taxonID	type	description	language	source
732487F5BD4C6E79EBEFA1B7FD8EFD35.taxon	materials_examined	Type locality. Yemen, Socotra Island, Al Haghier Mts., wadi Madar. Type material. Holotype — (13 NMPC): YEMEN, SOCOTRA Island, Al Haghier Mts., wadi Madar, 1180 – 1230 m, 12 ° 33.2 ’ N, 54 ° 00.4 ’ E, 13 – 14. xi. 2010, J. Bezdĕk lgt. The paratypes — (1 Ƥ NMPC, 13 + 3 Ƥ LPCB): same data as holotype; (23 + 1 Ƥ LPCB): same data as holotype, L. Purchart lgt.; (13 JFCP): Yemen, Soqotra Is.: 28. – 29. xi. 2003, HOMHIL protected area, N 12 ° 34 ’ 27 ” E 54 ° 18 ’ 32 ”, 364 m [GPS]; Jan Farkač lgt. // YEMEN — SOQOTRA 2003, Expedition; Jan Farkač, Petr Kabátek & David Král. Note. The altitudes of the type locality and the locality of the last mentioned paratype differ markedly. Therefore some doubts exist regarding the paratype´s locality data. Differential diagnosis. Beside the characters stated in the key, Deretus bezdeki sp. nov. can easily be distinguished from the remaining species of the genus by the presence of large tubercles in the elytral interstriae. D. hulai sp. nov. also possesses tubercles but very small ones and furthermore differs from D. bezdeki sp. nov. in the shape of pronotum. In the latter species the pronotum is distinctly transverse with extremely produced anterior corners and with five large acute teeth situated on each side of pronotum (Fig. 13). In D. hulai sp. nov. the pronotum is not transverse but more or less square-formed, anterior angles not extended and lateral margin of pronotum with only several small spine-like teeth (Fig. 15).	en	Purchart, Luboš (2012): Biodiversity research of darkling beetles on Socotra Island. Part I. The genus Deretus Gahan, 1900 (Coleoptera: Tenebrionidae). Zootaxa 3153: 57-68, DOI: 10.5281/zenodo.211332
732487F5BD4C6E79EBEFA1B7FD8EFD35.taxon	description	Description. Holotype, body length 5.8 mm, width 2.5 mm. Variability of size in paratypes — females: 5.3 – 7.5 x 2.3 – 3.3 mm; males: 5.0 – 5.4 x 2.2 – 2.3 mm. Body dark brown, shiny (Fig. 1). Head densely punctate with punctures as large as eye facet. The space between punctures narrower or equal to their diameter. Eyes reniform, not divided by genae. Clypeus with almost straight anterior margin, covered with yellowish setae anteriorly. Labrum transverse, covered with long yellowish setae. Ventral side of head shiny and punctate. Mentum transverse with midlongitudinal and smooth gibbosity. Apical labial palpomere broadened, triangular in both sexes. Antennae filiform, relatively long, covered by whitish setae. Third antennomere prolonged, distinctly longer than the following two joints taken together. Apical antennomere pale brown, longer in males. Pronotum (Fig. 13) transverse, punctate with same punctation as head. Anterior corners extremely extended (elongated), spine-like. Each lateral margin with five very large and acute teeth. Base of pronotum distinctly bordered, lateral and anterior margins bordered slightly. Anterior margin medially obliterated. Elytra ovoid, sparsely covered with short yellowish setae, slightly convex (lateral view) with gentle apical declivity (lateral view). Base of elytra bordered only around humeri. Scutellum subtriangular. Elytral striae conspicuous, composed of punctures which are slightly smaller than eye facet. All elytral interstriae shagreened, covered irregularly, especially in lateral and apical parts of elytra, with two kinds of tubercles — very small and large ones. Elytral epipleuron relatively broad and very finely punctate. Prosternum, hypomeron, mesoventrite, mesepisternum, mesepimeron and metaventrite glabrous, smooth, densely and coarsely punctate. Prosternal process widened behind coxae (ventral view). Abdominal ventrites finely and shallowly punctate, pubescent, similarly to surface of elytra. In males without patches of sensillae. Legs covered with yellowish setae. Tibiae densely and roughly punctate. In males first three protarsomeres wider than penultimate and apical protarsomeres. Further modifications not present in either sex. Aedeagus see Fig. 19, 24.	