taxonID	type	description	language	source
B55459FED26655CB892501A02113017D.taxon	diagnosis	Diagnosis. Head hypognathous, globular; compound eye oval, not covering head length, without postocular carina; flagellomeres cylindrical, longer than wide; subantennal groove without dorsal carina; mandibles symmetrical, with three or four teeth. Posterolateral margin of pronotum with notch accommodating anterior thoracic spiracle, not surrounding spiracle posteriorly; mesoscutum large, around half length of mesosoma, parapsidal lines present; median mesoscutal sulcus and axillar grooves present, deeply crenulate or smooth. Metasoma elongate. Fore wing hyaline; C, Sc + R, M + Cu, A, Rs + M, cu-a, Cu, Rs and R 1 pigmented; costal, radial, first cubital, submarginal, and marginal cells closed by tubular veins. Hind leg stout, metafemur and metatibia swollen; at least either inner margin of metatrochanter, metacoxa, metatibia, or metabasitarsus bearing row of comb-like, thick setae along ventral surface.	en	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
3F5C11CCF27B5F0391E4012059EA0EB3.taxon	etymology	Etymology. Combination of Creto - referring to the Cretaceous age of the Burmese amber deposit and the suffix - lyra often used in megalyrid genus names. Gender feminine.	en	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
3F5C11CCF27B5F0391E4012059EA0EB3.taxon	diagnosis	Diagnosis. Compound eye oval, higher than long; frons longitudinally divided by sulcus anterior to median ocellus; flagellomere 1 shorter than individual length of flagellomeres 2 - 5; occipital carina crenulate. Fore wing with M + Cu, Sc + R, A, M and Cu fully pigmented; cu-a almost branching with A, absent for short distance; medial cell rectangular; marginal cell narrow, closed in straight line (Figs 2 A, B, 3 C and 4 A). Median mesoscutal sulcus smooth; parapsidal lines present, diverging anteriorly (Figs 3 B and 4 A). Metacoxa long; metafemur and metatibia swollen; two mesotibial and one metatibial spurs; row of comb-like setae along ventral surface of metatibia and metabasitarsus. Metasoma elongate and narrowed at apex, longer than mesosoma.	en	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
982B71A3D015503EACAFCDC54F6F2963.taxon	description	Figures 2 A, 3	en	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
982B71A3D015503EACAFCDC54F6F2963.taxon	etymology	Etymology. The specific epithet is an adjective referring to the type locality where the amber pieces were collected.	en	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
982B71A3D015503EACAFCDC54F6F2963.taxon	materials_examined	Material studied. Holotype male IGR. BU- 067; housed in the amber collection of the Geology Department and Museum of the University of Rennes, France (IGR).	en	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
982B71A3D015503EACAFCDC54F6F2963.taxon	diagnosis	Diagnosis. Antennae inserted on edge of clypeus (Fig. 3 A; vs. inserted above clypeal margin in Cretolyra shawi gen. et sp. nov.); axillae contiguous medially and axillar grooves crenulated (Fig. 3 B; vs axillae separated medially and axillar grooves smooth in Cretolyra shawi gen. et sp. nov.).	en	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
982B71A3D015503EACAFCDC54F6F2963.taxon	description	Description. Body length 3.33 mm; body very pubescent, especially on metasoma where setae conceal margins of tergites; the setae have agglomerated small bubbles that make observation difficult. - Head globular, higher than long (length 0.55 mm; height 0.75 mm); frons convex, smooth, divided by thin median sulcus; compound eye oval, higher than long; vertex convex, with smooth ocellar triangle; clypeus well-developed; torulus inserted closer to clypeus than to eye; shallow but distinct subantennal groove adjoining ventral margin of eye; antenna half as long as body, almost reaching metasoma; scape twice as long as wide (length 0.16 mm); pedicel shorter than scape (length 0.11 mm); flagellomeres cylindrical, elongate, longer than wide; flagellomere 1 shortest (length 0.15 mm); following flagellomeres longer (length ca. 0.21 mm); occipital carina crenulate. - Mesosoma almost half body length (length 1.29 mm; height 0.65 mm); mesoscutum convex, shagreened, divided by smooth median mesoscutal sulcus (mesoscutum length 0.58 mm; width ~ 0.60 mm); parapsidal lines present, diverging anteriorly; axillae contiguous medially, axillar groove crenulate; mesoscutellum diamond-shaped; pronotum smooth, not visible dorsally, with posteromedial part moderately high as viewed laterally; anterior thoracic spiracle not fully surrounded by pronotal cuticle; mesometapectal sulcus crenulate; propodeum shorter than mesoscutellum, with incomplete pairs of median and submedian carinae, two complete lateral longitudinal carinae, anterior region of propodeum with