.taxon	description	Figs 44 – 46; Tables 3 – 4	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF9FEC06FE0BFE15FC4DF80A.taxon	etymology	Etymology The name ʻchiosensisʼ is proposed based on the island of Chios, where it occurs in its deep subterranean habitat.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF9FEC06FE0BFE15FC4DF80A.taxon	materials_examined	Material examined Holotype GREECE • ♀; Chios Island, Sykias; 10 Jun. 2006; Fulvio Gasparo leg.; MZB 2019 - 1030. Paratypes GREECE • 2 ♂♂; same collection data as for holotype; Coll. AS.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF9FEC06FE0BFE15FC4DF80A.taxon	description	Description BODY. Body length 4.4 and 4.9 mm (males, paratypes) and 7.8 mm (female, holotype). Epicuticle smooth under optical microscope; body with sparse thin, long clothing, covered or not by a few thin distal barbs. HEAD. Holotype with two intact antennae of 54 antennomeres, 14.8 mm long. Small coniform sensillum of third antennomere located in ventral position between c and d macrosetae. Central antennomeres 3.8 × as long as wide, apical anntenomere 3.6 × as long as wide. Small cupuliform organ occupying 1 / 9 of total length of apical antennomere, with 6 – 7 complex olfactory chemoreceptors. Thin and long gouge sensilla (44 – 47 µm long) in a single distal whorl of 10 – 13 sensilla on each medial and distal antennomere. Frontal process developed, with tubercular setae and macrosetae with thin barbs. Three macrosetae along line of insertion of antennae poorly differentiated, only anterior one with 1 – 2 thin apical barbs; x setae longer than these macrosetae, with a couple of thin apical barbs (Fig. 44). Suboval labial palps with latero-external sensillum larger than sensillum of third antennomere, with two guard setae, up to 12 setae on anterior border and up to 130 neuroglandular setae. THORAX. Distribution of thoracic macrosetae: pronotum with 1 + 1 ma, 4 + 4 – 3 + 3 la 1,2,3,4, 2 + 2 lp 2,3 (additionally in the three type specimens, 1 + 1 extra ma near the sagittal plane); mesonotum with 1 + 1 ma, 1 + 1 (0 + 1) la 2, 2 + 2 lp 2, 3, 1 + 1 mp; metanotum with 1 + 1 ma, 1 + 1 lp 2 and 1 + 1, 0 + 1 or 0 + 0 mp. All notal macrosetae long, thin, covered by very thin barbs along distal third up to half; marginal setae longer than clothing setae, covered by thin barbs along distal half up to two thirds; spare clothing setae, with 14, 21 and 23 clothing setae on each heminotum of pronotum, mesonotum and metanotum, respectively (holotype). Legs elongated, metathoracic legs overpassing end of abdomen in middle of tibia; measurements as in Table 3. Femora II – III with one long dorsal macroseta and one longer ventral macroseta. Tibiae I – III with two ventral macrosetae surrounded all along by thin, short barbs. Calcars surrounded from base to tip by thin, short barbs. Dorsal and lateral subapical tarsal setae completely covered by thin barbs almost from base and almost to tip. Subequal claws (posterior claw 1.0 – 1.05 × as long as anterior claw); posterior claw with a small backward overhang; lateral crests well developed, noticeably ridged on ventral side. Pretarsal lateral process setiform, with plenty of very short proximal barbs. ABDOMEN. Distribution of abdominal macrosetae on tergites: 1 + 1 post 1 on I – III; 1 + 1 la, 2 + 2 lp 1,2 on IV; 1 + 1 la, 5 + 5 post 1 – 5 on V – VII, 8 + 8 post on VIII; 11 + 11 (11 + 12) post on abdominal segment IX. All post urotergal macrosetae thick, long, covered by thin barbs along distal four-fifths; la urotergal macrosetae shorter than post macrosetae, covered by barbs along distal half. Urosternite I with 7 + 7, urosternites IIVII with 5 + 5, urosternite VIII with 2 + 2 macrosetae; all urosternal macrosetae robust and large, covered by long barbs along distal half up to four-fifths. Apical subapical and ventro-medial setae completely surrounded by short abundant barbs. CERCI. One complete cercus in holotype with eight articles in addition to basal article, 19.70 mm long (Table 4), 2.53 mm longer than body length; articles show whorls of long macrosetae covered by thin barbs along distal two-thirds, combined with whorls of smooth, thin setae shorter than macrosetae. SECONDARY SEX CHARACTERS. Female urosternite I (Fig. 45) with enlarged subcylindrical appendages, each bearing up to 16 glandular a 1 setae in a distal area. Male urosternite I (Fig. 46) without glandular g 1 setae, with moderated large subcylindrical appendages thicker than those of female, each bearing up to 33 glandular a 1 setae. Phyletic affinities, habitat and distribution Although P. (P.) chiosensis sp. nov. has medial posterior macrosetae on the mesonotum and metanotum (sometimes absent in one or two heminota), as do many previously described species of Plusiocampa s. str., a similar reduction in the number of lateral anterior and lateral posterior macrosetae on the mesonotum and metanotum is present in another troglomorphic species, P. (P.) glagra from continental Greece (Condé 1984 a). Plusiocampa (P.) chiosensis sp. nov. also shares a couple of noticeable taxonomical features with P. (P.) glabra in their subequal claws and the number of urosternal macrosetae. Nevertheless, the presence of 1 + 1 lateral anterior macrosetae on the mesonotum, the more abundant macrosetae on all urotergites, including the eighth abdominal segment, and the absence of glandular g 1 setae in males in P. (P.) chiosensis sp. nov. distinguish it from P. (P.) glabra. Plusiocampa (P.) chiosensis sp. nov. is only known from one cave and occupies the deep subterranean spaces, showing highly troglomorphic features (elongation of appendages, antennomeres and articles, and the strengthening of sensorial equipment of the antennae).	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF90EC07FE12FC3FFB93FB3A.taxon	diagnosis	Differential diagnosis against the type form (Condé 1984 a) Antennae with 22 antennomeres; urotergite III with 1 + 1 post macrosetae, instead of 2 + 2; urotergite IV with 2 + 2 post macrosetae, instead of 3 + 3; urotergites V – VII with 3 + 3 post macrosetae, instead of 4 + 4. Habitat and distribution Soil-dwelling subspecies found only on Ithaca and other large Ionian islands.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF90EC07FE1EFEB3FAD2FCBC.taxon	diagnosis	Diagnosis (Silvestri 1912; Condé 1947 a, 1984 a) Non-troglomorphic species. Antennae with 18 – 21 antennomeres. Pronotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp macrosetae; metanotum with 1 + 1 ma, 1 + 1 la, 2 + 2 lp macrosetae; long, barbed notal macrosetae. Two ventral tibial macrosetae; small subequal claws with small lateral crests, posterior claw with no backward overhang. Urotergites I – II with 1 + 1 post macrosetae; urotergite III with 2 + 2 post macrosetae; urotergite IV with 1 + 1 la, 3 + 3 post macrosetae; urotergites V – VII with 1 + 1 la, 4 + 4 post macrosetae; urotergite VIII with 6 + 6 post macrosetae; abdominal segment IX with 8 + 8 post macrosetae; urosternite I with 7 + 7, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae. Male unknown; female with enlarged appendages with glandular a 1 setae. Habitat and distribution Soil-dwelling species found at two localities on Kerkyra (Corfu), one of the main Ionian islands, and at three localities in continental Greece, in the Epirus region in the east of the Balkan Peninsula.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF90EC07FE62FAB5FC86F9B0.taxon	diagnosis	Differential diagnosis against P. (P.) corcyraea corcyraea (Condé 1953 a) Non-troglomorphic species. Antennae with 20 – 21 antennomeres. Mesonotum with 2 + 2 la macrosetae, instead of 3 + 3; metanotum without la macrosetae, instead of 1 + 1 la. Habitat and distribution Soil-dwelling species found at two localities on Corsica.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF90EC04FE47F92BFD03FDB4.taxon	diagnosis	Diagnosis (Bareth & Condé 1984) Troglomorphic species. Antennae with 32 antennomeres; 8 complex olfactory chemoreceptors with an atypical multi-sheet shape in cupuliform organ; protruding frontal process with tubercular setae. Pronotum with 1 + 1 ma, 2 + 2 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 2 + 2 la, 2 + 2 lp macrosetae; metanotum with 1 + 1 ma, 1 + 1 la, 2 + 2 lp macrosetae; long, barbed notal macrosetae; thin, barbed notal clothing setae. Three ventral tibial macrosetae; unequal claws (1.25) with large lateral crests, posterior claw with a backward overhang. Urotergites I – III with 1 + 1 post macrosetae; urotergite IV with 1 + 1 la, 5 + 5 post macrosetae; urotergites V – VII with 2 + 2 la, 5 + 5 post macrosetae; urotergite VIII with 6 + 6 post macrosetae; abdominal segment IX with 8 + 8 post macrosetae; urosternite I with 7 + 7, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae. Male unknown, female with subcylindrical appendages with glandular a 1 setae. Habitat and distribution Subterranean species only known from Grotta di Montenero, in the Gargano Promontory part of the Apulia Carbonate Platform, Italian Peninsula.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF93EC04FE50FD28FD51F951.taxon	diagnosis	Diagnosis (Denis 1923; Condé 1946; Bareth 2006) Troglomorphic species. Antennae with 40 – 48 antennomeres; eight to nine complex olfactory chemoreceptors in cupuliform organ; protruding frontal process with tubercular setae. Pronotum with 1 + 1 ma, 4 + 4 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 2 + 2 la, 2 + 2 lp macrosetae; metanotum with 1 + 1 ma, 2 + 2 la, 2 + 2 – 1 + 1 lp macrosetae; thin, poorly barbed notal macrosetae; sparse long, thin, barbed notal clothing setae. No ventral tibial macrosetae; unequal claws with large lateral crests, posterior claw with a backward overhang. Urotergite III with 1 + 1 la macrosetae; urotergite IV with 1 + 1 la, 1 + 1 post macrosetae; urotergites V – VII with 1 + 1 la, 3 + 3 – 4 + 4 post macrosetae; urotergite VIII with 6 + 6 post macrosetae; abdominal segment IX with 8 + 8 post macrosetae; urosternite I with 11 + 11, urosternites II – VII with 9 + 9, urosternite VIII with 3 + 3 macrosetae. Male with a narrow area of glandular g 1 setae; subtrapezoidal appendages with an area of glandular a 2 setae; female with subcylindrical appendages with glandular a 1 setae. Spermatozoid fascicles 60 – 65 µm in diameter and 25 – 30 µm wide, wheels with 2 ½ – 3 turns of 650 µm long and 13 µm diameter filament.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF93EC04FE50FD28FD51F951.taxon	discussion	Remarks The study of one female from Puech Cave, Grands Causses, France (see Table S 2 in Supplementary Material) has allowed us to supplement the former description, including regarding the presence of only 1 + 1 lp metanotal macrosetae. Habitat and distribution Subterranean species, well-spread around the Grands Causses, where it is found in eight caves, living alongside the troglobite P. (P.) balsani.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF93EC05FE5BF90CFC94FDB4.taxon	diagnosis	Diagnosis (Condé 1984 b) Troglomorphic species. Antennae with 34 – 35 and 38 antennomeres; 28 simple olfactory chemoreceptors in cupuliform organ; protruding frontal process with tubercular setae. Pronotum with 1 + 1 ma, 3 + 3 – 2 + 2 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 2 + 2 la, 2 + 2 lp macrosetae; metanotum with 1 + 1 ma, 1 + 1 la, 2 + 2 lp macrosetae; long, barbed macrosetae; barbed notal clothing setae. Two or three ventral tibial macrosetae; unequal claws (1.4) with large lateral crests, posterior claw with a backward overhang. Urotergites I – III with 1 + 1 post macrosetae; urotergite IV with 0 + 0 – 1 + 1 la, 2 + 2 – 3 + 3 post macrosetae; urotergites V – VII with 1 + 1 la, 5 + 5 post macrosetae; urotergite VIII with 6 + 6 post macrosetae; abdominal segment IX with 8 + 8 post macrosetae; urosternite I with 7 + 7, urosternites II – VII with 5 + 5 – 6 + 6, urosternite VIII with 2 + 2 macrosetae. Young male with an area of glandular g 1 setae, enlarged appendages with glandular a 1 setae; female with subcylindrical appendages with glandular a 1 setae. Habitat and distribution Subterranean species found in two caves, Ifri Semedane and Ifri Maareb, in the Djurdjura Mountains, Tell Atlas, Algeria, on the former Kabylian microplate.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF92EC05FE7EFD28FBD1FAAE.taxon	diagnosis	Diagnosis (Condé 1993 a; Sendra & Weber 2018) Troglomorphic species. Antennae with 27 – 29 antennomeres; 7 – 9 complex olfactory chemoreceptors on cupuliform organ; protruding frontal process characterized by a frontal and lateral rim with nontubercular setae. Pronotum with 1 + 1 ma, 3 + 3 la, 2 + 2 – 3 + 3 lp macrosetae; mesonotum with 1 + 1 ma, 2 + 2 – 3 + 3 la, 2 + 2 lp macrosetae; metanotum with 1 + 1 ma, 1 + 1 – 2 + 2 (2 + 1) la, 2 + 2 lp macrosetae; long, barbed notal macrosetae; smooth notal clothing setae. Two ventral tibial macrosetae; unequal claws (1.3) with large lateral crests, posterior claw with a backward overhang; lateral processes smooth and short. Urotergites I – II with 1 + 1 post macrosetae; urotergite III with 2 + 2 (2 + 1) post macrosetae; urotergite IV with 0 + 0 – 1 + 1 (0 + 1) la, 2 + 2 (1 + 2) post macrosetae; urotergite V with 1 + 1 (1 + 2) la, 5 + 5 post macrosetae; urotergite VI with 2 + 2 (1 + 2, 2 + 3) la, 5 + 5 post macrosetae; urotergite VII with 2 + 2 (1 + 2) la, 5 + 5 post macrosetae; urotergite VIII with 6 + 6 post macrosetae; abdominal segment IX with 8 + 8 post macrosetae; urosternite I with 7 + 7, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae. Male without glandular g 1 setae; short enlarged subcylindrical appendages with glandular a 1 setae; female with subcylindrical appendages, with glandular a 1 setae. Habitat and distribution Subterranean species found in 13 caves around Swabian Alb, Germany.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF92EC05FE1CFA21FC32F817.taxon	diagnosis	Diagnosis (Bareth & Condé 1984) Troglomorphic species. Frontal process with non-tubercular setae. Pronotum with 1 + 1 ma, 2 + 2 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 1 + 1 la, 2 + 2 lp macrosetae; metanotum with 1 + 1 ma, 2 + 2 lp macrosetae; long, barbed notal macrosetae. One ventral tibial macroseta; subequal claws with small lateral crests, posterior claw with no backward overhang. Urotergites I – IV with 1 + 1 post macrosetae; urotergites V – VII with 4 + 4 post macrosetae; urotergite VIII with 6 + 6 post macrosetae; abdominal segment IX with 8 + 8 post macrosetae; urosternite I with 7 + 7, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae. Adults unknown. Habitat and distribution Description based on a single juvenile from only one cave, Grotta Trojano, Jacotenente, the Gargano Promontory part of the Apulia Carbonate Platform, Italian Peninsula.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF95EC02FE09FEB3FE6FFDB6.taxon	diagnosis	Differential diagnosis against the type form (Bareth 2002) Pronotum with 3 + 3 lateral posterior macrosetae, instead of 2 + 2 in the type form. Habitat and distribution Subterranean subspecies inhabiting one cave, Grotte Scaloria, Manfredonia, Gargano, Italian Peninsula, near the type locality.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF95EC00FE02FD29FEE7FE30.taxon	description	Figs 47 – 48	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF95EC00FE02FD29FEE7FE30.taxon	etymology	Etymology This species is named in honour of Professor V. N. Dublyansky (1930 – 2012), who was the instigator of complex karstological and speleological studies of the Kizil-Koba (= Krasnaya) Cave.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF95EC00FE02FD29FEE7FE30.taxon	materials_examined	Material examined Holotype CRIMEA • 1 ♂; Simferopol District, Dolgorukovsky karst massif, Kizil-Koba (= Krasnaya) Cave; 5 Nov. 2014; I. S. Turbanov leg.; ZMMU Dip 0006.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF95EC00FE02FD29FEE7FE30.taxon	description	Description BODY. Body length 2.7 mm. Epicuticle smooth under optical microscope but with a reticulate surface under SEM (Fig. 47); body with sparse thin and middle-sized clothing setae with 1 – 3 thin distal barbs. HEAD. Two intact antennae with 35 antennomeres, longer than body length (3.2 mm); thin subcylindrical sensillum on third antennomere in ventral position between macrosetae d and e; central antennomeres 2.1 × as long as wide, apical antennomere 3.0 × as long as wide; cupuliform organ occupying 1 / 7 of its length, with nine complex olfactory chemoreceptors; gouge sensilla 40 – 50 µm long in a single whorl of 10 – 12 sensilla on each medial and distal antennomere; a small, thin subcylindrical sensillum on third antennomere. Non-protruding frontal process with very slightly tubercular frontal macrosetae. Frontal process plain, with very slight tubercular frontal macrosetae; macrosetae along line of insertion of antennomere and x setae similar in length and with a few distal barbs (a / i / p / x with relative lengths of 29 / 19 / 24 / 25). Small suboval labial palps with a large subcylindrical sensillum, two guard setae, five normal setae and up to 20 neuroglandular setae. THORAX. Thoracic macrosetal distribution: pronotum with 1 + 1 ma, 2 + 2 la 2, 2 + 2 lp 2,3; mesonotum with 1 + 1 ma, 2 + 2 la 2,3, 2 + 2 lp 2,3, 1 + 1 mp; metanotum with 1 + 1 ma, 2 + 2 lp 2,3, 1 + 1 mp. All notal macrosetae long and barbed along distal two-thirds; marginal setae similar to clothing setae, but a bit longer. Legs elongated (1.53 mm long), overpassing end of abdomen. Femur I without dorsal macrosetae; femora II – III with one dorsal macroseta inserted on distal third. No ventral tibial macrosetae. Calcars with 2 – 3 rows of long barbs, from base with a smooth and narrow lateral side along each calcar. Dorsal and lateral subapical tarsal macrosetae smooth. Very unequal claws, posterior claw 1.80 – 2.05 × as long as anterior one, including a large backward overhang in posterior claw; ventral surface of claws with grooves, dorsal surface including large lateral crests with thin stripes. Posterior claw with a long, setiform pretarsal process reaching claw; anterior claw with short pretarsal processes without reaching extremity of this short claw (Figs 48 – 49). ABDOMEN. Distribution of abdominal macrosetae on tergites: 1 + 1 post 1 on I – III; 1 + 1 la, 1 + 1 post 1 on IV; 1 + 1 la, 2 + 2 post 1,2 on V; 1 + 1 la, 3 + 3 post 1,2,3 on VI; 1 + 1 la, 4 + 4 post 1,2,3,4 on VII; 5 + 5 post on VIII; 8 + 8 post on abdominal segment IX. All post urotergal macrosetae long, robust, barbed; la urotergal macrosetae much shorter, with thin distal barbs. Urosternite I with 7 + 7, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae; all urosternal macrosetae robust, large, covered by long barbs along distal half to three-fourths. Thick stylus with almost smooth stylar setae. SECONDARY SEX CHARACTERS. Male urosternite I without glandular g 1 setae, with subcylindrical, short and enlarged appendages with up to 15 large glandular a 1 setae, with spermatozoid fascicles. Female unknown.