identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
E4F76AB906955F789E8C10683883212D.text	E4F76AB906955F789E8C10683883212D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Beraea jennyae Rasmussen & Harris 2025	<div><p>Beraea jennyae Rasmussen &amp; Harris sp. nov.</p><p>Figs 6, 7</p><p>Beraea sp. nov.; original collection records, Rasmussen (2004).</p><p>Beraea sp. nov.; distributional record, Pescador et al. (2004).</p><p>Type material.</p><p>Holotype male (CMNH) • Florida, Okaloosa County, Turkey Hen Creek, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-86.55639&amp;materialsCitation.latitude=30.646667" title="Search Plazi for locations around (long -86.55639/lat 30.646667)">East branch steephead, Eglin Air Force Base, 0.3 km W of Okaloosa Lookout Tower, off SR- 85</a>, 30°38'48"N, 86°33'23"W, 8 April 1999, A. Rasmussen and M. Pescador, UV pan trap . Paratypes • Same data as holotype, 1 male (CMNH) • Same data as holotype except 9 April 1999, beating sheet, 1 female (CMNH) • Same data as holotype except 10 April 2001, 1 male (FAMU) .</p><p>Diagnosis.</p><p>Of the North America species, male and female genitalia of Beraea jennyae sp. nov. are more similar to B. fontana Wiggins (Figs 8 A – D, 9 A – D) and B. nigritta Banks (Figs 8 E – H, 9 E – H) than to B. gorteba Ross (Figs 8 I – L, 9 I – L). Males of the new species are easily distinguished from the three other North American species by the lyre-shaped intermediate appendages, which are short, not reaching the apex of the X th tergum; the intermediate appendages of B. fontana and B. nigritta in dorsal view are straighter and extend slightly beyond the X th tergum, while in B. gorteba the intermediate appendages are long and sinuous in dorsal view extending well past the X th tergum. Females of Beraea jennyae sp. nov. are differentiated from those of other North American species by the presence of a triangular, inward projecting process located along the midlength of each lateral lobe of the IX th sternite.</p><p>Description.</p><p>Fig. 6. Total length male 4.4–4.6 mm (mean = 4.5 mm, n = 3), female 4.0 mm (n = 1). In alcohol, body dark brown, legs somewhat lighter brown; wings light brown with dark brown hairs. General structure typical of the family and genus, as described by Ross (1944), Wiggins (1954), and Schmid (1998).</p><p>Male genitalia. Fig. 7 A – D. Segment VII annular, ventral mesal process elongate directed posteriorly. Segment VIII annular, posterior band of long setae dorsally and ventrally. Segment IX rectangular in lateral view; in ventral view posterior margin deeply concave. Preanal appendages clavate, about half length of intermediate appendages, stout setae posteriorly. Intermediate appendages in dorsal view lyre-shaped, curved outward basally, distally narrowed, terminating well short of Segment X apex. Segment X deeply incised dorsally forming two semi-membranous lobes, apex triangular with mesal margin angled outward; in lateral view rectangular, apex slightly rounded dorsally. Inferior appendages in lateral view base rectangular with membranous dorsal lobe directed posteroventrally, bearing many setae; in ventral view base narrow strap-like with curved mesal filament; stout circular hook posteriorly, with thumb-like basal process bearing setae. Phallus in ventral view thick, vasiform basally, single membranous lobe ventrally, paired lateral membranous lobes dorsally with pair of sclerotized styles running through middle terminating slightly beyond apex of lobes.</p><p>Female genitalia. Fig. 7 E – H. Segment VII, VIII in lateral view with tergite and sternite separated by wide membranous pleuron, sternum VII with prominent mesal process directed posteriorly. Segment VIII tergum with posterior band of long setae; pleuron with small setal-bearing sclerite; sternum forming ventral flap covering vaginal apparatus in ventral view. Segments IX + X fused dorsally, apically with pair of posterolateral processes each side of anal opening, processes triangular in dorsal and ventral views, dorsal process broader than ventral process, ventral process in lateral view with posteroventral sharp point. Vaginal apparatus in ventral view with pair of sclerotized lobes (Segment IX sternites) lateral to genital chamber, apex of each lobe bifurcated with mesal process pointed, lateral process broadly rounded; triangular inward projecting process midlength along mesal margin of each sternal lobe; genital chamber with sclerotized ventral pouch 8 - shaped, pointed posterior end.</p><p>Larva and pupa. Unknown.</p><p>Distribution.</p><p>The new species is known from only a single locality, a spring-fed ravine stream on Eglin Air Force Base in the western panhandle of Florida. Three other species are known in North America: Beraea nigritta and B. fontana each from a few localities, primarily in the northeast, and B. gorteba known from one stream in central Georgia.