transversal row of foveae adjacent to concealed metanotum, posterior region of propodeum with three large foveae. - Fore wing hyaline and covered with microtrichiae, about two thirds of body length (length 2.50 mm); C, Sc + R, M + Cu, A, Rs fully pigmented; R 1 extending beyond marginal cell; Rs closing marginal cell in straight line; M pigmented to apex; medial cell rectangular, narrow; Cu almost reaching posterior wing margin. - Legs covered with setae; two mesotibial and one metatibial spurs; metacoxa elongate; metafemur and metatibia swollen; metabasitarsus three times as long as following metatarsomeres and with row of short comb-like setae along ventral surface. - Metasoma half body length (length 1.48 mm), elongate and narrowed at apex; seven smooth tergites with pubescent posterior margin; first tergite largest; second tergite one quarter of metasoma length; remaining tergites shorter.	en	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
3743BE522B3758F59647816F3AE30EA8.taxon	description	Figures 2 B, 4	en	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
3743BE522B3758F59647816F3AE30EA8.taxon	etymology	Etymology. The specific epithet is a patronym honoring Scott R. Shaw, for his contributions to the knowledge of Megalyridae.	en	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
3743BE522B3758F59647816F3AE30EA8.taxon	materials_examined	Material studied. Holotype female NIGP 203545; housed in the Nanjing Institute of Geology and Paleontology (NIGP), Chinese Academy of Sciences, Nanjing, China.	en	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
3743BE522B3758F59647816F3AE30EA8.taxon	diagnosis	Diagnosis. Pedicel bilobed, long and thick (Figs 4 D-E; vs. calciform and elongate in Cretolyra noijebumensis gen. et sp. nov.); axillae almost rounded and reduced on edges of mesoscutellum, not contiguous medially (vs. contiguous medially in Cretolyra noijebumensis gen. et sp. nov.); axillar grooves smooth (Fig. 4 A; vs. axillar grooves crenulated in Cretolyra noijebumensis gen. et sp. nov.); medial cell nearly triangular, with 1 m-cu very short (Fig. 2 B; vs. medial cell rectangular, with 1 m-cu distinct in Cretolyra noijebumensis gen. et sp. nov.).	en	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
3743BE522B3758F59647816F3AE30EA8.taxon	description	Description. Body length 2.82 mm. - Head globular, higher than long (length ca. 0.40 mm), covered with short setae; frons convex, smooth, divided by sulcus; compound eye oval, higher than long; vertex convex; clypeus smooth, apically rounded; mandibles symmetrical, with three teeth; toruli separated from each other by less than their own diameter; subantennal groove present; antenna about half body length; scape shorter than pedicel; pedicel thick, bilobed; flagellomeres cylindrical, longer than wide; flagellomere 1 shorter than flagellomeres 2 - 5 (length 0.15 mm vs. ca. 0.21 mm); flagellomeres 6 - 11 shorter (length ca. 0.10 mm); flagellomere 12 as long as flagellomere 1 (length 0.15 mm); occipital carina minutely crenulate. - Mesosoma one third of body length (length 0.98 mm); mesoscutum convex, divided by smooth median mesoscutal sulcus; parapsidal lines present; axillae small, almost rounded and not contiguous medially; axillar groove smooth; mesoscutellum smooth, convex, diamond-shaped; pronotum smooth, not visible dorsally, with posteromedial part moderately high as viewed laterally; anterior thoracic spiracle not surrounded by pronotal cuticle posteriorly; mesometapectal sulcus crenulate; propodeum carinate. - Fore wing hyaline and covered with microtrichiae, two thirds of body length (length 2.57 mm); pterostigma reduced; C, Sc + R, M + Cu, A, Rs fully pigmented; R 1 extending beyond marginal cell; Rs closing marginal cell in straight line; M pigmented to apex; medial cell small, elongate and rectangular, nearly triangular by side of 1 m-cu. - Legs with two mesotibial and one metatibial spurs; metacoxa elongate; metafemur and metatibia swollen; metabasitarsus two to three times as long as remaining metatarsomeres; row of comb-like setae along ventral surface of metatibia and metabasitarsus. - Metasoma longer than mesosoma (length 1.44 mm); elongate and narrowed at apex; tergites smooth; hypopygium well-developed; ovipositor shorter than metasoma (length 1.33 mm; OL / BL ratio 0.43), sheaths fully preserved, transversely striated.	en	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
6E91A808941E580DB5C2EB6B37060D64.taxon	etymology	Etymology. Anagram of Ga Hkyeng, meaning red soil in Jinghpaw, from which the term Kachin is derived. Gender masculine.	en	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
