taxonID	type	description	language	source
7B7E785C64649F1457EEFAD2FD17EF52.taxon	discussion	Remarks. Heleodromia immaculata was reported from Novaya Zemlya by Sack (1923), but this record needs to be confirmed. Species identification of Heleodromia is dependent on careful study of the complex male terminalia and the shape of the male tergite 8 (Sinclair et al. 2011).	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64649F1557EEF9B7FC5DE885.taxon	materials_examined	Material examined. RUSSIA. Chukotka AO (Wrangel Island): middle flow of river Neozhidannaya, 71 ° 01 ′ N 179 ° 09 ′ E, YPT, BT 4, 12 – 16. vii. 2016, LFV (1 ♂, ZIN). Recognition. Small greyish delicate flies (body length 3 – 3.5 mm) with very slender legs almost lacking distinct setae, eyes dichoptic in both sexes, separated by broad V-shaped flattened frons; legs black to reddish brown, “ knees ” often somewhat paler; habitually very similar to H. pullata (Fig. 2, see below). Male terminalia large, reniform; tergite 8 not broader than sternite 8, only produced medially as a shallow bilobed to forked projection and lacking posterolateral projection (Sinclair et al. 2011: 641, fig. 10); epandrial lobe slender, finger-like, bearing apical seta; surstylus slender, finger-like, with scattered setae, with small preapical thumb-like projection medially; apex with short dentiform process; phallus with slender, needle-like apex.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64649F1557EEF9B7FC5DE885.taxon	distribution	Distribution. Holarctic. The distribution of this species appears to be represented by widely disjunct populations exhibiting boreo-alpine connections in Europe and North America (Plant 2005; Plant et al. 2017; Sinclair et al. 2011). Our record of H. irwini from Wrangel Island may support this suggestion. In Eurasia, this species is known from Austria, Germany, Italy, Switzerland, Scotland and the Far East of Russia (Amurskaya and Magadanskaya Provinces); in North America, it is recorded from western part of Canada and USA. It seems that H. irwini is found here for the first time from the tundra zone. Habitat. The species was collected in wetlands in a river valley.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64659F1557EEF953FC06EED3.taxon	discussion	Remarks. The identification history of C. appendiculata is somewhat confusing (see Sinclair (1999) for details), consequently the record of this species from Novaya Zemlya (Sack 1923) requires confirmation. We expect this record to possibly be Clinocera aucta (Zetterstedt, 1849) (Sinclair 2008).	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64659F1557EEFD82FE68ECAA.taxon	description	(Fig. 2)	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64659F1557EEFD82FE68ECAA.taxon	materials_examined	Material examined. RUSSIA. Chukotka AO (Wrangel Island): Somnitelnaya Bay, on stony river shore, 23. vii. 1971, KBG (1 ♂, ZIN); upper flow of river Khischniki, 7 km SE Sovetskaya Mtn, 200 m, dryad tundra near river, 11. vii. 1972, KBG (3 ♂, 1 ♀, ZIN); middle flow of river Mamontovaya, 71 ° 10 ′ N 179 ° 45 ′ W, BT 2, YPT, 2 – 4. vii. 2015, OAK (2 ♂, 2 ♀, ZIN); same locality, 71 ° 10 ′ N 179 ° 45 ′ W, BT 3, Sw, 1. vii. 2015, OAK (1 ♀, ZIN); upper flow of river Neizvestnaya, 71 ° 13 ′ N 179 ° 19 ′ W, BT 2 В, 8 – 29. vii. 2015, OAK (1 ♂, 1 ♀, ZIN); same locality, 71 ° 13 ′ N 179 ° 19 ′ W, BT 13, YPT, 6 – 15. vii. 2015, OAK (1 ♂, ZIN). Recognition. Small greyish delicate flies (body length 3 – 3.5 mm) with very slender legs almost lacking distinct setae, eyes dichoptic in both sexes, separated by broad V-shaped flattened frons; legs black to reddish brown, “ knees ” often somewhat paler (Fig. 2). Male terminalia large, reniform; tergite 8 simple, lacking dorsomedial elongation or process (Sinclair et al. 2011: 644, fig. 11 B); epandrial lobe slender, digitiform; surstylus slightly shorter than epandrial lobe, with apical third slender and digitiform; phallus with apex membranous and broad.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64659F1557EEFD82FE68ECAA.taxon	distribution	Distribution. Holarctic; in North America, H. pullata is widely distributed in northern Canada and Alaska, occurring on tundra and in boreal forests, with isolated populations in the Rocky Mountains of Arizona, Colorado and New Mexico (Sinclair et al. 2011); in Eurasia, it is known from Taymyr Peninsula and Magadanskaya Province of Russia. Among the Russian Arctic islands, H. pullata is recorded only from Wrangel Island. Habitat. This species was collected on Wrangel Island mainly in river beds, where it was found in biotopes with different moisture levels (also on stones). Family EMPIDIDAE	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C646A9F1B57EEF8DEFDCEEB80.taxon	description	(Fig. 3)	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C646A9F1B57EEF8DEFDCEEB80.taxon	materials_examined	Material examined. RUSSIA. Archangelskaya Prov. (Novaya Zemlya Archipelago): Southern coast near Cape Cherny, 21, 29. vii. 1911, Rusakov (1 ♂, 2 ♀, ZIN); Eastern coast, Karskie vorota — Matochkin Shar, viii – ix, Rusakov (1 ♂, 1 ♀, ZIN); Matochkin Shar, brook Nochuev, 22. v. 1925 (1 ♀, ZIN), Vakulenko; Matochkin Shar, 10. vii. 1925, Vakulenko (1 ♀, ZIN). Chukotka AO (Wrangel Island): SW mountain Sovetskaya, upper flow of Khrustalnaya River, under snowfield on detritus, 3. viii. 1971, KBG (1 ♀, ZIN); Mineev Mtns, 5 km N Somnitelnaya, slope, swamp spring, 26. vii. 1971, KBG (1 ♂, 1 ♀, ZIN); lower flow of Khischniki River, foothills of Mineev Mtns, swamp tundra, 11. vii. 1972, KBG (1 ♂, ZIN); Rogers Bay, small swamp, on surface of water, 20. vii. 1971, KBG (2 ♂, 1 ♀, ZIN). Recognition. This species of Clinocera is readily distinguished by the presence of an auxiliary crossvein between R 2 + 3 and R 4, lower half of face with silvery pruinescence and a dark ring around the anterior spiracle (Fig. 3).	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C646A9F1B57EEF8DEFDCEEB80.taxon	distribution	Distribution. A Holarctic species distributed across the subarctic and arctic areas of Eurasia and North America (Sinclair 2008). In Russia, C. nivalis is known from Murmanskaya Province, Novaya Zemlya, north of Yakutia, Chukotka and Wrangel Island. Habitat. On Wrangel Island, this species was collected in various wet habitats. Genus Trichoclinocera Collin	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C646B9F1B57EEFCDAFAC3EF6D.taxon	description	(Figs 4, 5)	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C646B9F1B57EEFCDAFAC3EF6D.taxon	materials_examined	Material examined. CANADA. Quebec [new records since Sinclair (1994)]: 10 km SW Tasiujaq, trib. of Finger River, 17. viii. 1996, B. J. Sinclair (7 ♂, CNC). RUSSIA. Chukotka AO (Wrangel Island): middle flow of Neozhidannaya River, 71 ° 01 ′ N, 179 ° 09 ′ W, BT, YPT, 8 – 9. vii. 2019, OAK (1 ♀, ZIN); Somnitelnaya River, on water, water temperature + 5, air temperature + 2.5, 3. viii. 1971, KBG (10 ♂, 3 ♀, ZIN); same locality, under pebble near water, 3. viii. 1971, KBG (2 ♂, 1 ♀, ZIN); SW Sovetskaya Mtn, upper flow of Khrustalnaya River, 3. viii. 1971, KBG (2 ♂, 12 ♀, ZIN); same locality, snowfield, on water, air temperature + 3, water temperature + 7, snowfall, 4. viii. 1971, KBG, Chelnokov (1 ♂, 7 ♀, ZIN); environs of Tundrovaya Mtn, 71 ° 17.863 ′ N, 179 ° 47.875 ′ W, BT 2, YPT, 7 – 11. vii. 2019, U. V. Babiy (1 ♀, ZIN); middle flow of Mamontovaya River, N Perkatkun, on emergent stone in river, 29. vii. 1972, KBG (3 ♂, 1 ♀, ZIN). Recognition. Among clinocerines, Trichoclinocera is distinguished by the presence of setae on vein R 1. This species is distinguished readily by its dorsoventrally flattened tarsomeres 4 and 5, convex face and small eyes (Figs 4, 5).	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C646B9F1B57EEFCDAFAC3EF6D.taxon	distribution	Distribution. Holarctic species known from northern Finland and Sweden as well as it is widespread throughout the arctic region of North America (Sinclair 1994); recorded for the first time from the territory of Russia. Habitat. On Wrangel Island, this species was collected mainly on pebble floodplains, near water.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C646B9F1C57EEF92FFA56EE94.taxon	description	(Figs 6, 7) urn: lsid: zoobank. org: act: 1 E 05618 C-B 300 - 4 E 56 - B 381 - 1 A 0865 B 7462 E	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C646B9F1C57EEF92FFA56EE94.taxon	materials_examined	Type material. HOLOTYPE ♂, labelled: [RUSSIA. Chukotka AO (Wrangel Island)] “ [printed in Cyrillic, Russian] Chukotskiy AO [= Autonomous Okrug] o. [= ostrov, island] Wrangel / Verch. tech. [= verkhnee techenie, upper flow] river Neizvest- / naya 71 ° 13 ′ N 179 ° 19 ′ W / O. A. Khruleva 8.7.2015 / 2 B ”; “ Holotypus / Empis / septentrionalis / Shamshev, Sinclair sp. nov. [red label] ” (INS _ DIP _ 0000612, ZIN). PARATYPES: RUSSIA. Chukotka AO (Wrangel Island): same locality as holotype, BT 7 A, 5. vii. 2006, OAK (1 ♂, 1 ♀, CNC); same locality as holotype, BT 7 A, 10. vii. 2015, OAK (1 ♂, ZIN); same locality as holotype, BT 7 B, 10. vii. 2015, OAK (7 ♀, ZIN); same locality as holotype, BT C 1, YPT, 4 – 15. vii. 2015, OAK (1 ♂, ZIN); same locality as holotype, BT 1 A, 6 – 13. vii. 2015, OAK (1 ♂, ZIN); Mineev Mtns, 71 ° 00 ′ N 179 ° 31 ′ W, BT 14 B, 14. vii. 2006, OAK (1 ♂, ZIN); environs of Pervaya Mtn (71 ° 10 ′ N 179 ° 27 ′ W), BT A, hand collecting, 21. vii. 2019, OAK (1 ♀, ZIN).	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C646B9F1C57EEF92FFA56EE94.taxon	diagnosis	Diagnosis. Mid-sized (wing length 5 – 5.5 mm) robust flies with brownish pruinescent thorax and abdomen and almost entirely dark brown legs. Male dichoptic; occiput slightly convex with numerous long, flattened setae laterally behind eye; thorax black setose, scutum with 4 vittae; legs with only hind tarsomeres 1 – 3 brownish yellow, hind femur somewhat thickened, hind tibia very slender; apical part of phallus uniformly tubular. Female with brownish infuscate wing; abdomen densely brown pruinescent on tergites dorsally.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C646B9F1C57EEF92FFA56EE94.taxon	description	Description. Body length 5.2 – 5.9 (holotype 5.3); wing length 5 – 5.5 (holotype 5.1) mm. Male (Fig. 6). Head with dense brownish pruinescence on face, frons, postgena, ocellar triangle and occiput; entirely dark setose; clypeus shiny. Dichoptic; ommatidia equally small. Frons broad, below ocellar tubercle broader than distance between outer margins of posterior ocelli; with marginal setulae. Face broad, bare. Ocellar triangle with several long fine setae. Occiput slightly convex laterally behind eyes; with numerous long mostly flattened setae on convex part, fine long postoculars and similar setae on upper part; postgena with numerous hair-like setae. Antenna black; scape short, slightly longer than pedicel, both with short setulae; postpedicel conical, with slightly concave ventral margin on apical part, nearly 2 X longer than wide; stylus rather long, slightly longer (1.3 – 1.4 X) than postpedicel basal width. Palpus dark (sometimes yellowish, 2 ♂); with several black moderately long fine setae. Proboscis with labrum mostly brownish, reddish-brown apically, about 1.5 X longer than head height. Thorax dark in ground-colour, brownish pruinescent; with black setation; scutum with 4 indistinct narrow blackish brown vittae (dorsal view). Proepisternum with tuft of numerous mostly long and slightly flattened setae (with some additional shorter fine setae) on lower section; upper proepisternum in front of spiracle bare. Prosternum bare. Antepronotum with numerous short fine setae. Postpronotal lobe covered with very numerous subequally long setae. Mesonotum with hardly differentiated setae; several long fine intra-alars just behind postpronotal lobe; supra-alar space with several similar setae before suture and less numerous setae behind suture, notopleuron with numerous long setae, some posterior setae somewhat stronger; 2 long strong pal (sometimes with additional setulae), 8 – 10 sctl; acr long, fine, arranged in 2 irregular rows, lacking on prescutellar depression; presutural dc similar to acr, multiserial, postsutural dc less numerous, 3 – 4 prescutellars longest. Laterotergite with numerous long fine setae. Anterior and posterior spiracles brown. Legs robust, almost entirely dark brown, only hind tarsomeres 1 – 3 (sometimes 1 – 4) brownish yellow; black setose; hind femur somewhat thickened, mid tibia slightly arcuate, hind tibia very slender. Coxae and trochanters with simple setae. Femora whitish pilose ventrally (less distinct on fore femur); fore femur with long fine setae anteroventrally, posteroventrally and posteriorly; mid femur with numerous spine-like setae of different lengths ventrally, posteroventrals longer, some moderately long anterodorsal setae; hind femur with numerous long flattened setae anterodorsally (except short subapical part), only pale pilose along lower half anteriorly (except several setae near extreme base), rows of long, strong anteroventral and posteroventral setae, short spine-like setae ventrally. Tibiae devoid of strong setae (including setae of subapical circlet); fore tibia with some short, fine setae posteriorly; mid tibia with erect setulae ventrally; hind tibia pale pilose ventrally, no seta in posteroapical comb. Hind tarsomeres with somewhat finer and sparser setulae. Wing membrane very faintly infuscate, with brownish veins; CuA + CuP (anal vein) complete; cell dm with elongate apex. Pterostigma brownish yellow. Basal costal seta absent. Anal lobe well-developed; axillary incision acute. Squama dirty yellow, brownish fringed. Halter with yellow knob and brownish stem. Abdomen dark, brownish pruinescent (somewhat denser on tergites laterally and sternites); covered with numerous long, dark fine setae (shorter on tergites dorsally). Segment 8 with tergite and sternite separated; tergite 8 mostly membranous, represented by two weakly sclerotized, narrow lateral sclerites each bearing 0 – 1 setae; sternite simple, nearly as long as sternite 7 (lateral view), with numerous long setae along posterior margin. Terminalia (Fig. 7). Cerci broadly yellowish to brownish yellow, narrowly brownish along upper margin and anteriorly; separated from each other and from epandrium; cercus elongate oval (lateral view), with rounded apex, slightly extended beyond apex of epandrium, covered with short dark setae. Epandrium almost entirely brownish, narrowly yellowish brown along lower margin; subtriangular (lateral view), with dark setae more numerous and longer along lower margin. Hypandrium mostly membranous, rim-like sclerotized along upper margin, bare. Phallus almost uniformly broad, slightly narrowed near middle, S-like bend, sulcate on subapical portion, with pair of small projections near apex beneath. Ejaculatory apodeme extended far beyond basal curvature of phallus, with subequal lateral and vertical wings and somewhat narrower lower wing. Female. Occiput not convex laterally, without flattened setae, ocellar tubercle with short setae; palpus somewhat paler, yellowish brown, sometimes with yellowish apex or yellowish. Thorax and legs with shorter setation; hind femur not thickened, hind tibia simple; hind femur without flattened setae and pale pilosity along lower half anteriorly, entirely covered with setulae anteriorly; fore tibia without fine setae posteriorly, mid and hind tibiae with usual setulae ventrally; hind tarsus brownish, with similar setation to fore and mid tarsi. Wing darker, brownish infuscate, pterostigma brownish. Abdomen densely brownish grey pruinescent on tergites laterally and sternites, brown pruinescent on tergites dorsally (except noted); tergites 7 – 8 extensively shiny (except posterior margin), also sternites 7 – 8 usually subshiny to shiny anteriorly; only tergites 1 – 2 with some short setae laterally, otherwise tergites covered with scattered setulae; cercus black, with dark setulae.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C646B9F1C57EEF92FFA56EE94.taxon	distribution	Distribution. Palaearctic, known only from Wrangel Island.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C646B9F1C57EEF92FFA56EE94.taxon	etymology	Etymology. The specific name is from the Latin septentrionalis (north, northern), in reference to the distribution of this species.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C646B9F1C57EEF92FFA56EE94.taxon	discussion	Remarks. Empis septentrionalis sp. nov. belongs to the group of species with uncertain subgeneric position within Empis (Shamshev 2001 a). The group is known almost exclusively from the Asiatic part of the Palaearctic (except one species) and is especially diverse in Middle Asia. In addition, three undescribed species are known from Yukon in Canada. The new species is very similar to E. jacutiensis Shamshev occurring in Yakutia and Chukotka of Russia (Shamshev 2016). Empis septentrionalis sp. nov. can be readily distinguished from E. jacutiensis by uniformly tubular apical part of the phallus (cup-shaped apically in E. jacutiensis) (Fig. 8) and dark brown hind tibia (yellowish to brownish yellow in E. jacutiensis). In addition, the male of E. septentrionalis sp. nov. usually has dark palpi (yellowish in E. jacutiensis), but this character appears to be somewhat variable. The female of the new species differs from the female of E. jacutiensis by dark brown hind tibia and tarsus and by brown pruinescence on abdominal tergites dorsally (abdomen uniformly light grey pruinescent in the female of E. jacutiensis). Habitat. Almost all specimens were collected in the warmer central region of Wrangel Island, mainly in dry habitats. Genus Rhamphomyia Meigen	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C646D9F0257EEFF53FC18ECED.taxon	description	(Fig. 9)	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C646D9F0257EEFF53FC18ECED.taxon	materials_examined	Material examined. RUSSIA. Chukotka AO (Wrangel Island): Somnitelnaya Bay, valley of river Somnitelnaya, Mineev Mtns, 150 m, shingle, on flower of Potentilla emarginata, 22. vii. 1966, KBG (1 ♀, ZIN); same locality, on inflorescence of Polygonum ovalifolium, 19. vii. 1966, KBG (1 ♂, ZIN); same locality, on flower Dryas integrifolia, 19, 22. vii. 1966, KBG (2 ♂, ZIN); 5 km N Somnitelnaya Bay, valley of river Somnitelnaya, foothills, 150 m, 25. vii. 1972, KBG (7 ♂, 11 ♀, ZIN); near north slope of Berri Peak, 6 km ESE mountain Sovetskaya, meadow near stream, 300 m, 12. vii. 1972, KBG (1 ♂, ZIN); upper flow of river Khischniki, 7 km SE mountain Sovetskaya, Dryas tundra near river, 200 m, 11. vii. 1972, KBG (1 ♀, ZIN); same locality, meadow near stream, 200 m, 12. vii. 1972, KBG (1 ♂, ZIN); same locality, on Dryas, 14. vii. 1972, KBG (1 ♂, 1 ♀, ZIN); same locality, 15. vii. 1972, KBG (1 ♂, 3 ♀, ZIN); basin of river Khischniki, 8 km SE mountain Sovetskaya, meadow near stream, 250 m, 13. vii. 1972, KBG (8 ♂, 16 ♀, ZIN); environs of mountain Tundrovaya, near stream, 18. vii. 1972, KBG (13 ♂, 7 ♀, ZIN); same locality, BT 3, PT, 1 – 19. vii. 2015, LFV (1 ♀, ZIN); middle flow of river Mamontovaya, N Perkatkun, osier-bed in river valley, 17,21,29. vii. 1972, KBG (10 ♂, 1 ♀, ZIN); same locality, 71 ° 10 ′ N 179 ° 45 ′ W, BT 2 ’, 8. vii. 2006, OAK (1 ♂, 1 ♀, ZIN); same locality, BT 14, 9. vii. 2006, OAK (1 ♀, ZIN); same locality and biotope, Sw, 2. vii. 2015, OAK (2 ♂, 1 ♀, ZIN); same locality, Sw, BT 20 A, 1. vii. 2015, OAK (1 ♀, ZIN); same locality and biotope, YPT, 2 – 4. vii. 2015, OAK (2 ♂, ZIN); same locality and biotope, PT, 1 – 23. vii. 2015, OAK (1 ♀, ZIN); same locality, Sw, BT 1 A, 2. vii. 2015, OAK (1 ♀, ZIN); same locality, litter, BT 7, 1. vii. 2015, OAK (1 ♂, ZIN); same locality, Sw, BT 6, 2. vii. 2015, OAK (1 ♂, ZIN); same locality, Sw, BT 11, 30. vi. 2015, OAK (1 ♀, ZIN); same locality, Sw, BT 5, 30. vi. 2015, OAK (1 ♀, ZIN); same locality, in house, 2. vii. 2015, OAK (2 ♂, 1 ♀, ZIN); upper flow of river Neizvestnaya, 71 ° 13 ′ N 179 ° 19 ′ W, BT 7 А, 5. vii. 2006, OAK (4 ♂, 5 ♀, ZIN); same locality, Sw, BT 4 A, 22. vi. 2015, OAK (1 ♀, ZIN); spures of Pervaya Mtn, Sw, BT 11, 28. vi. 2015, OAK (2 ♂, 1 ♀, ZIN); same locality, Sw, BT 13, 28. vi. 2015, OAK (1 ♂, ZIN); middle flow of river Neizvestnaya, 71 ° 20.287 ′ N, 179 ° 29.779 ′ W, BT 1, YPT, 4 – 5. vii. 2019, U. V. Babiy (1 ♂, ZIN).	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C646D9F0257EEFF53FC18ECED.taxon	description	Recognition. Large (wing length 6 – 7 mm) species with setose prosternum; thorax and abdomen covered with long pale setae, legs long slender. Male (Fig. 9) holoptic; scutum mostly subshiny (dorsal view); mesonotal setae mostly hair-like, 4 – 6 strong npl, at least 10 sctl, acr biserial, presutural dc multiserial; legs dark, fore basitarsus distinctly thickened (slightly broader than tibia width at apex), with long setae (except dorsal and ventral face), hind basitarsus usually slightly thickened but often quite slender, with long setae dorsally (more numerous and longer in southern populations); wing infuscate, basal costal seta absent, CuA + CuP reaching wing margin but sometimes weakened on about middle, halter with yellowish knob; abdomen with faintly pruinescent tergites dorsally; segments 6 – 8 modified, tergite 6 broadened, deeply concave posteriorly, sternite 6 inflated, with dense transverse cluster of strong golden setae, sternite 7 with pair of horn-like projections ventrally. Female legs without pennate setae; wing brownish, broadened.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C646D9F0257EEFF53FC18ECED.taxon	distribution	Distribution. Holarctic; in North America, R. albopilosa is widespread across the low arctic, with isolated southern records (Sinclair et al. 2019). In Eurasia, the species occurs in Eastern and Western Siberia (Shamshev 2016). Among the Russian Arctic islands, R. albopilosa is known only from Wrangel Island.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C646D9F0257EEFF53FC18ECED.taxon	biology_ecology	Habitat. On Wrangel Island this species is found in various biotopes.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64729F0357EEFA2AFBD1E8F9.taxon	description	(Fig. 10)	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64729F0357EEFA2AFBD1E8F9.taxon	materials_examined	Material examined. RUSSIA. Archangelskaya Prov. (Novaya Zemlya Archipelago): Beluzhja Bay, 5. vii. 1904, Sokolovsky (1 ♂, 1 ♀, ZIN); foot of Pila Mtn, Matochkin Shar, 9. viii. 1901, Timofeev (1 ♂, ZIN); Matochkin Shar, winter hut Razmyslova, 28. viii. 1923, Lozina-Lozinsky (1 ♂, 1 ♀, ZIN); Matochkin Shar, brook Nochuev, 18 – 19. vii. 1925, Vakulenko (18 ♂, 10 ♀, ZIN). Krasnoyarskiy Terr. (Dikson Island): wet marsh in lowland between hills (some specimens on Dryas punctata), 28 – 30. vii. 1948, Korotkevich (6 ♂, 5 ♀, ZIN).	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64729F0357EEFA2AFBD1E8F9.taxon	description	Recognition. Large, blackish brown species (wing length 6 – 6.5 mm). Male (Fig. 10) eyes dichoptic, separated by very narrow frons; thorax black setose, scutum densely brownish pruinescent, without distinct vittae; all mesonotal setae strong, bristle-like, long, presutural dc 2 – 3 - serial, acr biserial, 10 – 12 sctl; legs robust, entirely dark brown, hind femur entirely setose, hind tibia slender; wing brownish infuscate, halter brown; cerci directed anteriorly, short, prolonged nearly to middle of tergite 5, epandrium brownish to yellowish brown. Female legs not pennate.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64729F0357EEFA2AFBD1E8F9.taxon	distribution	Distribution. Eurasia. This species is known only from the Russian Arctic territories, including Novaya Zemlya Archipelago, Taymyr Peninsula (Shamshev & Sinclair 2018) and Dikson Island.