taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
53CC67C870175165968AD28480EFC5C1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510474	https://doi.org/10.3897/zookeys.1018.61033.figure10	Figure 10. Holotype Cryptophyllium animatum gen. et sp. nov., photographs by RTC A habitus, dorsal B details of the protibial and profemoral lobes C details of the head and thorax, dorsal D genitalia, ventral E head-thorax, lateral.	Figure 10. Holotype Cryptophyllium animatum gen. et sp. nov., photographs by RTC A habitus, dorsal B details of the protibial and profemoral lobes C details of the head and thorax, dorsal D genitalia, ventral E head-thorax, lateral.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
53CC67C870175165968AD28480EFC5C1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510525	https://doi.org/10.3897/zookeys.1018.61033.figure61	Figure 61. Cryptophyllium rarum comb. nov. male from Cuc Phuong N. P., Ninh Binh Province, Vietnam collected by Jerome Constant and Joachim Bresseel (RBINS), photographs by Jerome Constant (RBINS) A habitus, dorsal B habitus, ventral C details of the base of antennae, head, and thorax, dorsal D details of the base of antennae, head, and thorax, lateral E pro- femoral and tibial lobes, dorsal F terminalia, dorsal G genitalia, ventral.	Figure 61. Cryptophyllium rarum comb. nov. male from Cuc Phuong N. P., Ninh Binh Province, Vietnam collected by Jerome Constant and Joachim Bresseel (RBINS), photographs by Jerome Constant (RBINS) A habitus, dorsal B habitus, ventral C details of the base of antennae, head, and thorax, dorsal D details of the base of antennae, head, and thorax, lateral E pro- femoral and tibial lobes, dorsal F terminalia, dorsal G genitalia, ventral.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
53CC67C870175165968AD28480EFC5C1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510495	https://doi.org/10.3897/zookeys.1018.61033.figure31	Figure 31. Holotype Cryptophyllium faulkneri gen. et sp. nov., photographs by RTC A details of the base of antennae, head, and thorax, dorsal B habitus, dorsal C details of thorax, ventral D details of head and thorax, lateral.	Figure 31. Holotype Cryptophyllium faulkneri gen. et sp. nov., photographs by RTC A details of the base of antennae, head, and thorax, dorsal B habitus, dorsal C details of thorax, ventral D details of head and thorax, lateral.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
53CC67C870175165968AD28480EFC5C1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510471	https://doi.org/10.3897/zookeys.1018.61033.figure7	Figure 7. Typical tegmina and alae venation present in the Cryptophyllium gen. nov., photographed by RTC A female: Cryptophyllium celebicum comb. nov. (Coll RC 16 - 075) B male: Cryptophyllium yunnanense comb. nov. (Coll RC 16 - 120). Abbreviations: C (costa); Sc (subcosta); R (radius); R 1 (radius 1); R 2 (radius 2); Rs (radial sector); M (media); MA (media anterior); MP (media posterior); MP 1 (first media posterior); MP 2 (second media posterior); MA + MP (media anterior fused with media posterior); Cu (cubitus); CuA (cubitus anterior); CuP (cubitus posterior); Cu + 1 AA (cubitus and first anterior anal); 1 A (first anal); 1 AA- 7 AA (first-seventh anterior anal); 1 PA- 7 PA (first-seventh posterior anal).	Figure 7. Typical tegmina and alae venation present in the Cryptophyllium gen. nov., photographed by RTC A female: Cryptophyllium celebicum comb. nov. (Coll RC 16 - 075) B male: Cryptophyllium yunnanense comb. nov. (Coll RC 16 - 120). Abbreviations: C (costa); Sc (subcosta); R (radius); R 1 (radius 1); R 2 (radius 2); Rs (radial sector); M (media); MA (media anterior); MP (media posterior); MP 1 (first media posterior); MP 2 (second media posterior); MA + MP (media anterior fused with media posterior); Cu (cubitus); CuA (cubitus anterior); CuP (cubitus posterior); Cu + 1 AA (cubitus and first anterior anal); 1 A (first anal); 1 AA- 7 AA (first-seventh anterior anal); 1 PA- 7 PA (first-seventh posterior anal).	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
FD7965C10ABE5CC29D3C500CF043194D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510475	https://doi.org/10.3897/zookeys.1018.61033.figure11	Figure 11. Cryptophyllium athanysus comb. nov. holotype female, photographs by Paul Brock (NHMUK) A habitus, dorsal view B details of the thorax, dorsal C genitalia details, ventral D illustration of a Cryptophyllium athanysus comb. nov. egg from Greene (1906).	Figure 11. Cryptophyllium athanysus comb. nov. holotype female, photographs by Paul Brock (NHMUK) A habitus, dorsal view B details of the thorax, dorsal C genitalia details, ventral D illustration of a Cryptophyllium athanysus comb. nov. egg from Greene (1906).	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
FD7965C10ABE5CC29D3C500CF043194D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510476	https://doi.org/10.3897/zookeys.1018.61033.figure12	Figure 12. Live Cryptophyllium athanysus comb. nov. observed in the Sinharaja Forest Reserve, Sri Lanka A, B female photographed by Alberto Carrera (A photograph purchased from Alamy photo stock website B photograph purchased from Adobe Stock) C first record of a live male observed and photographed by Angela Christine Chua (Singapore) in January 2020.	Figure 12. Live Cryptophyllium athanysus comb. nov. observed in the Sinharaja Forest Reserve, Sri Lanka A, B female photographed by Alberto Carrera (A photograph purchased from Alamy photo stock website B photograph purchased from Adobe Stock) C first record of a live male observed and photographed by Angela Christine Chua (Singapore) in January 2020.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
803CD1D08BC65EBB869D05E0B758D8B4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510469	https://doi.org/10.3897/zookeys.1018.61033.figure5	Figure 5. Details of the Cryptophyllium gen. nov. male vomer. All from the RBINS collection, prepared and photographed by Jerome Constant (RBINS). Views 1 ventral 2 right lateral of the ventrally oriented vomer 3 left lateral of the ventrally oriented vomer A Cryptophyllium westwoodii comb. nov., Thailand, Na Haeo B Cryptophyllium khmer gen. et sp. nov. Cambodia, Kbal Spean C Cryptophyllium phami gen. et sp. nov., Vietnam, Cat Tien D Cryptophyllium khmer gen. et sp. nov., Cambodia, Tatai E Cryptophyllium bollensi gen. et sp. nov., Vietnam, Phuoc Binh F Cryptophyllium oyae comb. nov., Laos, Mt. Phu Phan G Cryptophyllium bankoi gen. et sp. nov., Vietnam, Ba Na-Nui Chua H Cryptophyllium icarus gen. et sp. nov., Vietnam, Bidoup-Nui Ba I Cryptophyllium rarum comb. nov., Vietnam, Cuc Phuong J Cryptophyllium limogesi gen. et sp. nov., Vietnam, Ta Dung. Scale bars, 1.0 mm.	Figure 5. Details of the Cryptophyllium gen. nov. male vomer. All from the RBINS collection, prepared and photographed by Jerome Constant (RBINS). Views 1 ventral 2 right lateral of the ventrally oriented vomer 3 left lateral of the ventrally oriented vomer A Cryptophyllium westwoodii comb. nov., Thailand, Na Haeo B Cryptophyllium khmer gen. et sp. nov. Cambodia, Kbal Spean C Cryptophyllium phami gen. et sp. nov., Vietnam, Cat Tien D Cryptophyllium khmer gen. et sp. nov., Cambodia, Tatai E Cryptophyllium bollensi gen. et sp. nov., Vietnam, Phuoc Binh F Cryptophyllium oyae comb. nov., Laos, Mt. Phu Phan G Cryptophyllium bankoi gen. et sp. nov., Vietnam, Ba Na-Nui Chua H Cryptophyllium icarus gen. et sp. nov., Vietnam, Bidoup-Nui Ba I Cryptophyllium rarum comb. nov., Vietnam, Cuc Phuong J Cryptophyllium limogesi gen. et sp. nov., Vietnam, Ta Dung. Scale bars, 1.0 mm.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
803CD1D08BC65EBB869D05E0B758D8B4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510477	https://doi.org/10.3897/zookeys.1018.61033.figure13	Figure 13. Cryptophyllium bankoi gen. et sp. nov. paratype female nymph collected in Bach Ma N. P. on Rubus sp. (Rosaceae), Vietnam, photographed by Jerome Constant (RBINS) on 11 th July 2011. Molecular sample RBINS 06 within this study A dorsal view with abdomen curled B dorsal view with abdomen held flat.	Figure 13. Cryptophyllium bankoi gen. et sp. nov. paratype female nymph collected in Bach Ma N. P. on Rubus sp. (Rosaceae), Vietnam, photographed by Jerome Constant (RBINS) on 11 th July 2011. Molecular sample RBINS 06 within this study A dorsal view with abdomen curled B dorsal view with abdomen held flat.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
803CD1D08BC65EBB869D05E0B758D8B4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510478	https://doi.org/10.3897/zookeys.1018.61033.figure14	Figure 14. Cryptophyllium bankoi gen. et sp. nov. holotype male, photographs by RTC A front leg details, dorsal B details of the base of antennae, head, thorax C habitus, dorsal D details of genitalia, ventral.	Figure 14. Cryptophyllium bankoi gen. et sp. nov. holotype male, photographs by RTC A front leg details, dorsal B details of the base of antennae, head, thorax C habitus, dorsal D details of genitalia, ventral.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
803CD1D08BC65EBB869D05E0B758D8B4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510479	https://doi.org/10.3897/zookeys.1018.61033.figure15	Figure 15. Cryptophyllium bankoi gen. et sp. nov. paratype male from the Ba Na-Nui Chua Nature Reserve (RBINS), photographs by Jerome Constant (RBINS) A habitus, dorsal B habitus, ventral C pro- femoral and tibial, dorsal D details of the base of antennae, head, and thorax, dorsal E details of the base of antennae, head, and thorax, lateral F terminalia, dorsal G genitalia, ventral.	Figure 15. Cryptophyllium bankoi gen. et sp. nov. paratype male from the Ba Na-Nui Chua Nature Reserve (RBINS), photographs by Jerome Constant (RBINS) A habitus, dorsal B habitus, ventral C pro- femoral and tibial, dorsal D details of the base of antennae, head, and thorax, dorsal E details of the base of antennae, head, and thorax, lateral F terminalia, dorsal G genitalia, ventral.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
803CD1D08BC65EBB869D05E0B758D8B4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510523	https://doi.org/10.3897/zookeys.1018.61033.figure59	Figure 59. Cryptophyllium rarum comb. nov. male and female A male, habitus, dorsal, molecular sample within this study Coll RC 16 - 115 from Vietnam, Da Nang Province, photograph by RTC B details of the head and thorax, dorsal, Coll RC 16 - 115, photograph by RTC C female, habitus, dorsal, from China: Guangxi Province, Nanning city, Shuangding town, molecular sample DZW 08 within this study, Photographs by Zhiwei Dong (KIZ) D female, genitalia, ventral, photographs by Zhiwei Dong (KIZ).	Figure 59. Cryptophyllium rarum comb. nov. male and female A male, habitus, dorsal, molecular sample within this study Coll RC 16 - 115 from Vietnam, Da Nang Province, photograph by RTC B details of the head and thorax, dorsal, Coll RC 16 - 115, photograph by RTC C female, habitus, dorsal, from China: Guangxi Province, Nanning city, Shuangding town, molecular sample DZW 08 within this study, Photographs by Zhiwei Dong (KIZ) D female, genitalia, ventral, photographs by Zhiwei Dong (KIZ).	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