6E91A808941E580DB5C2EB6B37060D64.taxon	diagnosis	Diagnosis. Compound eye oval, higher than long; antenna inserted half way between eye and clypeus; flagellomeres cylindrical, subequal in length (Figs 5 C, D). Mesosoma less than half body length, shorter than metasoma; mesoscutum hiding pronotum in dorsal view; median mesoscutal sulcus and axillar groove crenulate; parapsidal lines present, parallel to median mesoscutal sulcus and diverging posteriorly; axillae not contiguous medially; mesoscutellum short (Figs 5 A, B); propodeum carinate. Fore wing hyaline, with microtrichiae; venation almost fully pigmented, with only Cu nebulous; medial cell trapezoidal and broad; marginal cell closed without bend; R 1 extending beyond this cell (Figs 2 C, 5 A, B, and E). Legs with one mesotibial and two metatibial spurs. Metasoma elongate, fusiform.	en	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
FCC83238CF44597C9C8CB733D9D5D303.taxon	description	Figures 2 C, 5	en	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
FCC83238CF44597C9C8CB733D9D5D303.taxon	etymology	Etymology. From Latin ' nebula, In nebula', meaning " in the fog ", in reference to the amber pieces where the specimens are found, that contains numerous terrigenous inclusions, sometimes partially hiding the specimens. It is a noun in apposition.	en	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
FCC83238CF44597C9C8CB733D9D5D303.taxon	materials_examined	Material studied. Holotype female NIGP 203546; housed in the Nanjing Institute of Geology and Paleontology (NIGP), Chinese Academy of Sciences, Nanjing, China. Three paratypes male FAI-BI 11324 a, male FAI-BI 11324 b, FAI-BI- 11324 c of unknown sex; housed in the Fushun Amber Institute.	en	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
FCC83238CF44597C9C8CB733D9D5D303.taxon	diagnosis	Diagnosis. As for genus.	en	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
FCC83238CF44597C9C8CB733D9D5D303.taxon	description	Description. Holotype female incompletely preserved (body length 3.12 mm), in a piece with multiple terrigenous inclusions. Paratypes preserved in a single piece: FAI-BI 11324 a visible in dorsal and frontal views, FAI-BI 11324 b in lateral view (body length ca. 3.45 mm), FAI-BI 11324 c being one complete fore and hind wings and a partial fore wing. - Head globular, apparently entirely glabrous (length for holotype ca. 0.47 mm); frons shagreened, convex, divided by a faintly impressed sulcus; compound eye oval, without postocular carina; vertex convex, shagreened; clypeus short; subantennal groove present; toruli inserted half way between clypeus and eyes; antenna short, barely reaching propodeum; scape globular; pedicel slightly longer; flagellomeres cylindrical, longer than wide; flagellomeres 7 - 11 shorter than flagellomeres 1 - 6; flagellomere 12 longest; occipital carina crenulate. - Mesosoma slightly shorter than metasoma (length for holotype 1.12 mm; for paratype 11324 a 1. 29 mm; for paratype 11324 b 1.57 mm); pronotum not visible dorsally; mesoscutum shagreened, half as long as mesosoma, median mesoscutal sulcus crenulate (mesoscutum length for holotype 0.51 mm; for paratype 11324 a 0.61 mm; width for holotype 0.65 mm; for paratype 11324 a 0.71 mm); parapsidal lines present; axillae barely contiguous medially and inner grooves crenulate; mesoscutellum wider than long, shorter than mesoscutum; pronotum smooth, not visible dorsally, with posteromedial part moderately high as viewed laterally; anterior thoracic spiracle not fully surrounded by pronotal cuticle; propodeum carinate, carinae delimitating row of small foveae in anterior region and larger foveae in posterior region. - Fore wing hyaline, covered with microtrichiae (length for holotype 1.84 mm; for paratype 11324 a 2.10 mm; for paratype 11324 b 2.11 mm; for paratype 11324 c 2.86 mm); venation almost complete, Sc + R, M + Cu, A, Rs + M and M fully pigmented; Cu spectral; marginal cell very narrow, closed in straight line; R 1 barely extending beyond marginal cell; medial cell rectangular and relatively broad. - Hind wing hyaline; venation reduced to Sc + R and R 1, Sc + R almost not diverging from margin, R 1 pigmented almost to tip; at least seven hamuli, located distal to middle of hind wing. - Legs with one mesotibial and two metatibial spurs; metafemur swollen; ventral surface of metatibia and metabasitarsus with row of comb-like setae, metabasitarsus at least three times as long as following metatarsomeres. - Metasoma elongate, oval, with sparse short setae (length for holotype 1.43 mm); tergites smooth; ovipositor shorter than metasoma (length for holotype 1.34 mm; OL / BL ratio 0.43).	en	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
7DDF1391E8C854A8A428E13D42F4B321.taxon	etymology	Etymology. Anagram of Taimyrski, the Russian name of the Taimyr peninsula from where the amber pieces containing the specimens derive. Gender masculine.	en	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