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF97EC00FE62FDB3FB88FA76.taxon	description	Fig. 49 – 50	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF97EC00FE62FDB3FB88FA76.taxon	materials_examined	Material examined WESTERN CAUCASUS • 1 ♂; Abkhazia, Ochamchira District, near Otap Village, Abrskila Cave; 9 Sep. 2014; I. S. Turbanov leg.; ZMMU.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF97EC00FE62FDB3FB88FA76.taxon	discussion	Remarks Body length 2.8 mm. It shows some differences from the type form in the number of macrosetae on the pronotum (1 + 2 la 2,3), metanotum (1 + 1 la), urotergite IV (without la macrosetae), urotergite V (without la and 1 + 1 post 1), urotergite VI (1 + 1 la, 2 + 2 post 1,2), urotergite VII (2 + 2 la 2,3, 2 + 2 post 1,2), urotergite VIII (4 + 4 post) and abdominal segment IX (7 + 7 post). Phyletic affinities, habitat and distribution Among the described species of Plusiocampa s. str., P. (P.) bulgarica, P (P.) elongata, P. (P.) vodniensis and P. (P.) beroni from the Balkan karst region (Condé 1954 a; Bareth & Condé 2001; Sendra et al. 2012 b), P. (P.) strouhali, P. (P.) caprai, P. (P.) grandii and P. (P.) hoelzeli from the Alps (Condé 1954 a; Neuherz 1984) and P. (P.) fagei from caves on Mallorca (Condé 1955 b) share two important taxonomic characters with P. (P.) dublanskii sp. nov.: the distinctly unequal claws with large lateral crests and an evident backward overhang on the posterior claw, and the presence of medial posterior macrosetae on the mesonotum and metanotum. Nonetheless, the absence of ventral tibial macrosetae or the extremely unequal claws with unequal lateral pretarsal processes suggest a completely different relationship with the unclear subgenus Venetocampa from the Veneto region, northern Italian Peninsula. These new troglomorphic taxa live in two separated karst regions in the north (Crimea) and the east (Western Caucasus) of the Black Sea. The important differences between the two studied specimens are remarkable, in particular the number and distribution of the macrosetae on the urotergites and abdominal segment, something that should be reviewed in the future with further samples.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF97EC01FE64FA72FAA2FE94.taxon	diagnosis	Diagnosis (Ionescu 1955, Sendra et al. 2012 b) Troglomorphic species. Antennae with 30 – 35 antennomeres; cupuliform organ with four complex olfactory chemoreceptors; protuding frontal process with very slightly tubercular setae. Pronotum with 1 + 1 ma, 4 + 4 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp, 1 + 1 mp macrosetae; metanotum with 1 + 1 ma, 2 + 2 la, 2 + 2 lp, 1 + 1 mp macrosetae; long, barbed notal macrosetae; abundant notal clothing setae. Two or three ventral tibial macrosetae; unequal claws (1.15 – 1.30) with large lateral crests, posterior claw with a backward overhang. Urotergites I – II with 1 + 1 post macrosetae; utrotergite III with 0 + 0 – 1 + 1 la, 3 + 3 – 4 + 4 post macrosetae; urotergite IV with 1 + 1 – 2 + 2 – 3 + 3 la, 4 + 4 post macrosetae; urotergite V with 2 + 2 – 3 + 3 la, 5 + 5 post macrosetae; urotergites VI – VII with 3 + 3 la, 5 + 5 post macrosetae; urotergite VIII with 6 + 6 post macrosetae; abdominal segment IX with 8 + 8 post macrosetae; urosternite I with 8 + 8, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae. Male without glandular g 1 setae, with enlarged, thick subtrapezoidal appendages with a large area of glandular a 1 setae; female with enlarged, thick subcylindrical appendages with a few glandular a 1 setae. Spermatozoid fascicles 70 µm in diameter, wheels with 2 ½ turns of 400 µm long and 7 µm wide filament.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF96EC01FE4DFEC8FC0CFD46.taxon	diagnosis	Differential diagnosis against the type form (Sendra et al. 2012 b) Urotergite I with 0 + 0 – 0 + 1 – 1 + 1 post macrosetae; urotergites II – III with 1 + 1 post macrosetae; urotergite IV with 2 + 2 post macrosetae; urotergites V – VII with 1 + 1 la, 4 + 4 post macrosetae. Habitat and distribution Subterranean species found in nine caves around the Carpathian Mountains in Romania; P. (P.) aff. elongata has only been found in one cave, the Peştera Cloşani cave.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF96EC01FE6FFD79FEE9F918.taxon	diagnosis	Diagnosis (Condé 1947 a, 1955 a, 1966; Bareth 1988) Non-troglomorphic species. Antennae with 21 – 25 antennomeres; four simple olfactory chemoreceptors in cupuliform organ; non-protruding frontal process with non-tubercular setae. Pronotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp macrosetae; metanotum with 1 + 1 ma, 1 + 1 la, 2 + 2 lp macrosetae; long, barbed notal macrosetae; smooth notal clothing setae. Two ventral tibial macrosetae. Small subequal claws (1.05) with middle-sized lateral crests, posterior claw with no backward overhang. Urotergites I – II with 1 + 1 post macrosetae; urotergite III with 2 + 2 post macrosetae; urotergites IV – V with 1 + 1 la, 5 + 5 post macrosetae; urotergites VI – VII with 2 + 2 (2 + 1) la, 5 + 5 post macrosetae; urotergite VIII with 6 + 6 post macrosetae; abdominal segment IX with 8 + 8 post macrosetae; urosternite I with 7 + 7, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae. Male with an area of glandular g 1 setae, subcylindrical enlarged appendages with glandular a 1 setae; female with slender subcylindrical appendages with glandular a 1 setae. Spermatozoid fascicle wheels with 1 ½ turns of 150 µm long and 10 µm diameter filament.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF96EC01FE6FFD79FEE9F918.taxon	discussion	Remarks The study of five specimens from two caves in Slovenia and Trieste, Italy, and one from Faedo San Michele, Trento, Italy (see Table S 2 in Supplementary Material) allowed the addition of information to the diagnosis. Habitat and distribution Hypogean species found in caves and frequently in soil. A total of 20 localities are reported from around Central Europe, from the Czech Republic and Slovakia to the Italian and Dinaric Mountains, reaching Croatia and Bosnia and Herzegovina towards the east and France towards the west, where it has been found in garden soils. This species is often found in urban soils, e. g., at ca 20 sites in Vienna (Christian 1992). The presence of P. (P.) cf. exsulans on the Anatolian Peninsula (Sendra et al. 2006) must be confirmed.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF96EC1EFE79F953FD7DFA02.taxon	description	Figs 51 – 52	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF96EC1EFE79F953FD7DFA02.taxon	diagnosis	Diagnosis (Condé 1955 b) Troglomorphic species. Antennae with 39 – 48 antennomeres; 21 complex olfactory chemoreceptors in cupulifrom organ; protruding frontal process with non-tubercular setae. Pronotum with 1 + 1 ma, 2 + 2 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 2 + 2 la, 2 + 2 lp, 1 + 1 mp macrosetae; metanotum with 1 + 1 ma, 1 + 1 la, 2 + 2 lp, 1 + 1 mp macrosetae; long, barbed notal macrosetae; sparse notal clothing setae. Two or three ventral tibial macrosetae; unequal claws (1.25) with large lateral crests, posterior claw with a backward overhang. Urotergites I – II with 1 + 1 post macrosetae; urotergite III with 0 + 0 – 1 + 1 la, 2 + 2 post macrosetae; urotergite IV with 1 + 1 la, 2 + 2 – 4 + 4 post macrosetae; urotergites V – VII with 1 + 1 la, 5 + 5 post macrosetae; urotergite VIII with 6 + 6 post macrosetae; abdominal segment IX with 8 + 8 post macrosetae; urosternite I with 6 + 6 – 8 + 8, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae. Male with a narrow area of glandular g 1 setae and very long cylindrical appendages with glandular a 1 setae; female with thin subcylindrical appendages with glandular a 1 setae.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF96EC1EFE79F953FD7DFA02.taxon	discussion	Remarks Two specimens from Cova Sa Gleda, Manacor, Mallorca (see Table S 2 in Supplementary Material) were used for SEM to look at some features that are difficult to observe under an optical microscope, such as their thin gouge sensilla, 48 – 50 µm long with an external transversal striped surface (Figs 51 – 52). Habitat and distribution Subterranean species inhabiting 19 caves in Mallorca, Balearic Islands, mainly in the Tramuntana mountain chain on the north of the island.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF89EC1FFE76F9BDFA84FCF2.taxon	diagnosis	Diagnosis (Silvestri 1933 a; Condé 1984 a) Slightly troglomorphic species. Antennae with 25 and 35 antennomeres. Non-protruding frontal process with non-tubercular setae. Pronotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp, 1 + 1 mp macrosetae; metanotum with 1 + 1 ma, 2 + 2 lp, 1 + 1 mp macrosetae; long, barbed notal macrosetae; abundant notal clothing setae. Two (rarely three) ventral tibial macrosetae; subequal claws (1.1) with large lateral crests; posterior claw with no backward overhang. Urotergites I – II with 1 + 1 post macrosetae; urotergite III with 1 + 0 la, 1 + 1 post macrosetae; urotergite IV with 1 + 1 la, 2 + 2 – 3 + 3 post macrosetae; urotergites V – VII with 1 + 1 la, 5 + 5 post macrosetae; urotergite VIII with 7 + 7 post macrosetae; abdominal segment IX with 9 + 9 post macrosetae; urosternite I with 7 + 7, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae. Male with an area of glandular g 1 setae and enlarged appendages with glandular a 1 setae.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF89EC1FFE76F9BDFA84FCF2.taxon	discussion	Remarks The studied material (see Table S 2 in Supplementary Material) from MSS habitats was collected from inland Greece. Three specimens from Évia show variability in the number of post macrosetae on the third urotergite; they have 2 + 1, 2 + 2 and 3 + 3 post macrosetae, showing the dubious value of the subspecies P. (P.) festae coi. In addition, a juvenile from Montes Orno, Crete, has also been studied and it matches in all chaetotaxic characters with P. (P.) festae festae. In conclusion, we suggest that P. (P.) festae coi should be synonymized with the nominate subspecies. Habitat and distribution Soil-dwelling species previously found on Karpathos and Leros, islands of the Dodecanese Archipelago; also collected in endogean environments at two sites on Crete and at a locality near Izmir, Anatolia, Turkey. It seems to inhabit MSS, where it has also been found at 14 localities in continental Greece.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF88EC1FFE2DFCDCFEA5F9E6.taxon	diagnosis	Diagnosis (Bareth & Condé 1984) Slightly troglomorphic species. Antennae with 27 antennomeres; seven simple olfactory chemoreceptors in cupuliform organ; frontal process with non-tubercular setae. Pronotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp, 1 + 1 mp macrosetae; metanotum with 1 + 1 ma, 1 + 1 la, 2 + 2 lp, 1 + 1 mp macrosetae; long, barbed notal macrosetae. Two ventral tibial macrosetae. Unequal claws (more than 1.2) with large lateral crests, posterior claw with a backward overhang. Urotergites III with 1 + 1 post macrosetae; urotergite III with 2 + 2 post macrosetae; urotergite IV with 1 + 1 la, 4 + 4 post macrosetae; urotergite V with 1 + 1 la, 5 + 5 post macrosetae; urotergites VI – VII with 2 + 2 la, 5 + 5 post macrosetae; urotergite VIII with 6 + 6 post macrosetae; abdominal segment IX with 8 + 8 post macrosetae; urosternite I with 7 + 7, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae. Male with a large area of glandular g 1 setae and enlarged appendages with a large area of glandular a 1 setae; female appendages thinner than those of male, with glandular a 1 setae. Habitat and distribution Subterranean species found at Inghiottitoio di Val dei Pai, Montereale Valcellina, Pordenone, Carnic Pre-Alps, Italy.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF88EC1CFE5CF9C8FB5AFD6A.taxon	diagnosis	Diagnosis (Sendra 2001) Troglomorphic species. Antennae with 46 – 52 antennomeres; 15 – 18 complex olfactory chemoreceptors in cupuliform organ; small protruding frontal process with a few slightly tubercular setae. Pronotum with 1 + 1 ma, 2 + 2 – 3 + 3 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 2 + 2 la, 2 + 2 (2 + 3) lp macrosetae; metanotum with 1 + 1 ma, 2 + 2 la, 2 + 2 (2 + 3) lp macrosetae; long, barbed notal macrosetae; short notal clothing setae. Two or three ventral tibial macrosetae; slightly unequal claws (1.15) with large lateral crests, posterior claw with no backward overhang. Urotergite II with 0 + 0 – 1 + 1 (0 + 1) post macrosetae; urotergite III with 0 + 0 – 1 + 1 la, 1 + 1 – 2 + 2 (0 + 1) post macrosetae; urotergite IV with 1 + 1 (1 + 2) la, 3 + 3 – 4 + 4 (4 + 5) post macrosetae; urotergite V with 1 + 1 (1 + 2) la, 4 + 4 – 5 + 5 (4 + 5) post macrosetae; urotergite VI with 1 + 1 (1 + 2) la, 5 + 5 (4 + 5) post macrosetae; urotergite VII with 1 + 1 la, 5 + 5 (4 + 5) post macrosetae; urotergite VIII with 6 + 6 (6 + 5) post macrosetae; abdominal segment IX with 8 + 8 post macrosetae; urosternite I with 7 + 7 – 8 + 8 (8 + 9), urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae. Male with an area of glandular g 1 setae, subcylindrical appendages with glandular a 1 setae; female and male appendages identical. Habitat and distribution Subterranean species endemic to the isolated karst of Sierra de Gádor, in the Peni-Baetic System, southeastern Iberian Peninsula. It is known from seven natural caves and one mine.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF8BEC1CFE70FD5DFE44FA67.taxon	diagnosis	Diagnosis (Condé 1984 a) Troglomorphic species. Antennae with 43 antennomeres; 14 olfactory chemoreceptors within cupuliform organ; non-protruding frontal process with non-tubercular setae. Pronotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 2 + 2 lp, 1 + 1 mp macrosetae; metanotum with 1 + 1 ma, 1 + 1 lp, 1 + 1 mp or smp macrosetae; thin, poorly barbed notal macrosetae; very sparse notal and urotergal clothing setae. Two ventral tibial macrosetae; subequal claws (1.0 – 1.1) with large lateral crests, posterior claw with a small backward overhang. Urotergites I – IV without macrosetae; urotergite V with 2 + 2 (1 + 2, 2 + 3) post macrosetae; urotergite VI with 1 + 1 la, 2 + 2 to 4 + 4 post macrosetae; urotergite VII with 1 + 1 la, 3 + 3 – 4 + 4 post macrosetae; urotergite VIII with 7 + 7 post macrosetae; abdominal segment IX with 9 + 9 post macrosetae; urosternite I with 7 + 7, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae. Male with a row of glandular g 1 setae and thin subcylindrical appendages with glandular a 1 setae; female and male appendages identical. Habitat and distribution Subterranean species found in Greece, namely at Spilaio Tripa tou Kalivaki, Klenia, Corinthos, southern Balkan Peninsula.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF8BEC1CFE61FA51FF2AF802.taxon	diagnosis	Diagnosis (Silvestri 1933 b; Condé 1954 a) Troglomorphic species. Antennae with 33 – 38 antennomeres. Pronotum with 1 + 1 ma, 2 + 2 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 2 + 2 la, 2 + 2 lp, 1 + 1 mp macrosetae; metanotum with 1 + 1 ma, 1 + 1 la, 2 + 2 lp, 1 + 1 mp macrosetae; long, barbed notal macrosetae; abundant clothing setae. Three ventral tibial macrosetae; unequal claws (1.25) with large lateral crests, posterior claw with a backward overhang. Urotergites I – II with 1 + 1 post macrosetae; urotergite III with 1 + 1 la, 1 + 1 post macrosetae; urotergite IV with 1 + 1 la, 5 + 5 post macrosetae; urotergites V – VII with 2 + 2 la, 5 + 5 post macrosetae; urotergite VIII with 6 + 6 post macrosetae; urosternite I with 7 + 7, urosternites II – VII with 6 + 6, urosternite VIII with 2 + 2 macrosetae. Male unknown; female with subcylindrical appendages bearing glandular a 1 setae.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF8AEC1DFE56FEB3FBE7FCF3.taxon	diagnosis	Differential diagnosis against the type form Observation of a single female from Inamila Cave in Evia, Greece (Coll. AS). It shows remarkable differences from the type form, such as the more anterior insertion position of mp mesonotal and metanotal macrasetae and post macrosetae on urotergites I – III. In addition, this Greek form shows important differences in the number and distribution of urotergal macrosetae: urotergite III with 1 + 0 la, 1 + 1 post; urotergite IV with 1 + 1 la, 4 + 5 post; urotegites VI – VII with 3 + 3 la, 5 + 5 post. Others features are: protruding frontal process with abundant tubercular setae; subequal claws with large lateral crests, posterior claw with a backward overhang; smooth subapical tarsal setae; stylus setae with abundant but separated barbs. Habitat and distribution Subterranean species only found in Bus del Globo Cave, in Trentino, Italian Alps. The aff. form from a cave in Greece should be accepted with caution, as it could be a new taxon.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF8AEC1DFE1AFCDCFACAFA0C.taxon	diagnosis	Diagnosis (Bareth & Condé 2001) Troglomorphic species. Antennae with 29 – 31 antennomeres; six simple olfactory chemoreceptors in cupuliform organ on probably regenerated antenna; protruding frontal process with tubercular setae. Pronotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 1 + 1 la, 2 + 2 lp macrosetae; metanotum with 1 + 1 ma, 0 + 1 la, 2 + 2 lp macrosetae; long, barbed notal macrosetae; barbed notal clothing setae. Two ventral tibial macrosetae; unequal claws (1.5) with large lateral crests, posterior claw with a backward overhang. Urotergite IV with 1 + 1 la, 2 + 2 post macrosetae; urotergites V – VII with 1 + 1 la, 4 + 4 post macrosetae; urotergite VIII with 6 + 6 post macrosetae; abdominal segment IX with 8 + 8 post macrosetae; urosternite I with 7 + 7, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae. Male with an area of glandular g 1 setae, enlarged appendages with glandular a 1 setae. Habitat and distribution Subterranean species found in Toplya Cave, Stara Planina Mountains (= Balkan Mts), Bulgaria.