74B70D7C21635BE19FE3D1049C04008F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510469	https://doi.org/10.3897/zookeys.1018.61033.figure5	Figure 5. Details of the Cryptophyllium gen. nov. male vomer. All from the RBINS collection, prepared and photographed by Jerome Constant (RBINS). Views 1 ventral 2 right lateral of the ventrally oriented vomer 3 left lateral of the ventrally oriented vomer A Cryptophyllium westwoodii comb. nov., Thailand, Na Haeo B Cryptophyllium khmer gen. et sp. nov. Cambodia, Kbal Spean C Cryptophyllium phami gen. et sp. nov., Vietnam, Cat Tien D Cryptophyllium khmer gen. et sp. nov., Cambodia, Tatai E Cryptophyllium bollensi gen. et sp. nov., Vietnam, Phuoc Binh F Cryptophyllium oyae comb. nov., Laos, Mt. Phu Phan G Cryptophyllium bankoi gen. et sp. nov., Vietnam, Ba Na-Nui Chua H Cryptophyllium icarus gen. et sp. nov., Vietnam, Bidoup-Nui Ba I Cryptophyllium rarum comb. nov., Vietnam, Cuc Phuong J Cryptophyllium limogesi gen. et sp. nov., Vietnam, Ta Dung. Scale bars, 1.0 mm.	Figure 5. Details of the Cryptophyllium gen. nov. male vomer. All from the RBINS collection, prepared and photographed by Jerome Constant (RBINS). Views 1 ventral 2 right lateral of the ventrally oriented vomer 3 left lateral of the ventrally oriented vomer A Cryptophyllium westwoodii comb. nov., Thailand, Na Haeo B Cryptophyllium khmer gen. et sp. nov. Cambodia, Kbal Spean C Cryptophyllium phami gen. et sp. nov., Vietnam, Cat Tien D Cryptophyllium khmer gen. et sp. nov., Cambodia, Tatai E Cryptophyllium bollensi gen. et sp. nov., Vietnam, Phuoc Binh F Cryptophyllium oyae comb. nov., Laos, Mt. Phu Phan G Cryptophyllium bankoi gen. et sp. nov., Vietnam, Ba Na-Nui Chua H Cryptophyllium icarus gen. et sp. nov., Vietnam, Bidoup-Nui Ba I Cryptophyllium rarum comb. nov., Vietnam, Cuc Phuong J Cryptophyllium limogesi gen. et sp. nov., Vietnam, Ta Dung. Scale bars, 1.0 mm.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
74B70D7C21635BE19FE3D1049C04008F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510470	https://doi.org/10.3897/zookeys.1018.61033.figure6	Figure 6. Details of female phylliid antennae illustrated from specimens within the RBINS collection, illustrations by Julien Caudron (RBINS). Note that the figures start with the scapus, therefore the diagnostic fourth antennomere is the third one shown; to ensure clarity A has the segment number listed to the right of the image. Note that the fourth antennomere of Phyllium (A) is longer than the fifth antennomere in comparison to the Cryptophyllium gen. nov. representatives (B-H), which have the fourth antennomere short and disk-like A Phyllium ericoriai, Philippines B Cryptophyllium bollensi gen. et sp. nov., Vietnam, Phuoc Binh C Cryptophyllium celebicum comb. nov., Indonesia, Sulawesi D Cryptophyllium khmer gen. et sp. nov., Cambodia, Siem Reap Province E Cryptophyllium nuichuaense gen. et sp. nov., Vietnam, Nui Chua F Cryptophyllium phami gen. et sp. nov., Vietnam, Cat Tien G Cryptophyllium westwoodii comb. nov., Laos, Bokeo Province H Cryptophyllium westwoodii comb. nov., Thailand, Chiang Mai. Drawn to relative scale.	Figure 6. Details of female phylliid antennae illustrated from specimens within the RBINS collection, illustrations by Julien Caudron (RBINS). Note that the figures start with the scapus, therefore the diagnostic fourth antennomere is the third one shown; to ensure clarity A has the segment number listed to the right of the image. Note that the fourth antennomere of Phyllium (A) is longer than the fifth antennomere in comparison to the Cryptophyllium gen. nov. representatives (B-H), which have the fourth antennomere short and disk-like A Phyllium ericoriai, Philippines B Cryptophyllium bollensi gen. et sp. nov., Vietnam, Phuoc Binh C Cryptophyllium celebicum comb. nov., Indonesia, Sulawesi D Cryptophyllium khmer gen. et sp. nov., Cambodia, Siem Reap Province E Cryptophyllium nuichuaense gen. et sp. nov., Vietnam, Nui Chua F Cryptophyllium phami gen. et sp. nov., Vietnam, Cat Tien G Cryptophyllium westwoodii comb. nov., Laos, Bokeo Province H Cryptophyllium westwoodii comb. nov., Thailand, Chiang Mai. Drawn to relative scale.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
74B70D7C21635BE19FE3D1049C04008F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510472	https://doi.org/10.3897/zookeys.1018.61033.figure8	Figure 8. Known eggs for the Cryptophyllium gen. nov. set in couplets of lateral and then dorsal views. All images except for K and L were taken by Bruno Kneubu ̈ hler (Switzerland), K and L images taken by the first author, Q and R images taken by Rene ́ Limoges (IMQC) A, B Cryptophyllium westwoodii comb. nov. C, D Cryptophyllium tibetense comb. nov. E, F Cryptophyllium bollensi gen. et sp. nov. G, H Cryptophyllium oyae comb. nov. I, J Cryptophyllium icarus gen. et sp. nov. K, L Cryptophyllium liyananae gen. et sp. nov. M, N Cryptophyllium chrisangi comb. nov. O, P Cryptophyllium celebicum comb. nov. Q, R Cryptophyllium limogesi gen. et sp. nov. Scale bars 2.0 mm long.	Figure 8. Known eggs for the Cryptophyllium gen. nov. set in couplets of lateral and then dorsal views. All images except for K and L were taken by Bruno Kneubu ̈ hler (Switzerland), K and L images taken by the first author, Q and R images taken by Rene ́ Limoges (IMQC) A, B Cryptophyllium westwoodii comb. nov. C, D Cryptophyllium tibetense comb. nov. E, F Cryptophyllium bollensi gen. et sp. nov. G, H Cryptophyllium oyae comb. nov. I, J Cryptophyllium icarus gen. et sp. nov. K, L Cryptophyllium liyananae gen. et sp. nov. M, N Cryptophyllium chrisangi comb. nov. O, P Cryptophyllium celebicum comb. nov. Q, R Cryptophyllium limogesi gen. et sp. nov. Scale bars 2.0 mm long.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
74B70D7C21635BE19FE3D1049C04008F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510473	https://doi.org/10.3897/zookeys.1018.61033.figure9	Figure 9. Freshly hatched nymphs known for the Cryptophyllium gen. nov., all except for C bred and photographed by Bruno Kneubu ̈ hler (Switzerland) A Cryptophyllium celebicum comb. nov. B Cryptophyllium chrisangi comb. nov. C Cryptophyllium khmer gen. et sp. nov. bred and photographed by Tim Bollens (Belgium) D Cryptophyllium tibetense comb. nov. E Cryptophyllium oyae comb. nov. F Cryptophyllium westwoodii comb. nov. G Cryptophyllium phami gen. et sp. nov. H Cryptophyllium bollensi gen. et sp. nov. I Cryptophyllium icarus gen. et sp. nov.	Figure 9. Freshly hatched nymphs known for the Cryptophyllium gen. nov., all except for C bred and photographed by Bruno Kneubu ̈ hler (Switzerland) A Cryptophyllium celebicum comb. nov. B Cryptophyllium chrisangi comb. nov. C Cryptophyllium khmer gen. et sp. nov. bred and photographed by Tim Bollens (Belgium) D Cryptophyllium tibetense comb. nov. E Cryptophyllium oyae comb. nov. F Cryptophyllium westwoodii comb. nov. G Cryptophyllium phami gen. et sp. nov. H Cryptophyllium bollensi gen. et sp. nov. I Cryptophyllium icarus gen. et sp. nov.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
74B70D7C21635BE19FE3D1049C04008F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510480	https://doi.org/10.3897/zookeys.1018.61033.figure16	Figure 16. Live Cryptophyllium bollensi gen. et sp. nov. paratype where she was found in Phuoc Binh N. P., Vietnam in July 2014 by Joachim Bresseel (RBINS) and Jerome Constant (RBINS). Photograph by Jerome Constant (RBINS).	Figure 16. Live Cryptophyllium bollensi gen. et sp. nov. paratype where she was found in Phuoc Binh N. P., Vietnam in July 2014 by Joachim Bresseel (RBINS) and Jerome Constant (RBINS). Photograph by Jerome Constant (RBINS).	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
74B70D7C21635BE19FE3D1049C04008F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510481	https://doi.org/10.3897/zookeys.1018.61033.figure17	Figure 17. Live Cryptophyllium bollensi gen. et sp. nov. bred and photographed by Bruno Kneubu ̈ hler (Switzerland) A adult female, dorsal B adult male, dorsal C genitalia, female, ventral.	Figure 17. Live Cryptophyllium bollensi gen. et sp. nov. bred and photographed by Bruno Kneubu ̈ hler (Switzerland) A adult female, dorsal B adult male, dorsal C genitalia, female, ventral.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
74B70D7C21635BE19FE3D1049C04008F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510482	https://doi.org/10.3897/zookeys.1018.61033.figure18	Figure 18. Cryptophyllium bollensi gen. et sp. nov. paratype female, photographs by Jerome Constant (RBINS) A habitus, dorsal B habitus, ventral C details of the antennae, dorsal D front leg details, dorsal E details of the antennae, head, and thorax, dorsal F details of the antennae, head, and thorax, lateral G terminalia, dorsal H genitalia, ventral.	Figure 18. Cryptophyllium bollensi gen. et sp. nov. paratype female, photographs by Jerome Constant (RBINS) A habitus, dorsal B habitus, ventral C details of the antennae, dorsal D front leg details, dorsal E details of the antennae, head, and thorax, dorsal F details of the antennae, head, and thorax, lateral G terminalia, dorsal H genitalia, ventral.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
74B70D7C21635BE19FE3D1049C04008F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510483	https://doi.org/10.3897/zookeys.1018.61033.figure19	Figure 19. Cryptophyllium bollensi gen. et sp. nov. holotype male, photographs by Jerome Constant (RBINS) A habitus, dorsal B habitus, ventral C pro- tibial and femoral lobes, dorsal D details of the base of the antennae, head, and thorax, lateral E details of the antennae, head, and thorax, dorsal F terminalia, dorsal G genitalia, ventral.	Figure 19. Cryptophyllium bollensi gen. et sp. nov. holotype male, photographs by Jerome Constant (RBINS) A habitus, dorsal B habitus, ventral C pro- tibial and femoral lobes, dorsal D details of the base of the antennae, head, and thorax, lateral E details of the antennae, head, and thorax, dorsal F terminalia, dorsal G genitalia, ventral.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
74B70D7C21635BE19FE3D1049C04008F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510484	https://doi.org/10.3897/zookeys.1018.61033.figure20	Figure 20. Abnormal male vomer found in Cryptophyllium bollensi gen. et sp. nov. from Vietnam, Phuoc Binh, observed in the RBINS collection bred from the original limited stock. Prepared and photographed by Jerome Constant (RBINS). Scale bars, 1.0 mm. Views 1 ventral 2 right lateral of the ventrally oriented vomer 3 left lateral of the ventrally oriented vomer A abnormal singularly hooked vomer B abnormal three hooked vomer.	Figure 20. Abnormal male vomer found in Cryptophyllium bollensi gen. et sp. nov. from Vietnam, Phuoc Binh, observed in the RBINS collection bred from the original limited stock. Prepared and photographed by Jerome Constant (RBINS). Scale bars, 1.0 mm. Views 1 ventral 2 right lateral of the ventrally oriented vomer 3 left lateral of the ventrally oriented vomer A abnormal singularly hooked vomer B abnormal three hooked vomer.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
74B70D7C21635BE19FE3D1049C04008F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510485	https://doi.org/10.3897/zookeys.1018.61033.figure21	Figure 21. Cryptophyllium bollensi gen. et sp. nov. egg (RBINS), photographs by Jerome Constant A lateral view B dorso-lateral view C dorsal view D opercular (anterior) view E posterior view F ventral view.	Figure 21. Cryptophyllium bollensi gen. et sp. nov. egg (RBINS), photographs by Jerome Constant A lateral view B dorso-lateral view C dorsal view D opercular (anterior) view E posterior view F ventral view.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
74B70D7C21635BE19FE3D1049C04008F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510468	https://doi.org/10.3897/zookeys.1018.61033.figure4	Figure 4. Phylogenetic relationships among 35 phylliid species (67 specimens). The species of the Cryptophyllium gen. nov. are highlighted with colors and symbols corresponding to the distribution map (Fig. 2). Support values based on maximum likelihood (UFBoot, IQ-TREE) are indicated at each node.	Figure 4. Phylogenetic relationships among 35 phylliid species (67 specimens). The species of the Cryptophyllium gen. nov. are highlighted with colors and symbols corresponding to the distribution map (Fig. 2). Support values based on maximum likelihood (UFBoot, IQ-TREE) are indicated at each node.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