</p><p>Etymology.</p><p>This species is named for Xiaojing (Jenny) Wang, in appreciation of her love and support of the senior author throughout their marriage.</p><p>Remarks.</p><p>Specimens of adult males and females of the Beraea fontana, B. gorteba, and B. nigritta species were obtained for study from the U. S. National Museum, University of Minnesota Insect Collection, and the Clemson University Collection of Arthropods. Male and female genitalia for each species were illustrated (Figs 8, 9) and compared with B. jennyae sp. nov. These results may be used in a future review of the genus in North America.</p></div>	https://treatment.plazi.org/id/E4F76AB906955F789E8C10683883212D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Rasmussen, Andrew K.;Denson, Dana R.;Orfinger, Alexander B.;Harris, Steven C.	Rasmussen, Andrew K., Denson, Dana R., Orfinger, Alexander B., Harris, Steven C. (2025): Diversity and distribution of the Trichoptera of Florida, United States, with descriptions of five new species. ZooKeys 1263: 389-439, DOI: 10.3897/zookeys.1263.147317
96E070901E8B5FD285F758D6F19B6E36.text	96E070901E8B5FD285F758D6F19B6E36.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ceraclea pescadori Rasmussen & Harris 2025	<div><p>Ceraclea pescadori Rasmussen &amp; Harris sp. nov.</p><p>Figs 10, 11</p><p>Ceraclea sp. nov.; larval illustration, Pescador et al. 2004: fig. 168.</p><p>Ceraclea (Ceraclea) sp. nov. Glover, Carnagey, &amp; Morse; description of female, Carnagey and Morse 2006: 29–30, fig. 28.</p><p>Ceraclea sp. nov. (nr. maculata); collection record, Denson et al. (2016).</p><p>Type material.</p><p>Holotype male (CMNH) • Florida, Washington County: Lucas Lake at boat landing, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-85.69056&amp;materialsCitation.latitude=30.543612" title="Search Plazi for locations around (long -85.69056/lat 30.543612)">Lucas Lake Road, off County Road 279</a>, 30°32'37"N, 85°41'26"W, 17 April 2013, A. Rasmussen and N. Miller, UV pan trap . Paratypes • Same data as holotype, 10 males and 6 females (CMNH) • 10 males and 6 females (NMNH) • 10 males and 6 females (FAMU) • 10 males and 6 females (CUAC) • 5 males and 6 females (UMSP); Calhoun County • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-85.1975&amp;materialsCitation.latitude=30.539165" title="Search Plazi for locations around (long -85.1975/lat 30.539165)">Page Pond at Page Pond Assembly of God</a>, 30°32'21"N, 85°11'51"W, 7 May 2011, D. Denson, UV pan trap, 15 males (FAMU) .</p><p>Diagnosis.</p><p>Ceraclea pescadori sp. nov. is placed in the Senilis group of Morse (1975) based on genitalic features of the male and female (Carnagey and Morse 2006). The male is most similar to C. maculata, but is easily distinguished by the more elongate tergum X and the presence of the much larger spinous, knob-like median lobe of the inferior appendages. Additionally, the basoventral lobe of the inferior appendages in ventral view is broadly triangular in C. pescadori sp. nov., versus thumb-like in C. maculata . The female is distinguished from other Senilis group members by the prominent and elongate sclerotic bulge and preanal appendages, and in the short wedge-shaped median plate (see also Carnagey and Morse 2006: fig. 28 a, c).</p><p>Description.</p><p>Forewing length male 7.9–9.1 mm (mean = 8.3 mm, n = 10), female 7.7–8.1 mm (mean = 7.9 mm, n = 10). In alcohol, head and thorax reddish brown, abdomen light brown. Fore- and middle legs brown, hind legs light brown. Antennal scapes and pedicels brown, flagella light brown. Setal warts of head and thorax covered with primarily white hairs. Forewing R thickened near stigma, small translucent spots scattered anteriorly. Hind wings broad, unpatterned.</p><p>Male genitalia. Fig. 10. Segment IX in lateral view narrow, rounded posteriorly; in ventral view excised mesally. Superior appendages in lateral view divided into dorsal beak-like process and larger ventral flap; in dorsal view dorsomesal process pointed, lateral lobes widely lobate. Segment X elongate dorsally, extended as undivided process well past superior appendages; in lateral view concave along dorsal margin, strongly convex along ventral margin. Inferior appendages in lateral view with acute triangular basal plate; basoventral lobes prominent, bearing long setae, in lateral view extending posteriorly, in ventral view broadly triangular, directed posteromesally; in caudal view with prominent, spinous median knob; subapical dorsal lobes elongate, straight in lateral view, curved slightly inward in ventral and caudal views, bearing short setae basally, mix of long and short setae distally; harpagos slender, nearly as long as subapical dorsal lobes, curved apically, bearing small setae subapically. Phallus phallobase with broad dorsal lobe extending to about mid-length of phallus, divided apically in dorsal view; in lateral view elongate ventral lip longer than phalicata; parameres absent.