identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
B55459FED26655CB892501A02113017D.text	B55459FED26655CB892501A02113017D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Megazarinae Brazidec & Vilhelmsen & Boudinot & Richter & Hammel & Perkovsky & Fan & Wang & Wu & Wang & Perrichot 2024	<div><p>3.1. Subfamily † Megazarinae Perrichot, 2009 stat. nov.</p><p>Type genus.</p><p>† Megazar Perrichot, 2009.</p><p>Diagnosis.</p><p>Head hypognathous, globular; compound eye oval, not covering head length, without postocular carina; flagellomeres cylindrical, longer than wide; subantennal groove without dorsal carina; mandibles symmetrical, with three or four teeth. Posterolateral margin of pronotum with notch accommodating anterior thoracic spiracle, not surrounding spiracle posteriorly; mesoscutum large, around half length of mesosoma, parapsidal lines present; median mesoscutal sulcus and axillar grooves present, deeply crenulate or smooth. Metasoma elongate. Fore wing hyaline; C, Sc+R, M+Cu, A, Rs+M, cu-a, Cu, Rs and R1 pigmented; costal, radial, first cubital, submarginal, and marginal cells closed by tubular veins. Hind leg stout, metafemur and metatibia swollen; at least either inner margin of metatrochanter, metacoxa, metatibia, or metabasitarsus bearing row of comb-like, thick setae along ventral surface.</p><p>Genera included.</p><p>† Cretolyra gen. nov., † Genkyhag gen. nov., † Megacoxa gen. nov., † Megalava Perrichot, 2009 [† Megalavini trib. nov.]; † Megazar Perrichot, 2009 [† Megazarini Perrichot, 2009].</p><p>Stratigraphic extension.</p><p>Albian to Cenomanian.</p><p>Comments.</p><p>The tribe Megazarini Perrichot, 2009 is here elevated to subfamily level, as † Megazarinae stat. nov. based on the results of our phylogenetic analyses. The monophyly of the subfamily is supported by three synapomorphies under parsimony: (1) metatibia and/or metatarsus with comb-like setae along ventral surface (31:1; Fig. 3A); (2) fore wing Rs+M located posteriorly, with the submarginal cell being at least twice the size of the medial cell (42:1; Fig. 3C); and (3) fore wing pterostigma absent or reduced (46:0; Fig. 3C). Within the subfamily, a new tribe is created to accommodate three of the newly described genera and † Megalava .</p></div>	https://treatment.plazi.org/id/B55459FED26655CB892501A02113017D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Brazidec, Manuel;Vilhelmsen, Lars;Boudinot, Brendon E.;Richter, Adrian;Hammel, Joerg U.;Perkovsky, Evgeny E.;Fan, Yong;Wang, Zhen;Wu, Qiong;Wang, Bo;Perrichot, Vincent	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
3F5C11CCF27B5F0391E4012059EA0EB3.text	3F5C11CCF27B5F0391E4012059EA0EB3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cretolyra Brazidec & Vilhelmsen & Boudinot & Richter & Hammel & Perkovsky & Fan & Wang & Wu & Wang & Perrichot 2024	<div><p>3.1.1.1. Genus † Cretolyra gen. nov.</p><p>Type species.</p><p>† Cretolyra noijebumensis sp. nov.</p><p>Etymology.</p><p>Combination of Creto - referring to the Cretaceous age of the Burmese amber deposit and the suffix - lyra often used in megalyrid genus names. Gender feminine.</p><p>Diagnosis.</p><p>Compound eye oval, higher than long; frons longitudinally divided by sulcus anterior to median ocellus; flagellomere 1 shorter than individual length of flagellomeres 2-5; occipital carina crenulate. Fore wing with M+Cu, Sc+R, A, M and Cu fully pigmented; cu-a almost branching with A, absent for short distance; medial cell rectangular; marginal cell narrow, closed in straight line (Figs 2A, B, 3C and 4A). Median mesoscutal sulcus smooth; parapsidal lines present, diverging anteriorly (Figs 3B and 4A). Metacoxa long; metafemur and metatibia swollen; two mesotibial and one metatibial spurs; row of comb-like setae along ventral surface of metatibia and metabasitarsus. Metasoma elongate and narrowed at apex, longer than mesosoma.</p></div>	https://treatment.plazi.org/id/3F5C11CCF27B5F0391E4012059EA0EB3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Brazidec, Manuel;Vilhelmsen, Lars;Boudinot, Brendon E.;Richter, Adrian;Hammel, Joerg U.;Perkovsky, Evgeny E.;Fan, Yong;Wang, Zhen;Wu, Qiong;Wang, Bo;Perrichot, Vincent	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
982B71A3D015503EACAFCDC54F6F2963.text	982B71A3D015503EACAFCDC54F6F2963.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cretolyra noijebumensis Brazidec & Vilhelmsen & Boudinot & Richter & Hammel & Perkovsky & Fan & Wang & Wu & Wang & Perrichot 2024	<div><p>3.1.1.1.1. † Cretolyra noijebumensis sp. nov.</p><p>Figures 2A, 3</p><p>Etymology.</p><p>The specific epithet is an adjective referring to the type locality where the amber pieces were collected.</p><p>Material studied.</p><p>Holotype male IGR.BU-067; housed in the amber collection of the Geology Department and Museum of the University of Rennes, France (IGR).</p><p>Type locality.</p><p>Noije Bum Hill, Hukawng Valley, Kachin State, Myanmar.</p><p>Age.</p><p>Upper Albian to lower Cenomanian, mid-Cretaceous.</p><p>Diagnosis.</p><p>Antennae inserted on edge of clypeus (Fig. 3A; vs. inserted above clypeal margin in Cretolyra shawi gen. et sp. nov.); axillae contiguous medially and axillar grooves crenulated (Fig. 3B; vs axillae separated medially and axillar grooves smooth in Cretolyra shawi gen. et sp. nov.).</p><p>Description.</p><p>Body length 3.33 mm; body very pubescent, especially on metasoma where setae conceal margins of tergites; the setae have agglomerated small bubbles that make observation difficult. - Head globular, higher than long (length 0.55 mm; height 0.75 mm); frons convex, smooth, divided by thin median sulcus; compound eye oval, higher than long; vertex convex, with smooth ocellar triangle; clypeus well-developed; torulus inserted closer to clypeus than to eye; shallow but distinct subantennal groove adjoining ventral margin of eye; antenna half as long as body, almost reaching metasoma; scape twice as long as wide (length 0.16 mm); pedicel shorter than scape (length 0.11 mm); flagellomeres cylindrical, elongate, longer than wide; flagellomere 1 shortest (length 0.15 mm); following flagellomeres longer (length ca. 0.21 mm); occipital carina crenulate. - Mesosoma almost half body length (length 1.29 mm; height 0.65 mm); mesoscutum convex, shagreened, divided by smooth median mesoscutal sulcus (mesoscutum length 0.58 mm; width ~0.60 mm); parapsidal lines present, diverging anteriorly; axillae contiguous medially, axillar groove crenulate; mesoscutellum diamond-shaped; pronotum smooth, not visible dorsally, with posteromedial part moderately high as viewed laterally; anterior thoracic spiracle not fully surrounded by pronotal cuticle; mesometapectal sulcus crenulate; propodeum shorter than mesoscutellum, with incomplete pairs of median and submedian carinae, two complete lateral longitudinal carinae, anterior region of propodeum with transversal row of foveae adjacent to concealed metanotum, posterior region of propodeum with three large foveae. - Fore wing hyaline and covered with microtrichiae, about two thirds of body length (length 2.50 mm); C, Sc+R, M+Cu, A, Rs fully pigmented; R1 extending beyond marginal cell; Rs closing marginal cell in straight line; M pigmented to apex; medial cell rectangular, narrow; Cu almost reaching posterior wing margin. - Legs covered with setae; two mesotibial and one metatibial spurs; metacoxa elongate; metafemur and metatibia swollen; metabasitarsus three times as long as following metatarsomeres and with row of short comb-like setae along ventral surface. - Metasoma half body length (length 1.48 mm), elongate and narrowed at apex; seven smooth tergites with pubescent posterior margin; first tergite largest; second tergite one quarter of metasoma length; remaining tergites shorter.</p></div>	https://treatment.plazi.org/id/982B71A3D015503EACAFCDC54F6F2963	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Brazidec, Manuel;Vilhelmsen, Lars;Boudinot, Brendon E.;Richter, Adrian;Hammel, Joerg U.;Perkovsky, Evgeny E.;Fan, Yong;Wang, Zhen;Wu, Qiong;Wang, Bo;Perrichot, Vincent	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
