taxonID	type	description	language	source
76261A0AD054FFDDFF2E12D8FAD1FD28.taxon	diagnosis	Diagnosis. Members of this genus are distinguished by having a strong median longitudinal carina on propodeum; vein cu-a (nervellus) in hind wing straight or sinuate near posterior end; T 1 moderately to strongly narrowed apically, without a median depression; T 2 much wider than long, with strongly diverging lateral margins and much shorter than T 3; hypopygium large; ovipositor valves long and hairy and tapering gradually (Mason 1981).	en	Zargar, Mohammad, Gupta, Ankita, Talebi, Asghar, Farahani, Samira (2019): A review of the Iranian species of genus Iconella Mason (Hymenoptera: Braconidae: Microgastrinae) with description of two new species. Zootaxa 4586 (3): 491-504, DOI: 10.11646/zootaxa.4586.3.6
76261A0AD054FFDDFF2E107BFCB5FA5D.taxon	materials_examined	Material examined: Southwestern Iran, Khuzestan province: Ramhormoz, Gharabad, (30 ° 59 ’ 37.73 ” N, 49 ° 46 ′ 50.63 ” E, 126 m), 4. v. 2016; Malaise trap in olive orchard; leg. M. Zargar, 2 ♀ (TMUC); Khoramshahr, Sheneh, (30 ° 25 ’ 32.24 ” N, 48 ° 11 ′ 20.83 ” E, 2 m), 2. iii. 2017; Malaise trap in palm orchard; leg. M. Zargar, 1 ♀ (ICAR ̄ NBAIR); Ramhormoz, Gharabad, (30 ° 59 ’ 37.73 ” N, 49 ° 46 ′ 50.63 ” E, 126 m), 6. vii. 2017; Malaise trap in olive orchard; leg. M. Zargar, 1 ♀ (ICAR ̄ NBAIR).	en	Zargar, Mohammad, Gupta, Ankita, Talebi, Asghar, Farahani, Samira (2019): A review of the Iranian species of genus Iconella Mason (Hymenoptera: Braconidae: Microgastrinae) with description of two new species. Zootaxa 4586 (3): 491-504, DOI: 10.11646/zootaxa.4586.3.6
76261A0AD054FFDDFF2E107BFCB5FA5D.taxon	diagnosis	Diagnosis. Width of face 1.5 × as long as height (Fig. 3 A); ocelli large, POL: MOD: OOL – 5: 3: 5 (Fig. 3 B); penultimate flagellomere 1.3 × as long as wide; Mesoscutum densely evenly punctate and setose, scutellum smooth and shiny (Fig. 3 C); propodeum with strong medial longitudinal carina with fine short radiating lines, smooth and shiny centrally on both the sides of median carina (Fig. 3 C); 1 - R 1 1.4 × as long as pterostigma, 1 - R 1 3.1 × as long as distance from end of 1 - R 1 to tip of radial cell, pterostigma 3 × as long as wide (Fig. 3 D); TI subparallel sided at base, converging from middle to apex, 2.1 × longer than basal width, basal width 1.8 × as long as apical width (Fig. 3 E); ovipositor sheath 1.2 × as long as hind tibia (Fig. 3 F); body black.	en	Zargar, Mohammad, Gupta, Ankita, Talebi, Asghar, Farahani, Samira (2019): A review of the Iranian species of genus Iconella Mason (Hymenoptera: Braconidae: Microgastrinae) with description of two new species. Zootaxa 4586 (3): 491-504, DOI: 10.11646/zootaxa.4586.3.6
76261A0AD054FFDDFF2E107BFCB5FA5D.taxon	distribution	Distribution. Hungary, Israel, Jordan, Romania, Turkey (Yu et al., 2016), Iran (Khuzestan province), new record.	en	Zargar, Mohammad, Gupta, Ankita, Talebi, Asghar, Farahani, Samira (2019): A review of the Iranian species of genus Iconella Mason (Hymenoptera: Braconidae: Microgastrinae) with description of two new species. Zootaxa 4586 (3): 491-504, DOI: 10.11646/zootaxa.4586.3.6
76261A0AD054FFDDFF2E107BFCB5FA5D.taxon	discussion	Comments. One character was found to be different from the original description of I. meruloides; Nixon (1965) indicated that the ovipositor is as long as hind tibia, however in the specimens examined in the present study, the ovipositor is 1.2 × as long as hind tibia (Fig. 3 F).	en	Zargar, Mohammad, Gupta, Ankita, Talebi, Asghar, Farahani, Samira (2019): A review of the Iranian species of genus Iconella Mason (Hymenoptera: Braconidae: Microgastrinae) with description of two new species. Zootaxa 4586 (3): 491-504, DOI: 10.11646/zootaxa.4586.3.6