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64739F0357EEFE1EFD40EEF5.taxon	description	(Fig. 11)	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64739F0357EEFE1EFD40EEF5.taxon	materials_examined	Material examined. RUSSIA. Archangelskeya Prov. (Novaya Zemlya Archipelago): Beluzhya Bay, 5. vii. 1904, Sokolovskij (3 ♂, 1 ♀, ZIN); foot of Pila Mtn, Matochkin Shar, 9. viii. 1901, Timofeev (1 ♂, 1 ♀, ZIN). Yakutia (New Siberian Islands): centre of Kotelny Island, upper flow of river Balykhtakh near mouth of river Tuguttakh, stream valley on flowers of Caltha, 15 – 16. vii. 1973, KBG (34 ♂, 18 ♀, ZIN); same locality, arctic desert on flowers, 15. vii. 1973, KBG (1 ♂, 1 ♀, ZIN). Chukotka AO (Wrangel Island): middle flow of Neozhidannaya River, 71 ° 01 ′ N 179 ° 09 ′ E, BT 1, YPT, 12 – 16. vii. 2016, LFV (1 ♀, ZIN); same locality, BT 3, YPT, 12 – 16. vii. 2016, LFV (1 ♂, 8 ♀, ZIN); same locality, BT 4, YPT, 12 – 16. vii. 2016, LFV (4 ♀, ZIN); same locality, BT A, Sw, 8. vii. 2019, OAK (1 ♀, ZIN); same locality, BT W, Sw, 8. vii. 2019, OAK (1 ♂, ZIN); same locality, BT B, Sw, 8. vii. 2019, OAK (1 ♀, ZIN); same locality, BT G, Sw, 8 – 9. vii. 2019, OAK (3 ♀, ZIN); 2 km W Somnitelnaya Bay, spotted tundra, 9. vii. 1972, KBG (3 ♂, 6 ♀, ZIN); Somnitelnaya Bay, on flowers of Dryas integrifolia, 21 – 26. vii. 1966, KBG (12 ♂, 16 ♀, ZIN); near northern slope of Berri Peak, 6 km ESE Sovetskaya Mtn, 300 m, swamp on slope, 12. vii. 1972, KBG (1 ♂, ZIN); Sovetskaya Mtn, snowfield near top, 1000 m, 14. vii. 1972, KBG (1 ♀, ZIN); SE part of island, tundra, 2. viii. 1931 & 26. vii. 1933, А. Mineev (1 ♂, 1 ♀, ZIN); Rogers Bay, 20. vii. 1932, А. Mineev (6 ♂, 4 ♀, ZIN); same locality, 19. vii. 1972, KBG (1 ♀, ZIN); same locality, 70 ° 59 ′ N 178 ° 29 ′ W, BT 1, YPT, 10 – 13. vii. 2019, OAK (1 ♂, 1 ♀, ZIN); same locality, Ushakovskoe vill., 70 ° 59 ′ N 178 ° 30 ′ W, BT A, YPT, 10 – 13. vii. 2019, OAK (2 ♀, ZIN); environs of Tundrovaya Mtn, tundra and near stream, 18. vii. 1972, KBG (2 ♂, 1 ♀, ZIN); same locality, 71 ° 18.469 ′ N 179 ° 44.327 ′ W, BT 3, 5 – 15. vii. 2018, U. V. Babiy (1 ♂, ZIN); same locality, 71 ° 17.788 ′ N 179 ° 47.81 ′ W, BT 5, 15 – 26. vii. 2018, U. V. Babiy (1 ♂, ZIN). USA. Alaska: Barrow, ex. Dryas, 18 – 26. vii. 1955, sweeping arctic tundra (13 ♂, 10 ♀, CNC); Inaru River, 25 – 28. vii. 1956; ex. Polygonum and dandelion flowers (6 ♂, 1 ♀, CNC). Recognition. Mid-sized blackish species (wing length about 5 mm). Male (Fig. 11) dichoptic, frons very broad; thorax black setose, scutum densely brownish pruinescent, without distinct vittae; all mesonotal setae fine, hair-like, long, presutural dc multiserial, acr 2 – 4 - serial, at least 10 sctl; legs robust, entirely dark brown, all tibiae only pilose ventrally, hind femur without setae on about apical 3 / 4 ventrally, only pale pilose, hind tibia very slender; wing faintly infuscate, halter brown; cerci prolonged anteriorly, short, extended nearly to middle of tergite 5, epandrium brown. Female legs not pennate.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64739F0357EEFE1EFD40EEF5.taxon	distribution	Distribution. Holarctic species widely distributed across extreme northern Russia, including Novaya Zemlya Islands, Taymyr Peninsula, New Siberian Islands and Wrangel Island. In North America, R. erinacioides is recorded on Barter Island in Camden Bay, Alaska, the nearby mainland coast (Sinclair et al. 2019) and newly recorded here from the region surrounding Barrow. Habitat. On Wrangel Island, in the 1960 s and 70 s this species was most numerous on the coastal Southern plane; in the mountains at around 1000 m. In the 2000 s, the greatest abundance of this species was observed in areas with more severe climatic conditions; e. g., in the northern variant of the Arctic tundra subzone (Neozhidannaya River), where it was found in various habitats, including zonal spotted tundra. In the warmer central region of the island, only a single specimen was captured.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64709F0757EEFA3CFEF8ED94.taxon	description	(Figs 14 – 19) urn: lsid: zoobank. org: act: 34153559 - 3823 - 4932 - A 856 - 693999 B 0 C 7 A 4	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64709F0757EEFA3CFEF8ED94.taxon	materials_examined	Type material. HOLOTYPE ♂, labelled: [RUSSIA, Chukotka AO:] “ [in Cyrillic, Russian] Apapelkhin, 15 km / NOO Pevek, Chukot. [= Chukotka] / Gorodkov 28. vi. [1] 963 ”; “ shrubby tundra ”; “ Holotypus / Rhamphomyia / gorodkovi Shamshev, Sinclair, Saigusa sp. nov. [red label] ” (INS _ DIP _ 0000613, ZIN). PARATYPES: RUSSIA. Chukotka AO (Wrangel Island): upper flow of river Neizvestnaya, 71 ° 13 ′ N 179 ° 19 ′ W, BT 3 G, 4. vii. 2006, OAK (1 ♂, ZIN); same locality, 71 ° 12.96 ′ N, 179 ° 20.09 ′ W, 125 m, BT 12, Sw, 24. vi. 2015, OAK (1 ♀, ZIN) same locality, 71 ° 13.02 ′ N, 179 ° 19.508 ′ W, 120 m, BT 3, Sw, 21. vi. 2015, OAK (1 ♂, ZIN). Chukotka mainland: same locality as holotype, 28. vi. 1963, KBG (10 ♂, 6 ♀, ZIN; 2 ♂, 1 ♀, CNC); Iultin, waterlogged sedge tundra, 21. vii. 1963, KBG (3 ♂, 2 ♀, ZIN); Krasnoarmeiskiy, Chaunskiy District, osier-bed in valley, 8. vii. 1963, KBG (1 ♂, ZIN); 24 km SE Pevek, 150 m, osier-bed in valley, 1. vii. 1963, KBG (1 ♂, ZIN); valley of river Ichuvisi, Komsomolskiy mine, Chaunskiy District, shrubby tundra, 5. vii. 1963, KBG (3 ♂, 2 ♀, ZIN); Komsomolskiy mine, Chaunskiy District, tundra, 4. vii. 1963, KBG (17 ♂, 3 ♀, ZIN); Schmidt, southern slope, tundra with willow, 18. vii. 1963, KBG (1 ♀, ZIN); 20 km SSE Iultin, river Amguema, bush of Alnaster fruticosa, 22. vii. 1963, KBG (3 ♂, ZIN); Schmidt, on snowfield, 16. vii. 1966, KBG (1 ♂, ZIN); Schmidt, 5 km SW of village, southern slope, tundra with willow, 11. vii. 1971, KBG (3 ♂, 3 ♀, ZIN). Krasnoyarskiy Terr.: Taymyr, village Agapa, Pyasina, moist shrubby tundra, 14. vii. 1967, KBG (1 ♀, ZIN). Yakutia: Chokurdakh, Indigirka River, river bank, slope, shrubby tundra, 12. vii. 1966, KBG (1 ♀, ZIN); lower flow of Indigirka River, 71 ° N, border of tundra and forest-tundra, 20. vi. 1973, V. Flint (2 ♂, 1 ♀, ZMMU; 1 ♂, 1 ♀, CULSP). USA. Alaska: Cape Thompson, 29. vii. 1960, H. E. Erdman (1 ♀, CNC). Additional material. RUSSIA. Chukotka AO: Wrangel Island: middle flow of river Neozhidannay, 71 ° 01 ′ N 179 ° 09 ′ E, BT A, YPT, 8 – 9. vii. 2019, OAK (1 ♂, ZIN); same locality, BT B, Sw, 8 – 9. vii. 2019, OAK (1 ♂, ZIN); same locality, BT W, YPT, 8 – 9. vii. 2019, OAK (1 ♂, ZIN); same locality, BT G, 8 – 9. vii. 2019, OAK (5 ♂, 4 ♀, ZIN); same locality, YPT, BT 4, 12 – 16. vii. 2016, LFV (2 ♂, 3 ♀, ZIN); same locality, YPT, BT 1, 12 – 16. vii. 2016, LFV (1 ♂, ZIN); same locality, YPT, BT 3, 12 – 16. vii. 2016, LFV (2 ♂, 2 ♀, ZIN).	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64709F0757EEFA3CFEF8ED94.taxon	diagnosis	Diagnosis. A large species (body about 6.5 mm) with body and abdomen densely light grey pruinescent, halteres yellow, acrostichals multiserial. Male: holoptic, legs uniformly brownish, hind femur with numerous short setae over entire length ventrally, hind tibia thickened towards apex; wing whitish; cerci short, extended only to middle of tergite 6. Female: legs extensively yellowish, with simple setae, hind tibia broadened toward apex, wings faintly infuscate.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64709F0757EEFA3CFEF8ED94.taxon	description	Description. Male (Fig. 14). Body length 6.1 – 6.4 (lectotype 6.3); wing length 6.0 – 6.2 (lectotype 6.2) mm. Head with dense light greyish pruinescence on face, frons, postgena, ocellar triangle and occiput; dark setose. Holoptic, eye with upper ommatidia enlarged. Frons represented by very small triangular space below ocellar tubercle and larger subtriangular space above antennae, with scattered marginal setulae above antennae. Face parallel-sided, bare. Ocellar triangle with several fine setae of different lengths. Occiput with numerous moderately long setae laterally, including postoculars; postgena with numerous finer setae than occiput. Antenna with scape and pedicel brownish, postpedicel somewhat darker, blackish brown; scape and pedicel short, scape slightly longer, both with short setulae; postpedicel conical, 2.4 – 2.6 longer than wide; stylus short, 2.5 – 2.6 times shorter than postpedicel. Palpus dark; with long, dark fine setae. Proboscis with labrum dark reddish brown, 1.2 – 1.3 X head height. Thorax dark in ground-colour, largely densely light grey pruinescent; with black setation. Scutum almost uniformly light grey, with 2 indistinct, narrow, brownish grey vittae along dc rows (dorsal view), sometimes, hardly visible vitta along acr rows present. Proepisternum with tuft of numerous long setae on lower section; upper proepisternum in front of spiracle with 3 – 5 fine setae. Prosternum bare. Antepronotum with numerous short setae. Postpronotal lobe with numerous almost uniform fine setae. Mesonotum mostly with fine setae, including numerous undifferentiated similar presut spal, npl and psut spal (some posterior npl and psut spal somewhat longer and stronger), 2 – 3 pal with several shorter setae (number and robustness variable); 6 – 10 of subequally long sctl (sometimes with additional shorter setae); acr short, multiserial, absent on prescutellar depression; presutural dc short and multiserial, postsutural dc less numerous, long and arranged in 2 – 3 irregular rows. Laterotergite with numerous long fine setae. Anterior and posterior spiracles yellowish. Legs almost uniformly brownish, only extreme base of tibiae somewhat yellowish, mostly subshiny, coxae and trochanters with denser greyish pruinescence; black setose. Femora slender, of subequal width; mid tibia slightly stouter than fore tibia, hind tibia evenly thickened toward apex; all tarsomeres slender. Mid and hind femora whitish pilose ventrally (more distinctly on hind femur); fore and mid femora clothed in moderately long, thin setae, mid femur with more numerous and denser anteroventral setae near base; mid femur with similar setation but anteroventral setae more numerous and denser near base; hind femur entirely covered with setulae ventrally, bearing complete rows of short setae anteroventrally (shorter than half of femur width). All tibiae covered with moderately long, thin setae posterodorsally (at most as long as corresponding tibia width), without strong setae; no seta in posteroapical comb of hind tibia. All tarsomeres covered with short, simple setae (except somewhat longer setae of subapical circlet). Wing membrane somewhat whitish, mostly with brownish veins; all veins complete (except Sc); CuA + CuP (anal vein) mostly weakened but short subapical portion normally sclerotised. Pterostigma brownish yellow; basal costa seta absent. Anal lobe well-developed; axillary incision acute but almost 90 °. Squama yellow, dark fringed. Halter yellow. Abdomen dark, densely light grey pruinescent (concolorous with thorax); covered with black setae longer on tergites laterally and shorter on tergites dorsally. Segments before segment 8 unmodified. Segment 8 with tergite and sternite separated. Tergite 8 strongly upturned, laying almost vertically to body axis, with posterior margin slightly recurved, with numerous setae along posterior margin. Sternite 8 simple, scoop-like, with numerous long setae along posterior margin. Terminalia (Figs 15 – 18). Cerci yellowish; fused to epandrium (but suture distinct), broadly concave and flattened medially, strongly prolonged anteriorly, forming small subglobular window-like space subapically (dorsal view), without pair of subepandrial lobes beneath; apical part of cercus (before epandrium) short, extended only to tergite 6, subtriangular viewed laterally, rounded apically, covered with dark setulae; posterior part of cercus broadly digitiform, rounded apically, only slightly extended beyond epandrium, with numerous dark spinule-like setulae ventrally. Epandrium usually brownish, sometimes yellowish or reddish brown, faintly greyish pruinescent, mostly with short dark fine setae longer on posterior projection; rather subglobular (lateral view), with short, rounded apically projection on lower part posteriorly. Hypandrium yellowish, subtriangular (ventral view), entirely sclerotized, bare; gonocoxal apodeme directed posteriorly, large, broadened apically. Phallus (Fig. 18) yellowish; short, mostly hidden (lateral view); gently arched, broad on about basal half, becoming evenly slenderer beyond epandrial projection; with two minute lateral “ spinules ” on apical part. Ejaculatory apodeme extended far beyond basal curvature of phallus, with narrow lateral wings and very broad vertical wing. Female (Fig. 19). Body length 6.4; wing length 5.9 mm. Similar to male, except eyes dichoptic, ommatidia of equal size; frons broad, somewhat widened toward ocellar tubercle; mesonotum, legs and abdomen with distinctly shorter setation; legs paler, largely rather brownish yellow, with simple setae (sometimes posteroventral setae on mid and hind femora slightly flattened); wing membrane faintly brownish infuscate; abdominal tergite 8 pointed posteriorly; cercus black, long, slender, covered with dark setulae.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64709F0757EEFA3CFEF8ED94.taxon	distribution	Distribution. Holarctic; in Eurasia, the new species is broadly distributed across subarctic areas of Russia (from Taymyr to Chukotka). Among the Russian Arctic islands the species is known only from Wrangel Island. In North America, this species is known from Alaska and Yukon, but unknown from Canadian Arctic islands.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64709F0757EEFA3CFEF8ED94.taxon	etymology	Etymology. The specific name is a patronym in honour of the late Kirill Borisovich Gorodkov, Russian Dipterist (ZIN, St. Petersburg), who collected many of the specimens used in this study, as well as sorted and identified many Palaearctic Rhamphomyia housed in the ZIN.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64709F0757EEFA3CFEF8ED94.taxon	discussion	Remarks. The new species resembles strongly the Holarctic species R. villipes Coquillett, 1900 (known for a long time in Eurasia as R. hambergi Frey). However, R. gorodkovi sp. nov. can be readily distinguished from R. villipes primarily by the whitish wings of the male (hyaline to very faintly infuscate in R. villipes) and simple setose mid and hind legs of the female (mid, hind femora and hind tibia with pennate setae in R. villipes). In addition, in the male of R. villipes the legs are covered with long hair-like setae, fore and mid tibiae somewhat thickened and mid femur slightly arcuate. In the Chukotka Peninsula, both species appear to occur as sympatric populations. Among the material from North America, this species was recognised as Rhamphomyia (Dasyrhamphomyia) sp. 8 (Saigusa unpubl. data).	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64779F0457EEFAE3FBF2EA65.taxon	description	(Fig. 12)	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64779F0457EEFAE3FBF2EA65.taxon	materials_examined	Material examined. RUSSIA. Krasnoyarskiy Terr. (Dikson Island): small island “ Konus ”, 31. vii. 1948, Korotkevich (1 ♂, ZIN); marsh in lowland between hills, 30. vii. 1948, Korotkevich (1 ♀, ZIN). Chukotka AO (Wrangel Island): middle flow of Neozhidannaya River, 71 ° 01 ′ N 179 ° 09 ′ E, BT 4, YPT, 12 – 16. vii. 2016, LFV (1 ♀, ZIN); environs of Tundrovaya Mtn, tundra nesting-place of gooses, 18. vii. 1972, KBG (1 ♀, ZIN); middle flow of Mamontovaya River, N Perkatkun, waterlogged coast, 17. vii. 1972, KBG (2 ♂, 3 ♀, ZIN); same locality, small swamp, 29. vii. 1972, KBG (1 ♂, 3 ♀, ZIN); same locality, 71 ° 10 ′ N 179 ° 45 ′ W, BT 20, 12. vii. 2006, OAK (3 ♀, ZIN); same locality and biotope, Sw, 1. vii. 2015, OAK (2 ♂, 3 ♀, ZIN); upper flow of Neizvestnaya River, 71 ° 13 ′ N 179 ° 19 ′ W, BT 12, PT, 24. vi. – 6. vii. 2015, OAK (1 ♀, ZIN). Recognition. Large, blackish brown species (wing length 6.3 – 7 mm). Male (Fig. 12) eyes holoptic; thorax black setose, scutum densely brownish pruinescent, without distinct vittae; almost all mesonotal setae fine, hair-like (npl usually slightly stronger), long, presutural dc 3 – 5 - serial, acr 2 – 3 - serial; legs robust, entirely dark brown, hind femur entirely setose, hind tibia slender; wing brownish infuscate, halter brown; cerci directed anteriorly, short, prolonged nearly to middle of tergite 5, epandrium brownish. Female wing slightly darker and broader; mid femur with pennate posteroventral setae, hind femur with pennate anterodorsal and posteroventral setae, hind tibia with pennate setae dorsally and ventrally (on about middle).	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64779F0457EEFAE3FBF2EA65.taxon	distribution	Distribution. Holmgren (1880) described R. hovgaardii from Novaya Zemlya, however, later he noted also Vaygach Island (Holmgren & Aurivillius 1883; see Shamshev & Sinclair 2018). Holarctic species; in Eurasia, R. hovgaardii extends across arctic and subarctic Russia, including both continental areas (Taymyr, Yakutia, Chukotka) and islands (Novaya Zemlya, Vaygach, Wrangel). In North America, this species ranges in the low arctic from the western edge of Hudson Bay to Alaska (Sinclair et al. 2019). Habitat. On Wrangel Island, this species was collected mainly in wet habitats.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64749F0B57EEFCA2FDC9EBBD.taxon	description	(Figs 20 – 25) urn: lsid: zoobank. org: act: 26 E 7 A 46 E- 1 A 47 - 4193 - A 5 FF-FA 5618 C 8 E 98 E	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64749F0B57EEFCA2FDC9EBBD.taxon	materials_examined	Type material. HOLOTYPE ♂, labelled: [RUSSIA, Chukotka AO:] “ [in Cyrillic, Russian] Pevek [69 ° 42 ′ N 170 ° 19 ′ N], Chukotskij nats. [= natsionalny, national] okr. [= okrug, name of administrative unit] archadovaya / tundra. / Gorodkov, 28. vi. [1] 963 ”; “ Holotypus / Rhamphomyia / mallochi / Shamshev, Sinclair, Saigusa sp. nov. [red label] ” (INS _ DIP _ 0000614, ZIN). PARATYPES: CANADA. Yukon: Dawson, 10,11. vii. 1949, W. W. Judd (3 ♀, CNC); km 465 Dempster Hwy, 23 – 25. vi. 1980, D. M. Wood & D. Lafontaine (1 ♂, CNC); Richardson Mts, 66 ° 26.5 ′ N 135 ° 48 ′ W, 3000 ft, 8. vii. 1982, D. Wood (2 ♂, CNC). RUSSIA. Chukotka AO (Wrangel Island): SE part of Rogers Bay, 20. vii. 1932, A. Mineev (1 ♀, ZIN); SE part of Rogers Bay, 24. vii. 1932, A. Mineev (1 ♀, ZIN); environs of Somnitelnaya Bay, valley of river Somnitelnaya, Mineev Mtns, 150 m, shingle, on flower of Dryas integrifolia, 19. vii. 1966, KBG (1 ♂, ZIN); lower flow of river Khischniki, foothills of Mineev Mtns, marshy tundra, 11. vii. 1972, KBG (1 ♂, 1 ♀, CNC); middle flow of river Neozhidannay, 71 ° 01 ′ N 179 ° 09 ′ E, BT G, 8 – 9. vii. 2019, OAK (3 ♂, 1 ♀, ZIN); same locality, BT A, YPT, 8 – 9. vii. 2019, OAK (2 ♀, ZIN); same locality, BT W, YPT, 8 – 9. vii. 2019, OAK (1 ♂, ZIN); same locality, BT D, YPT, 8 – 9. vii. 2019, OAK (1 ♀, ZIN); Inkali Mtn, 71 ° 05 ′ N 179 ° 42 ′ W, damp tundra, Sw, 17. vii. 2019, OAK (1 ♀, ZIN). Chukotka AO (Kolyuchin Island): Kolyuchin Island, 25 – 27. vii. 1938, Hecker (5 ♂, 5 ♀, ZIN; 1 ♂, 1 ♀, CULSP). Chukotka mainland: Apuka, island in delta of river Apuka, on flowers of cloudberry Rubus chamaemorus, 2. vii. 1959 KBG (1 ♂, ZIN); Apuka, Pakhachinskij Range, mountain tundra, 7. vii. 1959, KBG (1 ♂, ZIN); Pevek, 200 – 300 m, moist tundra, 29. vi. 1963, KBG (1 ♂, ZIN); same locality, dryad tundra, 3. vii. 1963, KBG (2 ♀, ZIN); same locality, osier-bed in valley, 8. vii. 1963, KBG (1 ♂, ZIN); same locality, moist sedge tundra, 11. vii. 1963, KBG (1 ♂, ZIN); same locality, mossy-sedge tundra, 18. vii. 1963, KBG (7 ♂, 10 ♀, ZIN); Schmidt, on snow, 9. viii. 1966, KBG (5 ♂, 1 ♀, ZIN); same locality, on snowfield, 16. vii. 1966, KBG (6 ♂, 7 ♀, ZIN); same locality, polygon tundra, 11. vii. 1971, KBG (2 ♂, 1 ♀, ZIN); same locality, spotted tundra, 17. vii. 1971, KBG (9 ♂, 5 ♀, ZIN); same locality, S slope tundra with willow, 11. vii. 1971, KBG (1 ♀, ZIN); valley of river Ichuvisi, Komsomolskij mine, Chaunskij District, shrubby tundra, 5. vii. 1963, KBG (3 ♂, 2 ♀, ZIN); same locality, sedge tundra and osier-bed in river valley, 4. vii. 1963, KBG (10 ♂, 3 ♀, ZIN); Iultin, water-logged sedge tundra, 21. vii. 1963, KBG (11 ♂, 2 ♀, ZIN); 5 km N of Egvekinot, 27. vii. 1963, KBG (1 ♂, ZIN). USA. Alaska: Cape Thompson, 29. vii. 1960, H. E. Erdman (1 ♀, CNC); Schrader Lake, 3 – 5. vii. 1972, K. W. Philip (2 ♂, CNC). Additional material: RUSSIA. Chukotka AO (Wrangel Island): middle flow of river Neozhidannaya, 71 ° 01 ′ N 179 ° 09 ′ E, YPT, BT 4, 12 – 16. vii. 2016, LFV (1 ♂, ZIN); same locality, YPT, BT 3, 12 – 16. vii. 2016, LFV (3 ♀, ZIN).	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64749F0B57EEFCA2FDC9EBBD.taxon	diagnosis	Diagnosis. A large species (body about 7 mm) with body and abdomen densely brownish pruinescent, halteres yellow, acrostichals biserial. Male: eyes separated by very narrow frons, legs yellowish to brownish yellow, hind femur with numerous short setae over entire length ventrally, hind tibia slender; wing membrane slightly infuscate; terminalia yellowish, cerci extended only to tergite 5. Female: mid and hind femora with posteroventral pennate setae.