9E1ECF645E4D5FD6AAC328B2E0E41250.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510470	https://doi.org/10.3897/zookeys.1018.61033.figure6	Figure 6. Details of female phylliid antennae illustrated from specimens within the RBINS collection, illustrations by Julien Caudron (RBINS). Note that the figures start with the scapus, therefore the diagnostic fourth antennomere is the third one shown; to ensure clarity A has the segment number listed to the right of the image. Note that the fourth antennomere of Phyllium (A) is longer than the fifth antennomere in comparison to the Cryptophyllium gen. nov. representatives (B-H), which have the fourth antennomere short and disk-like A Phyllium ericoriai, Philippines B Cryptophyllium bollensi gen. et sp. nov., Vietnam, Phuoc Binh C Cryptophyllium celebicum comb. nov., Indonesia, Sulawesi D Cryptophyllium khmer gen. et sp. nov., Cambodia, Siem Reap Province E Cryptophyllium nuichuaense gen. et sp. nov., Vietnam, Nui Chua F Cryptophyllium phami gen. et sp. nov., Vietnam, Cat Tien G Cryptophyllium westwoodii comb. nov., Laos, Bokeo Province H Cryptophyllium westwoodii comb. nov., Thailand, Chiang Mai. Drawn to relative scale.	Figure 6. Details of female phylliid antennae illustrated from specimens within the RBINS collection, illustrations by Julien Caudron (RBINS). Note that the figures start with the scapus, therefore the diagnostic fourth antennomere is the third one shown; to ensure clarity A has the segment number listed to the right of the image. Note that the fourth antennomere of Phyllium (A) is longer than the fifth antennomere in comparison to the Cryptophyllium gen. nov. representatives (B-H), which have the fourth antennomere short and disk-like A Phyllium ericoriai, Philippines B Cryptophyllium bollensi gen. et sp. nov., Vietnam, Phuoc Binh C Cryptophyllium celebicum comb. nov., Indonesia, Sulawesi D Cryptophyllium khmer gen. et sp. nov., Cambodia, Siem Reap Province E Cryptophyllium nuichuaense gen. et sp. nov., Vietnam, Nui Chua F Cryptophyllium phami gen. et sp. nov., Vietnam, Cat Tien G Cryptophyllium westwoodii comb. nov., Laos, Bokeo Province H Cryptophyllium westwoodii comb. nov., Thailand, Chiang Mai. Drawn to relative scale.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
9E1ECF645E4D5FD6AAC328B2E0E41250.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510472	https://doi.org/10.3897/zookeys.1018.61033.figure8	Figure 8. Known eggs for the Cryptophyllium gen. nov. set in couplets of lateral and then dorsal views. All images except for K and L were taken by Bruno Kneubu ̈ hler (Switzerland), K and L images taken by the first author, Q and R images taken by Rene ́ Limoges (IMQC) A, B Cryptophyllium westwoodii comb. nov. C, D Cryptophyllium tibetense comb. nov. E, F Cryptophyllium bollensi gen. et sp. nov. G, H Cryptophyllium oyae comb. nov. I, J Cryptophyllium icarus gen. et sp. nov. K, L Cryptophyllium liyananae gen. et sp. nov. M, N Cryptophyllium chrisangi comb. nov. O, P Cryptophyllium celebicum comb. nov. Q, R Cryptophyllium limogesi gen. et sp. nov. Scale bars 2.0 mm long.	Figure 8. Known eggs for the Cryptophyllium gen. nov. set in couplets of lateral and then dorsal views. All images except for K and L were taken by Bruno Kneubu ̈ hler (Switzerland), K and L images taken by the first author, Q and R images taken by Rene ́ Limoges (IMQC) A, B Cryptophyllium westwoodii comb. nov. C, D Cryptophyllium tibetense comb. nov. E, F Cryptophyllium bollensi gen. et sp. nov. G, H Cryptophyllium oyae comb. nov. I, J Cryptophyllium icarus gen. et sp. nov. K, L Cryptophyllium liyananae gen. et sp. nov. M, N Cryptophyllium chrisangi comb. nov. O, P Cryptophyllium celebicum comb. nov. Q, R Cryptophyllium limogesi gen. et sp. nov. Scale bars 2.0 mm long.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
9E1ECF645E4D5FD6AAC328B2E0E41250.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510473	https://doi.org/10.3897/zookeys.1018.61033.figure9	Figure 9. Freshly hatched nymphs known for the Cryptophyllium gen. nov., all except for C bred and photographed by Bruno Kneubu ̈ hler (Switzerland) A Cryptophyllium celebicum comb. nov. B Cryptophyllium chrisangi comb. nov. C Cryptophyllium khmer gen. et sp. nov. bred and photographed by Tim Bollens (Belgium) D Cryptophyllium tibetense comb. nov. E Cryptophyllium oyae comb. nov. F Cryptophyllium westwoodii comb. nov. G Cryptophyllium phami gen. et sp. nov. H Cryptophyllium bollensi gen. et sp. nov. I Cryptophyllium icarus gen. et sp. nov.	Figure 9. Freshly hatched nymphs known for the Cryptophyllium gen. nov., all except for C bred and photographed by Bruno Kneubu ̈ hler (Switzerland) A Cryptophyllium celebicum comb. nov. B Cryptophyllium chrisangi comb. nov. C Cryptophyllium khmer gen. et sp. nov. bred and photographed by Tim Bollens (Belgium) D Cryptophyllium tibetense comb. nov. E Cryptophyllium oyae comb. nov. F Cryptophyllium westwoodii comb. nov. G Cryptophyllium phami gen. et sp. nov. H Cryptophyllium bollensi gen. et sp. nov. I Cryptophyllium icarus gen. et sp. nov.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
9E1ECF645E4D5FD6AAC328B2E0E41250.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510486	https://doi.org/10.3897/zookeys.1018.61033.figure22	Figure 22. Holotype, Cryptophyllium celebicum (de Haan, 1842), comb. nov. the type species for the Cryptophyllium gen. nov. Photographs by Luc Willemse, Naturalis Biodiversity Center (RMNH) A dorsal, habitus B genitalia, ventral C ventral, habitus, and original collection label inset to left D details of the front legs, head, and thorax, dorsal E details of the antennae and thorax, dorsal.	Figure 22. Holotype, Cryptophyllium celebicum (de Haan, 1842), comb. nov. the type species for the Cryptophyllium gen. nov. Photographs by Luc Willemse, Naturalis Biodiversity Center (RMNH) A dorsal, habitus B genitalia, ventral C ventral, habitus, and original collection label inset to left D details of the front legs, head, and thorax, dorsal E details of the antennae and thorax, dorsal.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
9E1ECF645E4D5FD6AAC328B2E0E41250.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510487	https://doi.org/10.3897/zookeys.1018.61033.figure23	Figure 23. Live Cryptophyllium celebicum comb. nov. A green form female, dorsal, bred and photographed by Thomas Stijnts (Belgium; Flanders) B-D bred and photographed by Bruno Kneubu ̈ hler (Switzerland) B orange form adult female, dorsal C lateral view of the female genitalia holding an egg ready to be flicked away. Note the large gonapophyses VIII and the smaller gonapophyses IX holding the egg. D orange form adult male, dorsal.	Figure 23. Live Cryptophyllium celebicum comb. nov. A green form female, dorsal, bred and photographed by Thomas Stijnts (Belgium; Flanders) B-D bred and photographed by Bruno Kneubu ̈ hler (Switzerland) B orange form adult female, dorsal C lateral view of the female genitalia holding an egg ready to be flicked away. Note the large gonapophyses VIII and the smaller gonapophyses IX holding the egg. D orange form adult male, dorsal.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
9E1ECF645E4D5FD6AAC328B2E0E41250.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510468	https://doi.org/10.3897/zookeys.1018.61033.figure4	Figure 4. Phylogenetic relationships among 35 phylliid species (67 specimens). The species of the Cryptophyllium gen. nov. are highlighted with colors and symbols corresponding to the distribution map (Fig. 2). Support values based on maximum likelihood (UFBoot, IQ-TREE) are indicated at each node.	Figure 4. Phylogenetic relationships among 35 phylliid species (67 specimens). The species of the Cryptophyllium gen. nov. are highlighted with colors and symbols corresponding to the distribution map (Fig. 2). Support values based on maximum likelihood (UFBoot, IQ-TREE) are indicated at each node.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
9E1ECF645E4D5FD6AAC328B2E0E41250.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510506	https://doi.org/10.3897/zookeys.1018.61033.figure42	Figure 42. Holotype female Cryptophyllium limogesi, photographs by Rene Limoges (IMQC) A habitus, dorsal B habitus, ventral C head and antennae dorsal D antennae detail dorsal E thorax dorsal F front leg, dorsal G terminalia, dorsal H genitalia, ventral.	Figure 42. Holotype female Cryptophyllium limogesi, photographs by Rene Limoges (IMQC) A habitus, dorsal B habitus, ventral C head and antennae dorsal D antennae detail dorsal E thorax dorsal F front leg, dorsal G terminalia, dorsal H genitalia, ventral.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
9E1ECF645E4D5FD6AAC328B2E0E41250.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510539	https://doi.org/10.3897/zookeys.1018.61033.figure75	Figure 75. Male Cryptophyllium yunnanense comb. nov. (Coll RC 16 - 120), photographed by RTC A habitus, dorsal B left front leg details C head-thorax details D genitalia details, ventral E head-thorax details, lateral.	Figure 75. Male Cryptophyllium yunnanense comb. nov. (Coll RC 16 - 120), photographed by RTC A habitus, dorsal B left front leg details C head-thorax details D genitalia details, ventral E head-thorax details, lateral.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
9E1ECF645E4D5FD6AAC328B2E0E41250.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510514	https://doi.org/10.3897/zookeys.1018.61033.figure50	Figure 50. Live Cryptophyllium oyae comb. nov. bred and photographed by Bruno Kneubu ̈ hler (Switzerland) A female B male C lateral view of female with tegmina held up to expose the alae D female genitalia, ventral E male genitalia, ventral.	Figure 50. Live Cryptophyllium oyae comb. nov. bred and photographed by Bruno Kneubu ̈ hler (Switzerland) A female B male C lateral view of female with tegmina held up to expose the alae D female genitalia, ventral E male genitalia, ventral.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
A16BAF5141C05232A40975226EEEBA20.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510472	https://doi.org/10.3897/zookeys.1018.61033.figure8	Figure 8. Known eggs for the Cryptophyllium gen. nov. set in couplets of lateral and then dorsal views. All images except for K and L were taken by Bruno Kneubu ̈ hler (Switzerland), K and L images taken by the first author, Q and R images taken by Rene ́ Limoges (IMQC) A, B Cryptophyllium westwoodii comb. nov. C, D Cryptophyllium tibetense comb. nov. E, F Cryptophyllium bollensi gen. et sp. nov. G, H Cryptophyllium oyae comb. nov. I, J Cryptophyllium icarus gen. et sp. nov. K, L Cryptophyllium liyananae gen. et sp. nov. M, N Cryptophyllium chrisangi comb. nov. O, P Cryptophyllium celebicum comb. nov. Q, R Cryptophyllium limogesi gen. et sp. nov. Scale bars 2.0 mm long.	Figure 8. Known eggs for the Cryptophyllium gen. nov. set in couplets of lateral and then dorsal views. All images except for K and L were taken by Bruno Kneubu ̈ hler (Switzerland), K and L images taken by the first author, Q and R images taken by Rene ́ Limoges (IMQC) A, B Cryptophyllium westwoodii comb. nov. C, D Cryptophyllium tibetense comb. nov. E, F Cryptophyllium bollensi gen. et sp. nov. G, H Cryptophyllium oyae comb. nov. I, J Cryptophyllium icarus gen. et sp. nov. K, L Cryptophyllium liyananae gen. et sp. nov. M, N Cryptophyllium chrisangi comb. nov. O, P Cryptophyllium celebicum comb. nov. Q, R Cryptophyllium limogesi gen. et sp. nov. Scale bars 2.0 mm long.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