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF8AEC1BFE5CF9BEFEEAF970.taxon	description	Figs 53 – 58; Table 5	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF8AEC1BFE5CF9BEFEEAF970.taxon	diagnosis	Diagnosis (Neuherz 1984) Troglomorphic species. Antennae with 34 – 35 antennomeres; 10 – 12 complex olfactory chemoreceptors in cupuliform organ; protruding frontal process with non-tubercular setae. Pronotum with 1 + 1 ma, 2 + 2 – 3 + 3 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 sma, 1 + 1 la, 2 + 2 lp, 1 + 1 (0 + 1) smp macrosetae; metanotum with 1 + 1 lp macrosetae; long, barbed notal macrosetae; long, abundant notal clothing setae. One ventral tibial macroseta (two and three in Renejevo brezno); unequal claws (1.5) with large lateral crests, posterior claw with a backward overhang. Urotergite IV with 0 + 0 – 1 + 1 la, 1 + 1 post macrosetae; urotergite V with 1 + 1 la, 2 + 2 – 3 + 3 post macrosetae; urotergites VI – VII with 1 + 1 la, 4 + 4 post macrosetae; urotergite VIII with 5 + 5 post macrosetae; abdominal segment IX with 7 + 7 – 8 + 8 post macrosetae; urosternite I with 7 + 7, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae. Male with glandular g 1 setae, slightly enlarged subcylindrical appendages with abundant glandular a 1 setae; female with subcylindrical appendages with glandular a 1 setae.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF8AEC1BFE5CF9BEFEEAF970.taxon	discussion	Remarks Five specimens (three females, one male and one juvenile) of Plusiocampa were collected at a very deep location,> 1180 m deep in Renejevo brezno, Kanin Mountain, Slovenia (see Table S 2 in Supplementary Material). No other dipluran has yet been found at such a depth, with the exception of specimens of Plusiocampa (Stygiocampa) sp. presently under study from the Lukina jama – Trojama cave system in Croatia, collected by Branko Jalžić (Kazimir Micilinić, pers. obs.). These specimens match the thoroughly described population of P. (P.) hoelzeli found only in Hafnerhöhle, Karawanken Mountains in Carinthia. Nevertheless, there is some variability in important taxonomic features. First, the number of ventral tibial macrosetae on the metathoracic leg, with two (three in one leg) rather than only one; presumably, only the prothoracic legs were observed in the original description. In addition, there is variability in the antennomeres and cercal articles; their mesurements are recorded (Table 5). Body length from 3.4 to 5.1 mm; antennae shorter than body length, with 10 – 12 complex olfactory chemoreceptors in cupuliform organ; a whorl of gouge sensilla, 34 – 38 µm long, on each medial and distal antennomere, with an oblique striation (Figs 53 – 54); small protruding frontal process with slightly tuberculate anterior macrosetae; pronotum with 1 + 1 ma, 1 + 1 – 2 + 2 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 la, 2 + 2 lp macrosetae; metanotum with 1 + 1 – 0 + 0 lp (0 + 1 – 1 + 1 slp) macrosetae; unequal claws, posterior claw 1.4 – 1.5 × as long as anterior claw; posterior claw with a large backward overhang (Figs 55 – 57); urotergite IV with 1 + 1 la, 1 + 1 (2 + 1) post macrosetae; urotergite V with 1 + 1 la, 2 + 2 – 3 + 3 (3 + 1) post macrosetae; urotergites VI – VII with 1 + 1 la, 4 + 4 (4 + 3) post macrosetae; urotergite VIII with 6 + 6 post macrosetae; abdominal segment IX with 7 + 7 – 8 + 8 (8 + 7) post macrosetae; apical stylus setae with one or two medial thin barbs, subapical and medial setae with a few thin barbs (Fig. 58); cerci longer than body length in adult specimens (Table 5), with several whorls of long, barbed macrosetae on each article in addition to a whorl of smooth, shorter setae in distal position and a characteristic whorl of short, thin setae with a few thin distal barbs in apical position. Habitat and distribution Subterranean species found in a cave in Carinthia (Austria) and another one in the Kanin Mountains (Slovenia).	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF8CEC10FEF1F96AFC61FB90.taxon	description	Figs 59 – 94, 204; Tables 6 – 8	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF8CEC10FEF1F96AFC61FB90.taxon	etymology	Etymology This species is named in honour of Luc Hoffmann (1923 – 2016), an extraordinary man, a naturalist and philanthropist, who helped create many international conservation organisations and supported conservation research and actions in the Mediterranean and elsewhere.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF8CEC10FEF1F96AFC61FB90.taxon	materials_examined	Material examined Holotype GREECE • ♀; Crete, Marathos nomos Iraklion, Spilaio Doxa (n. 1065); 490 m a. s. l.; 11 May 2005; Fulvio Gasparo leg.; MZB. Paratypes GREECE – Crete • 2 ♂♂, 2 ♀♀; same collection data as for holotype but 2 Jun. 2002; Coll. AS • 1 ♀; Zoniana nomos Rethymon, Spilaio Sfento Trypa (n. 701); 630 m a. s. l.; 11 Apr. 2003; Kaloust Paragamian leg.; Coll. AS • 6 ♀♀; Krousonas, Malevizi, Krousaniotiko Livadi, Varathro Stou Bokon ton Poro; 1 Jul. 1990; Kaloust Paragamian leg.; Coll. AS • 1 ♀; Mylopotamos, Livadia, Varathro Sipouli; 24 Mar. 1990; Kaloust Paragamian leg.; Coll. AS • 1 ♂, 2 ♀♀; Malevizi, Gonies, Varathro Psistraki; 20 Nov. 2005; Kaloust Paragamian and S. Paragamian leg.; Coll. AS.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF8CEC10FEF1F96AFC61FB90.taxon	description	Description BODY. Body length: 4.8 mm (juvenile), 4.7 and 5.4 mm (males from Spilio Doxa), 5.9 and 6.4 mm (females from Spilio Doxa), and 7.4 mm (male from Sfentoni Trypa). Epicuticle smooth under optical microscope, but reticulate at high magnification (Figs 68 – 69); body with thin, long clothing, smooth or with a few thin distal barbs on dorsal side and well barbed on ventral side, legs and urosternal appendages. HEAD. Antennae with 44 – 53 antennomeres; central antennomeres 2.5 – 3 × as long as wide, apical antennomere 2.5 × as long as wide (Fig. 61). Large cupuliform organ occupying ¼ of total length of apical antennomere, with 8 – 10 complex olfactory chemoreceptors (Figs 59 – 60). Small coniform sensillum of third antennomere located in ventral position between c and d macrosetae. Thin and long gouge sensilla, 38 – 50 µm long, in a single distal whorl of 10 – 14 sensilla on each medial and distal antennomere (Figs 62 – 64). Protruding frontal process with tubercular setae and macrosetae with thin barbs. Three macrosetae along line of insertion of antennomere and x setae relatively long, covered from distal third to fourth (Figs 65 – 66). THORAX. Thoracic macrosetal distribution (Fig. 67): pronotum with 1 + 1 ma, 4 + 4 la 1,2,3,4, 2 + 2 lp 2,3; mesonotum with 1 + 1 ma, 3 + 3 la 1,2,3, 2 + 2 lp 2,3, 1 + 1 mp; metanotum with 1 + 1 ma, 0 + 0, 0 + 1, 1 + 1, 1 + 2 la or 0 + 1, 1 + 1 sla, 2 + 2 lp 2,3, 1 + 1 mp. All medial posterior macrosetae in a forward position. All long, thin notal macrosetae with very thin barbs along distal third or half; marginal setae longer than clothing setae, covered by a few thin distal barbs. Legs elongated, metathoracic legs overpassing end of abdomen at beginning of tarsus. Tibia longest part of leg, 1.11 – 1.30 × as long as tarsus and 1.21 – 1.37 × as long as femur (Table 7). Ratios between lengths of tibia, tarsus and femur seem more or less fixed (Table 7). Femora II – III with one long, thin dorsal macroseta and one shorter ventral macroseta. Tibiae I – III with two or three ventral macrosetae surrounded by thin, short barbs. Calcars surrounded from base almost to tip by thin, short barbs. Metatarsus with a pseudo-articulation on distal third. Typical double rows of thick, completely barbed setae along ventral side of tarsus. Dorsal and lateral subapical tarsal setae completely covered by thin barbs nearly from base almost to tip. Subequal to slightly unequal claws (posterior claw 1.05 – 1.15 × as long as anterior one), with large lateral crests well developed on posterior claw prolongated backwards. Ventral side of claws with longitudinal ridges separated by a surface of micro-hemispherical structures. Pretarsal processes setiform, with abundant, very short proximal barbs (Figs 71 – 82). ABDOMEN. Distribution of abdominal macrosetae on tergites: 1 + 1 post 1 on I – II; 1 + 1 – 0 + 0 la, 1 + 1 post 1 on III; 1 + 1 la, 3 + 3 – 4 + 4 post 1,2,4,5 on IV; 1 + 1 la, 5 + 5 post 1 – 5 on V – VII, 7 + 7 post on VIII; 8 + 8 – 9 + 9 post on abdominal segment IX. Urosternite I with 7 + 7 – 9 + 9, urosternites II – VII with 5 + 5 – 7 + 7, urosternite VIII with 2 + 2 macrosetae; all urosternal macrosetae robust, large, covered by long barbs along distal half to four-fifths (Figs 87 – 88). Stylus with subapical setae longer and more barbated, in fact completely surrounded by short, abundant barbs, more numerous on apical and ventromedial setae (Figs 92 – 93); large eversible vesicles (Figs 92, 94). Two complete cerci, 2.37 and 1.71 × as long as body; this ratio higher in an incomplete 14.50 mm long cercus of 5.4 mm long male (Table 8); up to 11 articles, covered with numerous disorganized, long, thin macrosetae with thin barbs along distal half, in addition to a distal whorl of longer, thicker macrosetae on each article plus an apical whorl of barbed, thin, short clothing setae (Fig. 70). SECONDARY SEX CHARACTERS. Female urosternite I with subcylindrical appendages, each bearing up to 18 glandular a 1 setae in a distal area (Figs 89 – 91). Male urosternite I (Figs 83 – 86) with a posterior area of up to 160 glandular g 1 setae in larger male from Sfedoni Trypa (absent in smaller male); with thick, large, subcylindrical appendages, much thicker than female appendages, each bearing up to 100 glandular a 1 setae in a large distal area in two largest adults. Spermatozoid fascicles with a thick spiral filament. Phyletic affinities, habitat and distribution The presence of medial posterior macrosetae on the mesonotum and metanotum relates P. (P.) hoffmanni sp. nov. to almost half of the species in the subgenus Plusiocampa, all living in the European region, including most of the Mediterranean islands. Among them, P (P) festae is probably the closest related species due to the distribution pattern of their notal macrosetae; in spite of the fact that both species live in Crete (Condé 1984 a; Silvestri 1933 a), they occupy different habitats, P. (P.) hoffmanni sp. nov. in deep subterranean habitats, P. (P.) festae in soil and subterranean superficial habitats. In addition to this ecological difference, there are several important features that allow them to be separated. Plusiocampa (P.) hoffmanni sp. nov. has troglomorphic traits, with longer appendages than P. (P.) festae and more antennomeres and cercal articles, with more complex and more numerous olfactory chemoreceptors within the cupuliform organ. In P. (P.) hoffmanni sp. nov., the claws are slightly unequal to subequal, but with large lateral crests and a backward prolongation of the posterior claws. Additionally, P. (P.) hoffmanni sp. nov. shows a protruding frontal process covered by tuberculated setae, 8 + 8 macrosetae on the first urosternite and 6 + 6 on the second to seventh urosternites. Some of these features also separate P. (P.) hoffmanni sp. nov. from P. (P.) rybaki, a poorly described troglomorphic species from the southern Balkan Peninsula (Condé 1956). Plusiocampa (P.) hoffmanni sp. nov. has been collected from the deep zone of 14 caves in the numerous karst areas of Crete (Fig. 204).	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF87EC10FE5BFB3AFDE0F88B.taxon	diagnosis	Diagnosis (Ionescu 1951; Sendra et al. 2012 b) Non-troglomorphic species. Antennae with 21 – 23 doliiform antennomeres; four or five olfactory chemoreceptors; frontal process without non-tubercular setae. Pronotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp, 1 + 1 mp macrosetae; metanotum with 1 + 1 ma, 1 + 1 la, 2 + 2 lp, 1 + 1 mp macrosetae; long, barbed notal macrosetae. Two ventral tibial macrosetae; slightly unequal claws (1.1 – 1.2) with large lateral crests, posterior claw without backward overhang. Urotergites I – II with 1 + 1 post macrosetae; urotergite III with 2 + 2 post macrosetae; urotergite IV with 1 + 1 la, 4 + 4 post macrosetae; urotergite V with 1 + 1 la, 5 + 5 post macrosetae; urotergites VI – VII with 2 + 2 la, 5 + 5 post macrosetae; urotergite VIII with 6 + 6 post macrosetae; abdominal segment IX with 8 + 8 post macrosetae; urosternite I with 7 + 7, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae. Male without glandular g 1 setae; subcylindrical appendages with glandular a 1 setae. Appendages of female slightly thinner, subcylindrical, with glandular a 1 setae. Habitat and distribution Soil-dwelling species found at four sites around the South Carpathian Mountains, including the Transylvanian reliefs, Romania.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF87EC11FE21F835FBE3FCD4.taxon	diagnosis	Diagnosis (Bareth & Condé 1984) Troglomorphic species. Antennae with 31 antennomeres (regenerated antenna); frontal process with non-tubercular setae. Pronotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp, 1 + 1 mp macrosetae; metanotum with 1 + 1 ma, 2 + 2 la, 2 + 2 lp, 1 + 1 mp macrosetae; long, barbed notal macrosetae; abundant notal clothing setae. Two ventral tibial macrosetae; slightly unequal claws (1.2) with large lateral crests, posterior claw with a small backward overhang. Urotergite I with 2 + 2 post macrosetae; urotergite II with 2 + 3 post macrosetae; urotergite III with 3 + 3 post macrosetae; urotergite IV with 1 + 1 la, 5 + 5 post macrosetae; urotergites V – VII with 3 + 3 la, 5 + 5 post macrosetae; urotergite VIII with 7 + 7 post macrosetae; abdominal segment IX with 9 + 9 post macrosetae; urosternite I with 7 + 7, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae. Male unknown; female with subcylindrical appendages bearing glandular a 1 setae. Habitat and distribution Subterranean species, found only in Bus de Bec Cave, Brescia, Italian Alps.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF86EC11FE76FC86FE26FAA6.taxon	diagnosis	Diagnosis (Condé 1993 a, 1996) Non-troglomorphic species. Antennae with 23 antennomeres; four simple olfactory chemoreceptors in cupuliform organ; slightly protruding frontal process with some slightly tubercular setae. Pronotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp macrosetae; metanotum with 1 + 1 ma, 1 + 1 (2 + 1) la, 2 + 2 lp macrosetae; long, thin, barbed notal macrosetae; short, smooth clothing setae. Two ventral tibial macrosetae; unequal claws (1.29) with large lateral crests, posterior claw with a backward overhang; pretarsal processes short, with a few sparse barbs. Urotergites I – II with 1 + 1 post macrosetae; urotergite III with 2 + 2 post macrosetae; urotergites IV – V with 1 + 1 la, 5 + 5 post macrosetae; urotergites VI – VII with 2 + 2 la, 5 + 5 post macrosetae; urotergite VIII with 6 + 6 post macrosetae; abdominal segment IX with 8 + 8 post macrosetae; urosternite I with 7 + 7, urosternites IIVII with 5 + 5, urosternite VIII with 2 + 2 macrosetae. Male unknown; subcylindrical female appendages bearing glandular a 1 setae.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FF86EC11FE5FFA17FAD9F850.taxon	diagnosis	Differential diagnosis against the type form This record is based on seven specimens found in Ledenjača Cave, Bosnia and Herzegovina (see Table S 2 in Supplementary Material). Due to their clear differences from the type of P. (P.) isterina, described from Movile Cave, they have been identified as P. (P.) aff. isterina. Specimens from Ledenjača Cave have a higher number of antennomeres (25 – 28), six olfactory chemoreceptors in a relatively large cupuliform organ (occupying one third of a short antennomere) and a clearer protruding frontal process with remarkable tuberculate setae. Habitat and distribution Non-troglomorphic species. The nominate form is found in Peştera Movile Cave, Southern Dobruja, Romania. The aff. form is found in a cave in the Dinaric Mountains, Bosnia and Hercegovina.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFF9EC6AFE38FEB3FB65FD73.taxon	description	Figs 95 – 118, 203	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFF9EC6AFE38FEB3FB65FD73.taxon	diagnosis	Diagnosis (Sendra & Condé 1987; Sendra 2001; Sendra et al. 2004) Troglomorphic species. Antennae with 32 – 34 antennomeres; 15 – 18 complex olfactory chemoreceptors in cupuliform organ; slightly protruding frontal process with non-tubercular setae. Pronotum with 1 + 1 ma, 3 + 3 (4 + 3) la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 2 + 2 la, 2 + 2 lp, 1 + 1 mp macrosetae; metanotum with 1 + 1 ma, 1 + 1 la, 2 + 2 lp (1 + 1 lp supplementary in some populations), 1 + 1 mp macrosetae. Two or three ventral tibial macrosetae; subequal claws (1.1) with large lateral crests, posterior claw with a small backward overhang. Urotergites I – III with 1 + 1 post macrosetae; urotergite IV with 1 + 1 la, 2 + 2 – 3 + 3 – 4 + 3 post macrosetae; urotergites V – VII with 1 + 1 la, 5 + 5 (4 + 5) post macrosetae; urotergite VIII with 6 + 6 (6 + 5, 6 + 7) post macrosetae; abdominal segment IX with 8 + 8 (8 + 9) post macrosetae; urosternite I with 7 + 7, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae. Male with a narrow area of glandular g 1 setae, subclylindrical appendages with glandular a 1 setae; female with subcylindrical appendages bearing glandular a 1 setae. Spermatozoid fascicle wheels with 1 ½ turns of 110 – 140 µm long and 6 µm wide filament.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFF9EC6AFE38FEB3FB65FD73.taxon	discussion	Remarks Ten specimens from the Chorros and Farallón Caves in Riopar, Albacete, Spain (see Table S 2 in Supplementary Material) were used for SEM to study some features that are difficult to observe under an optical microscope. Epicuticle slightly reticulate, with large polygonal pattern (Fig. 107); antennomeres with a whorl of thin gouge sensilla, 34 – 36 µm long, with an external transversal striped surface (Figs 96 – 100). Cupuliform organ with spheroidal olfactory chemoreceptors in a complex spiral shape (Fig. 95). Head suboval with slightly protruding frontal process (Figs 101 – 102). Labial palp with short neuroglandular setae (Figs 103 – 106). Tarsus with two ventral barbed rows (Fig. 112). Claws with large lateral crest with dorsal striated surface and with ventral longitudinal grooves separated by a multihemispherical surface (Figs 108 – 111). Male with a large area of glandular g 1 setae and slightly thicker subcylindrical appendages (Figs 113 – 117). Stylar setae covered by a few thin, short barbs (Fig. 118). Habitat and distribution Subterranean species found in 31 caves in the external zone of the Baetic Mountain Range, around the carbonate areas of the Cazorla, Segura, Alcaraz and Castril Mountains and the peripheral karst areas of Filabres Mountain and the Yesos de Sorbas, southeastern Iberian Peninsula (Fig. 203).	