en	Purchart, Luboš (2012): Biodiversity research of darkling beetles on Socotra Island. Part I. The genus Deretus Gahan, 1900 (Coleoptera: Tenebrionidae). Zootaxa 3153: 57-68, DOI: 10.5281/zenodo.211332
732487F5BD4C6E79EBEFA1B7FD8EFD35.taxon	etymology	Etymology. Dedicated to my dear colleague and friend Dr. Jan Bezdĕk (Czech Republic), expert in Chrysomelidae and collector of type specimens.	en	Purchart, Luboš (2012): Biodiversity research of darkling beetles on Socotra Island. Part I. The genus Deretus Gahan, 1900 (Coleoptera: Tenebrionidae). Zootaxa 3153: 57-68, DOI: 10.5281/zenodo.211332
732487F5BD4E6E7EEBEFA673FF0CFA39.taxon	materials_examined	Type locality. Yemen, Socotra Island, Jena-agahan. Type material examined. Deretus denticollis: Lectotype (designated here) — (Ƥ BMNH): Type {white and red, rounded, printed} // Jena-agahan, Sokotra. 1200 feet, Jan. 99, W. R. O. Grant, 99 - 85 {white, printed} // Deretus denticollis Gahan, Type {white, handwritten}. Deretus wraniki: Holotype — (13 SMNS): ZOOLOGISCHE EXKURSION, Socotra, Diksam, 11. 1997, leg. Wranik {white label, handwritten} // HOLOTYPUS, Deretus wraniki n. sp., Schawaller 2000 {red label, printed}; Paratype [actually D. maderai sp. n.] – (1 Ƥ SBSC): YEMEN E, Socotra Isl., 6. – 24. ix. 1999, V. Bejček & K. Šťastný lgt. {printed} // COLLECTION STANISLAV BEČVÁŘ {yellow, printed} // PARATYPUS, Deretus wraniki n. sp., SCHAWALLER 2000 {red, printed}. Additional material. (43 + 4 Ƥ LPCB): Yemen, Socotra Island, wadi Ayhaft, 12 ° 36.5 ’ N, 53 ° 58.9 ’ E, 200 m, 7 – 8. xi. 2010, L. Purchart lgt.; (43 + 6 Ƥ LPCB): same data as previous, but J. Bezdĕk lgt.; (1 Ƥ JBCP): same data as previous, but Jan Batelka leg.; (53 + 4 Ƥ NMPC): same data as previous, but Jiří Hájek leg.; (13 LPCB): same data as previous, but J. Bezdĕk lgt., Alcohol 70 % + acetic acid; (1 Ƥ SBCB): Yemen, Socotra Isl., Hadiboh env., 6. – 24.1999, V. Bejček & K. Šťastný lgt.; (13 SBCB): Socotra Isl. (Yemen), Hamadero, 20. – 21. xi. 2000, lgt. V. Bejček + K. Šťastný; (13 SBCB): Socotra Isl. (Yemen), Ayhaft, 3. xi. 2000, lgt. V. Bejček + K. Šťastný; (23 SBCB): Socotra Isl. (Yemen), Lahas, 17. – 18. xi. 2000, lgt. V. Bejček + K. Šťastný; (23 + 2 Ƥ NMPC): Yemen, Soqotra Is., 21. xi. – 12. xii. 2003, Hadiboh env., ca 10 – 100 m, N 12 ° 65 ’ 02 ’’ E 54 ° 02 ’ 04 ’’ [GPS], David Král lgt. // Yemen – Soqotra, 2003, Expedition; Jan Farkač, Petr Kabátek & David Král; (23 + 4 Ƥ NMPC): Yemen, Soqotra Is., 24 – 26. xi. 2003, wadi Ayhaft, 190 m, N 12 ° 36 ’ 38 ” E 53 ° 58 ’ 49 ” [GPS], David Král lgt. // Yemen – Soqotra, 2003, Expedition; Jan Farkač, Petr Kabátek & David Král; (13 JFCP): same as previous, but Jan Farkač lgt.; (13 NMPC): Yemen, Soqotra Is., Homhil protected area, 28 – 29. xi. 2003, 364 m, N 12 ° 34 ’ 27 ” E 54 ° 18 ’ 32 ” [GPS], David Král lgt. // Yemen – Soqotra, 2003, Expedition; Jan Farkač, Petr Kabátek & David Král; (33 JFCP): same as previous, but Jan Farkač lgt.; (1 Ƥ JFCP): Yemen, Soqotra Is., 6. – 7. xii. 2003, Noged plain: wadi Ireeh, N 12 ° 23 ’ 11 ” E 53 ° 59 ’ 47 ”, 95 m [GPS]; Jan Farkač lgt. // Yemen – Soqotra, 2003, Expedition; Jan Farkač, Petr Kabátek & David Král; (23 NMPC): Yemen, Socotra Island, Aloove area, Hassan vill. env., 12 ° 31.2 ’ N, 54 ° 07.4 ’ E, 221 m, Jiří Hájek leg., 9. – 10. xi. 2010; (1 ƤSMNS): Socotra Archipelago, Socotra, Hadibo, 1. xi. 2008, leg. T. van Harten; (33 + 1 Ƥ SMNS): Socotra Archipelago, Socotra, Wadi Danegan, 90 m, 30. x. 2000, leg. T. van Harten; (1 Ƥ SMNS): Yemen, Soqotra Archipelago, Soqotra, Noged, Farnihin, Steroh wadi, 12 ° 24 ’ 26 ‘‘ N, 54 ° 08 ‘ 10 ‘‘ E, 24. x. 2010, leg. T. van Harten, SOQ 2000 / 06.	en	Purchart, Luboš (2012): Biodiversity research of darkling beetles on Socotra Island. Part I. The genus Deretus Gahan, 1900 (Coleoptera: Tenebrionidae). Zootaxa 3153: 57-68, DOI: 10.5281/zenodo.211332
732487F5BD4E6E7EEBEFA673FF0CFA39.taxon	discussion	Remarks. D. denticollis was described by Gahan (1900) probably based on a single female, as the only known type is a female deposited in the BMNH. However, Gahan stated neither the number of described specimens nor their deposition. For that reason I designate this specimen as a lectotype. Koch (1970) made a detailed description of a single male specimen deposited in the Frey collection (Natural History Museum, Basel) and designated it as the allotype of D. denticollis. However, this is not a valid name-bearing type (ICZN 1999: 72 A) and therefore it cannot be used as a type specimen. Koch undoubtedly believed that this male belonged to the same species as the above mentioned female type specimen in the BMNH. Much later Schawaller (2004) listed an additional 4 females as D. denticollis that were compared with this non-type male described by Koch. The same author also described Deretus wraniki Schawaller, 2004 which was differentiated from D. denticollis mainly by its very small body size. During expeditions to Socotra Island the author collected, with his colleagues, a larger series of Deretus specimens comprising males and females of several species. These specimens were compared with the type material of D. denticollis and D. wraniki and the non-type male of D. denticollis from the Frey collection. The author discovered that the male holotype of D. wraniki is identical to the lectotype of D. denticollis and therefore D. wraniki should be synonymised. He also observed, based on a large series of this species, that the body size of this species is strongly variable and cannot be used for differential diagnoses. The holotype of D. wraniki simply represents one extreme of the body-size range. It was also found that the female paratype of D. wraniki is a different species (see remarks in D. maderai sp. nov.). The most surprising discovery is that all specimens previously identified by Koch and Schawaller as D. denticollis differ from the lectotype (designated in this paper) and from new specimens of D. denticollis collected in the field. These specimens represent a new species, which has been named Deretus necopinatus sp. nov. Differential diagnosis. The type specimen was redescribed by Gahan (1903) and figured by Koch (1970). For this reason, and as the male and female do not differ significantly from each other in the shape of habitus (in males the apical antennomere is somewhat longer than in females), the description is not repeated here. Size of lectotype 11.0 x 4.0 mm. Variability of size in females — 5.7 – 11.0 x 2.1 – 4.1 mm; males — 4.7 – 9.5 x 1.8 – 3.8 mm. This species, in its general appearance, most resembles D. necopinatus sp. nov. from which it is differentiated by the shiny integument, the broadly rounded