3743BE522B3758F59647816F3AE30EA8.text	3743BE522B3758F59647816F3AE30EA8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cretolyra shawi Brazidec & Vilhelmsen & Boudinot & Richter & Hammel & Perkovsky & Fan & Wang & Wu & Wang & Perrichot 2024	<div><p>3.1.1.1.2. † Cretolyra shawi sp. nov.</p><p>Figures 2B, 4</p><p>Etymology.</p><p>The specific epithet is a patronym honoring Scott R. Shaw, for his contributions to the knowledge of Megalyridae .</p><p>Material studied.</p><p>Holotype female NIGP203545; housed in the Nanjing Institute of Geology and Paleontology (NIGP), Chinese Academy of Sciences, Nanjing, China.</p><p>Type locality.</p><p>Noije Bum Hill, Hukawng Valley, Kachin State, Myanmar.</p><p>Age.</p><p>Upper Albian to lower Cenomanian, mid-Cretaceous.</p><p>Diagnosis.</p><p>Pedicel bilobed, long and thick (Figs 4D-E; vs. calciform and elongate in Cretolyra noijebumensis gen. et sp. nov.); axillae almost rounded and reduced on edges of mesoscutellum, not contiguous medially (vs. contiguous medially in Cretolyra noijebumensis gen. et sp. nov.); axillar grooves smooth (Fig. 4A; vs. axillar grooves crenulated in Cretolyra noijebumensis gen. et sp. nov.); medial cell nearly triangular, with 1m-cu very short (Fig. 2B; vs. medial cell rectangular, with 1m-cu distinct in Cretolyra noijebumensis gen. et sp. nov.).</p><p>Description.</p><p>Body length 2.82 mm. - Head globular, higher than long (length ca. 0.40 mm), covered with short setae; frons convex, smooth, divided by sulcus; compound eye oval, higher than long; vertex convex; clypeus smooth, apically rounded; mandibles symmetrical, with three teeth; toruli separated from each other by less than their own diameter; subantennal groove present; antenna about half body length; scape shorter than pedicel; pedicel thick, bilobed; flagellomeres cylindrical, longer than wide; flagellomere 1 shorter than flagellomeres 2-5 (length 0.15 mm vs. ca. 0.21 mm); flagellomeres 6-11 shorter (length ca. 0.10 mm); flagellomere 12 as long as flagellomere 1 (length 0.15 mm); occipital carina minutely crenulate. - Mesosoma one third of body length (length 0.98 mm); mesoscutum convex, divided by smooth median mesoscutal sulcus; parapsidal lines present; axillae small, almost rounded and not contiguous medially; axillar groove smooth; mesoscutellum smooth, convex, diamond-shaped; pronotum smooth, not visible dorsally, with posteromedial part moderately high as viewed laterally; anterior thoracic spiracle not surrounded by pronotal cuticle posteriorly; mesometapectal sulcus crenulate; propodeum carinate. - Fore wing hyaline and covered with microtrichiae, two thirds of body length (length 2.57 mm); pterostigma reduced; C, Sc+R, M+Cu, A, Rs fully pigmented; R1 extending beyond marginal cell; Rs closing marginal cell in straight line; M pigmented to apex; medial cell small, elongate and rectangular, nearly triangular by side of 1m-cu. - Legs with two mesotibial and one metatibial spurs; metacoxa elongate; metafemur and metatibia swollen; metabasitarsus two to three times as long as remaining metatarsomeres; row of comb-like setae along ventral surface of metatibia and metabasitarsus. - Metasoma longer than mesosoma (length 1.44 mm); elongate and narrowed at apex; tergites smooth; hypopygium well-developed; ovipositor shorter than metasoma (length 1.33 mm; OL/BL ratio 0.43), sheaths fully preserved, transversely striated.</p></div>	https://treatment.plazi.org/id/3743BE522B3758F59647816F3AE30EA8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Brazidec, Manuel;Vilhelmsen, Lars;Boudinot, Brendon E.;Richter, Adrian;Hammel, Joerg U.;Perkovsky, Evgeny E.;Fan, Yong;Wang, Zhen;Wu, Qiong;Wang, Bo;Perrichot, Vincent	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
6E91A808941E580DB5C2EB6B37060D64.text	6E91A808941E580DB5C2EB6B37060D64.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Genkyhag Brazidec & Vilhelmsen & Boudinot & Richter & Hammel & Perkovsky & Fan & Wang & Wu & Wang & Perrichot 2024	<div><p>3.1.1.2. Genus † Genkyhag gen. nov.</p><p>Type species.</p><p>† Genkyhag innebula sp. nov.</p><p>Etymology.</p><p>Anagram of Ga Hkyeng, meaning red soil in Jinghpaw, from which the term Kachin is derived. Gender masculine.</p><p>Diagnosis.</p><p>Compound eye oval, higher than long; antenna inserted half way between eye and clypeus; flagellomeres cylindrical, subequal in length (Figs 5C, D). Mesosoma less than half body length, shorter than metasoma; mesoscutum hiding pronotum in dorsal view; median mesoscutal sulcus and axillar groove crenulate; parapsidal lines present, parallel to median mesoscutal sulcus and diverging posteriorly; axillae not contiguous medially; mesoscutellum short (Figs 5A, B); propodeum carinate. Fore wing hyaline, with microtrichiae; venation almost fully pigmented, with only Cu nebulous; medial cell trapezoidal and broad; marginal cell closed without bend; R1 extending beyond this cell (Figs 2C, 5A, B, and E). Legs with one mesotibial and two metatibial spurs. Metasoma elongate, fusiform.</p><p>Comments.</p><p>One of the main differences compared to † Cretolyra gen. nov. is the presence of one mesotibial and two metatibial spurs (instead of two mesotibial and one metatibial). The presence of two metatibial spurs seems to be an apomorphy among megalyrids with reversion in some fossil taxa (Vilhelmsen et al. 2010a).</p></div>	https://treatment.plazi.org/id/6E91A808941E580DB5C2EB6B37060D64	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Brazidec, Manuel;Vilhelmsen, Lars;Boudinot, Brendon E.;Richter, Adrian;Hammel, Joerg U.;Perkovsky, Evgeny E.;Fan, Yong;Wang, Zhen;Wu, Qiong;Wang, Bo;Perrichot, Vincent	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
