identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
7B6687A7FFEFFF87FF12D75EFA1CFE9E.text	7B6687A7FFEFFF87FF12D75EFA1CFE9E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Seirinae Yosii, 1961 sensu Zhang & Deharveng 2015	<div><p>Key to Seirinae genera</p><p>1 Th II with one anterior extra group (a5 group) of at least 8 mac, p5 chaeta always as mic (Figs 13, 15A, 20A, 47A); unguis outer side with a pair of large laterobasal teeth (Figs 12C, 28E, 37F); Brazil and Paleotropical region..................................................................................................... Lepidocyrtinus B̂rner, 1903</p><p>- Th II with one anterior extra group (a5 group) with 5 or less mac, p5 chaeta mostly as mac; unguis outer side with a pair of normal laterobasal teeth................................................................................ 2</p><p>2 Inner labral papillae pointed; leg I of males generally without modifications, if present then femur normal and with elongated spine-like mac displaced in one longitudinal row on the median half; Worldwide................... Seira Lubbock, 1870</p><p>- Inner labral papillae bifurcate distally; leg I of males with femur internally strongly enlarged and with one subdistal group of short pointed spines displaced on transverse rows; Brazil ....................... Tyrannoseira Bellini &amp; Zeppelini, 2011</p></div>	https://treatment.plazi.org/id/7B6687A7FFEFFF87FF12D75EFA1CFE9E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cipola, Nikolas Gioia;Morais, José Wellington De;Bellini, Bruno Cavalcante	Cipola, Nikolas Gioia, Morais, José Wellington De, Bellini, Bruno Cavalcante (2020): Review of Lepidocyrtinus Börner, 1903 (Collembola, Entomobryidae, Seirinae): the African species. Zootaxa 4898 (1): 1-110, DOI: 10.11646/zootaxa.4898.1.1
7B6687A7FFFAFF90FF12D290FF71FD4C.text	7B6687A7FFFAFF90FF12D290FF71FD4C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lepidocyrtinus annulicornis (Borner 1903) sensu Coates 1968	<div><p>Lepidocyrtinus annulicornis (Börner, 1903) sensu Coates, 1968</p><p>Figs 2, 11, Tables 1 and 3</p><p>Lepidocyrtus (Lepidocyrtinus) annulicornis B̂rner, 1903: 154, Mozambique, Lake Malawi (as Nyassa See), not examined.</p><p>Pseudosira (Mesira) annulicornis; B̂rner 1908: 56.</p><p>Lepidocyrtus annulicornis; Womersley 1934: 463. Lepidocyrtinus (Mesira) annulicornis; Denis 1935: 5, Mozambique, stations: Bas Sangadzé, Vila Pery, Nova Choupanga. Seira annulicornis; Coates 1968: 449, figs 43-44, Mozambique and Africa South: Pongola and Pretoria. Coates 1970: 182,</p><p>South Africa, Kruger National Park (catalog). Jacquemart 1974: 7 (cited). Janion-Scheepers et al. 2015: 68, Africa South</p><p>(catalog).</p><p>Diagnosis. Body pale white with dark violet pigments on Ant I–IV, proximal leg III, femur and tibiotarsus distally; Th II projected anteriorly and with 6 median (m1ip present) in m1–2 complex and 11 posterior mac (p3 absent); Th III–Abd II with 9, 5, 4 central mac respectively (Fig. 11A); unguis a.t. present; unguiculus pe lamella serrated but toothless; manubrial plate with 2–3 blunt mac; dens with 4 blunt mac (Fig. 11B) (modified from B̂rner 1903; Coates 1968).</p><p>Remarks. The specimen of L. annulicornis described by Yosii (1959) from Cape Town does not show the same morphology of specimens from Mozambique (sensu B̂rner 1903; Denis 1935; Coates 1968) (Fig. 2), and for this reason they are herein designated as L. ryozoi nom. nov. In this same sense the chaetotaxy description of Szeptycki (1979), as well as the interpretation by Soto-Adames (2008) of Seira annulicornis (locality not specified) also do not fit neither the description of Yosii (1959) or Coates (1968), and consequently should not be considered as identity of this species. Even so, L. annulicornis differs from these species by body with dark violet pigments on Ant I–IV, proximal leg III, femur and tibiotarsus distally, while L. ryozoi nom. nov. has pigments on Ant III–IV, Ant II and tibiotarsus I–III distally (unknow in Szeptycki’s specimens). Lepidocyrtinus annulicornis differs in chaetotaxy by Th II with 6 median mac in m1–2 complex (5 in L. ryozoi and Szeptycki’s specimens) and posteriorly without p3 mac (present in L. ryozoi). They also differ by Th III with p1i mac and p1i2 mac absent (opposite in L. ryozoi), and dens with 4 blunt mac (9–10 in L. ryozoi). Other differences among similar species to L. annulicornis are listed in Table 3.</p></div>	https://treatment.plazi.org/id/7B6687A7FFFAFF90FF12D290FF71FD4C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cipola, Nikolas Gioia;Morais, José Wellington De;Bellini, Bruno Cavalcante	Cipola, Nikolas Gioia, Morais, José Wellington De, Bellini, Bruno Cavalcante (2020): Review of Lepidocyrtinus Börner, 1903 (Collembola, Entomobryidae, Seirinae): the African species. Zootaxa 4898 (1): 1-110, DOI: 10.11646/zootaxa.4898.1.1
7B6687A7FFFBFF90FF12D35CFDC1FA9A.text	7B6687A7FFFBFF90FF12D35CFDC1FA9A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lepidocyrtinus ryozoi Cipola & Morais & Bellini 2020	<div><p>Lepidocyrtinus ryozoi nom. nov. Cipola &amp; Bellini</p><p>Figs 2, 12, Tables 1 and 3</p><p>Seira (Lepidocyrtinus) annulicornis; Yosii 1959: 10, figs 5A–H, South Africa, Cape Town, Table Mountain (Fig. 2), not examined; Janion-Scheepers et al. 2015: 68, Africa South (catalog).</p><p>Diagnosis. Body pale white with violet pigments on Ant III–IV, Ant II and femur I–III distally; Th II projected anteriorly and with 14 anterior, 7 median (m1ip absent, m4 and m4i present) and 11 posterior mac (p3 present); Th III–Abd II with 9, 5, 4 central mac respectively (Fig. 12A); Abd IV with 17 central mac and 8 lateral mac (Fig. 12B); unguis a.t. present; unguiculus with all lamella acuminate and toothless (Fig. 12C); manubrial plate with 3 blunt mac; dens with 9–10 blunt mac apically nipple-like (Fig. 12D).</p><p>Etymology. It is named after Dr. Ryozo Yoshii (in memoriam), an important Collembola researcher.</p><p>Remarks. Lepidocyrtinus ryozoi nom. nov. resembles L. annulicornis in general morphology (see the remark sections of these species and Table 3).</p></div>	https://treatment.plazi.org/id/7B6687A7FFFBFF90FF12D35CFDC1FA9A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cipola, Nikolas Gioia;Morais, José Wellington De;Bellini, Bruno Cavalcante	Cipola, Nikolas Gioia, Morais, José Wellington De, Bellini, Bruno Cavalcante (2020): Review of Lepidocyrtinus Börner, 1903 (Collembola, Entomobryidae, Seirinae): the African species. Zootaxa 4898 (1): 1-110, DOI: 10.11646/zootaxa.4898.1.1
7B6687A7FFFBFF99FF12D569FB91F991.text	7B6687A7FFFBFF99FF12D569FB91F991.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lepidocyrtinus barnardi Womersley 1934	<div><p>Lepidocyrtinus barnardi Womersley, 1934</p><p>Figs 2, 13–17, Tables 1 and 4</p><p>Lepidocyrtinus cooperi var. barnardi Womersley, 1934: 461, fig. 10, South Africa, Cape Town, Kirstenbosch, not examined. Seira (Lepidocyrtinus) barnardi; Yosii 1959: 12, figs 6A–I, South Africa, Cape Town, Kirstenbosch.</p><p>Seira barnardi; Coates 1968: 454; Janion-Scheepers et al. 2015: 68, Africa South (catalog).</p><p>Typological note. One cotype of four specimens used in the original description of Lepidocyrtinus cooperi var. barnardi by Womersley (1934) is deposited at SAM/IZIKO (Scheepers, personal communication). We could not analyse this material, but our specimens fit the descriptions of Womersley (1934) and Yosii (1959) and they were collected from the type locality (Fig. 2). For this reason, the redescription is justified.</p><p>Examined material. 1 male and 4 females in slides and 1 specimen in alcohol (INPA): South Africa, Cape Town, Table Mountain National Park, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=18.440304&amp;materialsCitation.latitude=-33.964417" title="Search Plazi for locations around (long 18.440304/lat -33.964417)">Newlands</a>, 33°57’51.9”S, 18°26’25.1”E (Fig. 2), in Pine Plantation, 200 m, 12.ix.2008, Leaf Litter, C Uys coll. 3 females in slides and 170 specimens in alcohol (CC / UFRN): 33°57’58.3”S, 18°26’31.6”E, in <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=18.44211&amp;materialsCitation.latitude=-33.966194" title="Search Plazi for locations around (long 18.44211/lat -33.966194)">Afrotemperate Forest</a>, 230 m, 15.i.2009, Sugar-baited Ant Trap, C Uys coll. 23 specimens in alcohol (INPA): Spilhaus, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=18.414333&amp;materialsCitation.latitude=-33.99825" title="Search Plazi for locations around (long 18.414333/lat -33.99825)">Cecilia Ravine</a>, 33°59’53.7”S, 18°24’51.6”E, in Fynbos, 520 m, 23.i.2009, Sugar-baited Ant Trap, C Uys coll.</p><p>Diagnosis. Body with dark blue pigments on Ant III–IV, Th II laterally up to one transverse band on Abd III, and spots on Abd IV–V and legs (Fig. 13); head mac A0, A2–3, A5, M1–2, M4i, S0–6 and Pp5 present; prelabral chaetae smooth; labial papilla E with l.p. apically pointed and not reaching the base of a.a. (Figs 14 D–F); basolateral and basomedian labial fields with chaetae R (smaller than others) ciliated, e and l2 smooth, L1 ciliated or smooth; Th II not projected anteriorly and with 17 anterior, 9 median (m1ip, m4i–4p present) and 15–16 posterior mac (p2ep, p2ep2, p3 as mac); Th III–Abd II with 10, 5–6, 4 central mac respectively; Abd IV with 10–11 central mac and 11 lateral mac (Figs 15, 16B); unguis a.t. present or absent; unguiculus with pe lamella serrated and with small proximal tooth; collophore anteriorly with 4–5 spine-like chaetae and 4 ciliated chaetae apically acuminate, posteriorly without spines, smooth chaeta present (Fig. 17E); manubrium ventrally with 2/2 subapical chaetae; manubrial plate without blunt mac; dens with with 13–19 blunt mac apically nipple-like (Figs 17 F–G).</p><p>Description. Total length (head + trunk) of specimens 2.21–3.22 (n=4). Specimens pale white with dark blue pigment on distal Ant II to Ant IV, anterior head, lateral Th II to Abd II which extends to most of the Abd III, central spots on Abd IV and V, coxa II–III, and trochanter to proximal half of tibiotarsus; eyepatches black (Fig. 13). Scales present on Ant I to basis of Ant III, dorsal and ventral head, dorsal thorax and abdomen, legs (except empodia), anterior collophore, ventral and dorsal manubrium and dens ventrally.</p><p>Head. Antennae subequal to trunk length (Fig. 13); ratio antennae: trunk = 1: 1.04; antennal segments ratio as I: II: III: IV = 1: 1.57–1.82: 2.92–3.24: 2.97 (Ant IV present in one specimens). Ant IV annulated on distal one seventh, with apical bulb apically bilobed. Ant III annulated on distal two thirds, apical organ with 2 rods elongated, 3 spiny guard sens, s-blunt sens of different sizes and ciliated chaetae (Fig. 14A). Ant I dorsally with 3–4 median mac and 3 smooth mic at the base, without spines (Fig. 14B). Eyes A and B larger, G and H smaller, others subequal, and with 5 ‘IO’ chaetae; head dorsal macrochaetotaxy with 9–10 An, 4 A, 4 M, 8 S and 1 Pp mac (Fig. 14F). Clypeal formula with 4 (l1–2), 8 (ft), 5 (pf1–2) ciliated chaetae, l1–2 and pf0–1 larger, l1 apically acuminate, others subequal (Fig. 14C). Prelabral chaetae smooth, inner chaetae eventually with two small median filaments (Fig. 14D). Labral p0 chaeta with median filament unequal in length. Labral inner and outer papillae rounded, not projected (as in Fig. 19C). Maxillary palp with smooth t.a. and b.c. weakly ciliated, thicker and 1.19 longer than t.a. Labial papilla E with l.p. apically pointed and not reaching the base of a.a. (Fig. 14E). Basolateral and basomedian labial fields with chaetae M1–2, R (smaller than others) ciliated, e and l2 smooth, L1 ciliated or smooth. Ventral head with about 25 ciliated chaetae, postlabial formula with 4 (G1–4), 2 (H2, H4), 3 (J1–3) chaetae, 5 thin posterior chaetae with different lengths, and 3 larger chaetae, 1 b.c. surrounding the cephalic groove (Fig. 14G).</p><p>Thorax chaetotaxy (Figs 15 A–B). Th II not projected anteriorly, a, m and p series with 17, 9 and 15–16 mac, respectively. Th III a, m and p series with 4, 1 and 8 mac, respectively. Ratio Th II: III = 1.69–153: 1 (n=4).</p><p>Abdomen chaetotaxy (Figs 15 C–D, 16A–C). Abd I a, m and p series with 0–1, 5 and 0 mac, respectively. Abd II a, m and p series with 1, 4 and 0 mac, respectively. Abd III a, m and p series with 0–1, 3 and 1 mac, respectively. Abd IV with 10–11 central mac on A–T series and 11 lateral mac on E– Fe series; at least 12 posterior sens (ps type I, others type II) and 11 posterior mes. Abd V a, m and p series with 1, 4 and 6 mac, respectively. Ratio Abd III : IV = 1: 3.87–4.23 (n=4).</p><p>Legs. Subcoxa I with 5 chaetae and 3 psp; subcoxa II with an anterior row of 12 chaetae, posterior row of 7 chaetae, 1 posterior chaeta and 8 psp; subcoxa III with one row of 12 chaetae, 2 anterior chaetae and 2 posterior psp (Figs 17 A–C). Trochanteral organ with about 94 spine-like chaetae (Fig. 17D). Tibiotarsus normal or subdivided on distal two thirds, outer side with 3 large chaetae apically acuminate, inner side with 8 mac weakly ciliated. Unguis with 3–4 inner teeth, b.t. smaller than m.t. in length, and a minute a.t., present or absent. Unguiculus with all lamellae smooth and acuminate, except pe serrated and with a small proximal tooth; ratio unguis: unguiculus = 1: 0.52. Tibiotarsal smooth chaeta 1.32 larger than unguiculus and tenent hair 0.88 smaller than unguis outer edge.</p><p>Collophore. Anterior side with 19–22 chaetae, 4–5 proximal spine-like chaetae, 1 thin and 7–9 normal ciliated chaetae, 4 ciliated chaetae apically acuminate and 3 mac distally; posterior side with 12 chaetae, 8 thin ciliated chaetae widely distributed and 3 ciliated and 1 smooth chaetae distally; lateral flap with about 42–44 chaetae, 4–5 smooth and 38–39 ciliated (Fig. 17E).</p><p>Furcula. Manubrium ventral formula with 0, 0, 0, 2/2 (subapical), 24–28 (apical) ciliated chaetae plus approximately 18 elongated apical scales per side (Fig. 17F); manubrial plate with 3 ciliated mac apically acuminate (no blunt), 11 ciliated chaetae of different sizes and 3 psp. Dens dorsally with one proximal row of 13–19 blunt mac finely ciliated and apically nipple-like (Fig. 17G).</p><p>Remarks. Lepidocyrtinus barnardi resembles L. dayi in Th II normal (not projected anteriorly) and with anterior mac group sparce, prelabral chaetae mostly smooth, labral papillae rounded and discrete, dorsal chaetotaxy of Th II–III, and manubrial plate without blunt mac (Tables 3–4). However, L. barnardi differs from this species by color pattern with one transversal band on Abd III and one central spot on Abd IV, while in L. dayi most of the body is pigmented (Figs 13, 18). They also differ in head by basomedian labial field with chaeta e smooth (ciliated in L. dayi), labial papilla E with l.p. not reaching the base of a.a. (surpassing the base in L. dayi), and dorsal chaetotaxy with S4 mac and Pa3, Pm3 and Pp3 absent (opposite in L. dayi) (Figs 14 E–F, 19D–E). In dorsal chaetotaxy they differ by Th III with 10 central and 3 lateral mac (11 and 2 respectively in L. dayi), Abd I generally with 6 central mac (a2 present or absent, m4 present) and Abd II with 4 central mac, while in L. dayi there are 5 central mac (m4 absent) on Abd I and 3 central mac (m3e absent) on Abd II (Figs 15 B–D, 20B–D). Other features that differ are: unguis a.t. often present (absent in L. dayi), manubrium ventral formula with 0, 0, 0, 2/2 chaetae (1, 2, 2, 0/ 2 in L. dayi), and dens with 13–19 blunt mac (only 1 in L. dayi) (Figs 17 F–G, 22F–G).</p></div>	https://treatment.plazi.org/id/7B6687A7FFFBFF99FF12D569FB91F991	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cipola, Nikolas Gioia;Morais, José Wellington De;Bellini, Bruno Cavalcante	Cipola, Nikolas Gioia, Morais, José Wellington De, Bellini, Bruno Cavalcante (2020): Review of Lepidocyrtinus Börner, 1903 (Collembola, Entomobryidae, Seirinae): the African species. Zootaxa 4898 (1): 1-110, DOI: 10.11646/zootaxa.4898.1.1
