taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
7B3FB814D321473F93BFFE10FDA5FCC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11450110/files/figure.png	https://doi.org/10.5281/zenodo.11450110	Figure 1. Onthophagus (Indonthophagus) mopsus (Fabricius, Figure 2. Onthophagus (Indonthophagus) turbatus Walker, 1792) [Pakistan, Islamabad Capital Territory]. 1a) ♀ head, 1858 [Sri Lanka, Uva prov.]. Detail of pronotal posterolateral from above. 1b) Epipharynx. Photos by A. Degiovanni, ed- area. Photos by A. Degiovanni, edited by G. Fiumi.	Figure 1. Onthophagus (Indonthophagus) mopsus (Fabricius, Figure 2. Onthophagus (Indonthophagus) turbatus Walker, 1792) [Pakistan, Islamabad Capital Territory]. 1a) ♀ head, 1858 [Sri Lanka, Uva prov.]. Detail of pronotal posterolateral from above. 1b) Epipharynx. Photos by A. Degiovanni, ed- area. Photos by A. Degiovanni, edited by G. Fiumi.	2024-04-05	Ziani, Stefano		Zenodo	biologists	Ziani, Stefano			
7B3FB814D323473193BFFC42FB06FB4F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11450112/files/figure.png	https://doi.org/10.5281/zenodo.11450112	Figure 3. Onthophagus (Indonthophagus) aeneopiceus d’Orbigny, 1902 [major ♂: Yemen, Ibb gov.]. 3a) Dorsal habitus. 3b) Parameres, lateral view. 3c) Parameres, dorsal view. 3d) Endophallus. AcS: accessory endophallites. LC: lamella copulatrix. Rs: raspulae. Photos by A. Degiovanni, edited by G. Fiumi [from Ziani and Keith (2023)].	Figure 3. Onthophagus (Indonthophagus) aeneopiceus d’Orbigny, 1902 [major ♂: Yemen, Ibb gov.]. 3a) Dorsal habitus. 3b) Parameres, lateral view. 3c) Parameres, dorsal view. 3d) Endophallus. AcS: accessory endophallites. LC: lamella copulatrix. Rs: raspulae. Photos by A. Degiovanni, edited by G. Fiumi [from Ziani and Keith (2023)].	2024-04-05	Ziani, Stefano		Zenodo	biologists	Ziani, Stefano			
7B3FB814D32D473293BFFACAFC34FC6B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11450114/files/figure.png	https://doi.org/10.5281/zenodo.11450114	Figure 4. Onthophagus (Indonthophagus) ensifer Boucomont, 1914 [major ♂: India, Tamil Nadu st.]. 4a) Dorsal habitus. 4b) Parameres, lateral view. 4c) Parameres, dorsal view. 4d) Endophallus. AcS: accessory endophallites. LC: lamella copulatrix. Rs: raspula. Photos by A. Degiovanni, edited by G. Fiumi.	Figure 4. Onthophagus (Indonthophagus) ensifer Boucomont, 1914 [major ♂: India, Tamil Nadu st.]. 4a) Dorsal habitus. 4b) Parameres, lateral view. 4c) Parameres, dorsal view. 4d) Endophallus. AcS: accessory endophallites. LC: lamella copulatrix. Rs: raspula. Photos by A. Degiovanni, edited by G. Fiumi.	2024-04-05	Ziani, Stefano		Zenodo	biologists	Ziani, Stefano			
7B3FB814D32E473793BFFC17FAEFF92B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11450116/files/figure.png	https://doi.org/10.5281/zenodo.11450116	Figure 5. Onthophagus (Indonthophagus) hastifer von Lansberge, 1914 [holotype, major ♂: Myanmar, Minhla]. 5a) Dorsal habitus.5b) Parameres, lateral view. 5c) Parameres, dorsal view. 5d) Endophallus. AcS: accessory endophallites. LC: lamella copulatrix. Rs: raspulae. Photos by A. Degiovanni, edited by G. Fiumi.	Figure 5. Onthophagus (Indonthophagus) hastifer von Lansberge, 1914 [holotype, major ♂: Myanmar, Minhla]. 5a) Dorsal habitus.5b) Parameres, lateral view. 5c) Parameres, dorsal view. 5d) Endophallus. AcS: accessory endophallites. LC: lamella copulatrix. Rs: raspulae. Photos by A. Degiovanni, edited by G. Fiumi.	2024-04-05	Ziani, Stefano		Zenodo	biologists	Ziani, Stefano			
