taxonID	type	description	language	source
75255D75FF8CCF2FFE85E9EDFED8F92F.taxon	materials_examined	TYPE SPECIES. — Murex messorius Sowerby II, 1841, Western Atlantic (by original designation).	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF8CCF2EFEB6E84EFF55FC3C.taxon	description	(Figs 3 A-G; 5 A; 14 A)	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF8CCF2EFEB6E84EFF55FC3C.taxon	materials_examined	TYPE MATERIAL. — Holotype (lv) MNHN-IM- 2009 - 14495 BOLD ID BOMGA 023 - 15, GenBank accession number KP 697994 for COI gene KC 860506 (Fig. 3 A, B) and 7 paratypes MNHN: 1 (lv) IM- 2009 - 14501 BOLD ID BOMGA 022 - 15, GenBank accession number KP 697995 for COI gene KC 860506; 3 (lv) & 3 (dd) IM- 2000 - 30001; 1 paratype coll. RH (all from type locality). TYPE LOCALITY. — Southern Madagascar, Southwest of Pointe Barrow, 25 ° 03 ’ S, 44 ° 00 ’ E, 132 - 153 m [ATIMO VATAE: stn CP 3589]. MATERIAL EXAMINED. — Known only from the type material.	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF8CCF2EFEB6E84EFF55FC3C.taxon	distribution	DISTRIBUTION. — Southern Madagascar, taken alive at 132 - 153 m.	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF8CCF2EFEB6E84EFF55FC3C.taxon	etymology	ETYMOLOGY. — Latin, rectaspira, rectus (straight) and spira (spire). This species is named for its flattened spire outline.	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF8CCF2EFEB6E84EFF55FC3C.taxon	description	DESCRIPTION Shell small for the genus, up to 41.2 mm in height at maturity (holotype MNHN). Height / width ratio 2.9 (holotype MNHN). Slender, lanceolate, weakly spinose, lightly built. Shoulder ramp strongly sloping, weakly concave. Beige or light tan with brown or dark brown spiral cords. Aperture glossy white. Spire high with 2.10 protoconch whorls and teleoconch with up to six weakly convex and weakly shouldered whorls. Suture slightly impressed. Protoconch moderately broad, weakly conical, whorls rounded, last whorl minutely punctate, glossy, maximum width of last whorl 1000 µm and first whorl 700 µm (holotype). Terminal lip heavy, narrow, erect and curved. Axial sculpture of teleoconch whorls consisting of high, rounded varices and ribs, each varix with a single, short, acute, narrow, narrowly open primary shoulder spine (P 1). First whorl with 11 or 12 ribs, second with 9 - 12, third with 8 - 10, fourth to last whorl with three narrow, rounded, high varices and two intervariceal, high, narrow ribs. Spiral sculpture of narrow, smooth, primary, secondary and tertiary cords. First and second teleoconch whorls with IP, P 1, P 2, (S 2), P 3, P 4, third whorl with (adis), IP, P 1, (s 1), P 2, s 2, P 3, (s 3), P 4, fourth and fifth with adis, IP, abis, P 1, S 1, P 2, s 2, P 3, s 3, P 4, last whorl with adis, IP, abis, P 1, s 1, P 2, s 2, P 3, s 3, P 4, s 4, P, s 5, P 6, s 6, P 7, s 7, ADP, ads, MP, ms, ABP, abs, EAB 1, eabs 1, EAB 2, eabs 2, EAB 3, eabs 3, followed by several shallower spiral cords. Only P 1 ending as short, acute, narrow variceal shoulder spine. Aperture moderately small, roundly-ovate. Columellar lip narrow, flaring, smooth. Rim erect, a small portion adherent at adapical extremity. Low parietal tooth at extreme adapical extremity. Anal notch shallow, broad. Outer lip erect, smooth within or with seven very weak, elongate denticles: ID, D 1 - D 6. Siphonal canal long, narrow, narrowly open. Operculum dark brown, roundly-ovate with subapical nucleus. Attached with about four growth lines and broad, callused rim. Radula (Fig. 14 A) typical for the genus, with a rachidian bearing a long, triangular central cusp, a small lateral denticle and a broad, triangular, medium sized lateral cusp, marginal area smooth. Lateral tooth sickle shaped, broad.	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF8CCF2EFEB6E84EFF55FC3C.taxon	discussion	REMARKS Vokesimurex rectaspira n. sp. differs from the superficially similar V. gallinago fernandesi Houart, 1990 from Mozambique (Fig. 3 H, I) by having a more regularly-shaped protoconch, with a narrower, more strongly curved terminal lip with slightly more numerous whorls, 2.10 - 2.15 as opposed to 1.75 - 2.0 in V. gallinago fernandesi and with a narrower first whorl (width 700 µm vs 900 - 1100 µm). Vokesimurex rectaspira n. sp. is also more slender with a higher spire consisting of more numerous teleoconch whorls for a similar shell height and also differs by having a flatter as opposed to a more rounded or occasionally weakly concave spire outline in V. gallinago fernandesi, and by having narrower primary spiral cords on the siphonal canal. Another Vokesimurex species occurring in northern Madagascar, namely V. dolichourus (Ponder & Vokes, 1988) (Fig. 3 J-L) differs also in many ways in its protoconch and teleoconch characters.	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF8DCF2EFE86EB4BFC50F839.taxon	description	(Figs 3 M, N; 4 A-E; 5 B)	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF8DCF2EFE86EB4BFC50F839.taxon	materials_examined	TYPE MATERIAL. — Holotype (lv), MNHN-IM- 2007 - 36971 BOLD ID BOMGA 020 - 15, GenBank accession number KP 697992 for COI gene KC 860506 and 5 paratypes (lv) MNHN: 3 MNHN-IM- 2000 - 30002, 1 MNHN-IM- 2007 - 36950, 1 MNHN-IM- 2007 - 36958 BOLD ID BOMGA 021 - 15, GenBank accession number KP 697993 for COI gene KC 860506; 1 paratype coll. RH (all from type locality). TYPE LOCALITY. — Madagascar, between Nosy Be and Banc du Leven, 12 ° 39 ’ S, 48 ° 22 ’ E, 70 - 74 m [MIRIKY: stn CP 3206]. MATERIAL EXAMINED. — Known only from the type material.	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF8DCF2EFE86EB4BFC50F839.taxon	distribution	DISTRIBUTION. — Northern Madagascar, taken alive at 70 - 74 m.	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF8DCF2EFE86EB4BFC50F839.taxon	etymology	ETYMOLOGY. — Latin, aliquantulus, little, small. This species is named for its small size relative to the other Murex s. s. species.	