7DDF1391E8C854A8A428E13D42F4B321.taxon	diagnosis	Diagnosis. As for tribe with the following additional characters: head shagreened; compound eye higher than long; occipital carina foveate (Figs 9 C and F); flagellomeres cylindrical and elongate. Mesosoma less than half of metasoma length; mesoscutum reduced on anterior part of mesosoma, hiding pronotum dorsally (Figs 9 B and F). Fore wing with medial cell pentagonal with 1 m-cu, Cu 1 and M + Cu and Cu spectral, Rs + M not aligned with M + Cu (Figs 2 G, H, 9 B and D). Hind leg with scattered erect setae; one mesotibial and one metatibial spur.	en	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
6586EC0085A85D39BE714BC02321F92A.taxon	description	Figures 2 G, 9 A-C	en	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
6586EC0085A85D39BE714BC02321F92A.taxon	etymology	Etymology. " exfrigore " means " from the cold " in Latin as this species is the northernmost record of a megalyrid, extant and extinct, in a region where the temperature is currently below zero most of the year. The specific epithet is to be treated as an adjective.	en	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
6586EC0085A85D39BE714BC02321F92A.taxon	materials_examined	Material studied. Holotype female PIN 3730 / 411; housed in the collection of the Paleontological Institute of the Russian Academy of Sciences (PIN).	en	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
6586EC0085A85D39BE714BC02321F92A.taxon	diagnosis	Diagnosis. Eye nearly round (vs. oval, higher than long in † Kamyristi yantardakhensis gen. et sp. nov.); axillae contiguous medially with axillar groove smooth (Fig. 9 B; vs. separated medially with axillar groove crenulated in † Kamyristi yantardakhensis gen. et sp. nov.).	en	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
6586EC0085A85D39BE714BC02321F92A.taxon	description	Description. Body length 2.03 mm. Almost complete, except ovipositor. - Head globular, higher than long (length 0.38 mm; height 0.43); slightly pubescent on vertex and frons; frons convex, shagreened; compound eye rounded, slightly higher than long, not covering head length, postocular carina absent; vertex convex, shagreened; torulus inserted very close to ventral margin of eye; subantennal groove present; mandible with three teeth; scape short and thick (length 0.09 mm); pedicel same length; flagellomeres cylindrical, longer than wide, elongate (length ca. 0.08 mm); flagellomere 12 longest; occipital carina crenulate. - Mesosoma shorter than metasoma (length 0.69 mm; height 0.40 mm), almost fully shagreened; mesoscutum shagreened, short (length 0.20 mm; width 0.47 mm), one third of mesosoma length, divided by crenulate median mesoscutal sulcus, with sparse erect setae; axillae continuous medially, with inner axillar groove smooth; mesoscutellum as long as mesoscutum, posterior margin slightly outcurved; pronotum not visible dorsally; anterior thoracic spiracle fully surrounded by pronotal cuticle; propodeum areolate-rugose. - Fore wing hyaline (length ca. 1.50 mm) and covered with microtrichia; C, Sc + R, Rs + M, basal segments of Rs and M; M + Cu, Cu and 1 m-cu spectral; Rs not closing first submarginal cell and pigmented for two thirds between r-rs and Rs + M; M pigmented to level of 2 r-m and then spectral to wing tip; pterostigma broad; R 1 extending beyond marginal cell; Rs pigmented, arched toward stigma, closing marginal cell. - Legs bearing many scattered erect setae; one mesotibial spur, one metatibial spur present; metafemur swollen, metatibia notched before spur; metabasitarsus three times longer than following, remainder same length. - Metasoma almost half body length (length 0.96 mm), elongate, smooth; segments subequal in length; hypopygium well-developed, concave; ovipositor incomplete, covered by sheaths only at base; cerci triangular, exserted.	en	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
9292FEA4705259A6A6FB6637A3FB90C9.taxon	description	Figures 2 H, 9 D-F	en	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
9292FEA4705259A6A6FB6637A3FB90C9.taxon	etymology	Etymology. The specific epithet refers to the locality where the specimen was collected. The specific adjective is to be treated as a noun in a genitive case.	en	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
9292FEA4705259A6A6FB6637A3FB90C9.taxon	materials_examined	Material studied. Holotype male PIN 3311 / 2718; housed in the collection of the Paleontological Institute of the Russian Academy of Sciences (PIN).	en	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
9292FEA4705259A6A6FB6637A3FB90C9.taxon	diagnosis	Diagnosis. Axillae not contiguous medially, grooves crenulated (Fig. 9 F; vs. contiguous medially, with axillar grooves smooth in † Kamyristi exfrigore gen. et sp. nov.).	en	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