pronotal anterior corners (Fig. 14) and by the absence of patchs of sensillae on the abdominal ventrites together with the unmodified metatibia in males. From D. spinicollis and D. bezdeki sp. nov. it can be distinguished mainly by the shape of the pronotum, which in the latter two species is characterized by extremely elongated (extended) anterior corners (Fig. 13, 18). It differs from D. hulai sp. nov. by its smooth, almost glabrous (only ultra-microscopically and very sparsely pubescent), elytra (conspicuously pubescent and sculptured with small tubercles in the latter species). From D. maderai sp. nov. it can be separated by its almost flat pronotum (lateral view) and by its elytra length / pronotum width ratio, that is larger than 2.0. In D. maderai sp. nov. the pronotum is distinctly convex in lateral view and elytra length / pronotum width ratio is less than 1.8.	en	Purchart, Luboš (2012): Biodiversity research of darkling beetles on Socotra Island. Part I. The genus Deretus Gahan, 1900 (Coleoptera: Tenebrionidae). Zootaxa 3153: 57-68, DOI: 10.5281/zenodo.211332
732487F5BD496E7FEBEFA347FAB7FC15.taxon	materials_examined	Type locality. Yemen, Socotra Island, Firmihin. Type material. Holotype — (1 Ƥ NMPC): Republic of Yemen, Socotra Isl., Firmihin plato, Dracena [Dracaena] tree forest, N 12 ° 28 ’ 465 ’’, E 54 ° 00 ’ 89830 ’’, V. Hula lgt., 22. – 25.6.2009. The paratypes — (1 Ƥ LPCB): same as holotype; (1 Ƥ LPCB): YEMEN, SOCOTRA Island, Firmihin, 400 – 500 m, N 12 ° 28´27 ´´, E 54 ° 0´54 ´´, 22 – 25. vi. 2009, L. Purchart lgt. Differential diagnosis. Deretus hulai sp. nov. is the only species in the genus to possess no emargination on the anterior margin of pronotum. The margin is concave in dorsal view. Moreover, the pronotum is subquadrate and not distinctly transverse, while in the rest of the species it is transverse. With D. bezdeki sp. nov. it shares dentate sides of the pronotum and sculptured elytra. However, D. hulai sp. nov. possesses small spine-like teeth on lateral margin of pronotum (Fig. 15) and very small tubercles on the elytra. D. bezdeki sp. nov. has large tubercles in the elytral intervals, large acute teeth on the pronotal sides and strongly elongated anterior corners of the pronotum (Fig. 13). The remaining four species have no spine-like or acute teeth, and lack tubercles on the elytra.	en	Purchart, Luboš (2012): Biodiversity research of darkling beetles on Socotra Island. Part I. The genus Deretus Gahan, 1900 (Coleoptera: Tenebrionidae). Zootaxa 3153: 57-68, DOI: 10.5281/zenodo.211332
732487F5BD496E7FEBEFA347FAB7FC15.taxon	description	Description. Holotype, body length 5.3 mm, width 2.1 mm. Variability of size in paratypes — females: 5.7 – 5.8 x 2.15 – 2.3 mm; male: unknown. Body subcylindric, dark brown, shiny (Fig. 5). Head densely punctate, punctures as large as eye facets. Space between punctures narrower or equal to their diameter. Eyes reniform, not divided by genae. Clypeus with almost straight anterior margin. The latter with several long yellowish setae. Labrum transverse, punctate, covered with long yellowish setae. Ventral side of head shiny and punctate. Mentum transverse with midlongitudinal and smooth gibbosity. Apical labial palpomere broadened, triangular. Antennae filiform, relatively short, covered by yellowish setae. Third antennomere prolonged, longer than fourth antennomere but shorter than the two following joints combined. Pronotum (Fig. 15) subquadrate, punctate, punctures same as on head. Lateral margin of pronotum with several (4 – 6) small spine-like teeth. Base of