7B6687A7FFF2FF9FFF12D660FAECFBAD.text	7B6687A7FFF2FF9FFF12D660FAECFBAD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lepidocyrtinus dayi (Yosii 1959)	<div><p>Lepidocyrtinus dayi (Yosii, 1959)</p><p>Figs 2, 18–22, Tables 1 and 3</p><p>Seira (Lepidocyrtinus) dayi Yosii, 1959: 14, Figs 7 A–G, South Africa, Cape Town, Skeleton Gorge.</p><p>Seira dayi; Coates 1968: 454 (combination); Janion-Scheepers et al. 2015: 69, Africa South (catalog).</p><p>Typological note. The type material of L. dayi stat. nov. (Scheepers, personal communication) is probably lost. Thus, here its neotype is designated, according to ICZN (2000, chapter 16, article 72.4.5.), since the specimen fits all morphological characteristics presented in the original description and was collected from the type locality (Fig. 2), Kirstenbosch, Cape Town.</p><p>Examined type material. Neotype female designed on slide (SAM / IZIKO): South Africa, Cape Town, Ta-ble Mountain National Park, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=18.42361&amp;materialsCitation.latitude=-33.981945" title="Search Plazi for locations around (long 18.42361/lat -33.981945)">Kirstenbosch</a>, 33°58’55.0”S, 18°25’25.0”E (Fig. 2), <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=18.42361&amp;materialsCitation.latitude=-33.981945" title="Search Plazi for locations around (long 18.42361/lat -33.981945)">Afrotemperate Forest</a>, 400 m, 12.ii.2009, Pitfall trap, C Uys coll.</p><p>Other examined material. 2 females in slides (INPA): same data as neotype . 1 female in slide and 1 specimens in alcohol (INPA): South Africa, Cape Town, Table Mountain National Park, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=18.3905&amp;materialsCitation.latitude=-34.003418" title="Search Plazi for locations around (long 18.3905/lat -34.003418)">Orange Kloof</a>, 34°00’12.3”S, 18°23’25.8”E, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=18.3905&amp;materialsCitation.latitude=-34.003418" title="Search Plazi for locations around (long 18.3905/lat -34.003418)">Afrotemperate Forest</a>, 130 m, 28.i.2009, Sugar-baited Ant trap, C Uys coll. 6 specimens in alcohol (CC / UFRN): <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=18.419722&amp;materialsCitation.latitude=-33.993584" title="Search Plazi for locations around (long 18.419722/lat -33.993584)">Cecilia-Rooikat</a>, 33°59’36.9”S, 18°25’11.0”E, in Fynbos, 430 m, 21.ii.2009, Sugar-baited Ant trap, C Uys coll. 4 females in slides and 11 specimens in alcohol (CC / UFRN): <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=18.402832&amp;materialsCitation.latitude=-34.065083" title="Search Plazi for locations around (long 18.402832/lat -34.065083)">Tokai</a>, 34°03’54.3”S, 18°24’10.2”E, in Pine Plantation, 300 m, 19.i.2009, Sugar-baited Ant trap, C Uys coll.</p><p>Diagnosis. Body all dark blue but depigmented in dorsal head, few trunk parts, tibiotasus and dens distally (Fig. 18); head mac A0, A2–3, A5, M1–2, S0–3, S5–6, Pa3, Pm3, Pp3 and Pp5 present, M4i as mac or mic; prelabral chaetae smooth; labial papilla E with l.p. apically pointed and surpassing the base of a.a. (Figs 19B, D–E); Basolateral and basomedian labial fields with chaetae R, E, L1 ciliated, R subequal or slightly smaller than the others, l2 smooth; Th II not projected anteriorly and with 32–34 anterior, 9 median (m1ip, m4i–4p present) and 18–20 posterior mac (p2ep, p2ep2, p3 mac); Th III–Abd II with 11, 5, 3 central mac respectively; Abd IV with 11–12 central mac and 14 lateral mac (Figs 20, 21B); unguis a.t. absent; unguiculus with pe lamella serrated and with small proximal tooth; collophore anteriorly with 5 spine-like chaetae and 3 ciliated chaetae apically acuminate, posteriorly without spines, smooth chaeta absent (Fig. 22E); manubrium ventrally with 0/2 subapical chaetae; manubrial plate without blunt mac; dens with 1 blunt mac apically nipple-like (Figs 22 F–G).</p><p>Description. Total length (head + trunk) of specimens 2.20–2.55 (n=4). Specimens with dark blue pigments all over the body, except for posterior dorsal head, trunk intersegments membranae, Abd IV anteriorly, tibiotasus, and collophore and dens distally; eyepatches black (Fig. 18). Scales present on Ant I to proximal half of Ant III, dorsal and ventral head, dorsal thorax and abdomen, legs (except empodia), anterior collophore, ventral and dorsal manubrium and dens ventrally.</p><p>Head. Antennae smaller to trunk length; ratio antennae: trunk = 1: 1.98 (n=1); antennal segments ratio as I: II: III: IV = 1: 1.42: 1.83: 2.14. Ant IV annulated on distal fourth-fifths, apical bulb apically bilobed. Ant III annulated on distal half, apical organ with 2 rods, 3 spiny guard sens, s-blunt sens of different sizes and ciliated chaetae. Ant I dorsally with 1 inner mac and 3 smooth mic at base, without spines (Fig. 19A). Eyes A, B and C larger, G and H smaller, others subequal, and with 5 ‘IO’ chaetae; head dorsal macrochaetotaxy with 9–10 An, 4 A, 3–4 M, 7 S, 1 Pa, 1 Pm and 2 Pp mac (Fig. 19E). Clypeal formula with 4 (l1–2), 4 (ft), 3 (pf0–1) ciliated chaetae, l1–2 larger than others and apically acuminate, 1 ft smaller, others subequal. Prelabral chaetae smooth and not bifurcated. Labral p0 chaeta generally with median filament (Fig. 19B). Labral inner and outer papillae rounded, not projected (Fig. 19C). Maxillary palp with smooth t.a. and b.c. weakly ciliated, thicker and 1.13 longer than the t.a. Labial papilla E with l.p. apically pointed and surpassing the base of a.a. (Fig. 19D). Basolateral and basomedian labial fields with chaetae M1–2, R, E, L1 ciliated, R subequal or slightly smaller than others, l2 smooth. Ventral head with about 30–31 ciliated chaetae, postlabial formula with 4 (G1–4), 3 (H2–4), 4 (J1–4) chaetae, about 14 thin posterior chaetae with different lengths, and 1 larger b.c. surrounding the cephalic groove (Fig. 19F).</p><p>Thorax chaetotaxy (Fig. 20 A–B). Th II not projected anteriorly, a, m and p series with 32–34, 9 and 18–20 mac, respectively. Th III a, m and p series with 4, 1 and 8 mac, respectively. Ratio Th II: III = 1.95–1.26: 1 (n=4).</p><p>Abdomen chaetotaxy (Figs 20 C–D, 21A–C). Abd I a, m and p series with 1, 4 and 0 mac, respectively. Abd II a, m and p series with 1, 3 and 0 mac, respectively, m3e absent. Abd III a, m and p series with 0, 3 and 1 mac, respectively. Abd IV with 11–12 central mac on A–T series and 14 lateral mac on E– Fe series; at least 9 sens (ps type I, others type II) and 10 posterior mes. Abd V a, m and p series with 1, 3 and 5 mac, respectively. Ratio Abd III : IV = 1: 4.26–5.25 (n= 4).</p><p>Legs. Subcoxa I with 5 chaetae, 1 anterior chaeta present or absent and 2 psp; subcoxa II with an anterior row of 11–12 chaetae, posterior row of 6 chaetae and 4 psp; subcoxa III with one row of 13 chaetae, 3 anterior chaetae and 2 posterior psp (Fig. 22 A–C). Trochanteral organ with about 38 spine-like chaetae (Fig. 22D). Tibiotarsus not subdivided, modified chaetae absent. Unguis with 3 inner teeth, b.t. subequal to m.t. in length, a.t. absent. Unguiculus with all lamellae smooth and acuminate, except pe serrated and with a small proximal tooth.</p><p>Collophore. Anterior side with 18–19 chaetae, 5 proximal spine-like chaetae, 1 thin and 7–8 normal ciliated chaetae, 3 ciliated chaetae apically acuminate and 2 mac distally; posterior side with 15 chaetae, 13 thin ciliated chaetae widely distributed and 2 ciliated chaetae distally; lateral flap with about 30–31 chaetae, 4–5 smooth and 26 ciliated (Fig. 22E).</p><p>Furcula. Manubrium ventral formula with 1, 2, 2, 0/2 (subapical), 12–20 (apical) ciliated chaetae plus approximately 16 elongated apical scales per side (Fig. 22F); manubrial plate with 5 mac ciliated and apically acuminate (no blunt), 7–11 ciliated chaetae of different sizes and 3 psp. Dens dorsally with 1 proximal blunt mac finely ciliated and apically nipple-like (Fig. 22G).</p><p>Remarks. Lepidocyrtinus dayi resembles L. barnardi (see the remarks of this species and Tables 3 and 4).</p></div>	https://treatment.plazi.org/id/7B6687A7FFF2FF9FFF12D660FAECFBAD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cipola, Nikolas Gioia;Morais, José Wellington De;Bellini, Bruno Cavalcante	Cipola, Nikolas Gioia, Morais, José Wellington De, Bellini, Bruno Cavalcante (2020): Review of Lepidocyrtinus Börner, 1903 (Collembola, Entomobryidae, Seirinae): the African species. Zootaxa 4898 (1): 1-110, DOI: 10.11646/zootaxa.4898.1.1
7B6687A7FFF4FFA6FF12D443FCEDFA8E.text	7B6687A7FFF4FFA6FF12D443FCEDFA8E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lepidocyrtinus africanus Cipola & Morais & Bellini 2020	<div><p>Lepidocyrtinus africanus sp. nov. Cipola &amp; Bellini</p><p>Figs 2, 23–31, Tables 1 and 3</p><p>Type Material. Holotype female on slide (INPA-CLL 0000110): South Africa, KwaZulu-Natal province, Ndumo Game Reserve, Ezulweni Hide near to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=32.310192&amp;materialsCitation.latitude=-26.886389" title="Search Plazi for locations around (long 32.310192/lat -26.886389)">Nyamiti lake</a>, 26°53’11.0”S, 32°18’36.7”E (Fig. 2), 35 m., 27.xi-3.xii.2014, Yellow Pan Trap, SL Winterton &amp; PH Kerr coll. 11 paratypes: 1 male, 2 females and 1 juvenile on slides and 2 specimens in alcohol (INPA-CLL 0000110), 1 female on slide (CC / UFRN), and 2 females and 2 specimen in alcohol (SAM / IZIKO), all with the same data of the holotype.</p><p>Diagnosis. Body with dark pigments on inner side of Ant II–III, all head and proximal legs; Abd III with one lateral spot in club-shape, and all Abd V to proximal manubrium (Fig. 23A); head mac A0, A2–3, A5, M1–2, M4i, S0–3, S5–6 and Pp5 present; prelabral chaetae ciliated; labial papilla E with l.p. apically pointed and reaching the base of a.a.; basolateral and basomedian labial fields with chaetae R (subequal to others), E, L1–2 ciliated (Figs 24 C–D, F–G); Th II projected anteriorly and with 14–15 anterior, 8 median (m1ip, m4–4p present, m4i mac absent) and 12 posterior mac (p2ep as mac, p3 as mic); Th III–Abd II with 9, 5, 4 central mac respectively; Abd IV with 16 central mac and 13 lateral mac, B6 mac modified (Figs 25–27); unguis a.t. present; unguiculus with pe lamella smooth and toothless (Fig. 28E); collophore anteriorly with 10 spine-like chaetae and 2 ciliated chaetae apically acuminate, posteriorly without spines, smooth chaeta absent (Fig. 29A); manubrium ventrally with 0/2 subapical chaetae; manubrial plate with 3 blunt mac abruptly acuminate at the apex; dens with 5–7 blunt mac apically bulletlike (Figs 29 B–C).</p><p>Description. Total length (head + trunk) of specimens 2.63–2.68 (n=4), holotype 2.68 mm. Specimens yellowish (except for tibiotarsus III and furcula part, pale white) with dark pigments on Ant I that extends in an inner strip to the Ant III distally, Ant IV (apex depigmented); head (more pigmented dorso-anteriorly laterally), one lateral spot in club-shape on Abd III, Abd V to proximal manubrium and coxae I–III; Abd IV posteriorly with vestigial ventrolateral pigments; eyepatches black (Fig. 23A). Scales present on Ant I to proximal one fifth of Ant III, dorsal and ventral head, dorsal thorax and abdomen, legs (except empodia), anterior collophore, ventral and dorsal manubrium and dens ventrally.</p><p>Head. Antennae subequal or slighty smaller than trunk length (Fig. 23A); ratio antennae: trunk = 1: 1.11–1.46 (n=3), holotype 1: 1.11; antennal segments ratio as I: II: III: IV = 1: 1.52–1.67: 2.9–3.17: 2.63–3.08, holotype 1: 1.54: 3.17: 2.63. Ant IV almost entirely annulated (except for a small proximal part) with apical bulb apically bilobed. Ant III annulated on distal three fourths, apical organ with 2 oval sens, 2 spiny guard sens, s-blunt sens of different sizes and ciliated chaetae (Fig. 24A). Ant I dorsally with 3 median mac and 3 smooth mic at the base, without spines (Fig. 24B). Eye A larger, G and H smaller, others subequal, and with 5 ‘IO’ chaetae; head dorsal macrochaetotaxy with 10–12 An, 4 A, 4 M, 7 S and 1 Pp mac (Fig. 24F). Clypeal formula with 4 (l1–2), 2 (ft), 3 (pf0–1) ciliated chaetae, l1–2 larger than the others, others subequal (Fig. 24C). Prelabral chaetae ciliated and not bifurcated. Labral p0–1 chaeta larger than the others, others subequal, p0 without filament (Fig. 24C). Labral papillae with two inner rounded projections, outer papillae absent (Fig. 24C). Maxillary palp with t.a. and b.c. both smooth, b.c. 1.09 larger than t.a. (Fig. 24E). Labial papilla E with l.p. apically pointed and almost reaching the base of a.a. (Fig. 24D). Basolateral and basomedian labial fields with chaetae M1–2, R, E, L1–2 ciliated, R subequal to others (Fig. 24G). Ventral head with about 20 ciliated chaetae, postlabial formula with 4 (G1–4), 3 (H2–4), 3 (J1–3) chaetae, 1 thin posterior chaeta and 2 larger chaetae, 1 b.c. surrounding the cephalic groove (Fig. 24G).</p><p>Thorax chaetotaxy (Fig. 25 A–B). Th II projected anteriorly, a, m and p series with 14–15, 8 and 12 mac, respectively. Th III a, m and p series with 2, 0 and 8 mac, respectively. Ratio Th II: III = 2.24–1.50: 1 (n=4), holotype 1.53: 1.</p><p>Abdomen chaetotaxy (Figs 25 C–D, 26–27). Abd I a, m and p series with 1, 4 and 0 mac, respectively, m4 absent. Abd II a, m and p series with 1, 4 and 0 mac, respectively. Abd III a, m and p series with 0, 3 and 1 mac, respectively. Abd IV with 16 central mac on A–T series and 13 lateral mac on E–Fe series, B6 mac modified; at least 4 posterior sens (ps type I, others type II) and 8 posterior mes. Abd V a, m and pa / p series with 1, 4 and 3/6 mac, respectively. Ratio Abd III: IV = 1: 3.12–4.74 (n= 4), holotype 1: 3.92.</p><p>Legs. Subcoxa I with 4–5 chaetae on a row and 2 psp; subcoxa II with an anterior row of 8 chaetae, posterior row of 5–6 chaetae and 4 psp; subcoxa III with one row of 8–9 chaetae, 0–1 posterior chaeta and 2 posterior psp (Fig. 28 A–C). Trochanteral organ with about 27 spine-like chaetae (Fig. 28D). Tibiotarsus subdivided or not, outer side with 4 proximal larger ciliated chaetae apically acuminate, inner side with 6 mac weakly ciliated, anterior side with 2 mac on proximal half. Unguis with 4 inner teeth, b.t. subequal to m.t. in length, a.t. smaller. Unguiculus with all lamellae smooth and acuminate; ratio unguis: unguiculus = 1: 0.56. Tibiotarsal smooth chaeta 1.54 larger than unguiculus and tenent hair 0.86 smaller than unguis outer edge (Fig. 28E).</p><p>Collophore. Anterior side with 16 chaetae, 10 spine-like chaetae, 1 proximal ciliated chaeta, 3 ciliated chaetae apically acuminate and 2 mac distally; posterior side with 10 ciliated chaetae (2 unpaired), 9 thin widely distributed and 1 thicker distally; lateral flap with about 29 chaetae, 4 smooth and 25 ciliated (Fig. 29A).</p><p>Furcula. Manubrium ventral formula with 1, 0, 0, 0/2 (subapical), 16–18 (apical) ciliated chaetae plus approximately 13 elongated apical scales per side (Fig. 29B); manubrial plate with 3 blunt mac ciliated and abruptly acuminate at apex (1 smaller), 5 ciliated chaetae of different sizes and 3 psp. Dens dorsally with one proximal row of 5–7 blunt mac weakly ciliated and apically bullet-like, holotype with 7 (Fig. 29C).</p><p>Etymology. Refers to the continent where the species was found, Africa.</p><p>Remarks. Lepidocyrtinus africanus sp. nov. resembles L. annulicornis, L. botswanensis sp. nov. and L. capensis Womersley, 1934 by Ant III–IV annulated, prelabral chaetae ciliated (at least in L. botswanensis), Th II with 6 median mac in m1–2 complex, and PmB group with 3 mac, Th III to Abd II with 9, 5 and 4 central mac, respectively, unguis with 4 inner teth, and furcula with modified mac (see Table 2). However, L. africanus differs from these species by Abd III with one lateral spot in club-shape and Abd V to proximal manubrium completely pigmented, while in L. annulicornis there are violet pigments on femur III distally, in L. botswanensis the posterior head and coxae are weakly pigmented (Fig. 32), and in L. capensis there are dark pigments on head and Th II anteriorly (see Womersley 1934: 462). Concerning the chaetotaxy, L. africanus differs by Th II with 7 mac in PmA group (6 in L. annulicornis and L. botswanensis) and 2 in PmC group (4 in L. capensis), Th III with 9 central mac (8 in L. capensis), and Abd IV with B6 mac modified (Fig. 27), a feature never reported before to any species of Lepidocyrtinus or even other Seirinae . Finally, L. africanus also differs by unguiculus pe lamella smooth and toothless (serrated in L. annulicornis and L. botswanensis and with a proximal tooth in L. botswanensis and L. capensis), and manubrial plate and dens respectively with 3 and 5–7 blunt mac (less in the other species).</p></div>	https://treatment.plazi.org/id/7B6687A7FFF4FFA6FF12D443FCEDFA8E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cipola, Nikolas Gioia;Morais, José Wellington De;Bellini, Bruno Cavalcante	Cipola, Nikolas Gioia, Morais, José Wellington De, Bellini, Bruno Cavalcante (2020): Review of Lepidocyrtinus Börner, 1903 (Collembola, Entomobryidae, Seirinae): the African species. Zootaxa 4898 (1): 1-110, DOI: 10.11646/zootaxa.4898.1.1