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF8DCF2EFE86EB4BFC50F839.taxon	description	DESCRIPTION Shell small for the genus, up to 33.1 mm in height. Height / width ratio 2.3 (holotype). Slender, lanceolate, narrowlyovate, nodose, lightly built. Shoulder ramp strongly sloping, weakly convex. White with occasionally a very light brown darker band on P 1 and P 2, more obvious on P 2 and on axial varices, and between P 6 and P 7. Aperture white. Spire high with 1.75 - 2.00 protoconch whorls and teleoconch up to 6.25, convex, narrow, weakly shouldered, nodose whorls. Suture impressed. Protoconch small, whorls rounded, smooth, occasionally with a narrow keel abapically, width 700 µm. Terminal lip heavy, broad, erect, weakly curved. Axial sculpture of teleoconch whorls consisting of low, narrow, rounded ribs and high, rounded, weakly spinose varices. First whorl with 11 or 12 ribs, second with 11 - 13, third with 10 - 12, early varices on fourth whorl with two or three intervariceal ribs. Last whorl with three high, narrow, rounded varices and two broad intervariceal ribs, occasionally with a third narrow one. Spiral sculpture of moderately high, narrow, weakly nodose primary, secondary and tertiary cords. First whorl with P 1 - P 4, P 4 partially covered by the following teleoconch whorl, second and third whorls with IP, P 1 - P 4, fourth with adis, IP, P 1, P 2, P 2, P 3, s 3, P 4, fifth with adis, IP, abis, P 1, P 2, P 3, s 3, P 4, sixth and seventh with t, adis, (t), IP, abis, P 1, s 1, P 2, s 2, P 3, s 3, P 4, s 4, P 5, s 5, P 6, s 6, P 7, s 7, ADP, ads, MP, ms, ADP, ads. Other spiral cords decreasing in strength and height abapically. Intersection of axial varices and P 1 giving rise to a short, narrow, narrowly open, acute shoulder spinelet from fourth or fifth to last teleoconch whorl. Aperture narrow, roundly-ovate. Columellar lip narrow, strongly flaring with two or three elongate, weak knobs abapically. Rim erect, a small portion adherent adapically. Low parietal tooth at extreme adapical extremity. Anal notch shallow, broad. Outer lip strongly erect, weakly crenulated, bearing eight weak, elongate, narrow denticles within: ID, D 1 - D 7. Siphonal canal narrow, straight, weakly dorsally recurved, weakly open, occasionally with a single, acute, short spine adapically. Operculum and radula not examined.	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF8DCF2EFE86EB4BFC50F839.taxon	discussion	REMARKS Vokesimurex aliquantulus n. sp. differs from V. gallinago fernandesi (Fig. 3 H, I) in being more lightly-built, in having a narrower and more acute spire, a narrower siphonal canal and a comparatively narrower aperture. The protoconch is also much smaller, 700 µm wide × 750 µm high as opposed to 1000 - 1200 µm wide × 1000 - 1100 µm high. It also has a very narrow spiral keel abapically, not observed in V. gallinago fernandesi.	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF8DCF2EFE86EB4BFC50F839.taxon	description	Vokesimurex aliquantulus n. sp. differs from Vokesimurex rectaspira n. sp. (Fig. 3 A-G) in the same characters and in having a spire that is weakly incurved and not almost straight as in Vokesimurex aliquantulus n. sp. It differs from Vokesimurex gallinago gallinago (Sowerby III, 1903) and V. dentifer (Watson, 1883), not present in the western Indian Ocean, in also having a smaller protoconch (700 µm wide vs 1100 - 1200), a narrower aperture and a narrower siphonal canal. From V. gallinago gallinago it also differs in having almost spineless varices, rather than the more spinose, occasionally weakly webbed varices of V. gallinago gallinago. Vokesimurex rectirostris (Sowerby II, 1841) (Fig. 4 F, G) and V. sobrinus (A. Adams, 1863) (Fig. 4 H, I) both have a rounded protoconch with a strong keel abapically but in V. aliquantulus n. sp. the keel is more delicate, narrower and situated more abapically, and the protoconch is also relatively smaller (700 vs 1100 - 1400 µm wide). Both these species, restricted to the region from Taiwan to southern Japan, are also more spinose.	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF8BCF28FE94EFA8FE0DFB9C.taxon	materials_examined	TYPE SPECIES. — Murex latifolius Bellardi, 1872, Middle Miocene, Italy (by subsequent designation; Vokes 1964: 14).	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF8BCF28FE94EFA8FE0DFB9C.taxon	discussion	REMARKS The genus Timbellus was reinstated by Merle et al. (2011) for the species generally classified in Pterynotus s. s. Swainson, 1833 but usually with a less scabrous shell morphology and with three major axial varices “ appearing early during the ontogeny ”. This distinction from the Pterynotus s. s. group was noted by Barco et al. (2010), with Pterynotus fulgens Houart, 1988 in their DNA analysis even falling outside the Muricinae Rafinesque, 1815. The classification used here follows Barco et al. (2010) pending availability of more material for further DNA research and the publication of additional results. On the other hand, the classification of Merle et al. (2011) mixes the species with a trivaricate axial sculpture starting already on the first teleoconch whorl, such as Timbellus latifolius, the type species, and some Recent species like T. fulgens (Houart, 1988). In Timbellus goniodes n. sp., here described, and a few other species such as T. bednalli (Brazier, 1878), T. phyllopterus (Lamarck, 1822) and Pterynotus alatus (Röding, 1798), the type species of Pterynotus, the trivaricate sculpture appears only on the third or fourth teleoconch whorl. Their generic placement awaits confirmation by genetic analysis.	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF8BCF2AFEA3EBECFD1CFD82.taxon	description	(Figs 4 J-O; 7 A)	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF8BCF2AFEA3EBECFD1CFD82.taxon	materials_examined	TYPE MATERIAL. — Holotype (lv) MNHN-IM- 2007 - 36940 BOLD ID BOMGA 026 - 15, GenBank accession number KP 697989 for COI gene KC 860506 and 5 paratypes MNHN, 1 paratype coll. RH (as listed below). TYPE LOCALITY. — Madagascar, between Nosy Be and Banc du Leven, 12 ° 40 ’ S, 48 ° 12 ’ E, 492 - 524 m [MIRIKY: stn CP 3184]. MATERIAL EXAMINED. — Northwest Madagascar. MIRIKY: stn CP 3184, between Nosy Be and Banc du Leven, 12 ° 40 ’ S, 48 ° 12 ’ E, 492 - 524 m, 2 lv, 1 dd (holotype MNHN-IM- 2007 - 36940 and 1 paratype (lv) MNHN-IM- 2007 - 38031 BO LD ID BOMGA 025 - 15, GenBank accession number KP 697990 for COI gene KC 86050, 1 paratype (dd) MNHN-IM 2000 - 30003. — Stn CP 3188, between Nosy Be and Banc du Leven, 12 ° 31 ’ S, 48 ° 22 ’ E, 298 - 301 m, 2 lv (1 paratype MNHN-IM- 2000 - 30004; 1 paratype Coll. RH). — Stn DW 3197, West of Cap d’Ambre, 12 ° 07 ’ S, 48 ° 58 ’ E, 362 - 431 m, 3 dd. — Stn CP 3209, between Nosy Be and Banc du Leven, 12 ° 43 ’ S, 48 ° 14 ’ E, 291 - 353 m, (2 lv paratypes MNHN-IM- 2000 - 30005). — Stn CP 3223, between Nosy Be and Banc du Leven, 12 ° 46 ’ S, 48 ° 11 ’ E, 430 - 488 m, 1 dd.	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF8BCF2AFEA3EBECFD1CFD82.taxon	distribution	DISTRIBUTION. — North-western Madagascar, taken alive at 301 - 492 m, shells at 431 - 492 m.	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF8BCF2AFEA3EBECFD1CFD82.taxon	etymology	ETYMOLOGY. — Latin, pannuceus, wrinkled. This species is named for the wrinkled surface of the shell.	