taxonID	type	description	language	source
B659B9ABAF9759E792C7553D71C2FEA3.taxon	diagnosis	Diagnosis. Differs from other members of the subgenus Tetrardisia in having linear-oblong leaves, with a long, acuminate-caudate apex, and finely serrulate margins.	en	Julius, Avelinah, Kajita, Tadashi, Utteridge, Timothy M. A. (2020): Two new species of Ardisia subgenus Tetrardisia (Primulaceae - Myrsinoideae) from Borneo. PhytoKeys 145: 139-148, DOI: http://dx.doi.org/10.3897/phytokeys.145.48573, URL: http://dx.doi.org/10.3897/phytokeys.145.48573
B659B9ABAF9759E792C7553D71C2FEA3.taxon	materials_examined	Type. INDONESIA. Borneo: Central Kalimantan, Kotawaringan [Kotawaringin] Timur, S. Mentaya, km 92 from Sangai, Plot 8, [1 ° 18 ' S, 112 ° 32 ' E], 100 m elevation, 18 May 1993, Argent et al. 93187 (holotype E!; iso: BO).	en	Julius, Avelinah, Kajita, Tadashi, Utteridge, Timothy M. A. (2020): Two new species of Ardisia subgenus Tetrardisia (Primulaceae - Myrsinoideae) from Borneo. PhytoKeys 145: 139-148, DOI: http://dx.doi.org/10.3897/phytokeys.145.48573, URL: http://dx.doi.org/10.3897/phytokeys.145.48573
B659B9ABAF9759E792C7553D71C2FEA3.taxon	description	Description. Shrub ca. 1 m high; stems sparsely scaly when young, soon glabrous, slightly flexuous, winged between the nodes with raised lines running along the internodes between the petiole bases. Indumentum of sessile, circular, peltate scales up to 0.05 mm in diameter, on young parts, leaves and inflorescence. Leaves alternate; petioles 3 - 5 mm long, winged by the decurrent leaf base, glabrous; lamina chartaceous, linear-oblong, (7 -) 10 - 11.5 x 1 - 2 cm, with dense gland-dots throughout apex acuminate-caudate, acumen 1 - 2.5 cm long; base cuneate, between the higher order venation, lacking hairs, sparsely scaly on the lower midrib, glabrous above, midrib sunken above, raised beneath; lateral veins ca. 48 pairs, semicraspedodromus, and with 1 - 2 intersecondary veins within each pair; intercostal veins obscure. Inflorescences terminal on lateral branches, proximally laxly paniculate with subumbellate branches with 2 - flowers distally racemose; peduncle and rachis (1.5 -) 2.5 - 4 cm long, slightly flexuose, sparsely scaly; bracts linear-lanceolate, (0.4 -) 1.5 - 1.7 x 0.4 mm, very sparsely ciliate with few scattered hairs. Flowers ca. 8; pedicels slender, ca. 1.5 cm long, sparsely scaly; calyx-lobes 4, green, gland-dotted, lacking hairs, sparsely scaly outside, ovate, ca. 1 x 0.8 mm, apex acute, margin very sparsely ciliate with few scattered short hairs; corolla-lobes 4, reflexed, twisted apically, white with orange-brown elongated-dots, lanceolate, 6.4 - 7 x 1.7 - 2.3 mm, glabrous on both surfaces; stamens 4, spreading upright (in open flower), filament + / - sessile, anthers twisted apically, lanceolate, ca. 5 - 6.2 x 0.8 mm, apex elongated into a hyaline tip, thecae open by longitudinal-slits, with scattered, lineate, orange brown dots behind, glabrous; ovary ovoid, ca. 7 x 6 mm, style and stigma filiform, ca. 6.8 mm long, ovules ca. 6 in 1 - series. Fruits n. v.	en	Julius, Avelinah, Kajita, Tadashi, Utteridge, Timothy M. A. (2020): Two new species of Ardisia subgenus Tetrardisia (Primulaceae - Myrsinoideae) from Borneo. PhytoKeys 145: 139-148, DOI: http://dx.doi.org/10.3897/phytokeys.145.48573, URL: http://dx.doi.org/10.3897/phytokeys.145.48573
B659B9ABAF9759E792C7553D71C2FEA3.taxon	distribution	Distribution. Endemic to Borneo; known only from Sungai Mentaya, Kotawaringin Timur, Central Kalimantan.	en	Julius, Avelinah, Kajita, Tadashi, Utteridge, Timothy M. A. (2020): Two new species of Ardisia subgenus Tetrardisia (Primulaceae - Myrsinoideae) from Borneo. PhytoKeys 145: 139-148, DOI: http://dx.doi.org/10.3897/phytokeys.145.48573, URL: http://dx.doi.org/10.3897/phytokeys.145.48573
B659B9ABAF9759E792C7553D71C2FEA3.taxon	etymology	Etymology. The species epithet commemorates Graham Charles George Argent (1941 - 2019), a prominent botanist on tropical botany in South-East Asia and a leading expert on Ericaceae, especially the tropical ' Vireya' rhododendrons, and collector of the type material.	en	Julius, Avelinah, Kajita, Tadashi, Utteridge, Timothy M. A. (2020): Two new species of Ardisia subgenus Tetrardisia (Primulaceae - Myrsinoideae) from Borneo. PhytoKeys 145: 139-148, DOI: http://dx.doi.org/10.3897/phytokeys.145.48573, URL: http://dx.doi.org/10.3897/phytokeys.145.48573