76261A0AD054FFD9FF2E1564FE2EFA77.taxon	materials_examined	Type material. Holotype: ♀ (mounted on card); Southwestern Iran: Khuzestan, Baghmalek, Ghale tol, (31 ° 37 ’ 49.70 ” N, 49 ° 52 ′ 53.35 ” E, 880 m), 4. iv. 2017; Malaise trap in citrus orchard; leg. M. Zargar (ICAR ̄ NBAIR). Paratypes: 2 ♀ Khuzestan, Baghmalek, Ghale tol, (31 ° 37 ’ 49.70 ” N, 49 ° 52 ′ 53.35 ″ E, 880 m), 21. iv. 2017, 5. v. 2017; Malaise trap in citrus orchard; leg. M. Zargar (TMUC ICAR ̄ NBAIR).	en	Zargar, Mohammad, Gupta, Ankita, Talebi, Asghar, Farahani, Samira (2019): A review of the Iranian species of genus Iconella Mason (Hymenoptera: Braconidae: Microgastrinae) with description of two new species. Zootaxa 4586 (3): 491-504, DOI: 10.11646/zootaxa.4586.3.6
76261A0AD054FFD9FF2E1564FE2EFA77.taxon	diagnosis	Diagnosis. Propodeum with strong medial longitudinal carina with fine radiating lines / carinae on lateral sides at an angle, running from apex towards base on the lateral sides with granular sculpture in between carinae; median area on both sides shallowly and sparsely punctate with setae; apical one third with fine pitted appearance (Fig. 4 C); penultimate flagellomere 1.6 × as long as wide; TI parallel sided at base, converging from middle to apex, TI 2.1 × longer than basal width, basal width 2.4 × as long as apical width (Fig. 4 E); 1 - R 1 1.3 × as long as pterostigma, 3.2 × as long as distance from end of 1 - R 1 to tip of radial cell (Fig. 4 D).	en	Zargar, Mohammad, Gupta, Ankita, Talebi, Asghar, Farahani, Samira (2019): A review of the Iranian species of genus Iconella Mason (Hymenoptera: Braconidae: Microgastrinae) with description of two new species. Zootaxa 4586 (3): 491-504, DOI: 10.11646/zootaxa.4586.3.6
76261A0AD054FFD9FF2E1564FE2EFA77.taxon	description	Description. Female. Body length 2.9 mm, fore wing length 3.1 mm (Fig. 4 F). Head (Figs 4 A – 4 B). Confluent punctations on face and shallowly rugose on vertex and temple region; width of head 1.2 × as long as height; width of face 1.4 × as long as height; ocelli large, POL: MOD: OOL – 6: 4: 5; eyes large, 1.5 × as high as wide; antenna as long as body; flagellomeres finely setose, apical flagellomere 1.6 × longer than penultimate, penultimate flagellomere 1.6 × as long as wide. Mesosoma (Fig. 4 C). Mesoscutum densely and evenly punctate, setose and dull; notauli indistinct; scutellum smooth and shiny, scutellar sulcus crenulate; postscutellum smooth and shiny; prepectal carina absent; precoxal sulcus absent; propodeum with strong medial longitudinal carina with fine radiating lines on lateral sides at an angle from apex towards base; basal margin densely punctate, remaining propodeum not smooth, shallowly punctate. Wings (Fig. 4 D). Areolet absent, vein r arising little after from middle of pterostigma; pterostigma 3 × as long as wide; vein 1 - R 1 1.3 × as long as pterostigma, 3.2 × as long as distance from end of 1 - R 1 to tip of radial cell, 3.2 × as long as r; vein 1 - cu 1 1.1 × as long as 2 - cu 1; width of discoidal cell 1.4 × as long as height. Legs (Fig. 4 F). Metafemur length 3.3 × as long as median width; hind basitarus 1.9 × longer than inner hind tibial spur, 3.1 × longer than telotarsus. Metasoma (Figs 4 E – F). TI parallel sided at base, converging from middle to apex; TI 2.1 × longer than basal width, basal width 2.4 × as long as apical