7B6687A7FFCDFFAEFF12D59CFBC8FE49.text	7B6687A7FFCDFFAEFF12D59CFBC8FE49.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lepidocyrtinus botswanensis Cipola & Morais & Bellini 2020	<div><p>Lepidocyrtinus botswanensis sp. nov. Cipola &amp; Bellini</p><p>Figs 2, 32–38, Tables 1 and 3</p><p>Type Material. Holotype female on slide (NHMLA): Botswana, South-East District, Lobatse, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=25.1&amp;materialsCitation.latitude=-25.166666" title="Search Plazi for locations around (long 25.1/lat -25.166666)">Moroekwe</a> farm, 25°10’S; 25°06’E (Fig. 2), 1.240 m, 8–22.xi.2005, Malaise trap, M Bing coll. 4 paratypes: 2 females on slides and 1 specimen in alcohol (NHMLA), and 1 female on slide (INPA-CLL 000034), same data as holotype.</p><p>Diagnosis. Body with purple pigments on Ant II internally to IV and dark blue on coxa I (Fig. 32); head mac A0, A2–3, A5, M1–2, M4i, S0–3, S5–6 and Pp5 present; prelabral chaetae ciliated; labial papilla E with l.p. apically pointed and surpassing the base of a.a; basolateral and basomedian labial fields with chaetae R (smaller than others), E, L1–2 ciliated (Figs 33 C–D, 34A and C); Th II projected anteriorly and with 13–14 anterior, 8 median (m1ip, m4-4p present, m4i mac absent) and 11 posterior mac (p2ep as mac, p3 as mic); Th III–Abd II with 9, 5, 4 central mac respectively; Abd IV with 17 central mac and 13 lateral mac (Figs 35, 36B); unguis a.t. present; unguiculus pe lamella serrated and with a small proximal tooth (Fig. 37F); collophore anteriorly with 7 spine-like chaetae and 4 ciliated chaetae apically acuminate, posteriorly without spines, smooth chaeta present (Fig. 38A); manubrium ventrally with 0/0 subapical chaetae; manubrial plate with 1 blunt mac abruptly acuminate at the apex; dens with 3–4 blunt mac apically bullet-like (Figs 38 B–C).</p><p>Description. Total length (head + trunk) of specimens 1.83–2.62 (n=4), holotype 2.61 mm. Specimens pale white with purple pigments on Ant IV, internal side of the Ant II–III, and weakly present on anterior and posterior head, and basal part of tibiotarsus I. Pigments dark blue intensely on coxa I, and weakly on coxa II–III and Th II; eyepatches black (Fig. 32). Scales present on Ant I to proximal one quarter of Ant III, proximal Ant IV, dorsal and ventral head, dorsal thorax and abdomen, legs (except empodia), anterior and posterior collophore, ventral and dorsal manubrium and dens ventrally.</p><p>Head. Antennae smaller than trunk length (Fig. 32); ratio antennae: trunk = 1: 1.35–1.84 (n=4), holotype 1: 1.40; antennal segments ratio as I: II: III: IV = 1: 1.43–1.59: 2.2–2.44: 2.38–3.34, holotype 1: 1.43: 2.2: 2.38. Ant IV completely annulated with apical bulb apically unilobed or bilobed. Ant III annulated on distal three fourths, apical organ with 2 club-shaped sens, 3 spiny guard sens, s-blunt sens of different sizes and ciliated chaetae (Fig. 33A). Ant I dorsally with 3 median mac and 3 smooth mic at the base, apparently without spines (Fig. 33B). Eyes A and B larger, G and H smaller, others subequal, and with 5 ‘IO’ chaetae; head dorsal macrochaetotaxy with 10 An, 4 A, 4 M, 7 S and 1 Pp mac. (Fig. 33D). Clypeal formula with 4 (l1–2), 3 (ft), 3 (pf0–1) ciliated chaetae, l1–2 larger than others, l1 apically acuminate, others subequal (Fig. 33C). Prelabral chaetae ciliated and not bifurcated. Labral p0 chaeta without median filament, p0–1 larger than others. Labral papillae with two inner conical projections, outer papillae absent (Fig. 33C). Maxillary palp with smooth t.a and b.c. finely ciliated, thicker and 1.09 longer than the t.a. (Figs. 34B). Labial papilla E with l.p. apically pointed and surpassing the base of a.a. (Fig. 34A). Basolateral and basomedian labial fields with chaetae M1–2, R (smaller than others), E, L1–2 ciliated (Fig. 34C). Ventral head with about 22 ciliated chaetae, postlabial formula with 4 (G1–4), 3 (H2–4), 4 (J1–4) chaetae, 5 thin posterior chaetae, and 3 larger chaetae, 1 b.c. surrounding the cephalic groove (Fig. 34C).</p><p>Thorax chaetotaxy (Fig. 35 A–B). Th II projected anteriorly, a, m and p series with 13–14, 8 and 11 mac, respectively. Th III a, m and p series with 2, 0 and 8 mac, respectively. Ratio Th II: III = 1.90–1.40: 1 (n=4), holotype 1.40: 1.</p><p>Abdomen chaetotaxy (Figs 35 C–D, 36A–C). Abd I a, m and p series with 1, 4 and 0 mac, respectively, m4 absent. Abd II a, m and p series with 1, 4 and 0 mac, respectively. Abd III a, m and p series with 0, 2–3 and 0–1 mac, respectively. Abd IV with 17 central mac on A– T series and 13 lateral mac on E– Fe series; at least 2 posterior sens (ps type I, other type II) and 6 posterior mes. Abd V a, m and pa / p series with 1, 4 and 1/5 mac, respectively. Ratio Abd III: IV = 1: 3.53–4.48 (n= 4), holotype 1: 4.14.</p><p>Legs. Subcoxa I with 5 chaetae on a row and 2 psp; subcoxa II with an anterior row of 9 chaetae, posterior row of 4 chaetae, 2 extra chaetae near p row and 6 psp; subcoxa III with one row of 9 chaetae and 2 posterior psp (Figs 37 A–C). Trochanteral organ with about 23 spine-like chaetae (Fig. 37D). Tibiotarsus not subdivided, outer side with 3 proximal larger ciliated chaetae apically acuminate (e), inner side with 9 mac weakly ciliated (ai and pi), and anterior side with 2 mac (ae) on proximal half (Fig. 37E). Unguis with 4 inner teeth, b.t. subequal to m.t. in length, a.t. smaller. Unguiculus with all lamellae smooth and acuminate, except pe serrated and with a small proximal tooth; ratio unguis: unguiculus = 1: 0.57. Tibiotarsal smooth chaeta 1.08 larger than unguiculus and tenent hair 0.90 smaller than unguis outer edge (Fig. 37F).</p><p>Collophore. Anterior side with 14 chaetae, 7 proximal spine-like chaetae, 1 ciliated chaeta, 4 ciliated chaetae apically acuminate and 2 mac distally; posterior side with 5 chaetae, 4 thin ciliated chaetae widely distributed and 1 smooth chaeta distally; lateral flap with about 19 chaetae, 5 smooth and 14 ciliated (Fig. 38A).</p><p>Furcula. Manubrium ventral formula with 1, 0, 0, 0/0 (subapical), 14 (apical) ciliated chaetae plus approximately 15 elongated apical scales per side (Fig. 38B); manubrial plate with 1 blunt mac ciliated abruptly acuminate at the apex, 5 ciliated chaetae of different sizes and 2 psp. Dens dorsally with one proximal row of 3–4 blunt mac weakly ciliated and apically bullet-like, holotype with 3 (Fig. 38C).</p><p>Etymology. Refers to the type locality of the new species, Botswana.</p><p>Remarks. Lepidocyrtinus botswanensis sp. nov. resembles L. annulicornis in antennae subequal to trunk length, Th II with 6 median mac in m1–2 complex and 11 posterior mac, Th III to Abd II with 9, 5 and 4 central mac, respectively, and unguiculus pe lamellae serrated. However, L. botswanensis differs from this species by color pattern (see remarks of L. africanus), unguiculus pe lamella with a small proximal tooth (absent in L. annulicornis) and manubrial plate with 1 mac abruptly acuminate at apex (2–3 in L. annulicornis).</p></div>	https://treatment.plazi.org/id/7B6687A7FFCDFFAEFF12D59CFBC8FE49	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cipola, Nikolas Gioia;Morais, José Wellington De;Bellini, Bruno Cavalcante	Cipola, Nikolas Gioia, Morais, José Wellington De, Bellini, Bruno Cavalcante (2020): Review of Lepidocyrtinus Börner, 1903 (Collembola, Entomobryidae, Seirinae): the African species. Zootaxa 4898 (1): 1-110, DOI: 10.11646/zootaxa.4898.1.1
7B6687A7FFC5FFB1FF12D258FBD0F8DD.text	7B6687A7FFC5FFB1FF12D258FBD0F8DD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lepidocyrtinus fabioi Cipola & Morais & Bellini 2020	<div><p>Lepidocyrtinus fabioi sp. nov. Cipola &amp; Bellini</p><p>Figs 2, 39–44, Tables 1–2</p><p>Type Material. Holotype female on slide (NHMLA): Republic of the Congo, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=15.466666&amp;materialsCitation.latitude=-3.2666667" title="Search Plazi for locations around (long 15.466666/lat -3.2666667)">Poll Departament</a>, Nagbe, LesioLouna Reserve, 03°16’S, 15°28’E (Fig. 2), 364 m., 18.viii.2008, Malaise trap, B Brown coll. 7 paratypes: 2 females on slides (NHMLA and INPA-CLL 000035), same data as holotype; 1 male, 2 females and 2 juveniles on slides (NHMLA), same data as holotype, except 25.viii.2008.</p><p>Diagnosis. Body with dark bands on proximal Ant IV and femur III distally (Fig. 39); head mac A0, A2, A5, M1–2, S3, S5, S6 and Pp5 present; prelabral chaetae ciliated; labial papilla E with l.p. apically finger-shaped and surpassing the base of a.a. (Figs 40 D–E); basolateral and basomedian labial fields with chaetae E, L1–2 ciliated and r reduced in spine; Th II projected anteriorly and with 5–8 anterior, 5–6 median (m4–4p present, m1ip, m2i2 and m4i mac absent) and 4–5 posterior mac (p2ep as mic, p3 as mac); Th III–Abd II with 4, 1, 3 central mac respectively; Abd IV with 10 central mac and 13–14 lateral mac (Figs 41, 42B); unguis a.t. present; unguiculus pe lamella serrated and with a small proximal tooth; collophore anteriorly with 8 spine-like chaetae and 3 ciliated chaetae apically acuminate, posteriorly with 5 spines, smooth chaeta present (Fig. 43E); manubrium ventrally with 2/4 subapical chaetae; manubrial plate with 2 blunt mac abruptly acuminate at the apex; dens without blunt mac (Figs 43 F–G).</p><p>Description. Total length of body (head + trunk) 2.02–3.05 mm (n=4), holotype 2.92 mm. Specimens pale white with dark blue pigments on proximal Ant IV, distal and proximal Ant I, femur II–III distally and proximal tibiotarsus; eyepatches and basal Ant I dark (Fig. 39). Scales present on Ant I to proximal one fifth of Ant IV, ventral and dorsal head, dorsal trunk, legs (except empodium), anterior and posterior collophore, ventral and dorsal manubrium and dens ventrally.</p><p>Head. Antennae smaller than the trunk length (Fig. 39); ratio antennae: trunk = 1: 1.33–1.63 (n=2); antennal segments ratio as I: II: III: IV = 1: 1.23–1.5: 1.20–1.39: 2.01–2.48 (present in two paratypes), holotype 1: 1.25: 1.30: missing. Ant IV annulated on distal three quarters, with apical bulb apically unilobed. Ant III not annulated, apical organ with 2 club-shaped sens curved, 3 spiny guard sens, s-blunt sens of different sizes and ciliated chaetae (Fig. 40A). Ant I dorsally with 4 median mac and 6 smooth mic at the base, without spines (Fig. 40B). Eyes A and B larger, G and H smaller, others subequal, and with 5 ‘IO’ chaetae; head dorsal macrochaetotaxy with 8–9 An, 3 A, 3 M, 3 S, 1 Ps and 1 Pp mac. (Fig. 40E). Clypeal formula with 4 (l1–2), 2 (ft), 3 (pf0–1) ciliated chaetae, l1–2 larger than the others, l1 apically acuminate, others subequal (Fig. 40C). Prelabral chaetae ciliated and not bifurcated. Labral p0 chaeta without median filament. Labral papillae with two inner conical projections, outer papillae absent. Labial papilla E with l.p. apically finger-shaped and surpassing the base of a.a. (Fig. 40D). Basolateral and basomedian labial fields with chaetae M1–2, E, L1–2 ciliated, r reduced in spine. Ventral head with about 14 ciliated chaetae, postlabial formula with 4 (G1–4), 3 (H2–4), 4 (J1–4) chaetae, and 2 larger chaetae, 1 b.c. surrounding the cephalic groove (Fig. 40F).</p><p>Thorax chaetotaxy (Fig. 41 A–B). Th II projected anteriorly, a, m and p series with 5–8, 5–6 and 4–5 mac, respectively. Th III a, m and p series with 0, 1 and 4 mac, respectively. Ratio Th II: III = 1.90–1.66: 1 (n=4), holotype 1.68: 1.</p><p>Abdomen chaetotaxy (Figs 41 C–D, 42A–C). Abd I a, m and p series with 0, 1 and 0 mac, respectively, m2i and m4i absent. Abd II a, m and p series with 1, 3 and 0 mac, respectively, m3ep as mic. Abd III a, m and p series with 0, 3 and 2 mac, respectively. Abd IV with 10 central mac on A– T series and 13–14 lateral mac on E– Fe series; at least 9 posterior sens (ps type I, others type II) and 8 posterior mes. Abd V a, m and p series with 1, 4 and 6 mac, respectively. Ratio Abd III: IV = 1: 3.31–4.82 (n= 4), holotype 1: 4.82.</p><p>Legs. Subcoxa I with 6 chaetae on a row and 2 psp; subcoxa II with an anterior row of 10 chaetae, posterior row of 2 chaetae and 4 psp; subcoxa III with one row of 8 chaetae and 2 posterior psp (Fig. 43 A–C). Trochanteral organ with about 37 spine-like chaetae (Fig. 43D). Tibiotarsus subdivided on distal two thirds, outer side with 2 proximal larger ciliated chaetae apically acuminate, inner side with 7 mac weakly ciliated. Unguis with 4 inner teeth, b.t. subequal to m.t. in length, a.t. smaller. Unguiculus with all lamellae smooth and acuminate, except pe serrated and with a small proximal tooth; ratio unguis: unguiculus = 1: 0.64. Tibiotarsus smooth chaetae 0.86 smaller than unguiculus; tenent hair 0.91 smaller than unguis outer edge.</p><p>Collophore. Anterior side with 14 chaetae, 8 proximal spine-like chaetae, 1 ciliated chaeta, 3 ciliated chaetae apically acuminate and 2 mac distally; posterior side with 9 chaetae, 5 spines, 3 thin ciliated chaetae and 1 smooth chaeta distally; lateral flap with about 17 chaetae, 5 smooth and 12 ciliated (Fig. 43E).</p><p>Furcula. Manubrium ventral formula with 1, 2 2, 2/4 (subapical), 12 (apical) ciliated chaetae of different sizes plus approximately 11 elongated apical scales per side (Fig. 43F); manubrial plate with 2 blunt mac ciliated abruptly acuminate at the apex, 4–5 ciliated chaetae of different sizes and 3 psp. Dens dorsally without blunt mac (Fig. 43G).</p><p>Etymology. It is named after our friend, the Collembola researcher Dr. Fabio Gonçalves de Lima Oliveira (USP/RP, Brazil), who helped with this study providing specimens from Africa.</p><p>Remarks. Lepidocyrtinus fabioi sp. nov. resembles especially L. rykei (Coates 1968) comb. nov. in reduced macrochaetotaxy, as Th II with 4 mac in m1–2 complex, and Abd I–II with 1 and 3 central mac, respectively (Figs 41A, C–D). However, L. fabioi differs from this species by Ant III not annulated (annulated in L. rykei), Th II with 4–5 posterior mac (8 in L. rykei), Th III with 4 central mac (6 in L. rykei), and dens devoided of blunt mac (present in L. rykei). Other differences among similar species to L. fabioi are listed in Table 2.</p></div>	https://treatment.plazi.org/id/7B6687A7FFC5FFB1FF12D258FBD0F8DD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cipola, Nikolas Gioia;Morais, José Wellington De;Bellini, Bruno Cavalcante	Cipola, Nikolas Gioia, Morais, José Wellington De, Bellini, Bruno Cavalcante (2020): Review of Lepidocyrtinus Börner, 1903 (Collembola, Entomobryidae, Seirinae): the African species. Zootaxa 4898 (1): 1-110, DOI: 10.11646/zootaxa.4898.1.1