</p><p>Female genitalia. Fig. 11. Segment IX in lateral view rectangular, dorsal sclerotic bulge prominent; preanal appendages setaceous, posterior margin narrowly rounded. In dorsal view tergum IX arched, separated with creases from sclerotic bulge with median finger-like process usually bifid. Preanal appendages broad, fused mesally. Lamellae crescent shaped in lateral view; pointed in dorsal and ventral views. In ventral view, lateral gonopod plates isolated, anteriorly broad, tapered posteriorly; median plate with anterior half triangular, forming wedge between lateral plates, rounded posteriorly; longitudinal striations present posterolaterally. Spermathecal sclerite in ventral view generally vase-shaped, pointed anteriorly, deltoid sclerite absent, lateral arms somewhat sinuous, truncate posterolaterally, posterior bridge prominent, projecting past median plate; in lateral view acute anteriorly, clavate posteriorly with serrate posteroventral margin.</p><p>Larva and pupa. Illustrations of the head and pro- and mesonotum prepared by Dr James Glover were presented by Pescador et al. (2004: fig. 168). Pupa unknown.</p><p>Distribution.</p><p>This species is known from only two natural lakes located in the western Florida panhandle.</p><p>Etymology.</p><p>This species is named for Dr Manuel (Manny) Pescador in recognition of his contributions to trichopterology in Florida. His mentorship of the senior author and leadership of the aquatic entomology program at Florida A &amp; M University are greatly appreciated.</p><p>Remarks.</p><p>This species was collected in the largest numbers from Lucas Lake, Washington County, Florida. Prior to this collection, Dr James Glover collected larvae and pupae of Ceraclea from Lucas Lake that he reared to adulthood in March / April of 2000. A preliminary description and figures of the male (Glover and Morse In Litt) confirm that the specimens collected by Glover represent the same species as we report here. Based on a single female pupa that was reared to adulthood by Glover, the female was described as Ceraclea sp. nov. (Carnagey and Morse 2006). Although we noted differences between their published description and the specimens we examined, especially in regard to the spermathecal sclerite, we contend that those differences are probably an artifact of lab rearing and specimen preparation or in interpretation, rather than the reared female specimen representing a different species. Also notable, single males of two other rare leptocerid caddisflies, Ceraclea limnetes and Oecetis parva, were collected in the same sample as the holotype.</p></div>	https://treatment.plazi.org/id/96E070901E8B5FD285F758D6F19B6E36	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Rasmussen, Andrew K.;Denson, Dana R.;Orfinger, Alexander B.;Harris, Steven C.	Rasmussen, Andrew K., Denson, Dana R., Orfinger, Alexander B., Harris, Steven C. (2025): Diversity and distribution of the Trichoptera of Florida, United States, with descriptions of five new species. ZooKeys 1263: 389-439, DOI: 10.3897/zookeys.1263.147317
11D995E5C36B52F09DBEAD4F5C0958F0.text	11D995E5C36B52F09DBEAD4F5C0958F0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydroptila aviforma Rasmussen & Harris 2025	<div><p>Hydroptila aviforma Rasmussen &amp; Harris sp. nov.</p><p>Fig. 5</p><p>Type material.</p><p>Holotype male (CMNH) • Florida, Washington County, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-85.69056&amp;materialsCitation.latitude=30.543612" title="Search Plazi for locations around (long -85.69056/lat 30.543612)">Lucas Lake at boat landing, Lucas Lake Road, off County Road 279</a>, 30°32'37"N, 85°41'26"W, 17 April 2013, A. Rasmussen and N. Miller, UV pan trap . Paratypes • Same data as holotype, 3 males (FAMU) .</p><p>Diagnosis.</p><p>Hydroptila aviforma sp. nov. shares a number of characters with several southeastern species notably, H. cretosa Harris, H. metteei, and H. wakulla . All have an elongate tenth segment and inferior appendages which are equal in length in lateral view; a tenth tergite which is split distally; segment IX completely enclosed within segment VIII, which is tapered and rounded distally in lateral aspect; and a phallus which is narrow over nearly the entire length with the ejaculatory duct protruding apically. Hydroptila aviforma sp. nov. differs from these species in the tenth segment being wide and bulbous at midlength in lateral aspect, with a setal-bearing sclerite; the tenth tergite being deeply divided with each side thin and truncate distally; the inferior appendages having elongate, thick setae subapically; and segment VII lacking a ventromesal process.</p><p>Description.