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64749F0B57EEFCA2FDC9EBBD.taxon	description	Description. Male (Fig. 20). Body length 6.9 – 7.3 (lectotype 7.1); wing length 6.9 – 7.4 (lectotype 7.3) mm. Head with dense brownish pruinescence on face, frons, postgena, ocellar triangle and occiput; dark setose. Eyes separated by frons; upper ommatidia slightly enlarged. Frons very narrow on middle part, at most as broad as anterior ocellus, slightly broader just below ocellar tubercle, forming larger subtriangular space above antennae, with scattered marginal setulae. Face broad, bare. Ocellar triangle with several fine setae of different lengths. Occiput with numerous long, thin setae, including postoculars; postgena with numerous finer setae than occiput. Antenna blackish brown; scape and pedicel short, scape slightly longer, both with short setulae; postpedicel conical, nearly 3 X longer than wide; stylus rather long, nearly 1.5 X shorter than postpedicel. Palpus dark; with long, dark fine setae. Proboscis with labrum dark reddish-brown, 1.2 – 1.3 X head height. Thorax dark in ground-colour, densely brownish pruinescent; with black setation; scutum brownish, without vittae. Proepisternum with tuft of numerous long setae on lower section; upper proepisternum in front of spiracle with 3 – 5 fine setae. Prosternum bare. Antepronotum with numerous short setae. Postpronotal lobe with numerous almost uniform fine setae of different lengths, sometimes 1 – 2 setae somewhat stronger. Mesonotum mostly with fine setae, including numerous undifferentiated similar presut spal, npl and psut spal (some posterior npl and psut spal somewhat longer and stronger but their number and position are very variable, often even on right and left sides), 2 – 3 pal with several shorter setae (number and robustness variable); 8 – 10 of subequally long sctl and some number of additional finer setae (number and robustness very variable); acr rather long, thin, arranged in 2 irregular rows, absent on prescutellar depression; presutural dc long, arranged in 3 – 4 irregular rows, postsutural dc less numerous, longer. Laterotergite with numerous long fine setae. Anterior and posterior spiracles yellowish. Legs with coxae, trochanters and tarsi (except basal part of hind basitarsus) brownish, fore femur, mid femur ventrally, fore tibia toward apex, mid and hind tibiae at apex brownish yellow, remaining parts yellowish; mostly subshiny, coxae and trochanters with denser greyish pruinescence; black setose. Femora, tibiae and tarsomeres slender; hind femur of subequal thickness to fore and mid femora. Mid and hind femora whitish pilose ventrally (more distinctly on hind femur); fore femur with rather thin, moderately long setae anteroventrally and posteroventrally, longest setae closer to base slightly longer than half of femur basal width; mid femur with numerous strong, short setae anteroventrally and posteroventrally; hind femur entirely covered with setae ventrally (at most as long as half of femur width). All tibiae with some short ad and pd setae stronger on hind tibia; no seta in posteroapical comb of hind tibia. All tarsomeres covered with short, simple setae (except somewhat longer setae of subapical circlet); mid and hind basitarsi with short, strong setae ventrally. Wing membrane faintly infuscate, mostly with brownish veins; all veins complete (except Sc). Pterostigma brownish yellow; basal costa seta absent. Anal lobe well-developed; axillary incision acute but closer to 90 °. Squama yellow, dark fringed. Halter yellow. Abdomen dark, densely brownish pruinescent; covered with uniform black setae shorter on tergites dorsally. Segments before segment 7 unmodified. Tergite 7 broadly concave posteriorly. Segment 8 with tergite and sternite separated. Tergite 8 upturned, with posterior margin recurved anteriorly, with numerous setae along posterior margin. Sternite 8 simple, scoop-like, with numerous long setae along posterior margin. Terminalia (Figs 21 – 24). Cerci yellowish; fused to epandrium (but suture distinct), broadly concave and flattened medially, strongly prolonged anteriorly, without pair of subepandrial lobes beneath; apical part of cercus (before epandrium) short, extended to tergite 5, subrectangular viewed laterally, subtriangular, with straight inner margin viewed dorsally, mostly bare, with some dark scattered setulae dorsally; posterior part of cercus very short, rounded, not extended beyond epandrium, covered with fine setulae. Epandrium almost entirely yellowish, only posterior digitiform projection brownish yellow (very rarely epandrium dark, 1 ♂ from Wrangel Island), faintly greyish pruinescent, with moderately long dark setae on upper part and longer setae along lower margin, posterior projection covered with black spinules dorsally; rather subrectangular (lateral view), somewhat humped dorsally, with short, digitiform projection on lower part posteriorly. Hypandrium yellowish, subtriangular (ventral view), entirely sclerotized, bare; gonocoxal apodeme large, broadened apically. Phallus (Fig. 24) yellowish; short, mostly hidden (lateral view); gently arched, thick on about basal half, becoming abruptly slenderer beyond epandrial projection; with two minute lateral “ spinules ” slightly beyond thickened portion. Ejaculatory apodeme extended far beyond basal curvature of phallus, with narrow lateral wings and very broad vertical wing. Female (Fig. 25). Frons broad, parallel-sided; ocellar tubercle and occiput with somewhat shorter and stronger setae. Thorax with greyish pruinescence, scutum with 3 indistinct brownish vittae along rows of acr and dc; mesonotum with shorter setae. Legs often somewhat paler than in male (except coxae and trochanters) with only fore femur and tarsomeres darkened; mid and hind femora with complete rows of moderately long, pennate posteroventral setae. Abdomen with shorter setae; cercus long, slender, with dark setulae.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64749F0B57EEFCA2FDC9EBBD.taxon	distribution	Distribution. Holarctic; in Eurasia, the new species is known from Chukotka mainland and nearby Arctic islands (Kolyuchin, Wrangel); in North America, this species is known from Alaska and Yukon, but unknown from the Canadian Arctic islands.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64749F0B57EEFCA2FDC9EBBD.taxon	etymology	Etymology. The specific name is a patronym in honour of the J. R. Malloch, a world-renowned dipterist in the first half of the 20 th century, and among his numerous contributions were publications on Arctic Diptera (including Empididae) of North America.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64749F0B57EEFCA2FDC9EBBD.taxon	discussion	Remarks. The new species is most similar to R. hovgaardii differing from the latter primarily by brownish grey pruinescent thorax and abdomen (dark brown in R. hovgaardii), yellowish halteres, extensively yellowish legs and yellowish epandrium (brown in R. hovgaardii). The female of R. hovgaardii has pennate setae on the hind tibia. In addition, the new species could be compared with R. plumipes (Meigen) and R. vesiculosa (Fallén). However, both these species can be readily distiguished from R. mallochi sp. nov. primarily by the much longer cerci of the male terminalia extending to abdominal tergite 2 and dark halteres. Among material from North America, the species was recognised as Rhamphomyia (Dasyrhamphomyia) sp. 12 (Saigusa unpubl. data). Habitat. Judging by the material of K. B. Gorodokov from 1950 s – 1970 s, this species is most numerous in coastal areas in northern Chukotka (Pevek, Schmidt), where it was recorded in various habitats. On Wrangel Island, the largest series of specimens was collected in the colder regions of the island (Neozhidannaya River), where it was most numerous in the zonal spotted tundra. Moreover, in the warmer central region, the only specimen was found in a wetland below a snow field.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C647B9F0857EEFCDAFE4EEAB9.taxon	description	(Fig. 13)	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C647B9F0857EEFCDAFE4EEAB9.taxon	materials_examined	Material examined. RUSSIA. Chukotka AO (Wrangel Island): middle flow of Neozhidannaya River, 71 ° 01 ′ N 179 ° 09 ′ E, BT 2, YPT, 12 – 16. vii. 2016, LFV (1 ♂, ZIN); environs of Somnitelnaya Bay, 6. vii. 1972, KBG (1 ♀, ZIN); 4 km N of Somnitelnaya Bay, Somnitelnaya River, shore slope and flood-lands, 28. vii. 1972, KBG (2 ♂, 1 ♀, ZIN); 5 km N of Somnitelnaya Bay, valley of Somnitelnaya River, foothills 150 m, on flowers of Dryas, 25, 28. vii. 1972, KBG (2 ♀, ZIN); Somnitelnaya Bay, valley of Somnitelnaya River, Mineev Mtns, 150 m, pebbles, on flower of Potentilla emarginata, 22. vii. 1966, KBG (1 ♀, ZIN); same locality, south slope of Mineev Mtns, 100 m, pebbles, rubbly placer, 26. vii. 1966, KBG (1 ♀, ZIN); same locality, Sw on Epilobium latifolium L., 23. vii. 1971, KBG (1 ♀, ZIN); basin of Khischniki River, 8 km SE of Sovetskaya Mtn, lawn near stream, 250 m, 13. vii. 1972, KBG (1 ♂, ZIN); environs of Tundrovaya Mtn, near stream, 18. vii. 1972, KBG (1 ♀, ZIN); same locality, tundra, 18. vii. 1972, KBG (2 ♂, 2 ♀, ZIN); same locality, 71 ° 18 ′ N, 179 ° 48 ′ W, BT 3, PT, 1 – 19. vii. 2015, LFV (1 ♀, ZIN); same locality, BT 5, 1 – 19. vii. 2015, PT, LFV (1 ♀, ZIN); same locality, BT 1, 71 ° 18.342 ′ N 179 ° 48.367 ′ W, 30. vi. – 6. vii. 2018, U. V. Babiy (1 ♀, ZIN); middle flow of Mamontovaya River, N Perkatkun, osier-bed in river valley, 17, 21. vii. 1972, KBG (4 ♂, 2 ♀, ZIN); same locality and collector, 21. vii. 1972 (19 ♂, 11 ♀, ZIN); same locality and collector, 29. vii. 1972 (1 ♂, ZIN); same locality and collector, 18. vii. 1972 (1 ♂, 2 ♀, ZIN); same locality, BT 4, 71 ° 09 ′ N 179 ° 45 ′ W, 26. vi. – 19. vii. 2011, А. А. Rodionov (1 ♀, ZIN); same locality, 71 ° 10 ′ N 179 ° 45 ′ W, in house, 29. vi., 1. vii. 2015, OAK (2 ♀, ZIN); same locality, BT 20 A, Sw, 1 – 2. vii. 2015, OAK (2 ♂, 3 ♀, ZIN); same locality, BT 14, Sw, 2. vii. 2015, OAK (7 ♂, 19 ♀, ZIN); same locality, BT 3, Sw, 1. vii. 2015, OAK (1 ♂, 4 ♀, ZIN); same locality, BT 2, Sw, 2. vii. 2015, OAK (2 ♂, ZIN); same locality, BT 1 A, Sw, 2. vii. 2015, OAK (1 ♂, 1 ♀, ZIN); same locality and biotope, YPT, 2 – 4. vii. 2015, OAK (1 ♂, 1 ♀, ZIN); same locality, BT 7, Sw, 1. vii. 2015, OAK (1 ♂, 6 ♀, ZIN); same locality, BT 6, Sw, 2 – 3. vii. 2015, OAK (5 ♂, 5 ♀, ZIN); same locality, BT 6 A, Sw, 1. vii. 2015, OAK (1 ♀, ZIN); same locality, BT 11, Sw, 30. vi. 2015, OAK (1 ♀, ZIN); same locality, BT 5, Sw, 30. vi. 2015, OAK (2 ♀, ZIN); same locality, BT 10, Sw, 30. vi. 2015, OAK (1 ♀, ZIN); same locality, BT 17, Sw, 3. vii. 2015, OAK (9 ♀, ZIN); same locality, Arctic fox burrow, Sw, 1. vii. 2015, OAK (1 ♂, 1 ♀, ZIN); same locality and biotope, PT, 1 – 23. vii. 2015, OAK (1 ♀, ZIN); same locality and biotope, Sw, 18. vii. 2019, OAK (1 ♀, ZIN); brook Vesely, 71 ° 11 ′ N, 179 ° 43 ′ W, BT A, Sw, 18. vii. 2019, OAK (1 ♂, ZIN); spurs of Pervaya Mtn, 71 ° 09 ′ N, 179 ° 27 ′ W, BT 11, Sw, 28. vi. 2015, OAK (5 ♂, 3 ♀, ZIN); same locality and biotope, Sw, 13. vii. 2015, OAK (1 ♀, ZIN); upper flow of Neizvestnaya River, 71 ° 13 ′ N, 179 ° 19 ′ W, BT 4 A, Sw 5. vii. 2006, OAK (11 ex., SZM); same locality, BT 7 A, 5. vii. 2006, OAK (1 ex., SZM); same locality, BT 3 A, 6. vii. 2006, OAK (1 ex., SZM); same locality, BT 12, Sw, 6. vii. 2015, OAK (3 ♂, 2 ♀, ZIN); same locality, BT 5 A, PT, 26. vi. – 10. vii. 2015, OAK (1 ♀, ZIN); same locality and biotope, Sw, 12. vii. 2015, OAK (1 ♀, ZIN); same locality, BT 3 B, Sw, 7. vii. 2015, OAK (1 ♀, ZIN); same locality, BT C 1, Sw, 21. vi. 2015, OAK (3 ♀, ZIN); same locality, wormwoods near house, Sw, 24. vi. 2015, OAK (1 ♂, ZIN); middle flow of Neizvestnaya River, BT 1, YPT, 71 ° 20.287 ′ N 179 ° 29.779 ′ W, 5 – 14. vii. 2018, U. V. Babiy (1 ♀, ZIN); same locality, BT 2, YPT, 71 ° 20.416 ′ N 179 ° 29.651 ′ W, 13 – 14. vii. 2018, U. V. Babiy (2 ♀, ZIN). Recognition. Large (wing length 6 – 7 mm) brownish grey species. Male (Fig. 13) holoptic; thorax black setose, scutum densely brownish grey pruinescent, with dark vittae beneath acrostichal and dorsocentral rows; mesonotal setae mostly fine, long, presutural dc and acr multiserial; legs robust, entirely dark brown, hind femur without setae on about apical 2 / 3 ventrally, only densely pale pilose, hind tibia evenly thickened toward apex; wing brownish infuscate, halter yellowish; cerci directed anteriorly, very long, extended nearly to middle of abdominal tergite 3, epandrium dark. Female wing broadened; legs without pennate setae.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C647B9F0857EEFCDAFE4EEAB9.taxon	distribution	Distribution. A Holarctic species with huge area of distribution; in Eurasia, R. nigrita is widespread across the low and high arctic of Norway, Sweden and Russia (with some records from the boreal zone; e. g., Altay, Magadanskaya Province); in North America, from Greenland to Alaska and as far south as Churchill, Manitoba and northern British Columbia (Sinclair et al. 2019). However, among the Russia Arctic islands, R. nigrita was found only from Wrangel Island. Habitat. On Wrangel Island this species is most common in the warmer central region of the island, where it is found in various biotopes. In colder areas (Neozhidannaya River, coastal plain in Somnitelnaya Bay), a single specimen was found.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64789F0957EEFBC9FB7CE9A5.taxon	description	(Fig. 26)	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64789F0957EEFBC9FB7CE9A5.taxon	materials_examined	Material examined. RUSSIA. Chukotka AO (Wrangel Island): middle flow of Mamontovaya River, N of Perkatkun, osier-bed in river valley, 17. vii. 1972, KBG (1 ♀, ZIN); same locality, 71 ° 10 ′ N 179 ° 45 ′ W, BT 3, 29. vi. – 9. vii. 2006, OAK (1 ♂, ZIN); same locality and biotope, 9. vii. 2006, OAK (1 ♀, ZIN); same locality and biotope, 1. vii. 2015, leg. OAK (3 ♀, ZIN); same locality, BT 1 А, 2. vii. 2015, OAK (1 ♂, ZIN); same locality, BT 20 А, 2 – 4. vii. 2015, OAK (2 ♂, ZIN); same locality, 71 ° 10 ′ N 179 ° 45 ′ W, BT 7, 1. vii. 2015, OAK (1 ♂, ZIN); upper flow of river Neizvestnaya, 71 ° 13 ′ N 179 ° 19 ′ W, BT С 1, 24. vi. 2015, OAK (1 ♀, ZIN); same locality, BT 7 А, 25. vi. – 10. vii. 2015, OAK (1 ♀, ZIN); middle flow of Neizvestnaya River, 71 ° 20.287 ′ N 179 ° 29.779 ′ W, BT 1, YPT, 5 – 14. vii. 2018, U. V. Babiy (1 ♀, ZIN); same locality, 71 ° 20.416 ′ N 179 ° 29.651 ′ W, BT 2, YPT, 13 – 14. vii. 2018, U. V. Babiy (3 ♂, 2 ♀, ZIN); same locality and biotope, YPT, 4 – 5. vii. 2019, U. V. Babiy (4 ♂, 4 ♀, ZIN). Recognition. Mid-sized (wing length 4.7 – 5.3 mm) blackish species. Male (Fig. 26) holoptic; thorax black setose, scutum with pair of pruinescent vittae anteriorly, sometimes lustrous posteriorly; supra-alar region with broad shiny stripe, mesonotal setae well differentiated, presutural dc 3 – 4 - serial, acr biserial, 6 – 8 sctl (rarely 10); legs slender, entirely dark brown, no modified podomeres, hind femur with minute anteroventral setulae (except several longer setae near extreme base), all tibiae with rows of long ad and pd setae; wing faintly infuscate, axillary incision very acute, at most 45 °, halter yellow; abdomen pruinose dorsally and narrowly shiny laterally; phallus short, almost entirely hidden within epandrium and cerci, at most very short portion between lower margin of epandrium and hypandrium visible, hypandrium well exposed, subtriangular. Female with somewhat darker wing, legs without pennate setae. Besides the species noted in the key, R. shewelli should be compared primarily with R. dorsata Becker known from Arctic continental territories of Eurasia. However, in the latter species the scutum is broadly shiny laterally leaving only a very narrow greyish vitta beneath the acrostichals and the abdomen is entirely shiny in dorsal view (Sinclair et al. 2019).	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64789F0957EEFBC9FB7CE9A5.taxon	distribution	Distribution. Holarctic; in North America, this low arctic species is widespread across western and central arctic region (Sinclair et al. 2019); in Eurasia, it is known only from Wrangel Island. Habitat. On Wrangel Island this species was collected only in the warmer central part of the island. The majority of specimens were found in wet habitats with willow bushes, located in river valleys.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64799F0E57EEFEE2FBC9E8FA.taxon	description	(Fig. 27)	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64799F0E57EEFEE2FBC9E8FA.taxon	description	(catalogue); Barták, 1999: 99 (type data); Shamshev, 2001 b: 320 (key); Barták, 2006: 508 (key); 2007: 103 (catalogue); Yang et al., 2007: 157 (catalogue); Shamshev, 2016: 56 (checklist).	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64799F0E57EEFEE2FBC9E8FA.taxon	materials_examined	Material examined. RUSSIA. Murmanskaya Prov. (Kildin Island): 13. vii. 1908, L. Bianki (1 ♀, ZIN). Recognition. Small, blackish, pale-legged, delicate flies (body length about 3 mm). Male (Fig. 27) dichoptic, eyes separated by broad shiny frons, face narrower than frons; occiput shiny on upper part and faintly pruinescent on lower part; antenna black, postpedicel long; palpus usually yellowish with darkened apex; labrum about 2 X longer than head height, extensively yellowish, only apex brownish; thorax black, pale setose, 1 npl; postpronotal lobe and almost entire mesoscutum shiny (except faintly pruinescent narrow marginal edging); legs extensively yellow, only apical part of femora and tibiae and all tarsi more or less intensively darkened, fore basitarsus slightly incrassate; wing faintly brownish infuscate, axillary incision very obtuse, anal lobe almost absent, anal vein incomplete, halter yellow; abdomen shiny, brownish to black dorsally and usually yellowish to brownish yellow ventrally, pale setose; terminalia directed upwards; cerci separated from each other and from epandrium; cercus with straight upper margin, apex slightly produced beyond apex of epandrium, covered with setulae; epandrium fused with hypandrium, with somewhat produced truncate apical part, covered long pale setae, some spinule-like setulae at apex; subepandrial sclerite entire, with apex recurved as slender projection, extended beyond lower margin of epandrium; hypandrium deeply concave medially, with very narrow slightly sclerotised rim anteriorly, produced as digitiform lobes posterolaterally, lobes covered with long, pale setae; phallus well exposed, pale yellow, mostly uniformly thin, somewhat thicker on basal part, gently curved; ejaculatory apodeme far before basal curvature of phallus, very close to base of cerci, with two small lateral wings. Female similar to male, face somewhat broader.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64799F0E57EEFEE2FBC9E8FA.taxon	distribution	Distribution. Palaearctic. Fennoscandia, mountains of Europe with the exception of southernmost parts, northwest and north parts of European Russia (Barták 2007; Shamshev 2016).	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C647E9F0C57EEFE09FD15E8A2.taxon	description	(Figs 28 – 31)	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C647E9F0C57EEFE09FD15E8A2.taxon	materials_examined	Type material examined. LECTOTYPE (here designated in order to fix identity of the species) ♂ (Fig. 28), labelled (Fig. 29): “ Typus [blue label] ”; “ Novaj. Semlja. Ökland / Østre Korsä / (Achaugel bűgt.) / No 224 / 15 aűg. ”; “ Rhamphomyia / armipes Sack / nov. spec. ”; “ Gl. 1695 / ZM. Oslo ”; “ pinx [pink label] ”; “ Typus [pink label] ”; “ NHMO: type collection / 1000327600 ”; “ NHMO / Norway [data matrix code] ”; “ LECTOTYPE / Rhamphomyia / armipes Sack / des. Sinclair 2020 [red label] ” (NHMO). PARALECTOTYPE: Berek (Briska) Ösla [?], 18 Aug. No. 235 (1 ♀, NHMO). Note on types. This species was originally described on the basis of one male and two female specimens.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C647E9F0C57EEFE09FD15E8A2.taxon	diagnosis	Diagnosis. This dark, shiny-legged and pale brown setose species is distinguished from other Pararhamphomyia by the male hind femur and tibia with triangular projections and apex of subepandrial lobe pointed. Female legs without pennate setae.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C647E9F0C57EEFE09FD15E8A2.taxon	description	Redescription. Wing length 4.5 – 5.5 mm. Male (Fig. 28). Head with greyish pruinescence on face, frons, postgena and occiput. Holoptic, eye with ommatidia larger on upper half, smaller on lower half. Frons represented by very small triangular space below ocellar tubercle and larger subtriangular space above antennae, bare. Face slightly divergent towards mouthparts; bare, with narrow oral margin shiny. Ocellar triangle dark, pruinescent, cellar setae broken; 2 – 3 pairs of postocellar setae slightly shorter and finer than posterior ocellar setae. Occiput with rows of dark, long, curved postocular setae; postgena with finer, sinuous setae than occiput. Antenna dark brown; scape and pedicel of equal lengths; postpedicel broken off. Palpus dark, with long, dark setae. Clypeus with greyish pruinescence; labrum lustrous and dark reddish-brown, 1.5 X head height; labellum dark and bearing dark setulae. Thorax dark in ground-colour, largely densely grey pruinescent; setae long, fine, brownish. Scutum with 2 indistinct, dark grey vittae between acr and dc rows. Pleura clothed in grey pruinescence. Proepisternum with several long, dark setae on lower section; upper proepisternum in front of spiracle with several long, dark setae; prosternum bare. Antepronotum with row of short, stout dark setae. Postpronotum with numerous long setae. Scutum with biserial acr; dc multi-serial, subequal in length to acr, prescutellar setae longer, not differentiated from lateral setae; numerous undifferentiated presut spal; numerous undifferentiated npl; numerous prealar setae; 4 or more psut spal similar to prealars; 1 pal with several shorter setae; 4 pairs of sctl. Laterotergite with several long, dark setae. Anterior and posterior spiracles dark brown with outer black ring. Legs dark reddish brown, shiny; coxae with faint greyish pruinescence. Coxae with dark setae. All femora with distinct white ventral pile; fore femur with anteroventral row of setae, with longer posterior setae; mid and hind femora with 1 anteroventral and 1 posteroventral row of setae, stronger than on fore femur, less than half corresponding femur width. Hind femur with triangular posteroventral projection beyond mid-length. Fore tibia and tarsomere lost. Mid tibia with long, erect setae; posterior setae twice as long than anterior setae; with posteroventral row of strong setae, longer than tibial width. Hind tibia (Fig. 30) with triangular posteroventral projection on basal third and long posterodorsal setae; 1 long seta in posteroapical comb. Hind tarsomere 1 similar in size to apical width of hind tibia, dorsal setae shorter than tibial setae; with dense, dark, stout ventral setae. Wing lightly infuscate with brown veins; all veins complete (except Sc); CuA + CuP at apical fourth extended to wing margin as faint fold. Pterostigma brownish; basal costa seta absent; anal lobe well-developed. Axillary incision right angle. Halter dark brown. Abdomen dark brown with greyish pruinescence; covered with long, pale brown setae laterally, shorter on tergites dorsally. Tergite 8 similar in length to tergite 7. Terminalia (Fig. 31, undissected): Epandrium strongly tapered apically; long setae on apical half dorsally and ventrally. Cercus with broad base, tapered, shorter than subepandrial lobe [right cercus broken at base]. Subepandrial lobe cylindrical, with pointed apex; clothed in pruinescence. Hypandrium not visible. Phallus with swollen cylindrical base; lustrous and reddish brown, not forming loops, gently arched between epandrial lamellae [apex not visible]. Female. Similar to male, except basal costa seta present; postoccipital setae stronger; thoracic setae stronger, more differentiated, with more numerous sctl; mid tibia slightly arched, distinctly shorter than femur, with 1 anteroventral and 1 posteroventral row of short, strong setae; without pennate setae; CuA + CuP strong to wing margin.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C647E9F0C57EEFE09FD15E8A2.taxon	discussion	Remarks. Sack (1923) included illustrations of the male hindleg and male terminalia, the latter upside down. The basal costal seta is apparently absent in the male lectotype, but is present in the female. Certain features of the female (presence of basal costal seta; strong CuA + CuP, midleg chaetotaxy) suggest that the female paralectotype is not conspecific with the male lectotype. Further collections of both sexes of this species are required for confirmation. The cylindrical subepandrial lobe with pointed apex is similar in form to that of R. hoeli.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C647E9F0C57EEFE09FD15E8A2.taxon	distribution	Distribution. This species is known only after the type series collected from Novaya Zemlya Archipelago (Krestovy Island, Serebryanka Fjord, Berkh Island).	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C647C9F0C57EEFDE6FCDFEDB9.taxon	discussion	Remarks. Rhamphomyia caudata has a somewhat complicated taxonomic history. As a consequence, some distributional records of this species may be misleading. Rhamphomyia caudata was identified as the single species of Empididae from Spitsbergen (Boheman 1866; Holmgren 1869; Collin 1923) and was reflected in Empididae Catalogues and faunal checklists (e. g., Hackman 1968; Chvála & Wagner 1989: 299; Coulson & Refseth 2004: 104; Yang et al. 2007: 169). We have examined the specimens of R. caudata noted by Holmgren (housed in NHRS) and found that, actually, they belong to R. longestylata Frey (see below). It should be noted that Collin was somewhat confused about the synonymy of R. caudata. He believed that R. longestylata Frey was conspecific with R. caudata (Zetterstedt) (see Collin, 1961: 378). However, they are distinctly two different species (see Frey 1955 a: 472; Barták & Danielson 2008: 107). A note in Collin’s paper on flies collected from Spitsbergen (Collin 1923: 116) indicates clearly that he in fact had in front of him a male of R. longestylata rather than R. caudata. Our study does not confirm the presence of R. caudata on the Arctic islands.