pronotum and anterior corners bordered. Pronotal base rounded. Anterior margin not emarginated, convex (dorsal view). Elytra sub parallel, sparsely covered with short and erected yellowish setae, slightly flattened (lateral view) with steep apical declivity (lateral view). Base of elytra bordered only around humeri. Scutellum subtriangular. Elytral striae conspicuous, composed of relatively deep punctures which are little larger than eye facet. All nine elytral interstriae flat, shagreened, covered irregularly with small shiny tubercles. Elytral epipleuron narrow and very finely punctate. Prosternum, hypomeron and mesepisternum glabrous, smooth, densely and coarsely punctate. Mesoventrite, mesepimeron and metaventrite finely pubescent, smooth, densely and coarsely punctate. Abdominal ventrites shallowly punctate, pubescent. Legs punctate and covered with yellowish setae.	en	Purchart, Luboš (2012): Biodiversity research of darkling beetles on Socotra Island. Part I. The genus Deretus Gahan, 1900 (Coleoptera: Tenebrionidae). Zootaxa 3153: 57-68, DOI: 10.5281/zenodo.211332
732487F5BD496E7FEBEFA347FAB7FC15.taxon	etymology	Etymology. Dedicated to my dear colleague and friend Dr. Vladimír Hula (Czech Republic), zoologist and ecologist, specialist in Lepidoptera and Aranea, and collector of type specimens, for his long-term support.	en	Purchart, Luboš (2012): Biodiversity research of darkling beetles on Socotra Island. Part I. The genus Deretus Gahan, 1900 (Coleoptera: Tenebrionidae). Zootaxa 3153: 57-68, DOI: 10.5281/zenodo.211332
732487F5BD486E7CEBEFA113FEECFBA1.taxon	materials_examined	Type locality. Yemen, Socotra Island, Shibhon. Type material. Holotype — (13 NMPC): Yemen, Socotra Island, Shibhon, 680 m, N 12 ° 28´1.5 ´´, E 53 ° 58´31.4 ´´, 13. vi. 2009, L. Purchart lgt. The paratypes — (23 + 2 Ƥ LPCB): same data as holotype; (1 ƤLPCB, 1 Ƥ NMPC): Yemen, Socotra Island, Firmihin plato, 400 – 500 m, N 12 ° 28´45 ´´, E 54 ° 00´89 ´´, 18. – 19. vi. 2010, V. Hula & J. Niedobová leg.; (1 Ƥ SBCB): YEMEN E, Socotra Isl., 6. – 24. ix. 1999, V. Bejček & K. Šťastný lgt. // COLLECTION STANISLAV BEČVÁŘ {yellow label} // PARATYPUS, Deretus wraniki n. sp., SCHAWALLER 2000 {red label} // Collection Stanislav Bečvář {green label}. Differential diagnosis. This species can be separated from D. hulai sp. nov. by its transverse pronotum (Fig. 16) and smooth elytra (in D. hulai sp. nov. subquadrate — Fig. 15 and sculptured respectively). From D. bezdeki sp. nov. it differs by having smooth elytra, a non dentate pronotum which also lacks extended anterior corners (elytra sculptured, pronotal sides dentate and with extremely protruded anterior corners of pronotum in D. bezdeki sp. nov.). From D. denticollis and D. necopintaus sp. nov. it differs by having a distinctly convex pronotum when viewed laterally (more or less flat in D. denticollis and D. necopinatus sp. nov.), by elytra length / pronotum width ratio less than 1.8 and by the sixth elytral interval with a longitudinal impression at the elytral base (no impressions on elytra in the latter two species). It also differs from D. necopinatus sp. nov. by its shiny body (dull in the latter species). From D. spinicollis it can be distinguished by its much smaller size and unextended anterior pronotal corners.	en	Purchart, Luboš (2012): Biodiversity research of darkling beetles on Socotra Island. Part I. The genus Deretus Gahan, 1900 (Coleoptera: Tenebrionidae). Zootaxa 3153: 57-68, DOI: 10.5281/zenodo.211332