</p><p>Male. Total length 2.8–3.0 mm (mean = 2.9 mm, n = 4). Antennae with 31 segments, brown in alcohol. Genitalia as in Fig. 5. Segment VII annular without a ventromesal process. Segment VIII wide anteriorly, rounded posteriorly in lateral view; dorsally incised on posteromesal margin; quadrate ventrally. Segment IX in lateral view short, incised posteroventrally, completely enclosed within segment VIII; dorsally divided mesally, posterolateral margins pointed distally; ventrally deeply incised on anterior margin. Segment X in lateral view widening at midlength, bulbous ventrally and bearing short, stout setae, tapering posteriorly to downturned pointed apex; dorsally deeply divided mesally, each lateral branch parallel-sided over length, truncate apically; ventrally with setal-bearing plate anterior of split. Inferior appendages thin and sickle-shaped in lateral aspect, bearing cluster of thickened, elongate setae posterodorsally, equal in length to segment X; in ventral view narrow and parallel-sided over much of length, sinuate posteriorly, pointed apically, with thickened setae subapically. Phallus elongate and thin over length, thin paramere encircling shaft below midlength, ejaculatory duct protruding apically.</p><p>Adult female, larva, and pupa. Unknown.</p><p>Distribution.</p><p>Known from only the type locality, a natural lake in the Florida panhandle.</p><p>Etymology.</p><p>Latin avis (bird) and forma (shape) referring to the bird-shape form of the apex of the tenth segment as seen in lateral view.</p><p>Remarks.</p><p>Hydroptila quinola (11 individuals) was the only other species of Hydroptila collected in the same sample as the new species. The addition of this new species brings the total number of Florida endemics in the microcaddisflies to 21, 14 of which are in the genus Hydroptila .</p></div>	https://treatment.plazi.org/id/11D995E5C36B52F09DBEAD4F5C0958F0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Rasmussen, Andrew K.;Denson, Dana R.;Orfinger, Alexander B.;Harris, Steven C.	Rasmussen, Andrew K., Denson, Dana R., Orfinger, Alexander B., Harris, Steven C. (2025): Diversity and distribution of the Trichoptera of Florida, United States, with descriptions of five new species. ZooKeys 1263: 389-439, DOI: 10.3897/zookeys.1263.147317
BA399CFD7D615651AFF1771EF9897FAA.text	BA399CFD7D615651AFF1771EF9897FAA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Oecetis densoni Rasmussen & Harris 2025	<div><p>Oecetis densoni Rasmussen &amp; Harris sp. nov.</p><p>Figs 12, 13</p><p>Oecetis sp. nov. (nr. cinerascens); collection records, Rasmussen et al. (2008 a).</p><p>Oecetis sp. nov. (nr. cinerascens); collection records, Denson et al. (2016).</p><p>Type material.</p><p>Holotype male (CMNH) • Florida, Lake County: Sellars Lake, Ocala National Forest, southeast lobe of lake, near canoe launch, 28 April 2007, D. Denson, A. Rasmussen, UV pan trap . Paratypes • Same data as holotype, 39 males, 1 female (CUAC) • unnamed lake, Ocala National Forest, jeep trail South off State Road 40, just East of Wildcat Lake, 17 May 2009, D. Denson, A. Rasmussen, UV pan trap, 84 males and 21 females (CMNH) ; Marion County • Lake Delancy, Ocala National Forest, Lake Delancy campground, 27 April 2007, D. Denson, A. Rasmussen, UV pan trap, 14 males and 1 female (USNM) • Fore Lake, Ocala National Forest, Fore Lake Recreation Area, 16 May 2008, D. Denson, A. Rasmussen, UV pan trap, 6 males (FAMU) ; Calhoun County • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-85.14278&amp;materialsCitation.latitude=30.426111" title="Search Plazi for locations around (long -85.14278/lat 30.426111)">Wildcat Creek at State Road 20</a>, 30°25'34"N, 85°08'34"W, 6 May 2011, D. Denson, 1 male (FAMU) • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-85.1975&amp;materialsCitation.latitude=30.539165" title="Search Plazi for locations around (long -85.1975/lat 30.539165)">Page Pond at Page Pond Assembly of God</a>, 30°32'21"N, 85°11'51"W, 7 May 2011, D. Denson, UV pan trap, 6 males (FAMU) ; Jackson County • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-85.276115&amp;materialsCitation.latitude=30.588057" title="Search Plazi for locations around (long -85.276115/lat 30.588057)">Porter Pond at Pittman Hall Road</a>, 30°35'17"N, 85°16'34"W, 20 May 2010, D. Denson, A. Rasmussen, UV pan trap, 1 male (FAMU) ; Leon County • Lofton Ponds, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-84.390274&amp;materialsCitation.latitude=30.36111" title="Search Plazi for locations around (long -84.390274/lat 30.36111)">Apalachicola National Forest, at Sam Allen Road</a>, 30°21'40"N, 84°23'25"W, 24 April 2007, M. Heyn, A. Wilson, UV-Light, 1 male (FAMU) • same as above except 25 June 2007, D. Denson, UV pan trap, 1 male (FAMU); Liberty County • Camel Pond, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-84.988884&amp;materialsCitation.latitude=30.276945" title="Search Plazi for locations around (long -84.988884/lat 30.276945)">Apalachicola National Forest, Camel Pond campground, Forest Road 105</a>, 30°16'37"N, 84°59'20"W, 16 May 2006, R. Flowers, A. Rasmussen, B. Richard, Mercury-vapor light, 3 males (UMSP) ; Washington County • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-85.69056&amp;materialsCitation.latitude=30.543612" title="Search Plazi for locations around (long -85.69056/lat 30.543612)">Lucas Lake at boat landing, Lucas Lake Road</a>, 30°32'37"N, 85°41'26"W, 17 April 2013, N. Miller, A. Rasmussen, UV pan trap, 4 males (UMSP) .