A16BAF5141C05232A40975226EEEBA20.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510473	https://doi.org/10.3897/zookeys.1018.61033.figure9	Figure 9. Freshly hatched nymphs known for the Cryptophyllium gen. nov., all except for C bred and photographed by Bruno Kneubu ̈ hler (Switzerland) A Cryptophyllium celebicum comb. nov. B Cryptophyllium chrisangi comb. nov. C Cryptophyllium khmer gen. et sp. nov. bred and photographed by Tim Bollens (Belgium) D Cryptophyllium tibetense comb. nov. E Cryptophyllium oyae comb. nov. F Cryptophyllium westwoodii comb. nov. G Cryptophyllium phami gen. et sp. nov. H Cryptophyllium bollensi gen. et sp. nov. I Cryptophyllium icarus gen. et sp. nov.	Figure 9. Freshly hatched nymphs known for the Cryptophyllium gen. nov., all except for C bred and photographed by Bruno Kneubu ̈ hler (Switzerland) A Cryptophyllium celebicum comb. nov. B Cryptophyllium chrisangi comb. nov. C Cryptophyllium khmer gen. et sp. nov. bred and photographed by Tim Bollens (Belgium) D Cryptophyllium tibetense comb. nov. E Cryptophyllium oyae comb. nov. F Cryptophyllium westwoodii comb. nov. G Cryptophyllium phami gen. et sp. nov. H Cryptophyllium bollensi gen. et sp. nov. I Cryptophyllium icarus gen. et sp. nov.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
A16BAF5141C05232A40975226EEEBA20.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510488	https://doi.org/10.3897/zookeys.1018.61033.figure24	Figure 24. Live Cryptophyllium chrisangi comb. nov. A female bred and photographed by Bruno Kneubu ̈ hler (Switzerland) B male genitalia, ventral view, bred by Bruno Kneubu ̈ hler (Switzerland) C male observed and photographed by Mathieu MJP Van Goethem (South Africa) on Weh Island off the north shore of Sumatra D female observed and photographed in Thailand: Nakhon Si Thammarat Province, Thung Song District, by Tatsatorn Dharithai (Thailand) in August 2020.	Figure 24. Live Cryptophyllium chrisangi comb. nov. A female bred and photographed by Bruno Kneubu ̈ hler (Switzerland) B male genitalia, ventral view, bred by Bruno Kneubu ̈ hler (Switzerland) C male observed and photographed by Mathieu MJP Van Goethem (South Africa) on Weh Island off the north shore of Sumatra D female observed and photographed in Thailand: Nakhon Si Thammarat Province, Thung Song District, by Tatsatorn Dharithai (Thailand) in August 2020.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
A16BAF5141C05232A40975226EEEBA20.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510489	https://doi.org/10.3897/zookeys.1018.61033.figure25	Figure 25. Wood-Mason's 1875 plate XVII with the beautifully illustrated ' Phyllium westwoodii ' female syntype from South Andaman. Note the morphological similarity to Cryptophyllium chrisangi comb. nov. and given the wide range of Cryptophyllium chrisangi comb. nov. outside of the type locality of Singapore, this record may represent a range expansion. Public domain work downloaded from the Biodiversity Heritage Library (https: // www. biodiversitylibrary. org / item / 114410  page / 227 / mode / 1 up)	Figure 25. Wood-Mason's 1875 plate XVII with the beautifully illustrated ' Phyllium westwoodii ' female syntype from South Andaman. Note the morphological similarity to Cryptophyllium chrisangi comb. nov. and given the wide range of Cryptophyllium chrisangi comb. nov. outside of the type locality of Singapore, this record may represent a range expansion. Public domain work downloaded from the Biodiversity Heritage Library (https: // www. biodiversitylibrary. org / item / 114410  page / 227 / mode / 1 up)	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
A16BAF5141C05232A40975226EEEBA20.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510490	https://doi.org/10.3897/zookeys.1018.61033.figure26	Figure 26. Tentative female Cryptophyllium chrisangi comb. nov. from Kuching, Sarawak, Malaysia which represents a range expansion for this species, photographs by Jerome Constant (RBINS) A habitus, dorsal B details of the anterior of the head, dorsal C habitus, ventral D front leg details, dorsal E details of the head and thorax, dorsal F details of the head and thorax, lateral G genitalia details, dorsal H genitalia details, ventral I data label.	Figure 26. Tentative female Cryptophyllium chrisangi comb. nov. from Kuching, Sarawak, Malaysia which represents a range expansion for this species, photographs by Jerome Constant (RBINS) A habitus, dorsal B details of the anterior of the head, dorsal C habitus, ventral D front leg details, dorsal E details of the head and thorax, dorsal F details of the head and thorax, lateral G genitalia details, dorsal H genitalia details, ventral I data label.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
A16BAF5141C05232A40975226EEEBA20.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510466	https://doi.org/10.3897/zookeys.1018.61033.figure2	Figure 2. Distribution map for the 24 Cryptophyllium gen. nov. species presently known (with solid lines from their name pointing to the type locality) as well as additional Cryptophyllium gen. nov. species which we could not herein describe / differentiate (indicated by dashed lines). Note that the line for Cryptophyllium westwoodii comb. nov. is pointing to the neotype locality and the type locality for Cryptophyllium athanysus comb. nov. is simply " Ceylon " therefore the line is pointing to the present-day localities we are aware of. Inset is of southern Vietnam showing the distributions of three additional species which could not fit within the main map. The colors in this map are noted to the left of the names within the phylogenetic tree in Fig. 4. Newly herein described species have names noted in bold. Note that with Cryptophyllium khmer sp. nov. not easily distinguishable from Cryptophyllium westwoodii comb. nov. from photos alone, only the locations for these two species where they were genetically sampled are solid colored, all observational images without genetic sampling have bicolored circles and could represent either of these species. Additionally, one symbol is split three ways for Cryptophyllium gen. nov. specimens from the Thai offshore islands of Ko Phangan and Ko Samui which could not be differentiated from C. chrisangi comb. nov., C. westwoodii comb. nov., and C. khmer sp. nov. from images alone. See Suppl. material 4 for a full list of the specimens / observations utilized to make the distribution map with deposition data for specimens and links to observational records>.	Figure 2. Distribution map for the 24 Cryptophyllium gen. nov. species presently known (with solid lines from their name pointing to the type locality) as well as additional Cryptophyllium gen. nov. species which we could not herein describe / differentiate (indicated by dashed lines). Note that the line for Cryptophyllium westwoodii comb. nov. is pointing to the neotype locality and the type locality for Cryptophyllium athanysus comb. nov. is simply " Ceylon " therefore the line is pointing to the present-day localities we are aware of. Inset is of southern Vietnam showing the distributions of three additional species which could not fit within the main map. The colors in this map are noted to the left of the names within the phylogenetic tree in Fig. 4. Newly herein described species have names noted in bold. Note that with Cryptophyllium khmer sp. nov. not easily distinguishable from Cryptophyllium westwoodii comb. nov. from photos alone, only the locations for these two species where they were genetically sampled are solid colored, all observational images without genetic sampling have bicolored circles and could represent either of these species. Additionally, one symbol is split three ways for Cryptophyllium gen. nov. specimens from the Thai offshore islands of Ko Phangan and Ko Samui which could not be differentiated from C. chrisangi comb. nov., C. westwoodii comb. nov., and C. khmer sp. nov. from images alone. See Suppl. material 4 for a full list of the specimens / observations utilized to make the distribution map with deposition data for specimens and links to observational records>.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
A16BAF5141C05232A40975226EEEBA20.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510532	https://doi.org/10.3897/zookeys.1018.61033.figure68	Figure 68. Live adult Cryptophyllium westwoodii comb. nov. bred and photographed by Bruno Kneubu ̈ hler (Switzerland) from stock collected in Tha Pla Duk subdistrict, Mae Tha district, Lamphun, Thailand, collected in May / June 2014 by Suttah Ek-Amnuay (Thailand) A female habitus B male habitus C female details of the thorax, dorsal D lateral view female with tegmina lifted to show long alae E female genitalia details, ventral.	Figure 68. Live adult Cryptophyllium westwoodii comb. nov. bred and photographed by Bruno Kneubu ̈ hler (Switzerland) from stock collected in Tha Pla Duk subdistrict, Mae Tha district, Lamphun, Thailand, collected in May / June 2014 by Suttah Ek-Amnuay (Thailand) A female habitus B male habitus C female details of the thorax, dorsal D lateral view female with tegmina lifted to show long alae E female genitalia details, ventral.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
4AF24F807CB058BD9C2BDB85EC87C7C1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510469	https://doi.org/10.3897/zookeys.1018.61033.figure5	Figure 5. Details of the Cryptophyllium gen. nov. male vomer. All from the RBINS collection, prepared and photographed by Jerome Constant (RBINS). Views 1 ventral 2 right lateral of the ventrally oriented vomer 3 left lateral of the ventrally oriented vomer A Cryptophyllium westwoodii comb. nov., Thailand, Na Haeo B Cryptophyllium khmer gen. et sp. nov. Cambodia, Kbal Spean C Cryptophyllium phami gen. et sp. nov., Vietnam, Cat Tien D Cryptophyllium khmer gen. et sp. nov., Cambodia, Tatai E Cryptophyllium bollensi gen. et sp. nov., Vietnam, Phuoc Binh F Cryptophyllium oyae comb. nov., Laos, Mt. Phu Phan G Cryptophyllium bankoi gen. et sp. nov., Vietnam, Ba Na-Nui Chua H Cryptophyllium icarus gen. et sp. nov., Vietnam, Bidoup-Nui Ba I Cryptophyllium rarum comb. nov., Vietnam, Cuc Phuong J Cryptophyllium limogesi gen. et sp. nov., Vietnam, Ta Dung. Scale bars, 1.0 mm.	Figure 5. Details of the Cryptophyllium gen. nov. male vomer. All from the RBINS collection, prepared and photographed by Jerome Constant (RBINS). Views 1 ventral 2 right lateral of the ventrally oriented vomer 3 left lateral of the ventrally oriented vomer A Cryptophyllium westwoodii comb. nov., Thailand, Na Haeo B Cryptophyllium khmer gen. et sp. nov. Cambodia, Kbal Spean C Cryptophyllium phami gen. et sp. nov., Vietnam, Cat Tien D Cryptophyllium khmer gen. et sp. nov., Cambodia, Tatai E Cryptophyllium bollensi gen. et sp. nov., Vietnam, Phuoc Binh F Cryptophyllium oyae comb. nov., Laos, Mt. Phu Phan G Cryptophyllium bankoi gen. et sp. nov., Vietnam, Ba Na-Nui Chua H Cryptophyllium icarus gen. et sp. nov., Vietnam, Bidoup-Nui Ba I Cryptophyllium rarum comb. nov., Vietnam, Cuc Phuong J Cryptophyllium limogesi gen. et sp. nov., Vietnam, Ta Dung. Scale bars, 1.0 mm.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
DB5E76716257505885921E4B977E7CF1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510491	https://doi.org/10.3897/zookeys.1018.61033.figure27	Figure 27. Holotype female Cryptophyllium daparo gen. et sp. nov., photographs by Zhiwei Dong (KIZ) A habitus, dorsal B details of the face and antennae, rostral view C profemoral lobes, dorsal D details of the head and thorax, dorsal E terminalia, dorsal F details of the head and thorax, lateral G genitalia, ventral.	Figure 27. Holotype female Cryptophyllium daparo gen. et sp. nov., photographs by Zhiwei Dong (KIZ) A habitus, dorsal B details of the face and antennae, rostral view C profemoral lobes, dorsal D details of the head and thorax, dorsal E terminalia, dorsal F details of the head and thorax, lateral G genitalia, ventral.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
37ECB48AD86050F7AB814CAADB108F15.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510492	https://doi.org/10.3897/zookeys.1018.61033.figure28	Figure 28. Cryptophyllium drunganum comb. nov. holotype female within the Beijing Agricultural University, photographed by Yu-Chen Zheng (China Agricultural University, China) A habitus, dorsal, inset specimen data labels B details of antennae, head, and anterior of the thorax.	Figure 28. Cryptophyllium drunganum comb. nov. holotype female within the Beijing Agricultural University, photographed by Yu-Chen Zheng (China Agricultural University, China) A habitus, dorsal, inset specimen data labels B details of antennae, head, and anterior of the thorax.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