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C647C9F0D57EEFADEFE96E8F9.taxon	description	(Fig. 32)	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C647C9F0D57EEFADEFE96E8F9.taxon	materials_examined	Material examined. RUSSIA. Chukotka AO (Wrangel Island): Somnitelnaya Bay, on flower of Dryas integrifolia, spotted tundra, 19. vii. 1966, KBG (1 ♀, ZIN); Somnitelnaya Bay, Mineev Mtns, S Sovetskaya Mtn, 400 m, 20. vii. 1966, KBG (1 ♀, ZIN); 9 km SE Sovetskaya Mtn, stream shore, 250 m, 12. vii. 1972, KBG (1 ♂, ZIN); environs of Tundrovaya Mtn, 71 ° 18.469 ′ N 179 ° 44.327 ′ W, BT 3, 5 – 15. vii. 2018, U. V. Babiy (1 ♂, ZIN); Perkatkun, about 30 km N Somnitelnaya Bay, 350 m, on flower of Dryas integrifolia, spotted tundra, 21. vii. 1966, KBG (6 ♂, 15 ♀, ZIN); middle flow of Mamontovaya River, 71 ° 10 ′ N 179 ° 45 ′ W, BT 6, 9. vii. 2006, OAK (1 ♂, 1 ♀, ZIN); same locality, BT 11, 8. vii. 2006, OAK (1 ♀, ZIN); same locality, BT 13, 30. vi. 2015, OAK (1 ♀, ZIN); spurs of Pervaya Mtn, 71 ° 09 ′ N 179 ° 45 ′ W, BT 12, 28. vi. 2015, OAK (1 ♀, ZIN). Recognition. Mid-sized (wing length 4.8 – 5.4 mm) blackish brown species. Male (Fig. 32) holoptic; anepisternum densely pruinescent, scutum densely brownish pruinescent, presutural dc multiserial, long, acr similar but 2 – 3 - serial; legs robust, dark brown, no modified podomeres, hind femur with short spine-like setae ventrally, all tibiae with several long bristle-like ad and pd setae, mid basitarsus with 1 long dorsal seta on about middle; wing faintly infuscate, CuA + CuP incomplete, halter brown; abdomen densely brownish pruinescent, black setose; epandrium elongate, narrow, constricted in middle, projecting far beyond cercus; cercus V-shaped, strongly constricted in middle; phallus well-exposed, filamentous, very long, without loops, gently curved beyond apex of epandrium. Female: anal lobe of wing darker than wing tip, legs without pennate setae.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C647C9F0D57EEFADEFE96E8F9.taxon	distribution	Distribution. Holarctic; in North America, R. filicauda occurs across the low and high arctic from Greenland to Yukon (Sinclair et al. 2019); in Eurasia, it is known only from Wrangel Island. Habitat. A large series of this species was collected only once (in 1966) in the warmer central region of the island, in dry spotted tundra. In the 2000 s, all collections of this species were also limited by dry biotopes of this part of the island.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C647D9F0D57EEFE1EFCFDEC03.taxon	description	(Fig. 33)	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C647D9F0D57EEFE1EFCFDEC03.taxon	materials_examined	Material examined. RUSSIA. Chukotka AO (Wrangel Island): middle flow of Neozhidannaya River, 71 ° 01 ′ N, 179 ° 09 ′ E, BT 1, YPT, 12 – 16. vii. 2016, LFV (12 ♂, 18 ♀, ZIN); same locality, BT 2, YPT, 12 – 16. vii. 2016, LFV (1 ♂, 1 ♀, ZIN); same locality, BT 3, YPT, 12 – 16. vii. 2016, LFV (1 ♂, 2 ♀, ZIN); same locality, BT 4, YPT, 12 – 16. vii. 2016, LFV (2 ♂, ZIN); same locality, BT A, YPT, 8 – 9. vii. 2019, OAK (1 ♂, ZIN); same locality, BT В, Sw, 8. vii. 2019, OAK (3 ♂, 1 ♀, ZIN); same locality and biotope, YPT, 8 – 9. vii. 2019, OAK (2 ♀, ZIN); basin of Khischniki River, 7 km SE Sovetskaya Mtn, 15. vii. 1972, KBG (1 ♂, ZIN). USA. Alaska: Inaru River, 28. vii. 1956; ex. dandelion flowers (1 ♀, CNC). Recognition. Mid-sized (wing length 4.2 – 5 mm), dark-legged, greyish species. Male (Fig. 33) holoptic; scutum densely greyish pruinescent, with two indistinct darker vittae between rows of acr and dc (dorsal view); thorax black setose, upper proepisternum in front of spiracle bare; main mesonotal setae well differentiated, strong, notopleuron with cluster of several long blunt-tipped setae, acr biserial, presutural dc 1 – 2 - serial, 6 sctl; spiracles brown; fore coxa with strong blunt-tipped setae anteriorly; hind tibia evenly thickened toward apex, hind basitarsus slightly thickened, both with similar long, dense fine setae dorsally; wing faintly infuscate, CuA + CuP incomplete, basal costal seta absent, halter with yellowish knob; abdomen faintly greyish pruinescent, with yellowish setae; terminalia with subepandrial lobe, phallus mostly exposed, thickened at base, without loops, evenly curved. Female similar to male, except eyes dichoptic, wing darker infuscate, legs and abdomen with much shorter setation; legs without pennate setae, hind tibia and basitarsus slender.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C647D9F0D57EEFE1EFCFDEC03.taxon	distribution	Distribution. Holarctic; in North America this species is known from a few localities along the Beaufort Sea and the Pribilof Islands in the Bering Sea (Sinclair et al. 2019); in Eurasia, it is recorded only from Wrangel Island. Habitat. Almost all specimens of this species were collected in the 2000 s in the colder region of Wrangel Island (Neozhidannaya River), in biotopes with different moisture.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C647D9F3357EEFA01FEE7E885.taxon	description	(Fig. 34)	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C647D9F3357EEFA01FEE7E885.taxon	materials_examined	Material examined. Russia. Chukotka AO (Wrangel Island): Somnitelnaya Bay, on flower of Dryas integrifolia, spotted tundra, 19. vii. 1966, KBG (2 ♂, 2 ♀, ZIN); environs of pass Vjuchny, NW Somnitelnaya Bay, 21. vii. 1972, KBG (1 ♂, ZIN); 5 km N Somnitelnaya Bay, valley of Somnitelnaya River, small meadow, 25. vii. 1972, KBG (2 ♀, ZIN); same locality, 7 – 8. vii. 1972, KBG (2 ♀, ZIN); Somnitelnaya Bay, valley of Somnitelnaya River, Mineev Mtns, 150 m, pebbles, on flower of Potentilla emarginata, 22. vii. 1966, KBG (7 ♂, 3 ♀, ZIN); near north slope of Berri Peak, 6 km ESE Sovetskaya Mtn, 15. vii. 1972, KBG (5 ♂, 2 ♀, ZIN); upper flow of Khischniki River, 7 km SE mountain Sovetskaya, Dryas tundra near river, 200 m, 11. vii. 1972, KBG (7 ♂, 2 ♀, ZIN); middle flow of Mamontovaya River, N Perkatkun, osier-bed in river valley, 17,22. vii. 1972, KBG (8 ♂, 1 ♀, ZIN); same locality, 71 ° 10 ′ N 179 ° 45 ′ W, BT 20 A, YPT, 2 – 4. vii. 2015, OAK (1 ♂, ZIN); spurs of Pervaya Mtn, 71 ° 09 ′ N, 179 ° 27 ′ W, BT 11, Sw, 28. vi. 2015, OAK (2 ♂, ZIN); same locality, BT 14, Sw, 28. vi. 2015, OAK (1 ♂, ZIN); upper flow of Neizvestnaya River, 71 ° 13 ′ N, 179 ° 19 ′ W, BT 5 А, 26. vi. 2006, OAK (1 ♂, ZIN); same locality and biotope, 26. vi. 2015, OAK (1 ♂, ZIN). Recognition. Rather small species (wing length 3.5 – 4.2 mm) with dark legs and densely pruinescent thorax and abdomen. Male (Fig. 34) holoptic; antenna dark brown; scutum brownish grey with two indistinct darker vittae between rows of acr and dc (dorsal view); thorax dark setose, upper proepisternum in front of spiracle with 0 – 2 short setae; mesonotal setae well differentiated, bristle-like, acrostichals and dorsocentrals long, 1 – 2 - serial, 2 – 3 npl, 4 sctl; spiracles dark brown; legs dark setose, fore coxa with fine setae anteriorly, tibiae and tarsomeres slender; mid tibia with 2 – 3 anterodorsal setae (except 1 long seta of subapical circlet), about 2 X as long as tibial width; hind tibia with moderately long strong setae dorsally, 1 seta in posteroapical comb; hind femur with complete row of anteroventral setae (longest subapical setae nearly as long as femur width), hind basitarsus only with short setae dorsally; wing whitish, CuA + CuP evanescent apically, not quite reaching wing-margin or extended as faint fold, basal costa seta present, axillary incision right-angled, halter pale brown to yellow; abdomen greyish, paler than thoracic pleuron, dark setose; terminalia small; cercus shorter than epandrium, with broad base, tapered at midlength to narrow apical portion, covered with short setulae; subepandrial lobe present, cylindrical, with pointed apex, projecting slightly farther than cercus; epandrium subtriangular, tapered to truncate, slightly bilobed setose apex, curved medially, outer apex with cluster of long, dark stout setae; hypandrium short, boot-shaped, ending in middle of swollen phallus base; phallus short, gently arched between epandrial lamellae, not forming loops, base swollen, apex with shallow S-shaped curvature. Female similar to male, except eyes dichoptic; wing slightly infuscate, CuA + CuP entirely sclerotized, brownish; leg chaetotaxy reduced, without pennate setae.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C647D9F3357EEFA01FEE7E885.taxon	distribution	Distribution. Holarctic; in North America, R. hoeli is known from the low and high arctic region east of Hudson Bay and Greenland (Sinclair et al. 2019); in Eurasia, it is recorded only from Wrangel Island. Habitat. On Wrangel Island this species was collected mainly in valleys of rivers and streams as well as some wet biotopes.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64439F3357EEFD82FCF9EEED.taxon	description	(Fig. 35)	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64439F3357EEFD82FCF9EEED.taxon	materials_examined	Material examined. RUSSIA. Archangelskaya Prov. (Novaya Zemlya Archipelago): steamer deck, in sea near western shore of Novaya Zemlya, 20. vii. 1948, Korotkevich (1 ♂, 1 ♀, ZIN). Chukotka AO (Wrangel Island): middle flow of Mamontovaya River, 71 ° 10 ′ N 179 ° 45 ′ W, BT 20 A, Sw, 2. vii. 2015, OAK (1 ♂, ZIN); same locality and biotope, YPT, 2 – 4. vii. 2015, OAK (3 ♂, ZIN); same locality, BT 3, Sw, 1. vii. 2015, OAK (3 ♂, ZIN); same locality, BT 2, YPT, 2 – 4. vii. 2015, OAK (1 ♂, ZIN); same locality, BT 1 A, Sw, 2. vii. 2015, OAK (1 ♂, ZIN); same locality and biotope, YPT, 2 – 4. vii. 2015, OAK (1 ♂, 2 ♀, ZIN); same locality, BT 7, Sw, 1. vii. 2015, OAK (1 ♂, ZIN); upper flow of Neizvestnaya River, 71 ° 13 ′ N, 179 ° 19 ′ W, BT 12, Sw, 6. vii. 2015, OAK (7 ♂, ZIN); same locality, BT 13, YPT, 6 – 15. vii. 2015, OAK (1 ♂, ZIN); same locality, BT 7 A, PT, 25. vi. – 10. vii. 2015, OAK (1 ♂, ZIN); same locality, BT C 1, Sw, 24. vi. 2015, OAK (1 ♂, ZIN). Chukotka AO (Kolyuchin Island): 25 – 27. vii. 1938, Hecker (4 ♂, 8 ♀, ZIN). Recognition. Mid-sized (wing length 4.2 – 4.5 mm) species. Male (Fig. 35) holoptic; scutum densely brownish grey pruinescent, dc uniserial, acr biserial, notopleuron with 4 – 6 long strong blunt-tipped setae, 6 sctl, laterotergite with pale setae; legs dark brown, shiny, fore coxa with strong blunt-tipped setae anteriorly, hind tibia clavate and clothed in dense long hair-like setae, hind tarsomeres 1 – 2 thickened, with similar setae dorsally; wing faintly infuscate, CuA + CuP incomplete, halter yellowish (sometimes dirty yellow); abdomen faintly greyish pruinescent, covered with pale setae; terminalia: cercus short, one-third length of epandrium, tapered apically; subepandrial lobe present, longer than cercus, clavate; epandrium gradually tapered, bearing numerous long, pale setae apically; phallus well exposed, slender, gently sinuate on about middle, looping beyond epandrium and curving back to cercus. Female wing faintly brownish infuscate, hind tibia and tarsus slender, legs with simple short setae.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64439F3357EEFD82FCF9EEED.taxon	distribution	Distribution. Holarctic; in Eurasia, R. kjellmanii extends across arctic and subarctic Russia, including both continental areas (Taymyr, Yakutia, Chukotka) and islands (Novaya Zemlya, Wrangel, Kolyuchin). In North America, this species is known only from the extreme northwestern region (Sinclair et al. 2019). Habitat. On Wrangel Island this species was collected mainly in wet and, less commonly, in moderately moistened habitats in the warmest central part of the island.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64409F3657EEFB3EFD8AE989.taxon	description	(Figs 36, 37)	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64409F3657EEFB3EFD8AE989.taxon	materials_examined	Material examined. NORWAY. Spitsbergen (Svalbard Archipelago): Advent Bay, Holmgren (5 ♂, 6 ♀, NHRS); Mimerdalen, Pyramiden, tundra, 29. viii. 1979, KBG; on flowers of Dryas octopetala (1 ♂, 1 ♀, ZIN).	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64409F3657EEFB3EFD8AE989.taxon	diagnosis	Diagnosis. Mid-sized (wing length about 4 mm), blackish, shiny flies. Male (Fig. 36) holoptic; thorax and abdomen black setose, scutum and anepisternum shiny; legs dark brown, hind basitarsus slightly uniformly thickened, all tibiae and tarsi clothed in long numerous fine setae but hind tibia with long setae on about apical half only; abdomen shiny, epandrium and cerci elongate, cercus finger-like beyond dorsal cavity; phallus well exposed, filamentous, long, gently curved. Female with brownish infuscate wing, short setose slender legs without pennate setae.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64409F3657EEFB3EFD8AE989.taxon	description	Redescription. Wing length 3.7 – 4.1 mm. Male (Fig. 36). Head with faint greyish pruinescence on face, frons, postgena, ocellar triangle and occiput; dark setose. Holoptic; upper ommatidia enlarged. Frons represented by very small subtriangular space below ocellar tubercle and larger subtriangular space above antennae, bare. Face broad, bare. Ocellar triangle with 2 long fine and some short setae. Occiput with numerous long fine setae, including postoculars; postgena with numerous hair-like setae. Antenna blackish; scape short, slightly longer than pedicel, both with short setulae; postpedicel conical, with slightly concave ventral margin on apical part, nearly 2.5 X longer than wide; stylus short, slightly shorter than postpedicel basal width. Palpus dark; with several long, dark fine setae. Proboscis with labrum dark reddish-brown, nearly 1.3 X as long as head height. Thorax dark in ground-colour, with black setation; prothoracic sclerites mostly densely greyish pruinescent, antepronotum and postpronotal lobe faintly pruinescent, subshiny; mesonotum almost entirely shiny, only notopleuron along lower margin, postalar tubercle, scutum in front of scutellum and scutellum very faintly pruinescent; mesopleuron mostly greyish pruinescent, anepisternum shiny, also usually katepisternum on upper part and anepimeron anteriorly shiny. Proepisternum with few moderately long fine setae on lower section; upper proepisternum in front of spiracle without setae. Prosternum bare. Antepronotum with several short setae. Postpronotal lobe with 1 – 2 long, stronger and several shorter fine setae. Mesonotal setation: presutural supra-alar space with several long fine setae, 3 – 4 npl (with numerous finer setae anteriorly), 1 – 2 psut spal (with additional setulae anteriorly), 2 pal (often with additional setulae), 4 – 6 sctl (usually 6, variable in length and robustness, sometimes asymmetrically set); acr short, fine, arranged in 2 close irregular rows, lacking on prescutellar depression; presutural dc longer than acr, fine, arranged in 2 irregular rows, postsutural dc 1 – 2 - serial, 3 – 4 prescutellars longest. Laterotergite with several long fine setae. Anterior and posterior spiracles brown. Legs slender, uniformly brownish; mostly shiny, coxae and trochanters faintly greyish pruinescence; black setose. Fore coxa covered with long fine setae anteriorly. All tibia slightly evenly thickened toward apex; hind basitarsus slightly uniformly thickened, nearly as broad as hind tibia width at apex. All femora whitish pilose ventrally, with very short setation; fore and mid femora with anteroventral and posteroventral rows of short fine setae; hind femur with row of short anteroventral setae. Fore and mid tibiae with numerous long fine setae dorsally and posteriorly (about 2 X tibial width); mid tibia with numerous stronger long anteroventral and posteroventral setae on about apical 2 / 3; hind tibia with long fine setae dorsally on about apical half, 1 long seta in posteroapical comb. Tarsomeres 1 – 2 of all legs with numerous long setae dorsally (somewhat longer on basitarsi). Wing membrane faintly infuscate, with brownish veins; CuA + CuP (anal vein) incomplete, evanescent apically. Pterostigma brownish yellow. Basal costa seta absent. Anal lobe well-developed; axillary incision right-angled or slightly obtuse. Squama brown, dark fringed. Halter brown. Abdomen dark brown, shiny; mostly with black fine moderately long setae. Segments 6 – 7 unmodified. Segment 8 with almost fused tergite and sternite; tergite 8 simple, entire, about half length of sternite 7, with straight posterior margin, with numerous long setae along posterior margin; tergite 8 simple, enlarged, subrectangular in lateral view, V-shaped viewed posteriorly, covered with numerous long setae posteriorly. Terminalia (Fig. 37) dark brown, directed oblique upstairs. Cerci separated from each other and from epandrium, not extended beyond tergite 8 anteriorly; cercus long, narrow, broadly concave on upper margin, apical portion finger like, parallel-sided (lateral view), at least as broad as middle portion, with small internal subtriangular projection slightly beyond middle (dorsal view); covered with short fine setae; without pair of subepandrial lobes beneath. Epandrium subrectangular, elongate, about 4 X as long as broad, broadly concave on upper margin, with straight lower margin, somewhat broadened and angularly expanded apically, extended slightly beyond cerci posteriorly; with cluster of numerous long dark setae near upper anterior corner, covered with dense fine setulae on broadened apical part, some numerous moderately long setae along lower margin. Hypandrium very narrow, rim-like, entire, well sclerotized; bare. Phallus well exposed, long; mostly hair-like, knob-like thickened just beyond hypandrium; with right-angle curvature shortly beyond basal thickening, otherwise gently arcuate. Ejaculatory apodeme not extended beyond basal curvature of phallus, with broad lateral wings and narrower vertical wing. Female. Similar to male, except dichoptic, frons broad, shiny, with marginal setulae; wing brownish uniformly infuscate; leg with tibiae and tarsi slender, with short setation, without pennate setae; abdomen covered with short setae; cercus long slender, with dark setulae.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64409F3657EEFB3EFD8AE989.taxon	distribution	Distribution. Palaearctic; R. longestylata is recorded only from northern Scandinavia (Finland, Norway, Sweden), nearby regions of Russia (Murmanskaya Province, Nenets) and Spitsbergen Island (Norway).	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64409F3657EEFB3EFD8AE989.taxon	discussion	Remarks. Rhamphomyia longestylata is assigned to the R. caudata group (Sinclair et al. 2019) or to the R. longestylata group (Barták & Kubík 2009). It is very similar to R. ursinella and the main differences between these species are provided in the key. We have examined the syntypes of R. longestylata (housed in MZH and NHRS), but the lectotype has not been designated.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64469F3757EEF991FE5DEA3B.taxon	description	(Fig. 38)	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64469F3757EEF991FE5DEA3B.taxon	description	Recognition. This species remains known by the female sex only. Small (wing length of lectotype 3.2 mm) blackish flies (Fig. 38); thorax black setose, upper proepisternum in front of spiracle with 3 short setae; scutum brownish grey pruinescent; main mesonotal setae fine, hardly distinguishable, 4 npl, 6 sctl (4 stronger), presutural dc rather long, 2 – 3 - serial, acr biserial; legs brownish, with simple setae, fore coxa with fine setae anteriorly; wing uniformly brownish infuscate, no distinct basal costal seta, CuA + CuP incomplete, axillary incision right-angled, halter brownish; abdomen dark brown, faintly greyish pruinescent, subshiny (lateral view), with brownish setae (extracted from Shamshev & Sinclair 2018: 313).	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64469F3757EEF991FE5DEA3B.taxon	distribution	Distribution. This species remains known only from Novaya Zemlya. In addition to the type material, Sack (1923: 7) noted a female of R. nordqvistii among his specimens of Empididae collected from Krestovy Island (Novaya Zemlya). However, this record needs verification because the original description of R. nordqvistii is noninformative and Sack never examined the types of this species. Also, Frey (1915: 8) identified as R. nordqvistii several females from the delta of Lena River (Yakutia). At least one specimen of this series is present in ZIN and it belongs to a different species.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64469F3757EEF991FE5DEA3B.taxon	discussion	Remarks. Shamshev & Sinclair (2018) have already noted that the synonymy of R. eunordquisti with R. nordqvistii is doubtful.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64479F3757EEFC58FDB6EE03.taxon	description	(Figs 39, 40)	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64479F3757EEFC58FDB6EE03.taxon	materials_examined	Material examined. RUSSIA. Chukotka AO (Wrangel Island): middle flow of Mamontovaya River, N Perkatkun, osier-bed in river valley, 17, 21. vii. 1972, KBG (17 ♂, 13 ♀, ZIN); same locality, 71 ° 10 ′ N 179 ° 45 ′ W, BT 14, 9. vii. 2006, OAK (4 ex., SZM); same locality and biotope, Sw, 2. vii. 2015, OAK (2 ♂, 5 ♀, ZIN); same locality, BT 3, 9. vii. 2006, OAK (1 ex., SZM); same locality and biotope, Sw, 1. vii. 2015, OAK (1 ♂, ZIN); same locality, BT 20 A, Sw, 2. vii. 2015, OAK (2 ♂, ZIN); spurs of Pervaya Mtn, 71 ° 09 ′ N, 179 ° 27 ′ W, BT 11, Sw, 28. vi. 2015, OAK (6 ♂, ZIN); upper flow of Neizvestnaya River, 71 ° 13 ′ N, 179 ° 19 ′ W, BT 4 A, 5. vii. 2006, OAK (1 ex., SZM); same locality and biotope, Sw, 22. vi. 2015, OAK (3 ♂, ZIN); same locality and biotope, 26. vi. 2015, OAK (1 ♂, ZIN). Recognition. Mid-sized (wing length 5.4 – 6.1) blackish slender flies. Male (Fig. 39) holoptic; scutum faintly greyish pruinescent, with 4 narrow shiny vittae; presutural dc pale 1 – 2 - serial, hair-like, moderately long, acr biserial, similar to but somewhat shorter than dc, laterotergite with pale setae; legs long, very slender, shiny, with strongly reduced setation; hind tibia gently curved inwards beyond middle, hind basitarsus long spindle-shaped, with dense long hair-like setae dorsally; wing faintly infuscate, CuA + CuP almost complete, well sclerotized, disappearing or evanescent just before wing margin, halter yellow; abdomen with shiny dorsum, covered with pale setae; terminalia longer than thorax; epandrium elongate, almost strap-like, somewhat broadly constricted on about middle, with broadly rounded apex; cercus long, but shorter than epandrium, narrow, constricted on about middle; phallus very long, hair-like, without loops, extended beyond epandrium. Female (Fig. 40) wing broad, brownish; hind tibia and tarsus simple; legs without pennate setae.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64479F3757EEFC58FDB6EE03.taxon	distribution	Distribution. Holarctic; R. septentrionalis occurs in the low arctic west of Hudson Bay of North America and in northern regions of Russia (Sinclair et al. 2019). Habitat. On Wrangel Island this species was collected in the central part of the island, where it was found mainly in river valleys with willow bushes.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64449F3A57EEFC00FF29E9A5.taxon	description	(Figs 41, 42) urn: lsid: zoobank. org: act: 27 AE 8916 - 35 F 9 - 42 A 2 - BD 16 - 692 DADF 550 F 9	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64449F3A57EEFC00FF29E9A5.taxon	materials_examined	Type material. HOLOTYPE ♂, labelled: [RUSSIA, Chukotka AO (Wrangel Island)] “ upper flow of river Neizvestnaya, 71 ° 13 ′ N 179 ° 19 ′ W, BT 2, 3. vii. 2006, O. A. Khruleva ”; “ Holotypus / Rhamphomyia / subfilicauda / Shamshev, Sinclair sp. n. [red label] ” (INS _ DIP _ 0000621, ZIN; terminalia dissected, in microvial pinned with specimen).	