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFFDEC6AFE73FD6EFD61FB45.taxon	diagnosis	Diagnosis (Silvestri 1932 a) Non-troglomorphic species. Antennae with 23 – 26 antennomeres; non-protruding frontal process with simple setae. Pronotum with 1 + 1 ma, 4 + 4 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp, 1 + 1 mp macrosetae; metanotum with 1 + 1 ma, 1 + 1 la, 2 + 2 lp, 1 + 1 mp macrosetae; long, barbed notal macrosetae. Two ventral tibial macrosetae; subequal claws (1.05) with large lateral crests without prolonged backward overhang. Urotergites I – II with 1 + 1 post macrosetae; urotergite III with 2 + 2 post macrosetae; urotergite IV with 1 + 1 la, 5 + 5 post macrosetae; urotergites V – VII with 3 + 3 la, 5 + 5 post macrosetae; urotergite VIII with 7 + 7 post macrosetae; urosternite I with 8 + 8, urosternite II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFFDEC6AFE5BFB07FAA2F9B5.taxon	diagnosis	Differential diagnosis against the type form One female and two juveniles (Coll. AS) collected in a couple of Croatian caves (see Table S 2 in Supplementary Material) show several differences from the type form in: the non-troglomorphic shape; antennae with 24 antennomeres and four simple olfactory chemoreceptors; the number of urotergal macrosetae, with 1 + 1 post macrosetae on urotergite III, 6 + 6 post macrosetae on urotergite VIII and 8 + 8 post macrosetae on abdominal segment IX; and a unique ventral tibial macroseta. Habitat and distribution Hypogean species found in edaphic habitats. The nominate form is known from a couple of localities in Marizza and Cremastò, Ródos, Greece. The aff. form was found in two caves in the Dinaric Mountains.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFFDEC6BFE7CF929FC62FD91.taxon	diagnosis	Diagnosis (Condé 1948 d) Slightly troglomorphic species. Antennae with 24 – 27 with antennomeres. Small protruding frontal process with slightly tuberculated setae. Pronotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp macrosetae; metanotum with 1 + 1 ma, 1 + 1 la, 2 + 2 lp macrosetae; long, barbed notal macrosetae. Two ventral tibial macrosetae; unequal claws (1.4) with large lateral crests, posterior claw with a backward overhang. Urotergites I – III with 1 + 1 post macrosetae; urotergite IV with 1 + 1 la, 3 + 3 post macrosetae; urotergites V – VII with 1 + 1 la, 5 + 5 post macrosetae; urotergite VIII with 6 + 6 post macrosetae; abdominal segment IX with 8 + 8 post macrosetae; urosternite I with 7 + 7, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae. Male with glandular g 1 setae, subcylindrical appendages with glandular a 1 setae; female and male appendages identical.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFFDEC6BFE7CF929FC62FD91.taxon	discussion	Remarks Among the studied material there is a male from Žira jama and another male from Crnulja ponor from Bosnia and Herzegovina (see Table S 2 in Supplementary Material) which match the original description and allowed the completion of the previous diagnosis (Coll. AS).	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFFCEC6BFE64FD3BFE18FAEF.taxon	diagnosis	Differential diagnosis against the type form One male and one female from Vidovića špilja, Croatia (see Table S 2 in Supplementary Material) show notable differences compared to the type form. Antennae with 25 doliiform antennomeres; nontubercular setae on simple frontal process. Pronotum with 1 + 1 sma, 1 + 1 ma, 3 + 3 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp macrosetae; metanotum with 1 + 1 ma, 1 + 1 la, 2 + 2 lp macrosetae. Urotergites I – II with 1 + 1 post macrosetae; urotergite III with 2 + 2 post macrosetae; urotergite IV with 0 + 1 la, 4 + 4 – 5 + 5 post macrosetae; urotergite V with 1 + 1 la, 5 + 5 post macrosetae; urotergites VI – VII with 2 + 2 la, 5 + 5 post macrosetae; urotergite VIII with 6 + 6 post macrosetae; abdominal segment IX with 8 + 7 post macrosetae. Urosternite I with 7 + 7, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae, mostly short with long barbs. Subequal claws (1.1) with small backward overhang. Male with glandular g 1 setae and enlarged appendages with thick glandular a 1 setae; female with enlarged subcylindrical appendages with glandular a 1 setae. Stylar setae with a few barbs. Habitat and distribution Hypogean species found in three caves (Baba pećina, Crnulja ponor and Žira jama) in Bosnia and Herzegovina for the type form and one cave for the aff. form (Vidovića špilja, Croatia); all localities in the Dinaric Mountains.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFFCEC68FE61FAD1FB01FD93.taxon	description	Fig 119	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFFCEC68FE61FAD1FB01FD93.taxon	diagnosis	Diagnosis (Condé 1956 a) Slightly troglomorphic species. Antennae with 34 – 37 antennomeres; five olfactory chemoreceptors in cupuliform organ of a possibly regenerated antenna; non-protruding frontal process with nontubercular setae. Pronotum with 1 + 1 ma, 4 + 4 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 2 + 2 la, 2 + 2 lp macrosetae; metanotum with 1 + 1 ma, 2 + 2 lp macrosetae; long, large, barbed notal macrosetae; smooth clothing setae. Two ventral tibial macrosetae; subequal claws (1.05 – 1.1) with middle-sized lateral crests, posterior claw with no backward overhang. Urotergites I – III with 1 + 1 post macrosetae; urotergite IV with 1 + 1 la, 2 + 2 post macrosetae; urotergites V – VII with 1 + 1 la, 4 + 4 post macrosetae; urotergite VIII with 6 + 6 post macrosetae; abdominal segment IX with 8 + 8 post macrosetae; urosternite I with 7 + 7, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae. Male with glandular g 1 setae, enlarged subcylindrical appendages with glandular a 1 setae; female with slender, subcylindrical appendages.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFFCEC68FE61FAD1FB01FD93.taxon	discussion	Remarks The study of one male adult of 4.6 mm body length collected from the MSS habitat in Xànthi, Gérakas, Greece (see Table S 2 in Supplementary Material) allows the addition of information on the urosternites of the male. It has a narrow area of 96 glandular g 1 setae, arranged in 2 – 3 rows; enlarged and subcylindrical appendages with a distal area of up to 60 glandular a 1 setae (Fig. 119). Habitat and distribution Subterranean species known from Saint-Jean-le-Prodrome Cave (Spilaio Pelade) and from the MSS habitat near Xánthi, both on the Balkan Peninsula, in the Macedonian and Greek regions.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFFEEC69FE3EFEB3FE5CFBC4.taxon	diagnosis	Diagnosis (Sendra & Condé 1986; Sendra et al. 2004) Troglomorphic species. Antennae with 37 – 40 antennomeres; 17 – 21 complex olfactory chemoreceptors in cupuliform organ; non-protruding frontal process with non-tubercular setae. Pronotum with 1 + 1 ma, 2 + 2 – 3 + 3 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 2 + 2 – 3 + 3 la, 2 + 2 lp, 1 + 1 (0 + 1) mi, 1 + 1 (0 + 1) li macrosetae; metanotum with 1 + 1 ma, 1 + 1 la, 2 + 2 lp, 1 + 1 mi, 1 + 1 – 0 + 0 li macrosetae; long, barbed notal macrosetae; sparse notal clothing setae. Two or three ventral tibial macrosetae; unequal claws (1.30) with large lateral crests, posterior claw with backward overhang. Urotergites I – II with 1 + 1 post macrosetae; urotergite III with 1 + 1 la, 1 + 1 post macrosetae; urotergite IV with 1 + 1 la, 2 + 2 post macrosetae; urotergites V – VII with 1 + 1 la, 4 + 4 post macrosetae; urotergite VIII with 6 + 6 post macrosetae; abdominal segment IX with 8 + 8 post macrosetae. Urosternite I with 7 + 7 (7 + 1 + 7), urosternites II – VII with 5 + 5 – 6 + 6, urosternite VIII with 2 + 2 macrosetae. Male without glandular g 1 setae, slightly subcylindrical appendages with glandular a 1 setae; female appendages like male appendages. Spermatozoid fascicle wheels with 2 turns of 140 – 160 µm long and 1.5 µm wide filament. Habitat and distribution Subterranean species inhabiting seven caves around the north-eastern Pre-Baetic reliefs on the Iberian Peninsula. Furthermore, it slightly extends its distribution to three more caves in the south of the Iberian Mountain Range.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFFEEC69FE52FBFEFD3CF888.taxon	diagnosis	Diagnosis (Condé 1957, 1961, 1983; Bareth 2006) Troglomorphic species. Antennae with 31 – 33 antennomeres; ten olfactory chemoreceptors in cupuliform organ; non-protruding frontal process with non-tubercular setae. Pronotum with 1 + 1 ma, 2 + 2 – 3 + 3 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 3 + 3 (2 + 2) la, 2 + 2 lp, 0 + 1 mp (1 + 0 smp) macrosetae; metanotum with 1 + 1 ma, 1 + 1 la, 2 + 2 lp, 1 + 1 smp macrosetae; long, barbed notal macrosetae; sparse notal clothing setae. Three or four ventral tibial macrosetae; slightly unequal claws with large lateral crests, posterior claw with a small backward overhang. Urotergite I with 1 + 1 post macrosetae; urotergite II with 1 + 1 – 2 + 2 post macrosetae; urotergite III with 2 + 2 – 3 + 4 post macrosetae; urotergite IV with 2 + 2 la, 5 + 5 post macrosetae; urotergite V with 3 + 3 la, 5 + 5 – 5 + 4 post macrosetae; urotergite VI with 3 + 3 la, 5 + 5 post macrosetae; urotergite VII with 3 + 3 la, 4 + 5 – 5 + 5 post macrosetae; urotergite VIII with 6 + 6 post macrosetae; abdominal segment IX with 8 + 8 post macrosetae; thick clothing setae on abdominal segments IX – X; urosternite I with 7 + 7, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae. Male unknown; female appendages slightly enlarged, from subcylindrical to conical in shape, with glandular a 1 setae. Habitat and distribution Subterranean species found in two adjacent caves, Grotte du Pertuis and Trou Miette, north of Méailles, Alpes de Haute-Provence, southeastern France.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFFEEC66FE59F83AFCA1FCDD.taxon	diagnosis	Diagnosis (Silvestri 1912; Condé 1978; Bareth 2006) Non-troglomorphic species. Antennae with 25 – 27 antennomeres. Pronotum with 1 + 1 ma, 4 + 4 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp, 1 + 1 mp macrosetae; metanotum with 1 + 1 ma, 1 + 1 la, 2 + 2 lp, 1 + 1 mp macrosetae; long, thin barbed macrosetae; sparse smooth, long notal clothing setae. Two ventral tibial macrosetae; small subequal claws with small lateral crests, posterior claw with no backward overhang. Urotergites I – II with 1 + 1 post macrosetrae; urotergite III with 2 + 2 post macrosetae; urotergites IV – VII with 1 + 1 la, 5 + 5 post macrosetae; urotergite VIII with 6 + 6 post macrosetae; abdominal segment IX with 8 + 8 post macrosetae; urosternite I with 7 + 7, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae. Male with glandular g 1 setae, subcylindrical appendages with glandular a 1 setae; female appendages like male appendages. Habitat and distribution Soil-dwelling species found at six edaphic sites on the island of Corsica, one on Isola Zannone and two at inland localities in Calabria and Lazio, Italian Peninsula.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFF1EC66FE0BFC97FE5BF914.taxon	description	Figs 120 – 123	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFF1EC66FE0BFC97FE5BF914.taxon	diagnosis	Diagnosis (Denis 1930; Condé 1948 b; Bareth 2006) Troglomorphic species. Antennae with 33 – 35 antennomeres, protruding frontal process with tubercular setae. Pronotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 3 + 3 la, 3 + 3 lp macrosetae; metanotum with 1 + 1 ma, 2 + 2 la, 3 + 3 lp macrosetae; long, barbed notal macrosetae; barbed notal clothing setae. Two ventral tibial macrosetae; unequal claws with large lateral crests, posterior claw with a backward overhang. Urotergites I – III with 1 + 1 post macrosetae; urotergite IV with 1 + 1 la, 5 + 5 post macrosetae; urotergites V – VI with 1 + 1 – 2 + 2 la, 5 + 5 post macrosetae; urotergite VII with 2 + 2 la, 5 + 5 post macrosetae; urotergite VIII with 6 + 6 post macrosetae; abdominal segment IX with 8 + 8 post macrosetae. Urosternite I with 8 + 8, urosternites II – VII with 6 + 6, urosternite VIII with 2 + 2 macrosetae. Male with a area of glandular g 1 setae, developed large appendages with a large area of glandular a 2 setae and an apical area of glandular a 1 setae; female with subcylindrical appendages with glandular a 1 setae.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFF1EC66FE0BFC97FE5BF914.taxon	discussion	Remarks The studied material from Pouade Cave (see Table S 2 in Supplementary Material) has allowed a better observation of the first urosternite with its very large appendages in males (Fig. 120), as well as a large area of glandular a 2 setae. The first urosternite of females has subcylindrical appendages (Figs 121, 123) and the surface is completely smooth (Fig. 122). Habitat and distribution Subterranean species only found in Grotte de Pouade, Banyuls de la Marenda, on the French side of the Eastern Pyrenees.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFF1EC67FE2AF94EFB0DFE8F.taxon	diagnosis	Differential diagnosis against the type form (Condé 1955 a; Bareth & Condé 1996) Urotergite III with 2 + 2 post macrosetae; urotergites V – VII with 2 + 2 la, 6 + 6 post macrosetae; urotergite VIII with 7 + 7 post macrosetae. Habitat and distribution Subterranean subspecies inhabiting two caves on the French side of the Eastern Pyrenees.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFF3EC64FE3EFEB3FB27FD54.taxon	diagnosis	Differential diagnosis against the type form (Condé 1951, 1955 a; Bareth & Condé 1996) Antennae with 28 – 33 antennomeres. Urotergite III with 2 + 2 post macrosetae; urotergite VIII with 7 + 7 post macrosetae. Habitat and distribution Subterranean subspecies inhabiting three caves on the Spanish side of the Eastern Pyrenees.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFF3EC64FE27FA59FD59F879.taxon	diagnosis	Differential diagnosis against the type form (Condé 1949, 1983; Bareth 2006) Protruding frontal process with slightly tubercular setae. Two ventral tibial macrosetae. Thick clothing setae on abdominal segments VIII – X. Male with slightly enlarged appendages with glandular a 1 setae; female appendages like male appendages. Habitat and distribution Subterranean subspecies found in Grotte du Saint-Trou in the Saint-Baume Massif, northeast of Évenos, Alps de Provence, southeastern France.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFF3EC64FE0AFD7EFD3CFA78.taxon	diagnosis	Diagnosis (Condé 1949, 1957, 1981 a, 1983; Bareth 2006) Troglomorphic species. Antennae with 30 – 36 antennomeres; sensillum of third antennomere very long, in ventral position; protruding frontal process with tubercular setae. Pronotum with 1 + 1 ma, 4 + 4 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp, 1 + 1 mp macrosetae; metanotum with 1 + 1 ma, 2 + 2 la, 2 + 2 lp, 1 + 1 mp macrosetae; long, barbed notal macrosetae; barbed notal clothing setae. Two ventral tibial macrosetae; subequal claws (1.06) with large lateral crests, posterior claw with a small backward overhang. Urotergites I – II with 2 + 2 post macrosetae; urotergite III with 2 + 2 – 3 + 3 post macrosetae; urotergite IV with 1 + 1 la, 5 + 5 post macrosetae; urotergite V with 2 + 2 la, 5 + 5 post macrosetae; urotergites VI – VII with 3 + 3 la, 5 + 5 post macrosetae; urotergite VIII with 7 + 7 post macrosetae; thick clothing setae on abdominal segments IX – X; urosternite I with 7 + 7, urosternites IIVII with 5 + 5, urosternite VIII with 2 + 2 macrosetae. Male with a large area of glandular g 1 setae and appendages with glandular a 2 setae. Habitat and distribution Subterranean species found in nine caves around Massif de Marseilleveyre and the Sainte-Baume Massif, Alps de Provence, southeastern France.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFF2EC63FE75FDF2FD94FBC5.taxon	description	Fig. 124; Table 9	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFF2EC63FE75FDF2FD94FBC5.taxon	etymology	Etymology This species is named in honour of Rhea, mother of the Greek god Zeus and daughter of Gaea, the Earth goddess. She gave birth and cared for her son Zeus in the Ideon Andron Cave located in Mount Psiloritis, the highest mountain in Crete, where this new species was found.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFF2EC63FE75FDF2FD94FBC5.taxon	materials_examined	Material examined Holotype GREECE • 1 ♂; Crete, Anogeia, Ideon Antron Cave (Zeus Cave) at twilight zone area, near entrance; 1174 m a. s. l.; 22 Mar. 2006; Alberto Sendra leg.; MZB 2019 - 1033. Paratypes GREECE – Crete • 1 ♂; Pombia; 211 m a. s. l.; Alberto Sendra leg.; MSS habitat; Coll. AS • 1 ♂; Sissarha; 601 m a. s. l.; MSS habitat; Alberto Sendra leg.; Coll. AS.