37ECB48AD86050F7AB814CAADB108F15.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510493	https://doi.org/10.3897/zookeys.1018.61033.figure29	Figure 29. Presumed male Cryptophyllium drunganum comb. nov. male used within our molecular analysis (sample DZW 05), from Du Longjiang Township, Nujian Lisu Autonomous Prefecture, photographs by Zhiwei Dong (KIZ) A habitus, dorsal B details of the legs and thorax, ventral.	Figure 29. Presumed male Cryptophyllium drunganum comb. nov. male used within our molecular analysis (sample DZW 05), from Du Longjiang Township, Nujian Lisu Autonomous Prefecture, photographs by Zhiwei Dong (KIZ) A habitus, dorsal B details of the legs and thorax, ventral.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
37ECB48AD86050F7AB814CAADB108F15.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510468	https://doi.org/10.3897/zookeys.1018.61033.figure4	Figure 4. Phylogenetic relationships among 35 phylliid species (67 specimens). The species of the Cryptophyllium gen. nov. are highlighted with colors and symbols corresponding to the distribution map (Fig. 2). Support values based on maximum likelihood (UFBoot, IQ-TREE) are indicated at each node.	Figure 4. Phylogenetic relationships among 35 phylliid species (67 specimens). The species of the Cryptophyllium gen. nov. are highlighted with colors and symbols corresponding to the distribution map (Fig. 2). Support values based on maximum likelihood (UFBoot, IQ-TREE) are indicated at each node.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
B7FA223E233555279DF4863A17B08329.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510494	https://doi.org/10.3897/zookeys.1018.61033.figure30	Figure 30. Cryptophyllium echidna gen. et sp. nov. holotype female, photographs by Rene ́ Limoges (IMQC) A habitus, dorsal B dorsal details of the antennae C habitus, ventral D dorsal details of the thorax E ventral details of the genitalia.	Figure 30. Cryptophyllium echidna gen. et sp. nov. holotype female, photographs by Rene ́ Limoges (IMQC) A habitus, dorsal B dorsal details of the antennae C habitus, ventral D dorsal details of the thorax E ventral details of the genitalia.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
B7FA223E233555279DF4863A17B08329.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510466	https://doi.org/10.3897/zookeys.1018.61033.figure2	Figure 2. Distribution map for the 24 Cryptophyllium gen. nov. species presently known (with solid lines from their name pointing to the type locality) as well as additional Cryptophyllium gen. nov. species which we could not herein describe / differentiate (indicated by dashed lines). Note that the line for Cryptophyllium westwoodii comb. nov. is pointing to the neotype locality and the type locality for Cryptophyllium athanysus comb. nov. is simply " Ceylon " therefore the line is pointing to the present-day localities we are aware of. Inset is of southern Vietnam showing the distributions of three additional species which could not fit within the main map. The colors in this map are noted to the left of the names within the phylogenetic tree in Fig. 4. Newly herein described species have names noted in bold. Note that with Cryptophyllium khmer sp. nov. not easily distinguishable from Cryptophyllium westwoodii comb. nov. from photos alone, only the locations for these two species where they were genetically sampled are solid colored, all observational images without genetic sampling have bicolored circles and could represent either of these species. Additionally, one symbol is split three ways for Cryptophyllium gen. nov. specimens from the Thai offshore islands of Ko Phangan and Ko Samui which could not be differentiated from C. chrisangi comb. nov., C. westwoodii comb. nov., and C. khmer sp. nov. from images alone. See Suppl. material 4 for a full list of the specimens / observations utilized to make the distribution map with deposition data for specimens and links to observational records>.	Figure 2. Distribution map for the 24 Cryptophyllium gen. nov. species presently known (with solid lines from their name pointing to the type locality) as well as additional Cryptophyllium gen. nov. species which we could not herein describe / differentiate (indicated by dashed lines). Note that the line for Cryptophyllium westwoodii comb. nov. is pointing to the neotype locality and the type locality for Cryptophyllium athanysus comb. nov. is simply " Ceylon " therefore the line is pointing to the present-day localities we are aware of. Inset is of southern Vietnam showing the distributions of three additional species which could not fit within the main map. The colors in this map are noted to the left of the names within the phylogenetic tree in Fig. 4. Newly herein described species have names noted in bold. Note that with Cryptophyllium khmer sp. nov. not easily distinguishable from Cryptophyllium westwoodii comb. nov. from photos alone, only the locations for these two species where they were genetically sampled are solid colored, all observational images without genetic sampling have bicolored circles and could represent either of these species. Additionally, one symbol is split three ways for Cryptophyllium gen. nov. specimens from the Thai offshore islands of Ko Phangan and Ko Samui which could not be differentiated from C. chrisangi comb. nov., C. westwoodii comb. nov., and C. khmer sp. nov. from images alone. See Suppl. material 4 for a full list of the specimens / observations utilized to make the distribution map with deposition data for specimens and links to observational records>.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
B7FA223E233555279DF4863A17B08329.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510486	https://doi.org/10.3897/zookeys.1018.61033.figure22	Figure 22. Holotype, Cryptophyllium celebicum (de Haan, 1842), comb. nov. the type species for the Cryptophyllium gen. nov. Photographs by Luc Willemse, Naturalis Biodiversity Center (RMNH) A dorsal, habitus B genitalia, ventral C ventral, habitus, and original collection label inset to left D details of the front legs, head, and thorax, dorsal E details of the antennae and thorax, dorsal.	Figure 22. Holotype, Cryptophyllium celebicum (de Haan, 1842), comb. nov. the type species for the Cryptophyllium gen. nov. Photographs by Luc Willemse, Naturalis Biodiversity Center (RMNH) A dorsal, habitus B genitalia, ventral C ventral, habitus, and original collection label inset to left D details of the front legs, head, and thorax, dorsal E details of the antennae and thorax, dorsal.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
C8E17D58FF8F562DAD931973898C355E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510495	https://doi.org/10.3897/zookeys.1018.61033.figure31	Figure 31. Holotype Cryptophyllium faulkneri gen. et sp. nov., photographs by RTC A details of the base of antennae, head, and thorax, dorsal B habitus, dorsal C details of thorax, ventral D details of head and thorax, lateral.	Figure 31. Holotype Cryptophyllium faulkneri gen. et sp. nov., photographs by RTC A details of the base of antennae, head, and thorax, dorsal B habitus, dorsal C details of thorax, ventral D details of head and thorax, lateral.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
C8E17D58FF8F562DAD931973898C355E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510474	https://doi.org/10.3897/zookeys.1018.61033.figure10	Figure 10. Holotype Cryptophyllium animatum gen. et sp. nov., photographs by RTC A habitus, dorsal B details of the protibial and profemoral lobes C details of the head and thorax, dorsal D genitalia, ventral E head-thorax, lateral.	Figure 10. Holotype Cryptophyllium animatum gen. et sp. nov., photographs by RTC A habitus, dorsal B details of the protibial and profemoral lobes C details of the head and thorax, dorsal D genitalia, ventral E head-thorax, lateral.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
875D19DBF6005EC786793DEA37E816C1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510469	https://doi.org/10.3897/zookeys.1018.61033.figure5	Figure 5. Details of the Cryptophyllium gen. nov. male vomer. All from the RBINS collection, prepared and photographed by Jerome Constant (RBINS). Views 1 ventral 2 right lateral of the ventrally oriented vomer 3 left lateral of the ventrally oriented vomer A Cryptophyllium westwoodii comb. nov., Thailand, Na Haeo B Cryptophyllium khmer gen. et sp. nov. Cambodia, Kbal Spean C Cryptophyllium phami gen. et sp. nov., Vietnam, Cat Tien D Cryptophyllium khmer gen. et sp. nov., Cambodia, Tatai E Cryptophyllium bollensi gen. et sp. nov., Vietnam, Phuoc Binh F Cryptophyllium oyae comb. nov., Laos, Mt. Phu Phan G Cryptophyllium bankoi gen. et sp. nov., Vietnam, Ba Na-Nui Chua H Cryptophyllium icarus gen. et sp. nov., Vietnam, Bidoup-Nui Ba I Cryptophyllium rarum comb. nov., Vietnam, Cuc Phuong J Cryptophyllium limogesi gen. et sp. nov., Vietnam, Ta Dung. Scale bars, 1.0 mm.	Figure 5. Details of the Cryptophyllium gen. nov. male vomer. All from the RBINS collection, prepared and photographed by Jerome Constant (RBINS). Views 1 ventral 2 right lateral of the ventrally oriented vomer 3 left lateral of the ventrally oriented vomer A Cryptophyllium westwoodii comb. nov., Thailand, Na Haeo B Cryptophyllium khmer gen. et sp. nov. Cambodia, Kbal Spean C Cryptophyllium phami gen. et sp. nov., Vietnam, Cat Tien D Cryptophyllium khmer gen. et sp. nov., Cambodia, Tatai E Cryptophyllium bollensi gen. et sp. nov., Vietnam, Phuoc Binh F Cryptophyllium oyae comb. nov., Laos, Mt. Phu Phan G Cryptophyllium bankoi gen. et sp. nov., Vietnam, Ba Na-Nui Chua H Cryptophyllium icarus gen. et sp. nov., Vietnam, Bidoup-Nui Ba I Cryptophyllium rarum comb. nov., Vietnam, Cuc Phuong J Cryptophyllium limogesi gen. et sp. nov., Vietnam, Ta Dung. Scale bars, 1.0 mm.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
875D19DBF6005EC786793DEA37E816C1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510472	https://doi.org/10.3897/zookeys.1018.61033.figure8	Figure 8. Known eggs for the Cryptophyllium gen. nov. set in couplets of lateral and then dorsal views. All images except for K and L were taken by Bruno Kneubu ̈ hler (Switzerland), K and L images taken by the first author, Q and R images taken by Rene ́ Limoges (IMQC) A, B Cryptophyllium westwoodii comb. nov. C, D Cryptophyllium tibetense comb. nov. E, F Cryptophyllium bollensi gen. et sp. nov. G, H Cryptophyllium oyae comb. nov. I, J Cryptophyllium icarus gen. et sp. nov. K, L Cryptophyllium liyananae gen. et sp. nov. M, N Cryptophyllium chrisangi comb. nov. O, P Cryptophyllium celebicum comb. nov. Q, R Cryptophyllium limogesi gen. et sp. nov. Scale bars 2.0 mm long.	Figure 8. Known eggs for the Cryptophyllium gen. nov. set in couplets of lateral and then dorsal views. All images except for K and L were taken by Bruno Kneubu ̈ hler (Switzerland), K and L images taken by the first author, Q and R images taken by Rene ́ Limoges (IMQC) A, B Cryptophyllium westwoodii comb. nov. C, D Cryptophyllium tibetense comb. nov. E, F Cryptophyllium bollensi gen. et sp. nov. G, H Cryptophyllium oyae comb. nov. I, J Cryptophyllium icarus gen. et sp. nov. K, L Cryptophyllium liyananae gen. et sp. nov. M, N Cryptophyllium chrisangi comb. nov. O, P Cryptophyllium celebicum comb. nov. Q, R Cryptophyllium limogesi gen. et sp. nov. Scale bars 2.0 mm long.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