width, dull punctations at apex; TII strongly constricted at base and triangular, its apical width 4 × longer than median length; tergites posterior to TI finely pitted and with scattered long setae; ovipositor sheath 1.1 × as long as hind tibia and setose; hypopygium strongly sclerotized, sharp edged and not truncated apically, with medium sized scattered setae. Colouration (Fig. 4 F). Body black; antenna black; first and second segment of labial palp and maxillary palp yellowish brown, remaining testaceous; tegula yellowish brown; profemur yellowish brown with blackish infuscation on anterior half; mesofemur brown, paler towards apex; metafemur black; protibia and mesotibia brown; metatibia brown on anterior half and black on posterior half; protarsus and mesotarsus yellowish brown; metatarsus dark brown, except for yellowish area on anterior half of first tarsomere; at least some fore wing veins with thin brown margins and interior of veins yellow to light brown, pterostigma brown. Male. Unknown Host. Unknown. Biology. Unknown.	en	Zargar, Mohammad, Gupta, Ankita, Talebi, Asghar, Farahani, Samira (2019): A review of the Iranian species of genus Iconella Mason (Hymenoptera: Braconidae: Microgastrinae) with description of two new species. Zootaxa 4586 (3): 491-504, DOI: 10.11646/zootaxa.4586.3.6
76261A0AD054FFD9FF2E1564FE2EFA77.taxon	distribution	Distribution. Khuzestan province (Iran).	en	Zargar, Mohammad, Gupta, Ankita, Talebi, Asghar, Farahani, Samira (2019): A review of the Iranian species of genus Iconella Mason (Hymenoptera: Braconidae: Microgastrinae) with description of two new species. Zootaxa 4586 (3): 491-504, DOI: 10.11646/zootaxa.4586.3.6
76261A0AD054FFD9FF2E1564FE2EFA77.taxon	etymology	Etymology. The name refers to mountainous region of Mongasht that it is considered to be one of the longest semi branches of the Zagross mountain ranges, which is also the distribution range of the species.	en	Zargar, Mohammad, Gupta, Ankita, Talebi, Asghar, Farahani, Samira (2019): A review of the Iranian species of genus Iconella Mason (Hymenoptera: Braconidae: Microgastrinae) with description of two new species. Zootaxa 4586 (3): 491-504, DOI: 10.11646/zootaxa.4586.3.6
76261A0AD054FFD9FF2E1564FE2EFA77.taxon	discussion	Comments. This new species is most similar to Iconella meruloides (Nixon 1965), from which it can be separated by (1) propodeum dull and not smooth in I. mongashtensis (Fig. 4 E) vs smooth and shiny in I. meruloides (Fig. 3 C) (2) first tergite basal width 2.4 × as long as apical width (Fig. 4 E) vs 1.8 × in I. meruloides (Fig. 3 E) (3) distance between hind ocelli visibly longer than distance between hind ocellus and marginal eye (Fig. 4 B) vs equal in I. meruloides (Fig. 3 B).	en	Zargar, Mohammad, Gupta, Ankita, Talebi, Asghar, Farahani, Samira (2019): A review of the Iranian species of genus Iconella Mason (Hymenoptera: Braconidae: Microgastrinae) with description of two new species. Zootaxa 4586 (3): 491-504, DOI: 10.11646/zootaxa.4586.3.6
76261A0AD050FFD7FF2E1682FEF8FEBC.taxon	materials_examined	Material examined: Southwestern Iran: Khuzestan, Lali, Taraz, (32 ° 20 ’ 49.70 ” N, 49 ° 05 ′ 11.31 ” E, 390 m), 21. v. 2016; Malaise trap in citrus orchard; leg. M. Zargar, 1 ♀ (ICAR ̄ NBAIR); Khuzestan, Lali, Taraz, (32 ° 20 ’ 49.70 ” N, 49 ° 05 ′ 11.31 ” E, 390 m), 21. iii. 2017, 22. v. 2017; Malaise trap in citrus orchard; leg. M. Zargar, 2 ♀ (TMUC).	en	Zargar, Mohammad, Gupta, Ankita, Talebi, Asghar, Farahani, Samira (2019): A review of the Iranian species of genus Iconella Mason (Hymenoptera: Braconidae: Microgastrinae) with description of two new species. Zootaxa 4586 (3): 491-504, DOI: 10.11646/zootaxa.4586.3.6