7B6687A7FFDCFFBDFF12D6D7FD46FB1D.text	7B6687A7FFDCFFBDFF12D6D7FD46FB1D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lepidocyrtinus paucisensillum Cipola & Morais & Bellini 2020	<div><p>Lepidocyrtinus paucisensillum sp. nov. Cipola &amp; Bellini</p><p>Figs 2, 45–49, Tables 1 and 3</p><p>Type Material. Holotype female on slide (NHMLA): Republic of the Congo, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=15.466666&amp;materialsCitation.latitude=-3.2666667" title="Search Plazi for locations around (long 15.466666/lat -3.2666667)">Poll Departament</a>, Nagbe, LesioLouna Reserve, 03°16’S, 15°28’E (Fig. 2), 18.viii.2008, Malaise trap, B Brown coll. 3 paratypes on slides: 1 male and 1 female (NHMLA), plus 1 female (INPA-CLL 000036), same data as holotype.</p><p>Diagnosis. Body with dark blue pigments on Ant II–III internally and posterior head to coxa I and Th II laterally (Fig. 45); head mac A0, A2–3, A5, M1–2, S0–3, S5–6 and Pp5 present; prelabral chaetae ciliated; labial papilla E with l.p. apically finger-shaped and reaching the base of a.a. (Figs 46 D–E); basolateral and basomedian labial fields with chaetae R (subequal to others), E, L1–2 ciliated; Th II projected anteriorly and with 13–14 anterior, 7 median (m4–4p present, m1ip and m4i mac absent) and 11 posterior mac (p2ep mac, p3 mic); Th III–Abd II with 8–9, 5, 3–4 central mac respectively; Abd IV with 16–18 central mac and 12–13 lateral mac (Figs 47, 48B); unguis a.t. present; unguiculus pe lamella serrated and with a small proximal tooth; collophore anteriorly with 12 spine-like chaetae and 4 ciliated chaetae apically acuminate, posteriorly without spines, smooth chaeta present (Fig. 49E); manubrium ventrally with 2/2 subapical chaetae; manubrial plate with 3 blunt mac abruptly acuminate at the apex; dens with 9 blunt mac apically nipple-like (Figs 49 F–G).</p><p>Description. Total length of body (head + trunk) 2.44–4.32 mm (n=4), holotype 4.32 mm. Specimen pale white with dark blue pigments present on inner side of the Ant I–III forming a longitudinal strip, posterolateral dorsal head, Th II lateral margin and all coxa I. Light blue pigments present on Th III laterally and femur III distally; eyepatches dark (Fig. 45). Scales present on Ant I to proximal half of Ant III, ventral and dorsal head, dorsal thorax and abdomen, legs (except empodium), anterior collophore, ventral and dorsal manubrium and dens ventrally.</p><p>Head. Ant IV missing in all specimens, antennae probably subequal or smaller than the trunk length (Fig. 45); ratio antennae: trunk = 1: 1.56–2.23 (n=3), holotype 1: 2.23 (Ant IV missing); antennal segments ratio as I: II: III = 1: 1.25–1.37: 2.26–2.65, holotype 1: 1.29: 2.28. Ant III annulated on distal three fourths, apical organ with 2 clubshaped sens, 3 spiny guard sens and s-blunt sens of different sizes (Fig. 46A). Ant I dorsally with 5 median mac and 5 smooth mic at base, without spines (Fig. 46B). Eyes A and B slightly larger, G and H smaller, others subequal, and with 5 ‘IO’ chaetae; head dorsal macrochaetotaxy with 10 An, 4 A, 3 M, 7 S, 1 Ps and 1 Pp mac (Fig. 46E). Clypeal formula with 4 (l1–2), 2 (ft), 3 (pf0–1) ciliated chaetae, l1–2 larger than others, l1 apically acuminate, others subequal (Fig. 46C). Prelabral chaetae ciliated and not bifurcated. Labral p0–1 larger than the others, p0 without median filament. Labral papillae with two inner conical projections, outer papillae absent. Maxillary palp with smooth t.a. and b.c. weakly ciliated, thicker and 1.14 longer than the t.a. Labial papilla E with l.p. finger-shaped and reaching the base of a.a. (Fig. 46D). Basolateral and basomedian labial fields with chaetae M1–2, R (subequal to others), E, L1–2 ciliated. Ventral head with about 23 ciliated chaetae, postlabial formula with 4 (G1–4), 3 (H2–4), 3 (J1–3) chaetae (H2 and J2 larger than the others), 7 thin posterior chaetae, and 3 larger chaetae, 1 b.c. surrounding the cephalic groove (Fig. 46F).</p><p>Thorax chaetotaxy (Fig. 47 A–B). Th II projected anteriorly, a, m and p series with 13–14, 7 and 11 mac, respectively. Th III a, m and p series with 2, 1 and 7–8 mac, respectively. Ratio Th II: III = 1.81–1.69: 1 (n=4), holotype 1.81: 1.</p><p>Abdomen chaetotaxy (Figs 47 C–D, 48A–C). Abd I a, m and p series with 1, 4 and 0 mac, respectively. Abd II a, m and p series with 1, 3–4 and 0 mac, respectively. Abd III a, m and p series with 0, 3 and 1 mac, respectively. Abd IV with 16–18 central mac on A– T series and 12–13 lateral mac on E– Fe series; at least 4 posterior sens (ps type I, others type II) and 9 posterior mes. Abd V a, m and p series with 1, 4 and 6 mac, respectively. Ratio Abd III: IV = 1: 3.64–4.98 (n= 4), holotype 1: 4.98.</p><p>Legs. Subcoxa I with 7 chaetae on a row and 2 psp; subcoxa II with an anterior row of 13 chaetae, posterior row of 5 chaetae, plus 1 extra chaeta and 7 psp; subcoxa III with one row of 12 chaetae and 2 posterior psp (Fig. 49 A–C). Trochanteral organ with about 63 spine-like chaetae (Fig. 49D). Tibiotarsus sometimes subdivided on distal two thirds, outer side with 4 larger ciliated chaetae apically acuminate, inner side with 6 mac weakly ciliated. Unguis with 4 inner teeth, b.t. subequal to m.t. in length, a.t. smaller. Unguiculus with all lamellae smooth and acuminate, except pe serrated and with a small proximal tooth; ratio unguis: unguiculus = 1: 0.65. Tibiotarsal smooth chaeta 0.83 smaller than unguiculus and tenent hair 0.86 smaller than unguis outer edge.</p><p>Collophore. Anterior side with 22 chaetae, 12 proximal spine-like chaetae, 3 ciliated chaetae, 5 ciliated chaetae apically acuminate and 2 mac distally; posterior side with 11 chaetae, 9 thin ciliated chaetae widely distributed and 1 ciliated and 1 smooth chaetae distally; lateral flap with about 42 chaetae, 11 smooth and 31 ciliated (Fig. 49E).</p><p>Furcula. Manubrium ventral formula with 1, 2 2, 2/2 (subapical), 2/16 (apical) ciliated chaetae of different sizes plus approximately 15 elongated apical scales per side (Fig. 49F); manubrial plate with 3 blunt mac weakly ciliated abruptly acuminate at the apex, 6 ciliated chaetae of different sizes and 3 psp. Dens dorsally with 9 blunt mac finely ciliated and apically nipple-like (Fig. 49G).</p><p>Etymology. The name of new species, paucisensillum, refers to the few (from Latin: pauci) sensilla seen in Abd IV.</p><p>Remarks. Lepidocyrtinus paucisensillum sp. nov. resembles L. ryozoi nom. nov. in Th II with 5 central mac in m1–2 group, Th III to Abd II with 9, 5, 4 central mac, respectively, unguis a.t. present and dens with at least 9 blunt mac (Table 3). However, L. paucisensillum differs from this species by dark blue pigments on Ant II–III internally and on posterior head to coxa I and Th II laterally (Fig. 45), while in L. ryozoi there are pigments on all Ant III–IV, and distally on femur and tibiotarsus I–III (see Yosii 1959). In chaetotaxy they differ by Th II with p3 mic and Th III without p1p2 mac (opposite in L. ryozoi), in Abd IV with 6–7 (anterior), 4 (median) and 6–7 (posterior) central mac (8, 5 and 4 in L. ryozoi), and manubrial plate with 3 blunt mac abruptly acuminate at the apex (apically nipple-like in L. ryozoi) (Figs 12 A–B, D, 47A–B, 48B, 49G).</p></div>	https://treatment.plazi.org/id/7B6687A7FFDCFFBDFF12D6D7FD46FB1D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cipola, Nikolas Gioia;Morais, José Wellington De;Bellini, Bruno Cavalcante	Cipola, Nikolas Gioia, Morais, José Wellington De, Bellini, Bruno Cavalcante (2020): Review of Lepidocyrtinus Börner, 1903 (Collembola, Entomobryidae, Seirinae): the African species. Zootaxa 4898 (1): 1-110, DOI: 10.11646/zootaxa.4898.1.1
7B6687A7FFD6FFC4FF12D4ECFE0FFA54.text	7B6687A7FFD6FFC4FF12D4ECFE0FFA54.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lepidocyrtinus scheepersae Cipola & Morais & Bellini 2020	<div><p>Lepidocyrtinus scheepersae sp. nov. Cipola &amp; Bellini</p><p>Figs 2, 50–54, Tables 1 and 4</p><p>Type Material. Holotype female on slide (SAM / IZIKO: SAM-ENW-C009786): South Africa, Cape Town, Somerset West, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=18.872223&amp;materialsCitation.latitude=-34.06111" title="Search Plazi for locations around (long 18.872223/lat -34.06111)">Helderberg Nature Reserve</a>, 34°03’40’’S, 18°52’20’’E (Fig. 2), <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=18.872223&amp;materialsCitation.latitude=-34.06111" title="Search Plazi for locations around (long 18.872223/lat -34.06111)">Southern Afrotemperate Forest</a>, 142 m, 13.xi.2017, beating and aspirating, A. Liu coll. 22 paratypes: 2 females on slides and 5 specimens in alcohol (SAM / IZIKO: SAM-ENW-C009786); 2 females on slides (SAM / IZIKO: SAM-ENW-C009786 donated to INPA- CLL 000033), same date as holotype; 2 females and 1 juvenile in slides and 8 specimens in alcohol (SAM / IZIKO: SAM-ENW-C009785), same data as holotype, except 17.x.2017.</p><p>Diagnosis. Body with dark bands on distal and proximal Ant III–IV and femur III distally (Fig. 50); head mac A0, A2–3, A5, M1–2, M4i and S0–6 present; prelabral chaetae ciliated; labial papilla E with l.p. apically fingershaped and not reaching the base of a.a. (Figs 51 C–D); basolateral and basomedian labial fields with chaetae R (subequal to others), E, L1–2 ciliated; Th II projected anteriorly and with 12–14 anterior, 8 median (m1ip, m4–4p present, m4i mac absent) and 12–13 posterior mac (p2ep and p3 as mac); Th III–Abd II with 11, 6, 4 central mac respectively; Abd IV with 16 central mac and 15–17 lateral mac (Figs 52, 53B); unguis a.t. present; unguiculus pe lamella serrated and with a small proximal tooth; collophore anteriorly with 11 spine-like chaetae and 4 ciliated chaetae apically acuminate, posteriorly without spines, smooth chaeta present (Fig. 54E); manubrium ventrally with 2/2 subapical chaetae; manubrial plate with 3 blunt mac apically nipple-like; dens with 11–13 blunt mac apically nipple-like (Figs 54 F–G).</p><p>Description. Total length of body (head + trunk) 3.25–3.71 mm (n=4), holotype 3.71 mm. Specimens pale white with dark brownish to bluish pigments on distal and proximal Ant III–IV, distal Ant II, anterior and posterior regions of eyepatches, and dark blue pigments on coxa I and tibiotarsus I–III distally; eyepatches dark (Fig. 50). Scales present on Ant I to proximal two thirds of Ant III, proximal one quarter of Ant IV, ventral and dorsal head, dorsal trunk, legs (except empodium), anterior collophore, ventral and dorsal manubrium and dens ventrally.</p><p>Head. Antennae longer than the trunk length (Fig. 50); ratio antennae: trunk = 1: 0.84–0.87 (n=4), holotype 1: 0.84; antennal segments ratio as I: II: III: IV = 1: 1.31–1.52: 2.29–2.74: 2.97–3.18, holotype 1: 1.35: 2.29: 2.97. Ant IV annulated on distal three quarters, with apical bulb apically unilobed. Ant III annulated on distal two thirds, apical organ with 2 elongated rods, 3 spiny guard sens, s-blunt sens of different sizes and ciliated chaetae. Ant I dorsally with 5 median mac and 3 smooth mic at the base, without spines (Fig. 51A). Eyes A and B larger, G and H slightly smaller, others subequal, and with 5 ‘IO’ chaetae; head dorsal macrochaetotaxy with 11–12 An, 4 A, 4 M and 8 S mac (Fig. 51D). Clypeal formula with 4 (l1–2), 13–17 (ft), 3 (pf0–1) ciliated chaetae, l1–2 and pf0–1 larger than others, l1 apically acuminate, 1–3 ft smaller, others subequal (Fig. 51B). Prelabral chaetae ciliated and not bifurcated. Labral p0 chaeta rarelly with minute median filament. Labral papillae with two inner conical projections, outer papillae absent. Maxillary palp with smooth t.a. and b.c. weakly ciliated, thicker and 1.08 longer than the t.a. Labial papilla E with l.p. apically finger-shaped and not reaching the base of a.a. (Fig. 51C). Basolateral and basomedian labial fields with chaetae M1–2, R (subequal to others), E, L1–2 ciliated. Ventral head with about 29 ciliated chaetae, postlabial formula with 4 (G1–4), 3 (H2–4), 3 (J1–3) chaetae, 13 thin posterior chaetae, and 1 larger b.c. surrounding the cephalic groove (Fig. 51E).</p><p>Thorax chaetotaxy (Fig. 52 A–B). Th II projected anteriorly, a, m and p series with 12–14, 8 and 12–13 mac, respectively. Th III a, m and p series with 4, 1 and 9 mac, respectively. Ratio Th II: III = 2.11–1.67: 1 (n=4), holotype 2.11: 1.</p><p>Abbreviations: (*) doubtful characteristic; (+) present; (–) absent; (?) unknown. References: (1) B̂rner, 1903; (2) Womersley, 1934; (3) Yosii, 1959; (4) Coates, 1968.</p><p>Abdomen chaetotaxy (Figs 52 C–D, 53A–C). Abd I a, m and p series with 1, 5 and 0 mac, respectively. Abd II a, m and p series with 1, 4 and 0 mac, respectively. Abd III a, m and p series with 1, 3 and 1 mac, respectively. Abd IV with 16 central mac on A– T series and 15–17 lateral mac on E– Fe series; at least 3 posterior sens (ps type I, others type II) and 8 posterior mes. Abd V a, m and p series with 1, 4 and 1/5 mac, respectively. Ratio Abd III: IV = 1: 3.86–4.62 (n= 4), holotype 1: 4.04.</p><p>Legs. Subcoxa I with 6 chaetae on a row and 2 psp; subcoxa II with an anterior row of 13 chaetae, posterior row of 7 chaetae and 6 psp; subcoxa III with one row of 12 chaetae, 4 anterior chaetae and 2 posterior psp (Fig. 54 A–C). Trochanteral organ with about 67 spine-like chaetae (Fig. 54D). Tibiotarsus not subdivided, outer side with 5 larger ciliated chaetae apically acuminate, inner side with 7 mac weakly ciliated. Unguis with 4 inner teeth, b.t. larger, m.t. subequal to a.t. in length. Unguiculus with all lamellae smooth and acuminate, except pe serrated and with a small proximal tooth; ratio unguis: unguiculus = 1: 0.54. Tibiotarsal smooth chaeta 1.79 larger than unguiculus and tenent hair 0.69 smaller than unguis outer edge.</p><p>Collophore. Anterior side with 21–23 chaetae, 11 proximal spine-like chaetae, 1 thin and 3–5 ciliated chaetae, 4 ciliated chaetae apically acuminate and 2 mac distally; posterior side with 15 chaetae (2 unpaired), 14 thin ciliated chaetae widely distributed and 1 smooth chaeta distally; lateral flap with about 44 chaetae, 8 smooth and 36 ciliated (Fig. 54E).</p><p>Abbreviations: (*) doubtful characteristic; (+) present; (–) absent; (?) unknown. References: (1) Coates, 1968.</p><p>Furcula. Manubrium ventral formula with 0, 0, 0, 2/2 (subapical), 20–22 (apical) ciliated chaetae of different sizes plus approximately 15 elongated apical scales per side (Fig. 54F); manubrial plate with 3 blunt mac finely ciliated and apically nipple-like, 14 ciliated chaetae of different sizes and 3 psp. Dens dorsally with one proximal row of 11–13 blunt mac finely ciliated and apically nipple-like, holotype with 11–13 (Fig. 54G)</p><p>Etymology. The new species is named after the Collembola researcher Dr. Charlene Janion-Scheepers (SAM/ IZIKO, South Africa), who provided us specimens of the new species.</p><p>Remarks. Considering the continental taxa from Africa, L. scheepersae sp. nov. resembles L. barnardi and L. marephila (Coates, 1968) comb. nov. in Th II with 6 median mac in m1–2 complex and Abd I–II with 6 and 4 central mac, respectively (Table 4). Lepidocyrtinus scheepersae differs from these species by head mac Pp5 absent (present in L. barnardi), Th II projected anteriorly (normal in L. barnardi and L. marephila) and with 12–13 posterior mac (15–16 in L. barnardi, 15 in L. marephila). In Th III it differs by 11 central mac (10 in L. barnardi, 12 in L. marephila), and Abd IV with 16 central mac (10–11 in L. barnardi). It also differs in ventral manubrium only with ciliated chaetae (3 distal smooth chaetae in L. marephila) and dens with 11–13 blunt chaetae (2–3 in L. marephila, 13–19 in L. barnardi).</p><p>Lepidocyrtinus scheepersae also resembles L. betamponensis sp. nov. from Madagascar in habitus, color pattern with distal spots of pigment on femur III, overall chaetotaxy pattern and other general morphology (Figs 50–54, 72–76). However, L. scheepersae differs from this species by dorsal head mac S4 present (absent in L. betamponensis), prelabral chaetae ciliated and basomedian labial fields with chaetae E ciliated (both smooth in L. betamponensis). It differs also by dorsal chaetotaxy of the Th III–Abd II with 11, 6, 4 central mac, respectively (8–10, 1–2, 3– 4 in L. betamponensis) and Abd IV with 16 central mac (12 in L. betamponensis). Other differences between them are: collophore anteriorly with 11 spine-like chaetae but absent on posterior side in the new species (9 anterior and 8 posterior in L. betamponensis) and manubrial plate and dens with blunt mac apically nipple-like (apically bullet-like and abruptly acuminate at the apex in L. betamponensis).