875D19DBF6005EC786793DEA37E816C1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510473	https://doi.org/10.3897/zookeys.1018.61033.figure9	Figure 9. Freshly hatched nymphs known for the Cryptophyllium gen. nov., all except for C bred and photographed by Bruno Kneubu ̈ hler (Switzerland) A Cryptophyllium celebicum comb. nov. B Cryptophyllium chrisangi comb. nov. C Cryptophyllium khmer gen. et sp. nov. bred and photographed by Tim Bollens (Belgium) D Cryptophyllium tibetense comb. nov. E Cryptophyllium oyae comb. nov. F Cryptophyllium westwoodii comb. nov. G Cryptophyllium phami gen. et sp. nov. H Cryptophyllium bollensi gen. et sp. nov. I Cryptophyllium icarus gen. et sp. nov.	Figure 9. Freshly hatched nymphs known for the Cryptophyllium gen. nov., all except for C bred and photographed by Bruno Kneubu ̈ hler (Switzerland) A Cryptophyllium celebicum comb. nov. B Cryptophyllium chrisangi comb. nov. C Cryptophyllium khmer gen. et sp. nov. bred and photographed by Tim Bollens (Belgium) D Cryptophyllium tibetense comb. nov. E Cryptophyllium oyae comb. nov. F Cryptophyllium westwoodii comb. nov. G Cryptophyllium phami gen. et sp. nov. H Cryptophyllium bollensi gen. et sp. nov. I Cryptophyllium icarus gen. et sp. nov.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
875D19DBF6005EC786793DEA37E816C1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510496	https://doi.org/10.3897/zookeys.1018.61033.figure32	Figure 32. Live Cryptophyllium icarus found in July 2014 by Joachim Bresseel (RBINS) and Jerome Constant (RBINS) in Bidoup Nui Ba N. P., Vietnam. Photographs by Jerome Constant (RBINS) A paratype female, dorsal closeup B male nymph, paratype C same female as in A but showing the host tree she was found on D disturbed area along the road where the original stock was found with arrows pointing out host trees.	Figure 32. Live Cryptophyllium icarus found in July 2014 by Joachim Bresseel (RBINS) and Jerome Constant (RBINS) in Bidoup Nui Ba N. P., Vietnam. Photographs by Jerome Constant (RBINS) A paratype female, dorsal closeup B male nymph, paratype C same female as in A but showing the host tree she was found on D disturbed area along the road where the original stock was found with arrows pointing out host trees.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
875D19DBF6005EC786793DEA37E816C1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510497	https://doi.org/10.3897/zookeys.1018.61033.figure33	Figure 33. Live adult Cryptophyllium icarus bred and photographed by Bruno Kneubu ̈ hler (Switzerland) A female B male C lateral view of female with tegmina held up to expose the dwarfed alae underneath.	Figure 33. Live adult Cryptophyllium icarus bred and photographed by Bruno Kneubu ̈ hler (Switzerland) A female B male C lateral view of female with tegmina held up to expose the dwarfed alae underneath.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
875D19DBF6005EC786793DEA37E816C1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510498	https://doi.org/10.3897/zookeys.1018.61033.figure34	Figure 34. Cryptophyllium icarus gen. et sp. nov. female paratype, photographs by Jerome Constant (RBINS) A habitus, dorsal B habitus, ventral C antennae and anterior of head capsule D profemoral and protibial lobes E head and thorax details dorsal F head and thorax details lateral G genitalia dorsal H genitalia ventral.	Figure 34. Cryptophyllium icarus gen. et sp. nov. female paratype, photographs by Jerome Constant (RBINS) A habitus, dorsal B habitus, ventral C antennae and anterior of head capsule D profemoral and protibial lobes E head and thorax details dorsal F head and thorax details lateral G genitalia dorsal H genitalia ventral.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
875D19DBF6005EC786793DEA37E816C1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510499	https://doi.org/10.3897/zookeys.1018.61033.figure35	Figure 35. Cryptophyllium icarus gen. et sp. nov. male holotype, photographs by Jerome Constant (RBINS) A habitus, dorsal B habitus, ventral C profemoral and protibial lobes D head and thorax details dorsal E head and thorax details lateral F genitalia dorsal G genitalia ventral.	Figure 35. Cryptophyllium icarus gen. et sp. nov. male holotype, photographs by Jerome Constant (RBINS) A habitus, dorsal B habitus, ventral C profemoral and protibial lobes D head and thorax details dorsal E head and thorax details lateral F genitalia dorsal G genitalia ventral.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
875D19DBF6005EC786793DEA37E816C1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510500	https://doi.org/10.3897/zookeys.1018.61033.figure36	Figure 36. Cryptophyllium icarus gen. et sp. nov. egg, RBINS collection, photographs by Jerome Constant A lateral view B dorso-lateral view C dorsal view D opercular (anterior) view E posterior view F ventral view.	Figure 36. Cryptophyllium icarus gen. et sp. nov. egg, RBINS collection, photographs by Jerome Constant A lateral view B dorso-lateral view C dorsal view D opercular (anterior) view E posterior view F ventral view.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
7F2F01FA305959E39E1EC7B005EBDF4C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510469	https://doi.org/10.3897/zookeys.1018.61033.figure5	Figure 5. Details of the Cryptophyllium gen. nov. male vomer. All from the RBINS collection, prepared and photographed by Jerome Constant (RBINS). Views 1 ventral 2 right lateral of the ventrally oriented vomer 3 left lateral of the ventrally oriented vomer A Cryptophyllium westwoodii comb. nov., Thailand, Na Haeo B Cryptophyllium khmer gen. et sp. nov. Cambodia, Kbal Spean C Cryptophyllium phami gen. et sp. nov., Vietnam, Cat Tien D Cryptophyllium khmer gen. et sp. nov., Cambodia, Tatai E Cryptophyllium bollensi gen. et sp. nov., Vietnam, Phuoc Binh F Cryptophyllium oyae comb. nov., Laos, Mt. Phu Phan G Cryptophyllium bankoi gen. et sp. nov., Vietnam, Ba Na-Nui Chua H Cryptophyllium icarus gen. et sp. nov., Vietnam, Bidoup-Nui Ba I Cryptophyllium rarum comb. nov., Vietnam, Cuc Phuong J Cryptophyllium limogesi gen. et sp. nov., Vietnam, Ta Dung. Scale bars, 1.0 mm.	Figure 5. Details of the Cryptophyllium gen. nov. male vomer. All from the RBINS collection, prepared and photographed by Jerome Constant (RBINS). Views 1 ventral 2 right lateral of the ventrally oriented vomer 3 left lateral of the ventrally oriented vomer A Cryptophyllium westwoodii comb. nov., Thailand, Na Haeo B Cryptophyllium khmer gen. et sp. nov. Cambodia, Kbal Spean C Cryptophyllium phami gen. et sp. nov., Vietnam, Cat Tien D Cryptophyllium khmer gen. et sp. nov., Cambodia, Tatai E Cryptophyllium bollensi gen. et sp. nov., Vietnam, Phuoc Binh F Cryptophyllium oyae comb. nov., Laos, Mt. Phu Phan G Cryptophyllium bankoi gen. et sp. nov., Vietnam, Ba Na-Nui Chua H Cryptophyllium icarus gen. et sp. nov., Vietnam, Bidoup-Nui Ba I Cryptophyllium rarum comb. nov., Vietnam, Cuc Phuong J Cryptophyllium limogesi gen. et sp. nov., Vietnam, Ta Dung. Scale bars, 1.0 mm.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
7F2F01FA305959E39E1EC7B005EBDF4C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510470	https://doi.org/10.3897/zookeys.1018.61033.figure6	Figure 6. Details of female phylliid antennae illustrated from specimens within the RBINS collection, illustrations by Julien Caudron (RBINS). Note that the figures start with the scapus, therefore the diagnostic fourth antennomere is the third one shown; to ensure clarity A has the segment number listed to the right of the image. Note that the fourth antennomere of Phyllium (A) is longer than the fifth antennomere in comparison to the Cryptophyllium gen. nov. representatives (B-H), which have the fourth antennomere short and disk-like A Phyllium ericoriai, Philippines B Cryptophyllium bollensi gen. et sp. nov., Vietnam, Phuoc Binh C Cryptophyllium celebicum comb. nov., Indonesia, Sulawesi D Cryptophyllium khmer gen. et sp. nov., Cambodia, Siem Reap Province E Cryptophyllium nuichuaense gen. et sp. nov., Vietnam, Nui Chua F Cryptophyllium phami gen. et sp. nov., Vietnam, Cat Tien G Cryptophyllium westwoodii comb. nov., Laos, Bokeo Province H Cryptophyllium westwoodii comb. nov., Thailand, Chiang Mai. Drawn to relative scale.	Figure 6. Details of female phylliid antennae illustrated from specimens within the RBINS collection, illustrations by Julien Caudron (RBINS). Note that the figures start with the scapus, therefore the diagnostic fourth antennomere is the third one shown; to ensure clarity A has the segment number listed to the right of the image. Note that the fourth antennomere of Phyllium (A) is longer than the fifth antennomere in comparison to the Cryptophyllium gen. nov. representatives (B-H), which have the fourth antennomere short and disk-like A Phyllium ericoriai, Philippines B Cryptophyllium bollensi gen. et sp. nov., Vietnam, Phuoc Binh C Cryptophyllium celebicum comb. nov., Indonesia, Sulawesi D Cryptophyllium khmer gen. et sp. nov., Cambodia, Siem Reap Province E Cryptophyllium nuichuaense gen. et sp. nov., Vietnam, Nui Chua F Cryptophyllium phami gen. et sp. nov., Vietnam, Cat Tien G Cryptophyllium westwoodii comb. nov., Laos, Bokeo Province H Cryptophyllium westwoodii comb. nov., Thailand, Chiang Mai. Drawn to relative scale.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
7F2F01FA305959E39E1EC7B005EBDF4C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510473	https://doi.org/10.3897/zookeys.1018.61033.figure9	Figure 9. Freshly hatched nymphs known for the Cryptophyllium gen. nov., all except for C bred and photographed by Bruno Kneubu ̈ hler (Switzerland) A Cryptophyllium celebicum comb. nov. B Cryptophyllium chrisangi comb. nov. C Cryptophyllium khmer gen. et sp. nov. bred and photographed by Tim Bollens (Belgium) D Cryptophyllium tibetense comb. nov. E Cryptophyllium oyae comb. nov. F Cryptophyllium westwoodii comb. nov. G Cryptophyllium phami gen. et sp. nov. H Cryptophyllium bollensi gen. et sp. nov. I Cryptophyllium icarus gen. et sp. nov.	Figure 9. Freshly hatched nymphs known for the Cryptophyllium gen. nov., all except for C bred and photographed by Bruno Kneubu ̈ hler (Switzerland) A Cryptophyllium celebicum comb. nov. B Cryptophyllium chrisangi comb. nov. C Cryptophyllium khmer gen. et sp. nov. bred and photographed by Tim Bollens (Belgium) D Cryptophyllium tibetense comb. nov. E Cryptophyllium oyae comb. nov. F Cryptophyllium westwoodii comb. nov. G Cryptophyllium phami gen. et sp. nov. H Cryptophyllium bollensi gen. et sp. nov. I Cryptophyllium icarus gen. et sp. nov.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
7F2F01FA305959E39E1EC7B005EBDF4C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510501	https://doi.org/10.3897/zookeys.1018.61033.figure37	Figure 37. Live Cryptophyllium khmer gen. et sp. nov. type material found by the joint effort of the RBINS, RUPP, and VNMN teams in Tatai, Cambodia in October 2016, all found as nymphs. Individuals in photographs A-H were found feeding on Guava trees (Psidium guajava) and the specimen in photograph I was found feeding on an unidentified species. Photographs by Jerome Constant (RBINS) A female B Hong Thai Pham (left) and Sisonila Kangsun (right), participants of the GTI project, photographing the leaf insects, which are very popular in Asia C male nymph D female nymph E male nymph F several nymphs G nymph H nymph I female nymph on unidentified host species.	Figure 37. Live Cryptophyllium khmer gen. et sp. nov. type material found by the joint effort of the RBINS, RUPP, and VNMN teams in Tatai, Cambodia in October 2016, all found as nymphs. Individuals in photographs A-H were found feeding on Guava trees (Psidium guajava) and the specimen in photograph I was found feeding on an unidentified species. Photographs by Jerome Constant (RBINS) A female B Hong Thai Pham (left) and Sisonila Kangsun (right), participants of the GTI project, photographing the leaf insects, which are very popular in Asia C male nymph D female nymph E male nymph F several nymphs G nymph H nymph I female nymph on unidentified host species.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