76261A0AD050FFD7FF2E1682FEF8FEBC.taxon	diagnosis	Diagnosis. Width of face 1.8 × as long as height (Fig. 5 A); ocelli large, POL: MOD: OOL − 7: 3: 5 (Fig. 5 B); penultimate flagellomere 1.4 × as long as wide; mesoscutum densely evenly punctate and setose, scutellum smooth and setose (Fig. 5 C); propodeum with strong medial longitudinal carina with fine radiating lines on lateral sides at an angle from apex toward base; basal margin densely punctate, remaining propodeum not smooth, punctuate (Fig. 5 C); vein 1 - R 1 1.3 × as long as pterostigma, 2.7 × as long as r, 2.4 × as long as distance from end of 1 - R 1 to tip of radial cell; pterostigma 4 × as long as wide (Fig. 5 D); TI parallel sided at base, converging from middle to apex; TI 2 × longer than basal width, basal width 2.0 × as long as apical width; in middle smooth, punctate on lateral sides and rugose at apical near to TII. TII strongly constricted at base and triangular, its apical width 2.7 × longer than median length (Fig. 5 E); ovipositor sheath 2.6 × as long as hind tibia (Fig. 5 F); body black.	en	Zargar, Mohammad, Gupta, Ankita, Talebi, Asghar, Farahani, Samira (2019): A review of the Iranian species of genus Iconella Mason (Hymenoptera: Braconidae: Microgastrinae) with description of two new species. Zootaxa 4586 (3): 491-504, DOI: 10.11646/zootaxa.4586.3.6
76261A0AD050FFD7FF2E1682FEF8FEBC.taxon	distribution	Distribution. Cyprus, Greece, Greece-main, Iran, Israel, Moldova, Spain, Spain-main, Turkey, Turkmenistan (Yu et al., 2016). This is a new record for Khuzestan province of Iran. Notes. In the original description (Wilkinson 1937) the ovipositor is about as long as combined length of trochanter, trochanellus, femur, tibia, and tarsus of hind leg, but in the key by Papp (1982) the ovipositor sheath is 2 × as long as hind tibia. However, in the specimens examined in the present study ovipositor sheath is 2.5 - 2.8 × as long as hind tibia (Fig. 5 F).	en	Zargar, Mohammad, Gupta, Ankita, Talebi, Asghar, Farahani, Samira (2019): A review of the Iranian species of genus Iconella Mason (Hymenoptera: Braconidae: Microgastrinae) with description of two new species. Zootaxa 4586 (3): 491-504, DOI: 10.11646/zootaxa.4586.3.6
76261A0AD05EFFD5FF2E12CFFE50FF1C.taxon	materials_examined	Type material: Holotype: ♀ (mounted on card); Southwestern Iran: Khuzestan, Baghmalek, Dobagh, (31 ° 31 ’ 16.14 ” N, 49 ° 52 ′ 53.00 ” E, 688 m), 4. iv. 2017; Malaise trap in citrus orchard; leg. M. Zargar; ICAR ̄ NBAIR. Paratypes: 4 ♀, 3 #, Khuzestan, Baghmalek, Dobagh, (31 ° 31 ’ 16.14 ” N, 49 ° 52 ′ 53.00 ” E, 688 m), 4. iv. 2017, 5. v. 2017; Malaise trap in citrus orchard; leg. M. Zargar; 1 ♀, Khuzestan, Baghmalek, Shang, (31 ° 31 ’ 46.00 ” N, 49 ° 53 ′ 14.61 ” E, 716 m), 22. v. 2017; Malaise trap in citrus orchard; leg. M. Zargar TMUC, ICAR ̄ NBAIR.	en	Zargar, Mohammad, Gupta, Ankita, Talebi, Asghar, Farahani, Samira (2019): A review of the Iranian species of genus Iconella Mason (Hymenoptera: Braconidae: Microgastrinae) with description of two new species. Zootaxa 4586 (3): 491-504, DOI: 10.11646/zootaxa.4586.3.6