9292FEA4705259A6A6FB6637A3FB90C9.taxon	description	Description. Body length 1.47 mm. Body mostly bare. - Head slightly higher than long (length 0.26 mm; height 0.28 mm), glabrous; frons convex, shagreened; compound eye almond-shaped, higher than long, not covering head length, postocular carina absent; vertex convex, minutely shagreened but sculpture not fully visible; toruli inserted closer to each other than to eyes; subantennal groove present; mandible with three teeth, decreasing in size from apex to base; scape short and thick (length 0.07 mm), pedicel of similar length with scape but thinner; 10 flagellomeres preserved, cylindrical, elongate, longer than wide; flagellomere 1 shortest flagellomere (length 0.04 mm), flagellomeres 2 - 10 similar in length (length ca. 0.06 mm); flagellomere 11 - 12 missing; occipital carina crenulate. - Mesosoma more than a third of body length (length 0.43 mm; height 0.22 mm); mesoscutum less than one third of mesosoma length (length 0.15 mm; width 0.31 mm), shagreened and divided by crenulate median mesoscutal sulcus; axillae not continuous medially, with inner groove crenulate; mesoscutellum shagreened; pronotum not visible dorsally; anterior thoracic spiracle fully surrounded by pronotal cuticle; propodeum areolate-rugose. - Fore wing hyaline and covered with microtrichia (length ca. 0.75 mm); C, Sc + R, A, Rs and basal segment of M pigmented; M + Cu, Cu and 1 m-cu spectral; Rs + M not aligned with M + Cu; medial cell pentagonal; Rs almost closing submarginal cell, not fully pigmented between r-rs and Rs + M; R 1 pigmented beyond marginal cell; marginal cell closed by Rs with a bend; M pigmented halfway to wing tip and then spectral. - Legs bearing numerous scattered setae; one long mesotibial spur, one shorter metatibial spur; basimetatrochanter long, half metacoxa length; metafemur slightly swollen; first metatarsomere longest and bearing small erect setae. - Metasoma almost half body length, longer than mesosoma (length 0.75 mm), elongate; smooth and bare; tergites nearly equal in length; hypopygium small, parameres triangular, projecting downward.	en	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
BC564AF72F8A5166903D7990FE9A477D.taxon	diagnosis	Diagnosis. Head prognathous; compound eye without setation, without posterior carina; subantennal groove without dorsal carina; mandible with three teeth; occipital carina present, curving towards mandible. Median mesoscutal sulcus crenulate; parapsidal line absent; propodeum areolate-rugose. Fore wing hyaline with C, Sc + R, A, cu-a, Rs + M, M, Rs and R 1 tubular; pterostigma large; M not reaching apical margin; Rs only present as a stub between Rs + M and r-rs; Rs apically strongly angled, arched towards stigma; Metasoma elongate, fusiform, longer than mesosoma; ovipositor sheaths only covering basal basal portion of ovipositor.	en	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
A5AC297D1FAF55B89C817FFB71BFF7D8.taxon	etymology	Etymology. The genus name refers to the size of the metacoxa, which is the key diagnostic feature of the genus; besides, the prefix Mega - alludes to the family name Megalyridae. Gender feminine.	en	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
A5AC297D1FAF55B89C817FFB71BFF7D8.taxon	diagnosis	Diagnosis. Compound eye oval; scape wider than long; thin occipital carina slightly crenulate. Parapsidal line present and located on posterolateral part of mesoscutum (Figs 6 C, 7 B and E); inner axillar groove crenulate (Fig. 8 F). Fore wing venation with R 1, M + Cu, Sc + R, A and Cu fully pigmented; A connected to medial cell with cu-a; Rs closing marginal cell without bend (Figs 2 D-F). Metacoxa much enlarged, with dorsal surface rounded to angular, outer metacoxal surface posteriorly foveate, inner metacoxal surface almost entirely accommodated in anterolateral, shallow concavity of metasoma (Figs 6 C, 7 B, 8 C and G). Metatibia and metabasitarsus with ventral row of comb-like setae; two mesotibial and two metatibial spurs (Figs 6 A and 7 A). Metasoma slightly shorter than mesosoma, tergites and sternites beveled (Figs 6 B and 7 A).	en	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
99C0269FADDF5E79BA4155230B90090D.taxon	description	Figures 2 D, 6	en	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
99C0269FADDF5E79BA4155230B90090D.taxon	etymology	Etymology. The specific epithet is a noun in apposition. It is the name of the legendary sword of Manjusrī, a bodhisattva in Buddhism, which allowed him to open the Kathmandu valley, and refers to the long, sword-like metatibial spurs of this species.	en	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