7B3FB814D32E473793BFFC17FAEFF92B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11450134/files/figure.png	https://doi.org/10.5281/zenodo.11450134	Figure 12. Head from above:12a) Onthophagus (Indonthophagus) hastifer von Lansberge, 1914 [major ♂: Taiwan, Pingtung]. 12b) Onthophagus (Indonthophagus) turbatus Walker, 1858 [major ♂: India, Arunachal Pradesh st.]. 12c) Onthophagus (Indonthophagus) hastifer von Lansberge, 1914 [♀: Taiwan, Pingtung]. 12d) Onthophagus (Indonthophagus) turbatus Walker, 1858 [♀: Pakistan, Islamabad Capital Territory]. Photos by A. Degiovanni, edited by G. Fiumi.	Figure 12. Head from above:12a) Onthophagus (Indonthophagus) hastifer von Lansberge, 1914 [major ♂: Taiwan, Pingtung]. 12b) Onthophagus (Indonthophagus) turbatus Walker, 1858 [major ♂: India, Arunachal Pradesh st.]. 12c) Onthophagus (Indonthophagus) hastifer von Lansberge, 1914 [♀: Taiwan, Pingtung]. 12d) Onthophagus (Indonthophagus) turbatus Walker, 1858 [♀: Pakistan, Islamabad Capital Territory]. Photos by A. Degiovanni, edited by G. Fiumi.	2024-04-05	Ziani, Stefano		Zenodo	biologists	Ziani, Stefano			
7B3FB814D32B472B93BFF950FA05F99E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11450110/files/figure.png	https://doi.org/10.5281/zenodo.11450110	Figure 1. Onthophagus (Indonthophagus) mopsus (Fabricius, Figure 2. Onthophagus (Indonthophagus) turbatus Walker, 1792) [Pakistan, Islamabad Capital Territory]. 1a) ♀ head, 1858 [Sri Lanka, Uva prov.]. Detail of pronotal posterolateral from above. 1b) Epipharynx. Photos by A. Degiovanni, ed- area. Photos by A. Degiovanni, edited by G. Fiumi.	Figure 1. Onthophagus (Indonthophagus) mopsus (Fabricius, Figure 2. Onthophagus (Indonthophagus) turbatus Walker, 1792) [Pakistan, Islamabad Capital Territory]. 1a) ♀ head, 1858 [Sri Lanka, Uva prov.]. Detail of pronotal posterolateral from above. 1b) Epipharynx. Photos by A. Degiovanni, ed- area. Photos by A. Degiovanni, edited by G. Fiumi.	2024-04-05	Ziani, Stefano		Zenodo	biologists	Ziani, Stefano			
7B3FB814D32B472B93BFF950FA05F99E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11450118/files/figure.png	https://doi.org/10.5281/zenodo.11450118	Figure 6. Onthophagus (Indonthophagus) mopsus (Fabricius, 1792) [major ♂: Pakistan, Islamabad Capital Territory]. 6a) Dorsal habitus; 6b) Parameres, lateral view. 6c) Parameres, dorsal view. 6d) Endophallus. AcS: accessory endophallites. LC: lamella copulatrix. Rs: raspulae. Photos by A. Degiovanni, edited by G. Fiumi.	Figure 6. Onthophagus (Indonthophagus) mopsus (Fabricius, 1792) [major ♂: Pakistan, Islamabad Capital Territory]. 6a) Dorsal habitus; 6b) Parameres, lateral view. 6c) Parameres, dorsal view. 6d) Endophallus. AcS: accessory endophallites. LC: lamella copulatrix. Rs: raspulae. Photos by A. Degiovanni, edited by G. Fiumi.	2024-04-05	Ziani, Stefano		Zenodo	biologists	Ziani, Stefano			
7B3FB814D32B472B93BFF950FA05F99E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11450120/files/figure.png	https://doi.org/10.5281/zenodo.11450120	Figure 7. Onthophagus (Indonthophagus) mopsus (Fabricius, 1792), lectotype labels. Photo by Katerine Child (OUMNH).	Figure 7. Onthophagus (Indonthophagus) mopsus (Fabricius, 1792), lectotype labels. Photo by Katerine Child (OUMNH).	2024-04-05	Ziani, Stefano		Zenodo	biologists	Ziani, Stefano			