7F2F01FA305959E39E1EC7B005EBDF4C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510502	https://doi.org/10.3897/zookeys.1018.61033.figure38	Figure 38. Live Cryptophyllium khmer gen. et sp. nov. type material bred and photographed by Tim Bolllens (Belgium), dorsal views A paler form female (paratype) B darker form female (paratype) C male (paratype).	Figure 38. Live Cryptophyllium khmer gen. et sp. nov. type material bred and photographed by Tim Bolllens (Belgium), dorsal views A paler form female (paratype) B darker form female (paratype) C male (paratype).	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
7F2F01FA305959E39E1EC7B005EBDF4C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510503	https://doi.org/10.3897/zookeys.1018.61033.figure39	Figure 39. Cryptophyllium khmer gen. et sp. nov. paratype female, molecular sample RBINS 01 in our analysis, from Tatai, Cambodia, photographs by Jerome Constant (RBINS) A habitus, dorsal B habitus, ventral C details of the antennae, dorsal D pro- tibial and femoral lobes, dorsal E details of the antennae, head, and thorax, dorsal F details of the antennae, head, and thorax, lateral G terminalia, dorsal H genitalia, ventral.	Figure 39. Cryptophyllium khmer gen. et sp. nov. paratype female, molecular sample RBINS 01 in our analysis, from Tatai, Cambodia, photographs by Jerome Constant (RBINS) A habitus, dorsal B habitus, ventral C details of the antennae, dorsal D pro- tibial and femoral lobes, dorsal E details of the antennae, head, and thorax, dorsal F details of the antennae, head, and thorax, lateral G terminalia, dorsal H genitalia, ventral.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
7F2F01FA305959E39E1EC7B005EBDF4C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510504	https://doi.org/10.3897/zookeys.1018.61033.figure40	Figure 40. Cryptophyllium khmer gen. et sp. nov. holotype male, from Tatai, Cambodia, photographs by Jerome Constant (RBINS) A habitus, dorsal B habitus, ventral C pro- tibial and femoral lobes, dorsal D details of the antennae, head, and thorax, lateral E details of the antennae, head, and thorax, dorsal F terminalia, dorsal G genitalia, ventral.	Figure 40. Cryptophyllium khmer gen. et sp. nov. holotype male, from Tatai, Cambodia, photographs by Jerome Constant (RBINS) A habitus, dorsal B habitus, ventral C pro- tibial and femoral lobes, dorsal D details of the antennae, head, and thorax, lateral E details of the antennae, head, and thorax, dorsal F terminalia, dorsal G genitalia, ventral.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
7F2F01FA305959E39E1EC7B005EBDF4C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510505	https://doi.org/10.3897/zookeys.1018.61033.figure41	Figure 41. Cryptophyllium khmer gen. et sp. nov. egg, RBINS collection, photographs by Jerome Constant (RBINS) A lateral view B dorso-lateral view C dorsal view D opercular (anterior) view E posterior view F ventral view.	Figure 41. Cryptophyllium khmer gen. et sp. nov. egg, RBINS collection, photographs by Jerome Constant (RBINS) A lateral view B dorso-lateral view C dorsal view D opercular (anterior) view E posterior view F ventral view.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
7F2F01FA305959E39E1EC7B005EBDF4C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510466	https://doi.org/10.3897/zookeys.1018.61033.figure2	Figure 2. Distribution map for the 24 Cryptophyllium gen. nov. species presently known (with solid lines from their name pointing to the type locality) as well as additional Cryptophyllium gen. nov. species which we could not herein describe / differentiate (indicated by dashed lines). Note that the line for Cryptophyllium westwoodii comb. nov. is pointing to the neotype locality and the type locality for Cryptophyllium athanysus comb. nov. is simply " Ceylon " therefore the line is pointing to the present-day localities we are aware of. Inset is of southern Vietnam showing the distributions of three additional species which could not fit within the main map. The colors in this map are noted to the left of the names within the phylogenetic tree in Fig. 4. Newly herein described species have names noted in bold. Note that with Cryptophyllium khmer sp. nov. not easily distinguishable from Cryptophyllium westwoodii comb. nov. from photos alone, only the locations for these two species where they were genetically sampled are solid colored, all observational images without genetic sampling have bicolored circles and could represent either of these species. Additionally, one symbol is split three ways for Cryptophyllium gen. nov. specimens from the Thai offshore islands of Ko Phangan and Ko Samui which could not be differentiated from C. chrisangi comb. nov., C. westwoodii comb. nov., and C. khmer sp. nov. from images alone. See Suppl. material 4 for a full list of the specimens / observations utilized to make the distribution map with deposition data for specimens and links to observational records>.	Figure 2. Distribution map for the 24 Cryptophyllium gen. nov. species presently known (with solid lines from their name pointing to the type locality) as well as additional Cryptophyllium gen. nov. species which we could not herein describe / differentiate (indicated by dashed lines). Note that the line for Cryptophyllium westwoodii comb. nov. is pointing to the neotype locality and the type locality for Cryptophyllium athanysus comb. nov. is simply " Ceylon " therefore the line is pointing to the present-day localities we are aware of. Inset is of southern Vietnam showing the distributions of three additional species which could not fit within the main map. The colors in this map are noted to the left of the names within the phylogenetic tree in Fig. 4. Newly herein described species have names noted in bold. Note that with Cryptophyllium khmer sp. nov. not easily distinguishable from Cryptophyllium westwoodii comb. nov. from photos alone, only the locations for these two species where they were genetically sampled are solid colored, all observational images without genetic sampling have bicolored circles and could represent either of these species. Additionally, one symbol is split three ways for Cryptophyllium gen. nov. specimens from the Thai offshore islands of Ko Phangan and Ko Samui which could not be differentiated from C. chrisangi comb. nov., C. westwoodii comb. nov., and C. khmer sp. nov. from images alone. See Suppl. material 4 for a full list of the specimens / observations utilized to make the distribution map with deposition data for specimens and links to observational records>.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
7F2F01FA305959E39E1EC7B005EBDF4C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510468	https://doi.org/10.3897/zookeys.1018.61033.figure4	Figure 4. Phylogenetic relationships among 35 phylliid species (67 specimens). The species of the Cryptophyllium gen. nov. are highlighted with colors and symbols corresponding to the distribution map (Fig. 2). Support values based on maximum likelihood (UFBoot, IQ-TREE) are indicated at each node.	Figure 4. Phylogenetic relationships among 35 phylliid species (67 specimens). The species of the Cryptophyllium gen. nov. are highlighted with colors and symbols corresponding to the distribution map (Fig. 2). Support values based on maximum likelihood (UFBoot, IQ-TREE) are indicated at each node.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
3634D0DD98725A759E2C550D8B673343.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510469	https://doi.org/10.3897/zookeys.1018.61033.figure5	Figure 5. Details of the Cryptophyllium gen. nov. male vomer. All from the RBINS collection, prepared and photographed by Jerome Constant (RBINS). Views 1 ventral 2 right lateral of the ventrally oriented vomer 3 left lateral of the ventrally oriented vomer A Cryptophyllium westwoodii comb. nov., Thailand, Na Haeo B Cryptophyllium khmer gen. et sp. nov. Cambodia, Kbal Spean C Cryptophyllium phami gen. et sp. nov., Vietnam, Cat Tien D Cryptophyllium khmer gen. et sp. nov., Cambodia, Tatai E Cryptophyllium bollensi gen. et sp. nov., Vietnam, Phuoc Binh F Cryptophyllium oyae comb. nov., Laos, Mt. Phu Phan G Cryptophyllium bankoi gen. et sp. nov., Vietnam, Ba Na-Nui Chua H Cryptophyllium icarus gen. et sp. nov., Vietnam, Bidoup-Nui Ba I Cryptophyllium rarum comb. nov., Vietnam, Cuc Phuong J Cryptophyllium limogesi gen. et sp. nov., Vietnam, Ta Dung. Scale bars, 1.0 mm.	Figure 5. Details of the Cryptophyllium gen. nov. male vomer. All from the RBINS collection, prepared and photographed by Jerome Constant (RBINS). Views 1 ventral 2 right lateral of the ventrally oriented vomer 3 left lateral of the ventrally oriented vomer A Cryptophyllium westwoodii comb. nov., Thailand, Na Haeo B Cryptophyllium khmer gen. et sp. nov. Cambodia, Kbal Spean C Cryptophyllium phami gen. et sp. nov., Vietnam, Cat Tien D Cryptophyllium khmer gen. et sp. nov., Cambodia, Tatai E Cryptophyllium bollensi gen. et sp. nov., Vietnam, Phuoc Binh F Cryptophyllium oyae comb. nov., Laos, Mt. Phu Phan G Cryptophyllium bankoi gen. et sp. nov., Vietnam, Ba Na-Nui Chua H Cryptophyllium icarus gen. et sp. nov., Vietnam, Bidoup-Nui Ba I Cryptophyllium rarum comb. nov., Vietnam, Cuc Phuong J Cryptophyllium limogesi gen. et sp. nov., Vietnam, Ta Dung. Scale bars, 1.0 mm.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
3634D0DD98725A759E2C550D8B673343.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510472	https://doi.org/10.3897/zookeys.1018.61033.figure8	Figure 8. Known eggs for the Cryptophyllium gen. nov. set in couplets of lateral and then dorsal views. All images except for K and L were taken by Bruno Kneubu ̈ hler (Switzerland), K and L images taken by the first author, Q and R images taken by Rene ́ Limoges (IMQC) A, B Cryptophyllium westwoodii comb. nov. C, D Cryptophyllium tibetense comb. nov. E, F Cryptophyllium bollensi gen. et sp. nov. G, H Cryptophyllium oyae comb. nov. I, J Cryptophyllium icarus gen. et sp. nov. K, L Cryptophyllium liyananae gen. et sp. nov. M, N Cryptophyllium chrisangi comb. nov. O, P Cryptophyllium celebicum comb. nov. Q, R Cryptophyllium limogesi gen. et sp. nov. Scale bars 2.0 mm long.	Figure 8. Known eggs for the Cryptophyllium gen. nov. set in couplets of lateral and then dorsal views. All images except for K and L were taken by Bruno Kneubu ̈ hler (Switzerland), K and L images taken by the first author, Q and R images taken by Rene ́ Limoges (IMQC) A, B Cryptophyllium westwoodii comb. nov. C, D Cryptophyllium tibetense comb. nov. E, F Cryptophyllium bollensi gen. et sp. nov. G, H Cryptophyllium oyae comb. nov. I, J Cryptophyllium icarus gen. et sp. nov. K, L Cryptophyllium liyananae gen. et sp. nov. M, N Cryptophyllium chrisangi comb. nov. O, P Cryptophyllium celebicum comb. nov. Q, R Cryptophyllium limogesi gen. et sp. nov. Scale bars 2.0 mm long.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
3634D0DD98725A759E2C550D8B673343.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510506	https://doi.org/10.3897/zookeys.1018.61033.figure42	Figure 42. Holotype female Cryptophyllium limogesi, photographs by Rene Limoges (IMQC) A habitus, dorsal B habitus, ventral C head and antennae dorsal D antennae detail dorsal E thorax dorsal F front leg, dorsal G terminalia, dorsal H genitalia, ventral.	Figure 42. Holotype female Cryptophyllium limogesi, photographs by Rene Limoges (IMQC) A habitus, dorsal B habitus, ventral C head and antennae dorsal D antennae detail dorsal E thorax dorsal F front leg, dorsal G terminalia, dorsal H genitalia, ventral.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
3634D0DD98725A759E2C550D8B673343.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510507	https://doi.org/10.3897/zookeys.1018.61033.figure43	Figure 43. Paratype male Cryptophyllium limogesi gen. et sp. nov. from Ta Dung N. P. reared to adulthood and photographed by Tim Bollens (Belgium) A habitus, dorsal B detail of the front legs and head C genitalia details, ventral.	Figure 43. Paratype male Cryptophyllium limogesi gen. et sp. nov. from Ta Dung N. P. reared to adulthood and photographed by Tim Bollens (Belgium) A habitus, dorsal B detail of the front legs and head C genitalia details, ventral.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