</p><p>Diagnosis.</p><p>Oecetis densoni sp. nov. belongs to the subgenus Pleurograpta, defined by Chen (1993) based on males having a very short IX th tergum and long IX th sternum. The new species is most similar to Oecetis (Pleurograpta) cinerascens in general appearance and shared characteristics of the male and female genitalia. The male of O. densoni sp. nov. and O. cinerascens differ from most species in the subgenus by the presence of one or no paramere spines in the phallus, versus many paramere spines typical of the subgenus. The new species differs from O. cinerascens in the absence of honeycomb sculpturing on the abdominal terga VI and VII and the dorsal margin of the inferior appendages having a less abrupt downturn midlength, as seen in lateral view. Females of both species have large circular gonopod plates with an anterior arcuate band of sclerotization and a posterior pair of concave sclerites. The new species female differs in having a more prominent mid-dorsal hump on segment IX and triangular preanal appendages versus in O. cinerascens a much less produced mid-dorsal hump on segment IX and less prominent, broadly rounded preanal appendages (in lateral view).</p><p>Description.</p><p>Fig. 12. Forewing length male 8.4–10.2 mm (mean = 9.1 mm, n = 10), female 7.8–8.9 mm (mean = 8.1 mm, n = 10). Forewing with transverse base of MA and MP stem crossveins aligned diagonally at approximately 60 o angle, veins slightly darkened at forks. Nigma of fore and hind wings located near crossvein forks. In alcohol, head, wings, thorax, and legs reddish brown in males and grayish brown in females. Antennal scapes and pedicels reddish brown, flagella pale brown. Abdomen cream color, without honeycomb sculpturing on abdominal terga V – VIII.</p><p>Male genitalia. Fig. 13 A – D. Segment IX narrow dorsally, widened dorsolaterally, posterior margin somewhat angulate in lateral view, ventrally with thick line of sclerotization separating posterior sternal area; in dorsal view sclerotized papillae resembling rabbit ears posteriorly; in ventral view each posterolateral corner with line of heavy sclerotization diagonally directed anterad. Preanal appendages ovoid in lateral view, digitate in dorsal view, flared posterolaterally. Segment X consisting of sclerotized mid-dorsal process subequal in length to preanal appendages, subtended on each side by longer membranous ventral process; in lateral view dorsal process arising from under base of preanal appendage, ventral process curved posteroventally, far surpassing dorsal process; in dorsal view apical margin of dorsal process variable among populations, grading from entire to deeply incised. Inferior appendages symmetric; in lateral view base with dorsomesal point, dorsal margin irregularly scalloped anteriorly, posteriorly angled downward and smoothly tapered, apically upturned; in dorsal view ridge separating lateral area from excavated mesal area; in ventral view mesal margins scalloped, slightly concave, outer margins parallel basally, distally tapered, curved inward. Phallus without paramere spines; in lateral view phallobase sharply angled anteriorly, rectangular along mid-length, sharply downturned distally with lip upturned apically, tear drop-shaped phallotremal sclerite near ventral downturn of phallobase dorsal margin.</p><p>Female genitalia. Fig. 13 E – G. Segment IX in lateral view narrow, truncate posteroventral extension above concave emargination; in dorsal view broadly rounded median hump with rabbit ear-like papillae produced posteriorly; in ventral view posterolateral corners each with triangular projection with sclerotized line connecting apex to perpendicular sclerotized line at base. Preanal appendages prominent, subtriangular in both lateral and dorsal views. Lamellae ovate in lateral view; in ventral view each with ear-like projection. Gonopod plate (sternum IX) in ventral view large, subcircular with anterior arcuate band of sclerotization, posterior pair of concave sclerites. Bursa copulatrix [spermathecal sclerite of Chen (1993)] in ventral view with median elongate ring surrounded by deltoid sclerite anteriorly, posterior margin sclerotized forming inverted V.</p><p>Larva and pupa. Unknown. Based on the high similarity of adults of Oecetis densoni sp. nov. with O. cinerascens and the collection of the two species from similar lake habitats in Florida, the larva of the new species is likely to also be similar to O. cinerascens, as described and illustrated by Floyd (1995).