7B3FB814D336472F93BFFF0EFD57FCFB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11450122/files/figure.png	https://doi.org/10.5281/zenodo.11450122	Figure 8. Onthophagus (Indonthophagus) nitidulus Klug, 1845 [major ♂: Yemen, Abyan gov.]. 8a) Dorsal habitus. 8b) Parameres, lateral view. 8c) Parameres, dorsal view. 8d) Endophallus. AcS: accessory endophallites. LC: lamella copulatrix. Rs: raspulae. Photos by A. Degiovanni, edited by G. Fiumi.	Figure 8. Onthophagus (Indonthophagus) nitidulus Klug, 1845 [major ♂: Yemen, Abyan gov.]. 8a) Dorsal habitus. 8b) Parameres, lateral view. 8c) Parameres, dorsal view. 8d) Endophallus. AcS: accessory endophallites. LC: lamella copulatrix. Rs: raspulae. Photos by A. Degiovanni, edited by G. Fiumi.	2024-04-05	Ziani, Stefano		Zenodo	biologists	Ziani, Stefano			
7B3FB814D333472393BFFC80FC7EFB6E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11450134/files/figure.png	https://doi.org/10.5281/zenodo.11450134	Figure 12. Head from above:12a) Onthophagus (Indonthophagus) hastifer von Lansberge, 1914 [major ♂: Taiwan, Pingtung]. 12b) Onthophagus (Indonthophagus) turbatus Walker, 1858 [major ♂: India, Arunachal Pradesh st.]. 12c) Onthophagus (Indonthophagus) hastifer von Lansberge, 1914 [♀: Taiwan, Pingtung]. 12d) Onthophagus (Indonthophagus) turbatus Walker, 1858 [♀: Pakistan, Islamabad Capital Territory]. Photos by A. Degiovanni, edited by G. Fiumi.	Figure 12. Head from above:12a) Onthophagus (Indonthophagus) hastifer von Lansberge, 1914 [major ♂: Taiwan, Pingtung]. 12b) Onthophagus (Indonthophagus) turbatus Walker, 1858 [major ♂: India, Arunachal Pradesh st.]. 12c) Onthophagus (Indonthophagus) hastifer von Lansberge, 1914 [♀: Taiwan, Pingtung]. 12d) Onthophagus (Indonthophagus) turbatus Walker, 1858 [♀: Pakistan, Islamabad Capital Territory]. Photos by A. Degiovanni, edited by G. Fiumi.	2024-04-05	Ziani, Stefano		Zenodo	biologists	Ziani, Stefano			
7B3FB814D333472393BFFC80FC7EFB6E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11450128/files/figure.png	https://doi.org/10.5281/zenodo.11450128	Figure 10. Onthophagus (Indonthophagus) turbatus Walker, 1858 [major ♂: Pakistan, Mansehra prov.]. Dorsal habitus. Photos by A. Degiovanni, edited by G. Fiumi.	Figure 10. Onthophagus (Indonthophagus) turbatus Walker, 1858 [major ♂: Pakistan, Mansehra prov.]. Dorsal habitus. Photos by A. Degiovanni, edited by G. Fiumi.	2024-04-05	Ziani, Stefano		Zenodo	biologists	Ziani, Stefano			
7B3FB814D333472393BFFC80FC7EFB6E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11450132/files/figure.png	https://doi.org/10.5281/zenodo.11450132	Figure 11. Onthophagus (incertae sedis) spinifex (Fabricius, 1781) [major ♂: Pakistan, Capital Territory].11a) Dorsal habitus. 11b) Parameres, lateral view.11c) Parameres, dorsal view.11d) Endophallus. AcS: accessory endophallites. LC: lamella copulatrix. Photos by A. Degiovanni, edited by G. Fiumi.	Figure 11. Onthophagus (incertae sedis) spinifex (Fabricius, 1781) [major ♂: Pakistan, Capital Territory].11a) Dorsal habitus. 11b) Parameres, lateral view.11c) Parameres, dorsal view.11d) Endophallus. AcS: accessory endophallites. LC: lamella copulatrix. Photos by A. Degiovanni, edited by G. Fiumi.	2024-04-05	Ziani, Stefano		Zenodo	biologists	Ziani, Stefano			