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFF2EC63FE75FDF2FD94FBC5.taxon	description	Description BODY. Body length 3.8 – 4.9 mm. Epicuticle smooth under optical microscope; body with thin, middlesized clothing, covered or not by a few thin barbs. HEAD. Three intact antennae in holotype and paratypes, with 30, 32 and 33 antennomeres, slightly shorter than body length (Table 9). Small coniform sensillum of third antennomere located in ventral position between c and d macrosetae. Central antennomeres 1.5 × as long as wide, apical antennomere 1.6 × as long as wide. Cupuliform organ occupying of total length of apical antennomere, with 8 – 9 complex olfactory chemoreceptors. Gouge sensilla 30 ⅕ – 32 µm long in a single distal whorl of 12 – 16 sensilla on each medial and distal antennomere. Frontal process plain, with non-tubercular setae and macrosetae with barbs along half to two-thirds; macrosetae along line of insertion of antennomere and x setae with similar lengths (a / i / p / x with relative lengths of 36 / 42 / 35 / 52 in holotype). Suboval labial palps with latero-external sensillum larger than sensillum of third antennomere, with two guard setae, up to 12 setae on anterior border and up to 180 neuroglandular setae. THORAX. Distribution of thoracic macrosetae (Fig. 124): pronotum with 1 + 1 ma, 2 + 2 la 3,4 and 2 + 2 sla 1,2, 2 + 2 lp 1,3 (with additional 1 + 1 extra sma near the sagittal plane); mesonotum with 1 + 1 ma, 3 + 3 la 1,2,3, 3 + 3 lp 1,2,3, 1 + 1 mp; metanotum with 1 + 1 ma, 3 + 3 lp 1,2,3, 1 + 1 mp. All notal macrosetae long, robust, covered by thin barbs along distal third or half; submacrosetae sla 1,2 and sma thinner and shorter than notal macrosetae, but with same large base; marginal setae longer than clothing setae, covered by thin barbs along distal three-fourths. Legs elongated, pretarsus of metathoracic legs slightly overpassing end of abdomen (Table 9). Femur I without dorsal macrosetae but with one long, barbed ventral macroseta; femora II – III with one long, barbed dorsal and one long, barbed ventral macroseta. Tibia I with one short, barbed ventral macroseta; tibiae II – III with two short, barbed ventral macrosetae. Calcars with 2 – 3 rows of long barbs from base to tip, smooth on one narrow side along calcar. Dorsal and lateral subapical tarsal setae smooth. Slightly unequal claws (posterior claw 1.15 – 1.2 × as long as anterior one); posterior claw without backward overhang; lateral crests well developed. Pretarsal processes long, setiform, reaching extremities of claws. ABDOMEN. Distribution of abdominal macrosetae on tergites: 1 + 1 post 1 on I – III (0 + 1 la on III); 1 + 1 la, 2 + 2 post 1,2 on IV; 1 + 1 la, 5 + 5 post 1 – 5 on V – VII; 7 + 7 post on VIII; 9 + 9 post on abdominal segment IX. All post urotergal macrosetae thick, long and covered by thin barbs along distal four-fifths; la urotergal macrosetae shorter than post macrosetae, covered by barbs along distal half. Urosternite I with 9 + 9, urosternites II – VII with 5 + 5; urosternite VIII with 2 + 2 macrosetae; all urosternal macrosetae robust and large, covered by long barbs along distal three-fourths to four-fifths. Apical, subapical and ventromedial setae with abundant long barbs. CERCI. One incomplete cercus on 4 mm long paratype with six articles in addition to basal article, 3.8 mm long; their articles show whorls of long macrosetae covered by thin barbs along distal twothirds, combined with whorls of smooth, thin setae, shorter than macrosetae, in addition to one apical whorl of short setae with a few distal barbs. GENITAL ORGANS. Male urosternite I with up to 160 glandular g 1 setae arranged in up to five rows with slightly enlarged subcylindrical appendages, each bearing up to 19 glandular a 1 setae. Phyletic affinities, habitat and distribution Plusiocampa (P.) rhea sp. nov. has 3 + 3 lateral posterior macrosetae on the mesonotum and metanotum, a feature only found in P. (P.) pouadensis (Condé 1948 c; Sendra & Escolà 2004). Nevertheless, P. (P.) rhea sp. nov. has medial posterior mesonotal and metanotal macrosetae, which are absent in P. (P.) pouadensis. Furthermore, P. (P.) pouadensis is a troglomorphic species from the Eastern Pyrenees, whereas P. (P.) rhea sp. nov. has been found in hypogean habitats, in soil, MSS and the twilight zones of caves at three locations on Crete.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFF4EC63FE68FBF8FAA0F900.taxon	diagnosis	Diagnosis (Condé 1954 b, 1983; Ramellini 1990) Troglomorphic species. Antennae with 26 antennomeres; 12 complex olfactory chemoreceptors in cupuliform organ; frontal process with non-tubercular setae. Pronotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp, 1 + 1 mp macrosetae; metanotum with 1 + 1 ma, 2 + 2 la, 2 + 2 lp, 1 + 1 mp macrosetae; thin, barbed notal macrosetae; sparse notal clothing setae. Two to four ventral tibial macrosetae; slightly unequal claws with large lateral crests, posterior claw with a backward overhang. Urotergite I with 1 + 1 post macrosetae; urotergite II with 2 + 2 post macrosetae; urotergite III with 4 + 4 post macrosetae; urotergite IV with 2 + 2 la, 5 + 5 post macrosetae; urotergite V with 2 + 2 – 3 + 3 la, 5 + 5 post macrosetae; urotergites VI – VII with 3 + 3 la, 5 + 5 post macrosetae; urotergite VIII with 6 + 6 post macrosetae; urosternite I with 7 + 7, urosternites II – VII with 5 + 5, urostenite VIII with 2 + 2 macrosetae. Male unknown. Habitat and distribution Subterranean species found on the Italian Peninsula in two caves, Grotta di Santa Liberata (Toscana) and Grotta dell’Inferniglio (Latium), and in soil habitat on Monti Ausoni, central Apennine Mountains.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFF4EC60FE76F8BBFA99FDF6.taxon	diagnosis	Diagnosis (Blesić 1992) Non-troglomorphic spceies. Antennae with 27 – 28 antennomeres. Pronotum with 1 + 1 ma, 2 + 2 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 2 + 2 la, 2 + 2 lp, 1 + 1 mp macrosetae; metanotum with 1 + 1 ma, 2 + 2 lp, 1 + 1 mp macrosetae. Two or three ventral tibial macrosetae. Subequal claws with large lateral crests, posterior claw with a small backward overhang. Urotergites I – III with 1 + 1 post macrosetae; urotergites IV – VII with 1 + 1 la, 4 + 4 post macrosetae; urotergite VIII with 6 + 6 post macrosetae; abdominal segment IX with 8 + 8 post macrosetae; urosternite I with 7 + 7, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae. This species should be judged with caution due to the fact that it is poorly described. Habitat and distribution Soil-dwelling species distributed in Rudnik Mountain (Serbia) on the west of the Balkan Peninsula.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFF7EC60FE71FDE0FC3BFB19.taxon	diagnosis	Diagnosis Plusiocampa (P.) rybaki was poorly described by Condé, based on specimens from three caves, one in Attica, Greece (Cave I in Rachi, northern part of Mt Ymittos) and two on Crete (Achyrospilo Cave on the Acrotiri Peninsula, Chania and Sarchos Cave, Irakleio). Troglomorphic species. Antennae with 40 – 44 antennomeres, large cupuliform organ with 13 – 21 olfactory chemoreceptors; frontal process with tubercular setae. Mesonotum with 3 + 3 la macrosetae, and mesonotum and metanotum with mp macrosetae. Pretarsus with large lateral crests. Urotergites V – VII with 5 + 5 post macrosetae.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFF7EC60FE71FDE0FC3BFB19.taxon	discussion	Remarks The specimens from Crete reported by Condé (1956 a) as P. (P.) rybaki, as well as two specimens collected by P. Strinati on Crete (deposited in the Natural History Museum of Geneva on two slides labelled by B. Condé as Plusiocampa rybaki: 1 male from Melidoni Spilia, 2 Aug. 1983 and 1 female from Grotta “ Sentoni ”), are here considered to belong to P. (P.) hoffmanni sp. nov.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFF7EC60FE5AFB4BFC21F83B.taxon	diagnosis	Diagnosis (Condé 1953 b, 1981 b, 1983) Troglomorphic species. Antennae with 33 – 38 antennomeres; protruding frontal process with tubercular setae. Pronotum with 1 + 1 ma, 4 + 4 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp, 1 + 1 mp macrosetae; metanotum with 1 + 1 ma, 2 + 2 la, 2 + 2 lp, 1 + 1 mp macrosetae. Two ventral tibial macrosetae; slightly unequal claws (1.15) with large lateral crests, posterior claw with a backward overhang. Urotergites I – III with 2 + 2 post macrosetae; urotergite IV with 1 + 1 (0 + 1) la, 5 + 5 post macrosetae; urotergite V with 1 + 1 (1 + 2) la, 5 + 5 post macrosetae; urotergite VI with 2 + 2 (1 + 2) la, 5 + 5 post macrosetae; urotergite VII with 2 + 2 – 3 + 3 la, 5 + 5 post macrosetae; urotergite VIII with 6 + 6 post macrosetae; abdominal segment IX with 8 + 8 post macrosetae; thick clothing setae on abdominal segments IX – X; urosternite I with 7 + 7, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae. Male without glandular g 1 setae or glandular a 2 setae, with the exception of 1 + 1 g 1 and 2 – 3 a 2 in one specimen; female with subcylindrical appendages with glandular a 1 setae. Habitat and distribution Subterranean species found only in the San Giovanni Caves, Sardinia.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFF6EC7EFE5DFEB3FB43FC1D.taxon	description	Figs 125 – 134	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFF6EC7EFE5DFEB3FB43FC1D.taxon	diagnosis	Diagnosis (Condé 1947 a) Troglomorphic species. Antennae with 22 – 29 antennomeres; cupuliform organ with seven or eight complex olfactory chemoreceptors. Protruding frontal process with tubercular setae. Pronotum with 1 + 1 ma, 2 + 2 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 2 + 2 – 2 + 3 la, 2 + 2 lp macrosetae; metanotum with 1 + 1 ma, 1 + 1 la, 2 + 2 lp macrosetae; long, barbed notal setae; abundant barbed notal clothing setae. Two ventral tibial macrosetae; unequal claws (1.3 – 1.5) with large lateral crests, posterior claw with a backward overhang. Urotergites I – III with 1 + 1 post macrosetae; urotergite IV with 1 + 1 la, 2 + 2 – 3 + 3 post macrosetae; urotergite V with 1 + 1 la, 4 + 4 – 5 + 5 post macrosetae; urotergites VI – VII with 1 + 1 – 2 + 2 la, 5 + 5 (4 + 5) post macrosetae; urotergite VIII with 6 + 6 macrosetae; abdominal segment IX with 8 + 8 – 8 + 9 macrosetae; urosternite I with 7 + 7, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae. Male with an area of glandular g 1 setae, subcylindrical and slightly enlarged appendages with glandular a 1 setae; female appendages like male appendages.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFF6EC7EFE5DFEB3FB43FC1D.taxon	discussion	Remarks The 17 specimens studied, from four caves in the Dinaric Mountains (Bosnia and Herzegovina) (see Table S 2 in Supplementary Material), allowed us to clarify some characters: five complete antennae preserved show 28 – 29 antennomeres, elongated antennomeres with a distal whorl of 12 – 14 long gouge sensilla (30 – 32 µm) (Figs 131 – 132); cupuliform organ with seven or eight spheroidal olfactory chemoreceptors, each with a jagged fold surrounding a central coniform structure covered by a highly visible, multiperforated polygonal net surface (Figs 125 – 130). Protruding frontal process with tubercular setae. Surface of epicuticle reticulate (Fig. 133). Smooth tarsal subapical setae and calcars with long barbs from base, pretarsus claws with large lateral crests (Fig. 134). Furthermore, they show a wider range in the number of urotergal macrosetae than mentioned in the diagnosis above.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFE8EC7FFE45FEB3FC3DFCC3.taxon	diagnosis	Differential diagnosis against the type form The five studied specimens from Karakaya Cave in Turkey (see Table S 2 in Supplementary Material) share the same macrosetal distribution found in the specimens from Bosnia and Herzegovina, with the exception of 7 + 7 post macrosetae on urotergite VIII and 9 + 9 post macrosetae on abdominal segment IX. Furthermore, two other important features separate these two forms, i. e., the shape of the claws (subequal claws and posterior claw with no backward overhang in the form from Karakaya Cave) and features of the frontal process (simple frontal process without tubercular setae in specimens from Karakaya Cave). However, more material is needed to describe these specimens with different features from Karakaya Cave, Mihalliçik as a new species. Habitat and distribution Subterranean species found in five caves of Bosnia and Herzegovina in the Dinaric Mountains. The aff. form was found in a cave in the northeast of the Anatolian Peninsula.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFE8EC7FFE79FCEDFB0DF9C8.taxon	diagnosis	Diagnosis (Condé 1983) Troglomorphic species. Antennae with 36 antennomeres; nine complex olfactory chemoreceptors in cupuliform organ; protruding frontal process with non-tubercular setae. Pronotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp, 1 + 1 mp macrosetae; metanotum with 1 + 1 ma, 2 + 2 la, 2 + 2 lp, 1 + 1 mp macrosetae; long, barbed notal macrosetae; abundant notal clothing setae, smooth or with a few distal barbs. Two ventral tibial macrosetae; subequal claws (1.05) with large lateral crests, posterior claw with a small backward overhang. Urosternite I with 2 + 1 post macrosetae; urotergite II with 2 + 2 post macrosetae; urotergite III with 0 + 1 la, 2 + 2 post macrosetae; urotergites IV – V with 1 + 1 la, 2 + 2 post macrosetae; urotergite VI with 3 + 3 – 2 + 2 la, 4 + 4 post macrosetae; urotergite VII with 3 + 3 la, 4 + 4 post macrosetae; urotergite VIII with 6 + 6 post macrosetae; urosternite I with 8 + 7, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae. Male unknown; subcylindrical appendages with glandular a 1 setae in females. Habitat and distribution Subterranean species found only in Grotta di Gana ‘ e Gortoe, Siniscola, Eastern Sardinia.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFE8EC7CFE6BF9F2FE1AFE95.taxon	diagnosis	Diagnosis (Silvestri 1933 a; Sendra et al. 2006) Non-troglomorphic species. Antennae with 22 – 25 antennomeres; frontal process with non-tubercular setae. Pronotum with 1 + 1 ma, 4 + 4 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 2 + 2 la, 2 + 2 lp macrosetae; metanotum with 1 + 1 ma, 1 + 1 la, 2 + 2 lp macrosetae; long, barbed notal macrosetae. Two ventral tibial macrosetae; slightly unequal claws (1.15) with large lateral crests, posterior claw with no backward overhang. Urotergites I – III with 1 + 1 post macrosetae; urotergite IV with 1 + 1 la, 2 + 2 post macrosetae; urotergites V – VII with 1 + 1 la, 4 + 4 post macrosetae; urotergite VIII with 6 + 6 post macrosetae; abdominal segment IX with 8 + 8 macrosetae; urosternite I with 7 + 7, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae. Male unknown; female with long subcylindrical appendages with glandular a 1 setae.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFEBEC7CFE50FEC7FBAFFC87.taxon	diagnosis	Differential diagnosis against the type form The only specimen collected is a female, a troglomorphic form found in a cave on the island of Cyprus (Coll. AS). This specimen shows several taxonomical differences from the type form. Body length 5.6 mm (metathoracic leg 3.2 mm long); 8.7 mm long antenna with 49 antennomeres; elongated antennomeres with 16 – 18 long gouge sensilla, more than eight complex sensilla in cupuliform organ; 2 or 3 ventral tibial macrosetae; 1 + 1 lateral anterior and 1 + 1 posterior macrosetae on fourth urotergite and 2 + 2 anterior lateral macrosetae on seventh urotergite. Habitat and distribution The type form was found in soil habitat on the island of Ródos (Greece) and at Izmir on the west coast of the Anatolian Peninsula, but the aff. troglomorphic form was found on Cyprus.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFEBEC7DFE70FC29FCF1FC54.taxon	diagnosis	Diagnosis (Stach 1930; Condé 1954 a; Paclt 1957) Non-troglomorphic species. Antennae with 20 – 22 antennomeres. Pronotum with 1 + 1 ma, 2 + 2 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp, 1 + 1 mp macrosetae; metanotum with 1 + 1 ma, 1 + 1 la, 2 + 2 lp, 1 + 1 mp macrosetae. Two ventral tibial macrosetae. Urotergites I – II with 1 + 1 post macrosetae; urotergite III with 2 + 2 post macrosetae; urotergites IV – VII with 1 + 1 la, 5 + 5 post macrosetae; urosternite I with 8 + 8?, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFEBEC7DFE70FC29FCF1FC54.taxon	discussion	Remarks The study of five females and one juvenile collected in four caves (see Table S 2 in Supplementary Material) has allowed the completion of the former description, although new specimens from the type locality are needed for a proper redescription of this species. Nevertheless, a complementary and differential description has been made using the studied material. Non-troglomorphic species (Table 10). Antennae with 21 and 23 doliiform antennomeres, apical antennomere 1.3 × as long as wide, with a small cupuliform organ occupying 1 / 7 of antennomere, with four simple olfactory chemoreceptors. Small ventral sensillum of third antennomere between d and e macrosetae. Non-protruding frontal process with non-tubercular setae. Labial palp suboval with a thick lateral anterior sensillum, two guard setae, up to five normal setae and 75 neuroglandular setae. Pronotum with 2 + 2 la to 3 + 3 la macrosetae; long, barbed notal macrosetae; notal clothing setae long, smooth or with one or two distal barbs. Calcars with two rows of long barbs. Smooth subapical tarsal setae. Slightly unequal claws (1.15 – 1.20) with large lateral crests and a small backward overhang. Urotergite IV with 4 + 4 post macrosetae (5 + 5 according to redescription in Paclt 1957); urotergites VIVII with 2 + 2 la macrosetae (1 + 1 la according to Paclt 1957); urotergite VIII with 6 + 6 post macrosetae; abdominal segment IX with 8 + 8 post macrosetae; urosternite I with 7 + 7, urosternites II – VII with 4 + 4, urosternite VIII with 2 + 2 macrosetae. Stylar setae with a few long, thin bars. Female with subcylindrical appendages bearing up to 9 glandular a 1 setae. Habitat and distribution Apparently, this species only lives in caves, but its non-troglomorphic shape reveals a more superficial habitat (MSS or soils). It has been found in six caves in the southern karst areas of the Western Carpathians, where it was described from the Domica-Baradla cave system in the Slovak and Aggtelek Karst, on the border between Slovakia and Hungary.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFEAEC7DFE24FC06FAF4FA23.taxon	diagnosis	Diagnosis (Silvestri 1933 c; Condé 1954 a) Troglomorphic species. Antennae with 40 – 41 antennomeres. Pronotum with 1 + 1 ma, 2 + 2 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp, 1 + 1 mp macrosetae; metanotum with 1 + 1 ma, 1 + 1 la, 2 + 2 lp, 1 + 1 mp macrosetae; long, barbed macrosetae; abundant notal clothing setae. Three ventral tibial macrosetae; unequal claws (1.4) with large lateral crests, posterior claw with a backward overhang. Urotergites I – III with 1 + 1 post macrosetae; urotergite IV with 1 + 1 la, 5 + 5 post macrosetae; urotergites V – VII with 2 + 2 la, 5 + 5 post macrosetae; urotergite VIII with 6 + 6 post macrosetae; urosternite I with 7 + 7, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae. Male without glandular g 1 setae on first urosternite. Female and male with subcylindrical appendages bearing glandular a 1 setae. Habitat and distribution Subterranean species reported from 10 caves, with a wide distribution area in the Austrian Alps.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFEAEC74FE15F98DFD5DFAD2.taxon	diagnosis	Differential diagnosis against the type form (Condé 1954 a) Troglomorphic subspecies. Antennae with 25 – 28 antennomeres. Until now, the only character used to separate P. (P.) strouhali strouhali from P. (P.) strouhali cavicola has been the number of antennomeres (Condé 1954 a). This characteristic had been used to sort out the subspecies (Vornatscher 1943; Strouhal & Vornatscher 1975), and it was applied to the study material (see Table S 2 in Supplementary Material). On the basis of this weak taxonomic character, the population from Odelsteinhöhle in the Gesäuse National Park, Austria has been classified as P. (P.) strouhali cavicola.