</p><p>Distribution.</p><p>Oecetis densoni sp. nov. appears to be endemic to Florida’s clear lakes and ponds where its known geographic range extends from the Ocala National Forest northward into the Florida panhandle as far west as Washington County.</p><p>Etymology.</p><p>This species is named in honor of Dana R. Denson, who helped discover the species and has contributed greatly to the study and protection of Trichoptera biodiversity in Florida.</p><p>DNA Barcoding.</p><p>To visualize putative phylogenetic relationships, a Neighbor Joining (NJ) tree was constructed (Suppl. material 3) based on barcoding sequences of mitochondrial cytochrome oxidase I (COI) nucleotides using the built-in tree-builder of the Barcode of Life Database (BOLD) using Florida specimens examined in the course of this study. Based on this tree and morphological affinity, O. cinerascens (Hagen) is hypothesized to be the sister species of Oecetis densoni sp. nov. For O. densoni sp. nov. and O. cinerascens, pairwise divergence distances (p-distances) of COI sequences were calculated in MEGA 11 (Tamura et al. 2021) using the Kimura 2 - parameter evolution model (K 2 P) (Kimura 1980) and pairwise deletion of missing sites. Sequences were mined from BOLD and include both available Oecetis densoni sp. nov. sequences from specimens included in the present study (BOLD FLCAD 120-09 and FLCAD 140-09) as well as specimens of O. cinerascens from the Ocala National Forest (FLCAD 130-09), northwest Florida (FLCAD 148-09 and FLCAD 056-08), and Virginia (PKCAD 021-07), to account for population-level genetic variation. Nucleotide sequences were aligned using ClustalW using default settings (Larkin et al. 2007). All COI sequences used consist of 658 bp and were generated from vouchered specimens identified by taxonomic experts. Interspecific pairwise p-distance between O. densoni sp. nov. and O. cinerascens ranged from 6.4 % to 6.9 % (Suppl. material 2). These values are substantially larger than intraspecific p-distance ranges for O. densoni sp. nov. (0.08 %) and O. cinerascens (0.2 % to 0.5 %), demonstrating a clear barcoding gap indicative of significant genetic differentiation between the congeners.</p><p>Remarks.</p><p>The presence of honeycomb sculpturing on abdominal terga V – VIII has been used to place Oecetis (Pleurograpta) species in the Testacea species group (Chen 1993). However, phylogenetic analysis by Quinteiro and Almeida (2021) of Neotropical Oecetis suggested that the Testacea species group is paraphyletic and that the honeycomb sculpturing of the abdominal terga probably originated more than once due to convergent evolution. In this study, the other Testacea species group members found in Florida, Oecetis georgia and O. persimilis, were shown to be closely related to each other (Suppl. material 3) but in a separate clade from O. cinerascens and O. densoni sp. nov. This is further evidence that honeycomb sculpturing is of limited use for placement of species into monophyletic groupings. Additionally, the presence of abdominal tergal sculpturing in O. cinerascens and the absence of it in O. densoni sp. nov, along with the relatively high degree of genetic and morphological similarities between the two species, further underscore the probable high evolutionary plasticity of abdominal tergal sculpturing in Trichoptera .</p></div>	https://treatment.plazi.org/id/BA399CFD7D615651AFF1771EF9897FAA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Rasmussen, Andrew K.;Denson, Dana R.;Orfinger, Alexander B.;Harris, Steven C.	Rasmussen, Andrew K., Denson, Dana R., Orfinger, Alexander B., Harris, Steven C. (2025): Diversity and distribution of the Trichoptera of Florida, United States, with descriptions of five new species. ZooKeys 1263: 389-439, DOI: 10.3897/zookeys.1263.147317
7BF5FD7A0FD55291A096A29CA9461062.text	7BF5FD7A0FD55291A096A29CA9461062.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protoptila chipolensis Rasmussen & Harris 2025	<div><p>Protoptila chipolensis Rasmussen &amp; Harris sp. nov.</p><p>Fig. 4</p><p>Protoptila sp. nov.; collection records, Denson et al. (2016).</p><p>Type material.