99C0269FADDF5E79BA4155230B90090D.taxon	materials_examined	Material studied. Holotype female IGR. BU- 068; housed in the amber collection of the Geology Department and Museum of the University of Rennes, France (IGR). Paratype female CNU-HYM-MA 2016207; housed in the Key Lab of Insect Evolution and Environmental Changes, College of Life Sciences, Capital Normal University, Beijing, China (CNU).	en	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
99C0269FADDF5E79BA4155230B90090D.taxon	diagnosis	Diagnosis. Median mesoscutal sulcus smooth (Fig. 6 F; vs. crenulate in Megacoxa synchrotron gen. et sp. nov.); axillae contiguous medially (Fig. 6 A; vs. separated medially in Megacoxa janzeni gen. et sp. nov.); medial cell elongate, almost triangular (Figs 2 D and 6 A; vs. broad and rectangular in Megacoxa synchrotron gen. et sp. nov. and Megacoxa janzeni gen. et sp. nov.); posterior margin of metacoxa outcurved and bordered with small foveae (Fig. 6 C; vs. bordered with large foveae in Megacoxa janzeni gen. et sp. nov.. and straight posterior metacoxal margin with large foveae in Megacoxa synchrotron gen. et sp. nov.); metatibial spurs very long (Figs 6 A, B and G); metasoma compressed, distinctly shorter than ovipositor; OL / BL ratio ~ 0.70.	en	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
99C0269FADDF5E79BA4155230B90090D.taxon	description	Description. Body length for holotype 2.64 mm; for paratype 4.80 mm. - Head globular, higher than long (length for holotype 0.52 mm; for paratype 0.93 mm), glabrous; frons convex, shagreened; compound eye oval, higher than long; vertex convex; clypeus short, shagreened; toruli inserted closer to each other than to eyes; subantennal groove present; antennae distorted, flagellomeres apparently cylindrical, longer than wide; occipital carina crenulate. - Mesosoma more than half body length (length for holotype 1.43 mm; for paratype 1.92 mm); covered with short setae; mesoscutum convex, wide (length for holotype 0.62 mm, width for holotype 0.74 mm; length for holotype 0.93 mm, width for holotype 0.90 mm), divided by smooth median mesoscutal sulcus; parapsidal lines present; axillae contiguous medially, inner axillar groove crenulate; pronotum not visible dorsally; anterior thoracic spiracle not fully surrounded by pronotal cuticle; mesometapectal sulcus crenulate; propodeum shorter than mesoscutellum, carinate. - Fore wing hyaline and covered with microtrichiae, slightly wrinkled at apex, as long as body (length for holotype 2.08 mm; for paratype 3.52 mm); venation complete with R 1, M + Cu, Sc + R, Rs, A and Cu fully pigmented; Rs closing marginal cell; medial cell elongate; A connected to medial cell by 1 cu-a. - Legs distorted; two mesotibial spurs; metacoxa huge, almost as high anteriorly as maximal propodeal height, with dorsal surface semicircular, posteroventral corner acute, ventral surface notched just anteriad trochanter insertion; outer metacoxal surface posteriorly with row of small foveae; dense row of short, comb-like setae along ventral surface of metatrochanter (full length), metafemur (basal half length), metatibia (distal half length), and metabasitarsus (full length); metafemur swollen; two long metatibial spurs, about twice as long as maximal tibial width; metabasitarsus three to four times longer than following metatarsomeres. - Metasoma relatively short (length for holotype 1.21 mm; for paratype 1.95 mm), distorted, partly intruding into propodeum during fossilization; tergites and sternites inserted in bevel, smooth; ovipositor longer than metasoma (length for holotype 1.87 mm, OL / BL ratio 0.70; length for paratype 3.24 mm, OL / BL ratio 0.68), sheaths transversely striated.	en	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
C65F8C4C37DD5B2B880B9F1B8220A4E3.taxon	description	Figures 2 E and 7	en	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
C65F8C4C37DD5B2B880B9F1B8220A4E3.taxon	etymology	Etymology. The specific epithet is a patronym honoring Jens-Wilhelm Janzen, former owner of the holotype specimen.	en	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
C65F8C4C37DD5B2B880B9F1B8220A4E3.taxon	materials_examined	Material studied. Holotype female IGR. BU- 069; housed in the amber collection of the Geology Department and Museum of the University of Rennes, France (IGR). One paratype male NIGP 203547; housed in the Nanjing Institute of Geology and Paleontology (NIGP), Chinese Academy of Sciences, Nanjing, China.	en	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
C65F8C4C37DD5B2B880B9F1B8220A4E3.taxon	diagnosis	Diagnosis. Median mesoscutal sulcus smooth (vs. crenulate in Megacoxa synchrotron gen. et sp. nov.); axillae not contiguous medially (vs. contiguous in † Megacoxa chandrahrasa gen. et sp. nov. and Megacoxa synchrotron gen. et sp. nov.); fore wing with very narrow marginal cell closed in straight line by Rs (vs. broad with Rs curved in † Megacoxa chandrahrasa gen. et sp. nov.); trapezoid medial cell (Fig. 2 E; vs. narrow in † Megacoxa chandrahrasa gen. et sp. nov.); posterior margin of metacoxa outcurved and bordered with large foveae (Fig. 7 B; vs. bordered with small foveae in † Megacoxa chandrahrasa gen. et sp. nov. and and straight posterior metacoxal margin in Megacoxa synchrotron gen. et sp. nov.).	en	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