732487F5BD486E7CEBEFA113FEECFBA1.taxon	description	Description. Holotype, body length 7.2 mm, width 2.7 mm. Variability of size in paratypes — females: 5.8 – 8.3 x 2.5 – 3.6 mm; males: 5.6 – 7.8 x 2.3 – 3.0 mm. Body dark brown, shiny (Fig. 3). Head densely punctate, punctures as large as eye facets. Space between punctures narrower or equal to their diameter. Eyes reniform, not divided by genae. Clypeus with almost straight anterior margin and with several long yellowish setae anteriorly. Labrum transverse, punctate and covered with long yellowish setae. Ventral side of head shiny and punctate. Mentum transverse with midlongitudinal and smooth gibbosity. Apical labial palpomere broadened, triangular in both sexes. Antennae filiform, relatively long, covered by whitish setae. Third antennomere prolonged, approximately as long as the two following joints combined or slightly shorter. Apical antennomere pale brown, longer in males. Pronotum (Fig. 16) transverse, distinctly convex (lateral view), punctate, punctures same as on head. Anterior margin broadly and shallowly emarginated. Both lateral margins irregularly serrate. Basal, lateral, and anterior margins of pronotum distinctly bordered. Anterior margin medially obliterated. Elytra subparallel in males and subovoid in females. Slightly convex (lateral view) with moderate apical declivity (lateral view). Base of elytra bordered only around humeri. Scutellum triangular, finely punctate. Elytral striae conspicuous, composed of punctures, puncture diameter subequal to eye facet. Elytral interstriae glabrous, convex and punctate with punctures smaller than eye facet. Sixth elytral interval with longitudinal impression at elytral base. Elytral epipleuron broad and very finely punctate. Prosternum, hypomeron, mesoventrite, mesepisternum, mesepimeron and metaventrite smooth, glabrous, densely and coarsely punctate. Prosternal process widened behind coxae (ventral view). Abdominal ventrites shallowly punctate, moderately pubescent. Males without patches of sensillae. Legs punctate, covered with yellowish setae. Males with first three protarsomeres distinctly wider than penultimate and apical protarsomeres. Further modifications not present in either sex. Aedeagus see Fig. 21, 26.	en	Purchart, Luboš (2012): Biodiversity research of darkling beetles on Socotra Island. Part I. The genus Deretus Gahan, 1900 (Coleoptera: Tenebrionidae). Zootaxa 3153: 57-68, DOI: 10.5281/zenodo.211332
732487F5BD486E7CEBEFA113FEECFBA1.taxon	etymology	Etymology. Dedicated to Dr. Petr Madĕra (Czech Republic), botanist and dendrologist. Without his generous offer to join his educational project I would never have had the possibility to visit Socotra Island.	en	Purchart, Luboš (2012): Biodiversity research of darkling beetles on Socotra Island. Part I. The genus Deretus Gahan, 1900 (Coleoptera: Tenebrionidae). Zootaxa 3153: 57-68, DOI: 10.5281/zenodo.211332
732487F5BD486E7CEBEFA113FEECFBA1.taxon	discussion	Remarks. Schawaller (2004) described D. wraniki Schawaller, 2004 based on two specimens (male holotype and female paratype), which were very similar to each other, and which he believed belonged together. During this work it was discovered that the holotype is identical to the lectotype of D. denticollis and therefore it is synonymised (see also remarks in D. denticollis). Further examination revealed that the female paratype of D. wraniki is a different species, identical to specimens collected during this study, which is described in this paper as D. maderai sp. nov.	en	Purchart, Luboš (2012): Biodiversity research of darkling beetles on Socotra Island. Part I. The genus Deretus Gahan, 1900 (Coleoptera: Tenebrionidae). Zootaxa 3153: 57-68, DOI: 10.5281/zenodo.211332