76261A0AD05EFFD5FF2E12CFFE50FF1C.taxon	diagnosis	Diagnosis. Width of face 1.7 × as long as height (Fig. 6 A); apical flagellomere 1.5 × longer than penultimate flagellomere; Propodeum with strong medial longitudinal (Fig. 6 C); 1 - R 1 1.1 × as long as pterostigma, 1 - R 1 4 × as long as distance from end of 1 - R 1 to tip of radial cell (Fig. 6 D); TI parallel sided at base, converging from middle to apex; TI 2.0 × longer than basal width, basal width 2.5 × as long as apical width (Fig. 6 E).	en	Zargar, Mohammad, Gupta, Ankita, Talebi, Asghar, Farahani, Samira (2019): A review of the Iranian species of genus Iconella Mason (Hymenoptera: Braconidae: Microgastrinae) with description of two new species. Zootaxa 4586 (3): 491-504, DOI: 10.11646/zootaxa.4586.3.6
76261A0AD05EFFD5FF2E12CFFE50FF1C.taxon	description	Description. Female. Body length 2.5 mm, fore wing length 3.0 mm (Fig. 6 F). Head (Figs 6 A – B). Shallowly rugose with sparse setation, setae denser near the eye margin; width of head 1.15 × as long as height, width of face 1.7 × as long as height; ocelli small, POL: MOD: OOL – 6.4: 3: 5; eyes large, 1.6 × as high as wide. Antenna as long as body; flagellomeres finely setose, apical flagellomere 1.5 × longer than penultimate; penultimate flagellomere length as long as wide. Mesosoma (Fig. 6 C). Mesoscutum densely and evenly punctate, pubescent; notauli indistinct; scutellum smooth, scutellar sulcus crenulate; postscutellum rugose; prepectal carina absent; precoxal sulcus absent; propodeum with strong medial longitudinal carina with fine short radiating lines, shallowly rugose on the both sides of median carina; laterally with fine transverse striations. Wings (Fig. 6 D). Areolet absent, vein r arising little after from middle of pterostigma. 1 - R 1 1.1 × as long as pterostigma, 1 - R 1 4.0 × as long as distance from end of 1 - R 1 to tip of radial cell, pterostigma 3.0 × as long as wide. 1 - R 1 3.0 × as long as r, r 2.1 × as long as 2 - SR. 1 - cu 1 as long as 2 - cu 1; width of discoidal cell 1.1 × as long as height. Legs (Fig. 6 F). Metafemur length 3.5 × as long as median width. Hind basitarus 1.7 × longer than inner hind tibial spur; hind basitarus 3.7 × longer than telotarsus. Metasoma (Figs 6 E – F). TI parallel sided at base, converging from middle to apex; TI 2 × longer than basal width, basal width 2.5 × as long as apical width and finely pitted; TII strongly constricted at base and triangular, its apical width 3.1 × longer than median length; tergites posterior to TI with rough finely pitted surface and scattered long setae; ovipositor sheath 1.3 × as long as hind tibia and setose; hypopygium strongly sclerotized, sharp edged and not truncated apically, scattered with medium sized setae. Colouration (Fig. 6 F). Body black; antenna black; palpi pale yellow; tegula yellowish brown and black at apex; profemur black on anterior half and yellowish brown on posterior half; mesofemur black except yellowish apex; metafemur black; protibia and mesotibia yellowish brown; metatibia yellowish brown on anterior half and black on posterior half; protarsus and mesotarsus yellow; metatarsus dark brown, except yellowish on anterior half of first tarsomere; fore wing with at least some veins with thin brown margins and interior of veins yellow to light brown, pterostigma pale yellow. Male. Unknown Host. Unknown. Biology. Unknown.	en	Zargar, Mohammad, Gupta, Ankita, Talebi, Asghar, Farahani, Samira (2019): A review of the Iranian species of genus Iconella Mason (Hymenoptera: Braconidae: Microgastrinae) with description of two new species. Zootaxa 4586 (3): 491-504, DOI: 10.11646/zootaxa.4586.3.6
76261A0AD05EFFD5FF2E12CFFE50FF1C.taxon	distribution	Distribution. Khuzestan province (Iran).	en	Zargar, Mohammad, Gupta, Ankita, Talebi, Asghar, Farahani, Samira (2019): A review of the Iranian species of genus Iconella Mason (Hymenoptera: Braconidae: Microgastrinae) with description of two new species. Zootaxa 4586 (3): 491-504, DOI: 10.11646/zootaxa.4586.3.6