C65F8C4C37DD5B2B880B9F1B8220A4E3.taxon	description	Description. Body length for holotype 5.20 mm; for paratype 3.72 mm. - Head glabrous, globular, higher than long (length for holotype 0.81 mm; for paratype 0.69 mm; height for holotype 1.04 mm; for paratype 0.75 mm); compound eye oval, higher than long, ocular carina absent; vertex convex, shagreened; clypeus short; subantennal groove present; antenna short, less than half as long as body; scape as long as wide (length for holotype 0.19 mm); pedicel wider than long (length for holotype 0.12 mm); flagellomeres cylindrical, elongate; mandibles symmetrical, with three teeth; occipital carina minutely crenulate. - Mesosoma half as long as body length (length for holotype 2.28 mm; for paratype 1.62 mm); mesoscutum one third as long as mesosoma (length for holotype 0.92 mm; for paratype 0.63 mm; width for holotype 1.03 mm), shagreened, overhanging pronotum almost at right angle, divided by smooth median mesoscutal sulcus; parapsidal line present; axillae not contiguous medially, inner axillar groove crenulate; pronotum with posteromedial part moderately high as viewed laterally; anterior thoracic spiracle not fully surrounded by pronotal cuticle; mesometapectal sulcus crenulate; propodeum carinate. - Fore wing hyaline and covered with microtrichiae, about two thirds of body length (length for holotype 3.08 mm; for paratype 2.58 mm); R 1, Sc + R and M + Cu pigmented, Rs present between r-rs and Rs + M, composing first submarginal cell, M and A pigmented, Rs closing in a straight line a very narrow medial cell, trapezoid medial cell composed by basal segments of M, Rs + M and Cu 1 (shortest side) and by 1 m-Cu, Cu pigmented to margin with a bend. - Legs with metacoxa enlarged, posteriorly curved, and notched just before trochanter insertion, bearing a row of large foveae along posterior margin; two mesotibial spurs present; metafemur swollen and bare; metatibia also swollen, with two spurs and bearing row of comb-like setae that continue on metabasitarsus; metatarsus almost as long as metatibia; metabasitarsus four times longer than following metatarsomeres. - Metasoma shorter than mesosoma, covered with short setae (length for holotype 2.12 mm; for paratype 1.41 mm); first tergite longest and second almost same length; third, fourth and fifth of similar length and shorter than first; last tergite twice as long as fifth; tergite and sternite inserted in bevel; hypopygium well-developed; ovipositor same length as metasoma (length for holotype 2.10 mm; OL / BL ratio 0.40), sheaths fully preserved in holotype, transversely striated.	en	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
46F7D5E75DAC5FD58D02AC732C02AC03.taxon	description	Figures 2 F and 8	en	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
46F7D5E75DAC5FD58D02AC732C02AC03.taxon	etymology	Etymology. The specific epithet is an adjective and refers to the µ-CT technique used at the Deutsches Elektronen-Synchrotron (DESY; Hamburg, Germany) to reconstruct the external surface of the specimen, as well as the internal features for a later study.	en	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
46F7D5E75DAC5FD58D02AC732C02AC03.taxon	materials_examined	Material studied. Holotype male CASENT 0753237; housed in the collection of the Phyletisches Museum Jena, Germany (PMJ).	en	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
46F7D5E75DAC5FD58D02AC732C02AC03.taxon	diagnosis	Diagnosis. Flagellomeres 1 - 7 distinctly longer than wide; flagellomeres 8 - 12 slightly longer than wide (Fig. 8 A); median mesoscutal sulcus crenulate (vs. smooth in † Megacoxa chandrahrasa gen. et sp. nov. and † Megacoxa janzeni gen. et sp. nov.); axillae contiguous medially (Figs 8 D and F; vs. separated medially in † Megacoxa janzeni gen. et sp. nov.); fore wing with medial cell wide (Fig. 2 F; vs. narrow in † Megacoxa chandrahrasa gen. et sp. nov.); posterior margin of metacoxa straight, with large foveae (Figs 8 C and G; vs. rounded with small foveae in † Megacoxa chandrahrasa gen. et sp. nov. and rounded with large foveae in † Megacoxa janzeni gen. et sp. nov.); comb-like setae reduced on the metabasitarsus (Fig. 8 A; vs. more developed on metatibia and metabasitarsus in † Megacoxa chandrahrasa gen. et sp. nov. and † Megacoxa janzeni gen. et sp. nov.).	en	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
