taxonID	type	description	language	source
7A2687556C79AF3AFDF4FEFFFE6DC81A.taxon	description	(Figs. 1 – 13)	en	Schawaller, Wolfgang, Purchart, Luboš (2012): Nanocaecus hlavaci gen. & sp. nov. - first record of the tribe Gnathidiini (Coleoptera: Tenebrionidae: Diaperinae) from the Socotra Archipelago. Acta Entomologica Musei Nationalis Pragae 52: 303-314, DOI: 10.5281/zenodo.5354846
7A2687556C79AF3AFDF4FEFFFE6DC81A.taxon	diagnosis	Diagnosis. Within the subtribe Anopidiina, Nanocaecus gen. nov. shares with Anopidium and Paranopidium the antenna with 10 antennomeres including a terminal club of 4 antennomeres, and the small but visible scutellum. For other genera see the key below. Anopidium can be separated by the antennomere 3 distinctly prolonged, about two times longer than antennomere 2 (contrary to antennomere 3 equal in length to antennomere 2 in Nanocaecus gen. nov. and Paranopidium); and by the round shape of pronotum and elytra combined with the pronotum widest at base (contrary to long parallel shape with the pronotum widest in the middle). In Paranopidium the last four antennomeres form a fused club with irregular overall setation (contrary to a compact club with terminal setation of each joint in Nanocaecus gen. nov.); the clypeal suture is present (absent); pronotum and elytra without any setation (with microsetae); pronotum with sinuated anterior margin and prominent anterior corners (not sinuated with rounded anterior corners); elytra with 8 irregular rows of punctures and with short scutellar striolae (with irregular punctation and without scutellar striolae); base of elytra bordered laterally (completely unbordered); meso-and metaventrite with feeble, nearly invisible punctation (with striking large but not confluent punctation); and first abdominal ventrite with finger-like apophysis between hind coxae (with broad rounded apophysis).	en	Schawaller, Wolfgang, Purchart, Luboš (2012): Nanocaecus hlavaci gen. & sp. nov. - first record of the tribe Gnathidiini (Coleoptera: Tenebrionidae: Diaperinae) from the Socotra Archipelago. Acta Entomologica Musei Nationalis Pragae 52: 303-314, DOI: 10.5281/zenodo.5354846
7A2687556C79AF3AFDF4FEFFFE6DC81A.taxon	type_taxon	Type species. Nanocaecus hlavaci sp. nov. by present designation. Tribal assignment. According to DOYEN & LAWRENCE (1979) the existence of an exposed and visible membrane between clypeus and labrum, as well as the long and acuminate last maxillary palpomere are synapomorphic characters of the subtribe Anopidiina (members of the subtribe Gnathidiina have no visible membrane between clypeus and labrum, and truncate last palpomere is broadened). Four African genera of the Anopidiina (Anopidium, Paranopidium, Peyrierasia Dajoz, 1974, Pseudanopidium Dajoz, 1973) are keyed by DAJOZ (1974), mainly using characters of the antennae, scutellum and body shape. However, this key is incomplete because DOYEN & LAWRENCE (1979) subsequently added two other genera with African species to this subtribe (Paralyreus Grouvelle, 1918, Tyrtaeus Champion, 1913, see the complete key and catalogue below).	en	Schawaller, Wolfgang, Purchart, Luboš (2012): Nanocaecus hlavaci gen. & sp. nov. - first record of the tribe Gnathidiini (Coleoptera: Tenebrionidae: Diaperinae) from the Socotra Archipelago. Acta Entomologica Musei Nationalis Pragae 52: 303-314, DOI: 10.5281/zenodo.5354846
7A2687556C79AF3AFDF4FEFFFE6DC81A.taxon	etymology	Etymology. The generic name is composed of the Latin words nanus (= dwarf) and caecus (= blind); gender masculine.	en	Schawaller, Wolfgang, Purchart, Luboš (2012): Nanocaecus hlavaci gen. & sp. nov. - first record of the tribe Gnathidiini (Coleoptera: Tenebrionidae: Diaperinae) from the Socotra Archipelago. Acta Entomologica Musei Nationalis Pragae 52: 303-314, DOI: 10.5281/zenodo.5354846