</p><p>Key to continental African species of Lepidocyrtinus</p><p>1 Abd I with 3 or less central mac (Fig. 41C)................................................................. 2</p><p>- Abd I with 5 or more central mac (Fig. 11A, 15C)........................................................... 9</p><p>2 Th III with 3 central; Abd I without mac................................................................... 3</p><p>- Th III with 4 or more central mac; Abd I with at least 1 central mac (Fig. 41 B–C).................................. 4</p><p>3 Th II with 3 central mac in a triangle shape; Th II normal; unguis a.t. present.......... L. metala (Coates, 1968) comb. nov.</p><p>- Th II with 3 central mac in a linear shape; Th II projected anteriorly; unguis a.t. absent. L. tsikama (Coates, 1968) comb. nov.</p><p>4 Abd I–II with 1 and 3 central mac, respectively (Figs 41 C–D).................................................. 5</p><p>- Abd I with 2–3 central mac, Abd II with 2 or 4 central mac (Fig. 35D)........................................... 6</p><p>5 Th II with 4–5 posterior mac (Fig. 41A); Th III with 4 central mac (Fig. 41B); unguiculus pe lamella serrated................................................................................................... L. fabioi sp. nov.</p><p>- Th II with 8 posterior mac; Th III with 6 central mac; unguiculus pe lamella smooth but with one proximal tooth.................................................................................. L. rykei (Coates, 1968) comb. nov.</p><p>6 Abd I with 2 central mac............................................................................... 7</p><p>- Abd I with 3 central mac............................................................................... 8</p><p>7 Th II with 4 mac in m1–2 complex and 8 posterior mac; Th III with 4 central mac (as Fig. 41B); Abd II with 4 central mac (as Fig. 35D)................................................................ L. annela (Coates, 1968) comb. nov.</p><p>- Th II with with 5 mac in m1–2 complex 10 posterior mac; Th III with 8 central mac; Abd II with 2 central mac................................................................................ L. mathewsi (Coates, 1968) comb. nov.</p><p>8 Th II with 10 posterior mac (PmA–C group); Th III with 9 central mac, p1i and p3 mac present; dens dorsally with 4 modified mac................................................................ L. metarsiosa (Coates, 1968) comb. nov.</p><p>- Th II with 9 posterior mac (PmA–C group); Th III with 7 central mac, p1i and p3 mac absent; dens dorsally with 2–3 modified mac................................................................. L. vaneedeni (Coates, 1968) comb. nov.</p><p>9 Abd I with 6–7 central mac (Fig. 52C).................................................................... 10</p><p>- Abd I with 5 central mac (Fig. 11A)..................................................................... 13</p><p>10 Th II with 6 median mac in m1–2 complex (Fig. 15A); Abd I with 6 central mac (Fig. 52C); unguiculus pe lamella serrated (as Fig. 37F)........................................................................................... 11</p><p>- Th II with 5 median mac in m1–2 complex (as in Fig. 12A); Abd I with 7 central mac; unguiculus pe lamella smooth (as Fig. 12C)................................................................... L. lindei (Coates, 1968) comb. nov.</p><p>11 Head with S4 mac (Fig. 14F); Th III with 11 or less central mac (Fig. 52B); dens with 10 or more modified mac (Fig. 17G).. .................................................................................................. 12</p><p>- Head devoid of S4 mac (as Fig. 19E); Th III with 12 central mac; dens with 2–3 modified mac............................................................................................. L. marephila (Coates, 1968) comb. nov.</p><p>12 Prelabral chaetae smooth (Fig. 14D); head with Pp5 mac (Fig. 14F); Abd IV with 10–11 central mac (Fig. 16B); manubrial plate without modified mac (Fig. 17G)................................................. L. barnardi (Womersley, 1934)</p><p>- Prelabral chaetae ciliated; head without Pp5 mac (Fig. 51D); Abd IV with 16 central mac (Fig. 53B); manubrial plate with 3 modified mac (Fig. 54G)............................................................. L. scheepersae sp. nov.</p><p>13 Th II with 5 median mac in m1–2 complex (Figs 12A, 47A).................................................. 14</p><p>- Th II with 6 median mac in m1–2 complex (Fig. 35A)....................................................... 15</p><p>14 Th II PmA and PmC groups with 6 and 2 mac (p3 as mic), respectively; Th III with p1p2 mac absent (Fig. 47 A–B); unguiculus pe lamella serrated (as in Fig. 37F)................................................... L. paucisensillum sp. nov.</p><p>- Th II PmA and PmC groups with 5 and 3 mac (p3 as mac), respectively; Th III with p1p2 mac (Fig. 12A); unguiculus pe lamella smooth (Fig. 12C)................................................................. L. ryozoi nom. nov.</p><p>15 Th III with 11 central mac (Fig. 20B).................................................................... 16</p><p>- Th III with 9 or less central mac (Fig. 25B)................................................................ 17</p><p>16 Th II with 18–20 posterior mac (Fig. 20A); unguis a.t. absent and unguiculus pe lamella serrated; manubrial plate devoid of modified mac (Fig. 22G)................................................................ L. dayi (Yosii, 1959)</p><p>- Th II with 16 or less posterior mac; unguis a.t. present and unguiculus pe lamella smooth (as in Fig. 12C); manubrial plate with 2–3 modified mac...................................................... L. grisea (Womersley, 1934) comb. nov.</p><p>17 Head dark and trunk with a pattern of spots of pigment (e.g. Fig. 23A; Womersley 1934, pag. 462); Th II PmA group with 7 mac............................................................................................... 18</p><p>- Head pale and trunk without a clear pattern of spots of pigment (e.g. Fig. 32); Th II PmA group with 6 mac............. 19</p><p>18 Th II PmC group with 2 mac; Th III with 9 central mac (Figs 25 A–B); manubrial plate and dens with 3 and 5–7 modified mac, respectively (Fig. 29C).................................................................. L. africanus sp. nov.</p><p>- Th II PmC group with 4 mac; Th III with 8 central mac; manubrial plate and dens with 1–2 and 2–3 modified mac, respectively......................................................................... L. capensis Womersley, 1934</p><p>19 Femur and tibiotarsus III distally with dark violet pigments; manubrial plate with 2–3 modified mac (Fig. 11B)........................................................................................ L. annulicornis (B̂rner, 1903)</p><p>- Legs pale (Fig. 32); manubrial plate with 1 modified mac (Fig. 38C).......................... L. botswanensis sp. nov.</p><p>Remarks. The species as L. armillatus (Wahlgren, 1908) and L. laetus (B̂rner, 1908), L. annulipes Handschin, 1929 and L. flavovirens (B̂rner, 1903), are herein transferred to Lepidocyrtinus due to the presence of Ant III–IV annulated, unguis with a pair of large laterobasal teeth, dens dorsally with modified mac (at least in the first two species), and overall macrochaetotaxy pattern (known only to the last two species) (see B̂rner 1903; Wahlgren 1908; Denis 1924; Handschin 1929; Yosii 1959; Coates 1968). However, such species are not included in the key because they are in need to be redescribed, since previous descriptions are insufficient to establish specific diagnoses to them.</p><p>In this sense, some Seira species from Africa which have Ant III and IV annulated and large laterobasal teeth on unguis, but also have poor descriptions unable to clarify their position among the Seirinae, such as S. addoensis Coates, 1968, S. eleana Coates, 1968, S. ethiopica (Denis, 1924) and S. laeta (B̂rner, 1908) may also represent Lepidocyrtinus taxa as well. For this dubious condition, they are herein maintained as Seira s. lat., as well as other species from India described in Lepidocyrtinus by Paliwal &amp; Baijal (1985).</p><p>Madagascar species</p></div>	https://treatment.plazi.org/id/7B6687A7FFD6FFC4FF12D4ECFE0FFA54	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cipola, Nikolas Gioia;Morais, José Wellington De;Bellini, Bruno Cavalcante	Cipola, Nikolas Gioia, Morais, José Wellington De, Bellini, Bruno Cavalcante (2020): Review of Lepidocyrtinus Börner, 1903 (Collembola, Entomobryidae, Seirinae): the African species. Zootaxa 4898 (1): 1-110, DOI: 10.11646/zootaxa.4898.1.1
7B6687A7FFAFFFCDFF12D608FEA0FB79.text	7B6687A7FFAFFFCDFF12D608FEA0FB79.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lepidocyrtinus voeltzkowi (Borner 1907)	<div><p>Lepidocyrtinus voeltzkowi (Börner, 1907)</p><p>Figs 55–60, Tables 1 and 5</p><p>Pseudosira (Mesira) voeltzkowi B̂rner, 1907: 152, figs 8–10, Madagascar, Fianarantsoa, Ankarimbelo (as Ankarimbela mahe Ikongo, SO. Madagascar).</p><p>Sira voeltzkowi; Denis 1925: 284 (as voeltzkovi)</p><p>Lepidocyrtus voeltzkowi; Womersley 1934: 463.</p><p>Lepidocyrtinus voeltzkowi; Delamare-Deboutteville 1948: 39; Salmon 1964: 490.</p><p>Seira voeltzkowi; Yosii 1959: 9; Thibaud 2008: 503.</p><p>Lepidocyrtinus pallidus B̂rner, 1907 syn. nov.: 154</p><p>Typological note. The type material of L. voeltzkowi is probably lost. Thus, here the neotype is designated, according to ICZN (2000, chapter 16, article 72.4.5.), as the specimen fits all morphological characteristics presented in the original description and was collected quite near (less than 50km) the type locality (Fig. 55).</p><p>Examined type material. Neotype female designed on slide (54322/ CAS): Madagascar, Fianarantsoa province, Haute Matsiatra Region, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=46.844444&amp;materialsCitation.latitude=-21.858889" title="Search Plazi for locations around (long 46.844444/lat -21.858889)">Anja Community Reserve</a>, 21°51’32”S, 046°50’40”E (Fig. 55), isleberg vegetation, 1097 m, 17-24.i.2013, M Irwin &amp; R Harin’Hala coll.</p><p>Additional material: 1 male, 6 females and 1 juvenile on slides and 15 specimens in alcohol (54322/ CAS), same data as neotype . 1 female on slide (54322/ CAS donated to INPA), same data as neotype . 11 specimens in alcohol (54321/ CAS donated to INPA), same data as neotype, except 27.x-07.xi.2013 . 31 specimens in alcohol (54342/ CAS), same data as neotype, except 30.iv-07.v.2012 . 3 females on slides (54015, 54057, 54059/ CAS): Toliara, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=44.660557&amp;materialsCitation.latitude=-20.069168" title="Search Plazi for locations around (long 44.660557/lat -20.069168)">Forêt of Kirindy</a>, 15.5 km 64° ENE Marofandilia, 20°04’09”S, 044°39’38”E (Fig. 64), Tropical dry forest, 60 m, 12-24.ix.2008, 24.ix-01.x.2008, 25.iii-01.iv.2009, Malaise trap, BL Fisher coll.</p><p>Diagnosis. Body with dark bluish to graish pigments internally on antennae, all head, trunk and femur, some specimens parcially pigmented or depigmented (Fig. 56); head mac A0, A2–3, A5, M1–2, S0–3, S5–6 and Pp5 present; prelabral chaetae ciliated; labial papilla E with l.p. apically pointed and reaching the base of a.a. (Figs 57 C–D); basolateral and basomedian labial fields with chaetae R (smaller than the others), E, L1–2 ciliated; Th II projected anteriorly and with 13–14 anterior, 8 median (m1ip, m4–4p present, m4i mac absent) and 10–11 posterior mac (p2ep as mac, p3 as mic); Th III–Abd II with 8–10, 4–5, 4 central mac, respectively; Abd IV with 16–17 central mac and 13 lateral mac (Figs 58, 59B); unguis a.t. present (Fig. 60E); unguiculus pe lamella serrated and with a small proximal tooth; collophore anteriorly with 16 spine-like chaetae and 3 ciliated chaetae apically acuminate, posteriorly with 2 spines, smooth chaeta absent (Fig. 60F); manubrium ventrally with 2/2 subapical chaetae; manubrial plate with 3 blunt mac apically bullet-like; dens with 6–10 blunt mac apically nipple-like (Figs 60 G–H).</p><p>Description. Total length (head + trunk) of specimens 2.31–3.61 (n=4), neotype 3.61 mm. Specimens ground pale white with variable colour patterns: with a dark bluish to grayish strip on Ant II that extends in an inner strip to Ant IV, or the entire Ant IV; pigments over all head and trunk, except for Th II–Abd II dorsally; or head and Th II–Abd III only partially pigmented; pigments on all femur I–III or only femur III distally; or head, trunk and legs depigmented; eyepatches black (Figs 56 A–D). Scales present on Ant I to proximal one-third of Ant IV, dorsal and ventral head, dorsal thorax and abdomen, legs (except empodia), anterior and posterior collophore, ventral and dorsal manubrium and dens ventrally.</p><p>Head. Antennae smaller to larger than trunk length (Fig. 56); ratio antennae: trunk = 1: 0.78–1.13 (n=4), neotype 1: 0.78; antennal segments ratio as I: II: III: IV = 1: 1.36–1.49: 2.75: 3.68–4.12, neotype 1: 1.38: 2.75: 3.68. Ant IV annulated on distal three-fourths, with apical bulb apically unilobed or bilobed. Ant III annulated on distal five-sixth, apical organ with 2 elongated rods, 3 spiny guard sens, s-blunt sens of different sizes and ciliated chaetae. Ant I dorsally with 4 median mac surrounded by ciliated guard mic, 2 outer spines and 4 smooth mic at the base (Fig. 57A). Eye B larger, G and H smaller, others subequal, and with 5 ‘IO’ chaetae; head dorsal macrochaetotaxy with 10–12 An, 4 A, 3 M, 7 S and 1 Pp mac (Fig. 57D). Clypeal formula with 4 (l1–2), 2 (ft), 3 (pf1) ciliated chaetae, l1–2 larger than the others, l1 apically acuminate, 2 ft smaller, others subequal (Fig. 57B). Prelabral chaetae ciliated and not bifurcated. Labral p0 chaeta without median filament. Labral papillae with two inner conical projections, outer papillae absent. Maxillary palp with smooth t.a. and b.c. weakly ciliated, thicker and 1.46 longer than the t.a. Labial papilla E with l.p. apically pointed and reaching the base of a.a. (Fig. 57C). Basolateral and basomedian labial fields with chaetae M1–2, R (smaller than the others), E, L1–2 ciliated. Ventral head with about 31–33 ciliated chaetae, postlabial formula with 4 (G1–4), 3 (H2–4), 4 (J1–4) chaetae, H3–4, J1–4 larger than the others, 11–12 thin posterior chaetae with different lengths, and 4–5 larger chaetae, 1 b.c. surrounding the cephalic groove (Fig. 57E).</p><p>Thorax chaetotaxy (Fig. 58 A–B). Th II projected anteriorly, a, m and p series with 13–14, 8 and 10–11 mac, respectively, and about 6 secondary psp. Th III a, m and p series with 3–4, 0 and 7–8 mac, respectively, and about 6 secondary psp. Ratio Th II: III = 2.06–1.64: 1 (n=4), neotype 1.73: 1.</p><p>Abdomen chaetotaxy (Figs 58 C–D, 59A–C). Abd I a, m and p series with 0–1, 4 and 0 mac, respectively, and about 3 secondary psp. Abd II a, m and p series with 1, 4 and 0 mac, respectively. Abd III a, m and p series with 0, 3 and 1 mac, respectively, and 1 lateral secondary psp. Abd IV with 16–17 central mac on A–T series and 13 lateral mac on E–Fe series; at least 2 posterior secondary psp, at least 4 posterior sens (ps type I, others type II) and 8 posterior mes. Abd V a, m and p series with 1, 4 and 2/6 mac, respectively. Ratio Abd III: IV = 1: 5.54–6.90 (n= 4), neotype 1: 5.54.</p><p>Legs. Subcoxa I with 7 chaetae on a row, 16 anterior chaetae and 2 psp; subcoxa II with an anterior row of 11 chaetae, posterior row of 6 chaetae and 4 psp; subcoxa III with one row of 13 chaetae, 2 anterior chaetae and 2 posterior psp (Figs 60 A–C). Trochanteral organ with about 54 spine-like chaetae (Fig. 60D). Tibiotarsus normal or subdivided on distal two thirds, outer side with 5 large chaetae apically acuminate, inner side with 8 mac weakly ciliated. Unguis with 4 inner teeth, b.t. subequal to m.t. in length, m.t. near to a.t., a.t. smaller (Fig. 60E). Unguiculus with all lamellae smooth and acuminate, except pe serrated and with a small proximal tooth; ratio unguis: unguiculus = 1: 0.69. Tibiotarsal smooth chaeta 1.07 larger than unguiculus and tenent hair 0.95 smaller than unguis outer edge.</p><p>Collophore. Anterior side with 24 chaetae, 16 spine-like chaetae, 1 thin and 1 normal ciliated chaeta, 3 ciliated chaetae apically acuminate and 3 mac distally; posterior side with 13 chaetae (1 unpaired), 2 spines and 11 thin ciliated chaetae widely distributed, smooth chaetae absent; lateral flap with about 35–36 chaetae, 10 smooth and 25–26 ciliated (Fig. 60F).</p><p>Furcula. Manubrium ventral formula with 0, 0, 0, 2/2 (subapical), 16 (apical) ciliated chaetae plus about 19 elongated apical scales and 1 ciliated chaeta per side (Fig. 60G); manubrial plate with 3 blunt mac ciliated apically bullet-like, 10 ciliated chaetae of different sizes and 3 psp. Dens dorsally with one proximal row of 6–10 blunt mac finely ciliated and apically nipple-like, neotype with 6+8 (Fig. 60H).</p><p>Remarks. After more than 100 years, L. voeltzkowi is herein redescribed for the first time. B̂rner (1907) in his original description of L. voeltzkowi reported different colour patterns to the species, with dark pigments on body, unguis with large outer teeth and presence of inner a.t. Such characteristics combined are diagnostic of L. voeltzkowi among the Madagascar species, and the chaetotaxy analysis allowed us to confirm its variable colour pattern, including a depigmented morphotype (Fig. 56C). These yellowish specimens with an internal strip of pigment on Ant IV and with unguis a.t. also fit B̂rner’s description of L. pallidus, and such combination of features are not seen in any other Madagascar taxon (see B̂rner 1907 and Table 5). So, we herein suggest L. pallidus syn. nov. as a junior synonym of L. voeltzkowi . Further comparisons of this species with similar taxa, see the remarks of L. iegoi sp. nov. and Table 5.</p></div>	https://treatment.plazi.org/id/7B6687A7FFAFFFCDFF12D608FEA0FB79	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cipola, Nikolas Gioia;Morais, José Wellington De;Bellini, Bruno Cavalcante	Cipola, Nikolas Gioia, Morais, José Wellington De, Bellini, Bruno Cavalcante (2020): Review of Lepidocyrtinus Börner, 1903 (Collembola, Entomobryidae, Seirinae): the African species. Zootaxa 4898 (1): 1-110, DOI: 10.11646/zootaxa.4898.1.1
