taxonID	type	description	language	source
7316879532290571D1CAFAECFC29FD0D.taxon	description	E t y m o l o g y: The specific epithet is an adjective derived from Hellenes, the ancient name of the Greek. D e s c r i p t i o n: Measurements (in mm) and ratios (range, arithmetic mean) (n = 49): TL: 7.5 - 11.5, 9.7; HL: 1.16 - 1.44, 1.31; HW: 1.32 - 1.72, 1.55; PL: 1.51 - 1.97, 1.78; PW: 1.69 - 2.11, 1.92; EL: 1.00 - 1.28, 1.17; HTiL: 1.28 - 1.67, 1.54; HTaL: 1.07 - 1.32, 1.21; ML: 1.16 - 1.28, 1.21; HW / HL: 1.13 - 1.24, 1.18; PW / HW: 1.17 - 1.32, 1.24; PL / PW: 0.88 - 0.96, 0.93; EL / PL: 0.60 - 0.71, 0.66; HTiL / HTaL: 1.17 - 1.35, 1.27. Female of slightly larger average size than male. External characters (Figs 5 - 7) as in Q. coloratus, except for on average shorter legs (see measurements HTiL and HTaL) and an on average less transverse head. 3: protarsomeres I-IV and shape of sternite VIII similar to those of Q. coloratus; aedeagus (Figs 24, 31 - 33, 43 - 45, 53) smaller than in Q. coloratus (see measuments ML and Fig. 58; practically no overlap) and with distinctly shorter, less slender, and more curved apex of the ventral process; subapical tooth closer to base of apex; paramere shorter and less slender than in Q. coloratus, with usually between 30 and 40 peg-setae. ♀: protarsomeres I-IV weakly dilated, of similar shape as in male; posterior excision of tergite X (Fig. 17) much smaller and less deep than in Q. coloratus; styli of segments IX and X shorter and less slender. C o m p a r a t i v e n o t e s: For characters distinguishing this species from Q. coloratus see the description above. From the species treated below, this species differs not only by the shape of the aedeagus, but also by the absence of additional punctures near the posterior frontal puncture of the head. C o m m e n t: In contrast to the material from all other localities, the male from Oros Pilion (Thessalía) has two additional punctures near one of the posterior frontal punctures. Unfortunately, the specimen had been dissected prior to the present study and the aedeagus is damaged, so that neither its original size nor its shape can be fully assessed. Other measurements and external characters are within the range of intraspecific variation of Q. hellenicus. D i s t r i b u t i o n a n d n a t u r a l h i s t o r y: The currently known distribution ranges from North Greece (Ioannina) to the south of the Pelopónnisos (Map 1). The records of Q. coloratus from Corfu, Thasos (COIFFAIT 1976), and Albania (GRIDELLI 1924) may refer to Q. hellenicus. Since there is no reference material in the museum in Vienna, the record of Q. coloratus from the Pelopónnisos by SCHEERPELTZ (1931) may be based on a misidentification. The vast majority of specimens was collected with subterranean pitfall traps. For details on this method and the habitat see GIACHINO & VAILATI (2010). Some specimens were sifted from litter, all of them as singletons. The remainder was most likely found under (large) stones. These observations suggest a subterranean habitat, possibly the MSS stratum, which would also explain the low number of records of a rather large species in a relatively well-studied region. The altitudes range from 100 to 2250 m.	en	Assing, Volker (2017): On Quedius coloratus FAUVEL, 1875 and allied species, with an appendix on Quedius species collected in Greece with subterranean pitfall traps, and a new synonymy (Coleoptera: Staphylinidae: Staphylininae). Linzer biologische Beiträge 49 (1): 207-228, DOI: 10.5281/zenodo.5356693
73168795322F0573D1CAFCE7FC19FE01.taxon	description	E t y m o l o g y: The specific epithet (Latin, adjective: obtuse) alludes to the apically convex median lobe of the aedeagus (lateral view). D e s c r i p t i o n: Measurements (in mm) and ratios (range, arithmetic mean) (n = 13): TL: 8.6 - 12.9, 10.7; HL: 1.21 - 1.48, 1.34; HW: 1.44 - 1.81, 1.63; PL: 1.67 - 2.04, 1.86; PW: 1.83 - 2.25, 2.03; EL: 1.09 - 1.32, 1.21; HTiL: 1.44 - 1.83, 1.63; HTaL: 1.16 - 1.37, 1.27; ML: 1.32 - 1.46, 1.39; HW / HL: 1.17 - 1.26, 1.21; PW / HW: 1.21 - 1.29, 1.25; PL / PW: 0.89 - 0.94, 0.92; EL / PL: 0.61 - 0.68, 0.65; HTiL / HTaL: 1.20 - 1.40, 