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF8BCF2AFEA3EBECFD1CFD82.taxon	description	DESCRIPTION Shell medium sized for the genus, up to 22.6 mm in height at maturity (paratype MNHN). Height / width ratio 1.4 (holotype). Slender, narrowly-ovate, lightly built. Shoulder ramp strongly sloping, weakly concave. Ivory-white, occasionally with a light tan spiral band between P 4 and P 5 and occasionally on shoulder ramp. Aperture white. Spire high with 1.5 protoconch whorls and teleoconch with up to five narrow, very weakly shouldered whorls. Suture weakly adpressed. Protoconch small, whorls rounded, smooth, glossy, maximum width 900 µm. Terminal lip narrow, low, weakly curved. Axial sculpture of teleoconch whorls consisting of very narrow, webbed varices starting from the first whorl. First to last teleoconch whorls with three varices. No intervariceal sculpture except narrow, low, growth striae. Spiral sculpture of moderately high, narrow, smooth primary and secondary cords. First whorl with very low P 1, second and third with P 1 and P 2, s 1 occasionally starting, fourth with P 1, (s 1), P 2, s 2, P 3, last whorl with P 1, (s 1), P 2, s 2, P 3, s 3, P 4, (s 4), P 5, P 6, ADP, MP, ABP, (ads). Aperture moderately large, roundly-ovate. Columellar lip narrow, more broadly expanded adapically, weakly flaring, rim partially weakly erect, adherent at adapical extremity. Anal notch shallow, broad. Outer lip smooth, occasionally with weak or strong denticles within: ID broad, low, D 1 - D 6 narrower; aperture occasionally smooth or with only a few denticles visible, others obsolete. Siphonal canal moderately long, broad, weakly bent dorsally and abaxially, narrowly open, with variceal webbing extending to 50 - 60 % of its length. Operculum and radula not examined.	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF8BCF2AFEA3EBECFD1CFD82.taxon	discussion	REMARKS Eight Indo-West Pacific species are included in Timbellus, all with a similar shell morphology consisting of a protoconch of 1.5 to 1.75 smooth whorls, smooth or almost smooth teleoconch whorls and thin, webbed variceal wings. These are T. vespertilio (Kuroda, 1959), T. flemingi (Beu, 1967), T. fulgens (Houart, 1988), T. levii (Houart, 1988), T. marshalli (Houart, 1989), T. stenostoma (Houart, 1991), T. crauropterus (Houart, 1991) and T. rubidus (Houart, 2001). Four of these species (T. flemingi, T. marshalli, T. crauropterus and T. rubidus) do not need to be compared with T. pannuceus n. sp. due to important differences in shell characters. The remaining four differ from T. pannuceus n. sp. as follows. Timbellus vespertilio (Fig. 6 A, B) differs in having more strongly shouldered teleoconch whorls with a relatively broader shoulder ramp and a more obvious and broader shoulder cord (P 1). It also has fewer obvious spiral cords on the teleoconch whorls and on the abapertural face of the variceal wings. Timbellus vespertilio is known only from southern Japan, the East China Sea and Taiwan.	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF8BCF2AFEA3EBECFD1CFD82.taxon	description	Timbellus fulgens (Fig. 4 P, Q) was described from New Caledonia and was identified from South Africa by Houart (1991). Similar specimens are now also recorded from Madagascar. However it differs from T. pannuceus n. sp. in having a much smoother shell and smoother variceal wings with much lesser obvious primary cords and obsolete secondary cords. Timbellus levii, a species currently only known from New Caledonia and Tonga (Houart & Héros 2008) has a larger shell, reaching 36 mm in height, with smooth teleoconch whorls, almost smooth, broader variceal wings and with a much narrower and higher aperture. Timbellus stenostoma from New Caledonia has a much smaller and smoother shell, reaching 15 mm in height, with a much smaller aperture and a longer siphonal canal.	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF89CF2AFEAFEDEAFB85FA7F.taxon	description	(Figs 6 C-G; 7 B)	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF89CF2AFEAFEDEAFB85FA7F.taxon	materials_examined	TYPE MATERIAL. — Holotype (dd) MNHN-IM- 2000 - 30006, 3 paratypes MNHN, 1 paratype coll. RH (as listed below). TYPE LOCALITY. — South Madagascar, South of Cap Sainte-Marie, 26 ° 14 ’ S, 45 ° 09 ’ E, 284 - 286 m [ATIMO VATAE: stn CP 3615]. MATERIAL EXAMINED. — South Madagascar. ATIMO VATAE: stn DW 3552, South of Faux-Cap, 26 ° 07 ’ S, 45 ° 39 ’ E, 264 - 280 m, 2 dd (1 paratype MNHN-IM- 2000 - 30007, 1 Coll. RH). — Stn DW 3553, South of Faux-Cap, 26 ° 08 ’ S, 45 ° 39 ’ E, 280 - 333 m, 1 dd fragment (paratype MNHN-IM- 2000 30008). — Stn CP 3615, South of Cap Sainte-Marie, 26 ° 14 ’ S, 45 ° 09 ’ E, 284 - 286 m, 2 dd (holotype MNHN-IM- 2000 - 30006, 1 paratype MNHN-IM- 2000 - 30009).	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF89CF2AFEAFEDEAFB85FA7F.taxon	distribution	DISTRIBUTION. — Southern Madagascar, 280 - 284 m (shells only).	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF89CF2AFEAFEDEAFB85FA7F.taxon	etymology	ETYMOLOGY. — Greek, goniodes, angular. This species is named for the angular shape of the teleoconch whorls.	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF89CF2AFEAFEDEAFB85FA7F.taxon	description	DESCRIPTION Shell medium sized for the genus, up to 23.7 mm in height at maturity. Height / width ratio 1.6 - 2.0. Slender, narrowlyovate, spinose, lightly built. Shoulder strongly sloping, weakly concave or straight. Milky white or pinkish white. Aperture white. Spire high, acute, with 1.5 protoconch whorls and teleoconch up to six weakly convex, angular, weakly shouldered whorls. Suture impressed. Protoconch small, whorls rounded, smooth, glossy, width 600 - 650 µm. Terminal lip delicate, thin, weakly raised, weakly curved. Axial sculpture of teleoconch whorls consisting of high, narrow, lamellose varices, each with a long, narrow, open shoulder spine. First whorl with six or seven varices, second with five or six, third with four. From fourth to last teleoconch whorl, three narrow, high, lamellose, webbed varices. Spiral sculpture of low and high, narrow, rounded primary cords. First whorl with P 1 - P 3 of approximately the same strength, second with P 1 - P 3, P 2 becoming broader and higher. Third and fourth whorls with shallow, almost obsolete P 1, P 2 and P 3 similar in strength and height, P 3 partially covered by following whorl, fifth whorl with very shallow P 1 and broad, high, rounded P 2 and P 3, last whorl with weak P 1, more obvious on lamellose, webbed varices, P 2 and P 3 higher and broader, P 4 - P 6 weak, decreasing in strength abapically, broader on webbed apertural varix. P 1 ending as a short, narrow, adapically bent shoulder spine. P 2 - P 6 ending as a shorter, backward recurved, narrowly open spine. Aperture ovate, angular. Columellar lip narrow, flaring, smooth or with two weak knobs abapically, rim partially erect, adherent at adapical extremity. Anal notch moderately deep, broad. Outer lip strongly erect, smooth with six weak or strong denticles within: ID, D 1 - D 5. ID very low, broad, D 1 - D 5 variable in strength and height. Siphonal canal moderately short, broad, weakly abaxially bent and dorsally recurved at tip, narrowly open, with variceal webbing extending to 80 - 90 % of its length. Operculum and radula unknown.	