732487F5BD4B6E7DEBEFA1FFFB20FB11.taxon	materials_examined	Type locality. Yemen, Socotra Island, Homhil area. Type material. Holotype — (13 NMPC): YEMEN, SOCOTRA Island E, Homhil area, 400 – 510 m, N 12 ° 34´25 ´´, E 54 ° 18´53 ´´, 9. – 10. ii. 2010, L. Purchart & J. Vybíral lgt. The paratypes — (1 Ƥ NMPC; 13 + 1 Ƥ SMNS; 53 + 13 Ƥ LPCB): same data as holotype; (33 + 8 Ƥ LPCB): YEMEN, SOCOTRA Island, Wadi Zirik, 650 – 670 m, N 12 ° 29 ’ 35 ’’, E 53 ° 59 ’ 28 ’’, 30. ii. 2010, L. Purchart lgt.; (23 + 2 Ƥ LPCB): YEMEN, SOCOTRA Island, Firmihin, 400 – 500 m, N 12 ° 28´27 ´´, E 54 ° 0´54 ´´, 6. – 7. ii. 2010, L. Purchart & J. Vybíral lgt.; (1 Ƥ LPCB): Yemen, Socotra Island, North-East “ arterial road ”, 970 m, N 12 ° 28´1.5 ´´, E 53 ° 58´31.4 ´´, 13. vi. 2009, L. Purchart lgt.; (13 + 1 Ƥ LPCB): Yemen, Socotra Island, Dixam plateau, Firmihin (Dracaena forest), 12 ° 28.6 ’ N, 54 ° 01.1 ’ E, 490 m, 15. – 16. xi. 2010, L. Purchart lgt.; (43 + 3 Ƥ NMPC): same data as previous, but Jiří Hájek leg.; (83 + 8 Ƥ JBCP): same data as previous, but Jan Batelka leg.; (1 Ƥ LPCB): Yemen, Socotra Island E, Kesa env., 220 – 300 m, N 12 ° 39´37 ´´, E 53 ° 26´42 ´´, 28. – 29. i. 2010, L. Purchart lgt.; (13 LPCB): Yemen, Socotra Island, Aloove area, 270 – 350 m, N 12 ° 30´58 ´´, E 54 ° 06´39 ´´, 3. – 4. ii. 2010, L. Purchart & J. Vybíral lgt.; (13 + 1 Ƥ NMPC): Yemen, Soqotra Is., 2003, 3. xii., Dixam plateau, WADI ZEERIQ, 750 m, N 12 ° 31 ’ 08 ’’ E 53 ° 59 ’ 09 ’’ [GPS], David Král lgt. / / YEMEN – SOQOTRA 2003 Expedition; Jan Farkač, Petr Kabátek & David Král; (1 SMNS): ZOOLOGISCHE EXKURSION, Diksam. März [March] 1999, leg. Wranik {handwritten}; (1 MNHN): Ille de SOCOTRA, 1993 – 1994, Canu Jean-Guy rec. {blue, printed} // Muséum Paris, 2001. Coll. générale {blue, printed}. Differential diagnosis. Deretus necopinatus sp. nov. can be distinguished from the remaining species, with the exception of D. spinicollis, by its conspicuously matt body and from all species by the presence of a patch of sensillae on males’ abdominal ventrites (Fig. 9). From D. spinicollis it differs by having an almost glabrous (only ultra-microscopically and very sparsely pubescent) elytra without impressions and only moderately extended anterior pronotal angles (Fig. 17). In the latter species the elytra are pubescent with yellowish hairs and possess shallow impressions near the humera. The anterior pronotal angles in D. spinicollis are strongly spine-like and elongate (Fig. 18).	en	Purchart, Luboš (2012): Biodiversity research of darkling beetles on Socotra Island. Part I. The genus Deretus Gahan, 1900 (Coleoptera: Tenebrionidae). Zootaxa 3153: 57-68, DOI: 10.5281/zenodo.211332
732487F5BD4B6E7DEBEFA1FFFB20FB11.taxon	description	Description. Holotype, body length 11.0 mm, width 4.0 mm. Variability of size in paratypes: females — 8.0 – 14.1 x 3.0 – 5.6 mm; males — 7.8 – 11.9 x 2.7 – 4.3 mm. Body subparallel, dark brown, dull (Fig. 8). Head coarsely and densely punctate, punctures subequal to eye facets or slightly larger. Space between punctures narrower or equal to their diameter. Eyes reniform, not divided by genae. Clypeus with straight anterior margin and with several long yellowish setae anteriorly. Labrum transverse, punctate, densely covered with long yellowish setae anteriorly. Ventral side of head shiny and coarsely punctate. Mentum transverse with midlongitudinal and smooth gibbosity. Apical labial palpomere broadened, triangular in both sexes. Antennae filiform, long, covered