2BA31A89C3645DCABCC32FA1601481A6.taxon	diagnosis	Diagnosis. Similar to Ardisia tetrasepala in its simple and compact inflorescence with corymbose flowers but differs mainly vegetatively in having leaves laxly arranged, longer petioles that are covered with glandular hairs, a chartaceous lamina without marginal secondary veins, surface with scattered stellate hairs near the margin beneath and leaf base obtuse or + / - cuneate. The corolla lobes are smooth without lepidote scales (vs. lepidote scales present in A. tetrasepala).	en	Julius, Avelinah, Kajita, Tadashi, Utteridge, Timothy M. A. (2020): Two new species of Ardisia subgenus Tetrardisia (Primulaceae - Myrsinoideae) from Borneo. PhytoKeys 145: 139-148, DOI: http://dx.doi.org/10.3897/phytokeys.145.48573, URL: http://dx.doi.org/10.3897/phytokeys.145.48573
2BA31A89C3645DCABCC32FA1601481A6.taxon	materials_examined	Type. MALAYSIA. Borneo: Sarawak, Tatau District, Ulu Merirai, Gua Naga, [02 ° 39 ' 12 " N, 113 ° 03 ' 05 " E], 11 July 2005, Julia et al. S 95726 (holotype KEP!; iso: SAR, SING)	en	Julius, Avelinah, Kajita, Tadashi, Utteridge, Timothy M. A. (2020): Two new species of Ardisia subgenus Tetrardisia (Primulaceae - Myrsinoideae) from Borneo. PhytoKeys 145: 139-148, DOI: http://dx.doi.org/10.3897/phytokeys.145.48573, URL: http://dx.doi.org/10.3897/phytokeys.145.48573
2BA31A89C3645DCABCC32FA1601481A6.taxon	description	Description. A small shrub, less than 1 m high; stems glabrescent, slightly flexuous, winged between the nodes with obscure raised lines running along the internodes between the petiole bases. Indumentum of simple, short (stalked), glandular, or stellate (star-liked), pale brown to rusty hairs (visible under microscope). Leaves alternate; petioles 1.5 - 3 cm long, sparsely to densely hairy towards midrib, simple, short, glandular hairs; lamina chartaceous, with dense gland-dots throughout the leaf, glabrous on both surfaces except for margin with scattered stellate hairs beneath, elliptic-lanceolate, 17 - 19 x 5 - 5.5 cm, apex long acuminate and slightly caudate with acumen 1.5 - 2.0 cm long; base obtuse or + / - cuneate, margin obscurely denticulate being entire in appearance, midrib flat or slightly sunken above, raised beneath, glabrous except hairy with glandular hairs beneath, denser near leaf base; lateral veins 13 - 15 pairs, distinct above, prominent beneath, inter-secondary veins present in between; intercostal veins percurrent, distinct beneath. Inflorescence terminal on lateral branches, simple with flowers arranged in corymbs; peduncle and rachis 4 - 5 cm long, densely hairy with glandular hairs; bracts ovate to elliptic, 0.5 - 1.5 mm long, hairy with glandular hairs. Flowers ca. 24; pedicels slender, 1.2 - 1.8 cm long, up to 2 cm long in fruiting, covered with dense, glandular hairs; calyx-lobes 4, purplish, gland-dotted, lacking hairs, broadly ovate, 1 - 1.5 x 1 - 1.5 mm, apex obtuse, margin incised, sparsely ciliate with short, thick hairs; corolla-lobes 4, reflexed, twisted apically, purplish with transparent margin, lanceolate, ca. 8 x 2.5 mm, glabrous on both surfaces; stamens 4, spreading upright (in open flower), filament + / - sessile, anthers twisted apically, lanceolate, ca. 6 x 1 mm, apex without a prolonged hyaline tip, thecae opening by longitudinal slits, with lineate, black dots behind, glabrous; ovary globose, ca. 1 x 0.8 mm, style and stigma filiform, ca. 6 mm long, ovules 4 - 6 in 1 - series. Fruits ripe bright red, globose, ca. 6 x 5 mm.	en	Julius, Avelinah, Kajita, Tadashi, Utteridge, Timothy M. A. (2020): Two new species of Ardisia subgenus Tetrardisia (Primulaceae - Myrsinoideae) from Borneo. PhytoKeys 145: 139-148, DOI: http://dx.doi.org/10.3897/phytokeys.145.48573, URL: http://dx.doi.org/10.3897/phytokeys.145.48573
2BA31A89C3645DCABCC32FA1601481A6.taxon	distribution	Distribution. Endemic in Borneo, Sarawak; known only from Gua Naga, Ulu Merirai area, Tatau District.	en	Julius, Avelinah, Kajita, Tadashi, Utteridge, Timothy M. A. (2020): Two new species of Ardisia subgenus Tetrardisia (Primulaceae - Myrsinoideae) from Borneo. PhytoKeys 145: 139-148, DOI: http://dx.doi.org/10.3897/phytokeys.145.48573, URL: http://dx.doi.org/10.3897/phytokeys.145.48573
2BA31A89C3645DCABCC32FA1601481A6.taxon	etymology	Etymology. Ardisia nagaensis is very local and was found at only one locality, Gua Naga, for which it is named.	en	Julius, Avelinah, Kajita, Tadashi, Utteridge, Timothy M. A. (2020): Two new species of Ardisia subgenus Tetrardisia (Primulaceae - Myrsinoideae) from Borneo. PhytoKeys 145: 139-148, DOI: http://dx.doi.org/10.3897/phytokeys.145.48573, URL: http://dx.doi.org/10.3897/phytokeys.145.48573