76261A0AD05EFFD5FF2E12CFFE50FF1C.taxon	etymology	Etymology. This name is chosen considering the similarity between this new species with Iconella isus (Nixon, 1965).	en	Zargar, Mohammad, Gupta, Ankita, Talebi, Asghar, Farahani, Samira (2019): A review of the Iranian species of genus Iconella Mason (Hymenoptera: Braconidae: Microgastrinae) with description of two new species. Zootaxa 4586 (3): 491-504, DOI: 10.11646/zootaxa.4586.3.6
76261A0AD05EFFD5FF2E12CFFE50FF1C.taxon	discussion	Comments. This new species comes near to Iconella isus (Nixon, 1965) from which it can be separated by the sculpture of mesoscutum that is densely and evenly punctate and dull in I. similus (Fig. 6 C) vs strongly shiny and very superficially punctate in I. isus; 1 - R 1 of fore wing 4 × as long as distance between its end to tip of the radial cell in I. similus (Fig. 6 D) vs 2.5 × in I. isus, and first tergite basal width 2.5 × as long as apical width in I. similus (Fig. 6 E) vs 2.0 × in I. isus.	en	Zargar, Mohammad, Gupta, Ankita, Talebi, Asghar, Farahani, Samira (2019): A review of the Iranian species of genus Iconella Mason (Hymenoptera: Braconidae: Microgastrinae) with description of two new species. Zootaxa 4586 (3): 491-504, DOI: 10.11646/zootaxa.4586.3.6
76261A0AD05CFFD5FF2E1221FEC1FCDD.taxon	materials_examined	Material examined: Southwestern Iran: Khuzestan, Ramhormoz, Gharabad, (30 ° 59 ’ 37.73 ” N, 49 ° 46 ′ 50.63 ” E, 126 m), 4. v. 2016; Malaise trap in olive orchard; leg. M. Zargar, 1 ♀ (ICAR ̄ NBAIR).	en	Zargar, Mohammad, Gupta, Ankita, Talebi, Asghar, Farahani, Samira (2019): A review of the Iranian species of genus Iconella Mason (Hymenoptera: Braconidae: Microgastrinae) with description of two new species. Zootaxa 4586 (3): 491-504, DOI: 10.11646/zootaxa.4586.3.6
76261A0AD05CFFD5FF2E1221FEC1FCDD.taxon	diagnosis	Diagnosis. Width of head 0.9 × as long as height, width of face 1.5 × as long as height (Fig. 7 A); POL: MOD: OOL – 4.1: 2: 3.5 (Fig. 7 B); eyes large, 1.7 × as high as wide; penultimate flagellomere length as long as wide; pterostigma 3.5 × as long as wide; 1 - R 1 as long as pterostigma, 1.16 × as long as distance from end of 1 - R 1 to tip of radial cell (Fig. 7 D); mesoscutum and scutellum densely evenly punctate and setose (Fig. 7 C); Propodeum smooth and shiny with fine transverse striations on lateral sides; TI parallel sided at base, converging from middle to apex, its median length 2 × longer than basal width, basal width 1.9 × as long as apical width (Fig. 7 E); TII strongly constricted at base and triangular, its apical width 4.5 × longer than median length (Fig. 7 E); ovipositor sheath 1.2 × as long as hind tibia; body black (Fig. 7 F).	en	Zargar, Mohammad, Gupta, Ankita, Talebi, Asghar, Farahani, Samira (2019): A review of the Iranian species of genus Iconella Mason (Hymenoptera: Braconidae: Microgastrinae) with description of two new species. Zootaxa 4586 (3): 491-504, DOI: 10.11646/zootaxa.4586.3.6
76261A0AD05CFFD5FF2E1221FEC1FCDD.taxon	distribution	Distribution. Azerbaijan, Cape Verde Islands, Iran, Israel (Yu et al., 2016). This is new record for Khuzestan province of Iran.	en	Zargar, Mohammad, Gupta, Ankita, Talebi, Asghar, Farahani, Samira (2019): A review of the Iranian species of genus Iconella Mason (Hymenoptera: Braconidae: Microgastrinae) with description of two new species. Zootaxa 4586 (3): 491-504, DOI: 10.11646/zootaxa.4586.3.6