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64449F3A57EEFC00FF29E9A5.taxon	diagnosis	Diagnosis. Mid-sized (wing length 4.3 mm) blackish flies of the R. pusilla group. Scutum uniformly velvety brown, without vittae; postpronotal lobe and mesonotum with numerous uniformly very long fine setae (including acr and dc), main mesonotal setae undifferentiated; hind tibia curved inwards closer to base; halter brownish; wing whitish, CuA + CuP complete; phallus well exposed, very long, mostly hair-like.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64449F3A57EEFC00FF29E9A5.taxon	description	Description. Male (Fig. 41). Wing 4.3 mm, body 4 mm. Head with brownish pruinescence on face, frons, postgena, ocellar triangle and occiput; dark setose. Eyes holoptic, touching on frons. Ocellar triangle with several long fine setae. Occiput with numerous uniformly long thin setae, including postoculars. Antenna blackish brown; scape and pedicel short, scape slightly longer, both with short setulae; postpedicel conical, 3 X longer than wide; stylus very short, slightly shorter than postpedicel basal width. Palpus dark; with numerous long, dark fine setae. Labrum 1.3 X head height. Thorax densely brownish pruinescent; with black setation; scutum uniformly velvety brown pruinescent, without vittae. Proepisternum with tuft of several long fine setae on lower section; upper proepisternum in front of spiracle without setae. Prosternum bare. Antepronotum with numerous short setae. Postpronotal lobe and mesonotum with numerous uniformly very long fine setae (including acr and dc), main mesonotal setae undifferentiated; 6 sctl; acr arranged in 2 irregular rows, absent on prescutellar depression; presutural dc separated by bare space from supra-alars, arranged in 3 – 4 irregular rows, postsutural dc 1 – 2 - serial. Laterotergite with numerous long fine setae. Anterior and posterior spiracles brown. Legs rather robust; mid legs and hind tarsi missing, remaining parts uniformly dark brown, black setose. Fore basitarsus slightly expanded; hind tibia straight, evenly expanded towards apex. Fore femur with rows of moderately long, fine anteroventral (becoming longer toward base of femur) and mostly minute (3 – 4 moderately long setae near extreme base) posteroventral setae, some moderately long setae posteriorly; hind femur with moderately long to long (on about apical half of femur) setae anteroventrally, some moderately long setae anteriorly. Fore tibia with numerous long setae dorsally and posterodorsally (longest setae nearly 1.5 X longer than tibia width); fore basitarsus with similar setae; hind tibia with numerous very long setae dorsally and similar setae anteroventrally (except extreme apex and base), 1 seta in posteroapical comb. Wing membrane whitish; CuA + CuP (anal vein) incomplete; 1 basal costal seta present; axillary incision almost 90 °. Squama brownish, dark fringed. Halter brownish. Abdomen densely brownish grey pruinescent, covered with long black setae (shorter on tergites dorsally). Segments 6 – 7 with unmodified structure; tergite 6 covered with only scattered minute setulae; tergite 7 mostly with minute setulae, bearing several fine short setae posteromarginally and laterally. Segment 8 with tergite and sternite separated; tergite 8 simple, entire, subtriangular viewed laterally, very narrow mid-dorsally, broadly concave anteriorly, with almost straight posterior margin, numerous very long setae along posterior margin; sternite 8 simple, enlarged, subrectangular in lateral view, V-shaped viewed posteriorly, covered with numerous very long setae. Terminalia (Fig. 42) dark brown, black setose. Cerci separated from each other and from epandrium, not extended beyond tergite 8 anteriorly; cercus rather short, deeply cleft mid-dorsally and divided almost entirely into two parts: basal part subrectangular (lateral view), apical part rather elongate oval, both covered with short fine setae; without pair of subepandrial lobes beneath. Epandrium subtriangular, with strongly produced narrow apical part, slightly concave on upper margin, with more or less straight lower margin, extended far beyond cerci posteriorly; with row of 6 long fine closely set setae near upper anterior corner, cluster of short, very closely set spine-like setae on upper part slightly beyond middle, numerous long setae over lower margin becoming denser and stronger slightly beyond cluster of spine-like setae but short and scattered on remaining part of epandrium. Subepandrial sclerite produced slightly beyond lower margin of epandrium. Hypandrium very narrow, rim-like, entire, well sclerotized; bare; gonocoxal apodeme directed anteriorly. Phallus well exposed, very long; mostly hair-like, knob-like thickened just beyond hypandrium; slightly sinuate and with broad even curvature shortly beyond basal expansion, gently sinuate on remaining part. Ejaculatory apodeme not extended beyond basal curvature of phallus, with broad lateral wings and somewhat narrower vertical wing. Female. Unknown.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64449F3A57EEFC00FF29E9A5.taxon	distribution	Distribution. Eurasia; only Wrangel Island.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64449F3A57EEFC00FF29E9A5.taxon	etymology	Etymology. The specific name refers to the similarity of the new species to R. filicauda Henriksen & Lundbeck.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64449F3A57EEFC00FF29E9A5.taxon	discussion	Remarks. The new species belongs to the R. pusilla group sensu Barták & Kubík (2009) (see also Sinclair et al. 2019: 65). It is similar to R. filicauda Henriksen & Lundbeck, the main differences between these species have been provided in the key. In addition, the new species could be compared with R. aversa Frey known only from Tajikistan. However, R. aversa differs from the new species primarily by faintly infuscate wings (vs. whitish). Habitat. The single specimen was collected in the moss-herb-dryad tundra in the warmer region of Wrangel Island.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C644A9F3857EEFEE2FC49EBBD.taxon	description	(Figs 43, 44) urn: lsid: zoobank. org: act: 6 D 5 C 99 D 5 - DC 42 - 4972 - 857 E-D 9 A 31 B 87234 A	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C644A9F3857EEFEE2FC49EBBD.taxon	materials_examined	Type material. HOLOTYPE ♂, labelled: [RUSSIA: Chukotka AO (Wrangel Island)] “ Somnitelnaya Bay, / south o. [= ostrov, island] Wrangel / Gorodkov 20. vii. [1] 966 ”; “ 400 m / Mineev Mountains, S g. [= gora, mountain] Sovetskaya ”; “ Holotypus / Rhamphomyia / submacrura / Shamshev, Sinclair sp. n. [red label] ” (INS _ DIP _ 0000615, ZIN; terminalia dissected, in microvial pinned with specimen).	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C644A9F3857EEFEE2FC49EBBD.taxon	diagnosis	Diagnosis. Rather large brownish grey species of the R. pusilla group with long, slender legs, curiously long epandrium and, especially, phallus; presutural dc setae multiserial, halter brownish; hind tibia and basitarsus only slightly thickened; phallus extremely long, hair-like.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C644A9F3857EEFEE2FC49EBBD.taxon	description	Description. Male (Fig. 43). Wing length 6.7 mm, body length about 6.5 mm. Head with brownish grey pruinescence on face, frons, postgena, ocellar triangle and occiput; dark setose. Eyes holoptic, touching on frons. Ocellar triangle with several long fine setae. Occiput with numerous uniformly long thin setae, including postoculars. Antenna blackish brown; scape and pedicel short, scape slightly longer, both with short setulae; postpedicel conical, 3 X longer than wide; stylus short, slightly shorter than postpedicel basal width. Palpus dark; with numerous long, dark fine setae. Labrum nearly of head height. Thorax densely brownish grey pruinescent; with black setation; scutum uniformly brownish pruinescent, without vittae. Proepisternum with tuft of several long fine setae on lower section; upper proepisternum in front of spiracle without setae. Prosternum bare. Antepronotum with numerous fine setae shorter dorsally and longer laterally. Postpronotal seta hardly distinguishable; postpronotal lobe with numerous setae of different lengths and robustness (1 – 2 long setae slightly stronger). Mesonotal setation: presutural supra-alar space covered with fine mostly short setae (1 – 2 setae longer), 4 npl (with numerous shorter and finer setae anteriorly), 1 sut spal (with several shorter setae anteriorly), 2 pal, 14 sctl; acr short fine, arranged in 2 – 3 irregular rows, lacking on prescutellar depression; presutural dc longer than acr, multiserial, not separated by bare space from supra-alars, postsutural dc arranged in 2 – 3 irregular rows, several prescutellars longest. Laterotergite with numerous long fine setae. Anterior and posterior spiracles brown. Legs long, slender, uniformly brownish, subshiny, black setose. Hind tibia somewhat evenly thickened toward apex; hind basitarsus slightly thickened but somewhat narrower that hind tibia at apex, cylindrical, 1.7 X shorter than hind tibia. Fore coxa with numerous long simple setae anteriorly. Femora whitish pilose ventrally. Fore femur with rows of fine moderately long anteroventral and posteroventral setae. Mid femur with numerous short strong setae ventrally. Hind femur with row of anteroventral setae, row with short, fine setae on basal half but longer and stronger setae on apical half; spinule-like ventral setae on basal part. Fore tibia with numerous long, quite strong setae posterodorsally and posteriorly (longest setae nearly 1.5 X tibia width). Mid tibia with several moderately long anteroventral and posteroventral setae, some strong spine-like setae toward apex ventrally. Hind tibia with numerous moderately long, rather strong setae over entire length; 1 seta in posteroapical comb. Fore basitarsus with some fine moderately long setae posterodorsally, remaining fore tarsomeres, mid and hind tarsi covered with simple short setae. Wing membrane faintly infuscate; CuA + CuP (anal vein) complete, well sclerotised; basal costal seta absent; axillary incision acute. Squama brownish, dark fringed. Halter brownish. Abdomen dark brown, brownish grey pruinescent, tergites somewhat lustrous; covered with long, dense, fine black setae. Segments 6 – 7 with unmodified structure; tergite 6 only with scattered minute setulae dorsally and several short fine posteromarginal setae; tergite 7 almost bare, with some fine setae posteromarginally (longer middorsally) and laterally. Segment 8 with tergite and sternite separated; tergite 8 simple, entire, subtriangular viewed laterally, very narrow mid-dorsally, broadly concave anteriorly, with almost straight posterior margin, numerous very long setae along posterior margin; sternite 8 simple, enlarged, subrectangular in lateral view, V-shaped viewed posteriorly, covered with numerous very long setae. Terminalia (Fig. 44) dark brown, black setose. Cerci separated from each other and from epandrium, not extended beyond tergite 8 anteriorly, without pair of subepandrial lobes beneath; cercus rather short, deeply cleft middorsally and divided almost entirely into two parts; basal part subquadrate (lateral view); apical part subrectangular, elongate, narrower on basal section, somewhat broadened apically, apex slightly concave; both parts covered with short fine setae. Epandrium subtriangular, with strongly produced narrow apical part, nearly 4 X as long as basal portion width, extended far beyond cerci posteriorly, slightly concave on upper margin, with more or less straight lower margin; with several long closely set setae near upper anterior corner, numerous long setae over lower margin becoming denser and stronger on about middle, moderately long and scattered setae on apical part of epandrium. Subepandrial sclerite produced slightly beyond lower margin of epandrium. Hypandrium very narrow, rim-like, entire, well sclerotized; bare; gonocoxal apodeme directed anteriorly. Phallus well exposed, extremely long; mostly hair-like, knob-like thickened just beyond hypandrium; slightly sinuate, with broad even curvature shortly beyond basal thickening, gently curved on remaining part. Ejaculatory apodeme not extended beyond basal curvature of phallus, with broad lateral wings and similar vertical wing. Female. Unknown.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C644A9F3857EEFEE2FC49EBBD.taxon	distribution	Distribution. Eurasia; known only from Wrangel Island.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C644A9F3857EEFEE2FC49EBBD.taxon	etymology	Etymology. The specific name refers to the similarity of the new species to R. macrura Loew.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C644A9F3857EEFEE2FC49EBBD.taxon	discussion	Remarks. The new species belongs to the R. pusilla group sensu Barták & Kubík (2009) (see also Sinclair et al. 2019: 65). It resembles R. macrura Loew described from East Siberia (Irkutsk) (Loew 1871). Rhamphomyia macrura is somewhat smaller (wing length 4.8 mm vs. 6.7 mm in R. submacrura sp. nov.), has brush-like arranged long setae on the mid basitarsus and much longer epandrial lamellae (Frey 1954: 427, textfig. 214), which are nearly as long as the abdomen. The females of both species are unknown.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64489F3E57EEFCDAFAF8EF21.taxon	description	(Figs 45 – 47) urn: lsid: zoobank. org: act: A 3936415 - AFA 7 - 4469 - 8 A 77 - E 07648 EAAAF 3	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64489F3E57EEFCDAFAF8EF21.taxon	materials_examined	Type material. HOLOTYPE male, labelled: [Chukotka AO (Wrangel Island):] “ [printed in Cyrillic, Russian] bass. [= basseyn, basin] r. [= reka, river] Khischniki / 7 km SO g. [= gora, mountain] Sovetskaya / o. [= ostrov, island] Wrangel / Gorodkov 15. vii. [1] 972 ”; “ Holotypus / Rhamphomyia / subtenuiterfilata / Shamshev, Sinclair sp. n. [red label] ” (INS _ DIP _ 0000616, ZIN; terminalia dissected, in microvial pinned with specimen). PARATYPES: RUSSIA. Chukotka AO (Wrangel Island): Somnitelnaya Bay, 400 m, S Sovetskaya Mtn, 20. vii. 1966, KBG (1 ♀, ZIN). Yakutia: Indigirka River, lower flow of river Ystan-Yuryakh, Momskiy District, 29. vi. 1976, V. Kovalev (1 ♂, ZMMU).	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64489F3E57EEFCDAFAF8EF21.taxon	diagnosis	Diagnosis. Mid-sized (wing length about 4.5 mm), blackish, subshiny flies. Male holoptic; thorax and abdomen black setose, scutum subshiny, anepisternum pruinescent; legs dark brown, hind basitarsus clavate, hind tibia with moderately long setae and hind tarsomeres 1 – 2 with numerous long fine setae dorsally; abdomen almost shiny, epandrium and cerci elongate, cercus spatulate beyond dorsal cavity; phallus well exposed, filamentous, long, gently curved. Female with brownish infuscate wing, short setose slender legs without pennate setae.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64489F3E57EEFCDAFAF8EF21.taxon	description	Description. Wing length 4.7 mm. Male (Fig. 45). Head with faint greyish pruinescence on face, frons, postgena, ocellar triangle and occiput; dark setose. Holoptic; upper ommatidia enlarged. Frons represented by very small subtriangular space below ocellar tubercle and larger subtriangular space above antennae, bare. Face broad, bare. Ocellar triangle with 2 moderately long fine and some short setae. Occiput with numerous long fine setae, including postoculars; postgena with numerous hair-like setae. Antenna blackish; scape short, slightly longer than pedicel, both with short setulae; postpedicel conical, nearly 2 X longer than wide; stylus very short, slightly shorter than half of postpedicel basal width. Palpus dark; with several long, dark fine setae. Proboscis with labrum dark brown, nearly as long as head height. Thorax dark in ground-colour, faintly greyish pruinescent (except noted), with black setation; postpronotal lobe and mesonotum almost uniformly shiny, very faintly pruinescent (somewhat denser on scutellum). Proepisternum with few moderately long fine setae on lower section; upper proepisternum in front of spiracle without setae. Prosternum bare. Antepronotum with several short setae. Postpronotal lobe with 1 long stronger and several shorter fine setae. Mesonotal setation: 1 presut spal (with some additional short fine setae), 3 npl (with several additional fine setae anteriorly), 1 psut spal (with additional setulae anteriorly), 1 pal, 4 sctl (in holotype 5); acr very short, cilia-like, arranged in 2 close irregular rows, lacking on prescutellar depression; presutural dc similar to acr, fine, arranged in 2 irregular rows, postsutural dc 1 – 2 - serial, 3 – 4 prescutellars longest. Laterotergite with several long fine setae. Anterior and posterior spiracles brown. Legs slender (except noted), uniformly brownish; mostly shiny, coxae and trochanters faintly greyish pruinescence; black setose. Fore coxa covered with long fine setae anteriorly. Hind femur slightly thickened on about apical half; hind tibiae slightly evenly thickened toward apex; hind basitarsus clavate, about 1.5 X broader at middle than apex of hind tibia; hind tarsomere 2 expanded. All femora whitish pilose ventrally, short setose (except noted). Fore femur with minute anteroventral and posteroventral setae. Mid femur with rows of moderately long anteroventrals (4 – 5 setae near base longer) and posteroventrals (somewhat longer on about apical half). Hind femur with numerous moderately long setae anteroventrally on about apical third. Fore and mid tibiae with some slightly longer fine setulae posteriorly, mid tibia with numerous stronger long anteroventral and posteroventral setae on about apical 2 / 3; hind tibia with moderately long fine setae dorsally (longest setae only slightly longer than tibia width), 1 long seta in posteroapical comb. Fore and mid tarsomeres short setose; hind tarsomeres 1 – 2 with long, dense erect fine setae dorsally, tarsomere 3 with slightly longer setae dorsally. Wing membrane faintly infuscate, with brownish veins; CuA + CuP (anal vein) incomplete, evanescent apically, almost reaching wing margin as fold. Pterostigma brownish yellow. Basal costal seta absent. Anal lobe welldeveloped; axillary incision right-angled. Squama brown, dark fringed. Halter brownish. Abdomen dark brown, almost shiny, very faintly pruinescent; mostly with black fine moderately long setae. Segments 6 – 7 unmodified. Segment 8 with separated tergite and sternite; tergite 8 simple, entire, subtriangular viewed laterally, very narrow mid-dorsally, broadly concave anteriorly, with almost straight posterior margin, numerous moderately long setae along posterior margin; sternite 8 simple, enlarged, subrectangular in lateral view, V-shaped viewed posteriorly, covered with numerous long setae posteriorly. Terminalia (Fig. 46) dark brown, directed obliquely upwards. Cerci (Fig. 47) separated from each other and from epandrium, not extended beyond tergite 8 anteriorly; cercus long, broadly deeply concave on upper margin about middle, its middle portion very narrow, apical portion spatulate (lateral view), about 2.5 X as broad as middle portion, with small internal subrectangular projection slightly beyond middle of cercus (dorsal view); covered with short fine setae on broadened anterior and posterior parts, lacking setation on narrowed middle portion; without pair of subepandrial lobes beneath. Epandrium subrectangular, elongate, about 4 X as long as broad, broadly concave on upper margin, with straight lower margin, somewhat broadened and angularly expanded apically, extended slightly beyond cerci posteriorly; with row of 6 – 7 long dark setae near upper anterior corner, covered with dense fine setulae on broadened apical part, some numerous moderately long setae along lower margin. Hypandrium very narrow, rim-like, entire, well sclerotized; bare. Phallus well exposed, long; mostly hair-like, knob-like thickened just beyond hypandrium; with right-angle curvature shortly beyond basal thickening, otherwise gently sinuate. Ejaculatory apodeme slightly extended beyond basal curvature of phallus, with narrow lateral wings and broad vertical wing. Female. Frons broad, greyish pruinescent, with marginal setulae; body with shorter setation; hind femur uniformly slender, with short anteroventral setae over entire length; hind tibia and tarsi slender, covered with short setae; wing darker, distinctly brownish infuscate; abdominal segments 5 – 8 somewhat denser pruinescent (lateral view); cercus long, slender, with dark setulae.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64489F3E57EEFCDAFAF8EF21.taxon	distribution	Distribution. The new species is currently known only from Yakutia and Wrangel Island of Russia.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64489F3E57EEFCDAFAF8EF21.taxon	etymology	Etymology. The specific name refers to the similarity of the new species to R. tenuiterfilata Becker, 1900.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64489F3E57EEFCDAFAF8EF21.taxon	discussion	Remarks. The new species is very similar to R. tenuiterfilata (known from Taymyr Peninsula) in the structure of the male hind leg and the terminalia. However, R. subtenuiterfilata differs from R. tenuiterfilata primarily by brownish halteres (vs. yellowish) and black setose abdomen (vs. pale setose). Habitat. On Wrangel Island, the species is known from single specimens collected in the mountains.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C644E9F3F57EEF966FF23E885.taxon	description	(Fig. 48)	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C644E9F3F57EEF966FF23E885.taxon	materials_examined	Material examined. RUSSIA. Yakutia (New Siberian Islands): centre of Kotelny Island, upper flow of river Balykhtakh near mouth of river Tuguttakh, arctic desert on flowers, 15. vii. 1973, KBG (20 ♂, 20 ♀, ZIN). Recognition. Mid-sized (wing length 3.6 – 4.5 mm) blackish flies. Male (Fig. 48) holoptic; anepisternum with shiny spot; scutum with 2 narrow shiny median vittae (sometimes with less distinct subshiny vitta on supra-alar space); thorax black setose, presutural dc long, fine, 2 – 3 - serial, acr similar to dc, biserial, 10 – 12 sctl; legs dark brown, hind basitarsus thickened apically, all tibiae and basitarsi clothed in dense long fine setae, hind tibia covered with these setae over entire dorsal face; wing infuscate, CuA + CuP incomplete, halter brown; abdomen very faintly pruinescent, almost shiny, with black setae; epandrium elongate, slightly constricted at middle; cercus elongate, about 2 / 3 length of epandrium, slightly constricted medially; phallus well exposed, filamentous, long, gently curved slightly beyond epandrium. Female with darker wing, short setose slender legs without pennate setae.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C644E9F3F57EEF966FF23E885.taxon	distribution	Distribution. Holarctic; R. ursinella is widespread across the low to high arctic region of North America, east of Hudson Bay (Sinclair et al. 2019). In Eurasia, this species was found only from New Siberian Islands of Arctic Russia.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C644F9F2257EEF961FD0CEA65.taxon	description	(Figs 49 – 53) urn: lsid: zoobank. org: act: D 1 A 835 B 2 - FA 2 B- 4104 - B 593 - F 6 A 43 BDDFA 89	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C644F9F2257EEF961FD0CEA65.taxon	materials_examined	Type material. HOLOTYPE ♂, labelled [RUSSIA. Chukotka AO (Wrangel Island)]: “ [printed in Cyrillic, Russian] bukh. [= bukhta, bay] Somnitelnaya, / south of o. [= ostrov, island] Wrangel / Gorodkov 19. vii. [1] 966 ”; “ on flower / Dryas integrifolia, / spotted / tundra ”; “ Holotypus / Rhamphomyia / wrangeli / Shamshev, Sinclair sp. n. [red label] ” (INS _ DIP _ 0000617, ZIN). PARATYPES: RUSSIA: Chukotka AO (Wrangel Island): same data as holotype, (1 ♂, 7 ♀, ZIN; 1 ♂, 1 ♀, CULSP); environs of Somnitelnaya Bay, south of Wrangel Island, valley of river Somnitelnaya, Mineev Mtns, 150 m, pebbles, on flower of Dryas integrifolia, 19. vii. 1966, KBG (1 ♂, 4 ♀, ZIN; 1 ♂, 1 ♀, CNC); same locality, on flower of Potentilla emarginata, 22. vii. 1966, KBG (2 ♂, ZIN).	