1.29. Female of slightly larger average size than male. External and sexual characters similar to those of Q. coloratus, except as follows: Legs somewhat darker, dark-reddish to reddish-brown; antennae dark-brown with the basal three antennomeres reddish. Head (Figs 11 - 12) with additional 1 - 3 smaller punc- tures near posterior frontal puncture and postero-dorsal margin of eye on either side (occasionally, these additional punctures may be completely absent on one side). Body on average larger (Fig. 58). Elytra (Fig. 10) on average slightly shorter (see ratio EL / PL). Protarsomeres more strongly dilated both in male and in female (Fig. 13). 3: aedeagus (Figs 25, 34 - 36, 46 - 47, 54) larger than in Q. coloratus (see measuments ML and Fig. 58; little overlap); median lobe apically stouter, not acute but convex, and somewhat curved dorsad in lateral view, broader in ventral view; paramere (Figs 36, 54) broader and with approximately 35 peg-setae arranged in a pair of distinctly separated clusters. ♀: tergite X (Fig. 18) apically with much smaller and less deep posterior excision; styli of segments IX-X shorter and less slender (similar to Fig. 22). HL * HW + PL * PW C o m p a r a t i v e n o t e s: Quedius hebes is reliably distinguished from Q. coloratus particularly by the presence of additional punctures near the posterior frontal puncture on the head, by the different morphology of the aedeagus (see above), and by the much smaller posterior excision of the female tergite X. For characters separating it from the geographically close Q. spiculatus see the following section. D i s t r i b u t i o n a n d n a t u r a l h i s t o r y: Thedistributionrangesacross southern Turkey from Muğla in the west to Adana in the east (Map 1), where it appears to parapatrically border on that of Q. coloratus. A previous record of Q. coloratus from Mersin (SMETANA 1967) refers to Q. hebes (see the paratype from " Bürücek "); the same probably applies to a record of Q. coloratus from Isparta (JAPOSHVILI & ANLAŞ 2011). The type specimens were found under stones or sifted from leaf litter at altitudes of 40 - 1870 m. All of them were collected as singletons, suggesting that the records are accidental and that this species is not an inhabitant of the leaf litter, but of a subterranean habitat. Three teneral specimens were collected in March, April, and May.	en	Assing, Volker (2017): On Quedius coloratus FAUVEL, 1875 and allied species, with an appendix on Quedius species collected in Greece with subterranean pitfall traps, and a new synonymy (Coleoptera: Staphylinidae: Staphylininae). Linzer biologische Beiträge 49 (1): 207-228, DOI: 10.5281/zenodo.5356693
73168795322D0573D1CAFDE4FD05FA01.taxon	description	E t y m o l o g y: The specific epithet is the past participle of the Latin verb spiculare (to sharpen) and alludes to the apically acute median lobe of the aedeagus in lateral view. D e s c r i p t i o n: Measurements (in mm) and ratios (range, arithmetic mean) (n = 8): TL: 9.3 - 12.3, 10.2; HL: 1.25 - 1.39, 1.34; HW: 1.53 - 1.69, 1.62; PL: 1.76 - 1.95, 1.85; PW: 1.90 - 2.11, 2.03; EL: 1.09 - 1.25, 1.18; HTiL: 1.55 - 1.69, 1.62; HTaL: 1.28 - 1.37, 1.30; ML: 1.35 - 1.42, 1.38; HW / HL: 1.19 - 1.23, 1.21; PW / HW: 1.24 - 1.27, 1.25; PL / PW: 0.89 - 0.94, 0.91; EL / PL: 0.59 - 0.67, 0.64; HTiL / HTaL: 1.18 - 1.31, 1.25. Female of slightly larger average size than male. External (Fig. 14) and sexual characters similar to those of Q. hebes, except as follows: 3: median lobe of aedeagus (Figs 26, 37 - 38, 48) apically acute and not bent dorsad in lateral view, also more acute in ventral view; paramere (Figs 39, 55) with more than 40 peg-setae. ♀: tergite X (Fig. 19) of similar shape as in Q. hebes, but posterior excision slightly larger; segments IX and X as in Fig. 22. C o m p a r a t i v e n o t e s: Quedius spiculatus is reliably distinguished from Q. hebes only by the shape of the median lobe of the aedeagus, especially in lateral view. It differs from Q. coloratus by the presence of additional punctures near the posterior frontal puncture on the head, by the different morphology of the larger aedeagus (little overlap; see