46F7D5E75DAC5FD58D02AC732C02AC03.taxon	description	Description. Body length 2.89 mm. - Head globular, higher than long (length 0.45 mm; height 0.53 mm), glabrous; frons convex, shagreened; compound eye oval, higher than long, without postocular carina; vertex convex, shagreened; clypeus short, shagreened; subantennal groove present; antenna filiform, reaching metacoxa; scape short and wide (length 0.09 mm); pedicel thinner than scape, shorter than flagellomeres (length 0.07 mm); flagellomeres cylindrical, elongate; flagellomeres 1 - 7 distinctly longer than wide (length ca. 0.11 - 0.14 mm); flagellomeres 8 - 11 slightly longer than wide (length 0.09 mm); flagellomere 12 longest (length 0.16 mm); mandible with three teeth; occipital carina minutely crenulate. - Mesosoma longer than metasoma (length 1.27 mm); mesoscutum (length 0.48 mm) convex, shagreened, divided by crenulate median mesoscutal sulcus; parapsidal lines present; axillae contiguous medially; inner axillar groove crenulate; mesoscutellum diamond-shaped, with blunt posterior margin; pronotum shagreened, with posteromedial part moderately high as viewed laterally; anterior thoracic spiracle not fully surrounded by pronotal cuticle; mesometapectal sulcus crenulate; propodeum carinate, carinae delimiting large fovea on median anterior region, smaller foveae laterally, large foveae on lateral median region and large triangular foveae on posterior region. - Fore wing hyaline, uniformly micropubescent (length 2.04 mm); venation complete with R 1, Sc + R, M + Cu, A, Rs + M, Rs, M and Cu fully pigmented; Rs + M aligned with M + Cu; medial cell wide; Rs present between Rs + M and r-rs, closing first submarginal cell; marginal cell narrow, closed in straight line by Rs. - Legs with metacoxa enlarged, posterior margin straight with upper corner angularly produced, bearing row of large foveae along posterior margin; metafemur and metatibia swollen; two mesotibial and two metatibial spurs; metabasitarsus with row of sparse comb-like setae along ventral margin. - Metasoma smooth, fusiform (length 1.17 mm); tergites inserted in bevel.	en	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
9F64490A6EBD5C7593FD9F398386375C.taxon	diagnosis	Diagnosis. Mandible with three teeth, decreasing in size from apex to base; occipital carina crenulate. Pronotum not visible dorsally; mesometapectal sulcus (sensu Vilhelmsen et al. 2010 b) crenulate. Fore wing venation most complete within Megalyridae; pterostigma reduced; C, M + Cu, Sc + R, A and M almost fully pigmented; R 1 extending beyond marginal cell; marginal cell closed by Rs in straight line; Rs present between r-rs and Rs + M, closing first submarginal cell; M + Cu aligned with Rs + M; medial cell rectangular or trapezoidal, located under Rs + M and closed by 1 m-cu and basal segments of Cu and Cu 1. Metasoma elongate, with long to very long ovipositor (OL / BL ~ 0.40).	en	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
E48F5B8FE6EB51A2AB715E51476381B2.taxon	diagnosis	Diagnosis. Head hypognathous; compound eye without setation, without posterior carina; subantennal groove without dorsal carina; mandible with three teeth; median sulcus on vertex present; occipital carina present, curving towards mandible. Median mesoscutal sulcus crenulate; parapsidal lines absent; axillae not connected at inner angles, separated by two triangular foveae; propodeum areolate-rugose. Fore wing hyaline; with C, Sc + R, A, cu-a, Rs + M, M, Rs and R 1 tubular; pterostigma large; M not reaching apical margin; Rs only present as a stub between Rs + M and r-rs; Rs apically evenly arched towards stigma; costal, marginal and radial + first cubital cells closed. Metasoma elongate, fusiform, longer than mesosoma.	en	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
06A075176FB150B7AFC036ADEF9DF13E.taxon	diagnosis	Diagnosis. Head hypognathous; compound eye without setation, without posterior carina; subantennal groove without dorsal carina; mandible with three teeth; occipital carina present, curving towards mandible. Median mesoscutal sulcus crenulate; parapsidal lines absent; axillae not connected at inner angles, separated by two triangular foveae; propodeum carinate. Protibia with stout apical spines; pro-, meso- and metatibiae shorter than combined length of tarsomeres. Fore wing hyaline, with C, Sc + R, A, Rs + M, cu-a, Rs, M and R 1 tubular; pterostigma reduced; M not reaching apical margin; Rs apically strongly angled, arched towards stigma; costal, submarginal, marginal and radial + first cubital cells closed. Hind wing with Rs short, barely projecting beyond R 1. Metasoma compact, shorter than mesosoma.	en	Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo, Perrichot, Vincent (2024): Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities. Arthropod Systematics & amp; Phylogeny 82: 151-181, DOI: http://dx.doi.org/10.3897/asp.82.e111148, URL: http://dx.doi.org/10.3897/asp.82.e111148