7B3FB814D33F472693BFFB14FC30FD40.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11450132/files/figure.png	https://doi.org/10.5281/zenodo.11450132	Figure 11. Onthophagus (incertae sedis) spinifex (Fabricius, 1781) [major ♂: Pakistan, Capital Territory].11a) Dorsal habitus. 11b) Parameres, lateral view.11c) Parameres, dorsal view.11d) Endophallus. AcS: accessory endophallites. LC: lamella copulatrix. Photos by A. Degiovanni, edited by G. Fiumi.	Figure 11. Onthophagus (incertae sedis) spinifex (Fabricius, 1781) [major ♂: Pakistan, Capital Territory].11a) Dorsal habitus. 11b) Parameres, lateral view.11c) Parameres, dorsal view.11d) Endophallus. AcS: accessory endophallites. LC: lamella copulatrix. Photos by A. Degiovanni, edited by G. Fiumi.	2024-04-05	Ziani, Stefano		Zenodo	biologists	Ziani, Stefano			
7B3FB814D33F472693BFFB14FC30FD40.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11450124/files/figure.png	https://doi.org/10.5281/zenodo.11450124	Figure 9. Onthophagus (Indonthophagus) turbatus Walker, 1858 [syntype, medium ♂: Sri Lanka]. 9a) Male dorsal habitus [photo and editing by G. Fiumi]. 9b) Parameres, lateral view. 9c) Parameres, dorsal view. 9d) Endophallus. AcS: accessory endophallites. LC: lamella copulatrix. Rs: raspulae. Photos by A. Degiovanni, edited by G. Fiumi.	Figure 9. Onthophagus (Indonthophagus) turbatus Walker, 1858 [syntype, medium ♂: Sri Lanka]. 9a) Male dorsal habitus [photo and editing by G. Fiumi]. 9b) Parameres, lateral view. 9c) Parameres, dorsal view. 9d) Endophallus. AcS: accessory endophallites. LC: lamella copulatrix. Rs: raspulae. Photos by A. Degiovanni, edited by G. Fiumi.	2024-04-05	Ziani, Stefano		Zenodo	biologists	Ziani, Stefano			
7B3FB814D305471893BFFA01FA04FB6F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11450132/files/figure.png	https://doi.org/10.5281/zenodo.11450132	Figure 11. Onthophagus (incertae sedis) spinifex (Fabricius, 1781) [major ♂: Pakistan, Capital Territory].11a) Dorsal habitus. 11b) Parameres, lateral view.11c) Parameres, dorsal view.11d) Endophallus. AcS: accessory endophallites. LC: lamella copulatrix. Photos by A. Degiovanni, edited by G. Fiumi.	Figure 11. Onthophagus (incertae sedis) spinifex (Fabricius, 1781) [major ♂: Pakistan, Capital Territory].11a) Dorsal habitus. 11b) Parameres, lateral view.11c) Parameres, dorsal view.11d) Endophallus. AcS: accessory endophallites. LC: lamella copulatrix. Photos by A. Degiovanni, edited by G. Fiumi.	2024-04-05	Ziani, Stefano		Zenodo	biologists	Ziani, Stefano			
7B3FB814D305471893BFFA01FA04FB6F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11450122/files/figure.png	https://doi.org/10.5281/zenodo.11450122	Figure 8. Onthophagus (Indonthophagus) nitidulus Klug, 1845 [major ♂: Yemen, Abyan gov.]. 8a) Dorsal habitus. 8b) Parameres, lateral view. 8c) Parameres, dorsal view. 8d) Endophallus. AcS: accessory endophallites. LC: lamella copulatrix. Rs: raspulae. Photos by A. Degiovanni, edited by G. Fiumi.	Figure 8. Onthophagus (Indonthophagus) nitidulus Klug, 1845 [major ♂: Yemen, Abyan gov.]. 8a) Dorsal habitus. 8b) Parameres, lateral view. 8c) Parameres, dorsal view. 8d) Endophallus. AcS: accessory endophallites. LC: lamella copulatrix. Rs: raspulae. Photos by A. Degiovanni, edited by G. Fiumi.	2024-04-05	Ziani, Stefano		Zenodo	biologists	Ziani, Stefano			