7B6687A7FFA6FFD2FF12D508FD7EFED9.text	7B6687A7FFA6FFD2FF12D508FD7EFED9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lepidocyrtinus andevo Cipola & Morais & Bellini 2020	<div><p>Lepidocyrtinus andevo sp. nov. Cipola &amp; Bellini</p><p>Figs 55, 61–66, Tables 1 and 5</p><p>Type Material. Holotype female on slide (54085/ CAS): Madagascar, Toamasina, Ankerana municipality, 18°24’23”S, 048°48’09”E (Fig. 55), <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=48.802498&amp;materialsCitation.latitude=-18.40639" title="Search Plazi for locations around (long 48.802498/lat -18.40639)">Montane forest</a>, 1108 m, 19-24.i.2012, Malaise trap, BL Fisher coll. 15 paratypes: 3 males, 4 females and 1 juvenile on slides and 5 specimens in alcohol (54085/ CAS); 1 male and 1 female on slides (54085/ CAS donated to INPA-CLL 000030), same data as holotype.</p><p>Diagnosis. Body with dark blue pigments all over Ant I–IV, Th II laterally and femur I–III (Fig. 61); head mac A0, A2–3, A5, M1–2, S0–3 and S5–6 present; prelabral chaetae smooth; labial papilla E with l.p. conical and reduced (Figs 62 D–F); basolateral and basomedian labial fields with chaetae R (smaller than the others), L1–2 ciliated and e smooth; Th II projected anteriorly and with 6–7 anterior, 8 median (m1ip, m4–4p present, m4i mac absent) and 9–11 posterior mac (p2ep absent, p3 as mac); Th III–Abd II with 9–10, 2–4, 3–4 central mac respectively; Abd IV with 10–12 central mac and 14–16 lateral mac (Figs 63, 64B); unguis a.t. absent; unguiculus pe lamella smooth and with a small proximal tooth (Fig. 65E); collophore anteriorly with 9 spine-like chaetae and 4 ciliated chaetae apically acuminate, posteriorly with 10 spines, smooth chaeta present (Fig. 66A); manubrium ventrally with 0/4 subapical chaetae; manubrial plate with 2 blunt mac abruptly acuminate at the apex; dens with 1–12 blunt mac, 3 apically bullet-like and up to 9 abruptly acuminate at the apex (Figs 66 B–C).</p><p>Description. Total length (head + trunk) of specimens 3.97–5.10 (n=4), holotype 5.10 mm. Specimens pale white with dark blue pigment on Ant I–IV, lateral Th II, anterior coxa III, trochanter and femur completely, lighter on tibiotarsus; eyepatches black (Fig. 61). Scales present on Ant I to proximal one quarter of Ant IV, dorsal and ven-tral head, dorsal thorax and abdomen, legs (except empodia), anterior and posterior collophore, ventral and dorsal manubrium and dens ventrally.</p><p>Head. Antennae subequal to trunk length (Fig. 61); ratio antennae: trunk = 1: 0.96–1.14 (n=4), holotype 1: 0.98; antennal segments ratio as I: II: III: IV = 1: 1.21–1.37: 1.18–1.47: 2.77–3.48, holotype 1: 1.23: 1.18: 2.77. Ant IV annulated on distal three fourths, with apical bulb apically unilobed, bilobed or trilobed.Ant III not annulated, apical organ with 2 elongated rods, 3 spiny guard sens, s-blunt sens of different sizes and ciliated chaetae (Fig. 62A). Ant I dorsally with 4 median mac surrounded by ciliated guard mic and 13 smooth mic at the base, without spines (Fig. 62B). Eyes A and B larger, G and H smaller, others subequal, and with 5 ‘IO’ chaetae; head dorsal macrochaetotaxy with 15–16 An, 4 A, 3 M, 7 S and 1 Ps mac (Fig. 62F). Clypeal formula with 6 (l12, l1–2), 15 (ft), 2 (pf1) ciliated chaetae, l1 larger than the others and apically acuminate, pf0 absent, others subequal (Fig. 62C). Prelabral chaetae smooth, sometimes bifurcated. Labral p0 chaeta sometimes with median filament, p1 larger than the others (Fig. 62D). Labral papillae with two rounded projections, outer papillae absent. Maxillary palp with smooth t.a. and b.c. weakly ciliated, thicker and 1.39 longer than the t.a. Labial papilla E with l.p. conical and reduced (Fig. 62E). Basolateral and basomedian labial fields with chaetae M1–2, R (smaller than the others), L1–2 ciliated, e smooth. Ventral head with about 13 ciliated chaetae, postlabial formula with 4 (G1–4), 3 (H2–4), 4 (J1–4) chaetae, H3 sometimes thin and smaller, and 1 b.c. larger than the others surrounding the cephalic groove (Fig. 62G).</p><p>Thorax chaetotaxy (Fig. 63 A–B). Th II projected anteriorly, a, m and p series with 6–7, 8 and 9–11 mac, respectively, and about 20 secondary psp. Th III a, m and p series with 5–6, 1 and 7 mac, respectively, and about 10 secondary psp. Ratio Th II: III = 2.93–1.95: 1 (n=4), holotype 2.93: 1.</p><p>Abdomen chaetotaxy (Figs 63 C–D, 64A–C). Abd I a, m and p series with 0–1, 2–3 and 0 mac, respectively, and about 5 secondary psp. Abd II a, m and p series with 1, 3–4 and 0 mac, respectively, and about 11 secondary psp. Abd III a, m and p series with 0, 3 and 1 mac, respectively, and about 16 secondary psp. Abd IV with 10–12 central mac on A–T series and 14–16 lateral mac on E–Fe series; about 11 secondary psp, at least 6 posterior sens (ps type I, others type II) and 5 posterior mes. Abd V a, m and p series with 1, 4 and 6 mac, respectively. Ratio Abd III: IV = 1: 6.65–4.78 (n= 4), holotype 1: 4.78.</p><p>Legs. Subcoxa I with 6 chaetae on a row, 12 anterior chaetae and 2 psp; subcoxa II with an anterior row of 17 chaetae, posterior row of 10 chaetae and 4 psp; subcoxa III with one row of 13 chaetae and 2 posterior psp (Fig. 65 A–C). Trochanteral organ with about 68 spine-like chaetae (Fig. 65D). Tibiotarsus III sometimes subdivided on distal two thirds, outer side with 3 proximal large chaetae apically acuminate, inner side with 7 mac weakly ciliated. Unguis with 3 inner teeth, b.t. smaller than m.t., a.t. absent. Unguiculus with all lamellae smooth and acuminate, except pe with a small proximal tooth; ratio unguis: unguiculus = 1: 0.49. Tibiotarsal smooth chaeta 1.12 larger than unguiculus and tenent hair 0.51 smaller than unguis outer edge (Fig. 65E).</p><p>Collophore. Anterior side with 20 chaetae, 9 proximal spine-like chaetae, 1 thin ciliated chaeta, 4 ciliated chaetae apically acuminate and 6 mac distally; posterior side with 28 chaetae (3 unpaired), 10 spines, 17 thin ciliated chaetae widely distributed and 1 smooth chaetae distally; lateral flap with about 88 chaetae, 11 smooth and 77 ciliated (Fig. 66A).</p><p>Furcula. Manubrium ventral formula with 0, 0, 0, 0/4 (subapical), 14–16 (apical) ciliated chaetae plus approximately 35 elongated apical scales per side (Fig. 66B); manubrial plate with 2–3 blunt mac ciliated abruptly acumi-nate at apex, 17 ciliated chaetae of different sizes and 2 psp. Dorsal dens with one row of 1–12 blunt mac weakly ciliated, 3 proximal apically bullet-like and up to 9 abruptly acuminate at the apex, holotype with 3+9 (Fig. 66C).</p><p>Etymology. The epithet “andevo” is proposed to honor the black slaves that constituted about a third of the Merina society of Madagascar during the 19th century.</p><p>Remarks. Lepidocyrtinus andevo sp. nov. resembles other species from Madagascar (Table 5). See the comparison among them in the remarks of L. iegoi sp. nov.</p></div>	https://treatment.plazi.org/id/7B6687A7FFA6FFD2FF12D508FD7EFED9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cipola, Nikolas Gioia;Morais, José Wellington De;Bellini, Bruno Cavalcante	Cipola, Nikolas Gioia, Morais, José Wellington De, Bellini, Bruno Cavalcante (2020): Review of Lepidocyrtinus Börner, 1903 (Collembola, Entomobryidae, Seirinae): the African species. Zootaxa 4898 (1): 1-110, DOI: 10.11646/zootaxa.4898.1.1
7B6687A7FFB9FFD9FF12D1A8FD53F901.text	7B6687A7FFB9FFD9FF12D1A8FD53F901.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lepidocyrtinus briani Cipola & Morais & Bellini 2020	<div><p>Lepidocyrtinus briani sp. nov. Cipola &amp; Bellini</p><p>Figs 55, 67–71, Tables 1 and 5</p><p>Type Material. Holotype female on slide (54095/ CAS): Madagascar, Antsiranana, Daraina municipality, Binara Forest, 13°15’50”S; 049°35’57”E (Fig. 55), <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=49.599167&amp;materialsCitation.latitude=-13.263889" title="Search Plazi for locations around (long 49.599167/lat -13.263889)">Rainforest</a>, 1065 m, 18.x.2013, Malaise trap, BL Fisher coll. 8 paratypes: 1 male and 2 females on slides and 3 specimens in alcohol (54095/ CAS); 2 females on slides (54095/ CAS donated to INPA-CLL 000031), same data as holotype.</p><p>Diagnosis. Body with dark pigments on Ant II–IV and Th II laterally (Fig. 67); head mac A0, A2–3, A5, M1–2, S0–3 and S5–6 present; prelabral chaetae smooth; labial papilla E with l.p. apically pointed and reaching the base of a.a.; basolateral and basomedian labial fields with chaetae R (subequal to others), E and L1–2 ciliated (Figs 68 B–D); Th II projected anteriorly and with 6–7 anterior, 8 median (m1ip, m4–4p present, m4i mac absent) and 9–10 posterior mac (p2ep absent, p3 as mac); Th III–Abd II with 5–7, 3, 3 central mac respectively; Abd IV with 11 central mac and 14 lateral mac (Figs 69, 70B); unguis a.t. present; unguiculus pe lamella serrated and with a small proximal tooth; collophore anteriorly with 8 spine-like chaetae and 5 ciliated chaetae apically acuminate, posteriorly with 8 spines, smooth chaeta present (Fig. 71E); manubrium ventrally with 0–2/2 subapical chaetae; manubrial plate with 2 blunt mac abruptly acuminate at the apex; dens without blunt mac (Figs 71 F–G).</p><p>Description. Total length (head + trunk) of specimens 2.48–3.43 (n=4), holotype 3.43 mm. Specimens pale white with dark blue pigment on lateral Th II and light blue pigment on Ant II–IV, femur and tibiotarsus; eyepatches black (Fig. 67). Scales present on Ant I to proximal one seventh of Ant IV, dorsal and ventral head, dorsal thorax and abdomen, legs (except empodia), anterior collophore, ventral and dorsal manubrium and dens ventrally.</p><p>Head. Antennae smaller than trunk length (Fig. 67); ratio antennae: trunk = 1: 1.78–1.80 (n=3), holotype 1: 1.78; antennal segments ratio as I: II: III: IV = 1: 1.22–1.54: 1.42: 1.98–3.09, holotype 1: 1.25: 1.42: 1.98 (Ant III only holotype). Ant IV annulated on distal six seventh, with apical bulb apically bilobed. Ant III not annulated, apical organ with 2 elongated rods, 3 spiny guard sens and s-blunt sens of different sizes. Ant I dorsally with 4 median mac not surrounded by ciliated guard mic and 3 smooth mic at the base, without spines. Eyes A and B larger, G and H smaller, others subequal, and with 5 ‘IO’ chaetae; head dorsal macrochaetotaxy with 10–12 An, 4 A, 3 M and 7 S mac (Fig. 68C). Clypeal formula with 4 (l1-2), 10 (ft), 3 (pf 0-2) ciliated chaetae, l1-2 larger than the others, l1 apically acuminate, 1 ft smaller, others subequal (Fig. 68A). Prelabral chaetae smooth, inner chaetae rarely bifurcated. Labral p0 chaeta sometimes with median filament, p1 larger than the others. Labral papillae with two inner rounded projections, outer papillae absent. Maxillary palp with smooth t.a. and b.c. weakly ciliated, thicker and 1.41 longer than the t.a. Labial papilla E with l.p. apically pointed and reaching the base of a.a. (Fig. 68B). Basolateral and basomedian labial fields with chaetae M1–2, R (subequal to others), E, L1–2 ciliated (Fig. 68D). Ventral head with about 17 ciliated chaetae, postlabial formula with 4 (G1–4), 3 (H2–4), 4 (J1–4) chaetae, and 2 larger b.c. surrounding the cephalic groove (Fig. 68D).</p><p>Thorax chaetotaxy (Fig. 69 A–B). Th II projected anteriorly, a, m and p series with 6–7, 8 and 9–10 mac, respectively. Th III a, m and p series with 3–4, 1 and 3–4 mac, respectively. Ratio Th II: III = 1.96–2.48: 1 (n=4), holotype 2.48: 1.</p><p>Abdomen chaetotaxy (Figs 69 C–D, 70A–C). Abd I a, m and p series with 0, 3 and 0 mac, respectively, and at least 1 secundary psp. Abd II a, m and p series with 1, 3 and 0 mac, respectively, m3e as mic. Abd III a, m and p series with 0, 2–3 and 1 mac, respectively. Abd IV with 11 central mac on A–T series and 14 lateral mac on E– Fe series; 2 secundary psp, at least 6 posterior sens (ps type I, others type II) and 8 posterior mes. Abd V a, m and p series with 1, 4 and 6 mac, respectively. Ratio Abd III: IV = 1: 4.49–5.39 (n= 4), holotype 1: 4.49.</p><p>Legs. Subcoxa I with 7–8 chaetae on a row, 5 anterior chaetae and 2 psp; subcoxa II with an anterior row of 14 chaetae, posterior row of 7 chaetae and 6 psp; subcoxa III with one row of 14 chaetae and 2 posterior psp (Fig. 71 A–C). Trochanteral organ with about 44 spine-like chaetae (Fig. 72D). Tibiotarsus not subdivided, outer side with 6 large chaetae apically acuminate, inner side with 13 mac weakly ciliated. Unguis with 4 inner teeth, b.t. smaller than m.t., a.t. smaller than the others. Unguiculus with all lamellae smooth and acuminate, except pe serrated and with a small proximal tooth; ratio unguis: unguiculus = 1: 0.63. Tibiotarsal smooth chaeta 1.18 larger than unguiculus and tenent hair 0.69 smaller than unguis outer edge.</p><p>Collophore. Anterior side with 18 chaetae, 8 proximal spine-like chaetae, 1 thin and 2 normal ciliated chaetae, 5 ciliated chaetae apically acuminate and 2 mac distally; posterior side with 22 chaetae (5 unpaired), 8 spines, 13 thin ciliated chaetae widely distributed and 1 smooth chaetae distally; lateral flap with about 50 chaetae, 7 smooth and 43 ciliated (Fig. 71E).</p><p>Furcula. Manubrium ventral formula with 0, 0, 0, 0–2/2 (subapical), 14–16 (apical) ciliated chaetae plus approximately 20 elongated apical scales per side (Fig. 71F); manubrial plate with 2 blunt mac ciliated abruptly acuminate at the apex, 16 ciliated chaetae of different sizes and 3 psp. Dens dorsally with ciliated chaetae, without modified mac (Fig. 71G).</p><p>Etymology. It is named after Dr. Brian L. Fisher (CAS), collector of the new species.</p><p>Remarks. Lepidocyrtinus briani sp. nov. resembles other species from Madagascar (Table 5). See the comparison among them in the remarks of L. iegoi sp. nov.</p></div>	https://treatment.plazi.org/id/7B6687A7FFB9FFD9FF12D1A8FD53F901	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cipola, Nikolas Gioia;Morais, José Wellington De;Bellini, Bruno Cavalcante	Cipola, Nikolas Gioia, Morais, José Wellington De, Bellini, Bruno Cavalcante (2020): Review of Lepidocyrtinus Börner, 1903 (Collembola, Entomobryidae, Seirinae): the African species. Zootaxa 4898 (1): 1-110, DOI: 10.11646/zootaxa.4898.1.1