7A2687556C79AF3EFE10FAA6FF42CE91.taxon	description	(Figs. 1 – 13)	en	Schawaller, Wolfgang, Purchart, Luboš (2012): Nanocaecus hlavaci gen. & sp. nov. - first record of the tribe Gnathidiini (Coleoptera: Tenebrionidae: Diaperinae) from the Socotra Archipelago. Acta Entomologica Musei Nationalis Pragae 52: 303-314, DOI: 10.5281/zenodo.5354846
7A2687556C79AF3EFE10FAA6FF42CE91.taxon	materials_examined	Type locality. Yemen, Socotra Island, Al Haghier Mts., Scant Mt. env. Type material. HOLOTYPE: ♁ (NMPC), labelled: YEMEN, SOCOTRA Island, Al Haghier Mts., Scant Mt. env., 1450 m, 12 ° 34.6 ′ N, 54 ° 01.5 ′ E, P. Hlaváč leg., 12 - 13. xi. 2010. Paratypes (27 specimens NMPC, 6 SMNS, 6 LPCB, 5 BMNH, 5 HNHM, 5 ZSMC): same data as holotype; (11 NMPC, 2 SMNS, 2 LPCB): same data as holotype, Jiří Hájek leg.; (11 NMPC, 2 SMNS, 2 LPCB): same data as holotype, Jan Bezděk leg.; (4 JBCP): same data as holotype, Jan Batelka leg.	en	Schawaller, Wolfgang, Purchart, Luboš (2012): Nanocaecus hlavaci gen. & sp. nov. - first record of the tribe Gnathidiini (Coleoptera: Tenebrionidae: Diaperinae) from the Socotra Archipelago. Acta Entomologica Musei Nationalis Pragae 52: 303-314, DOI: 10.5281/zenodo.5354846
7A2687556C79AF3EFE10FAA6FF42CE91.taxon	description	Description. Body length 1.9 – 2.1 mm, width of elytra 0.6 mm. Surface and all appendages unicoloured shining brown. Head without eyes, frons with regular, fine and separate punctation, each puncture bearing a small acute seta, setae distinctly longer at anterior margin of clypeus, clypeal suture absent; membrane between clypeus and labrum exposed; labrum of semicircular shape; mandibles bifid; last maxillary palpomere long oval with finger-like tip, ventrally with a row of 5 blunt sensillae, tip with field of smaller sensillae; last labial palpomere swollen in middle and tip with field of sensillae; mentum hexagonal; without transverse or medial gular impression; antenna with 10 antennomeres, antennomere 3 not prolonged, with apical four antennomeres forming dense but not fused oval club, club apically of each joint with dense and long acute setae. Pronotum with regular, fine punctation and setation similar as on head; disc without impressions, slightly convex until the lateral margin, lateral margin equally rounded and basally wider separated from disc than distally; anterior margin not sinuated and not protruding in middle, anterior corners rounded; basal (posterior) margin feebly sinuated, basal corners rectangular; all margins unbordered, before basal margin laterally with transverse row of larger punctures; epipleura smooth and unpunctured; prosternal apophysis flat and not projected. Meso- and metaventrite with extraordinary large but not confluent punctation. Scutellum small but visible. Wings completely absent. Elytra long oval, widest before middle, surface with punctures larger than those on pronotum and head, each puncture bearing acute microsetae, punctation irregular, without any rows or traces of rows, also without scutellar striolae; base of elytra completely unbordered; in dorsal view lateral margin visible only in anterior quarter; epipleura broad in anterior three quarters, abruptly narrower in posterior quarter. Abdominal ventrites with regular and separate punctation, punctures of first ventrite as large as punctures of metaventrite, punctures bearing acute microsetae; ventrites 3 / 4 and 4 / 5 somewhat more separated than basal ventrites, but membranes between them not exposed; last ventrite of semicircular shape and regularly bordered, without any impressions or other modifications. Femora claviform, tibiae rounded without any keels, tibial spurs short; tarsal formula 5 - 5 - 4. Tip of aedeagus see Fig. 12, tip of ovipositor see Fig. 13. No distinct external sexual dimorphism.	en	Schawaller, Wolfgang, Purchart, Luboš (2012): Nanocaecus hlavaci gen. & sp. nov. - first record of the tribe Gnathidiini (Coleoptera: Tenebrionidae: Diaperinae) from the Socotra Archipelago. Acta Entomologica Musei Nationalis Pragae 52: 303-314, DOI: 10.5281/zenodo.5354846