7B3FB814D305471893BFFA01FA04FB6F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11450114/files/figure.png	https://doi.org/10.5281/zenodo.11450114	Figure 4. Onthophagus (Indonthophagus) ensifer Boucomont, 1914 [major ♂: India, Tamil Nadu st.]. 4a) Dorsal habitus. 4b) Parameres, lateral view. 4c) Parameres, dorsal view. 4d) Endophallus. AcS: accessory endophallites. LC: lamella copulatrix. Rs: raspula. Photos by A. Degiovanni, edited by G. Fiumi.	Figure 4. Onthophagus (Indonthophagus) ensifer Boucomont, 1914 [major ♂: India, Tamil Nadu st.]. 4a) Dorsal habitus. 4b) Parameres, lateral view. 4c) Parameres, dorsal view. 4d) Endophallus. AcS: accessory endophallites. LC: lamella copulatrix. Rs: raspula. Photos by A. Degiovanni, edited by G. Fiumi.	2024-04-05	Ziani, Stefano		Zenodo	biologists	Ziani, Stefano			
7B3FB814D305471893BFFA01FA04FB6F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11450118/files/figure.png	https://doi.org/10.5281/zenodo.11450118	Figure 6. Onthophagus (Indonthophagus) mopsus (Fabricius, 1792) [major ♂: Pakistan, Islamabad Capital Territory]. 6a) Dorsal habitus; 6b) Parameres, lateral view. 6c) Parameres, dorsal view. 6d) Endophallus. AcS: accessory endophallites. LC: lamella copulatrix. Rs: raspulae. Photos by A. Degiovanni, edited by G. Fiumi.	Figure 6. Onthophagus (Indonthophagus) mopsus (Fabricius, 1792) [major ♂: Pakistan, Islamabad Capital Territory]. 6a) Dorsal habitus; 6b) Parameres, lateral view. 6c) Parameres, dorsal view. 6d) Endophallus. AcS: accessory endophallites. LC: lamella copulatrix. Rs: raspulae. Photos by A. Degiovanni, edited by G. Fiumi.	2024-04-05	Ziani, Stefano		Zenodo	biologists	Ziani, Stefano			
7B3FB814D305471893BFFA01FA04FB6F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11450112/files/figure.png	https://doi.org/10.5281/zenodo.11450112	Figure 3. Onthophagus (Indonthophagus) aeneopiceus d’Orbigny, 1902 [major ♂: Yemen, Ibb gov.]. 3a) Dorsal habitus. 3b) Parameres, lateral view. 3c) Parameres, dorsal view. 3d) Endophallus. AcS: accessory endophallites. LC: lamella copulatrix. Rs: raspulae. Photos by A. Degiovanni, edited by G. Fiumi [from Ziani and Keith (2023)].	Figure 3. Onthophagus (Indonthophagus) aeneopiceus d’Orbigny, 1902 [major ♂: Yemen, Ibb gov.]. 3a) Dorsal habitus. 3b) Parameres, lateral view. 3c) Parameres, dorsal view. 3d) Endophallus. AcS: accessory endophallites. LC: lamella copulatrix. Rs: raspulae. Photos by A. Degiovanni, edited by G. Fiumi [from Ziani and Keith (2023)].	2024-04-05	Ziani, Stefano		Zenodo	biologists	Ziani, Stefano			
7B3FB814D305471893BFFA01FA04FB6F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11450134/files/figure.png	https://doi.org/10.5281/zenodo.11450134	Figure 12. Head from above:12a) Onthophagus (Indonthophagus) hastifer von Lansberge, 1914 [major ♂: Taiwan, Pingtung]. 12b) Onthophagus (Indonthophagus) turbatus Walker, 1858 [major ♂: India, Arunachal Pradesh st.]. 12c) Onthophagus (Indonthophagus) hastifer von Lansberge, 1914 [♀: Taiwan, Pingtung]. 12d) Onthophagus (Indonthophagus) turbatus Walker, 1858 [♀: Pakistan, Islamabad Capital Territory]. Photos by A. Degiovanni, edited by G. Fiumi.	Figure 12. Head from above:12a) Onthophagus (Indonthophagus) hastifer von Lansberge, 1914 [major ♂: Taiwan, Pingtung]. 12b) Onthophagus (Indonthophagus) turbatus Walker, 1858 [major ♂: India, Arunachal Pradesh st.]. 12c) Onthophagus (Indonthophagus) hastifer von Lansberge, 1914 [♀: Taiwan, Pingtung]. 12d) Onthophagus (Indonthophagus) turbatus Walker, 1858 [♀: Pakistan, Islamabad Capital Territory]. Photos by A. Degiovanni, edited by G. Fiumi.	2024-04-05	Ziani, Stefano		Zenodo	biologists	Ziani, Stefano			