by yellowish setae. Third antennomere prolonged, approximately as long as the two following joints combined. Apical antennomere pale brown, longer in males. Pronotum (Fig. 17) transverse, punctate, punctures same as on head, except space between punctures slightly larger. Anterior margin broadly and deeply emarginate, anterior corners extended. Lateral margin of pronotum slightly and irregularly waved, arcuate before posterior angles. Basal, lateral, and anterior pronotal margins distinctly bordered. Anterior margin medially obliterated. Elytra subparallel, shagreened. Flattened (lateral view), with gentle apical declivity (lateral view). Base of elytra bordered only around humeri. Scutellum subtriangular, sparsely punctate. Elytral striae conspicuous, composed of slightly longitudinal punctures, puncture diameter approximately twice that of one eye facet. Elytral interstriae almost glabrous (only ultra-microscopically and very sparsely pubescent), flat, finely and irregularly punctate, puncture diameter less than eye facet diameter. Elytral epipleuron broad, very finely and sparsely punctate, with weak transversal wrinkles. Prosternum, hypomeron, mesoventrite, mesepisternum, mesepimeron and metaventrite glabrous and densely punctate. Prosternal process widened behind coxae (ventral view). Abdominal ventrites shallowly punctate and finely pubescent. Males with patches of sensillae (Fig. 9). Legs punctate, covered with yellowish setae. Metatibia medially dentated in males, medially widened in females (Fig. 8, 10). Aedeagus see Fig. 22, 27.	en	Purchart, Luboš (2012): Biodiversity research of darkling beetles on Socotra Island. Part I. The genus Deretus Gahan, 1900 (Coleoptera: Tenebrionidae). Zootaxa 3153: 57-68, DOI: 10.5281/zenodo.211332
732487F5BD4B6E7DEBEFA1FFFB20FB11.taxon	etymology	Etymology. Latin expression for surprising or unexpected. Adjective — necopinatus, - a, - um.	en	Purchart, Luboš (2012): Biodiversity research of darkling beetles on Socotra Island. Part I. The genus Deretus Gahan, 1900 (Coleoptera: Tenebrionidae). Zootaxa 3153: 57-68, DOI: 10.5281/zenodo.211332
732487F5BD4A6E7DEBEFA06FFE62F84B.taxon	materials_examined	Type locality. Yemen, Socotra Island, Haghier mountains. Type material examined. (3 SMNS): Yemen, Socotra Island, Haghier Mountains, 23. II. 1999, K. Van Damme, HLMD-Col- 920 - PT {white, printed} // PARATYPUS, Deretus spinicollis n. sp., Schawaller 2000 {red label, printed}. Additional material. (43 + 2 Ƥ LPCB): Yemen, Socotra Island, Skant area, 1300 – 1500 m, N 12 ° 34 ’ 33 ’’, E 54 ° 01 ’ 31 ’’, 31. i. – 1. ii. 2010, L. Purchart lgt.; (13 LPCB): Yemen, Socotra Island, Al Haghier Mts., Scant Mt. env., 12 ° 34.6 ’ N, 54 ° 01.5 ’ E, 1450 m, 12 – 13. xi. 2010, J. Bezdĕk leg.; (5 Ƥ + 4 3 NMPC): same data as previous, Jiří Hájek leg.; (13 + 1 Ƥ JBCP): same data as previous, but Jan Batelka leg., ex larvae. Differential diagnosis. Variability of size in females — 13.3 – 16.1 x 5.1 – 6.5 mm; males — 11.9 – 13.4 x 4.6 – 5.4 mm. This species can easily be separated from the other Deretus species by its extremely elongate spine-like anterior pronotal corners (Fig. 18) and, usually, by its larger size. D. bezdeki sp. nov., which also possesses spine-like anterior pronotal corners, is much smaller and has large acute teeth on the sides of the pronotum (Fig. 13) and large tubercles on the elytra.	en	Purchart, Luboš (2012): Biodiversity research of darkling beetles on Socotra Island. Part I. The genus Deretus Gahan, 1900 (Coleoptera: Tenebrionidae). Zootaxa 3153: 57-68, DOI: 10.5281/zenodo.211332