measurements ML and Fig. 58), and by the much smaller posterior excision of the female tergite X. D i s t r i b u t i o n a n d n a t u r a l h i s t o r y: This species is currently known only from four localities in Samsun and Tokat provinces, North Turkey. All the specimens were sifted from leaf litter of mixed deciduous forests (mostly composed of Quercus and Carpinus) at altitudes of 800 - 1000 m.	en	Assing, Volker (2017): On Quedius coloratus FAUVEL, 1875 and allied species, with an appendix on Quedius species collected in Greece with subterranean pitfall traps, and a new synonymy (Coleoptera: Staphylinidae: Staphylininae). Linzer biologische Beiträge 49 (1): 207-228, DOI: 10.5281/zenodo.5356693
73168795322D0572D1CAF9E4FDAFFB7D.taxon	description	C o m m e n t: The female paratype was recorded as Q. coloratus by ASSING (2016 a). E t y m o l o g y: The specific epithet is an adjective derived from Carpathus, the Latin name for Karpathos. D e s c r i p t i o n: Measurements (in mm) and ratios (range, arithmetic mean) (n = 5): TL: 9.1 - 10.0, 9.6; HL: 1.23 - 1.32, 1.27; HW: 1.46 - 1.58, 1.50; PL: 1.67 - 1.79, 173; PW: 1.76 - 1.95, 1.86; EL: 1.07 - 1.28, 1.15; HTiL: 1.37 - 1.53, 1.47; HTaL: 1.16 - 1.23, 1.19; ML: 1.21 - 1.28, 1.24; HW / HL: 1.18 - 1.19, 1.19; PW / HW: 1.21 - 1.27, 1.24; PL / PW: 0.89 - 0.95, 0.93; EL / PL: 0.64 - 0.71, 0.67; HTiL / HTaL: 1.10 - 1.32, 1.27. External (Fig. 15) and sexual characters similar to those of Q. hebes, except as follows: Body on average smaller (see measurements; little overlap). 3: aedeagus (Figs 27, 40, 49, 56) distinctly smaller than in Q. hebes (see measuments ML and Fig. 58; no overlap); median lobe apically more acute and not curved dorsad in lateral view; paramere constricted medially. ♀: tergite X (Fig. 20) apically with conspicuously small posterior excision. C o m p a r a t i v e n o t e s: Among the species with additional punctures near the posterior frontal puncture on the head and with more strongly dilated protarsomeres I-IV, Quedius carpathius is characterized by relatively small body size, a small pronotum, short tibiae, and particularly by the shape of the aedeagus (paramere constricted medially) and by the smaller posterior excision of the female tergite X. D i s t r i b u t i o n a n d n a t u r a l h i s t o r y: Currently available evidence suggests that Q. carpathius is endemic to Karpathos (Map 1). Previously, ten endemic species of Staphylinidae (three of them undescribed) belonging to eight genera were known to occur in this island (ASSING 2016 a). The type specimens were collected in five localities, four of them in the north of Karpathos, at altitudes between 250 and at least 450 m. The female paratype was found under a stone in a dry ruderal habitat. The remaining specimens may have been collected from under stones, too. One specimen taken at the end of April is teneral.	en	Assing, Volker (2017): On Quedius coloratus FAUVEL, 1875 and allied species, with an appendix on Quedius species collected in Greece with subterranean pitfall traps, and a new synonymy (Coleoptera: Staphylinidae: Staphylininae). Linzer biologische Beiträge 49 (1): 207-228, DOI: 10.5281/zenodo.5356693
73168795322C056DD1CAFB28FE6EFD93.taxon	description	C o m m e n t: The above male may represent a distinct species. However, with only one male in somewhat poor condition available and since a male (in even worse condition) without additional punctures and longer elytra was found in Oros Pelion, too, the status of this material requires clarification based on more material, to rule out the possibility that the above male represents an aberrant specimen of Q. hellenicus. It is distinguished from the material of the described species treated in the present paper by shorter elytra, hind wings of reduced length, and by the morphology of the aedeagus, additionally also by smaller body size (Fig. 58) and shorter tarsi. The locality is situated in Oros Pelion [39 ° 24 ' N, 23 ° 02 ' E] (Greece: Thessalia) (Map 1) at an altitude of 400 m. D e s c r i p t i o n: Measurements (in mm) and ratios: TL: 10.2; HL: 1.21; HW: 1.44; PL: 1.65; PW: 1.76; EL: 0.86; HTiL: 1.39; HTaL: 1.00; ML: 1.18; HW / HL: 1.19; PW / HW: 1.23; PL / PW: 0.93; EL / PL: 0.52; HTiL / HTaL: 1.40. External (Figs 8 - 9) and sexual characters similar to those of Q. hebes, Q. spiculatus, and Q. carpathius, except as follows: Body smaller (practically no overlap) (see measurements) and with very short tarsi (see measument HTaL and ratio HTiL / HTaL). Elytra (Fig. 8) very short (see measurement EL and ratio EL / PL). Hind wings present, but of reduced length. 