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF89CF2AFEAFEDEAFB85FA7F.taxon	discussion	REMARKS Timbellus goniodes n. sp. differs in many ways from the other Indo-West Pacific Timbellus species. From T. fulgens (Fig. 4 P, Q) living in the same geographical area, T. goniodes n. sp. differs in having a narrower shell with a higher spire, a smaller protoconch (600 - 650 µm wide vs 1000 µm), a narrower subsutural ramp, strong and relatively broader spiral cords with higher P 2 and P 3, a more angular aperture and more strongly adapically bent shoulder spines, and chiefly in having four to seven varices on the three adapical teleoconch whorls as opposed to three in T. fulgens.	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF89CF2AFEAFEDEAFB85FA7F.taxon	description	From T. crauroptera (Fig. 6 H) from New Caledonia, a species that also has upward bent shoulder spines and a more angular aperture, T. goniodes n. sp. differs in having somewhat narrower shoulder spines, more strongly obvious spiral cords, less convex teleoconch whorls without intervariceal varices, a higher spire and variceal webbing extending to almost 80 % of the length of the siphonal canal rather than extending to 10 or 20 % of its length in T. crauroptera with the additional presence of ADP and MP. Moreover T. crauroptera also has three variceal wings starting at the first teleoconch whorl.	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF89CF24FC02EA8CFAACFB63.taxon	materials_examined	TYPE SPECIES. — Poirieria (Paziella) vaubanensis Houart, 1986, New Caledonia (by original designation).	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF89CF24FC02EA8CFAACFB63.taxon	discussion	REMARKS Although molecular phylogenetic research is improving our understanding of the classification of supraspecific taxa in the Muricidae (Barco et al. 2010), based on shell morphology, Bouchetia and Flexopteron Shuto, 1969 (see below) were treated by Merle et al. (2011) as subgenera of Paziella Jousseaume, 1880. Pending the result of DNA analyses we feel it premature to adopt this new classification based only on shell characters. We will therefore continue to treat Bouchetia as a full genus as used by Houart & Héros (2008: 450). FIG. 5. — Distributions of: A, Vokesimurex rectaspira n. sp.; B, Vokesimurex aliquantulus n. sp. Symbols: ★, type locality.	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF87CF20FED7EB8CFCA7FA1F.taxon	description	(Figs 2 A; 6 I-M; 9 A)	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF87CF20FED7EB8CFCA7FA1F.taxon	materials_examined	TYPE MATERIAL. — Holotype (lv) MNHN-IM- 2007 - 36941 and 3 paratypes MNHN, 1 paratype coll. RH (listed as below). TYPE LOCALITY. — Madagascar, between Nosy Be and Banc du Leven, 12 ° 30 ’ S, 48 ° 18 ’ E, 346 - 376 m [MIRIKY: stn CP 3189]. MATERIAL EXAMINED. — Mozambique Channel. BENTHEDI: stn DS 72, Mayotte, NNE du Récif Nord, 12 ° 32 ’ S, 45 ° 02 ’ E, 300 - 350 m, 2 dd. Northwest of Madagascar. MIRIKY: stn CP 3183, between Nosy Be and Banc du Leven, 12 ° 38 ’ S, 48 ° 14 ’ E, 420 - 436 m, 1 lv (paratype MNHN-IM- 2007 - 36979 BOLD ID BOMGA 024 - 15, GenBank accession number KP 697988 for COI gene KC 860506). — Stn CP 3184, between Nosy Be and Banc du Leven, 12 ° 40 ’ S, 48 ° 12 ’ E, 492 - 524 m, 1 lv (paratype Coll. RH). — Stn CP 3189, between Nosy Be and Banc du Leven, 12 ° 30 ’ S, 48 ° 18 ’ E, 346 - 376 m, 1 lv (holotype MNHN-IM- 2007 - 36941). — Stn DW 3197, West of Cap d’Ambre, 12 ° 07 ’ S, 48 ° 58 ’ E, 362 - 431 m, 1 dd. — Stn DW 3217, between Nosy Be and Banc du Leven, 12 ° 33 ’ S, 47 ° 56 ’ E, 391 - 438 m, 1 dd (paratype MNHN-IM- 2000 - 30010). — Stn DW 3226, between Nosy Be and Banc du Leven, 12 ° 51 ’ S, 48 ° 12 ’ E, 341 - 345 m, 1 dd (paratype MNHN-IM- 2000 - 30011). South Madagascar. ATIMO VATAE: stn DW 3524, sector of Manantenina, 24 ° 23 ’ S, 47 ° 32 ’ E, 307 - 319 m, 1 dd, juv. — Stn DW 3534, sector of Sainte Luce, 24 ° 43 ’ S, 47 ° 32 ’ E, 296 - 307 m, 1 lv, juv. — Stn DW 3552, South of Faux-Cap, 26 ° 07 ’ S, 45 ° 39 ’ E, 264 - 280 m, 2 dd, juv. — R. V. Vauban, 1972 - 73, 22 ° 17 ’ S, 43 ° 03 ’ E, 600 - 605 m, 1 dd. DISTRIBUTION. — Madagascar and Mayotte, live in 307 - 492 m, shells in 280 - 600 m. ETYMOLOGY. — This species is named after Anders Warén, in recognition of his help in extracting, mounting and imaging numerous radulae for the senior author. He also has participated in many expeditions organized by MNHN, including ATIMO VATAE. DESCRIPTION Shell medium sized for the genus, with up to 18.5 mm in height at maturity (paratype MNHN). Height / width ratio 1.4 - 1.7. Broad, spinose, lightly built. Subsutural band narrow, weakly sloping, weakly concave. White or very light tan. Spire high with 1.60 - 1.75 protoconch whorls and teleoconch up to six broad, weakly convex, strongly shouldered, spinose whorls. Suture impressed, partially obscured by small axial lamellae of following whorl. Protoconch small, whorls rounded, smooth. Maximum width 700 - 800 µm. Terminal lip delicate, thin, very weakly curved. FIG. 7 — Distributions of: A, Timbellus pannuceus n. sp.; B, Timbellus goniodes n. sp. Symbols: ★, type locality; ■, other occurences. Axial sculpture of teleoconch whorls consisting of low, narrow lamellae, more strongly developed at intersection of axial lamellae and spiral cords, producing short or long, narrow, spine-like projections. Shoulder spine longest. Axial sculpture of first whorl with 8 - 10 lamellae, second with 11 - 13, third with 13 or 14, fourth 12 - 15, fifth 15 and last whorl with 10 - 13. Spiral sculpture of moderately high, rounded, narrow, smooth primary cords. First to penultimate teleoconch whorl with P 1 - P 3, P 3 partially covered by following whorl. Last whorl with P 1 - P 6, ADP, (MP). Secondary cords s 1 or s 2 rarely present. Primary cords decreasing in strength abapically. P 1 broadest with long shoulder spine. Other spiral sculpture consisting of numerous lirae. Aperture moderately small, broad, rounded. Columellar lip broad, flaring, smooth. Rim partially strongly erect, adherent at adapical extremity. Anal notch shallow, broad. Outer lip weakly erect, smooth with 4 - 6 weak, elongate, narrow denticles within: D 1 (occasionally split), D 2, D 3, D 4, D 5 (D 4 and D 5 occasionally fused), rarely with low, weak, narrow ID. Siphonal canal moderately long or long, up to 33 % of total shell height, narrow, weakly