7B6687A7FFB2FFDBFF12D710FCAFF8CD.text	7B6687A7FFB2FFDBFF12D710FCAFF8CD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lepidocyrtinus betamponensis Cipola & Morais & Bellini 2020	<div><p>Lepidocyrtinus betamponensis sp. nov. Cipola &amp; Bellini</p><p>Figs 55, 72–76, Tables 1 and 5</p><p>Type Material. Holotype female on slide (54128/ CAS): Madagascar, Toamasina, Réserve Nationale Intégrale Betampona, 17°54’58”S, 049°12’07”E (Fig. 55), <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=49.201946&amp;materialsCitation.latitude=-17.916111" title="Search Plazi for locations around (long 49.201946/lat -17.916111)">Rainforest</a>, 550 m, 24.iv-04.v.2008, Malaise trap, BL Fisher coll. 67 paratypes: 1 male and 5 females on slides and 30 specimens in alcohol (54128/ CAS); 1 male and 2 females on slides and 15 specimens in alcohol (54128/ CAS donated to INPA-CLL 000032), same data as holotype; 13 specimens in alcohol (54120, 54153-54, 56/ CAS), same data as holotype, except 3-10.viii.2008, 31.viii-07.ix.2008, 28.ix-05- x.2008, 21-28.xii.2008.</p><p>Diagnosis. Body with dark pigments on Ant II–IV and Th II laterally (Fig. 72); head mac A0, A2–3, A5, M1–2, S0–3 and S5–6 present; prelabral chaetae smooth; labial papilla E with l.p. conical and not reaching the base of a.a.; basolateral and basomedian labial fields with chaetae R (smaller than the others) and L1–2 ciliated, e smooth (Figs 73 B–E); Th II projected anteriorly and with 5–7 anterior, 8 median (m1ip, m4–4p present, m4i mac absent) and 9–10 posterior mac (p2ep absent, p3 as mac); Th III–Abd II with 8–10, 1–2, 3–4 central mac respectively; Abd IV with 12 central mac and 12–15 lateral mac (Figs 74, 75B); unguis a.t. absent; unguiculus pe lamella smooth and with a small proximal tooth; collophore anteriorly with 9 spine-like chaetae and 5–6 ciliated chaetae apically acuminate, posteriorly with 8 spines, smooth chaeta present (Fig. 76E); manubrium ventrally with 0/4 subapical chaetae; manubrial plate with 2 blunt mac abruptly acuminate at the apex; dens with 5–9 blunt mac apically bullet-like abruptly acuminate at the apex (Figs 76 F–G).</p><p>Description. Total length (head + trunk) of specimens 3.03–3.70 (n=4), holotype 3.70 mm. Specimens pale white with dark blue pigment on Ant I–IV (darker on distal Ant II, proximal and distal Ant III and IV), lateral Th II–III, femur I–II with one distal band and III with proximal and distal bands, and tibiotarsus almost completely (except distally); eyepatches black (Fig. 72). Scales present on Ant I to proximal one quarter of Ant IV, dorsal and ventral head, dorsal thorax and abdomen, legs (except empodia), anterior collophore, ventral and dorsal manubrium and dens ventrally.</p><p>Head. Antennae subequal or smaller than trunk length (Fig. 72); ratio antennae: trunk = 1: 1.10–1.21 (n=3), holotype 1: 1.21; antennal segments ratio as I: II: III: IV = 1: 1.28–1.48: 1.41–1.48: 2.83–3.00, holotype 1: 1.40: 1.48: 2.85. Ant IV annulated on distal three fourths, with apical bulb apically bilobed. Ant III not annulated, apical organ with 2 short rods, 3 spiny guard sens and s-blunt sens of different sizes. Ant I dorsally with 4–5 median mac and 3 smooth mic at the base, without spines. Eyes A and B larger, G and H smaller, others subequal, and with 5–6 ‘IO’ chaetae; head dorsal macrochaetotaxy with 10–13 An, 4 A, 3 M and 7 S mac (Fig. 73D). Clypeal formula with 4 (l1–2), 7 (ft), 5 (pf0–2) ciliated chaetae, l1–2 larger than the others, l1 apically acuminate, 1 ft smaller, others subequal (Fig. 73A). Prelabral chaetae smooth but sometimes with a small median filament, outer chaetae rarely with two median filaments (Fig. 73B). Labral p0 chaeta sometimes with median filament or rarely bifurcated, p1 larger than the others. Labral papillae with two inner rounded projections, outer papillae absent. Maxillary palp with smooth t.a. and b.c. weakly ciliated, thicker and 1.29 longer than the a.a. Labial papilla E with l.p. conical and not reaching the base of a.a. (Fig. 73C). Basolateral and basomedian labial fields with chaetae M1–2, R (smaller than the others), L1–2 ciliated, e smooth (Fig. 73E). Ventral head with about 16 ciliated chaetae, postlabial formula with 4 (G1–4), 3 (H2–4), 4 (J1–4) chaetae, and 3 larger chaetae, 2 b.c. surrounding the cephalic groove (Fig. 73E).</p><p>Thorax chaetotaxy (Fig. 74 A–B). Th II projected anteriorly, a, m and p series with 5–7, 8 and 9–10 mac, respectively, and about 11 secundary psp. Th III a, m and p series with 4–5, 1 and 7–8 mac, respectively, and about 6 secundary psp. Ratio Th II: III = 2.25–1.92: 1 (n=4), holotype 2.13: 1.</p><p>Abdomen chaetotaxy (Figs 74 C–D, 75A–C). Abd I a, m and p series with 0, 1–2 and 0 mac, respectively, and at least 2 secundary psp. Abd II a, m and p series with 0–1, 4 and 0 mac, respectively, and at least 2 secundary psp. Abd III a, m and p series with 0, 3 and 1 mac, respectively, and about 8 secundary psp. Abd IV with 12 central mac on A–T series and 12–15 lateral mac on E– Fe series; about 9 secundary psp, at least 8 posterior sens (ps type I, others type II) and 6 posterior mes. Abd V a, m and p series with 1, 4 and 6 mac, respectively. Ratio Abd III: IV = 1: 4.51–5.44 (n= 4), holotype 1: 4.53.</p><p>Legs. Subcoxa I with 6 chaetae on a row, 2 anterior chaetae and 2 psp; subcoxa II with an anterior row of 12 chaetae, posterior row of 7 chaetae and 6 psp; subcoxa III with one row of 12 chaetae and 2 posterior psp (Figs 76 A–C). Trochanteral organ with about 78 spine-like chaetae (Fig. 76D). Tibiotarsus III sometimes subdivided on distal two thirds, outer side with 4 proximal large chaetae apically acuminate, inner side with 7 mac weakly ciliated. Unguis with 3 inner teeth, b.t. subequal to m.t. in length, a.t. absent. Unguiculus with all lamellae smooth and acuminate, except pe with a small proximal tooth (as in Fig. 51E); ratio unguis: unguiculus = 1: 0.63. Tibiotarsal smooth chaeta 1.18 larger than unguiculus and tenent hair 0.72 smaller than unguis outer edge.</p><p>Collophore. Anterior side with 18–20 chaetae, 9 proximal spine-like chaetae, 1 thin and 1–2 normal ciliated chaetae, 5–6 ciliated chaetae apically acuminate and 2 mac distally; posterior side with 18–19 chaetae (2 unpaired), 8 spines, 9–10 thin ciliated chaetae widely distributed and 1 smooth chaeta distally; lateral flap with about 67 chaetae, 10 smooth and 57 ciliated (Fig. 76E).</p><p>Furcula. Manubrium ventral formula with 0, 0, 0, 0/4 (subapical), 12–16 (apical) ciliated chaetae plus approximately 24 elongated apical scales per side (Fig. 76F); manubrial plate with 2 blunt mac ciliated abruptly acuminate at the apex, 12 ciliated chaetae of different sizes and 3 psp. Dens dorsally with one proximal row of 5–9 blunt mac weakly ciliated, 3 proximal apically bullet-like and up to 6 abruptly acuminate at the apex, holotype with 7+7 (Fig. 76G).</p><p>Etymology. Refers to its type locality: Betampona Reserve, Madagascar.</p><p>Remarks. Lepidocyrtinus betamponensis sp. nov. resembles others species from Madagascar (Table 5). See the comparison among them in the remarks of L. iegoi sp. nov.</p></div>	https://treatment.plazi.org/id/7B6687A7FFB2FFDBFF12D710FCAFF8CD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cipola, Nikolas Gioia;Morais, José Wellington De;Bellini, Bruno Cavalcante	Cipola, Nikolas Gioia, Morais, José Wellington De, Bellini, Bruno Cavalcante (2020): Review of Lepidocyrtinus Börner, 1903 (Collembola, Entomobryidae, Seirinae): the African species. Zootaxa 4898 (1): 1-110, DOI: 10.11646/zootaxa.4898.1.1
7B6687A7FFB5FFE9FF12D089FC2CF8B9.text	7B6687A7FFB5FFE9FF12D089FC2CF8B9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lepidocyrtinus iegoi Cipola & Morais & Bellini 2020	<div><p>Lepidocyrtinus iegoi sp. nov. Cipola &amp; Bellini</p><p>Figs 55, 77–80, Tables 1 and 5</p><p>Type Material. Holotype female on slide (54107/ CAS): Madagascar, Toamasina, Moramanga municipality, Andasibe-Mantadia National Park, 18°45’39”S, 048°22’17”E (Fig. 55), <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=48.371387&amp;materialsCitation.latitude=-18.760834" title="Search Plazi for locations around (long 48.371387/lat -18.760834)">Rainforest</a>, 968 m, 23-28.xi.2012, Malaise trap, BL Fisher coll. 1 paratype female on slide (54107/ CAS), same data as holotype.</p><p>Diagnosis. Body pale (Fig. 77); head mac A0, A2–3, A5, M1–2, S0–3 and S5–6 present; prelabral chaetae smooth; labial papilla E with l.p. conical and reduced (Figs 78 B–C); basolateral and basomedian labial fields with chaetae R (smaller than the others) and L1–2 ciliated, e smooth; Th II projected anteriorly and with 7 anterior, 8 median (m1ip, m4–4p present, m4i mac absent) and 10–11 posterior mac (p2ep absent, p3 as mac); Th III–Abd II with 11, 5, 4 central mac respectively; Abd IV with 10–12 central mac and 14–16 lateral mac (Figs 79 and as in 64B); unguis a.t. absent; unguiculus pe lamella smooth and with a small proximal tooth; collophore anteriorly with 8 spine-like chaetae and 6 ciliated chaetae apically acuminate, posteriorly with 6 spines, smooth chaeta present (Fig. 80E); manubrium ventrally with 0/4 subapical chaetae; manubrial plate with 2 blunt mac abruptly acuminate at the apex; dens with 4–7 blunt mac apically bullet-like (Figs 80 F–G).</p><p>Description. Total length (head + trunk) of specimens 2.97–3.61 (n=2), holotype 3.61 mm. Specimens pale white with vestigial dark blue pigment on distal and proximal Ant IV; eyepatches black (Fig. 77). Scales present on Ant I to proximal one fifth of Ant IV, dorsal and ventral head, dorsal thorax and abdomen, legs (except empodia), anterior and posterior collophore, ventral and dorsal manubrium and dens ventrally.</p><p>Head. Antennae subequal to trunk length (Fig. 77); ratio antennae: trunk = 1: 1.07; antennal segments ratio as</p><p>I: II: III: IV = 1: 1.38: 1.38: 2.70 in holotype (Ant III–IV missing in paratype). Ant IV annulated on distal four fifth, with apical bulb apically bilobed. Ant III not annulated, apical organ with 2 rods, 3 spiny guard sens, s-blunt sens of different sizes and ciliated chaetae.Ant I dorsally with 4 median mac surrounded by ciliated guard mic and 3 smooth mic at the base, without spines. Eyes A and B larger, G and H smaller, others subequal, and with 5 ‘IO’ chaetae; head dorsal macrochaetotaxy with 11–14 An, 4 A, 3 M and 7 S mac (Fig. 78C). Clypeal formula with 6 (l1–2), 8 (ft), 5 (pf0–2) ciliated chaetae, l1–2 larger than the others, l1 apically acuminate, 2 ft smaller, others subequal (Fig. 78A). Prelabral chaetae smooth and not bifurcated. Labral p0 chaeta with median filament, p1 larger than the others. Labral papillae with two inner rounded projections, outer papillae absent. Maxillary palp with smooth t.a. and b.c. weakly ciliated, thicker and 1.44 longer than the t.a. Labial papilla E with l.p. conical and reduced (Fig. 78B). Basolateral and basomedian labial fields with chaetae M1–2, R (smaller than the others), L1–2 ciliated, e smooth. Ventral head with about 15 ciliated chaetae, postlabial formula with 4 (G1–4), 3 (H2–4), 4 (J1–4) chaetae, and 1 larger b.c. surrounding the cephalic groove (Fig. 78D).</p><p>Thorax chaetotaxy (Fig. 79 A–B). Th II projected anteriorly, a, m and p series with 7, 8 and 10–11 mac, respectively, and about 5 secundary psp. Th III a, m and p series with 6, 1 and 8 mac, respectively, and about 6 secundary psp. Ratio Th II: III = 1.94–1.82: 1 (n=2), holotype 1.94: 1.</p><p>Abdomen chaetotaxy (Figs 79 C–D). Abd I a, m and p series with 1, 4 and 0 mac, respectively, and about 5 secundary psp. Abd II a, m and p series with 1, 4 and 0 mac, respectively, and about 5 secundary psp. Abd III a, m and p series with 0, 3 and 1 mac, respectively, and about 8 secondary psp. Abd IV with 10–12 central mac on A–T series (C1 as mac or mic) and 14–16 lateral mac on E–Fe series; about 9 secondary psp, at least 5 posterior sens (ps type I, others type II) and 3 posterior mes. Abd V a, m and p series with 1, 4 and 6 mac, respectively. Chaetotaxy of Abd III–V similar to L. andevo sp. nov. (Figs 64 A–C). Ratio Abd III: IV = 1: 5.15–5.44 (n= 2), holotype 1: 5.44.</p><p>Legs. Subcoxa I with 6 chaetae on a row, 3–5 anterior chaetae and 2 psp; subcoxa II with an anterior row of 16 chaetae, posterior row of 8 chaetae and 4 psp; subcoxa III with one row of 13 chaetae, 1 anterior chaeta and 2 posterior psp (Fig. 80 A–C). Trochanteral organ with about 51 spine-like chaetae (Fig. 80D). Tibiotarsus III normal or subdivided on slightly more than its distal half, outer side with 4 large chaetae apically acuminate, inner side with 7 mac weakly ciliated. Unguis with 3 inner teeth, b.t. smaller to m.t., a.t. absent. Unguiculus with all lamellae smooth and acuminate, except pe with a small proximal tooth; ratio unguis: unguiculus = 1: 0.69. Tibiotarsal smooth chaeta 0.82 smaller than unguiculus and tenent hair 0.73 smaller than unguis outer edge.</p><p>Collophore. Anterior side with 20 chaetae, 8 proximal spine-like chaetae, 4 ciliated chaetae, 6 ciliated chaetae apically acuminate and 2 mac distally; posterior side with 10 chaetae (1 unpaired), 6 spines, 3 thin ciliated chaetae widely distributed and 1 smooth chaeta distally; lateral flap with about 58 chaetae, 12 smooth and 46 ciliated (Fig. 80E).</p><p>Furcula. Manubrium ventral formula with 0, 0, 0, 0/4 (subapical), 12 (apical) ciliated chaetae plus approximately 26 elongated apical scales per side (Fig. 80F); manubrial plate with 2 blunt mac ciliated abruptly acuminate at the apex, 18 ciliated chaetae of different sizes and 3 psp. Dens dorsally with one proximal row of 4–7 blunt mac weakly ciliated and apically bullet-like, holotype with 4+6 (Fig. 80G).</p><p>Etymology. The name “iegoi” (in apposition) refers to nickname of our friend and collembologist Msc. Diego Dias da Silva (INPA), who helped us providing specimens from Madagascar.</p><p>Remarks. The four new species herein described from Madagascar are somehow similar to each other in habitus, general morphology and dorsal chaetotaxy patterns (Figs 61–80). However, they differ from each other by color pattern as: appendages heavily pigmented in L. andevo sp. nov., partially pigmented in L. briani sp. nov. and L. betamponensis sp. nov., and reduced in L. iegoi sp. nov. (Fig. 77). The head morphology of all new species from Madagascar lack Pp5 mac (present in L. voeltzkowi, Fig. 57D), labial papilla E with l.p. reduced in L. andevo, L. betamponensis and L. iegoi (reaching the base of a.a. in L. voeltzkowi and L. briani), basomedian labial field with chaeta E ciliated in L. voeltzkowi and L. briani (smooth in L. andevo, L. betamponensis and L. iegoi), and ventral cephalic groove with 1 b.c. in L. voeltzkowi, L. andevo and L. iegoi (2 in L. betamponensis and L. briani).</p><p>The dorsal chaetotaxy of the Madagascar species differs in: L. voeltzkowi Th II with more than 12 anterior mac in a5 group (7 or less on the four new species) and posteriorly with p3 as mic and p2ep as mac (the opposite in the four new species). In Th III there are 2 lateral mac in L. voeltzkowi and L. briani, while in the other new species there are 4 mac. Other differences in Th III are: a5 mac only present in L. iegoi sp. nov., a1 mac only absent in L. briani, p1i mac absent in L. andevo and L. briani, and a4 mac only absent in L. voeltzkowi (Figs 58 A–B, 63A–B, 69A–B, 74A–B, 79A–B).</p><p>......continued on the next page</p><p>Abbreviations: (C) ciliated chaetae; (S) smooth chaetae; (spn) spine; (+) present; (–) absent.