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C644F9F2257EEF961FD0CEA65.taxon	diagnosis	Diagnosis. Small dark flies (wing length about 3.5 mm). Male holoptic; scutum almost uniformly brownish grey pruinescent, acr and dc biserial, usually 6 sctl; wing faintly brownish, anal vein incomplete, halter brown; hind tibia strongly curved inward on about basal third; abdominal segments 7 and 8 asymmetrical; terminalia small, slightly rotated to right, phallus mostly hidden. Female legs with simple setae.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C644F9F2257EEF961FD0CEA65.taxon	description	Description. Body length 3.2 – 3.5 (holotype 3.3); wing length 3.2 – 3.4 (holotype 3.2) mm. Male (Fig. 49). Head with dense greyish brown pruinescence on face, frons, postgena, ocellar triangle and occiput; dark setose. Holoptic; upper ommatidia enlarged. Frons represented by very small subtriangular space below ocellar tubercle and larger subtriangular space above antennae, bare. Face broad, bare. Ocellar triangle with several long fine setae. Occiput with numerous long setae, including postoculars; postgena with numerous hair-like setae. Antenna blackish brown; scape short, slightly longer than pedicel, both with short setulae; postpedicel conical, with slightly concave ventral margin on apical part, nearly 2.5 X longer than wide; stylus short, nearly as long as postpedicel basal width. Palpus dark; with several long, dark fine setae. Proboscis with labrum dark reddish-brown, nearly as long as head height. Thorax dark in ground-colour, greyish brown pruinescent; with black setation; scutum almost uniformly brownish grey pruinescent (dorsal view), without vittae. Proepisternum with tuft of several moderately long fine setae on lower section; upper proepisternum in front of spiracle without setae. Prosternum bare. Antepronotum with several short setae. Postpronotal lobe with 1 long and several short setae. Mesonotum with well-differentiated setae but their number, position and robustness variable; 1 presutural intra-alar, 1 – 2 presut spal (with some additional short fine setae), 3 – 4 npl (with finer short setae anteriorly), 1 – 2 psut spal (with additional setulae anteriorly), 2 pal, 6 – 8 sctl (usually 6); acr short, arranged 2 close irregular rows, lacking on prescutellar depression; presutural dc longer than acr, arranged in 2 irregular rows, postsutural dc 1 – 2 - serial, 3 – 4 prescutellars longest. Laterotergite with numerous long fine setae. Anterior and posterior spiracles brown. Legs slender, uniformly brownish; mostly subshiny, coxae and trochanters with denser greyish pruinescence; black setose. Fore coxa covered with long fine setae anteriorly. Hind tibia strongly curved inward on about basal third (posterior view), remaining podomeres unmodified. Mid and hind femora whitish pilose ventrally. Fore femur with rows of anteroventral and posteroventral very short fine setae (somewhat longer near base of femur), some moderately long setae anteriorly toward apex; mid femur with rows of short anteroventral and posteroventral setae (longest setae at most as long as half of femur width), some moderately long setae anteriorly; hind femur with moderately long anteroventral setae somewhat denser and longer on about apical third, some similar setae anteriorly and posteriorly. Fore tibia covered with moderately long fine setae posterodorsally (some setae usually slightly stronger); mid tibia with 3 – 4 posterodorsal, short spine-like setae ventrally toward apex; hind tibia covered with moderately long setae dorsally (longest setae slightly longer than tibia median width); no seta in posteroapical comb. Fore basitarsus with similar to fore tibia setae posterodorsally; hind basitarsus with similar to hind tibia setae posterodorsally. Wing membrane faintly brownish, with brownish veins; CuA + CuP (anal vein) incomplete. Pterostigma brownish yellow. One basal costa seta present. Anal lobe well-developed; axillary incision right-angled. Squama brown, dark fringed. Halter brown. Abdomen (Fig. 50) with segments 1 – 6 brownish, segments 7 – 8 blackish brown, all segments faintly brownish grey pruinescent, subshiny; segments 1 – 6 mostly covered with moderately long dark setae, short setae on tergites dorsally. Tergite and sternite 6 slightly concave posteriorly. Sclerites of segments 7 and 8 strongly modified (Figs 51 – 53), asymmetrical and stronger sclerotized (darker) than preceded segments. Segment 7 with tergite and sternite narrowly fused anteriorly. Tergite 7 slightly asymmetrical, large, longer than tergite 6, subrectangular viewed dorsally, with straight posterior margin; minute setulae posteriorly. Sternite 7 strongly asymmetrical, inflated, narrowly deeply concave posteromedially, with produced left lateral margin, free laying internal lateral narrow sclerite supporting anterior lateral projections of segment 8; covered with numerous short setae denser and somewhat stronger around posterior concavity. Segment 8 slightly rotated to right, with fused tergite and sternite (no trace of suture), slightly asymmetrical ventrally; large subtriangular anterior dorsomedial projection and 2 broad digitiform anterior lateral projections inserted into tergite 7, subtriangular anterior ventromedial projection (corresponding to posterior concavity on sternite 7); numerous long setae along posterior margin laterally and ventrally, scattered minute setulae dorsally. Terminalia (Figs 51 – 53) slightly rotated to right, small, dark brown. Cerci separated from epandrium, not extended anteriorly beyond posterior margin of tergite 8, without pair of subepandrial lobes beneath; in lateral view cercus shorter than epandrium, subrectangular, with straight upper margin, slightly rounded apically; in dorsal view cercus strongly convex laterally and deeply concave on inner margin, with short broad subapical projection directed downward; mostly covered with minute setulae, subapical projection with dense spinule-like setulae. Epandrium narrowly subtriangular; with some moderately long setae along lower margin and closer to apex. Hypandrium invisible in situ, mostly membranous, only lateral arms sclerotised. Phallus mostly hidden, only short portion between posterior margin of sternite 8 and lower margin of epandrium visible; short, thicker on about basal third and uniformly slender on remaining portion, gently arcuate, hook-like curved to left at extreme apex. Ejaculatory apodeme slightly extended beyond basal curvature of phallus, with subequally narrow lateral and vertical wings. Female. Similar to male, except frons broad, with marginal setulae, ommatidia of equal size; sometimes upper proepisternum in front of spiracle with 1 seta; legs with short setae, no pennate setae, hind tibia simple; wing darker, uniformly brownish infuscate; cercus long slender, with dark setulae.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C644F9F2257EEF961FD0CEA65.taxon	distribution	Distribution. Russia (Chukotka Autonomous Okrug: Wrangel Island).	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C644F9F2257EEF961FD0CEA65.taxon	etymology	Etymology. The specific name is a patronym in honour of Ferdinand Petrovich Wrangel (Baron Ferdinand von Wrangel, a Baltic German), a Russian explorer and seaman, and one of the founders of the Russian Geographic Society. Wrangel Island was named in 1867 by Thomas W. Long, an American whaling captain.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C644F9F2257EEF961FD0CEA65.taxon	discussion	Remarks. The male of the new species can be readily distinguished from other groups of Pararhamphomyia recognised by Barták & Kubík (2009) in the characters noted in the key. Several undescribed species of this group occur in North America. Habitat. This species was collected from Wrangel Island only in 1966, and in one area (environs of Somnitelnaya Bay), mainly on the flowers of Dryas integrifolia.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64529F2357EEFCA2FD1FEB69.taxon	description	(Fig. 54)	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64529F2357EEFCA2FD1FEB69.taxon	materials_examined	Material examined. RUSSIA. Murmanskaya Prov. (Kildin Island): Kola Bay, 23. viii. – 1. ix. 1923, Serebrjansky (1 ♂, ZIN). Nenets AO (Kolguev Island): Bugrino, sea shore, meadow, 21 – 22. viii. 1970, KBG (3 ♂, 2 ♀, ZIN).	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64529F2357EEFCA2FD1FEB69.taxon	description	Recognition. Mid-sized (wing length 5 – 6.5 mm) greyish, dark-legged, black setose flies. Male (Fig. 54): holoptic; antennal postpedicel and stylus black, scape and pedicel somewhat paler; labrum 1.1 – 1.5 X longer than head height; palpus black, long, with several long strong setae. Thorax grey pruinose, scutum somewhat finer pruinescent, without vittae; prosternum bare, upper proepisternum in front of spiracle with a few fine setae; mesonotal setae strong bristle-like (including acr and dc), acr biserial, presutural dc 2 – 3 - serial, 4 – 6 sctl; legs blackish brown, long, slender, mostly shiny to subshiny (except coxae); fore femur with single row of very long posteroventral setae; hind femur with 3 – 4 anterodorsal setae on about apical half, 3 – 6 strong anteroventral setae of different lengths over entire femur length and 1 long strong posteroventral seta on about basal third; all tibiae with rows of numerous very long strong setae dorsally, hind tibia with 1 long seta in posteroapical comb; fore basitarsus with several long setae dorsally. Wing hyaline, CuA + CuP (anal vein) complete, entirely sclerotized, 1 long basal costal seta present, axillary incision very acute, halter brown. Abdomen (except terminalia) densely uniformly light grey pruinescent, tergites with very long posteromarginal setae laterally; segment 7 with unmodified structure and vestiture; segment 8 with tergite and sternite separated; tergite 8 upturned posteriorly, with slightly convex posterior margin, without projections; sternite 8 nearly as long as sternite 7. Terminalia with cerci and epandrium brownish to reddish brown, faintly pruinescent; cerci separated from each other and from epandrium, not extended anteriorly beyond tergite 8; cercus slightly produced anteriorly, prolonged slightly beyond epandrium middle posteriorly, with narrow cavity near base dorsally forming short finger-like basal projection, rather trapezoid-like beyond cavity, lower posterior corner slightly produced downward, mostly covered with minute setulae, lower posterior corner with dense spinules, without pair of subepandrial lobes beneath; epandrium subtriangular, narrow, rounded apically, with long to moderately long setae more numerous along lower margin; subepandrial sclerite divided; hypandrium yellow, shiny, subtriangular, bare; phallus thick, vesicular apically; ejaculatory apodeme extended far beyond basal curvature of phallus, without lateral wings. Female: similar to male except, eyes dichoptic, frons very broad, parallel-sided; femora mostly with short setulae, only hind femur with 2 – 4 strong anteroventral setae; wing faintly brownish infuscate; abdominal tergites 5 – 8 and sternites with shorter setation.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64529F2357EEFCA2FD1FEB69.taxon	distribution	Distribution. Palaearctic; Fennoscandia, Great Britain, mountains of Central Europe, across northern Russia from Murmanskaya Province to Taymyr Peninsula.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64539F2057EEFDAEFDCDEBBD.taxon	description	(Fig. 55)	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64539F2057EEFDAEFDCDEBBD.taxon	description	2007: 185 (catalogue); Shamshev, 2016: 78 (checklist).	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64539F2057EEFDAEFDCDEBBD.taxon	materials_examined	Material examined. RUSSIA. Murmanskaya Prov. (Kildin Island): 13. vii. 1908, L. Bianki (1 ♂, ZIN). Recognition. Large (wing length 6 – 7.5 mm) blackish grey, dark-legged, black setose flies. Male (Fig. 55): holoptic, eyes not quite touching; antenna black, scape nearly 3 X longer than pedicel, both with long setae; labrum about 1.5 X longer than head height; palpus black, long, with numerous long fine setae; thorax mostly densely greyish pruinescent, scutum greyish brown, somewhat lustrous; prosternum bare, upper proepisternum in front of spiracle with several long fine setae; mesonotal setae intermixed with additional long fine setae, often hardly differentiated, 6 – 8 sctl; acr long, fine, 2 – 3 - serial, presutural dc slightly longer than acr, 3 – 4 - serial, not separated by bare space from supra-alars anteriorly; legs black, long, slender, mostly shiny to subshiny (except coxae); fore femur covered with dense, very long, fine, irregularly spread setae posteroventrally; hind femur with several anterodorsal setae on about apical half, 4 – 8 strong short anteroventral setae over entire femur length and some similar setae posteriorly, only pale pilose ventrally; all tibiae with rows of numerous very long strong setae dorsally, hind tibia with 1 long seta in posteroapical comb. fore and hind basitarsi with some longer setae dorsally; wing faintly brownish infuscate, CuA + CuP (anal vein) complete, entirely sclerotized, 1 long basal costal seta present, axillary incision very acute, halter brown; abdomen uniformly light grey pruinescent, denser on sternites; tergites somewhat lustrous, with very long posteromarginal setae; terminalia (Barták 1981: 396, fig. 17 a) very similar to R. albosegmentata.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64539F2057EEFDAEFDCDEBBD.taxon	distribution	Distribution. Palaearctic; Fennoscandia, Great Britain, mountains of Central Europe, northern European Russia. Species unplaced to subgenus	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64509F2157EEFD3EFA49E9A5.taxon	description	(Fig. 56)	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64509F2157EEFD3EFA49E9A5.taxon	materials_examined	Material examined. RUSSIA. Archangelskaya Prov. (Novaya Zemlya Archipelago): Novaja Zemlja, acad. Baer, 1837 (2 ♂, ZIN); Propaschaya Guba, sea shore, in mountains, 18. vii. 1912, S. Skribova (1 ♀, ZIN). Recognition. Mid-sized (wing length 5.5 – 6 mm) blackish, dark-legged, black setose robust flies. Male (Fig. 56): holoptic; antenna dark, scape and pedicel with long fine setae on outer side, minute setulae on inner side; palpus dark with numerous long fine setae; labrum nearly 1.3 X longer than head height; thorax mostly greyish brown pruinose; scutum dark brown pruinescent, slightly lustrous, with two hardly visible and slightly paler vittae between rows of acr and dc; prosternum bare, upper proepisternum in front of spiracle with several long fine setae; mesonotal setae mostly fine (including acr and dc), hardly differentiated, acr 2 – 3 - serial, present on anterior half of prescutellar depression, presutural dc multiserial, not separated from supra-alars by bare space, 12 – 20 sctl; legs robust, hind tibia slightly uniformly incrassate; fore femur with numerous long fine setae anteroventrally, posteroventrally and posteriorly; mid femur with very short anteroventral (except near base) and somewhat longer posteroventral setae; hind femur with numerous moderately long setae anteroventrally and ventrally; fore and mid tibiae with rows of several moderately long strong setae dorsally and some fine setae posteriorly; hind tibia with numerous long strong setae dorsally and long fine setae anteroventrally, 1 short seta in posteroapical comb; fore basitarsus with some short fine setae posteriorly, hind tarsomeres 1 – 3 with several long setae dorsally; wing faintly infuscate, CuA + CuP (anal vein) complete, entirely sclerotized, 1 long basal costal seta present, axillary incision acute, halter with brown knob and reddish brown stem; abdomen densely uniformly greyish brown pruinescent, with numerous long setae; structure of segments 7 – 8 as in R. kaninensis but posteromarginal setae more numerous and stronger; terminalia almost identical to R. kaninensis (including shape of phallus) but larger, cercus more gently rounded apically, with broader basal projection, epandrium with more numerous strong setae and denser cluster of fine setae subapically. Female similar to male, except eyes dichoptic, frons very broad, parallel-sided, body (especially legs and abdomen) with much shorter setation; wing slightly darker, faintly brownish infuscate; legs without pennate setae, hind tibia somewhat dilated, almost bare anteriorly.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64509F2157EEFD3EFA49E9A5.taxon	distribution	Distribution. Holarctic; R. hirtula appears to have a disjunct distribution. In North America, this species is recorded from several localities in Yukon and Greenland; in Eurasia, it is known from Iceland and Scotland, as well as from Novaya Zemlya Archipelago, Taymyr, Chukotka and high mountains of Altay Republic of Russia (Shamshev & Barkalov 2009; Sinclair et al. 2019).	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64509F2157EEFD3EFA49E9A5.taxon	discussion	Remarks. Shamshev (2016: 64) has noted R. hirtula from Wrangel Island, but with a question mark. Our study did not confirm the presence of this species on this island. Rhamphomyia hirtula belongs to the R. hirtula group.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64519F2757EEFEE2FB4AEAD5.taxon	description	(Figs 57 – 61)	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64519F2757EEFEE2FB4AEAD5.taxon	materials_examined	Material examined. RUSSIA. Krasnoyarskiy Terr. (Dikson Island): small island “ Konus ”, 31. vii. 1948, Korotkevich (1 ♂, ZIN). Additional material. RUSSIA. Krasnoyarskiy Terr. (Taymyr Peninsula): Engelgardt Lake, NE shore, valley of brook, 2. vii. 1967, KBG (3 ♂, 5 ♀, ZIN; 1 ♂, 1 ♀, CNC).	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64519F2757EEFEE2FB4AEAD5.taxon	diagnosis	Diagnosis. Mid-sized (wing length about 4.5 mm), robust, black setose flies with brownish thorax and greyish abdomen. Male mid tibia and mid basitarsus with numerous very long setae dorsally; hind tibia slightly uniformly thickened, hind basitarsus slender; abdomen densely light grey pruinescent; wing whitish, CuA + CuP (anal vein) complete, halter brown; abdominal tergite 8 without projections; cercus with finger-like projection near base dorsally; phallus well exposed, slender, gently sinuate on about middle. Female wing uniformly faintly brownish infuscate, legs without pennate setae.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64519F2757EEFEE2FB4AEAD5.taxon	description	Redescription. Body length 4.5 – 4.9; wing length 4.3 – 4.5 mm. Male (Fig. 57). Head with dense greyish brown pruinescence on face, frons, postgena, ocellar triangle and occiput; dark setose. Holoptic; upper ommatidia enlarged; eyes touching. Frons represented by very small subtriangular space below ocellar tubercle and larger subtriangular space above antennae, bare. Face broad, bare. Ocellar triangle with 2 long and several shorter fine setae. Occiput with numerous long fine setae, including postoculars; postgena with numerous hair-like setae. Antenna blackish brown; scape slightly longer than pedicel, both with somewhat longer fine setae on outer side and short setulae on inner side; postpedicel conical, nearly 3 X longer than wide; stylus short, nearly as long as postpedicel basal width. Palpus dark, long; with numerous long, dark fine setae. Proboscis with labrum dark reddish-brown, nearly 1.3 X as long as head height. Thorax dark in ground-colour, mostly greyish brown pruinescent; with black setation; scutum almost uniformly brownish pruinescent (dorsal view), slightly lustrous, only some traces of two narrow dark vittae between rows of acr and dc. Proepisternum with tuft of several long fine setae on lower section; upper proepisternum in front of spiracle with 3 – 5 similar setae. Prosternum bare. Antepronotum with numerous short setae. Postpronotal lobe with 1 (sometimes 2) longer somewhat stronger and numerous shorter fine setae. Mesonotal setae generally quite well differentiated (number, position and robustness variable); 1 – 2 presut spal (with additional shorter setae), 3 – 4 npl (with additional shorter setae anteriorly), 1 – 2 psut spal (with additional setae anteriorly), 1 long and 1 short pal, 4 – 6 sctl; acr moderately long, fine, arranged in 2 irregular rows, lacking on prescutellar depression; presutural dc slightly longer than acr, 3 – 4 - serial, hardly separated by from supra-alar setae, postsutural dc 1 – 2 - serial, 3 – 4 prescutellars longest. Laterotergite with numerous long fine setae. Anterior and posterior spiracles brown. Legs robust, uniformly dark brown; mostly subshiny, coxae and trochanters with denser greyish pruinescence; black setose. Fore coxa covered with long fine setae anteriorly. Hind tibia slightly uniformly thickened. Femora whitish pilose ventrally. Fore femur covered with very short setae including rows of fine anteroventrals and posteroventrals. Mid femur with similar setation to fore femur but anteroventral and posteroventral setae somewhat longer. Hind femur mostly with very short setae anteroventrally, only 6 – 7 anteroventral setae on about apical third moderately long; covered with moderately long setae anteriorly and on basal part posteriorly. Fore tibia with numerous fine, moderately long setae over entire length posterodorsally, 2 – 3 similar anterodorsal setae on about basal third. Mid tibia (Fig. 58) with numerous dense very long (2 – 2.5 X longer than tibia width) setae dorsally and posterodorsally; numerous anteroventral and posteroventral setae becoming spine-like closer to apex. Hind tibia (Fig. 59) with numerous long setae dorsally and shorter setae anteroventrally; 1 seta in posteroapical comb. Fore basitarsus with some fine setae posterodorsally (as on fore tibia); mid basitarsus with several very long setae dorsally (as on mid tibia); hind basitarsus with several long setae dorsally. Wing membrane whitish, with brownish veins; all veins complete (except Sc), CuA + CuP (anal vein) weaker sclerotised on about basal half. Pterostigma brownish yellow. Basal costal seta present, long. Anal lobe welldeveloped; axillary incision right-angle. Squama brownish, dark fringed. Halter with brown knob and reddish brown stem. Abdomen dark, densely light grey pruinescent; covered with short to moderately long black fine setae (except noted). Tergite 7 with unmodified structure but bearing 4 erect medial posteromarginal setae longer and stronger than posteromarginal setae on tergite 6. Sternite 7 with similar structure and setation to sternite 6. Segment 8 with tergite and sternite separated but pleural space indistinct. Tergite 8 about 2 X shorter than tergite 7, somewhat upturned posteriorly, with truncate posterior margin, without projections; bearing 2 long, strong erect posteromarginal setae medially. Sternite 8 large, somewhat longer than sternite 7 (lateral view); with numerous dense very long setae near posterior margin. Terminalia (Figs 60, 61) brownish, black setose. Cerci separated from each other and from epandrium, not extended anteriorly beyond tergite 8; covered with mostly fine setulae, some marginal setulae on inner face stronger; cercus slightly shorter than epandrium, subrectangular, rounded apically (lateral view), with dorsal notch forming long pointed finger-like projection on about basal third; short broad inner projection subapically (dorsal view); without pair of subepandrial lobes beneath. Epandrium subtriangular, rounded apically, with several long setae on about middle and along lower margin, bearing cluster of somewhat finer numerous dense setae near apex. Hypandrium invisible in situ, rim-like, entirely sclerotized, bare. Phallus (Fig. 59) exposed on about middle third; expanded on basal part, gently sinuate near middle, not extended beyond cerci. Ejaculatory apodeme slightly extended beyond basal curvature of phallus, with broad lateral wings and somewhat narrower vertical wing. Female (described for first time). Similar to male, except frons very broad, parallel-sided, with marginal setulae; eye ommatidia equally small; ocellar tubercle and occiput with shorter and stronger setae; thorax with somewhat shorter setation; legs covered with short simple setae; wing membrane uniformly faintly brownish infuscate, anal vein uniformly sclerotised; abdomen slightly darker pruinescent; cercus long slender, with dark setulae.