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFEAEC74FE15F98DFD5DFAD2.taxon	description	Description Troglomorphic subspecies. Body length: females 3.4 – 7 mm, males 3.45 – 6.15 mm, juveniles 2.95 – 3.7 mm. Epicuticle smooth under optical microscope; clothing setae thin, middle-sized, smooth or with a few distal barbs (Fig. 144). Antennae with 30 – 35 antennomeres in adults (Table 11). Length of antennae equal to or slightly shorter than body length in adults and juveniles. Sensillum of third antennomere subcylindrical, sharply pointed, in ventral position between macrosetae d and e. Central antennomeres 1.6 × as long as wide. Twelve to twenty thin gouge sensilla (28 – 40 µm) in a single distal whorl on each medial and distal antennomere (Figs 137, 139 – 140). Cupuliform organ with 8 – 10 spheroidal olfactory chemoreceptors with a complex central structure surrounded by folds with visible polygonal net with porous surface (Figs 135 – 136, 138). Protruding frontal process with three distal, slightly tubercular macrosetae and a few posterior setae (Figs 142 – 143). Notal macrosetae (Fig. 141): pronotum with 1 + 1 ma, 3 + 3 (3 + 2) la, 2 + 2 lp 2,3; mesonotum with 1 + 1 ma, 3 + 3 la 1,2,3, 2 + 2 lp 2,3, 1 + 1 smp submacrosetae (or 1 + 1 mp); metanotum with 1 + 1 ma, 1 + 1 la 3, 2 + 2 lp 2,3, 1 + 1 mp; all notal macrosetae long, with long barbs along distal four-fifths (Fig. 144); marginal setae slightly longer than clothing setae, with thin barbs along distal half in lateral ones. Legs elongated, metathoracic legs reaching abdominal segment IX. Femur with one long dorsal macroseta, barbed along distal four-fifths. Tibia with 2 – 5 ventral macrosetae, barbed almost from base to tip. Calcars barbed from base to tip. Dorsal and lateral tarsal setae smooth. Unequal claws (1.1 – 1.4) with large lateral crests; ventral surface of claws noticeably ridged, dorsal surface slightly ridged but more evident on backward overhang of posterior claw; pretarsal processes smooth and setiform, with 0 – 4 thin, small basal barbs (Figs 145 – 146, 148). Distribution of abdominal macrosetae on tergites (Figs 149 – 150): 1 + 1 post 1 on I – II; 1 + 1 (0 + 0) la, 1 + 1 post 1 on III; 1 + 1 – 2 + 2 la, 5 + 5 (4 + 5) post 1 – 5 on IV; 2 + 2 la, 5 + 5 post 1 – 5 on V – VII; 6 + 6 (7 + 6) post on VIII; 8 + 8 – 9 + 9 post on abdominal segment IX. All post macrosetae long, covered with long barbs along distal four-fifths; la macrosetae shorter and less barbed than post macrosetae. Urosternite I with 7 + 7, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae. Apical setae of styli smooth, with basal tooth; subapical setae with 2 – 4 long proximal barbs; medial sternal setae with 4 – 6 long medial barbs (Fig. 147). Cerci up to 6.7 mm long (in 4.7 mm long female), with eight primary articles plus basal one, twice the length in juveniles (see Table 11). Articles with whorls of long macrosetae covered with thin barbs along distal two-thirds, combined with whorls of smooth, thin setae shorter than macrosetae In males, urosternite I (Fig. 151) with large subcylindrical appendages, each bearing up to 60 glandular a 1 setae in a distal area. In females, urosternite I (Fig. 152) with subcylindrical appendages thinner than male appendages, each bearing up to 20 glandular a 1 setae in a distal area.. Habitat and distribution Subterranean subspecies found in 21 caves in the Austrian Alps, where it is the common form in caves of the Northern Calcareous Alps. The areas of the two subspecies are overlapping, which casts serious doubt on the validity of the subspecies.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFE3EC74FE1AFA8DFCC2F83A.taxon	diagnosis	Diagnosis (Condé & Mathieu 1957) Non-troglomorphic species. Antennae with 20 – 21 antennomeres; four olfactory chemoreceptors in cupuliform organ; non-protruding frontal process with non-tubercular setae. Pronotum with 1 + 1 ma, 3 + 3 – 2 + 1 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp, 1 + 1 mp macrosetae; metanotum with 1 + 1 ma, 2 + 2 la, 2 + 2 lp, 1 + 1 mp macrosetae; long, barbed notal macrosetae; sparse smooth notal clothing setae. Two or three ventral tibial macrosetae; subequal small claws with small lateral crests, posterior claw with no backward overhang. Urotergites I – II with 2 + 2 post macrosetae; urotergite III with 4 + 4 post macrosetae; urotergites IV with 2 + 2 la, 5 + 5 post macrosetae; urotergites V – VII with 3 + 3 la, 5 + 5 post macrosetae; urotergite VIII with 6 + 6 post macrosetae; abdominal segment IX with 8 + 8 post macrosetae; urosternite I with 7 + 7, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae. Male unknown; female with subcylindrical to conical appendages with glandular a 1 setae. Habitat and distribution The locatity of this species is still not clear, as a label confusion may have occurred between two localities, one endogean habitat in France (Tarn-et-Garonne, La Capelle-Livran, Caylus) and a small cave in Sicily (Scaletta Zanclea, south of Messina).	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFE2EC73FE15FEB3FE3BFBA5.taxon	description	urn: lsid: zoobank. org: act: D 1 AE 6 C 14 - 8920 - 4505 - 9 B 42 - B 518 BB 1 A 6 FB 0 Figs 153 – 156; Table 12	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFE2EC73FE15FEB3FE3BFBA5.taxon	etymology	Etymology The specific name comes from the geographic area where this new species was found, the plateau called Trnovski gozd (Trnovo Forest).	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFE2EC73FE15FEB3FE3BFBA5.taxon	materials_examined	Material examined Holotype SLOVENIA • ♀; Ajdovšćina, Predmeja, Bela Griža 1 n. 7937; 2 Oct. 2014; Špela Borko leg., PMSL Diplura- 001. Paratypes SLOVENIA • 2 ♀♀, 2 ♂♂; same collection data as for holotype; Coll. AS • 7 ♀♀, 2 ♂♂, 3 juv.; Ajdovšćina, Predmeja, Velika ledena jama v Paradani n. 742; Coll. AS • 1 ♂; same collection data as for preceding; PMSL Diplura- 002 • 1 ♀; same collection data as for preceding; PMSL Diplura- 003. Other material See Table S 2 in Supplementary Material.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFE2EC73FE15FEB3FE3BFBA5.taxon	description	Description BODY. Body length 3.9 – 5.4 mm (females), 3.2 – 4.35 mm (males) and 2.7 – 3.7 (juveniles). Epicuticle smooth under optical microscope; body with thin, middle-sized barbed clothing. HEAD. Only two complete antennae, in a juvenile with 40 antennomeres. Bacilliform sensillum of third antennomere in ventral position between macrosetae d and e. Central antennomeres 1.3 × as long as wide. Eighteen to twenty thin gouge sensilla (30 – 34 µm) in a single distal whorl on each medial and distal antennomere; 2 – 3 coniform sensilla in same whorl, but also present in last antennomere. Cupuliform organ on apical antennomere with about eight complex olfactory chemoreceptors. Nonprotruding frontal process, with non-tubercular setae. THORAX. Pronotum with 1 + 1 ma, 4 + 4 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp macrosetae; metanotum with 1 + 1 ma, 2 + 2 (1 + 1, 3 + 2) la, 2 + 2 lp macrosetae; long, barbed macrosetae; long, thin, barbed notal macrosetae; clothing setae smooth or with up to four distal thin barbs. One dorsal femoral macroseta, barbed all along with thin, short barbs; one barbed tibial metathoracic macroseta and none on prothoracic or mesothoracic tibiae; calcars with thin, long barbs in several rows; very unequal claws (1.7 – 1.8) with large lateral crests; posterior claw with a large backward overhang (Figs 153 – 154). ABDOMEN. Urotergites I – III with 1 + 1 post macrosetae; urotergite IV with 1 + 1 (1 + 0) la, 3 + 3 – 2 + 2 post macrosetae; urotergite V with 1 + 1 (0 + 1) la, 4 + 4 post macrosetae; urotergite VI with 1 + 1 la, 4 + 4 post macrosetae; urotergite VII with 2 + 2 (2 + 1) la, 4 + 4 post macrosetae; urotergite VIII with 6 + 6 post macrosetae; abdominal segment IX with 8 + 8 post macrosetae; urosternite I with 7 + 7 – 8 + 8, urosternites II – VII with 6 + 6, urosternite VIII with 2 + 2 macrosetae. Stylus setae with a few barbs, smooth apically, with basal tooth; 3 – 5 thin distal barbs on subapical and mediosternal setae. CERCI. Incomplete cerci, a maximum of 5 primary articles have been observed in several specimens. Each primary cercus bears from six to ten disorganized whorls of long, barbed macrosetae, one distal whorl of smooth, long setae and an apical whorl of short, thin setae. GENITAL ORGANS. Male with two to four rows of up to 200 glandular g 1 setae, subcylindrical appendages with a small distal area of up to ten glandular a 1 setae; female with subcylindrical appendages with up to 8 glandular a 1 setae (Figs 155 – 156). Phyletic affinities, habitat and distribution Several species of Plusiocampa s. str. share important taxonomical features with P. (P.) ternovensis sp. nov., such as the clearly unequal claws with large lateral crests, the posterior claw with a backward overhang and the absence of medial posterior macrosetae on mesonotum and metanotum, but only P. (P.) balsani and P. (P.) affinis share a similar distribution pattern of notal and urotergal macrosetae. Nonetheless, the secondary sexual features of P. (P.) balsani, with the male having subtrapezoidal appendages and glandular a 2 setae, unambiguously isolate it from P. (P.) ternovensis sp. nov. Morphologically and geographically speaking, its closest related species is P. (P.) affinis, but some clear differences separate them. The very unequal claws (1.7 – 1.8), presence of only one ventral tibial macroseta and the 2 + 2 lateral macrosetae on the urotergites are features of the new species that are not present in P. (P.) affinis, which has unequal claws (1.3), two ventral tibial macrosetae and 1 + 1 lateral macrosetae on the urotergites. Plusiocampa (P.) ternovensis sp. nov. has been sampled in three caves at the edge of a karst plateau north of the town of Ajdovščina, in Slovenia. This karst area is in the northern part of the Dinaric Mountains, a transitional zone between the Mediterranean, Alpine, Dinaric and Central European biogeographic regions. In two of the caves, the species lives together with P. (Stygiocampa) nivea, although they segregate differently within the cave. Plusiocampa (P.) ternovensis sp. nov. lives at a wider range of depths, while P. (S.) nivea occupies a deeper zone of the cave, at a depth of 500 m in the Velika ledena jama v Paradani and Bela Griža 1 caves. The temperatures in these two deep caves increase with depth, from permanently under 0 ºC (Velika ledena jama v Paradani, with permanent ice in the upper parts) to more than 4 ºC in parts deeper than 500 m. In Ledenica pri Dolu, an ice cave with temperatures below 2 ºC, we found only P. (P.) ternovensis sp. nov., while in the Bošnarjev brezen Cave (5.9 ºC to 7.0 ºC) we found only P. (S.) nivea. This suggests that P. (P.) ternovensis sp. nov. may be considered better coldadapted than P. (S.) nivea.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFE7EC70FE13FB92FB95F906.taxon	diagnosis	Diagnosis (Sendra et al. 2019) Troglomorphic species. Antennae with 37 – 41 antennomeres. Very slight protruded frontal process with slightly differentiated smooth macrosetae. Pronotum with 1 + 1 ma, 4 + 4 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp, 1 + 1 mp macrosetae; metanotum with 1 + 1 ma, 1 + 1 la, 2 + 2 lp, 1 + 1 mp macrosetae; long, barbed notal macrosetae; body with a small number of thin clothing setae with thin distal barbs. Two or three ventral tibial macrosetae; slightly unequal claws (1.08 – 1.12) with large lateral crests, posterior claw with backward overhang. Urotergites I – III with 1 + 1 post macrosetae; urotergite IV with 1 + 1 la, 3 + 3 post macrosetae; urotergites V – VII with 1 + 1 la, 5 + 5 post macrosetae; urotergite VIII with 6 + 6 (6 + 5) post macrosetae; abdominal segment IX with 8 + 8 post macrosetae; urosternite I with 7 + 7, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae. Stylar setae with a few thin barbs. Male with an area of glandular g 1 setae, subcylindrical appendages with glandular a 1 setae; appendages of female slightly thinner than those of male. Habitat and distribution Subterranean species found in three caves near Siracusa, southeastern Sicily.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFE7EC71FE61F8B9FDDFFDB5.taxon	diagnosis	Diagnosis (Condé 1981 b) Troglomorphic species. Antennae with 39 antennomeres; eight simple olfactory chemoreceptors in cupuliform organ, highly likely regenerated; non-protruding frontal process with non-tubercular setae. Pronotum with 1 + 1 ma, 2 + 2 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 1 + 1 la, 2 + 2 lp macrosetae; metanotum with 1 + 1 ma, 1 + 1 lp macrosetae; long, barbed notal macrosetae. Two ventral tibial macrosetae; unequal claws with large lateral crests. Urotergite IV with 1 + 1 la, 1 + 0 post macrosetae; urotergites V – VII with 1 + 1 la, 3 + 3 post macrosetae; urotergite VIII with 6 + 6 post macrosetae; abdominal segment IX with 8 + 8 post macrosetae; urosternite I with 7 + 7, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae. Male unknown; female with subcylindrical appendages bearing glandular a 1 setae. Habitat and distribution Subterranean species found in Grouffre des Encanaux, Auriol, Sainte-Baume Massif, Bouches-duRhône, southeastern France.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFE6EC71FE22FD28FEFFFB25.taxon	diagnosis	Diagnosis (Bareth & Condé 2001) Troglomorphic species. Antennae with 31, 36 and 38 antennomeres; six simple olfactory chemoreceptors on cupuliform organ; protruding frontal process with tubercular setae. Pronotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp macrosetae; mesonotum 1 + 1 ma, 3 + 3 la, 2 + 2 lp, 1 + 1 mp macrosetae; metanotum with 1 + 1 ma, 2 + 2 la, 2 + 2 lp, 1 + 1 mp macrosetae; long, barbed notal macrosetae; clothing setae abundant, smooth or with a few distal barbs. Two or three ventral tibial macrosetae; unequal claws with large lateral crests, posterior claw with a backward overhang. Urotergites I – III with 1 + 1 post macrosetae; urotergites IV – V with 1 + 1 la, 4 + 4 post macrosetae; urotergites VI – VII with 2 + 2 la, 4 + 4 post macrosetae; urotergite VIII with 6 + 6 post macrosetae; abdominal segment IX with 8 + 8 post macrosetae; urosternite I with 7 + 7, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae. Male unknown. Habitat and distribution Subterranean species known only from the Vodni Pech Cave, Vidin town area, in the Balkan Mountains, Bulgaria.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFE6EC71FDB0FA98FE6BF96A.taxon	diagnosis	Diagnosis (Condé & Bareth 1996; Bareth & Condé 2001) No dorsal femoral macrosetae; without medial posterior notal macrosetae and with an overall reduction in mesonatal and metanotal macrosetae; extra macrosetae on first to eighth urosternites (with the exception of Plusiocampa (Stygiocampa) bureschi Silvestri, 1931 and P. (S.) denisi Condé (1947). Habitat and distribution Subterranean subgenus found almost exclusively in the Dinaric Mountains, but also in part of the Balkan Mountains (Fig. 211).	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFE6EC4EFE66F965FDA2FD73.taxon	diagnosis	Diagnosis (Silvestri 1931 b; Condé 1955 a, 1959; Condé & Bareth 1996; Bareth & Condé 2001) Troglomorphic species. Antennae with 39 – 54 antennomeres; seven to nine complex olfactory chemoreceptors in cupuliform organ. Pronotum with 1 + 1 ma, 2 + 2 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 1 + 1 lp macrosetae; metanotum without macrosetae; long, barbed notal macrosetae; abundant notal clothing setae. No ventral tibial macrosetae; slightly unequal claws (1.1) with large lateral crests, posterior claw with a small backward overhang. Urotergite VII with 2 + 2 post macrosetae; urotergite VIII with 4 + 4 post macrosetae; abdominal segment IX with 6 + 6 – 7 + 7 post macrosetae; urosternite I with 7 + 7, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae. Male without glandular g 1 setae, subcylindrical slightly enlarged appendages with glandular a 1 setae; female with subcylindrical appendages with glandular a 1 setae. Spermatozoid fascicles 50 µm in diameter, 25 µm wide; wheels with 3 ½ turns of 300 – 350 µm and 5.6 µm diamter filament. Habitat and distribution Subterranean species found in 13 caves in a karst region on the left bank of the Iskar River, Sofia District, Balkan Mountains, Bulgaria.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFD9EC4EFE2DFD6EFE27F997.taxon	description	Fig. 202	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFD9EC4EFE2DFD6EFE27F997.taxon	diagnosis	Diagnosis (Condé & Bareth 1996) Troglomorphic species. Antennae with 29 – 33 antennomeres; 12 – 14 complex olfactory chemoreceptors in cupuliform organ; protruding frontal process with non-tubercular setae. Pronotum with 1 + 1 ma, 2 + 2 la, 1 + 1 lp macrosetae; mesonotum with 0 + 0 – 1 + 1 sla macrosetae; metanotum without macrosetae; long, barbed macrosetae; abundant barbed notal clothing setae. No ventral tibial macrosetae; unequal claws (1.35) with large lateral crests, posterior claw with a large backward overhang. Urotergite VII with 0 + 0 – 1 + 1 post macrosetae; urotergite VIII with 3 + 3 – 2 + 2 post macrosetae; abdominal segment IX with 7 + 7 post macrosetae; urosternite I with 21 + 21 – 29 + 25, urosternites II – VII with 11 + 11 – 14 + 12, urosternite VIII with 4 + 4 (3 + 3, 4 + 5) macrosetae. Male without glandular g 1 setae, thick slightly short subcylindrical appendages without glandular a 2 setae; female with subcylindrical appendages without glandular a 2 setae. Spermatozoid fascicles 50 µm in diameter and 10 – 12 µm wide; wheels with 1 ½ turns of 140 – 150 µm long filament.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFD9EC4EFE2DFD6EFE27F997.taxon	discussion	Remarks The study of 26 specimens collected from three caves (see Table S 2 in Supplementary Material) allowed us to add characters to the diagnosis (Coll. AS). Habitat and distribution Subterranean species found in several caves in the Kučaj and Beljanica Mountains, eastern Serbia Balkan System (Fig. 202).	