FCC83238CF44597C9C8CB733D9D5D303.text	FCC83238CF44597C9C8CB733D9D5D303.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Genkyhag innebula Brazidec & Vilhelmsen & Boudinot & Richter & Hammel & Perkovsky & Fan & Wang & Wu & Wang & Perrichot 2024	<div><p>3.1.1.2.1. † Genkyhag innebula sp. nov.</p><p>Figures 2C, 5</p><p>Etymology.</p><p>From Latin 'nebula, In nebula’, meaning "in the fog", in reference to the amber pieces where the specimens are found, that contains numerous terrigenous inclusions, sometimes partially hiding the specimens. It is a noun in apposition.</p><p>Material studied.</p><p>Holotype female NIGP203546; housed in the Nanjing Institute of Geology and Paleontology (NIGP), Chinese Academy of Sciences, Nanjing, China. Three paratypes male FAI-BI 11324a, male FAI-BI 11324b, FAI-BI-11324c of unknown sex; housed in the Fushun Amber Institute.</p><p>Type locality.</p><p>Noije Bum Hill, Hukawng Valley, Kachin State, Myanmar.</p><p>Age.</p><p>Upper Albian to lower Cenomanian, mid-Cretaceous.</p><p>Diagnosis.</p><p>As for genus.</p><p>Description.</p><p>Holotype female incompletely preserved (body length 3.12 mm), in a piece with multiple terrigenous inclusions. Paratypes preserved in a single piece: FAI-BI 11324a visible in dorsal and frontal views, FAI-BI 11324b in lateral view (body length ca. 3.45 mm), FAI-BI 11324c being one complete fore and hind wings and a partial fore wing. - Head globular, apparently entirely glabrous (length for holotype ca. 0.47 mm); frons shagreened, convex, divided by a faintly impressed sulcus; compound eye oval, without postocular carina; vertex convex, shagreened; clypeus short; subantennal groove present; toruli inserted half way between clypeus and eyes; antenna short, barely reaching propodeum; scape globular; pedicel slightly longer; flagellomeres cylindrical, longer than wide; flagellomeres 7-11 shorter than flagellomeres 1-6; flagellomere 12 longest; occipital carina crenulate. - Mesosoma slightly shorter than metasoma (length for holotype 1.12 mm; for paratype 11324a 1. 29 mm; for paratype 11324b 1.57 mm); pronotum not visible dorsally; mesoscutum shagreened, half as long as mesosoma, median mesoscutal sulcus crenulate (mesoscutum length for holotype 0.51 mm; for paratype 11324a 0.61 mm; width for holotype 0.65 mm; for paratype 11324a 0.71 mm); parapsidal lines present; axillae barely contiguous medially and inner grooves crenulate; mesoscutellum wider than long, shorter than mesoscutum; pronotum smooth, not visible dorsally, with posteromedial part moderately high as viewed laterally; anterior thoracic spiracle not fully surrounded by pronotal cuticle; propodeum carinate, carinae delimitating row of small foveae in anterior region and larger foveae in posterior region. - Fore wing hyaline, covered with microtrichiae (length for holotype 1.84 mm; for paratype 11324a 2.10 mm; for paratype 11324b 2.11 mm; for paratype 11324c 2.86 mm); venation almost complete, Sc+R, M+Cu, A, Rs+M and M fully pigmented; Cu spectral; marginal cell very narrow, closed in straight line; R1 barely extending beyond marginal cell; medial cell rectangular and relatively broad. - Hind wing hyaline; venation reduced to Sc+R and R1, Sc+R almost not diverging from margin, R1 pigmented almost to tip; at least seven hamuli, located distal to middle of hind wing. - Legs with one mesotibial and two metatibial spurs; metafemur swollen; ventral surface of metatibia and metabasitarsus with row of comb-like setae, metabasitarsus at least three times as long as following metatarsomeres. - Metasoma elongate, oval, with sparse short setae (length for holotype 1.43 mm); tergites smooth; ovipositor shorter than metasoma (length for holotype 1.34 mm; OL/BL ratio 0.43).</p></div>	https://treatment.plazi.org/id/FCC83238CF44597C9C8CB733D9D5D303	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Brazidec, Manuel;Vilhelmsen, Lars;Boudinot, Brendon E.;Richter, Adrian;Hammel, Joerg U.;Perkovsky, Evgeny E.;Fan, Yong;Wang, Zhen;Wu, Qiong;Wang, Bo;Perrichot, Vincent	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
7DDF1391E8C854A8A428E13D42F4B321.text	7DDF1391E8C854A8A428E13D42F4B321.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kamyristi Brazidec & Vilhelmsen & Boudinot & Richter & Hammel & Perkovsky & Fan & Wang & Wu & Wang & Perrichot 2024	<div><p>3.2.1.1. Genus † Kamyristi gen. nov.</p><p>Type species.</p><p>† Kamyristi exfrigore sp. nov.</p><p>Etymology.</p><p>Anagram of Taïmyrski, the Russian name of the Taimyr peninsula from where the amber pieces containing the specimens derive. Gender masculine.</p><p>Diagnosis.</p><p>As for tribe with the following additional characters: head shagreened; compound eye higher than long; occipital carina foveate (Figs 9C and F); flagellomeres cylindrical and elongate. Mesosoma less than half of metasoma length; mesoscutum reduced on anterior part of mesosoma, hiding pronotum dorsally (Figs 9B and F). Fore wing with medial cell pentagonal with 1m-cu, Cu1 and M+Cu and Cu spectral, Rs+M not aligned with M+Cu (Figs 2G, H, 9B and D). Hind leg with scattered erect setae; one mesotibial and one metatibial spur.</p><p>Comments.</p><p>Two species from this genus are known, both displaying numerous similarities with the extant genus Cryptalyra from South America, e.g., the elongate smooth metasoma (Fig. 9A), the erect setae on the hind leg (Fig. 9A) or M half pigmented (Figs 2G and H). But † Kamyristi exfrigore gen. et sp. nov. displays the most surprising affinities to Cryptalyra: the ovipositor sheaths seem very reduced, only covering the basal part of the ovipositor (Fig. 9A). However, the closed marginal cell (Fig. 2G and H), the elongate flagellomeres (Fig. 9E) and the mesosoma being shagreened instead of foveate (Fig. 9C) indicate that it is not closely related to Cryptalyra . Examination of more female specimens will be necessary to confirm if reduced ovipositor sheaths are a diagnostic character for this genus. † Kamyristi gen. nov. species are the smallest megalyrids in Asia.</p><p>Due to the fore wing venation and absence of a sulcus on the vertex, this new genus resembles Dinapsini with Rs reduced. Another difference is the posterior head sculpture, distinctly shagreened, whereas it is reticulate for the extant dinapsine species. Finally, the lack of grooves behind the eyes and of a dorsal carina on the subantennal groove makes the placement in Dinapsini doubtful; they are similarly absent in † Valaa delclosi . According to Shaw (1990a), these characters are relatively homoplasious. The placement of † Kamyristi gen. nov. in the Dinapsini is not corroborated by the phylogenetic analyses (see below).</p></div>	https://treatment.plazi.org/id/7DDF1391E8C854A8A428E13D42F4B321	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Brazidec, Manuel;Vilhelmsen, Lars;Boudinot, Brendon E.;Richter, Adrian;Hammel, Joerg U.;Perkovsky, Evgeny E.;Fan, Yong;Wang, Zhen;Wu, Qiong;Wang, Bo;Perrichot, Vincent	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