</p><p>Holotype male (CMNH) • Florida, Jackson County, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-85.2225&amp;materialsCitation.latitude=30.661943" title="Search Plazi for locations around (long -85.2225/lat 30.661943)">Chipola River at Caverns Road (State Road 166)</a>, 30°39'43"N, 85°13'21"W, 6 May 2010, D. Denson, E. Denson, UV pan trap . Paratypes • Same data as holotype, 20 males and 10 females (CMNH) • 20 males and 10 Females (CUAC) • 20 males and 10 females (FAMU) • 20 males and 10 females (NMNH) • 20 males and 10 females (UMSP); Florida, Jackson County • Rocky Creek at Highway 71, 18 May 1994, M. Pescador, S. Harris, 3 males and 1 female (FAMU) • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-85.221664&amp;materialsCitation.latitude=30.793333" title="Search Plazi for locations around (long -85.221664/lat 30.793333)">Chipola River at State Road 167 near Marianna</a>, 30°47'36"N, 85°13'18"W, 4 October 2006, D. Denson, UV pan trap, 10 males (CMNH) • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-85.25889&amp;materialsCitation.latitude=30.87" title="Search Plazi for locations around (long -85.25889/lat 30.87)">Chipola River at State Road 162</a>, 30°52'12"N, 85°15'32"W, 2 October 2010, D. Denson, E. Denson, UV pan trap, 14 males and 18 females (CUAC) • same as previous, except 6 August 2011, D. Denson, 19 males and 22 females (UMSP) • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-85.16556&amp;materialsCitation.latitude=30.626667" title="Search Plazi for locations around (long -85.16556/lat 30.626667)">Chipola River at Peacock Bridge ramp</a>, 30°37'36"N, 85°09'56"W, 21 May 2010, D. Denson, A. Rasmussen, UV pan trap, 25 males and 44 females (FAMU) • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-85.2225&amp;materialsCitation.latitude=30.661943" title="Search Plazi for locations around (long -85.2225/lat 30.661943)">Pelt Creek at State Road 73</a>, 30°39'43"N, 85°13'21"W, D. Denson, UV pan trap, 12 males and 20 females (NMNH) • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-85.170555&amp;materialsCitation.latitude=30.55361" title="Search Plazi for locations around (long -85.170555/lat 30.55361)">Hollis Branch upstream of Chipola River</a>, 30°33'13"N, 85°10'14"W, 21 May 2010, D. Denson, A. Rasmussen, UV pan trap, 8 males, 13 females (UMSP) • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-85.1625&amp;materialsCitation.latitude=30.651388" title="Search Plazi for locations around (long -85.1625/lat 30.651388)">Bridge Creek at State Road 71</a>, 30°39'05"N, 85°09'45"W, 7 May 2011, D. Denson, UV pan trap, 1 male (FAMU) • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-85.27889&amp;materialsCitation.latitude=30.850834" title="Search Plazi for locations around (long -85.27889/lat 30.850834)">Waddell’s Mill Creek upstream of Chipola River</a>, 30°51'03"N, 85°16'44"W, D. Denson, E. Denson, UV pan trap, 18 males and 108 females (CMNH) ; Calhoun County • Chipola River at Highway 20, 27 September 1972, P. Carlson 2 males 3 females (CUAC) • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-85.165&amp;materialsCitation.latitude=30.534166" title="Search Plazi for locations around (long -85.165/lat 30.534166)">Chipola River at Highway 274</a>, 30°32'03"N, 85°09'54"W, 17 May 1994, M. Pescador, S. Harris, R. Flowers, UV pan trap, 78 males and 27 females (FAMU) • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-85.17139&amp;materialsCitation.latitude=30.551388" title="Search Plazi for locations around (long -85.17139/lat 30.551388)">Chipola River at boat ramp 5 km SW of Altha</a>, 30°33'05"N, 85°10'17"W, 28 March 1998, M. Pescador, A. Rasmussen, UV pan trap, 4 males and 1 female (FAMU) • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-85.158615&amp;materialsCitation.latitude=30.512222" title="Search Plazi for locations around (long -85.158615/lat 30.512222)">Chipola River at Laramore Landing</a>, 30°30'44"N, 85°09'31"W, 20 May 2010, A. Rasmussen, D. Denson, D. Ray, R. Abad, L. Brooks, Mercury vapor sheet, 29 males and 12 females (UMSP) • same as previous, except 6 May 2011, D. Denson, UV pan trap, 7 males and 137 females (CUAC) • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-85.16167&amp;materialsCitation.latitude=30.524721" title="Search Plazi for locations around (long -85.16167/lat 30.524721)">Chipola River at Look and Tremble Rapids</a>, 30°31'29"N, 85°09'42"W, 6 May 2011, D. Denson, UV pan trap, 4 males and 37 females (CMNM) • same as previous, except 28 October 2011, D. Denson, E. Denson, 9 males and 22 females (FAMU) .</p><p>Other material examined.</p><p>Florida, Calhoun County • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-85.17139&amp;materialsCitation.latitude=30.551388" title="Search Plazi for locations around (long -85.17139/lat 30.551388)">Chipola River at boat ramp 5 km SW of Altha</a>, 30°33'05"N, 85°10'17"W, 26 March 2010, A. Rasmussen, D. Denson, dipnet, 1 larva (FAMU) • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-85.165&amp;materialsCitation.latitude=30.534166" title="Search Plazi for locations around (long -85.165/lat 30.534166)">Chipola River at Highway 274</a>, 30°32'03"N, 85°09'54"W, 3 September 2010, A. Rasmussen, C. Zhou, dipnet, 3 larvae and 1 pupa (FAMU) .</p><p>Diagnosis.