dorsally recurved, open. REMARKS Bouchetia vaubanensis (Houart, 1986) (Figs 2 B; 6 N, O) described from New Caledonia and also known from Fiji (Houart & Héros 2008: 452) resembles the new species, but Bouchetia wareni n. sp. is distinct in having a comparatively higher spire, a longer siphonal canal, a broader aperture and also in having a different spiral sculpture morphology. Bouchetia wareni n. sp. possesses six spiral cords on the convex part of the teleoconch whorl (P 1 - P 6) and two cords on the siphonal canal (ADP and MP), while B. vaubanensis clearly has five spiral cords on the convex part of the last teleoconch whorl (P 1 - P 5) and a single one on the siphonal canal (ADP) (Fig. 2 A, B). Merle et al. (2011: 174) illustrated the canal cord as P 6 but this cord is clearly separated from the whorl, as in Paziella, for example Paziella pazi (Crosse, 1869) as shown by Merle et al. (2011: 162). This cord is here considered as a true siphonal canal spiral cord, here ADP. Merle et al. (2011: 174, text fig. 60 D) wrongly illustrated the radula of Pazinotus falcatiformis as Bouchetia vaubanensis. The radula of Bouchetia is peculiar in having a broad, long central cusp and long lateral cusps, but lacking lateral denticles (Fig. 14 G).	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF87CF20FED7EB8CFCA7FA1F.taxon	description	a species known only from the western Pacific. This juvenile specimen (8.9 mm high, with only 3.75 teleoconch whorls: BENTHEDI, stn DS 72) differs from T. polycyma in having a broader teleoconch whorl, a broader aperture with a wide, flared, columellar lip and a narrower, abapically tapered siphonal canal. Typical specimens from T. polycyma have been examined since then from Japan and from Fiji (Houart & Héros 2008).	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF83CF20FE8CE96CFBFDF8DF.taxon	materials_examined	TYPE SPECIES. — Flexopteron philippinensis Shuto, 1969, Late Miocene, Philippines (by original designation).	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF83CF20FE8CE96CFBFDF8DF.taxon	discussion	REMARKS There are currently two known Recent species of Flexopteron: F. primanova (Houart, 1985) from Madagascar and F. poppei (Houart, 1993) from the Philippines. Two other Recent species were included in Flexopteron by Merle et al. (2011): Poirieria (Paziella) tanaoa Houart & Tröndlé, 2008 from the Marquesas and Paziella (Flexopteron) sp. from the Philippines. Paziella (Flexopteron) sp. (Merle et al. 2011: pl. 141, figs 12 a, b) is a species of Coralliophilinae, probably in the genus Lamellatiaxis Habe & Kosuge, 1970. We do not accept the classification of Poirieria (Paziella) tanaoa in Flexopteron by Merle et al. (2011). The radula characters are not known for P. tanaoa but its shell is quite different from that of species of Flexopteron, being more elongate with fewer spiral cords, a relatively smaller aperture and a longer siphonal canal, shell characters that are closer to Paziella than to Flexopteron. In fact, P. tanaoa strongly resembles the specimen illustrated by Merle et al. (2011: pl. 144, figs 4 a, b) as Crassimurex (Eopaziella) capita (Philippi, 1844) from the Late Oligocene of Europe. Currently we do not see any reason to remove P. tanaoa from Paziella. The shell morphology of Flexopteron akainakares n. sp. is close to F. primanova Houart, 1985 but the species has different radula characters and is here only tentatively assigned to this genus. We have now used Flexopteron as a full genus since its radular morphology (Fig. 14 D-F) is different from Poirieria (Marshall & Houart, 1995).	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF83CF3DFC58E82EFCAAFB5C.taxon	description	(Figs 2 D, E; 6 P, Q; 8 A-E; 9 B; 14 B, C)	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF83CF3DFC58E82EFCAAFB5C.taxon	materials_examined	TYPE MATERIAL. — Holotype (dd) MNHN-IM- 2000 - 30012 and 4 paratypes MNHN, 1 paratype coll. RH (as listed below). FIG. 11. — Distributions of: A, Murexsul mananteninaensis n. sp.; B, Typhinellus laminatus n. sp. Symbols: ★, type locality; ■, other occurences. TYPE LOCALITY. — South Madagascar, Manantenina sector, 24 ° 23 ’ S, 47 ° 32 ’ E, 307 - 319 m [ATIMO VATAE: stn DW 3524]. MATERIAL EXAMINED. — South Madagascar. ATIMO VATAE: stn DW 3515, between Lokaro and Sainte Luce, 24 ° 53 ’ S, 47 ° 28 ’ E, 184 - 203 m, 1 lv, 2 dd (paratypes MNHN-IM- 2000 - 30013). — Stn DW 3524, Manantenina sector, 24 ° 23 ’ S, 47 ° 32 ’ E, 307 - 319 m, 1 dd (holotype MNHN-IM- 2000 - 30012), 3 dd, juv. — Stn DW 3528, Manantenina sector, 24 ° 24 ’ S, 47 ° 33 ’ E, 424 - 438 m, 13 dd (1 paratype Coll. RH). — Stn DW 3529, Manantenina sector, 24 ° 24 ’ S, 47 ° 33 ’ E, 402 - 407 m, 1 dd (paratype MNHN-IM- 2000 - 30014). — Stn DW 3534, Sainte Luce sector, 24 ° 43 ’ S, 47 ° 32 ’ E, 296 - 307 m, 2 dd, juv.	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF83CF3DFC58E82EFCAAFB5C.taxon	distribution	DISTRIBUTION. — South Madagascar, taken alive at 184 - 203 m, shells at 203 - 424 m.	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF83CF3DFC58E82EFCAAFB5C.taxon	etymology	ETYMOLOGY. — Greek, akaina, spine and akares, short, small. This species is named for the small and short spines on the axial lamellae.	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF83CF3DFC58E82EFCAAFB5C.taxon	description	DESCRIPTION Shell medium sized for the subgenus, up to 12.2 mm in height at maturity. Height / width ratio 1.5 - 1.7. Broadly ovate, weakly spinose, lightly built. Subsutural ramp narrow, weakly sloping on spire whorls, broader, strongly sloping on last teleoconch whorl, weakly concave. Shell entirely white. Spire high with 1.5 protoconch whorls and up to 4.15 broad, convex, angular, strongly shouldered, weakly spinose whorls. Suture impressed. Protoconch large, broad, whorls rounded, smooth, maximum width 900 - 1000 µm. Terminal lip weakly raised, narrow, prosocline. Axial sculpture of teleoconch whorls consisting of low, narrow, weakly webbed lamellae, more strongly developed at shoulder, producing short, narrow, broadly open spines at intersection of lamellae and primary spiral cords. First whorl with 6 or 7 lamellae, second with 7 or 8, third 7 - 9, fourth 6 - 9, last (apertural) with 6 - 7. Spiral sculpture of high, rounded, narrow, weakly squamous, primary cords, low, narrow secondary cords and numerous, narrow, low threads. First teleoconch whorl with P 1 and P 2, broadly separated by a large gap, second whorl with P 1 and P 2, starting weak s 1 and one or two threads, third whorl with P 1 and P 2, weak s 1 and narrow, weak threads of various strength, fourth and last whorls with P 1, s 1, P 2, P 3, P 4, (s 4), P 5, (s 5), P 6, s 6, adp and several threads of different magnitude. Aperture small, ovate. Columellar lip broad, flaring, smooth or occasionally with one or two weak knobs abapically, rim partially erect, adherent at adapical extremity. Occasionally with weak, very low parietal tooth at adapical extremity. Anal notch shallow, broad. Outer lip weakly erect, smooth with four weak or moderately strong, broad denticles within: D 1, D 2, D 3, D 4 - D 5 fused. Siphonal canal short, broad, weakly dorsally recurved, broadly open. Operculum ovate, light brown with apical nucleus. Radula (Fig. 14 B, C) muricine with broad rachidian bearing long, narrow, central cusp; short, triangular lateral denticles, almost half the size of central cusp; broad, long, triangular lateral cusps. Marginal area short, occasionally with small marginal denticle, almost indistinct when present. Marginal cusp short, outwardly bent. Lateral teeth sickle shaped, long, with broad base.	