3: aedeagus (Figs 60 - 61, 57) smaller than in other species; median lobe somewhat resembling that of Q. hellenicus in lateral view, but broader in ventral view; paramere (Fig. 57) broad and rather short, with approximately 30 peg-setae arranged in a pair of distinctly separated clusters.	en	Assing, Volker (2017): On Quedius coloratus FAUVEL, 1875 and allied species, with an appendix on Quedius species collected in Greece with subterranean pitfall traps, and a new synonymy (Coleoptera: Staphylinidae: Staphylininae). Linzer biologische Beiträge 49 (1): 207-228, DOI: 10.5281/zenodo.5356693
731687953233056DD1CAFD92FBA8FC41.taxon	description	C o m m e n t: The above females are clearly not conspecific with Q. coloratus, as which they were recorded by ASSING (2016 c). They are distinguished from Q. coloratus by a small posterior excision of the female tergite X and from the geographically close Q. hebes and Q. spiculatus by the absence of additional punctures near the posterior frontal puncture on the head. Males would be needed to clarify their specific identity.	en	Assing, Volker (2017): On Quedius coloratus FAUVEL, 1875 and allied species, with an appendix on Quedius species collected in Greece with subterranean pitfall traps, and a new synonymy (Coleoptera: Staphylinidae: Staphylininae). Linzer biologische Beiträge 49 (1): 207-228, DOI: 10.5281/zenodo.5356693
731687953232056FD1CAFE2CFB96FE90.taxon	description	C o m m e n t: Originally described from the Perister (= Pelister) range in Macedonia, this species was recorded from Greece (Oros Vitsi) for the first time by ASSING & WUNDERLE (2001). Additional records from the same region were reported by ASSING (2006) and SOLODOVNIKOV (2005). The currently known distribution is illustrated in Map 2. Whether this species is a true inhabitant of the MSS stratum is doubtful. The previously published material from Greece was collected by sifting leaf litter in calcareous beech forests and debris near snowfields. The altitudes range from 1340 to 2040 m.	en	Assing, Volker (2017): On Quedius coloratus FAUVEL, 1875 and allied species, with an appendix on Quedius species collected in Greece with subterranean pitfall traps, and a new synonymy (Coleoptera: Staphylinidae: Staphylininae). Linzer biologische Beiträge 49 (1): 207-228, DOI: 10.5281/zenodo.5356693
731687953231056FD1CAFE94FD9CFDD5.taxon	description	C o m m e n t: According to SCHÜLKE & SMETANA (2015), this widespread species had not been recorded from Greece.	en	Assing, Volker (2017): On Quedius coloratus FAUVEL, 1875 and allied species, with an appendix on Quedius species collected in Greece with subterranean pitfall traps, and a new synonymy (Coleoptera: Staphylinidae: Staphylininae). Linzer biologische Beiträge 49 (1): 207-228, DOI: 10.5281/zenodo.5356693
731687953231056FD1CAFDDFFC84FA65.taxon	description	C o m m e n t: The previously known distribution was confined to two localities in Evrytanía (ASSING 2007; SCHÜLKE & SMETANA 2015). The new records (see also Map 2) reveal that Q. endogeus is widespread in Greece (Pindos range, northern Pelopónnisos, Evvoia). The morphological adaptations (reduced eye size, pigmentation, and hind wings; slender habitus; slender pronotum), the complete absence of records other than those with subterranean traps, and the frequent catches with these traps suggest that Q. endogeus is a true and exclusive inhabitant of the MSS stratum.	en	Assing, Volker (2017): On Quedius coloratus FAUVEL, 1875 and allied species, with an appendix on Quedius species collected in Greece with subterranean pitfall traps, and a new synonymy (Coleoptera: Staphylinidae: Staphylininae). Linzer biologische Beiträge 49 (1): 207-228, DOI: 10.5281/zenodo.5356693