</p><p>The male of Protoptila chipolensis sp. nov. is most similar to P. maculata (Hagen) in the narrow, elongate similarly-shaped sternum VIII and the thick phallic parameres that are sharply turned apically; the parameres of P. chipolensis sp. nov., however, are less thick than those of P. maculata . The new species is most easily distinguished from all other Nearctic Protoptila by the shape of Segment X, the arms of which are stout and strongly decurved in lateral view and terminate in a distinctive cuspidate apex; in P. maculata the segment X arms are straighter and more slender. The female of P. chipolensis sp. nov. is similar to P. maculata and P. lega Ross in that all three species possess broad circular ventral plates on sternum VIII, but appears to be distinguishable by the shape and sclerotization of the internal structures of the vaginal apparatus. Additionally, P. chipolensis sp. nov. lacks a sclerotized dome-like band above the vaginal apparatus that is found in the other species [see Morse (1988: fig. 2 A)].</p><p>Description.</p><p>Total length male 3.3–3.8 mm (mean = 3.6 mm, n = 10), female 3.2–4.0 mm (mean = 3.6 mm, n = 10). General structure typical of genus, as described by Robertson and Holzenthal (2013). In alcohol, head and thorax reddish brown; antennae, wings, and legs tawny; abdomen pale brown. Forewings with transverse translucent line at mid-length. Abdominal sterna clothed with long hairs; sternum VI with midventral keel-like projection.</p><p>Male genitalia. Fig. 4 A – C. Segment VIII tergum with darkened narrow band with long setae along posterior margin; in lateral view sternum dark brown basally, ventral margin straight, posterior projection elongate, slightly upturned apically; in ventral view sternum elongate, apex with rounded V-shaped mesal incision. Segment IX sclerotized, mostly enclosed within segments VII and VIII, sternum apically divided into pair of setal bearing processes. Segment X posteriorly divided into pair of curved arms; in lateral view bulbous basally with broad setal-bearing lobe, distally arm wide, decurved, cuspidate apically. Phallic apparatus nested within Segment IX attached to rounded apodeme; basoventrally with paired digitate rods, apically setose, appressible into pockets on underside of phallus; in lateral view pair of intertwined processes form ventral cup near midlength; pair of stout parameres attached to phallus dorsolaterally, in lateral view each paramere usually tucked underneath phallus, abruptly curved dorsomesally near apex, lightly spinous at tips; phallus in lateral view terminating in rounded head (phallicata) with posteroventral projection.</p><p>Female genitalia. Fig. 4 D, E. Segment VII unmodified, tergal and sternal plates rectangular. Segment VIII in lateral view dorsally rounded, constricted laterally, ventrally ovate with sharply pointed posteroventral projection; in ventral view ventral plates broadly rounded with erect black setae. Venter with vulvar lip slightly bilobed, posterior membranous area terminating in paired slender projections similar to cerci. Segment IX + X fused, posterolateral lobes broad; in lateral view lobes rectangular, slender cercus extending from apex; in ventral view lobes broadly conical with sinuous posterior margin. Internal apodeme “ whip ” attached ventrally near vulvar lip, extended cephlad far into Segment VII. Internal vaginal apparatus box-like in ventral view with circular spermathecal duct, paired elliptical structures, two pairs sclerotized lateral bands, in lateral view outermost band dorsal to inner band.</p><p>Larva and pupa. Undescribed. Larvae and a pupa of P. chipolensis sp. nov. have been collected and preserved for future study (see above material examined).</p><p>Distribution.</p><p>Known from only the Chipola River and tributaries in Jackson and Calhoun counties in the western panhandle of Florida. Collection records of the species from 14 sites were presented by Denson et al. (2016) as part of a faunal survey of the basin, which documented 143 species of Trichoptera .</p><p>Etymology.</p><p>This species is named after the Chipola River basin where the species was collected.</p><p>Remarks.</p><p>The Chipola River flows nearly 100 miles through northwest Florida emptying into the Apalachicola River. In many ways the Chipola River is atypical of most Florida rivers as it is fed by 63 freshwater springs which prevents water temperatures from becoming inhospitably warm in the summer. Portions of the river flow over limestone outcroppings, which is the primary larval habitat of the new species (Pescador et al. 2004) and Setodes chipolanus (Rasmussen et al. 2008 b), which is also endemic to the Chipola River basin.</p></div>	https://treatment.plazi.org/id/7BF5FD7A0FD55291A096A29CA9461062	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Rasmussen, Andrew K.;Denson, Dana R.;Orfinger, Alexander B.;Harris, Steven C.	Rasmussen, Andrew K., Denson, Dana R., Orfinger, Alexander B., Harris, Steven C. (2025): Diversity and distribution of the Trichoptera of Florida, United States, with descriptions of five new species. ZooKeys 1263: 389-439, DOI: 10.3897/zookeys.1263.147317