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF83CF3DFC58E82EFCAAFB5C.taxon	discussion	REMARKS Flexopteron akainakares n. sp. differs in many ways from F. poppei and does not need to be compared here.	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF83CF3DFC58E82EFCAAFB5C.taxon	description	Flexopteron primanova (Fig. 8 F-I) was described from northern Madagascar but F. akainakares n. sp. is not sympatric, living more to the South. Flexopteron akainakares n. sp. differs in having a relatively smaller shell, a higher and larger protoconch (900 - 1000 wide and 900 µm high vs 850 - 900 wide and 700 - 800 µm high) with rounded first whorl vs flat in F. primanova, markedly shorter shoulder spines on the last teleoconch whorl in adult specimens, a shorter siphonal canal, a different morphology of the primary spiral cords of the teleoconch whorls, and a last teleoconch whorl with P 1, s 1, P 2, P 3, P 4, (s 4), P 5, (s 5) and P 6, with a broad gap between P 1 and P 2 as opposed to P 1, P 2, (s 2), P 3, P 4, P 5, (P 6). The radula of F. primanova (Fig. 14 D-F) is also different in having a very long, narrow central cusp, narrow lateral denticles approximately half the size of the central cusp, long, narrow, lateral cusps even longer than the central cusp, a smooth marginal area and acute, moderately long, marginal cusps. The operculum is similar, being ovate with an apical nucleus.	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF9ECF3CFEB5EBABFD6FFDE2.taxon	materials_examined	TYPE SPECIES. — Murex octogonus Quoy & Gaimard, 1832, New Zealand (by original designation). Murexsul mananteninaensis n. sp. Figs 2 C; 8 J-N; 11 A TYPE MATERIAL. — Holotype (lv) MNHN-IM- 2000 - 30015 and 3 paratypes MNHN, 1 paratype coll. RH (as listed below). TYPE LOCALITY. — South Madagascar, Manantenina sector, 24 ° 23 ’ S, 47 ° 32 ’ E, 307 - 319 m [ATIMO VATAE: stn DW 3524]. MATERIAL EXAMINED. — South Madagascar. ATIMO VATAE: stn DW 3523, Manantenina sector, 24 ° 23 ’ S, 47 ° 31 ’ E, 200 - 220 m, 2 dd (1 paratype MNHN-IM- 2000 - 30016, 1 paratype Coll. RH). — Stn DW 3524, Manantenina sector, 24 ° 23 ’ S, 47 ° 32 ’ E, 307 - 319 m, 1 lv (holotype MNHN-IM- 2000 - 30015), 1 dd (paratype MNHN- IM- 2000 - 30017). — Stn DW 3532, North of Sainte Luce, 24 ° 39.4 ’ S, 47 ° 31.7 ’ E, 86 - 87 m, 1 lv (paratype MNHN-IM- 2000 - 30018). — Stn CP 3615, South of Cap Sainte-Marie, 26 ° 14 ’ S, 45 ° 09 ’ E, 284 - 286 m, 1 dd.	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF9ECF3CFEB5EBABFD6FFDE2.taxon	distribution	DISTRIBUTION. — Southern Madagascar, taken alive at 87 - 307 m, shells at 220 - 307 m.	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF9ECF3CFEB5EBABFD6FFDE2.taxon	etymology	ETYMOLOGY. — Named after the type locality.	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF9ECF3CFEB5EBABFD6FFDE2.taxon	description	DESCRIPTION Shell very small for the genus, up to 5.3 mm in height at maturity. Height / width ratio 1.9 (holotype). Slender, lanceolate, biconical, narrow, very weakly spinose, lightly built. Substural ramp narrow, weakly sloping, very slightly convex. Entirely white, translucent white or orange (paratype MNHN). Spire high with 1.5 protoconch whorls and teleoconch of three broadly-convex, elongate, weakly shouldered whorls. Suture impressed. Protoconch large, broad, whorls rounded, smooth, maximum width 600 µm. Terminal lip delicate, narrow, weakly erect, very weakly curved. Axial sculpture of teleoconch whorls consisting of very low, weak, quite indistinguishable, narrow, lamellose ribs and low, weak, narrow varices. First whorl with eight or nine ribs, second and third with ribs and varices of same strength as each other. Apertural varix high, broad, rounded with five very short, blunt, broadly open spinelets, extending from P 1 - P 5. Spiral sculpture of low, rounded, narrow primary and secondary cords. First teleoconch whorl starting with P 3 only, starting P 1 and P 2 at the end of the whorl, second whorl with P 1 - P 3, occasionally with P 4 almost entirely covered by following whorl, last whorl with P 1, s 1, P 2, (s 2), P 3, P 4, s 4, P 5, s 5, P 6, ADP, (MP). P 1 - P 5 almost similar in strength and height, P 6 narrower. Aperture moderately large, ovate. Columellar lip narrow, strongly flaring, smooth, rim partially erect, adherent at adapical extremity. Anal notch shallow, broad. Outer lip erect, smooth within. Siphonal canal moderately long, narrow, weakly dorsally recurved, open. Operculum and radula not examined.	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF9ECF3CFEB5EBABFD6FFDE2.taxon	discussion	REMARKS	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF9ECF3CFEB5EBABFD6FFDE2.taxon	description	Murexsul queenslandicus (Fig. 8 O, P) is broader and heavier with broader spiral cords, no secondary spiral cords and a broader, higher apertural varix. Murexsul leonardi differs markedly and does not need to be compared here. Murexsul charcoti (Fig. 8 Q, R) is larger, reaching a height of 7.2 mm, and broader with broader, more obvious axial varices, broader spiral cords and small, blunt spinelets on axial varices and on the siphonal canal. Murexsul micra (Fig. 8 S) is smaller, reaching a height of 3.9 mm, with a lower spire, broader spiral cords, more obvious axial varices with short, blunt spinelets and a smaller aperture. Curiously, Murexsul mananteninaensis n. sp. also resembles species of Enatimene Iredale, 1929, a genus confined to eastern Australia, comprising three species. The shell morphology of E. lanceolatus Houart, 2004 is particularly close to Murexsul mananteninaensis n. sp., but E. lanceolatus is twice as large for the same number of teleoconch whorls, has a rounded protoconch and a first teleoconch whorl with P 1 - P 3. The genus Enatimene is currently classified in Trophoninae s. l. Cossmann, 1903 and its radula characters are distinct from those of the Muricopsinae Radwin & D’Attilio, 1971.	