6586EC0085A85D39BE714BC02321F92A.text	6586EC0085A85D39BE714BC02321F92A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kamyristi exfrigore Brazidec & Vilhelmsen & Boudinot & Richter & Hammel & Perkovsky & Fan & Wang & Wu & Wang & Perrichot 2024	<div><p>3.2.1.1.1. † Kamyristi exfrigore sp. nov.</p><p>Figures 2G, 9A-C</p><p>Etymology.</p><p>" exfrigore " means "from the cold" in Latin as this species is the northernmost record of a megalyrid, extant and extinct, in a region where the temperature is currently below zero most of the year. The specific epithet is to be treated as an adjective.</p><p>Material studied.</p><p>Holotype female PIN 3730/411; housed in the collection of the Paleontological Institute of the Russian Academy of Sciences (PIN).</p><p>Type locality.</p><p>Baikura, Russia.</p><p>Age.</p><p>Upper Albian to lower Cenomanian, mid-Cretaceous.</p><p>Diagnosis.</p><p>Eye nearly round (vs. oval, higher than long in † Kamyristi yantardakhensis gen. et sp. nov.); axillae contiguous medially with axillar groove smooth (Fig. 9B; vs. separated medially with axillar groove crenulated in † Kamyristi yantardakhensis gen. et sp. nov.).</p><p>Description.</p><p>Body length 2.03 mm. Almost complete, except ovipositor. - Head globular, higher than long (length 0.38 mm; height 0.43); slightly pubescent on vertex and frons; frons convex, shagreened; compound eye rounded, slightly higher than long, not covering head length, postocular carina absent; vertex convex, shagreened; torulus inserted very close to ventral margin of eye; subantennal groove present; mandible with three teeth; scape short and thick (length 0.09 mm); pedicel same length; flagellomeres cylindrical, longer than wide, elongate (length ca. 0.08 mm); flagellomere 12 longest; occipital carina crenulate. - Mesosoma shorter than metasoma (length 0.69 mm; height 0.40 mm), almost fully shagreened; mesoscutum shagreened, short (length 0.20 mm; width 0.47 mm), one third of mesosoma length, divided by crenulate median mesoscutal sulcus, with sparse erect setae; axillae continuous medially, with inner axillar groove smooth; mesoscutellum as long as mesoscutum, posterior margin slightly outcurved; pronotum not visible dorsally; anterior thoracic spiracle fully surrounded by pronotal cuticle; propodeum areolate-rugose. - Fore wing hyaline (length ca. 1.50 mm) and covered with microtrichia; C, Sc+R, Rs +M, basal segments of Rs and M; M+Cu, Cu and 1m-cu spectral; Rs not closing first submarginal cell and pigmented for two thirds between r-rs and Rs+M; M pigmented to level of 2r-m and then spectral to wing tip; pterostigma broad; R1 extending beyond marginal cell; Rs pigmented, arched toward stigma, closing marginal cell. - Legs bearing many scattered erect setae; one mesotibial spur, one metatibial spur present; metafemur swollen, metatibia notched before spur; metabasitarsus three times longer than following, remainder same length. - Metasoma almost half body length (length 0.96 mm), elongate, smooth; segments subequal in length; hypopygium well-developed, concave; ovipositor incomplete, covered by sheaths only at base; cerci triangular, exserted.</p></div>	https://treatment.plazi.org/id/6586EC0085A85D39BE714BC02321F92A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Brazidec, Manuel;Vilhelmsen, Lars;Boudinot, Brendon E.;Richter, Adrian;Hammel, Joerg U.;Perkovsky, Evgeny E.;Fan, Yong;Wang, Zhen;Wu, Qiong;Wang, Bo;Perrichot, Vincent	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
9292FEA4705259A6A6FB6637A3FB90C9.text	9292FEA4705259A6A6FB6637A3FB90C9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kamyristi yantardakhensis Brazidec & Vilhelmsen & Boudinot & Richter & Hammel & Perkovsky & Fan & Wang & Wu & Wang & Perrichot 2024	<div><p>3.2.1.1.2. † Kamyristi yantardakhensis sp. nov.</p><p>Figures 2H, 9D-F</p><p>Etymology.</p><p>The specific epithet refers to the locality where the specimen was collected. The specific adjective is to be treated as a noun in a genitive case.</p><p>Material studied.</p><p>Holotype male PIN 3311/2718; housed in the collection of the Paleontological Institute of the Russian Academy of Sciences (PIN).</p><p>Type locality.</p><p>Yantardakh, Russia.</p><p>Age.</p><p>Santonian, Upper Cretaceous.</p><p>Diagnosis.</p><p>Axillae not contiguous medially, grooves crenulated (Fig. 9F; vs. contiguous medially, with axillar grooves smooth in † Kamyristi exfrigore gen. et sp. nov.).</p><p>Description.</p><p>Body length 1.47 mm. Body mostly bare. - Head slightly higher than long (length 0.26 mm; height 0.28 mm), glabrous; frons convex, shagreened; compound eye almond-shaped, higher than long, not covering head length, postocular carina absent; vertex convex, minutely shagreened but sculpture not fully visible; toruli inserted closer to each other than to eyes; subantennal groove present; mandible with three teeth, decreasing in size from apex to base; scape short and thick (length 0.07 mm), pedicel of similar length with scape but thinner; 10 flagellomeres preserved, cylindrical, elongate, longer than wide; flagellomere 1 shortest flagellomere (length 0.04 mm), flagellomeres 2-10 similar in length (length ca. 0.06 mm); flagellomere 11-12 missing; occipital carina crenulate. - Mesosoma more than a third of body length (length 0.43 mm; height 0.22 mm); mesoscutum less than one third of mesosoma length (length 0.15 mm; width 0.31 mm), shagreened and divided by crenulate median mesoscutal sulcus; axillae not continuous medially, with inner groove crenulate; mesoscutellum shagreened; pronotum not visible dorsally; anterior thoracic spiracle fully surrounded by pronotal cuticle; propodeum areolate-rugose. - Fore wing hyaline and covered with microtrichia (length ca. 0.75 mm); C, Sc+R, A, Rs and basal segment of M pigmented; M+Cu, Cu and 1m-cu spectral; Rs+M not aligned with M+Cu; medial cell pentagonal; Rs almost closing submarginal cell, not fully pigmented between r-rs and Rs+M; R1 pigmented beyond marginal cell; marginal cell closed by Rs with a bend; M pigmented halfway to wing tip and then spectral. - Legs bearing numerous scattered setae; one long mesotibial spur, one shorter metatibial spur; basimetatrochanter long, half metacoxa length; metafemur slightly swollen; first metatarsomere longest and bearing small erect setae. - Metasoma almost half body length, longer than mesosoma (length 0.75 mm), elongate; smooth and bare; tergites nearly equal in length; hypopygium small, parameres triangular, projecting downward.</p></div>	https://treatment.plazi.org/id/9292FEA4705259A6A6FB6637A3FB90C9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Brazidec, Manuel;Vilhelmsen, Lars;Boudinot, Brendon E.;Richter, Adrian;Hammel, Joerg U.;Perkovsky, Evgeny E.;Fan, Yong;Wang, Zhen;Wu, Qiong;Wang, Bo;Perrichot, Vincent	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