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFD9EC4FFE65F9CAFEACFCB9.taxon	description	Figs 205, 208	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFD9EC4FFE65F9CAFEACFCB9.taxon	diagnosis	Diagnosis (Condé 1959; Bareth 1988; Condé & Bareth 1996) Troglomorphic species. Antennae with 41 – 47 antennomeres; numerous sheet-shaped complex olfactory chemoreceptors in cupuliform organ; protruding frontal process with tubercular setae. Pronotum with 1 + 1 ma, 3 + 3 – 4 + 4 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 2 + 2 la, 2 + 2 lp macrosetae; metanotum with 1 + 1 ma, 1 + 1 la, 2 + 2 lp macrosetae; long, barbed notal macrosetae; long, barbed notal clothing setae. No ventral tibial macrosetae; unequal claws with large lateral crests, posterior claw with a backward overhang. Urotergite III with 0 + 0 – 1 + 1 (1 + 0) la macrosetae; urotergite IV with 1 + 1 la, 0 + 1 post macrosetae; urotergite V with 1 + 1 la, 4 + 4 (3 + 4) post macrosetae; urotergites VI – VII with 1 + 1 la, 4 + 4 post macrosetae; urotergite VIII with 5 + 5 – 6 + 6 post macrosetae; abdominal segment IX with 8 + 8 – 9 + 9 (10 + 9) post macrosetae. Urosternite I with 14 + 15 – 21 + 20, urosternites II – VII with 9 + 9 – 12 + 12, urosternite VIII with 2 + 2 – 3 + 3 macrosetae. Male with lateral expansions of posterior margin occupied by an area of glandular g 1 setae in adults (no expansion in juveniles and little expansion in young males), subcylindrical appendages without glandular a 2 setae; female with subcylindrical appendages with glandular a 1 setae.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFD9EC4FFE65F9CAFEACFCB9.taxon	discussion	Remarks The studied material (see Table S 2 in Supplementary Material) allowed us to make additions to the diagnosis (Coll. AS). Habitat and distribution Subterranean species well distributed around the northwestern Dinaric Mountains, found in 18 caves (Figs 205, 208).	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFD8EC4FFE7CFC34FCD1FA06.taxon	diagnosis	Diagnosis (Condé 1947 a) Troglomorphic species. Antennae with 29 – 34 antennomeres. Pronotum with 1 + 1 ma, 3 + 3 la, 1 + 1 lp macrosetae; mesonotum with 1 + 1 ma, 1 + 1 – 2 + 1 – 2 + 2 la, 1 + 1 lp macrosetae; metanotum with 1 + 1 ma macrosetae; long, barbed notal macrosetae, abundant notal clothing setae, smooth or with a few distal barbs. Two ventral tibial macrosetae. Unequal claws with large lateral crests, posterior claw with a backward overhang. Urotergite IV with 1 + 1 la, 1 + 1 post macrosetae; urotergite V with 1 + 1 la, 2 + 2 post macrosetae; urotergites VI – VII with 1 + 1 la, 4 + 4 post macrosetae; urosternite I with 7 + 7, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae. Male with a large area with glandular g 1 setae, subcylindrical appendages with glandular a 1 setae; female appendages like those of male. Habitat and distribution Subterranean species found in the Pećina u Hamidovoj vrtači and Špela Hajnet (= Špela Maja Hajnet) caves in southwestern Serbia, Dinaric Mountains.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFD8EC48FE74F9B9FE57FE00.taxon	description	Figs 157 – 180, 207	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFD8EC48FE74F9B9FE57FE00.taxon	diagnosis	Diagnosis (Denis 1923; Condé 1947 a, 1959; Silvestri 1947; Bareth 1968) Troglomorphic species. Antennae with 28 – 36 antennomeres; abundant sheet-shaped olfactory chemoreceptors in cupuliform organ; protruding frontal process with tubercular setae. Pronotum with 1 + 1 ma, 3 + 3 – 4 + 4 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 la, 2 + 2 (3 + 2) lp macrosetae; metanotum with 2 + 2 lp macrosetae; long, barbed notal macrosetae; abundant barbed notal clothing setae. No ventral tibial macrosetae; unequal claws (1.35) with large lateral crests, posterior claw with a backward overhang. Urotergite IV with 0 + 0 – 1 + 1 la macrosetae; urotergites V – VII with 1 + 1 la, 4 + 4 post macrosetae; urotergite VIII with 5 + 5 – 6 + 6 post macrosetae; abdominal segment IX with 8 + 8 – 9 + 9 (9 + 10) post macrosetae; urosternite I with 11 + 1 + 11 – 15 + 15 (15 + 16), urosternites II – VII with 11 + 11 – 14 + 14, urosternite VIII with 2 + 2 – 4 + 4 macrosetae. Male without glandular g 1 setae, thick subcylindrical appendages without glandular a 2 setae; female with subcylindrical appendages without glandular a 2 setae.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFD8EC48FE74F9B9FE57FE00.taxon	discussion	Remarks The new material studied (see Table S 2 in Supplementary Material) allowed the addition of minor taxonomic details revealed by SEM microscopy. The epicuticle, apparently smooth in optical microscopy, shows a subtle reticulate surface (Figs 166, 168) covered by well-barbed clothing setae (Figs 167 – 168). The macrosetae are also well barbed, but the multi-barbed urosternal macrosetae (Figs 169 – 170) are remarkable. The gouge sensilla are abundant and as long as 28 µm, with transversal stripes in a V shape (Figs 161 – 162). The cupuliform organ has a narrow hole that does not allow an adequate observation of the olfactory chemoreceptors; however, these can be described under the optical microscope as having a multiple laminal shape and under SEM as a central structure with a pronounced terminal pore, semiradially and irregularly folded (Figs 157 – 160). The protruding frontal process with tubercular setae is clearly visible under SEM (Figs 163 – 165). Claws have well-developed lateral crests with a complex, grooved ventral surface (Figs 171 – 174). The completely barbed abundant macrosetae and setae cover the entire surface of the sternites (Figs 169 – 170). The entire length of the stylar setae is totally covered by barbs (Figs 177 – 178) and the eversible vesicles are large (Fig. 179). Males have large subcylindrical appendages with a large area of glandular a 1 setae (Figs 175 – 176, 180). Habitat and distribution Subterranean species, well distributed around the northwestern Dinaric Mountains, so far found in 75 caves (Fig. 207).	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFDFEC48FE63F99EFC48F883.taxon	diagnosis	Differential diagnosis against the type form (Condé & Bareth 1996) Antennae with 50 antennomeres; urosternite without lateral expansions, with a large area of glandular g 1 setae; enlarged subcylindrical appendages with a distal glandular a 1 seta. Habitat and distribution Subterranean species widespread in 21 caves of karst regions of the southeastern Dinaric Mountains. Plusiocampa (S.) aff. remyi occurs in Vranjača (cave), Croatia.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFDFEC48FE7AFDBBFD2BFA23.taxon	description	Figs 181 – 192	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFDFEC48FE7AFDBBFD2BFA23.taxon	diagnosis	Diagnosis (Condé 1947 a; Condé & Bareth 1996) Troglomorphic species. Antennae with 46 – 52 antennomeres; abundant sheet-shaped olfactory chemoreceptors in cupuliform organ; protruding frontal process with tubercular setae. Pronotum with 1 + 1 ma, 3 + 3 – 4 + 4 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 la, 2 + 2 lp macrosetae; metanotum with 2 + 2 lp macrosetae; barbed notal macrosetae; clothing setae abundant, completely barbed. No ventral tibial macrosetae; unequal claws with large crests, posterior claw with a backward overhang. Urotergite III with 1 + 1 (0 + 1) la macrosetae; urotergite IV with 1 + 1 la macrosetae; urotergites V – VII with 1 + 1 la, 3 + 3 post macrosetae; urotergite VIII with 5 + 5 post macrosetae; abdominal segment IX with 9 + 9 – 11 + 11 post macrosetae; urosternite I with 17 + 17 – 22 + 22, urosternites II – VII with 13 + 13 – 19 + 19, urosternite VIII with 3 + 3 – 6 + 6 macrosetae. Male with lateral expansions of posterior margin occupied by an area of glandular g 1 setae in adults (no expansions in juveniles and small expansions in young males), subcylindrical appendages without glandular a 2 setae; females with long subcylindrical appendages without glandular a 2 setae.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFDFEC48FE7AFDBBFD2BFA23.taxon	discussion	Remarks The studied material (see Table S 2 in Supplementary Material) allowed the addition of information to the diagnosis. The specimens from Njegoš pećina, Njeguši (Kotor, Montenegro) and Đatlo pećina, Lebršnik, Gacko (Bosnia and Herzegovina) revealed new diagnostic characters in SEM microscopy: antennomeres with a whorl of thin gouge sensilla, 40 – 47 µm long with an external transversal striped surface in a V pattern (Figs 181 – 182, 184), claws with large lateral crest with ventral longitudinal grooves separated by reticulate surfaces (Figs 183, 185 – 186). The first urosternite of the adult male has lateral expansions, bearing plenty of glandular g 1 setae (Figs 187 – 188), and abundantly long and completely barbed urosternal macrosetae (Figs 189, 192), like the stylar setae (Fig. 190), and similar to but thinner than the cercal macrosetae (Fig. 191).	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFDFEC47FD82F836FC6BFEAB.taxon	diagnosis	Diagnosis (Bareth & Condé 1984) Mesonotum with 1 + 1 ma, 2 + 2 la, 2 + 2 lp macrosetae; metanotum with 1 + 1 ma, 1 + 1 lp or 1 + 1 ma, 1 + 1 la, 2 + 2 lp macrosetae; without mp notal macrosetae. One dorsal femoral macroseta; without ventral tibial macrosetae. Very unequal, elbow-like claws with large lateral crests, posterior claw with a large backward overhang. Urotergites with at most 1 + 1 lateral anterior and 4 + 5 posterior macrosetae starting from sixth urotergite; urosternite I with 7 + 7, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae. Male with thick, enlarged appendages with a large area of glandular a 1 setae. Habitat and distribution The two species of the subgenus Venetocampa, which inhabit the subterranean habitat, are found in two caves of the Veneto Region, northern Italian Peninsula (Fig. 211).	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFD0EC45FE36FE26FC9DFCDC.taxon	description	urn: lsid: zoobank. org: act: 27 DC 5 E 1 D-B 0 E 4 - 4 BAD-B 12 E- 20 ADF 1 BF 16 FB Figs 193 – 195	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFD0EC45FE36FE26FC9DFCDC.taxon	etymology	Etymology This species is named after Milan Ferran, an enthusiastic caver, discoverer and explorer of the type locality in Ferranova buža (meaning Ferran’s hole), Slovenia.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFD0EC45FE36FE26FC9DFCDC.taxon	materials_examined	Material examined Holotype SLOVENIA • ♂; Vrhnika, Ferran buža; 16 Feb. 2013; T. Delić, T. Novak and M. Feron leg.; MZB 2019 - 1031. Paratype SLOVENIA • 1 juv.; same collecting data as for holotype; Coll. AS.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFD0EC45FE36FE26FC9DFCDC.taxon	description	Description BODY. Body length 3.0 mm (holotype) and 2.8 mm (paratype). Epicuticle smooth under optical microscope; body with sparse middle-sized clothing setae, distal two-thirds with thin barbs. HEAD. Two intact antennae in holotype, slightly longer than body length (3.5 / 3.6 mm), with 32 / 33 antennomeres, in paratype slightly shorter than body length (2.76 / 2.75 mm), with 32 / 32 antennomeres. Small, subcylindrical sensillum of third antennomere in ventral position between c and d macrosetae; central antennomeres 1.7 × as long as wide, apical antennomere 2.0 × as long as wide; cupuliform organ occupies of its length, with 7 or 8 spheroidal olfactory chemoreceptors, each with at least two folds, in cupuliform ⅙ organ; gouge sensilla starting on ventral side of third antennomere, with an apical row of eight gouge sensilla; fourth antennomere with 12 ventral gouge sensilla and 1 dorsal gouge sensillum; fifth antennomere with 12 ventral and 2 dorsal gouge sensilla; sixth antennomere with 13 ventral and 4 dorsal gouge sensilla; seventh antennomere with 15 ventral and 6 dorsal gouge sensilla; eighth antennomere with 14 ventral and 7 dorsal gouge sensilla; ninth antennomere with 12 ventral and 8 dorsal gouge sensilla; after tenth antennomere a complete apical whorl of gouge sensilla on each antennomere, with about 20 on medial antennomeres and 15 on distal ones. Frontal process with nontubercular macrosetae or setae; macrosetae along line of insertion of antennomere and x setae similar in length, with long, thin distal barbs, on distal two-thirds (a / i / p / x with relative lengths of 33 / 30 / 26 / 39 in holotype). Small suboval labial palps with a long subcylindrical sensillum, two guard setae, four normal setae and up to 72 neuroglandular setae. THORAX. Pronotum with 1 + 1 ma, 2 + 2 la 2,3, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 2 + 2 la 2,3, 2 + 2 lp 1,3 macrosetae; metanotum with 1 + 1 ma, 1 + 1 la, 2 + 2 lp 2,3 macrosetae; long notal macrosetae, barbed along distal four-fifths; long marginal setae with barbs along distal two-thirds (Fig. 193). Legs elongated (1.84 mm long in holotype and 1.54 mm in paratype), overpassing end of abdomen. Femur with one long dorsal macroseta, barbed almost from its base, inserted around middle of article. No ventral tibial macrosetae. Calcars with 2 – 3 rows of long barbs from base, with a smooth and narrow lateral side along each calcar. Dorsal and lateral subapical tarsal macrosetae smooth. Very unequal claws, posterior claw 1.6 – 1.8 × as long as anterior one, including a large backward overhang on posterior claw; ventral surface of claws with grooves, dorsal surface including large lateral crests with thin stripes. Setiform pretarsal processes reaching extremities of claws (Fig. 194). ABDOMEN. Distribution of abdominal macrosetae on tergites: 1 + 1 (holotype) or 0 + 0 (paratype) la, 2 + 3 post 3,4,5 on urotergite VII; 5 + 5 post on urotergite VIII; 7 + 7 post on abdominal segment IX. All post urotergal macrosetae long, robust, barbed on distal four-fifths; la urotergal macrosetae much shorter, barbed along distal half (Fig. 195). Urosternite I with 6 + 6, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae; all urosternal macrosetae robust, covered by very long barbs along distal half to three-fourths. Pseudoarticulated stylus with barbed setae with long barbs, apical setae shorter than medial and subapical setae. SECONDARY SEX CHARACTERS. Male urosternite I without glandular g 1 setae, subcylindrical short, enlarged appendages with up to 21 large glandular a 1 setae. Adult female unknown. Phyletic affinities, habitat and distribution Plusiocampa (V.) ferrani sp. nov. is the second representative of the subgenus Venetocampa. It is a cavedwelling species closely related to P. (V.) paoletii, found in the vinicity of Belluno, Italian Alps (Bareth & Condé 1984). Nonetheless, the macrosetal distribution patterns on the nota and urotergites show enough differences to describe these specimens as a new taxon.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFD2EC45FE36FC8EFAD0FA34.taxon	diagnosis	Diagnosis (Bareth & Condé 1984) Troglomorphic species. Antennae with 33 antennomeres; eight to nine simple olfactory chemoreceptors in perhaps a regenerated antennae; non-tubercular setae in frontal process. Pronotum with 1 + 1 ma, 2 + 2 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 2 + 2 la, 2 + 2 lp macrosetae; metanotum with 1 + 1 ma, 1 + 1 lp macrosetae; long, barbed notal macrosetae; abundant notal clothing setae. One dorsal femoral macroseta; no ventral tibial macrosetae; very unequal claws (2.5), posterior claw with a backward overhang, with large lateral crests; short anterior lateral process and long posterior lateral process. Urotergite V with 0 + 1 la macrosetae; urotergite VI with 1 + 1 la, 2 + 2 post macrosetae; urotergite VII with 1 + 1 la, 1 + 1 post macrosetae; urotergite VIII with 4 + 5 post macrosetae; abdominal segment IX with 7 + 7 post macrosetae; urosternite I with 7 + 7, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae. Male without glandular g 1 setae, short and very thick appendages with abundant glandular a 1 setae; female unknown. Habitat and distribution Subterranean species found in Biga de le Agnelezze Cave, Piani Eterni, Belluno, Italian Alps.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFD2EC45FDCAF9A7FD50F811.taxon	diagnosis	Diagnosis (Condé 1955 c) Monotypic genus with 1 + 1 medial anterior, (2 + 2) 3 + 3 lateral anterior, 1 + 1 medial posterior and 2 + 2 lateral posterior macrosetae on mesonotum and with 1 + 1 medial anterior, 1 + 1 (2 + 2) lateral anterior, 1 + 1 lateral posterior and 2 + 2 lateral posterior macrosetae on metanotum. One dorsal femoral macroseta. Two or three ventral tibial macrosetae. Subequal, elbow-like claws with large lateral crests. Laminar pretarsal process with a sharp ending covered with long ventral barbs. Urosternite I with 8 + 8, urosternites IIVII with 6 + 6, urosternite VIII with 2 + 2 macrosetae. Male without glandular g 1 setae, subcylindrical appendages without glandular a 2 setae.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFD5EC42FDB1FEB3FB52FCF2.taxon	diagnosis	Diagnosis (Condé 1955 c) Troglomorphic species. Antennae with 29 – 38 antennomeres; eight to ten olfactory chemoreceptors in cupuliform organ. Long, barbed notal macrosetae; sparse, smooth, long notal clothing setae. One dorsal femoral macroseta; two or three ventral tibial macrosetae; subequal claws (1.04) with large lateral crests, posterior claw without a backward overhang; laminar pretarsus processess with a sharp ending covered with long ventral barbs. Urotergites I – III with 1 + 1 post macrosetae; urotergite IV with 1 + 1 la, 3 + 3 post macrosetae; urotergites V – VII with 3 + 3 (2 + 2) la, 4 + 4 post macrosetae; urotergite VIII with 6 + 6 post macrosetae; urosternite I with 8 + 8, urosternites II – VII with 6 + 6, urosternite VIII with 2 + 2 macrosetae. Male with subcylindrical appendages of first urosternite similar to those of female. Habitat and distribution Subterranean species found in three caves in the karst areas of Lebanon (Fig. 210).	