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF9FCF3CFF6DED09FE8AFD2E.taxon	materials_examined	TYPE SPECIES. — Typhis sowerbyi Broderip, 1833 (= Murex labiatus Cristofori & Jan, 1832), Mediterranean, eastern Atlantic (by original designation).	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF9FCF3CFEBDEC4AFAF2FB3C.taxon	description	(Figs 10 A-F; 11 B)	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF9FCF3CFEBDEC4AFAF2FB3C.taxon	materials_examined	TYPE MATERIAL. — Holotype (lv) MNHN-IM- 2000 - 30019 and 1 (dd) paratype MNHN-IM- 2000 - 30020. TYPE LOCALITY. — Madagascar, off Baie Mahajamba, 14 ° 53 ’ S, 46 ° 56 ’ E, 90 - 257 m [MIRIKY: stn DW 3245]. MATERIAL EXAMINED. — Only known from the type material.	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF9FCF3CFEBDEC4AFAF2FB3C.taxon	distribution	DISTRIBUTION. — North-western Madagascar, taken alive at 90 - 257 m.	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF9FCF3CFEBDEC4AFAF2FB3C.taxon	etymology	ETYMOLOGY. — Latin, lamina, thin blade. This species is named for its thin axial lamellae.	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF9FCF3CFEBDEC4AFAF2FB3C.taxon	description	DESCRIPTION Holotype medium sized for the genus, 16.3 mm in height. Height / width ratio 1.7, narrowly-ovate, smooth, delicate. Subsutural ramp moderately broad, weakly sloping, weakly concave. Light ochre or light tan, brown on subsutural ramp on third, fourth and last teleoconch whorls, light brown spiral band at periphery. A single brown spot at adapical edge of aperture and three on edge of outer apertural lip. Three additional weak, light brown spots on left part of ventral side of siphonal canal. Spire high with 1.75 protoconch whorls and five weakly convex, angulate, strongly shouldered teleoconch whorls. Suture of whorls impressed. Protoconch small, whorls rounded, slightly eroded. Terminal lip unknown (eroded). Axial sculpture of teleoconch whorls consisting of four high, thin, sharp lamellate varices, each with a weakly open, short, inward curved shoulder spine. Apertural varix broad, flangelike, dorsally reflected, with two broadly open, short spinelets at outer edge. Variceal flange extending to tip of siphonal canal. Shoulder spine broad, triangular, very weakly dorsally curved, connected to last teleoconch whorl by a broad, high, lamellate partition. Spiral sculpture faint, consisting of P 1 with the anal tube, P 2 (shoulder spine) and probably P 3, P 4, P 5 and P 6 with very faint secondary cords. Spiral sculpture more obvious on adapertural part of apertural varix. P 1 with long, tapering, ventrally sealed anal tube, forming an angle of approximately 85 - 90 ° with axis of shell, only apertural tube hollow and functional, older tubes broken off and closed. P 2 with a weakly open, curved shoulder spine, P 4 and P 5 corresponding to broad, open spinelets at outer apertural margin. Very thin intritacalx consisting of simple axial striae present. Aperture small, ovate, forming a continuous peristome. Columellar lip narrow, outer lip smooth within. Siphonal canal long, broad, straight, ventrally sealed. Left side of canal weakly overlapping right side. Operculum and radula unknown.	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF9FCF3CFEBDEC4AFAF2FB3C.taxon	discussion	REMARKS Typhinellus laminatus n. sp. differs from Typhinellus bicolor Bozzetti, 2007 (Fig. 10 G, H) in having a smooth shell with the exception of a few, rather indistinct spiral cords, whereas T. bicolor has a microsculpture of irregular pits, visible even in the worn holotype. Typhinellus laminatus n. sp. also has a more fragile shell, sharper axial varices and a slightly smaller aperture, 17.1 % of shell height and 19 % of shell width as opposed to 17.6 % of shell height and 22.6 % of shell width in T. bicolor (measurements made on the holotype of T. laminatus n. sp. and specimens of T. bicolor from ATIMO VATAE, stn BP 11).	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF9FCF3CFEBDEC4AFAF2FB3C.taxon	description	Typhinellus laminatus n. sp. differs from T. androyensis Bozzetti, 2007 (Fig. 10 K) in having a more fragile shell with narrower teleoconch whorls, much thinner and sharper axial varices and a flatter siphonal canal. Typhinellus laminatus n. sp. differs from T. amoenus Houart, 1994 (Fig. 10 I, J) in the same characters as it differs from T. androyensis but also in having a smooth shell as opposed to the strongly spirally sculptured shell in T. amoenus.	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF9FCF3EFC43EA4CFEF5FAAD.taxon	description	(Figs 10 L-P; 13 A)	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF9FCF3EFC43EA4CFEF5FAAD.taxon	materials_examined	TYPE MATERIAL. — Holotype (lv) MNHN-IM- 2009 - 14446 and 5 paratypes MNHN; 1 paratype coll. RH (as listed below). TYPE LOCALITY. — Madagascar, sector of Fort-Dauphin, 25 ° 15.0 ’ S, 47 ° 14.5 ’ E, 97 - 98 m. [ATIMO VATAE: stn BS 09]. MATERIAL EXAMINED. — South Madagascar. ATIMO VATAE: stn BS 09, Rocher de l’Albatros, 25 ° 29 ’ S, 44 ° 57 ’ E, 11 - 13 m, 1 lv (holotype MNHN-IM- 2009 - 14446). — Stn BP 22, West sector of Lavanono, 25 ° 23 ’ S, 44 ° 52 ’ E, 20 - 22 m, 1 lv (paratype MNHN- IM- 2009 - 22522 BOLD ID BOMGA 027 - 15, GenBank accession number KP 697991 for COI gene KC 860506). — Stn BP 42, West sector of Lavanono, 25 ° 23 - 24 ’ S, 44 ° 51.1 ’ E, 18 - 21 m 1 dd (paratype MNHN-IM- 2000 - 30021). — Stn CP 3511, sector of Fort-Dauphin, 25 ° 15 ’ S, 47 ° 15 ’ E, 97 - 98 m, 1 dd (paratype MNHN- IM- 2000 - 30022). — Stn CP 3560, East of Faux-Cap, 25 ° 37 ’ S, 46 ° 10 ’ E, 102 - 106 m, 2 dd (2 paratypes MNHN-IM- 2000 - 30023). — Stn CP 3622, Southeast of Faux-Cap, 25 ° 43 ’ S, 45 ° 59 ’ E, 84 - 88 m, 1 dd (paratype Coll. RH). — No data, South Madagascar, sector of Fort Dauphin, 1 dd (MNHN).	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF9FCF3EFC43EA4CFEF5FAAD.taxon	distribution	DISTRIBUTION. — South Madagascar, taken alive at 13 - 20 m, empty shells at 21 - 102 m.	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF9FCF3EFC43EA4CFEF5FAAD.taxon	etymology	ETYMOLOGY. — Latin, constrictus, constricted. This species is named for the peculiar form of the teleoconch whorl.	