BC564AF72F8A5166903D7990FE9A477D.text	BC564AF72F8A5166903D7990FE9A477D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kamyristiini Brazidec & Vilhelmsen & Boudinot & Richter & Hammel & Perkovsky & Fan & Wang & Wu & Wang & Perrichot 2024	<div><p>3.2.1. Tribe † Kamyristiini trib. nov.</p><p>Type genus.</p><p>† Kamyristi gen. nov.</p><p>Diagnosis.</p><p>Head prognathous; compound eye without setation, without posterior carina; subantennal groove without dorsal carina; mandible with three teeth; occipital carina present, curving towards mandible. Median mesoscutal sulcus crenulate; parapsidal line absent; propodeum areolate-rugose. Fore wing hyaline with C, Sc+R, A, cu-a, Rs+M, M, Rs and R1 tubular; pterostigma large; M not reaching apical margin; Rs only present as a stub between Rs+M and r-rs; Rs apically strongly angled, arched towards stigma; Metasoma elongate, fusiform, longer than mesosoma; ovipositor sheaths only covering basal basal portion of ovipositor.</p><p>Genera and species included.</p><p>† Kamyristi exfrigore gen. et sp. nov., † Kamyristi yantardakhensis gen. et sp. nov.</p><p>Comments.</p><p>The tribe † Kamyristiini trib. nov. is erected based on the results of the phylogenetic analyses, to accommodate † Kamyristi gen. nov. and its two species. † Kamyristi gen. nov. is not retrieved within a larger clade, precluding its assignment to another monophyletic tribe.</p></div>	https://treatment.plazi.org/id/BC564AF72F8A5166903D7990FE9A477D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Brazidec, Manuel;Vilhelmsen, Lars;Boudinot, Brendon E.;Richter, Adrian;Hammel, Joerg U.;Perkovsky, Evgeny E.;Fan, Yong;Wang, Zhen;Wu, Qiong;Wang, Bo;Perrichot, Vincent	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
7D55FAD868145D9A93CDEF6A4E693D16.text	7D55FAD868145D9A93CDEF6A4E693D16.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Megalyrinae Brazidec & Vilhelmsen & Boudinot & Richter & Hammel & Perkovsky & Fan & Wang & Wu & Wang & Perrichot 2024	<div><p>3.2. Subfamily Megalyrinae Schletterer, 1889</p><p>Emended diagnosis.</p><p>Head globular; compound eye oval, with or without postocular carina; flagellomeres often elongate, sometimes compact; subantennal groove with or without dorsal carina, mandibles symmetrical with three teeth. Anterior thoracic spiracle fully surrounded by pronotal cuticle; median mesoscutal sulcus sometimes effaced or reduced; parapsidal lines sometimes absent. Fore wing with infumate banding patterns in Ettchellsia, Dinapsis, Megalyra and Megalyridia; at least Sc+R, Rs+M, basal segment of Rs and basal segment of R1 pigmented and no closed cell; at most with C, Sc+R, A, cu-a, Rs+M, Rs, R1 and basal segment of M pigmented, costal, submarginal, and marginal cells closed by tubular veins, subcostal and first cubital fused; M+Cu usually absent except in † Cretodinapsis; rarely brachypterous. Hind leg without row of erect setae along margin. Metasoma usually elongate but sometimes distinctly shorter than mesosoma.</p><p>Genera included.</p><p>Carminator Shaw, 1988, Cryptalyra Shaw, 1987 [ Cryptalyrini Shaw, 1988]; Dinapsis Waterston, 1922, Ettchellsia Cameron, 1909, Neodinapsis Shaw, 1987 [ Dinapsini Waterston, 1922]; † Cretodinapsis Rasnitsyn, 1977, Megalyra Westwood, 1832, † Prodinapsis Brues, 1923 [ Megalyrini Schletterer, 1889]; † Kamyristi gen. nov. [† Kamyristini trib. nov]; Megallica Perrichot, 2009 [† Megallicini trib. nov.]; Megalyridia Hedqvist, 1959 [ Megalyridiini Shaw, 1990]; Rigel Shaw, 1987 [ Rigelini Shaw, 1990]; † Valaa Perrichot, 2009 [† Valaaini trib. nov.].</p><p>Stratigraphic extension.</p><p>Albian to present.</p><p>Comments.</p><p>We redefine the subfamily Megalyrinae, according to the results of the phylogenetic analyses. The subfamily is composed of three non-monotypic tribes, namely Cryptalyrini, Dinapsini, and Megalyrini, as well as a number of monotypic tribes: Rigelini, † Kamyristini trib. nov., † Megallicini trib. nov., † Valaaini trib. nov., and Megalyridiini . The justification for redefining Megalyrinae is given in the Discussion.</p></div>	https://treatment.plazi.org/id/7D55FAD868145D9A93CDEF6A4E693D16	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Brazidec, Manuel;Vilhelmsen, Lars;Boudinot, Brendon E.;Richter, Adrian;Hammel, Joerg U.;Perkovsky, Evgeny E.;Fan, Yong;Wang, Zhen;Wu, Qiong;Wang, Bo;Perrichot, Vincent	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
A5AC297D1FAF55B89C817FFB71BFF7D8.text	A5AC297D1FAF55B89C817FFB71BFF7D8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Megacoxa Brazidec & Vilhelmsen & Boudinot & Richter & Hammel & Perkovsky & Fan & Wang & Wu & Wang & Perrichot 2024	<div><p>3.1.1.3. Genus † Megacoxa gen. nov.</p><p>Type species.</p><p>† Megacoxa janzeni sp. nov.</p><p>Etymology.</p><p>The genus name refers to the size of the metacoxa, which is the key diagnostic feature of the genus; besides, the prefix Mega - alludes to the family name Megalyridae . Gender feminine.</p><p>Diagnosis.</p><p>Compound eye oval; scape wider than long; thin occipital carina slightly crenulate. Parapsidal line present and located on posterolateral part of mesoscutum (Figs 6C, 7B and E); inner axillar groove crenulate (Fig. 8F). Fore wing venation with R1, M+Cu, Sc+R, A and Cu fully pigmented; A connected to medial cell with cu-a; Rs closing marginal cell without bend (Figs 2D-F). Metacoxa much enlarged, with dorsal surface rounded to angular, outer metacoxal surface posteriorly foveate, inner metacoxal surface almost entirely accommodated in anterolateral, shallow concavity of metasoma (Figs 6C, 7B, 8C and G). Metatibia and metabasitarsus with ventral row of comb-like setae; two mesotibial and two metatibial spurs (Figs 6A and 7A). Metasoma slightly shorter than mesosoma, tergites and sternites beveled (Figs 6B and 7A).</p><p>Comments.</p><p>The enlarged metacoxa accommodated in the anterior metasomal concavity is the key diagnostic feature for † Megacoxa gen. nov. This condition is present in all five specimens assigned to this genus and the metacoxae are clearly symmetrical in the synchrotron-scanned specimen, thus ruling out a preservational artefact.</p></div>	https://treatment.plazi.org/id/A5AC297D1FAF55B89C817FFB71BFF7D8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Brazidec, Manuel;Vilhelmsen, Lars;Boudinot, Brendon E.;Richter, Adrian;Hammel, Joerg U.;Perkovsky, Evgeny E.;Fan, Yong;Wang, Zhen;Wu, Qiong;Wang, Bo;Perrichot, Vincent	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
99C0269FADDF5E79BA4155230B90090D.text	99C0269FADDF5E79BA4155230B90090D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Megacoxa chandrahrasa Brazidec & Vilhelmsen & Boudinot & Richter & Hammel & Perkovsky & Fan & Wang & Wu & Wang & Perrichot 2024	<div><p>3.1.1.3.1. † Megacoxa chandrahrasa sp. nov.</p><p>Figures 2D, 6</p><p>Etymology.</p><p>The specific epithet is a noun in apposition. It is the name of the legendary sword of Mañjuśrī, a bodhisattva in Buddhism, which allowed him to open the Kathmandu valley, and refers to the long, sword-like metatibial spurs of this species.</p><p>Material studied.</p><p>Holotype female IGR.BU-068; housed in the amber collection of the Geology Department and Museum of the University of Rennes, France (IGR) . Paratype female CNU-HYM-MA2016207; housed in the Key Lab of Insect Evolution and Environmental Changes, College of Life Sciences, Capital Normal University, Beijing, China (CNU) .</p><p>Type locality.</p><p>Noije Bum Hill, Hukawng Valley, Kachin State, Myanmar.</p><p>Age.</p><p>Upper Albian to lower Cenomanian, mid-Cretaceous.</p><p>Diagnosis.