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64519F2757EEFEE2FB4AEAD5.taxon	distribution	Distribution. Holarctic; in Eurasia the species is known only from the arctic region of Russia. This species is also known from North America (T. Saigusa, unpubl. data), but further details were not available.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64519F2757EEFEE2FB4AEAD5.taxon	discussion	Remarks. Rhamphomyia kaninensis belongs to the R. hirtula group. Frey described this species after three males collected from the Kanin Peninsula (Arkhangelskaya Province, Russia), one male in MZH was examined by IVS. Frey (1913: 8) noted that this species was collected on bushes of Salix in the tundra.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64579F2A57EEFC32FACDE885.taxon	description	(Figs 62 – 65) urn: lsid: zoobank. org: act: 99 F 127 D 3 - A 192 - 4 D 59 - 9030 - EEEB 1 F 7 E 3827	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64579F2A57EEFC32FACDE885.taxon	materials_examined	Type material. HOLOTYPE ♂, labelled: [RUSSIA, Chukotka AO (Wrangel Island)] “ [printed in Cyrillic, Russian] bukh. [= bukhta, bay] Somnitelnaya / south of o. [= ostrov, island] Wrangel / Gorodkov 19. vii. [1] 966 ”; “ on flower / Dryas integrifolia, / spotted / tundra ”; “ Holotypus / Rhamphomyia / sublongiseta Shamshev, / Sinclair, Saigusa sp. n. [red label] ” (INS _ DIP _ 0000618, ZIN). PARATYPES: CANADA. Yukon: km 155 Dempster Hwy, 18 – 20. vi. 1980, 950 m, D. M. Wood & D. Lafontaine (1 ♂, CNC); Richardson Mts, 2800 ft, 66 ° 07 ′ N 136 ° 30 ′ W, 11. vi. 1982, D. M. Wood (1 ♂, 1 ♀, CNC). RUSSIA. Chukotka AO (Wrangel Island): SE part of Wrangel Island, Rogers Bay, tundra, 20. vii. 1932, A. Mineev (2 ♂, ZIN); same data as holotype, 19. vii. 1966, KBG (1 ♂, 1 ♀, ZIN; 1 ♂, 1 ♀, CNC); environs of Somnitelnaya Bay, south of Wrangel Island, valley of river Somnitelnaya, Mineev Mtns, 150 m, shingle, on flower of Dryas integrifolia, 19. vii. 1966, KBG (2 ♀, ZIN); same locality, on flower of Potentilla emarginata, 22. vii. 1966, KBG (1 ♀, ZIN); Somnitelnaya Bay, south of Wrangel Island, Mineev Mtns, 400 m, S of Sovetskaya Mtn, 20. vii. 1966, KBG (2 ♀, ZIN); 5 km N Somnitelnaya Bay, warmed up on stones river bank, 27. vii. 1966, KBG (1 ♀, ZIN); 5 km N Somnitelnaya Bay, valley of river Somnitelnaya, small meadow, 26. vii. 1972, KBG (1 ♀, ZIN); environs of Aterton Mtn, 70 ° 59 ′ N 178 ° 41 ′ W, BT B, Sw, 13. vii. 2019, OAK (1 ♂, 1 ♀, ZIN). Additional material. RUSSIA, Chukotka AO (Wrangel Island): middle flow of river Neozhidannaya, 71 ° 01 ′ N 179 ° 09 ′ E, YPT, BT 1, 12 – 16. vii. 2016, LFV (10 ♂, 10 ♀, ZIN; 1 ♂, 1 ♀, CULSP); same locality, YPT, BT 2, 12 – 16. vii. 2016, LFV (37 ♂, 28 ♀ ZIN); Thomas Mtn, 70 ° 56.964 ′ N 178 ° 43.026 ′ E, BT 16, 10 – 31. vii. 2016, LFV (2 ♂, ZIN).	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64579F2A57EEFC32FACDE885.taxon	diagnosis	Diagnosis. Mid-sized (wing length about 4.5 mm), robust, black setose, blackish brown flies. Male holoptic; hind tibia thickened, hind basitarsus strongly expanded, basal area broader than apex of hind tibia, elongate oval, somewhat narrowed apically, both clothed in very long, dense fine setae; abdominal tergite 8 without projections; cercus truncate at apex (lateral view), with notch slightly beyond middle dorsally; phallus well exposed, slender, gently sinuate near middle. Female wing darker near base, legs without pennate setae.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64579F2A57EEFC32FACDE885.taxon	description	Description. Body length 4.2 – 4.7 (holotype 4.7); wing length 4.3 – 4.5 (holotype 4.4) mm. Male (Fig. 62). Head with dense greyish brown pruinescence on face, frons, postgena, ocellar triangle and occiput; dark setose. Holoptic; upper ommatidia enlarged. Frons represented by very small subtriangular space below ocellar tubercle and larger subtriangular space above antennae, bare. Face broad, bare. Ocellar triangle with several long fine setae. Occiput with numerous long fine setae, including postoculars; postgena with numerous hair-like setae. Antenna blackish brown; scape and pedicel subequally short, both with long fine setae on outer side and short setulae on inner side; postpedicel conical, with straight margins, nearly 3 X longer than wide; stylus very short, shorter than postpedicel basal width. Palpus dark; with numerous long, dark fine setae. Proboscis with labrum dark reddish-brown, slightly longer than head height. Thorax dark in ground-colour, greyish brown pruinescent; with black setation; scutum almost uniformly brownish pruinescent (dorsal view), only some traces of two narrow vittae between rows of acr and dc. Proepisternum with tuft of numerous long fine setae on lower section; upper proepisternum in front of spiracle with numerous similar setae. Prosternum bare. Antepronotum with numerous short setae. Postpronotal lobe with numerous long fine setae. Mesonotal setae hardly differentiated, numerous, long, almost uniformly fine, only some npl, pal, sctl and prescutellar dorsocentrals somewhat stronger; 1 – 2 pal, 6 – 10 sctl (usually 6 – 8); acr long, fine, arranged in 2 irregular rows, lacking on prescutellar depression; presutural dc long, multiserial, not separated by bare space from supra-alar setae, some prescutellars somewhat stronger. Laterotergite with numerous long fine setae. Anterior and posterior spiracles brown. Legs robust, uniformly dark brown; mostly subshiny, coxae and trochanters with denser greyish pruinescence; black setose. Fore coxa covered with long fine setae anteriorly. Podomeres of fore and mid legs unmodified; hind femur somewhat broader near apical half; hind tibia expanded, slightly arcuate; hind basitarsus strongly expanded, basal area broader than apex of hind tibia, elongate oval, somewhat narrowed apically; hind tarsomeres 2 – 3 slightly expanded. Mid and hind femora whitish pilose ventrally. Legs mostly covered with fine setae, only mid and hind tibiae with some stronger setae. Fore femur clothed in long setae including anteroventral and posteroventral rows (about as long as femur width); mid femur with row of short anteroventrals and row of short posteroventral setae on basal half and longer on apical half, some short setae near extreme base ventrally; hind femur without setae ventrally, with several long av setae on about apical third (otherwise av minute), with more or less distinct row of long setae anteriorly and some long setae around apex and base. Fore tibia with 2 – 3 more or less distinct ad (except circlet of short subapicals), covered with moderately long fine setae posterodorsally; mid tibia with 2 – 3 ad, fine setae posterodorsally (sometimes 1 – 2 setae somewhat stronger), rows of numerous short spine-like av and pv setae; subapical circlet with 1 long ad seta; hind tibia clothed in very long dense curved setae; 1 seta in posteroapical comb. Fore and mid basitarsi with some longer fine setae posterodorsally, spine-like setae ventrally; hind basitarsus with numerous very long setae dorsally and ventrally (the latter stronger), hind tarsomeres 2 – 4 with long setae dorsally. Wing membrane faintly brownish, with brownish veins; all veins complete (except Sc). Pterostigma brownish yellow. Basal costal seta usually present, short (sometimes indistinct). Anal lobe well-developed; axillary incision acute but closer to 90 °. Squama brownish, dark fringed. Halter brown. Abdomen dark, densely brownish pruinescent; covered with uniform black fine setae shorter on tergites dorsally (except noted). Tergite 7 slightly convex, almost devoid of setation, with only 5 – 6 short posteromarginal setae medially and some setulae laterally. Sternite 7 with similar structure and setation to sternite 6. Segment 8 with tergite and sternite separated. Tergite 8 narrow, somewhat upturned posteriorly, slightly narrowly concave posteromedially; with cluster of several long setae near posterior margin laterally. Sternite 8 large, somewhat convex on posterior part ventrally, weaker sclerotized along midline (especially posteriorly); with numerous long setae along posterior margin, otherwise bare. Terminalia (Figs 63 – 65) brownish. Cerci separated from each other and from epandrium, not extended anteriorly beyond tergite 8; mostly grey pruinescent, covered with dark mostly fine setulae, spinule-like setulae on inner face subapically; cercus slightly shorter than epandrium, subrectangular and truncate at apex (lateral view), with notch forming small projection slightly beyond middle dorsally; without pair of subepandrial lobes beneath. Epandrium subtriangular, mostly grey pruinescent, subshiny anteriorly, with numerous long black setae along lower margin. Hypandrium small, subtriangular (ventral view), narrowly sclerotized along margin; bare. Phallus (Fig. 65) well exposed; mostly slender, evenly expanded on basal part, gently sinuate near middle, short apical part extended beyond cerci. Ejaculatory apodeme not extended beyond basal curvature of phallus, with broad lateral wings and narrow vertical wing. Female. Similar to male, except frons broad, with marginal setulae; eye ommatidia equally small; ocellar tubercle and occiput with shorter and stronger setae. Thorax with shorter setation; mesonotum with somewhat sparser and more differentiated setae, 3 – 4 npl, 6 – 8 sctl. Legs without pennate setae, shorter setose; mid femur with dense uniformly long posteroventral setae, mid tibia without av and pv spine-like setae; hind femur less distinctly narrowed on basal part, with longer av setae on basal part, dense long posteroventral setae; hind tibia and basitarsus only slightly expanded (latter as broad as hind tibia at apex), with rather short setae dorsally and ventrally (nearly as long as hind tibia or basitarsus width); tarsomeres 2 – 4 rather slender, with short setae. Wing membrane darker near wing base (about level of cells br and bm); basal costa seta usually distinct, short. Abdomen densely light grey pruinescent; cercus long slender, with dark setulae.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64579F2A57EEFC32FACDE885.taxon	distribution	Distribution. Holarctic; in Eurasia R. sublongiseta sp. nov. is recorded only from Wrangel Island; in North America this species is known only from Yukon.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64579F2A57EEFC32FACDE885.taxon	etymology	Etymology. The specific name refers to the similarity of the new species to R. longiseta Saigusa, 1964.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64579F2A57EEFC32FACDE885.taxon	discussion	Remarks. The new species belongs to the R. hirtula group (Saigusa 1964; Sinclair et al. 2019). In the structure and setation of the male hind leg, R. sublongiseta sp. nov. resembles R. longiseta known from the alpine region of Hokkaido, Japan (Saigusa 1964). However, the new species differs from R. longiseta by the more extensively setose hind tibia (with long setae only dorsally in R. longiseta), simple abdominal tergite 8 (with pair of short spatulate projections in R. longiseta), brownish pruinescent abdomen (ash grey in R. longiseta) and darker wing base of the female. Habitat. On Wrangel Island this species was collected mostly in the colder regions (Neozhidannaya River and some coastal areas), where it was captured in various dry biotopes. Apparently, the species is characterized by sharp interseasonal fluctuations in numbers, as evidenced by both Gorodkov’s data (who collected it only in 1966) and ours (on the Neozhidannaya River this species was numerous in 2016, but was not recorded in 2019).	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C645A9F2957EEFD82FD82E815.taxon	description	(Figs 66, 67)	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C645A9F2957EEFD82FD82E815.taxon	description	et al., 2007: 189 (catalogue); Shamshev, 2016: 82 (checklist).	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C645A9F2957EEFD82FD82E815.taxon	materials_examined	Material examined. RUSSIA. Arkhangelskaya Prov. (Novaya Zemlya Archipelago): Chekin Bay, 1. vii. 1901, Timofeev (1 ♂, 1 ♀, ZIN); Matochkin Shar, radio station, 8. vii. 1924, Tolmachow (1 ♀, ZIN). Chukotka AO (Wrangel Island): Somnitelnaya Bay, on flowers of Papaver, 18. vii. 1966, KBG (1 ♀, ZIN); same locality, on flowers of Dryas integrifolia, spotted tundra, 19. vii. 1966, KBG (8 ♂, 11 ♀, ZIN; 1 ♂, 1 ♀, CNC); same locality, sea shore at midnight on flowers of Papaver polaris, 25. vii. 1966, KBG (5 ♂, 3 ♀, ZIN); same locality, on flowers of Potentilla emarginata, 26. vii. 1966, KBG (3 ♀, ZIN); same locality, spotted tundra near village, 9. vii. 1972, KBG (5 ♂, 7 ♀, ZIN); 5 km N Somnitelnaya Bay, 28. vii. 1971, KBG (1 ♂, 1 ♀, ZIN); Somnitelnaya Bay, valley of river Somnitelnaya, southern slope of Mineev Mtn, 100 m, shingle, 26. vii. 1966, KBG (3 ♀, ZIN); environs of Somnitelnaya Bay, valley of river Somnitelnaya, Mineev Mtns, 150 m, shingle, on flower of Dryas integrifolia, 22. vii. 1966, KBG (1 ♂, ZIN); Somnitelnaya River, 71 ° 00 ′ N 179 ° 32 ′ W, BT 9 ВG, 14. vii. 2006, OAK (1 ♂, 1 ♀, ZIN); same locality and biotope, 18. vii. 2015, OAK (1 ♀, ZIN) environs of Tundrovaya Mtn, near stream, 18. vii. 1972, KBG (1 ♂, 2 ♀, ZIN); middle flow of Mamontovaya River, N Perkatkun, osier-bed in valley of river, 17. vii. 1972, KBG (2 ♂, 1 ♀, ZIN); same locality, 71 ° 10 ′ N 179 ° 45 ′ W, Arctic fox burrow, 11. vii. 2006, OAK (2 ♂, 2 ♀, ZIN); same locality and biotope, 11 – 21. vii. 2006, OAK (1 ♂, ZIN); same locality and biotope, 1. vii. 2015, OAK (1 ♂, ZIN); same locality, BT 6, 9. vii. 2006, OAK (2 ♂, 1 ♀, ZIN); same locality, BT 5 А, 20. vii. 2006, OAK (3 ♂, 15 ♀, ZIN); same locality, BT 5, 30. vi. 2015, OAK (3 ♂, 10 ♀, ZIN); same locality, BT 10, 8. vii. 2006, OAK (4 ♂, 3 ♀, ZIN); same locality and biotope, 30. vi. 2015, OAK (2 ♂, ZIN); same locality, BT 17, 3. vii. 2015, OAK (2 ♂, 1 ♀, ZIN); same locality, BT 1 А, YPT, 2 – 4. vii. 2015, OAK (1 ♂, ZIN); same locality, house, 30. vi. 2015, OAK (1 ♀, ZIN); spurs of Pervaya Mtn, 71 ° 09 ′ N 179 ° 27 ′ W, BT 13, 28. vi. 2015, OAK (2 ♂, 8 ♀, ZIN); same locality and biotope, 28. vi. – 13. vii. 2015, OAK (2 ♀, ZIN); same locality, BT 15, 28. vi. 2015, OAK (1 ♂, 1 ♀, ZIN); same locality and biotope, 28. vi. – 13. vii. 2015, OAK (1 ♀, ZIN); upper flow of Neizvestnaya River, 71 ° 13 ′ N 179 ° 19 ′ W, BT C 4, 3. vii. 2006, OAK (11 ♀, ZIN); same locality and biotope, 12 – 29. vii. 2015, OAK (1 ♀, ZIN); same locality, BT С 2, 21. vi. 2015, OAK (2 ♀, ZIN); same locality, BT С 3, 21. vi. 2015, OAK (2 ♀, ZIN); same locality, BT 10 А, 22. vi. 2015, OAK (1 ♂, ZIN).	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C645A9F2957EEFD82FD82E815.taxon	diagnosis	Diagnosis. Mid-sized (wing length 4.5 – 5.3 mm) blackish flies. Male holoptic; body black setose; scutum greyish brown pruinescent, without vittae; acr biserial, presutural dc 3 – 4 - serial; hind tibia curved inward closer to base, hind basitarsus expanded, nearly as broad as apex of hind tibia; halter with yellow knob; CuA + CuP complete; phallus well exposed, long, with narrow U-shaped loop just beyond cerci. Female mid and hind femora with anterodorsal and posteroventral rows of pennate setae (except extreme base); hind tibia with some subpennate setae dorsally and ventrally.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C645A9F2957EEFD82FD82E815.taxon	description	Redescription. Body length 4.6 – 5.5; wing length 4.5 – 5.3 mm. Male (Fig. 66, described for first time). Head with greyish brown pruinescence on face, frons, postgena, ocellar triangle and occiput; dark setose. Holoptic; upper ommatidia enlarged. Frons represented by very small subtriangular space below ocellar tubercle and larger subtriangular space above antennae, with scattered marginal setulae; with single setulae or bare. Face broad, bare. Ocellar triangle with 2 long and several short fine setae. Occiput with numerous long thin setae, including postoculars; postgena with numerous finer setae than occiput. Antenna blackish brown; scape and pedicel short, scape slightly longer, both with short setulae; postpedicel conical, slightly concave on apical part ventrally, nearly 2.5 – 3 X longer than wide; stylus rather long, 1.3 – 1.5 X shorter than postpedicel. Palpus dark; with long, dark fine setae. Proboscis with labrum dark reddish-brown, 1.3 – 1.5 X head height. Thorax dark in ground-colour, greyish brown pruinescent; with black setation; scutum greyish brown pruinescent, somewhat lustrous, without vittae. Proepisternum with tuft of numerous long setae on lower section; upper proepisternum in front of spiracle without setae. Prosternum bare. Antepronotum with numerous short setae. Postpronotal lobe usually with 2 long, stronger and several short fine setae (sometimes with additional long fine setae). Mesonotal setation (number, robustness and position of main mesonotal setae very variable, often even on right and left sides): several long fine setae on supra-alar space (psut spal more numerous), 3 – 5 npl (additional fine setae anteriorly), 1 long and 1 short pal, 6 – 8 sctl (often with additional short fine setae); acr long, thin, arranged in 2 irregular rows, absent on prescutellar depression; presutural dc long, arranged in 3 – 4 irregular rows, postsuturals 1 – 2 - serial, 3 – 4 prescutellars longest. Laterotergite with numerous long fine setae. Anterior and posterior spiracles brown. Legs uniformly dark brown; mostly subshiny, coxae and trochanters with denser greyish pruinescence; black setose. Fore coxa covered with long fine setae anteriorly. Femora and tibiae slender; hind femur only slightly thicker than fore and mid femora; hind tibia curved inward closer to base (posterior view); hind basitarsus thickened, nearly as broad as hind tibia at apex. Mid and hind femora whitish pilose ventrally; fore femur with rows of fine, long anteroventral and posteroventral setae (nearly as long as femur width) and similar setae anteriorly; mid femur with rows of short fine anteroventral and long posteroventral setae; hind femur with complete row of rather long fine anteroventral setae. Fore tibia with 2 – 3 short anterodorsal and long fine setae posterodorsally; mid tibia with 3 – 4 anterodorsal and 3 – 4 posterodorsal setae (except circlet of subapicals) becoming longer toward apex of tibia (longest setae nearly 1.5 X longer than tibia width), several short spine-like anteroventral and posteroventral setae on about apical half. Hind tibia with numerous long anteroventral and posteroventral setae, lacking setulae on concave space near base; 1 seta in posteroapical comb of hind tibia. Fore basitarsus with somewhat longer setae dorsally (nearly as long as basitarsus width); mid and hind basitarsi with numerous long setae dorsally (longest setae nearly twice longer that corresponding basitarsus width). Wing membrane faintly brownish, mostly with brownish veins; all veins complete (except Sc); CuA + CuP (anal vein) weakened on subapical portion. Pterostigma brownish yellow; 1 basal costa seta present, short. Anal lobe welldeveloped; axillary incision almost 90 °. Squama brownish, dark fringed. Halter with yellow knob and brownish yellow stem. Abdomen dark brown, faintly brownish pruinescent, subshiny; covered with uniform black fine setae shorter on tergites dorsally, sternite 8 with numerous very long posteromarginal setae. Segments 6 – 7 with unmodified vestiture and structure. Segment 8 with tergite and sternite separated, both normally sclerotized; tergite 8 concave posteromedially; sternite 8 unmodified, subrectangular viewed laterally, nearly as long as sternite 7. Terminalia dark brown (Fig. 67). Cerci directed posteriorly, separated from each other and from epandrium; covered with numerous fine dark setulae dorsally, numerous spinule-like setae along lower margin; cercus subrectangular (lateral view), with tapered apex, about as long as epandrium and half as broad as epandrium; without pair of subepandrial lobes beneath. Epandrium subrectangular, broadly rounded apically; shiny on basal region; covered with long, fine dark setae. Hypandrium mostly membraneous, only lateral arms sclerotized, bare. Phallus yellowish to brownish yellow, well exposed; evenly curved and uniformly slightly broader on about basal half until lower margin of epandrium, tapered apically; slightly curved between cerci, with narrow U-shaped loop just beyond cerci. Ejaculatory apodeme not extended beyond basal curvature of phallus, with narrow lateral wings and without prominent vertical wing. Female. Similar to male, except face broad with lateral setulae; mid and hind femora with anterodorsal and posteroventral rows of pennate setae (except extreme base); hind tibia with some subpennate setae dorsally and ventrally. Cercus long slender, covered with dark setulae.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C645A9F2957EEFD82FD82E815.taxon	discussion	Remarks. The male was associated with the female after a pair taken in copula, which was found among material collected from the type locality. The female was compared with the syntypes housed in ZIN. Frey (1950) placed R. taimyrensis in his subgenus Dasyrhamphomyia, but this is not confirmed by our discovery of a male of this species. Actually, R. taimyrensis is most closely related to R. laevigata Loew, revised very recently by Sinclair et al. (2019), which is widely distributed in North America and also was known for a long time from the extreme north of Russia as R. fridolini Frey. The subgeneric position of these species remains unclear.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C645A9F2957EEFD82FD82E815.taxon	distribution	Distribution. Palaearctic; R. taimyrensis is broadly distributed across the Russian Arctic including continental areas and islands (Novaya Zemlya Archipelago, Wrangel). Habitat. On Wrangel Island, this species was not found in the colder region (Neozhidannaya River), but in one year (1966) it was numerous in the coastal spotted tundra. In the 2000 s, this species was collected mainly in the warmer central region of the island, in dry habitats, especially often in tundra-steppe-like vegetation. Unnamed species of Rhamphomyia	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64599F2957EEFE56FE60ECFF.taxon	description	(Fig. 68)	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64599F2957EEFE56FE60ECFF.taxon	materials_examined	Material examined. RUSSIA. Chukotka AO (Wrangel Island): env. of Somnitelnaya Bay, south of Wrangel Island, 10. vii. 1966, KBG; upper flow of river Somnitelnaya, Mineev Mtns; on flower of Potentilla (1 ♀, ZIN).	