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF9FCF3EFC43EA4CFEF5FAAD.taxon	description	DESCRIPTION Shell large for the genus, up to 20.7 mm in height at maturity (holotype), although one incomplete paratype (MNHN) would have been larger. Height / width ratio 1.73 (holotype). Slender, narrowly ovate with expanded shoulder, lightly built. Subsutural ramp broad, almost horizontal, weakly convex. Tan or light brown, subsutural ramp darker. Abapertural side of apertural flange and ventral part of siphonal canal beige or light tan. Three or four axial, narrow, brown bands on partition. Aperture white. Spire high with paucispiral, weakly shouldered protoconch (observed in one specimen but partly broken). Teleoconch up to five angulate, strongly shouldered whorls. Abapical part of whorls constricted. Suture impressed. Axial sculpture of teleoconch whorls consisting of four, low, thin, sharp lamellate varices, each with a weakly open, inward curved shoulder spine. Apertural varix thin, broad, constricted medially, broader at abapical extremity, extending from shoulder spine almost to tip of siphonal canal. Apertural shoulder spine adapically bent, long, narrowly triangular, straight, connected to last teleoconch whorl by a broad, thin, lamellate partition. Partition with two or three backward bent, short, flattened, broad spinelets. Spiral sculpture very faint, consisting of P 1 with the anal tube, P 2 (shoulder), P 3, P 4 and P 5 only discernable on variceal flange and only very weakly between axial varices, corresponding to the broad, weakly curved abapertural spinelets at outer apertural margin. P 1 with long, rounded, ventrally sealed anal tube, forming an angle of approximately 85 - 90 ° with axis of shell. Only apertural tube hollow and functional, older tubes broken off and closed. Two narrow spiral cords between abapical part of aperture and variceal flange, occasionally obvious in some shells. Other sculpture of few, weak and very thin, low, broad scales, only visible on last teleoconch whorl, between axial varices. Aperture small, rounded, erect, forming a continuous peristome. Columellar lip narrow, outer lip smooth within. Siphonal canal long, broad, straight, ventrally sealed, dorsally bent at tip. Left side of canal weakly overlapping right side. Operculum and radula unknown.	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF9FCF3EFC43EA4CFEF5FAAD.taxon	discussion	REMARKS Typhinellus constrictus n. sp. differs from all the other Typhinellus species in having a broad, almost straight shoulder ramp, deeply constricted teleoconch whorls with an impressed suture and a rounded aperture.	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF9FCF3EFC43EA4CFEF5FAAD.taxon	description	FIG. 13. — Distributions of: A, Typhinellus constrictus n. sp.; B, Siphonochelus (Siphonochelus) aethomorpha n. sp. Symbols: ★, type locality; ■, other occurences.	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF9FCF3EFC43EA4CFEF5FAAD.taxon	materials_examined	TYPE SPECIES. — Typhis arcuatus Hinds, 1843, South Africa (by original designation).	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF9DCF38FF24EACDFD31FC1D.taxon	materials_examined	TYPE MATERIAL. — Holotype MNHN-IM- 2000 - 30024, 10 paratypes MNHN-IM- 2000 - 30025, 2 paratypes Coll. RH. TYPE LOCALITY. — Madagascar, between Majunga and Cap Saint- André, 15 ° 35 ’ S, 45 ° 43 ’ E, 177 - 199 m [ATIMO VATAE: stn DW 3259]. MATERIAL EXAMINED. — South Madagascar. MIRIKY: stn DW 3197, West of Cap d’Ambre, 12 ° 07 ’ S, 48 ° 58 ’ E, 362 - 431 m, 3 dd, juv. — Stn DW 3239, in front of Baie de Nazendry, 14 ° 30 ’ S, 47 ° 26 ’ E, 230 - 288 m, 1 dd, juv. — Stn CP 3241, in front of Baie de Nazendry, 14 ° 30 ’ S, 47 ° 27 ’ E, 274 - 325 m, 1 dd, juv. — Stn DW 3258, between Majunga and Cap Saint-André, 15 ° 34 ’ S, 45 ° 44 ’ E, 200 - 288 m, 7 dd, juv. — Stn DW 3259, between Majunga and Cap Saint-André, 15 ° 35 ’ S, 45 ° 43 ’ E, 177 - 199 m, 30 dd, ad & juv (holotype MNHN- IM- 2000 - 30024, 10 paratypes MNHN-IM- 2000 - 30025, 2 paratypes Coll. RH). — Stn CP 3260, between Majunga and Cap Saint-André, 15 ° 35 ’ S, 45 ° 45 ’ E, 179 - 193 m, 1 dd, juv. — Stn CP 3283, off Baie Mahajamba, 14 ° 52 ’ S, 46 ° 56 ’ E, 228 - 257 m, 5 dd, juv.	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF9DCF38FF24EACDFD31FC1D.taxon	distribution	DISTRIBUTION. — North-western Madagascar, 193 - 362 m (shells only).	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF9DCF38FF24EACDFD31FC1D.taxon	etymology	ETYMOLOGY. — Greek, morpho, form, shape and aethes, unusual, strange. This species is named for the strange and flattened form of the anal tubes.	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF9DCF38FF24EACDFD31FC1D.taxon	description	DESCRIPTION Shell medium sized for the subgenus, up to 11.1 mm in height at maturity (holotype). Height / width ratio 1.9 - 2.1. Slender, lanceolate, narrowly ovate, lightly built. Subsutural ramp narrow, weakly sloping, weakly concave. Shell entirely off white. Spire high with 1.25 - 1.5 protoconch whorls and up to 5 - 5.25 weakly convex, weakly shouldered, smooth whorls. Suture impressed. Protoconch moderately large, narrow, slightly elongate. Whorls rounded, smooth, maximum width 600 µm. Terminal lip delicate, thin, curved. Axial sculpture consisting of four, low, narrow, rounded, weakly shouldered varices from first to last whorl. Spiral sculpture of very shallow P 1 with anal tube, P 2 (shoulder) and very weak, narrow striae and lirae, occasionally extending on abapical part of anal tubes. Anal tubes rounded on first or first two teleoconch whorls, flattened from second or third whorl. Last whorl with very broad, very flattened tubes originating from succeeding varix, filling almost entire space between succeeding and preceding varix, forming an angle of approximately 30 - 40 ° with axis of shell. Last apertural anal tube strongly outward bent, forming an angle of approximately 70 ° with axis of shell. Only apertural tube hollow and functional, older tubes broken off and closed. Only small portion of last tube open (Fig. 12 H). Aperture moderately small, ovate, weakly erect, forming a continuous peristome. Columellar lip narrow, smooth. Outer lip smooth within. Siphonal canal moderately long, narrow, straight, ventrally sealed, weakly dorsally curved, strongly tapered abapically.	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF9DCF38FF24EACDFD31FC1D.taxon	discussion	REMARKS	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
75255D75FF9DCF38FF24EACDFD31FC1D.taxon	description	Siphonochelus pentaphasios differs in the same characters and in having teleoconch whorls with five varices per whorl instead of four. Siphonochelus stillacandidus also differs in having much narrower anal tubes, slightly more distant from the more strongly shouldered axial varices, and in having a shorter siphonal canal. The two other Indian Ocean species differ in many shell characters and do not need to be compared here.	en	Roland, Houart, Virginie, Héros (2015): New species of Muricidae Rafinesque, 1815 (Mollusca: Gastropoda) from the Western Indian Ocean. Zoosystema 37 (3): 481-503, DOI: 10.5252/z2015n3a4, URL: http://dx.doi.org/10.5252/z2015n3a4