</p><p>Median mesoscutal sulcus smooth (Fig. 6F; vs. crenulate in Megacoxa synchrotron gen. et sp. nov.); axillae contiguous medially (Fig. 6A; vs. separated medially in Megacoxa janzeni gen. et sp. nov.); medial cell elongate, almost triangular (Figs 2D and 6A; vs. broad and rectangular in Megacoxa synchrotron gen. et sp. nov. and Megacoxa janzeni gen. et sp. nov.); posterior margin of metacoxa outcurved and bordered with small foveae (Fig. 6C; vs. bordered with large foveae in Megacoxa janzeni gen. et sp. nov.. and straight posterior metacoxal margin with large foveae in Megacoxa synchrotron gen. et sp. nov.); metatibial spurs very long (Figs 6A, B and G); metasoma compressed, distinctly shorter than ovipositor; OL/BL ratio ~0.70.</p><p>Description.</p><p>Body length for holotype 2.64 mm; for paratype 4.80 mm. - Head globular, higher than long (length for holotype 0.52 mm; for paratype 0.93 mm), glabrous; frons convex, shagreened; compound eye oval, higher than long; vertex convex; clypeus short, shagreened; toruli inserted closer to each other than to eyes; subantennal groove present; antennae distorted, flagellomeres apparently cylindrical, longer than wide; occipital carina crenulate. - Mesosoma more than half body length (length for holotype 1.43 mm; for paratype 1.92 mm); covered with short setae; mesoscutum convex, wide (length for holotype 0.62 mm, width for holotype 0.74 mm; length for holotype 0.93 mm, width for holotype 0.90 mm), divided by smooth median mesoscutal sulcus; parapsidal lines present; axillae contiguous medially, inner axillar groove crenulate; pronotum not visible dorsally; anterior thoracic spiracle not fully surrounded by pronotal cuticle; mesometapectal sulcus crenulate; propodeum shorter than mesoscutellum, carinate. - Fore wing hyaline and covered with microtrichiae, slightly wrinkled at apex, as long as body (length for holotype 2.08 mm; for paratype 3.52 mm); venation complete with R1, M+Cu, Sc+R, Rs, A and Cu fully pigmented; Rs closing marginal cell; medial cell elongate; A connected to medial cell by 1cu-a. - Legs distorted; two mesotibial spurs; metacoxa huge, almost as high anteriorly as maximal propodeal height, with dorsal surface semicircular, posteroventral corner acute, ventral surface notched just anteriad trochanter insertion; outer metacoxal surface posteriorly with row of small foveae; dense row of short, comb-like setae along ventral surface of metatrochanter (full length), metafemur (basal half length), metatibia (distal half length), and metabasitarsus (full length); metafemur swollen; two long metatibial spurs, about twice as long as maximal tibial width; metabasitarsus three to four times longer than following metatarsomeres. - Metasoma relatively short (length for holotype 1.21 mm; for paratype 1.95 mm), distorted, partly intruding into propodeum during fossilization; tergites and sternites inserted in bevel, smooth; ovipositor longer than metasoma (length for holotype 1.87 mm, OL/BL ratio 0.70; length for paratype 3.24 mm, OL/BL ratio 0.68), sheaths transversely striated.</p><p>Comments.</p><p>We first interpreted the very high OL/BL ratio of the holotype of † Megacoxa chandrahrasa gen. et sp. nov. compared to other † Megalavini (0.70 vs. ~0.40) as a distortion of the specimen and the intruding of the metasoma into the mesosoma. However, the discovery of an additional specimen confirms that the metasoma is more compact in this species than in other species of the genus or the subfamily.</p></div>	https://treatment.plazi.org/id/99C0269FADDF5E79BA4155230B90090D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Brazidec, Manuel;Vilhelmsen, Lars;Boudinot, Brendon E.;Richter, Adrian;Hammel, Joerg U.;Perkovsky, Evgeny E.;Fan, Yong;Wang, Zhen;Wu, Qiong;Wang, Bo;Perrichot, Vincent	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
C65F8C4C37DD5B2B880B9F1B8220A4E3.text	C65F8C4C37DD5B2B880B9F1B8220A4E3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Megacoxa janzeni Brazidec & Vilhelmsen & Boudinot & Richter & Hammel & Perkovsky & Fan & Wang & Wu & Wang & Perrichot 2024	<div><p>3.1.1.3.2. † Megacoxa janzeni sp. nov.</p><p>Figures 2E and 7</p><p>Etymology.</p><p>The specific epithet is a patronym honoring Jens-Wilhelm Janzen, former owner of the holotype specimen.</p><p>Material studied.</p><p>Holotype female IGR.BU-069; housed in the amber collection of the Geology Department and Museum of the University of Rennes, France (IGR). One paratype male NIGP203547; housed in the Nanjing Institute of Geology and Paleontology (NIGP), Chinese Academy of Sciences, Nanjing, China.</p><p>Type locality.</p><p>Noije Bum Hill, Hukawng Valley, Kachin State, Myanmar.</p><p>Age.</p><p>Upper Albian to lower Cenomanian, mid-Cretaceous.</p><p>Diagnosis.</p><p>Median mesoscutal sulcus smooth (vs. crenulate in Megacoxa synchrotron gen. et sp. nov.); axillae not contiguous medially (vs. contiguous in † Megacoxa chandrahrasa gen. et sp. nov. and Megacoxa synchrotron gen. et sp. nov.); fore wing with very narrow marginal cell closed in straight line by Rs (vs. broad with Rs curved in † Megacoxa chandrahrasa gen. et sp. nov.); trapezoid medial cell (Fig. 2E; vs. narrow in † Megacoxa chandrahrasa gen. et sp. nov.); posterior margin of metacoxa outcurved and bordered with large foveae (Fig. 7B; vs. bordered with small foveae in † Megacoxa chandrahrasa gen. et sp. nov. and and straight posterior metacoxal margin in Megacoxa synchrotron gen. et sp. nov.).</p><p>Description.</p><p>Body length for holotype 5.20 mm; for paratype 3.72 mm. - Head glabrous, globular, higher than long (length for holotype 0.81 mm; for paratype 0.69 mm; height for holotype 1.04 mm; for paratype 0.75 mm); compound eye oval, higher than long, ocular carina absent; vertex convex, shagreened; clypeus short; subantennal groove present; antenna short, less than half as long as body; scape as long as wide (length for holotype 0.19 mm); pedicel wider than long (length for holotype 0.12 mm); flagellomeres cylindrical, elongate; mandibles symmetrical, with three teeth; occipital carina minutely crenulate. - Mesosoma half as long as body length (length for holotype 2.28 mm; for paratype 1.62 mm); mesoscutum one third as long as mesosoma (length for holotype 0.92 mm; for paratype 0.63 mm; width for holotype 1.03 mm), shagreened, overhanging pronotum almost at right angle, divided by smooth median mesoscutal sulcus; parapsidal line present; axillae not contiguous medially, inner axillar groove crenulate; pronotum with posteromedial part moderately high as viewed laterally; anterior thoracic spiracle not fully surrounded by pronotal cuticle; mesometapectal sulcus crenulate; propodeum carinate. - Fore wing hyaline and covered with microtrichiae, about two thirds of body length (length for holotype 3.08 mm; for paratype 2.58 mm); R1, Sc+R and M+Cu pigmented, Rs present between r-rs and Rs+M, composing first submarginal cell, M and A pigmented, Rs closing in a straight line a very narrow medial cell, trapezoid medial cell composed by basal segments of M, Rs+M and Cu1 (shortest side) and by 1m-Cu, Cu pigmented to margin with a bend. - Legs with metacoxa enlarged, posteriorly curved, and notched just before trochanter insertion, bearing a row of large foveae along posterior margin; two mesotibial spurs present; metafemur swollen and bare; metatibia also swollen, with two spurs and bearing row of comb-like setae that continue on metabasitarsus; metatarsus almost as long as metatibia; metabasitarsus four times longer than following metatarsomeres. - Metasoma shorter than mesosoma, covered with short setae (length for holotype 2.12 mm; for paratype 1.41 mm); first tergite longest and second almost same length; third, fourth and fifth of similar length and shorter than first; last tergite twice as long as fifth; tergite and sternite inserted in bevel; hypopygium well-developed; ovipositor same length as metasoma (length for holotype 2.10 mm; OL/BL ratio 0.40), sheaths fully preserved in holotype, transversely striated.