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64599F2957EEFE56FE60ECFF.taxon	description	Description. Female (Fig. 68). Length body 3.6 mm, wing 3.7 mm. Head with greyish brown pruinescence on face, frons, postgena, ocellar triangle and occiput; dark setose. Eyes dichoptic, separated by broad frons. Antenna blackish brown; scape and pedicel short, scape slightly longer, both with short setulae; postpedicel conical, 3 X longer than wide; stylus very short, nearly as long as postpedicel basal width. Palpus dark; with several moderately long, dark fine setae. Labrum short, nearly of head height. Thorax brownish grey pruinescent; with black setation; scutum almost uniformly faintly brownish grey pruinescent, without vittae in dorsal view, with some traces of darker narrow vittae between acr and dc rows viewed anterodorsally. Proepisternum with few moderately long fine setae on lower section; upper proepisternum in front of spiracle without setae. Prosternum bare. Antepronotum with several short setae. Postpronotal lobe with 2 long and several shorter fine setae. Mesonotal setae fine, hardly differentiated; 2 longer presut spal with numerous additional short setae, 3 – 4 npl (with additional short setae), 1 psut spal (with several additional short setae anteriorly), 1 – 2 pal, 6 sctl; acr arranged in 2 irregular rows, lacking on prescutellar depression, shorter than presutural dc; presutural dc 3 – 4 - serial, long, barely separated from numerous supra-alars, postsutural dc 1 – 2 - serial, longer, 3 – 4 prescutellars longest. Laterotergite with numerous long fine setae. Anterior and posterior spiracles brown. Legs long, slender, uniformly brownish, faintly greyish pruinescent, dark setose; fore coxa with fine setae anteriorly; femora and tibiae covered with short setae, no strong and pennate setae; hind femur with very short anteroventral setae; tibiae with some longer setae dorsally, hind tibia with 1 seta in posteroapical comb. Wing membrane uniformly brownish infuscate; CuA + CuP (anal vein) incomplete; 1 short basal costal seta; axillary incision right-angled. Squama brownish, dark fringed. Halter with brownish knob and reddish brown stem. Abdomen brownish, faintly greyish pruinescent, with short fine dark setae; cercus long, slender, with dark setulae.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64599F2957EEFE56FE60ECFF.taxon	discussion	Remarks. This species may belong to the R. pusilla species group of Pararhamphomyia, but this requires verification. According to Frey, this species would run to his group III. Family HYBOTIDAE	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64599F2C57EEF989FE5EE9A5.taxon	description	(Figs 69 – 72) urn: lsid: zoobank. org: act: 67 F 5 F 184 - B 4 CB- 4 F 4 E- 895 D- 491 D 576 A 1 A 5 E	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64599F2C57EEF989FE5EE9A5.taxon	materials_examined	Type material. HOLOTYPE ♂, labelled: [RUSSIA. Chukotka AO (Wrangel Island)] “ bukh. [= bukhta, bay] Somnitelnaya, / south o. [= ostrov, island] Wrangel / Gorodkov 22. vii. [1966] ”; “ dol. r. [= dolina reki, valley of river] Somnitelnaya, / Mineev Mountains, 150 m, / shingle ”; “ on flower of Potentilla emarginata ”; “ Holotypus / Platypalpus / septentrionalis / Shamshev, Sinclair sp. nov. [red label] ” (INS _ DIP _ 0000619, ZIN, pinned on one pin with a female). PARATYPES. RUSSIA. Chukotka AO (Wrangel Island): Pinned on one pin with the holotype male (1 ♀, ZIN); same data as holotype (1 ♂, 4 ♀, ZIN; 1 ♂, 1 ♀, CNC); same data as holotype, 19. vii. 1966 (1 ♀, ZIN).	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64599F2C57EEF989FE5EE9A5.taxon	diagnosis	Diagnosis. Small robust blackish species of the P. longicornis group; 2 pairs of black vertical setae; postpedicenearly 2.5 X longer than wide and 2 X longer than stylus; legs extensively dark brown, mid tibia with very short apical spur; halteres brown.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64599F2C57EEF989FE5EE9A5.taxon	description	Description. Body holotype 1.9 mm, wing 2.0 mm. Male (Fig. 69). Head black. Occiput entirely densely greyish pollinose; with 2 pairs of long, black vertical setae (inner pair proclinate and slightly inclinate; outer pair lateroclinate); very short dark setae on upper part and longer pale setae on lower part. Ocellar tubercle concolorous with occiput, with 2 long, black lateroclinate anterior setae and 2 minute posterior setae. Frons broad on upper part, narrowed toward antennae, below ocellar tubercle broader than distance between outer margins of posterior ocelli; densely greyish brown pruinose (anterior view). Face rather broad, nearly as broad as frons above antennae, concolorous with frons; clypeus shiny. Antenna uniformly dark brown; postpedicel conical, moderately long, nearly 2.5 X as long as basal width; stylus short, nearly 2 X shorter than postpedicel. Gena very narrow, shiny. Proboscis dark brown, short. Palpus black, small, oval, pale pubescent, with 2 pale setae apically. Thorax black, almost entirely uniformly greyish tomentose; katepisternum almost entirely shiny, only narrowly tomentose along upper margin and anteriorly. Postpronotal lobe with 1 long black seta and some minute setulae. Mesonotum with main setae black to brown; 2 long npl (with additional pale setulae), 2 pal (1 much longer), 4 sctl (apical pair long, lateral pair short); acr very short, fine, arranged in 2 broad irregular rows; dc mostly arranged in 2 irregular rows and almost similar to acr anteriorly (some setae very slightly longer), 2 prescutellar pairs long. Legs extensively dark brown, subshiny; fore femur narrowly yellowish brown apically, mid and hind femora reddish brown on extreme apex; fore tibia mostly yellowish brown, somewhat paler closer to base, mid and hind tibiae reddish brown near extreme base. Coxae broadly shiny laterally, otherwise faintly greyish pruinose; fore coxa with pale setae anteriorly; hind trochanter with fine setulae. Fore femur strongly thickened; pilose ventrally; with anteroventral and posteroventral mostly minute fine setae becoming slightly longer toward base. Mid femur slightly stouter than fore femur; with double row of blackish short spine-like setae ventrally, without row of posteroventral setae. Hind femur slender, with minute anteroventral setae. Fore tibia slightly uniformly thickened; with simple setulae dorsally. Mid tibia slightly arcuate; with row of ventral dark spinule-like setulae; apical spur very short, shorter than apical width of tibia. Hind tibia slender, straight, with simple setulae. Tarsi of all legs unmodified. Wing faintly infuscate, with pale brownish veins. Basal costal seta long, blackish. Veins R 4 + 5 and M 1 + 2 parallel near wing margin; CuA slightly recurrent; anal vein very faint; crossveins m-cu and r-m somewhat contiguous. Calypter brown, pale fringed. Halter brown. Abdomen entirely dark brown to black, almost entirely shiny, only tergites 1 and 2 laterally faintly greyish pruinose (tergite 2 only narrowly anteriorly); covered with scattered pale setulae. Terminalia (Figs 70 – 72) small, subglobular, concolourous with abdomen, epandrium mostly shiny. Right cercus (Fig. 71) digitiform with somewhat pointed apex, short, with small internal projection slightly beyond middle; covered with short simple setae. Left cercus of similar shape to right cercus viewed dorsally, broadly ovate viewed laterally, nearly as long and as broad as right cercus (dorsal view); with small internal projection slightly beyond middle; covered with short simple setae. Hypoproct subtriangular (dorsal view), with somewhat pointed apex. Right epandrial lamella (Fig. 70) subglobular viewed laterally, with row of several short to moderately long subapical setae; right surstylus hardly differentiated from apex of epandrial lamella, short, rounded apically. Left epandrial lamella (Fig. 72) rather subtriangular but with small rounded projection about mid-dorsally; with scattered short setae along ventral margin and some setulae ventrally. Female. Similar to male. Visible parts of tergites 7 and 8 densely greyish pruinescent; cercus long, slender.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64599F2C57EEF989FE5EE9A5.taxon	distribution	Distribution. Palaearctic: Russia (Chukotka Autonomous Okrug, Wrangel Island).	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64599F2C57EEF989FE5EE9A5.taxon	etymology	Etymology. The specific name is from the Latin septentrionalis (north, northern), in reference to the distribution of this species.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64599F2C57EEF989FE5EE9A5.taxon	discussion	Remarks. This new species can be readily distinguished from all other species of Holarctic Platypalpus by brown halteres. In having small body size, two pairs of vertical setae, very short midtibial spur and lacking posteroventral setae on the mid femur, the new species resembles P. vegrandis Frey, 1943 known from Europe (Chvála 1989). However, P. vegrandis has partly yellow antennae and legs. In addition, hind trochanters in the male of this species bear three black spine-like setae. It is interesting to note that both species are found on flowers and may have similar hunting behaviour (Grootaert et al. 2012). Also, the new species could be compared with P. palmeni Frey, 1943 known from the Alps (Chvála 1989). Both species are similar in body size, have two pairs of vertical setae and lack posteroventral setae on the mid femur. However, P. palmeni can be readily distinguished from the new species by pale yellow palps and extensively yellow legs. In addition, the new species was compared with the syntypes (currently in CNC) of the following species known from subarctic areas of North America in order to confirm its status (especially since the Nearctic fauna is not as well diagnosed in comparison to the Palaearctic): Platypalpus arcticus Melander, 1928, P. diversipes Coquillett, 1900, P. gilvipes Coquillett, 1900, P. lateralis Loew, 1864, P. vittiger Melander, 1928 (described as var. of P. flavirostris Loew, 1864). All these species differ from the new species primarily by a combination of characters noted in the diagnosis.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C645C9F5357EEFEE2FE1BE8F8.taxon	description	(Figs 73 – 76) urn: lsid: zoobank. org: act: FC 7 E 4 A 07 - 9126 - 4941 - A 797 - 489575 CA 0 E 21	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C645C9F5357EEFEE2FE1BE8F8.taxon	materials_examined	Type material. HOLOTYPE ♂, labelled: [RUSSIA. Chukotka AO (Wrangel Island)] “ sr. tech. r. [= srednee techenie reki, = middle flow of river] Mamontovaya, N / Perkatkun, o. [= ostrov, island] Wrangel / willows in river valley / Gorodkov 17. vii. [1] 972 ”; “ Holotypus / Platypalpus / subtectifrons / Shamshev, Sinclair sp. n. [red label] ” (INS _ DIP _ 0000620, ZIN). PARATYPES. CANADA. Northwest Territories: Aklavik, 2. vii. 1956, R. E. Leech (1 ♀, CNC); same data except, 25. vi. – 3. vii. 1956, E. F. Cashman (3 ♀, CNC); Fort McPherson, 17 – 19. vii. 1957, R. Hurley (2 ♀, CNC); 21 mi E Tuktoyaktuk, 8 – 12. vii. 1972, D. M. Wood (2 ♂, 1 ♀, CNC); Tuktoyaktuk, 69 ° 26 ′ 16.2 ″ N 133 ° 1 ′ 2.16 ″ W, pan traps, meadow, 15 – 18. vii. 2010, Goulet & Boudreault (12 ♂, 4 ♀, CNC). Yukon: British Mtns, Firth River, 25. vii. 1956, E. F. Cashman (2 ♀, CNC); Firth River, 14. vii. – 7. viii. 1956, E. F. Cashman (5 ♂, 7 ♀, CNC); same data except, 20. vii. – 3. viii. 1965, R. E. Leech (5 ♂, 8 ♀, CNC). RUSSIA. Chukotka AO (Wrangel Island): Same data as holotype (4 ♂, 1 ♀, ZIN; 2 ♂, 1 ♀, CNC); 21. vii. 1972, KBG (1 ♂, 4 ♀, ZIN); middle flow of river Neizvestnaya, 71 ° 20.287 ′ N 179 ° 29.779 ′ W, BT 1, 5 – 14. vii. 2018, U. V. Babiy (3 ♂, 3 ♀, ZIN); middle flow of river Mamontovaya, 71 ° 10 ′ N 179 ° 45 ′ W, BT 2, 30. vi. – 9. vii. 2006, OAK (1 ♂, ZIN); same locality and biotope, BT 2, YPT, 2 – 4. vii. 2015, OAK (2 ♂, ZIN); same locality, 71 ° 10 ′ N 179 ° 45 ′ W, BT 1 A, YPT, 2 – 4. vii. 2015, OAK (3 ♂, 3 ♀, ZIN); same locality, 71 ° 10 ′ N 179 ° 45 ′ W, BT 14, Sw, 2. vii. 2015, OAK (2 ♂, 1 ♀, ZIN); same locality, 71 ° 10 ′ N 179 ° 45 ′ W, 3. vii. 2015, OAK (1 ♂, ZIN); Tundrovaya Mnt, 71 ° 18 ′ N 179 ° 49 ′ W, BT 2, 1 – 19. vii. 2015, LFV (1 ♂, ZIN; dissected). USA. Alaska: Cape Thompson, 23 – 25. vii. 1961, R. Madge (2 ♀, CNC); Isabel Pass, mi 206 Richardson Hwy, 2900 ft, 18. vii. 1962, P. J. Skitsko (2 ♀, CNC); Nome Area, mi 46 Kougarok Rd, 64 ° 57 ′ 27 ″ N 168 ° 48 ′ 21 ″ W, 26 – 28. vi. 2005, MT, J. & R. Skevington (1 ♂, CNC); Prudhoe Bay, 9. vii. 1971, M. Deyrup (1 ♂, 6 ♀, CNC); Umalakleet, 18, 28. vi. 1961, B. S. Heming (1 ♂, 1 ♀, CNC); same data except, 28. vi., 8. vii. 1961, R. Madge (1 ♂, 1 ♀, CNC); Umiat, 3 – 25. vii. 1959, R. Madge (3 ♂, 38 ♀, CNC); same data except, 6 – 15. vii. 1959, J. E. H. Martin (26 ♂, 28 ♀, CNC); same data except, 4. viii. 1959, B. N. Smallman (1 ♂, CNC).	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C645C9F5357EEFEE2FE1BE8F8.taxon	diagnosis	Diagnosis. Small, blackish species of the P. pallidiventris group; one pair of vertical setae, postpedicel nearly 1.5 X longer than wide, palpus black, thorax with pale setae, scutum entirely pruinose, acr biserial, legs almost entirely dark brown, hind tibia with long apical spur, abdomen almost entirely shiny.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C645C9F5357EEFEE2FE1BE8F8.taxon	description	Description. Body length 2.4 – 2.5 (holotype 2.5), wing length 3.0 – 3.2 (holotype 3.1) mm. Male (Fig. 73). Head black, mostly pale setose, ocellar setae rather yellowish. Occiput entirely densely greyish pruinose; with one pair of moderately long, fine, wide apart vertical setae; numerous short setae on upper part and longer setae on lower part. Ocellar tubercle concolorous with occiput, with 2 long, lateroclinate anterior setae and 2 minute posterior setae. Frons broad, only slightly narrowed toward antennae, below ocellar tubercle broader than distance between outer margins of posterior ocelli; densely light grey pruinose (anterior view). Face broad, almost as broad as frons above antennae, densely whitish pruinose (including clypeus). Antenna uniformly dark brown; postpedicel short, nearly 1.5 X (in holotype 1.65 X) as long as basal width; stylus nearly 2 X (in holotype 2.2 X) longer than postpedicel. Gena narrow, shiny. Proboscis dark brown, elongate, nearly 1.5 X shorter of head height. Palpus black, small, oval, whitish pubescent, with few short setae. Thorax black, mostly pale setose, strong setae rather pale yellowish; prothoracic sclerites and mesopleuron densely greyish tomentose, scutum rather yellowish grey faintly tomentose; katepisternum mostly shiny, only tomentose on upper anterior corner and along posterior corner. Postpronotal lobe with 1 moderately long fine seta and some minute setulae. Mesonotum with 2 moderately long npl (with numerous additional setulae), 2 pal (1 longer), 4 sctl (apical pair long, lateral pair finer and shorter; sometimes additional setae present); some setulae behind postpronotal lobe and on supra-alar space; acr short, fine, arranged in 2 moderately broad irregular rows, on prescutellar depression hardly distinguishable from dc and sometimes more numerous; dc arranged in 2 – 3 irregular rows and mostly similar to acr, some prescutellars longer. Legs extensively dark brown, subshiny, pale setose (except noted), somewhat variable in yellowish pattern. In holotype and most paratypes fore femur very narrowly yellowish apically, mid and hind femora yellowish brown on extreme apex, fore tibia yellowish near base (becoming brownish toward apex); mid and hind tibiae slightly paler than corresponding femora, tarsomeres somewhat paler on basal part (broadly on basitarsus, extreme base on remaining tarsomeres); in addition, in some paratypes fore coxa on extreme apex and all femora on extreme base yellowish brown (1 ♂ and 5 ♀ from Perkatkun). Fore coxae broadly shiny laterally, otherwise densely greyish tomentose; mid and hind coxae almost entirely shiny; all coxae pale setose; hind trochanter with fine setulae. Fore femur strongly thickened; pilose ventrally; with rows of long, fine anteroventral and posteroventral setae (longest setae on basal part slightly longer than half of femur width). Mid femur slightly stouter than fore femur; with double row of short, blackish spine-like setae ventrally, row of long strong posteroventrals intermixed with shorter and finer setae. Hind femur slender, with somewhat longer anteroventral setae on about apical half. Fore tibia slightly uniformly thickened; with simple setulae dorsally, some anterodorsals slightly stronger; longer setae ventrally and posteroventrally (longest ones on subapical part slightly longer than tibia width). Mid tibia with row of ventral dark spinule-like setulae; apical spur long, pointed, with two setulae at tip. Hind tibia slender, straight, with simple setulae. Tarsi of all legs unmodified; fore basitarsus with longer setae ventrally and posteroventrally (similar to those on fore tibia). Wing faintly infuscate, with brownish veins. Basal costal seta moderately long, fine, pale. Veins R 4 + 5 and M 1 + 2 slightly convergent near wing margin; CuA straight, perpendicular, faint apically; anal vein very faint; crossveins m-cu and r-m slightly separated. Calypter dirty yellow, pale fringed. Halter yellow. Abdomen entirely blackish brown, almost entirely shiny, only tergite 1 faintly greyish pruinose laterally; covered with numerous pale hair-like setae longer on tergites laterally. Terminalia (Figs 74 – 76) small, concolourous with abdomen, epandrium shiny. Right cercus (Fig. 75) digitiform, long, almost parallel-sided toward apex, truncate; covered with short simple setae. Left cercus digitiform, slightly longer and narrower (on apical half) than right cercus (dorsal view), almost parallel-sided toward apex, rounded on apex; covered with short simple setae. Hypoproct subtriangular, narrow (dorsal view), with pointed apex almost reaching apex of cerci. Right epandrial lamella (Fig. 74) subtriangular viewed laterally, with row of several long subapical setae; right surstylus well differentiated from apex of epandrial lamella, long, slender, with some minute marginal setulae. Left epandrial lamella subtriangular (Fig. 76) somewhat concave about mid-ventrally; with numerous long pale yellow setae along about apical 2 / 3 of ventral margin. Female. Similar to male. Visible parts of tergites 7 and 8 densely greyish pruinescent (except narrow upper margin of sternite 8); cercus long, slender.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C645C9F5357EEFEE2FE1BE8F8.taxon	distribution	Distribution. Palaearctic: Russia (Chukotka Autonomous Okrug, Wrangel Island). In North America, this species is known from Alaska, Northwest Territories and Yukon.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C645C9F5357EEFEE2FE1BE8F8.taxon	etymology	Etymology. The specific name refers to the similarity of the new species to P. tectifrons (Becker).	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C645C9F5357EEFEE2FE1BE8F8.taxon	discussion	Remarks. This new species strikingly resembles the poorly known P. tectifrons (Becker, 1907) described from eastern Tibet of China (syntypes housed in ZIN), which has the same pattern of leg colour and other main characters. The redescription of P. tectifrons will be published in a forthcoming paper. Platypalpus tectifrons differs from the new species by the somewhat narrower, longer postpedicel (about 2.5 X longer than wide) and brownish to brownish yellow main mesonotal setae. Within the key to Platypalpus compiled by Chvála (1989), the new species would run to P. interstinctus (Collin, 1926) and P. pseudofulvipes (Frey, 1909) (P. pallidiventris group). Platypalpus interstinctus (somewhat variable in leg colour) differs from the new species by distinctly black annulated tarsi, brownish vertical, acrostichal and dorsocentral setae, as well as by details of the male terminalia (Chvála 1975: 178, figs 441 – 443). Platypalpus pseudofulvipes can be readily distinguished from the new species primarily by the spindle-shaped dilated fore tibiae. In addition, the new species could be compared with P. melancholicus (Collin, 1961), P. annulatus (Fallén, 1815) and P. carteri (Collin, 1926). Platypalpus melancholicus and P. annulatus differ from the new species primarily by 4 - serial acrostichals (at least on anterior part of mesoscutum). Platypalpus carteri can be distinguished from the new species by lateral pruinosity on all abdominal tergites. The male terminalia of the new species somewhat resemble those of P. notatus (Meigen, 1822) (especially cerci), however, P. notatus has longer postpedicel (about 2.5 X longer than wide) and entirely shiny abdomen (including lateral area of first two tergites). In addition, the new species was compared with the syntypes (currently in CNC) of the species known from subarctic areas of North America listed under above. All these species differ from the new species primarily by a combination of characters noted in the diagnosis. Habitat. On Wrangel Island, this species was collected along riverbeds. Genus Tachypeza Meigen	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64239F5357EEFE72FBAFEF46.taxon	description	(Figs 77 – 80)	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64239F5357EEFE72FBAFEF46.taxon	description	River (~ 71 ° 44 ′ N 128 ° 16 ′ E), Yakutia, Russia.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64239F5357EEFE72FBAFEF46.taxon	materials_examined	Material examined. CANADA. Northwest Territories: Ford Lake, 63 ° 11 ′ N 107 ° 19 ′ W, 23. vi. 1966, G. E. Shewell (1 ♂, 1 ♀, CNC). Nunavut: Whatever Lake, 62 ° 41 ′ N 97 ° 03 ′ W, vii. 1989, pitfall trap, J. Pilny (3 ♂, 1 ♀, CNC). RUSSIA. Chukotka AO (Wrangel Island): N Somnitelnaya Bay, 27. vii. 1966, KBG (1 ♀, ZIN); northern slope of Berry Peak, 7 km ESE Sovetskaya Mtn, 350 m, on stones, 12. vii. 1972, KBG (5 ♂, 6 ♀, ZIN). Recognition. Small blackish flies (body 2.5 – 3 mm) (Fig. 72); palpus pale to pale yellow, whitish pubescent, with short pale hair-like setae (sometimes 1 subapical seta brownish); prothoracic sclerites (except postpronotal lobe) densely silvery grey tomentose, mesopleuron almost entirely shiny, 1 strong npl, 2 broadly set similar sctl, 1 (sometimes 2) strong prescutellar dc; legs almost uniformly brown; halter with yellow knob. Male: mid femur with row of 6 – 7 closely set black short anteroventral spine-like setae on about basal 1 / 3 and 2 closely set, moderately long, spine-like setae near extreme base; mid tibia with shallow, ventral subapical excision. Male terminalia (Figs 78 – 80) large, brown, epandrium shiny. Cerci (Fig. 79) separated, shifted to inner margin of right epandrial lamella; right cercus digitiform, short, with apex slightly extended beyond apex of right epandrial lamella, covered with short setae stronger on inner margin; left cercus short, broad, subrectangular; with some short setae. Hypoproct weakly sclerotized, rather subtriangular, separated from both cerci, bare. Right epandrial lamella (Fig. 78) without ventral lobe, produced apicoventrally; with cluster of several spine-like short marginal setae apically; right surstylus separated from epandrial lamella, narrow, with two projections, outer projection very short, inner projection long, stylet-like; right bacilliform sclerite with 2 closely set spine-like setae. Left epandrial lamella narrow; with large apodeme; bearing some setulae apically; left surstylus (Fig. 80) large, rather subtriangular (lateral view) but with additional narrow pointed projection dorsally; with short marginal setae apically. Hypandrium hump-backed.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64239F5357EEFE72FBAFEF46.taxon	distribution	Distribution. Holarctic; T. sericeipalpis has a disjunct distribution occurring in Eurasia across the arctic region of Russia from Murmanskaya Province to Chukotka (including Wrangel Island) and is found locally in mountains of Altay Republic and Buryatia (Shamshev, unpubl. data). In North America, this species is newly recorded from the Northwest Territories and Nunavut of Canada.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
7B7E785C64239F5357EEFE72FBAFEF46.taxon	discussion	Remarks. The male terminalia of this species are here illustrated for the first time.	en	Shamshev, Igor V., Sinclair, Bradley J., Khruleva, Olga A. (2020): The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago. Zootaxa 4848 (1): 1-75, DOI: 10.11646/zootaxa.4848.1.1