7A2687556C79AF3EFE10FAA6FF42CE91.taxon	etymology	Etymology. Named in honour of Peter Hlaváč (Košice, Slovakia), specialist of Pselaphinae and main collector of the type series. Collection circumstances. The specimens of the type series were sifted from litter under shrubs and trees in high altitudes (above 1400 m) of the Haghier Mountains.	en	Schawaller, Wolfgang, Purchart, Luboš (2012): Nanocaecus hlavaci gen. & sp. nov. - first record of the tribe Gnathidiini (Coleoptera: Tenebrionidae: Diaperinae) from the Socotra Archipelago. Acta Entomologica Musei Nationalis Pragae 52: 303-314, DOI: 10.5281/zenodo.5354846
7A2687556C79AF3EFE10FAA6FF42CE91.taxon	distribution	Distribution. So far known only from the type locality in Haghier Mountains, Socotra, Yemen.	en	Schawaller, Wolfgang, Purchart, Luboš (2012): Nanocaecus hlavaci gen. & sp. nov. - first record of the tribe Gnathidiini (Coleoptera: Tenebrionidae: Diaperinae) from the Socotra Archipelago. Acta Entomologica Musei Nationalis Pragae 52: 303-314, DOI: 10.5281/zenodo.5354846
7A2687556C73AF31FF65FA08FF5BCF15.taxon	discussion	Remarks. Betschia was established by DAJOZ (1980) as a new genus of the family Colydiidae for Betschia minuta Dajoz, 1980 (by monotypy). IVIE & ŚLIPIŃSKI (1990) transferred the genus Betschia from the family Colydiidae to the Tenebrionidae. Betschia, along with its associated family-group name Betschiini Dajoz, 1980 were included under Tenebrionidae incertae sedis in BOUCHARD et al. (2011). The latter authors mentioned that its systematic placement could be close to Gnathidiini. Members of the tribe Gnathidiini possess the tarsal formula 5 - 5 - 4 or 4 - 4 - 4 (MATHEWS & BOUCHARD 2008) while the genus Betschia has the tarsal formula 3 - 3 - 3. Therefore its placement remains uncertain, and for this reason it is considered here as incertae sedis as well.	en	Schawaller, Wolfgang, Purchart, Luboš (2012): Nanocaecus hlavaci gen. & sp. nov. - first record of the tribe Gnathidiini (Coleoptera: Tenebrionidae: Diaperinae) from the Socotra Archipelago. Acta Entomologica Musei Nationalis Pragae 52: 303-314, DOI: 10.5281/zenodo.5354846
7A2687556C72AF31FF83FDD6FBB1C9B6.taxon	discussion	Remarks. DAJOZ (1977 b) described two genera: the genus Mauritianopidium Dajoz, 1977 b with species Mauritianopidium oculatum Dajoz, 1977 b, and the genus Mireanopidium Dajoz, 1977 b based on three species – Mireanopidium camerunensis Dajoz, 1977 b (type species of the genus), M. montanum Dajoz, 1977 b and M. distinctum Dajoz, 1977 b, and placed them in the tribe Anopidiini of the family Colydiidae. The tribe Anopidiini is presently recognised as the subtribe Anopidiina of the tribe Gnathidiini in the family Tenebrionidae. While Mauritianopidum undoubtedly belongs to Tenebrionidae, Mireanopidium does not. The main reason to exclude the latter genus from Tenebrionidae is the fact that it possesses 10 - segmented antenna with 3 - segmented club and the tarsal formula 4 - 4 - 3. We believe that it might be a member of the family Colydiidae as originally placed by Dajoz. However, the genus was not mentioned in the catalogue of this family presented by IVIE & ŚLIPIŃSKI (1990), so its systematic placement is uncertain. Therefore it is considered here as incertae sedis. Generic assignment unclear Anopidium errans Pope, 1962 Tanzania (Kilimanjaro, Mt. Meru) Remarks. The species Anopidium errans described by POPE (1962) was provisionally retained in the genus Anopidium by BREMER (1998), although he suggested, based on several morphological characters, that for this species a new genus should probably be established. To this date, no record of a transfer of A. errans to another (new) genus has been found in the literature, therefore this species is considered here as incertae sedis.	en	Schawaller, Wolfgang, Purchart, Luboš (2012): Nanocaecus hlavaci gen. & sp. nov. - first record of the tribe Gnathidiini (Coleoptera: Tenebrionidae: Diaperinae) from the Socotra Archipelago. Acta Entomologica Musei Nationalis Pragae 52: 303-314, DOI: 10.5281/zenodo.5354846