</p></div>	https://treatment.plazi.org/id/C65F8C4C37DD5B2B880B9F1B8220A4E3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Brazidec, Manuel;Vilhelmsen, Lars;Boudinot, Brendon E.;Richter, Adrian;Hammel, Joerg U.;Perkovsky, Evgeny E.;Fan, Yong;Wang, Zhen;Wu, Qiong;Wang, Bo;Perrichot, Vincent	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
46F7D5E75DAC5FD58D02AC732C02AC03.text	46F7D5E75DAC5FD58D02AC732C02AC03.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Megacoxa synchrotron Brazidec & Vilhelmsen & Boudinot & Richter & Hammel & Perkovsky & Fan & Wang & Wu & Wang & Perrichot 2024	<div><p>3.1.1.3.3. † Megacoxa synchrotron sp. nov.</p><p>Figures 2F and 8</p><p>Etymology.</p><p>The specific epithet is an adjective and refers to the µ-CT technique used at the Deutsches Elektronen-Synchrotron (DESY; Hamburg, Germany) to reconstruct the external surface of the specimen, as well as the internal features for a later study.</p><p>Material studied.</p><p>Holotype male CASENT0753237; housed in the collection of the Phyletisches Museum Jena, Germany (PMJ).</p><p>Type locality.</p><p>Noije Bum Hill, Hukawng Valley, Kachin State, Myanmar.</p><p>Age.</p><p>Upper Albian to lower Cenomanian, mid-Cretaceous.</p><p>Diagnosis.</p><p>Flagellomeres 1-7 distinctly longer than wide; flagellomeres 8-12 slightly longer than wide (Fig. 8A); median mesoscutal sulcus crenulate (vs. smooth in † Megacoxa chandrahrasa gen. et sp. nov. and † Megacoxa janzeni gen. et sp. nov.); axillae contiguous medially (Figs 8D and F; vs. separated medially in † Megacoxa janzeni gen. et sp. nov.); fore wing with medial cell wide (Fig. 2F; vs. narrow in † Megacoxa chandrahrasa gen. et sp. nov.); posterior margin of metacoxa straight, with large foveae (Figs 8C and G; vs. rounded with small foveae in † Megacoxa chandrahrasa gen. et sp. nov. and rounded with large foveae in † Megacoxa janzeni gen. et sp. nov.); comb-like setae reduced on the metabasitarsus (Fig. 8A; vs. more developed on metatibia and metabasitarsus in † Megacoxa chandrahrasa gen. et sp. nov. and † Megacoxa janzeni gen. et sp. nov.).</p><p>Description.</p><p>Body length 2.89 mm. - Head globular, higher than long (length 0.45 mm; height 0.53 mm), glabrous; frons convex, shagreened; compound eye oval, higher than long, without postocular carina; vertex convex, shagreened; clypeus short, shagreened; subantennal groove present; antenna filiform, reaching metacoxa; scape short and wide (length 0.09 mm); pedicel thinner than scape, shorter than flagellomeres (length 0.07 mm); flagellomeres cylindrical, elongate; flagellomeres 1-7 distinctly longer than wide (length ca. 0.11-0.14 mm); flagellomeres 8-11 slightly longer than wide (length 0.09 mm); flagellomere 12 longest (length 0.16 mm); mandible with three teeth; occipital carina minutely crenulate. - Mesosoma longer than metasoma (length 1.27 mm); mesoscutum (length 0.48 mm) convex, shagreened, divided by crenulate median mesoscutal sulcus; parapsidal lines present; axillae contiguous medially; inner axillar groove crenulate; mesoscutellum diamond-shaped, with blunt posterior margin; pronotum shagreened, with posteromedial part moderately high as viewed laterally; anterior thoracic spiracle not fully surrounded by pronotal cuticle; mesometapectal sulcus crenulate; propodeum carinate, carinae delimiting large fovea on median anterior region, smaller foveae laterally, large foveae on lateral median region and large triangular foveae on posterior region. - Fore wing hyaline, uniformly micropubescent (length 2.04 mm); venation complete with R1, Sc+R, M+Cu, A, Rs+M, Rs, M and Cu fully pigmented; Rs+M aligned with M+Cu; medial cell wide; Rs present between Rs+M and r-rs, closing first submarginal cell; marginal cell narrow, closed in straight line by Rs. - Legs with metacoxa enlarged, posterior margin straight with upper corner angularly produced, bearing row of large foveae along posterior margin; metafemur and metatibia swollen; two mesotibial and two metatibial spurs; metabasitarsus with row of sparse comb-like setae along ventral margin. - Metasoma smooth, fusiform (length 1.17 mm); tergites inserted in bevel.</p></div>	https://treatment.plazi.org/id/46F7D5E75DAC5FD58D02AC732C02AC03	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Brazidec, Manuel;Vilhelmsen, Lars;Boudinot, Brendon E.;Richter, Adrian;Hammel, Joerg U.;Perkovsky, Evgeny E.;Fan, Yong;Wang, Zhen;Wu, Qiong;Wang, Bo;Perrichot, Vincent	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
9F64490A6EBD5C7593FD9F398386375C.text	9F64490A6EBD5C7593FD9F398386375C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Megalavini Brazidec & Vilhelmsen & Boudinot & Richter & Hammel & Perkovsky & Fan & Wang & Wu & Wang & Perrichot 2024	<div><p>3.1.1. Tribe † Megalavini trib. nov.</p><p>Type genus.</p><p>† Megalava Perrichot, 2009.</p><p>Diagnosis.</p><p>Mandible with three teeth, decreasing in size from apex to base; occipital carina crenulate. Pronotum not visible dorsally; mesometapectal sulcus (sensu Vilhelmsen et al. 2010b) crenulate. Fore wing venation most complete within Megalyridae; pterostigma reduced; C, M+Cu, Sc+R, A and M almost fully pigmented; R1 extending beyond marginal cell; marginal cell closed by Rs in straight line; Rs present between r-rs and Rs+M, closing first submarginal cell; M+Cu aligned with Rs+M; medial cell rectangular or trapezoidal, located under Rs+M and closed by 1m-cu and basal segments of Cu and Cu1. Metasoma elongate, with long to very long ovipositor (OL/BL ~ 0.40).</p><p>Genera and species included.</p><p>† Megalava truncata Perrichot, 2009, † Cretolyra noijebumensis gen. et sp. nov., † Cretolyra shawi gen. et sp. nov., † Genkyhag innebula gen. et sp. nov., † Megacoxa chandrahrasa gen. et sp. nov., † Megacoxa janzeni gen. et sp. nov., † Megacoxa synchrotron gen. et sp. nov.</p><p>Comments.