</p><p>Concerning the body appendages, the Madagascar species differ in number of spine-like chaetae on trochanteral organ and unguis a.t. present in L. briani and L. voeltzkowi (Fig. 60E). The collophore anteriorly differs by 6 distal mac present in L. andevo (3 in L. voeltzkowi and 2 in the other three new species) and about 16 spine-like chaetae in L. voeltzkowi (less in the four new species). On the posterior face of the collophore, the species differ by 10 spines present in L. andevo (8 in L. briani and L. betamponensis, 6 in L. iegoi, 2 in L. voeltzkowi) and smooth chaeta present in the four new species (absent in L. voeltzkowi). Finally, in dens they differ by blunt mac: apically nipple-like in L. voeltzkowi, bullet-like and/or abruptly acuminate at the apex in L. andevo, L. betamponensis and L. iegoi (only bullet-like in this species), while such chaetae are completely absent in L. briani . More comparisons are listed in Table 5.</p><p>Key to Lepidocyrtinus species from Madagascar</p><p>1 Head without Pp5 mac (Fig. 62F); Th II with 7 or less anterior mac (a5 group), posteriorly with p3 mac, and p2ep absent (Fig. 63A)............................................................................................... 2</p><p>- Head with Pp5 mac (Fig. 57D); Th II with more than 12 anterior mac (a5 group), posteriorly with p3 as mic and p2ep as mac (Fig. 58A)..................................................................... L. voeltzkowi (B̂rner, 1907)</p><p>2 Basomedian labial field with chaeta e smooth (Fig. 73E); Th III with 4 lateral mac (Fig. 63B); Abd II with m3e mac (Fig. 63D); unguis a.t. absent (Fig. 65E)............................................................................. 3</p><p>- Basomedian labial field with chaeta E ciliated (Fig. 68D); Th III with 2 lateral mac (Fig. 69B); Abd II m3e mac absent (Fig. 69D); unguis a.t. present (as Fig. 60E)........................................................ L. briani sp. nov.</p><p>3 Body with pigments (Figs 61, 72); Th III a5 always as mic (Fig. 74B), Abd I with 4 or less central mac (Fig. 63C, 74C).... 4</p><p>- Body pale (Fig. 77); Th III with a5 mac (Fig. 79B), Abd I with 5 central mac Fig. 79C).................. L. iegoi sp. nov.</p><p>4 Femur I–III completely pigmented (Fig. 61); Abd I often with 3 central mac, rarely with 4 mac (a2 often mic) (Fig. 63C); collophore anteriorly with 6 distal mac (Fig. 66A)................................................ L. andevo sp. nov.</p><p>- Femur I–III distally pigmented (Fig. 72); Abd I often with 2 central mac, rarely with 1 mac (m3 often mac) (Fig. 74C); collophore anteriorly with 2 distal mac (Fig. 76E).......................................... L. betamponensis sp. nov.</p><p>Descriptions of intermediary stages of Lepidocyrtinus</p><p>The juvenile’s description of Lepidocyrtinus is based on L. africanus sp. nov. and L. fabioi sp. nov., both probably in 3 rd instar due to their similar dorsal chaetotaxy compared to the same instar of Seira (Soto-Adames 2008; Zhang et al. 2019). In L. africanus it was studied the development of the colour pattern, antennal segments, clypeus, all head and dorsal trunk chaetotaxy, trochanteral organ, collophore and furcula. For L. fabioi only the dorsal chaetotaxy was studied, since other characteristics are similar to the adult. The morphology of both species is compared to 3 rd instar Seira dowlingi (Wray, 1953) and adult, and to L. barnardi adult (Soto-Adames 2008; Zhang &amp; Deharveng 2015a, 2015b; Zhang et al. 2019).</p><p>Colour pattern. Specimen with weak dark pigments on Ant II to IV, and Abd III with 1 lateral spot similar to adult (in coxa-shape) (Fig. 23). In Lepidocyrtinus this gain of pigments in postembryonic development must be common, as observed in other Entomobryidae (Pan et al. 2011, 2015, 2019; Cipola et al. 2017, 2018a).</p><p>Scales. Present on all head, dorsal and lateral thorax and abdomen; few scales on coxa and femur II–III, and manubrium and dens ventrally; antennae and collophore unscaled. Such displacement was observed in 3 rd instar S. dowlingi (Soto-Adames 2008) .</p><p>Antennal segments. Antennae smaller than trunk length (Fig. 23B); antennal segments ratio as I: II: III: IV = 1: 1.89: 2.56: 3.79. Ant I–IV chaetotaxy reduced. Ant IV weakly annulated with apical bulb apically unilobed. Ant III not annulated. Ant II with 2 bothriotricha and 1 median mac, sensilar chaetotaxy unclear. Ant I dorsally with 13 chaetae (2 lateral mac), 1 distal sens (not seen in adults), and typically with 3 smooth mic at the base, bothriotricha absent (Fig. 30A). This same bothriotricha pattern on Ant I–II was observed in 3 rd instar S. dowlingi (Soto-Adames 2008), and so there are no clear generic differences regarding this topic.</p><p>Clypeus (Fig. 30B). Clypeal formula as in the adult, with 4 (l1–2), 2 (ft) and 3 (pf0–1) ciliated chaetae, but l1–2 apically acuminate. It is most likely all these chaetae are primary and homologous to other Entomobryoidea (Zhang et al. 2016). Consequently, extra chaetae present in adults of some species of Lepidocyrtinus (Figs 14C, 19B, 62C, 68A, 73A, 78A), as well as in other Entomobryoidea in general, are secondary chaetae of difficult homology to determine. The emergency of extra secondary clypeal chaetae during the development was also reported to Symphypleona (Betsch &amp; Waller 1994).</p><p>Interocellar chaetae ‘IO’ (Fig. 30C). Interocellar field with 3 chaetae (p, r and t, p as mac), all primary according to Soto-Adames (2008). Therefore, the secondary chaetae (at least q and s) should appear in Lepidocyrtinus probably from the 4 th instar, as observed in Seira .</p><p>Head dorsal chaetotaxy (Fig. 30C). Differs from adult by 5 An, 3 M (m4i as mic), 3 Pp (Pp2 and Pp3), and 1 Pe mac (Pe3). In adult or any instar after the 3 rd, the An series has additions of mac, m4i turns into mac, and there is also the reduction of 3 mac (Pa2, Pa3 and Pe3), at least in L. africanus sp. nov. (Fig. 24F). In S. dowlingi, only occurs transformations of mic to mac during the development (Soto-Adames 2008), and therefore, such reductions of postoccipital mac in Lepidocyrtinus can be an exclusive feature in the Seirinae . In Entomobryinae genera as Sinhomidia Zhang, 2009 (in Zhang et al. 2009) it was also observed the reduction in some postoccipital mac of 1 st to adult, while in Homidia B̂rner, 1906 the secondary mac emerges during other instars’ development (Pan et al. 2011, 2019). In this sense, a study with the complete development (1 st to adult) needs to be done contemplating more Seira and Lepidocyrtinus species to make sure if there is a generic pattern.</p><p>Basomedian and basolateral labial fields (Fig. 30D). Similar to the adult (Fig. 24G), except by M2 and R chaetae absent. Such condition of the 3 rd instar is shared with Seira (Soto-Adames 2008) . Chaetae a1–5, M1 (as M), E and L1–2 are primary of Entomobryoidea, and it is common the emergence of extra chaetae (at least M2, R) during the 3 rd or 4 th instar (Soto-Adames 2008; Pan et al. 2011, 2019; Zhang &amp; Pan 2020).</p><p>Postlabial field (Fig. 30D). Formula of the main chaetae as in the adult (Fig. 24G), with 4 (G1–4), 3 (H2–4), 3 (J1–3) chaetae, except by a sole larger chaeta posteriorly, between J1 and J2. Therefore, the secondary chaetae in this region appear at least after the 3 rd instar. Next to J3 and J4 chaetae, postero-laterally, there are two unnamed chaetae both in the juvenile and the adult of L. africanus sp. nov., as well as in other species like L. botswanensis sp. nov., L. voeltzkowi, L. betamponensis sp. nov. and L. iegoi sp. nov., and even Seira and Tyrannoseira taxa ( Cipola et al. 2018b, 2018c; 2019). Of such chaetae, the GHJ series are, quite probably, primary and present in all Entomobryoidea, since they were also observed in the 1 st instar of Homidia and Sinhomidia (Pan et al. 2011, 2019). However, so far little is known about the development of the other postlabial setae, since the development of this region has never been properly studied.</p><p>Th II dorsal chaetotaxy (Figs 31A, 44A). In L. africanus sp. nov., the juvenile’s Th II a series has 6 mac (a5p–a5i3, a5i2p), and 6 mic on a5 group (a5i4, a5i3p– 6p, a5ip), the latter probably gradually turning into mac until the adult, as well as the appearance of extra chaetae (a5ip2, a5i2p2–p3, a5i5) (Fig. 25A). These chaetae are extra rows of a5i, which are shared among Lepidocyrtinus species (Figs 35A, 41A, 47A, 52A, 58A). So, they are probably homologous in most species, except for L. barnardi and L. dayi (Figs 15A, 20A) which are atypical concerning such multiplets (Figs 15A, 20A). Regardless, in all African Lepidocyrtinus species there are commonly 8 mac (except for L. fabioi sp. nov. with 6 mac) in two transvesal rows posteriorly on a5 group (e.g. Fig. 35A), which are absent in Seira and Tyrannoseira (Soto-Adames 2008; Cipola et al. 2014a, 2014b, 2018b, 2018 c, 2019). These transvesal rows are reduced and some chaetae do not show clear homologies in L. fabioi (Fig. 41A), so it is not clear if they represent the same seen in other species or emerged independently in the latter species. Juveniles of L. fabioi have almost the same pattern as the adults in a5 group, except for two chaetae as mic (a5p and another anteriorly of uncertain homology), that become mac in adults (Figs 41A, 44A).</p><p>The juvenile’s m and p series of L. africanus are quite similar to the adult, except by m1ip mac absent and only 4 mac (p1i–p2p) present on PmA group (Figs 31A, 25A). However, the juvenile of L. fabioi only hold primary chaetae as mac in m and p series, and p3 emerges first as a mic and becomes a mac in the adult together with five other secondary chaetae (m1i, m2i, m4p, p1i and p1p) (Figs 41A, 44A). In p series, p5 is always mic and sometimes p3 too, in Lepidocyrtinus adults (e.g. Figs 25A, 35A, 47A). Both chaetae are also mic in the 3 rd instar, at leats in L. africanus and L. fabioi (Figs 31A, 41A). In the 1 st instar of S. dowlingi and L. barnardi, such chaetae are mac (Soto-Adames 2008; Zhang et al. 2019), but p5 becomes a mic in adult L. barnardi (Fig. 15A). Further juveniles of different taxa of Lepidocyrtinus need to be investigated to see if p3 is always a mic in the 1 st instar or if changes its state (from mac to mic) at any instar in different species. Thus, notably the chaetotaxy final pattern of occurs only in adults of both genera.</p><p>Th III dorsal chaetotaxy (Figs 31A, 44A). The juveniles of the 3 rd instar are similar to the adults in L. africanus sp. nov. (Figs 24B, 31A) and L. fabioi sp. nov. (Figs 41B, 44A), except by p1i mac absent and m7 as mac (as mes in adult) in L. africanus, and a1a absent and p1p as mic in L. fabioi . Such data support that in the 3 rd instar of Lepidocyrtinus the diagnostic chaetotaxy pattern of each species is not yet fully defined, while in Seira such pattern is well established at the 2 nd instar, at least in S. dowlingi (Soto-Adames 2008) .</p><p>Abd I–III dorsal chaetotaxy (Fig. 44B). The 3 rd instar Abd I–II chaetotaxy are the same as the adult in L. africanus sp. nov. and L. fabioi sp. nov. (Figs 41 C–D, 44B), and such similarity is also shared in the development of S. dowlingi (Soto-Adames 2008) .</p><p>The Abd III of the juvenile of L. africanus is the same as in the adult, but in L. fabioi some modifications occur, as two mac are reduced in mes (a8 and p7), a9 and p8 mic and one mes of uncertain homology disappear, and em and emp change into accessory chaetae (Figs 42A, 44B). In S. dowlingi, the am6 mac change only in the 5 th instar (Soto-Adames 2008), while in Lepidocyrtinus such change occurs in the 3 rd instar, at least in L. africanus and L. fabioi .</p><p>Abd IV dorsal chaetotaxy (Figs 31B, 44C). The 3 rd instar of L. africanus sp. nov. has 6 central mic (Ae6, B2, B4, B6, C1, T1) which become mac in the adult (or any stage before) (Figs 26B, 31B). In addition, in lateral series 3 mes are modified into mac during the development (E4, Ee10, Fe3), while Fe3 changes from mac to mes. Also, two accessory chaetae arise surrouding T4 bothriotrichum (Pe and other unnamed) and at least 2 sens next B6 mac (as sens not observed). On the other hand, in L. fabioi sp. nov., the macrochaetotaxy pattern of the 3 rd instar is most similar to the adult, at least in the central region (Figs 42B, 44C). On lateral series they differ only by the alteration of E4 mes into mac, and the emergence of Fe3 mac and s accessory chaeta surrouding T2 bothriotrichum, during the development. On Abd IV posterior region, more two mes occur only on the adult L. africanus, but in L. fabioi the number of 8 posterior mes seen in the adult is already established at the 3 rd instar. A similar development of the posterior chaetae was recorded to S. dowlingi, but the adult’s number is already reached during the 2 nd instar (SotoAdames 2008).</p><p>In the 3 rd instar as well as in the adult of all Lepidocyrtinus species, there may be one or two secondary mac on central series (Si and Sm), which most likely appear during the 2 nd instar, as in most Entomobryoidea, including Seira (Soto-Adames 2008; Zhang et al. 2019). Consequently, these mac are not useful to separate Lepidocyrtinus from other Seirinae, but to support interspecific diagnoses, as other secondary chaetae (e.g. A3a/5, Ae4–7, Be2) which can be present or absent in different species (Figs 36B, 48B, 53B, 59B, 64B). Compared to the 3 rd instar of S. dowlingi, at least 8 chaetae (A5, B3, T7, E2, E4, E4p2, Ee10, F1) change into mac in the 4 th instar, plus 4 more mac on lateral series (F2p, F3p, Fe2, Fe3) arise, and in the 5 th instar 4 more accessory chaetae (s, m, Pi, Pe,) surrouding T2 and T4 bothriotricha emerge (Soto-Adames 2008). In this sense, Lepidocyrtinus and Seira development support that the final chaetotaxy pattern of Abd IV is only reached at the final instars, or even in the adult.</p><p>Legs (Fig. 31E). Subcoxa and tibiotarsus chaetotaxy reduced in the juveniles. Tibiotarsal outer chaetae not clearly elongated and with few chaetae, similar to the 1 st instar of L. barnardi (Zhang &amp; Deharveng 2015a) . Thus, the elongation of these chaetae must occur at the final instars. The trochanteral organ has 4 spine-like chaetae (Fig. 31C). Legs subsegments chaetotaxy show a gradual increase in complexity (number and shape of chaetae) until the adult in Lepidocyrtinus, and this probably occurs in all Entomobryoidea as well (Pan et al. 2011, 2019). Unguis and unguiculus morphology of juveniles are as in the adults; ratio unguis: unguiculus = 1: 0.65. The tibiotarsal smooth chaeta is 1.49 larger than unguiculus and the tenent hair is 1.28 larger than unguis outer edge.</p><p>Collophore (Fig. 31D). Anterior side with 6 chaetae, 2 spine-like chaetae and 1 proximal ciliated chaeta, 1 median ciliated chaeta apically acuminate and 1 mac and 1 mes distally; posterior side with 1 ciliated chaeta distally; lateral flap with 5 chaetae, 3 smooth and 2 ciliated. From these chaetae, at least 1+1 distal on posterior side and 2 smooth on lateral flap are probably primary, since they were also observed in the 1 st instar of Homidia and Sinhomidia (Pan et al. 2011, 2019).</p><p>Furcula (Figs 31 E–F). Manubrium ventral formula as in the adult, except by 8 apical chaetae plus only 1 subapical scale per side (Fig. 31E); manubrial plate with 5 ciliated chaetae (2 inner larger but not blunt) and 2 psp apparently (Fig. 31F). Dens dorsally with 2 proximal blunt mac weakly ciliated and apically bullet-like (Fig. 31F). It is still necessary to investigate further juveniles of Lepidocyrtinus in in the 1 st instar to clearly understand the origin and pattern of increase of furcal modified mac up to the adult. Our data suggests there is also an increase in the number of furcal chaetae during the development, both in the manubrium and dens, as well the change of states of at least part of them, similarly to what is seen in the legs subsegments.</p></div>	https://treatment.plazi.org/id/7B6687A7FFB5FFE9FF12D089FC2CF8B9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cipola, Nikolas Gioia;Morais, José Wellington De;Bellini, Bruno Cavalcante	Cipola, Nikolas Gioia, Morais, José Wellington De, Bellini, Bruno Cavalcante (2020): Review of Lepidocyrtinus Börner, 1903 (Collembola, Entomobryidae, Seirinae): the African species. Zootaxa 4898 (1): 1-110, DOI: 10.11646/zootaxa.4898.1.1