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFD5EC43FF3BFCE6FAA2FBED.taxon	diagnosis	Other genera related to Plusiocampinae, the tachycampoid phyletic lineage Bareth & Condé (1981) originally proposed a unique and different tachycampoid phylogenetic lineage (“ lignée Tachycampoïde ”) for a number of previously described genera: Tachycampa Silvestri, 1936 with one species from a cave in the western Rif Mountains (Silvestri 1936); Paratachycampa Wygodzinsky, 1944 with three species, one from a cave in Nuevo León (Mexico) and two from several caves in the eastern Iberian Peninsula (Wygodzinsky 1944; Bareth & Condé 1981); Juxtlacampa Wygodzinsky, 1944 with two species, one from a cave in Guerrero (Mexico) and another from a cave in Alta Verapaz (Guatemala) (Wygodzinsky 1944); and Jeannelicampa Condé, 1955 with one species from caves in Tell Atlas (Algeria) (Condé 1952, 1954 c). Much later, Condé (1982) proposed a new genus within this phyletic lineage, Oncinocampa Condé, 1982, for a subterranean species from the north of the Iberian Peninsula and another one from Sardinia (Condé 1981 b, 1982). A few years later, the diversity of Oncinocampa was increased with four more species, two from the same karst region in the Cantabric Mountain Range on the northern Iberian Peninsula (Sendra & Condé 1988, Bareth 1989) and two from caves in the Americas, one near São Paulo, Brazil (Condé 1997) and another in Veracruz, Mexico (Sendra et al., 2016). The number of species in Tachycampa has also been expanded with three new cave-dwelling species from Mexico (Sendra et al. 2016) (Fig. 209). Diagnosis (Bareth & Condé 1981; Sendra et al. 2016) Three main morphological features support the phyletic line of the tachycampoid group, although exceptions exist. First, the elbow-like claws with lateral crests (absent in Jeannelicampa stygia Condé, 1952). Second, the notal macrosetal formula: pronotum with 3 + 3 ma, la and lp; mesontum with 2 + 2 ma and la; metanotum with 1 + 1 ma or fewer (with only 1 + 1 lp pronotal macrosetae in Oncinocampa genuitei (Bareth, 1989); Paratachycampa hispanica Bareth & Condé, 1981 and P. penyoensis Bareth & Condé, 1981 have an extra lateral anterior macroseta). Finally, dorsal macrosetae are absent from the femur, with the exception of a small, thin macroseta in O. asonensis Sendra & Condé, 1988 and O. paclti Condé, 1981. Additionally, all species inhabit subterranean ecosystems, except O. paclti, which has non-troglomorphic features.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFD4EC43FDC0FBD7FB54FA1B.taxon	diagnosis	Diagnosis (Condé 1952, 1954 c) Pronotum with 3 + 3 medial anterior, lateral anterior and lateral posterior macrosetae; mesonotum with 1 + 1 medial anterior macrosetae and with or without 1 + 1 lateral anterior submacrosetae; metanotum with or without 1 + 1 medial anterior submacrosetae. No dorsal femoral macrosetae; one ventral tibial macroseta. Simple and slightly curved claws with smooth, laminar lateral process. Urotergal macrosetae start from urotergite V or VII. 9 + 9 macrosetae on first urosternite. Male unknown.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFD4EC40FDCEFA46FEEFFEAB.taxon	diagnosis	Diagnosis (Condé 1952, 1954 c) Troglomorphic species. Antennae with 40 – 48 antennomeres; 20 – 28 olfactory chemoreceptors in cupuliform organ; protruding frontal process with non-tubercular setae. Pronotum with 1 + 1 ma, 1 + 1 la, 1 + 1 lp macrosetae; mesonotum with 1 + 1 short ma macrosetae and 0 + 0 – 1 + 1 sla submacrosetae; metanotum with 0 + 0 – 1 + 1 sma submacrosetae; long, barbed pronotal macrosetae, short mesonotal macrosetae; clothing setae barbed along distal three-fourths. No dorsal femoral macrosetae; one ventral tibial macroseta; subequal, slightly curved claws without lateral crests; narrow, laminar, smooth pretarsal processes. Urotergites V – VII with 0 + 0 – 1 + 1 la macrosetae; urotergite VIII with 2 + 2 – 3 + 3 (2 + 1) post macrosetae; urosternite I with 9 + 9, urosternites II – VII with 6 + 6, urosternite VIII with 1 + 1 – 2 + 2 macrosetae. Female appendages of urosternite I subcylindrical, with glandular a 1 setae; male unknown. Habitat and distribution Subterranean species located in two caves: Bou Iacor and Misserghin Ravine in Oran, Tell Atlas, Algeria (Fig. 210).	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFD7EC40FE7BFB92FBEBF919.taxon	diagnosis	Diagnosis (Sendra & Condé 1988) Troglomorphic species. Epicuticle smooth under optical microscope. Antennae with 53 – 64 antennomeres; protruding frontal process with non-tubercular setae; nine or ten complex olfactory chemoreceptors. Pronotum with 1 + 1 ma, 1 + 1 la, 1 + 1 lp macrosetae; mesonotum with 1 + 1 ma, 1 + 1 la macrosetae; metanotum with 1 + 1 ma macrosetae; long, barbed notal macrosetae; smooth, long clothing setae. Dorsal femoral macrosetae absent; ventral tibial macrosetae absent; subequal, elbow-like claws with small lateral crests; setiform, smooth pretarsal processes. Urotergites V – VII with 1 + 1 la, 1 + 1 lp macrosetae; urotergite VIII with 4 + 4 post macrosetae; abdominal segment IX with 6 + 6 post macrosetae; urosternite I with 7 + 1 + 7 – 11 + 1 + 11, urosternites II – VII with 6 + 6 – 7 + 7 (6 + 9), urosternite VIII with 1 + 1 macrosetae. Male with remarkable large lateral expansion on posterior border with glandular g 1 setae; piriform appendages with glandular a 1 and a 2 setae; subcylindrical appendages with glandular a 1 and a 2 setae. Habitat and distribution Subterranean species found in seven caves in the karst areas around Ason Valley, in the CantabroBasque sector of the Cantabric Mountain Range, northern Iberian Peninsula.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFD7EC40FDCBFE1EFB32FC27.taxon	diagnosis	Diagnosis (Sendra & Condé 1988; Condé 1997; Sendra et al. 2016) Pronotum with 3 + 3 medial anterior, lateral anterior and lateral posterior macrosetae; mesonotum with 2 + 2 medial anterior and lateral anterior macrosetae; metanotum with 1 + 1 medial anterior macrosetae. Dorsal femoral macrosetae absent (with the exception of O. asonensis and O. paclti); ventral tibial macrosetae absent (exceptions: one in O. paclti and O. trajanoae Condé, 1997); elbow-like claws with small lateral crest, without a backward overhang; setiform, smooth lateral process; one pair of lateral anterior and lateral posterior urotergal macrosetae on medial urotergites (absent in O. bolivarurrutiai Sendra & García, 2016); apical, subapical and medial setae of stylus with a few small barbs. Habitat and distribution The six current species of this genus have been found in caves, although two are non-troglomorphic. They have an amphi-Atlantic distribution, with one species in Mexico, one in Brazil, three in the center of the Cantabrian Mountain Range on the Iberian Peninsula and one in Sardinia (Fig. 210).	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFD7EC41FDC1F94CFAA2FE24.taxon	diagnosis	Diagnosis (Condé 1982) Troglomorphic species. Epicuticle smooth under optical microscope. Antennae with 39 – 40 antennomeres; six complex olfactory chemoreceptors in cupuliform organ; non-tubercular frontal process setae. Pronotum with 1 + 1 ma, 1 + 1 la, 1 + 1 lp macrosetae; mesonotum with 1 + 1 ma, 1 + 1 la macrosetae; metanotum with 1 + 1 ma macrosetae; long, thin macrosetae with a few barbs; short, smooth clothing setae. One weak dorsal femoral macroseta; ventral tibial macrosetae absent; unequal, elbow-like claws with very small lateral crests; setiform, smooth pretarsal process. Urotergites VI – VII with 1 + 1 la, 1 + 1 lp macrosetae; urotergite VIII with 3 + 3 post macrosetae; abdominal segment IX with 5 + 5 post macrosetae; urosternite I with 7 + 7, urosternites II – VII with 4 + 4, urosternite VIII with 1 + 1 macrosetae. Male without glandular g 1 setae; short, thick and enlarged appendages with glandular a 1 and a 2 setae; female unknown.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFD6EC41FDA9FD98FCC5FC31.taxon	diagnosis	Differential diagnosis against the type form Two specimens (Coll. AS), in poor condition and difficult to observe under the microscope, were collected in two adjacent caves, Sima de Landubia and Sistema de la Cubada Grande (Burgos) in the southeast of the Cantabrian Mountain Range. Mesonotum and metanotum with lp macrosetae, in addition to some variation in urotergal macrosetae. Nevertheless, new specimens should be collected before establishing a new species. Habitat and distribution Subterranean species known only from Cueva de la Marniosa (Treviso, Cantabria), Picos de Europa, Central Cantabrian Mountain Range. The aff. form was found in two caves in the southeast of the Cantabrian Mountains, northern Iberian Peninsula.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFD6EC41FDC7FBADFBCDF951.taxon	diagnosis	Diagnosis (Bareth 1989) Troglomorphic species. Epicuticle smooth under optical microscope. Non-protruding frontal process, non-tubercular setae. Pronotum with 1 + 1 lp macrosetae, long and poorly barbed; mesonotum and metanotum without macrosetae; smooth, short notal clothing setae. No dorsal femoral macrosetae; ventral tibial macrosetae absent; without developed calcars; very unequal, elbow-like claws with small lateral crests; setiform, smooth pretarsal process. Urotergite IV with 1 + 1 lp macrosetae; urotergites VVII with 1 + 1 la and 1 + 1 lp macrosetae; urotergite VIII with 3 + 3 post macrosetae; abdominal segment IX with 6 + 6 post macrosetae; urosternite I with 7 + 7, urosternites II – VII with 4 + 4, urosternite VIII with 1 + 1 macrosetae. Male without glandular g 1 setae; short, thick, enlarged appendages with glandular a 1 and a 2 setae; female unknown. Habitat and distribution Subterranean species known only from Sima del Trave, collected at a depth of 550 meters, Picos de Europa, Central Cantabrian Mountain Range, northern Iberian Peninsula.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFD6EC5EFDC1F90CFEF2FD21.taxon	diagnosis	Diagnosis (Paclt 1979; Condé 1981 a; Sendra & Condé 1988) Non-troglomorphic species; epicuticle with microdenticles; antennae with 18 – 25 doliiform antennomeres; gouge sensilla thick and short; four to six simple olfactory chemoreceptors in cupuliform organ; non-developed frontal process, with non-tubercular setae. Pronotum with 1 + 1 ma, 1 + 1 la, 1 + 1 lp macrosetae; mesonotum with 1 + 1 ma, 1 + 1 la macrosetae; metanotum with 1 + 1 ma macrosetae; middlesized notal macrosetae with a few distal long barbs, la pronotal and ma macrosetae with one or two distal barbs; short, smooth clothing setae. No dorsal femoral macrosetae; one ventral tibial macroseta; small, subequal, elbow-like claws with very small lateral crests; setiform, smooth pretarsal processes. Urotergites I – IV with 1 + 1 ma macrosetrae; urotergites V – VII with 1 + 1 ma, 1 + 1 la, 1 + 1 lp macrosetae; urotergite VIII with 1 + 1 ma, 3 + 3 post macrosetae; abdominal segment IX with 1 + 1 ma, 5 + 5 post macrosetae; urosternite I with 6 + 6, urosternites II – VII with 4 + 4, urosternite VIII with 1 + 1 macrosetae; male with glandular g 1 setae; small, subtrapezoidal appendages with glandular a 1 and a 2 setae; female with subcylindric appendages with glandular a 1 setae. Habitat and distribution Non-troglomorphic species found in one cave and within soil at two adjacent localities in the east of Sardinia.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFC9EC5EFD95FC8CFE66FAA8.taxon	diagnosis	Diagnosis (Wygodzinsky 1944; Bareth & Condé 1981) Pronotum with 1 + 1 ma, 1 + 1 – 2 + 2 la, 1 + 1 lp macrosetae; mesonotum with 1 + 1 mp, 1 + 1 la macrosetae; metanotum with 1 + 1 ma macrosetae. Dorsal femoral macrosetae absent; ventral tibial macrosetae absent. Claws with lateral crest, with a backward overhang. Pretarsal lateral processes from setiform to laminar and well barbed on ventral side. Urotergal macrosetae start on urotergite III or IV. Stylus with pubescent apical, subapical and medial setae. Extra urosternal macrosetae on first urosternite and 2 + 2 on eighth urosternite. Male with elongate appendages, covered with a large area of glandular a 2 setae. Habitat and distribution All three known species are highly cave-adapted to the deep subterranean ecosystems. The genus Paratachycampa has an amphi-Atlantic distribution, with one species in Mexico and two on the Iberian Peninsula (Fig. 210).	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFC9EC5CFE6BFA12FE66FE38.taxon	description	Figs 196 – 201, 206	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFC9EC5CFE6BFA12FE66FE38.taxon	diagnosis	Diagnosis (Bareth & Condé 1981; Sendra 1988; Sendra & Escolà 2004) Troglomorphic species. Epicuticle smooth. Antennae with 34 – 40 elongated antennomeres; 12 – 16 complex olfactory chemoreceptors in cupuliform organ. Protruding frontal process with abundant tubercular setae. Pronotum with 1 + 1 ma, 2 + 2 la, 1 + 1 lp macrosetae; mesonotum with 1 + 1 ma, 1 + 1 la macrosetae; metanotum with 1 + 1 ma, 1 + 1 la macrosetae; long, thin notal macrosetae with a very few barbs; long, thin smooth clothing setae. Subequal, elbow-like claws with large lateral crests, posterior claw with a backward overhang; laminar, barbed ventral pretarsal processes. Urotergites III – VII with 1 + 1 la macrosetae; urotergite VIII with 4 + 4 post macrosetae; abdominal segment IX with 7 + 7 post macrosetae; urosternite I with 22 + 22, urosternites II – VII with 7 + 7, urosternite VIII with 2 + 2 macrosetae. Male without glandular g 1 setae; overdeveloped appendages with a large area of glandular a 2 setae and an apical area of glandular a 1 setae; subcylindrical female appendages with glandular a 1 and a 2 setae.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFC9EC5CFE6BFA12FE66FE38.taxon	discussion	Remarks The studied material (see Table S 2 in Supplementary Material) allowed us to add taxonomic details revealed by SEM microscopy. Gouge sensilla abundant, as long as 28 µm, with thick transversal stripes with alternating rows of pores between them (Figs 196 – 197). Subequal claws with well-developed lateral crests (Figs 198 – 199). Lateral processes of pretarsus with S-shape, its ventral side covered with long, hooked barbs (Figs 200 – 201). Habitat and distribution Subterranean species found in nine caves throughout the coastal karst reliefs in the east of the Iberian Peninsula (Fig. 206).	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFCBEC5CFE60FDA1FC9DFADF.taxon	diagnosis	Diagnosis (Bareth & Condé 1981; Sendra 1988) Troglomorphic species. Epicuticle smooth under optical microscope. Antennae with 30 – 38 elongated antennomeres. Protruding frontal process well developed, with tuberculated, barbed setae. Pronotum with 1 + 1 ma, 2 + 2 la, 1 + 1 lp macrosetae; mesonotum with 1 + ma, 1 + 1 la macrosetae; metanotum with 1 + 1 ma macrosetae; middle-sized, barbed notal macrosetae; short, barbed clothing setae. Subequal, elbow-like claws with large lateral crests, with backward overhang; laminar, barbed ventral pretarsal processes. Urotergites III – V with 1 + 1 la macrosetae; urotergite VI with 1 + 1 la, 1 + 1 – 2 + 1 lp macrosetae; urotergite VII with 1 + 1 la, 2 + 2 – 3 + 4 lp macrosetae; urotergite VIII with 4 + 4 – 6 + 6 post macrosetae; abdominal segment IX with 6 + 6 – 7 + 7 post macrosetae; urosternite I with 21 + 21 – 22 + 22, urosternites IIVII with 5 + 5, urosternite VIII with 2 + 2 macrosetae. Male without glandular g 1 setae; developed, large appendages with a large area of glandular a 2 setae and an apical area of glandular a 1 setae; subcylindrical female appendages with glandular a 1 and a 2 setae. Habitat and distribution Subterranean species found in two caves, Cova del Penyó (type locality) and the Soterranya Cave (Serra, València), Calderona Massif, eastern Iberian Peninsula.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFCBEC5CFDCDFA82FDFEF801.taxon	diagnosis	Diagnosis (Sivestri 1936; Sendra et al. 2016) Epicuticle with microdenticles on surface. Pronotum with 3 + 3 medial anterior and lateral posterior macrosetae, and lateral anterior submacrosetae; mesonotum with 2 + 2 medial and lateral anterior submacrosetae; metanotum with 1 + 1 medial anterior submacrosetae. Dorsal femoral macrosetae absent. One small ventral tibial macroseta. Subequal, elbow-like claws with large lateral crests, posterior claw without a backward overhang; setiform, smooth pretarsal processes. With 3 + 3 and 6 + 6 posterior macrosetae on urotergite VIII and abdominal segment IX, respectively; urosternite I with 8 + 8 – 11 + 11, urosternites II – VII with 8 + 8, urosternite VIII with 1 + 1 macrosetae. Male urosternite I with glandular g 1 setae; short, thick, large appendages with a large area of glandular a 1 and a 2 setae; female with subcylindrical appendages with glandular a 1 setae. Habitat and distribution The four species of this genus are found in subterranean habitats, in the deep areas of caves. It has an amphi-Atlantic distribution, with three species in Mesoamerica and another one in the Rif Mountains, northwestern Africa (Fig. 210).	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFCAEC5DFDC4FEB3FEFAFB1D.taxon	diagnosis	Diagnosis (Sivestri 1936; Condé 1952) Troglomorphic species. Epicuticle with microdenticles. Antennae with 36 – 38 antennomeres; about 20 small complex olfactory chemoreceptors in cupuliform organ; protruding frontal process, with nontubercular setae. Pronotum with 1 + 1 ma and 1 + 1 lp macrosetae and 1 + 1 sla submacrosetae; mesonotum with 1 + 1 sma and 1 + 1 sla submacrosetae; metanotum with 1 + 1 sma submacrosetae; short, smooth notal macrosetae; short, smooth notal clothing setae. Dorsal femoral macrosetae absent; one small ventral tibial macroseta; subequal, elbow-like claws with large lateral crests, posterior claw without a backward overhang; setiform pretarsal processes. Urotergite VIII with 3 + 3 post macrosetae; abdominal segment IX with 6 + 6 post macrosetae; urosternite I with 8 + 8 – 11 + 11, urosternites II – VII with 8 + 8, urosternite VIII with 1 + 1 macrosetae. Male urosternite I with a wide area of glandular g 1 setae; short, thick, large appendages with a large area of glandular a 1 and a 2 setae; female with enlarged subcylindrical appendages with glandular a 1 setae.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
737287D7FFCAEC5DFDC4FEB3FEFAFB1D.taxon	discussion	Remarks The study of many specimens collected from five new caves (Coll. AS) (see Table S 2 in Supplementary Material) allowed the addition of information to the diagnosis and the inclusion of other minor taxonomic characters: ventral sensillum in third antennomere; labial palps with thick sensilla, two guard sensilla, nine simple setae and about 80 neuroglandular setae; metathoracic leg reaching end of abdomen; calcars with long barbs running up from base in two rows; smooth subapical tarsal setae; stylar setae covered with a few long barbs, apical setae with large basal tooth. Habitat and distribution Subterranean species found in nine caves in the karst areas near Taza in the Middle Atlas Mountains, Morocco.	en	Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante, Reboleira, Ana Sofia P. S. (2020): Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1-138, DOI: 10.5852/ejt.2020.591