</p><p>The † Megalavini trib. nov. retain a plesiomorphic fore wing venation (i.e., the most complete wing venation among megalyrids sensu stricto) as a key character with C, Sc+R, M+Cu, A, Rs+M, Cu, R1, Rs and M almost fully pigmented (Figs 2A-F). The main synapomorphy for this clade is the form of the medial cell, which is rectangular due to alignment of Rs+M with M+Cu (Figs 2A-F). Comprising exclusively Cretaceous taxa, † Megalavini trib. nov. also display an anterior thoracic spiracle that is not fully surrounded by pronotal cuticle (Fig. 3D), which we consider to be the second diagnostic feature (despite being plesiomorphic) for the tribe. When described, the monospecific genus † Megalava was stated to belong to the tribe † Megazarini with † Megazar, and this has been confirmed in Vilhelmsen et al. (2010a). However, based on the description of a new complete specimen ( Pérez-de la Fuente et al. 2012: fig. 2B), we propose to transfer † Megalava to the new tribe, due its more complete wing venation, the rectangular medial cell, the crenulate mesometapectal sulcus and the mandible configuration (three teeth while † Megazar has four).</p></div>	https://treatment.plazi.org/id/9F64490A6EBD5C7593FD9F398386375C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Brazidec, Manuel;Vilhelmsen, Lars;Boudinot, Brendon E.;Richter, Adrian;Hammel, Joerg U.;Perkovsky, Evgeny E.;Fan, Yong;Wang, Zhen;Wu, Qiong;Wang, Bo;Perrichot, Vincent	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
CA8E03AACD4E5FBEB6F80109E6CD78CE.text	CA8E03AACD4E5FBEB6F80109E6CD78CE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Megalyrini Perrichot & Perkovsky 2009	<div><p>3.2.3. Tribe Megalyrini Schletterer, 1889</p><p>Cretodinapsini Rasnitsyn, 1977 syn. nov.</p><p>Prodinapsini Shaw, 1990b</p><p>Type genus.</p><p>Megalyra Westwood, 1832.</p><p>Emended diagnosis.</p><p>Head hypognathous; compound eye often with distinct setation; flagellomeres elongate; median sulcus on vertex present; occipital carina present, curving towards mandible. Parapsidal lines absent. Fore wing with C, Sc+R, 1A, Rs+M, basal segment of M, R1, r-rs and small segment of Rs tubular. Metasoma elongate, subcylindrical, longer than mesosoma.</p><p>Genera and species included.</p><p>† C. caucasica Rasnitsyn, 1977, M. aquilonia Shaw, 1990, M. australia Girault, 1925, † M. baltica Poinar &amp; Shaw, 2007, M. brevicauda Shaw, 1990, M. caledonica Vachal, 1908, M. candata Szépligeti, 1902, M. exigua Shaw, 1990, M. fasciipennis Westwood, 1832, M. globula Shaw, 1990, M. gnoma Shaw, 1990, M. lilliputiana Turner, 1916, M. longiseta Szépligeti, 1902, M. minuta Froggatt, 1906, M. nanella Shaw, 1990, M. plana Shaw, 1990, M. pygmaea Shaw, 1990, M. reiki Shaw, 1990, M. rufipes Erichson, 1841, M. rufiventris Szépligeti, 1902, M. sedlaceki Shaw, 1990, M. shuckardi Westwood, 1851, M. spectabilis Shaw, 1990, M. tawiensis Petersen, 1966, M. testaceipes Turner, 1916, M. transversistriata Girault, 1925, M. troglodytes Naumann, 1987, M. viridescens Froggatt, 1906, M. wagneri Fahringer, 1928, † P. bruesi (Perrichot, 2009), † P. janzeni Perrichot, 2009, † P. minor Brues, 1933, † P. oesiensis Perrichot, 2009, † P. prolata (Perrichot &amp; Perkovsky, 2009), † P. pumilio Perrichot &amp; Perkovsky, 2009, † P. succinalis Brues, 1923.</p><p>Comments.</p><p>The tribe Megalyrini is expanded based on the results of the phylogenetic analyses. The genus † Cretodinapsis is retrieved to be sister of Megalyra, rendering the † Cretodinapsini paraphyletic. Therefore, † Cretodinapsini is synonymized with Megalyrini, with † Cretodinapsis and † Prodinapsis now components of the latter tribe.</p></div>	https://treatment.plazi.org/id/CA8E03AACD4E5FBEB6F80109E6CD78CE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Brazidec, Manuel;Vilhelmsen, Lars;Boudinot, Brendon E.;Richter, Adrian;Hammel, Joerg U.;Perkovsky, Evgeny E.;Fan, Yong;Wang, Zhen;Wu, Qiong;Wang, Bo;Perrichot, Vincent	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
E48F5B8FE6EB51A2AB715E51476381B2.text	E48F5B8FE6EB51A2AB715E51476381B2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Megallicini Brazidec & Vilhelmsen & Boudinot & Richter & Hammel & Perkovsky & Fan & Wang & Wu & Wang & Perrichot 2024	<div><p>3.2.2. Tribe † Megallicini trib. nov.</p><p>Type genus.</p><p>† Megallica Perrichot, 2009.</p><p>Diagnosis.</p><p>Head hypognathous; compound eye without setation, without posterior carina; subantennal groove without dorsal carina; mandible with three teeth; median sulcus on vertex present; occipital carina present, curving towards mandible. Median mesoscutal sulcus crenulate; parapsidal lines absent; axillae not connected at inner angles, separated by two triangular foveae; propodeum areolate-rugose. Fore wing hyaline; with C, Sc+R, A, cu-a, Rs+M, M, Rs and R1 tubular; pterostigma large; M not reaching apical margin; Rs only present as a stub between Rs+M and r-rs; Rs apically evenly arched towards stigma; costal, marginal and radial+first cubital cells closed. Metasoma elongate, fusiform, longer than mesosoma.</p><p>Genera and species included.</p><p>† Megallica parva Perrichot, 2009</p><p>Comments.</p><p>The tribe Megallicini trib. nov. is erected based on the results of the phylogenetic analyses, because Megallica is not retrieved within a clade with any other Megalyrinae genera.</p></div>	https://treatment.plazi.org/id/E48F5B8FE6EB51A2AB715E51476381B2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Brazidec, Manuel;Vilhelmsen, Lars;Boudinot, Brendon E.;Richter, Adrian;Hammel, Joerg U.;Perkovsky, Evgeny E.;Fan, Yong;Wang, Zhen;Wu, Qiong;Wang, Bo;Perrichot, Vincent	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
06A075176FB150B7AFC036ADEF9DF13E.text	06A075176FB150B7AFC036ADEF9DF13E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Valaaini Brazidec & Vilhelmsen & Boudinot & Richter & Hammel & Perkovsky & Fan & Wang & Wu & Wang & Perrichot 2024	<div><p>3.2.4. Tribe † Valaaini trib. nov.</p><p>Type genus.</p><p>† Valaa Perrichot, 2009.</p><p>Diagnosis.</p><p>Head hypognathous; compound eye without setation, without posterior carina; subantennal groove without dorsal carina; mandible with three teeth; occipital carina present, curving towards mandible. Median mesoscutal sulcus crenulate; parapsidal lines absent; axillae not connected at inner angles, separated by two triangular foveae; propodeum carinate. Protibia with stout apical spines; pro-, meso- and metatibiae shorter than combined length of tarsomeres. Fore wing hyaline, with C, Sc+R, A, Rs+M, cu-a, Rs, M and R1 tubular; pterostigma reduced; M not reaching apical margin; Rs apically strongly angled, arched towards stigma; costal, submarginal, marginal and radial+first cubital cells closed. Hind wing with Rs short, barely projecting beyond R1. Metasoma compact, shorter than mesosoma.</p><p>Genera and species included.</p><p>† Valaa delclosi Perrichot, 2009</p><p>Comments.</p><p>The tribe † Valaaini trib. nov. is erected based on the results of the phylogenetic analyses, because † Valaa is not retrieved within a clade with other Dinapsini s.s., contrary to what was suggested by Perrichot (2009).</p></div>	https://treatment.plazi.org/id/06A075176FB150B7AFC036ADEF9DF13E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Brazidec, Manuel;Vilhelmsen, Lars;Boudinot, Brendon E.;Richter, Adrian;Hammel, Joerg U.;Perkovsky, Evgeny E.;Fan, Yong;Wang, Zhen;Wu, Qiong;Wang, Bo;Perrichot, Vincent	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