3634D0DD98725A759E2C550D8B673343.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510508	https://doi.org/10.3897/zookeys.1018.61033.figure44	Figure 44. Paratype male Cryptophyllium limogesi gen. et sp. nov. same individual as in Figure 43 photographs by Jerome Constant (RBINS) A habitus, dorsal B habitus, ventral C details of the base of antennae, head, and thorax, dorsal D details of the base of antennae, head, and thorax, lateral E front leg details, dorsal F genitalia details, dorsal G genitalia details, ventral.	Figure 44. Paratype male Cryptophyllium limogesi gen. et sp. nov. same individual as in Figure 43 photographs by Jerome Constant (RBINS) A habitus, dorsal B habitus, ventral C details of the base of antennae, head, and thorax, dorsal D details of the base of antennae, head, and thorax, lateral E front leg details, dorsal F genitalia details, dorsal G genitalia details, ventral.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
3634D0DD98725A759E2C550D8B673343.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510509	https://doi.org/10.3897/zookeys.1018.61033.figure45	Figure 45. Paratype Cryptophyllium limogesi gen. et sp. nov. egg, laid by paratype females from Dak Lak Province, Vietnam. Photographs by Rene Limoges (IMQC) A lateral B dorsal C ventral D opercular (anterior) view.	Figure 45. Paratype Cryptophyllium limogesi gen. et sp. nov. egg, laid by paratype females from Dak Lak Province, Vietnam. Photographs by Rene Limoges (IMQC) A lateral B dorsal C ventral D opercular (anterior) view.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
3634D0DD98725A759E2C550D8B673343.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510468	https://doi.org/10.3897/zookeys.1018.61033.figure4	Figure 4. Phylogenetic relationships among 35 phylliid species (67 specimens). The species of the Cryptophyllium gen. nov. are highlighted with colors and symbols corresponding to the distribution map (Fig. 2). Support values based on maximum likelihood (UFBoot, IQ-TREE) are indicated at each node.	Figure 4. Phylogenetic relationships among 35 phylliid species (67 specimens). The species of the Cryptophyllium gen. nov. are highlighted with colors and symbols corresponding to the distribution map (Fig. 2). Support values based on maximum likelihood (UFBoot, IQ-TREE) are indicated at each node.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
3634D0DD98725A759E2C550D8B673343.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510466	https://doi.org/10.3897/zookeys.1018.61033.figure2	Figure 2. Distribution map for the 24 Cryptophyllium gen. nov. species presently known (with solid lines from their name pointing to the type locality) as well as additional Cryptophyllium gen. nov. species which we could not herein describe / differentiate (indicated by dashed lines). Note that the line for Cryptophyllium westwoodii comb. nov. is pointing to the neotype locality and the type locality for Cryptophyllium athanysus comb. nov. is simply " Ceylon " therefore the line is pointing to the present-day localities we are aware of. Inset is of southern Vietnam showing the distributions of three additional species which could not fit within the main map. The colors in this map are noted to the left of the names within the phylogenetic tree in Fig. 4. Newly herein described species have names noted in bold. Note that with Cryptophyllium khmer sp. nov. not easily distinguishable from Cryptophyllium westwoodii comb. nov. from photos alone, only the locations for these two species where they were genetically sampled are solid colored, all observational images without genetic sampling have bicolored circles and could represent either of these species. Additionally, one symbol is split three ways for Cryptophyllium gen. nov. specimens from the Thai offshore islands of Ko Phangan and Ko Samui which could not be differentiated from C. chrisangi comb. nov., C. westwoodii comb. nov., and C. khmer sp. nov. from images alone. See Suppl. material 4 for a full list of the specimens / observations utilized to make the distribution map with deposition data for specimens and links to observational records>.	Figure 2. Distribution map for the 24 Cryptophyllium gen. nov. species presently known (with solid lines from their name pointing to the type locality) as well as additional Cryptophyllium gen. nov. species which we could not herein describe / differentiate (indicated by dashed lines). Note that the line for Cryptophyllium westwoodii comb. nov. is pointing to the neotype locality and the type locality for Cryptophyllium athanysus comb. nov. is simply " Ceylon " therefore the line is pointing to the present-day localities we are aware of. Inset is of southern Vietnam showing the distributions of three additional species which could not fit within the main map. The colors in this map are noted to the left of the names within the phylogenetic tree in Fig. 4. Newly herein described species have names noted in bold. Note that with Cryptophyllium khmer sp. nov. not easily distinguishable from Cryptophyllium westwoodii comb. nov. from photos alone, only the locations for these two species where they were genetically sampled are solid colored, all observational images without genetic sampling have bicolored circles and could represent either of these species. Additionally, one symbol is split three ways for Cryptophyllium gen. nov. specimens from the Thai offshore islands of Ko Phangan and Ko Samui which could not be differentiated from C. chrisangi comb. nov., C. westwoodii comb. nov., and C. khmer sp. nov. from images alone. See Suppl. material 4 for a full list of the specimens / observations utilized to make the distribution map with deposition data for specimens and links to observational records>.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
3634D0DD98725A759E2C550D8B673343.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510527	https://doi.org/10.3897/zookeys.1018.61033.figure63	Figure 63. Cryptophyllium tibetense comb. nov. live female bred and photographed by Bruno Kneubu ̈ hler (Switzerland) A dorsal, habitus B ventral genitalia, note the subgenital plate which reaches all the way to the apex of the abdomen C dorsal head, thorax, and front legs D lateral view with tegmina raised to expose the alae length.	Figure 63. Cryptophyllium tibetense comb. nov. live female bred and photographed by Bruno Kneubu ̈ hler (Switzerland) A dorsal, habitus B ventral genitalia, note the subgenital plate which reaches all the way to the apex of the abdomen C dorsal head, thorax, and front legs D lateral view with tegmina raised to expose the alae length.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
DC10D79283405E2C8DC092214AC2E7B9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510510	https://doi.org/10.3897/zookeys.1018.61033.figure46	Figure 46. Female Cryptophyllium liyananae gen. et sp. nov. A live holotype female, photograph by Chengzhi Bian (China) B live female observed and photographed by Dr. Lu Qiu in Guangxi, Huaping Nature Reserve, 900 - 1000 m, August 2020 C paratype female Coll RC 19 - 182 shortly after being collected before the color faded, photograph by Chengzhi Bian.	Figure 46. Female Cryptophyllium liyananae gen. et sp. nov. A live holotype female, photograph by Chengzhi Bian (China) B live female observed and photographed by Dr. Lu Qiu in Guangxi, Huaping Nature Reserve, 900 - 1000 m, August 2020 C paratype female Coll RC 19 - 182 shortly after being collected before the color faded, photograph by Chengzhi Bian.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
DC10D79283405E2C8DC092214AC2E7B9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510511	https://doi.org/10.3897/zookeys.1018.61033.figure47	Figure 47. Holotype Cryptophyllium liyananae gen. et sp. nov., photographs by Rene ́ Limoges (IMQC) A habitus, dorsal B details of the antennae, dorsal C profemoral lobes, dorsal D details of the antennae, head, and thorax, dorsal E genitalia, ventral.	Figure 47. Holotype Cryptophyllium liyananae gen. et sp. nov., photographs by Rene ́ Limoges (IMQC) A habitus, dorsal B details of the antennae, dorsal C profemoral lobes, dorsal D details of the antennae, head, and thorax, dorsal E genitalia, ventral.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
DC10D79283405E2C8DC092214AC2E7B9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510472	https://doi.org/10.3897/zookeys.1018.61033.figure8	Figure 8. Known eggs for the Cryptophyllium gen. nov. set in couplets of lateral and then dorsal views. All images except for K and L were taken by Bruno Kneubu ̈ hler (Switzerland), K and L images taken by the first author, Q and R images taken by Rene ́ Limoges (IMQC) A, B Cryptophyllium westwoodii comb. nov. C, D Cryptophyllium tibetense comb. nov. E, F Cryptophyllium bollensi gen. et sp. nov. G, H Cryptophyllium oyae comb. nov. I, J Cryptophyllium icarus gen. et sp. nov. K, L Cryptophyllium liyananae gen. et sp. nov. M, N Cryptophyllium chrisangi comb. nov. O, P Cryptophyllium celebicum comb. nov. Q, R Cryptophyllium limogesi gen. et sp. nov. Scale bars 2.0 mm long.	Figure 8. Known eggs for the Cryptophyllium gen. nov. set in couplets of lateral and then dorsal views. All images except for K and L were taken by Bruno Kneubu ̈ hler (Switzerland), K and L images taken by the first author, Q and R images taken by Rene ́ Limoges (IMQC) A, B Cryptophyllium westwoodii comb. nov. C, D Cryptophyllium tibetense comb. nov. E, F Cryptophyllium bollensi gen. et sp. nov. G, H Cryptophyllium oyae comb. nov. I, J Cryptophyllium icarus gen. et sp. nov. K, L Cryptophyllium liyananae gen. et sp. nov. M, N Cryptophyllium chrisangi comb. nov. O, P Cryptophyllium celebicum comb. nov. Q, R Cryptophyllium limogesi gen. et sp. nov. Scale bars 2.0 mm long.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
9DD319B2D2195D248366348AE8D1C48F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510470	https://doi.org/10.3897/zookeys.1018.61033.figure6	Figure 6. Details of female phylliid antennae illustrated from specimens within the RBINS collection, illustrations by Julien Caudron (RBINS). Note that the figures start with the scapus, therefore the diagnostic fourth antennomere is the third one shown; to ensure clarity A has the segment number listed to the right of the image. Note that the fourth antennomere of Phyllium (A) is longer than the fifth antennomere in comparison to the Cryptophyllium gen. nov. representatives (B-H), which have the fourth antennomere short and disk-like A Phyllium ericoriai, Philippines B Cryptophyllium bollensi gen. et sp. nov., Vietnam, Phuoc Binh C Cryptophyllium celebicum comb. nov., Indonesia, Sulawesi D Cryptophyllium khmer gen. et sp. nov., Cambodia, Siem Reap Province E Cryptophyllium nuichuaense gen. et sp. nov., Vietnam, Nui Chua F Cryptophyllium phami gen. et sp. nov., Vietnam, Cat Tien G Cryptophyllium westwoodii comb. nov., Laos, Bokeo Province H Cryptophyllium westwoodii comb. nov., Thailand, Chiang Mai. Drawn to relative scale.	Figure 6. Details of female phylliid antennae illustrated from specimens within the RBINS collection, illustrations by Julien Caudron (RBINS). Note that the figures start with the scapus, therefore the diagnostic fourth antennomere is the third one shown; to ensure clarity A has the segment number listed to the right of the image. Note that the fourth antennomere of Phyllium (A) is longer than the fifth antennomere in comparison to the Cryptophyllium gen. nov. representatives (B-H), which have the fourth antennomere short and disk-like A Phyllium ericoriai, Philippines B Cryptophyllium bollensi gen. et sp. nov., Vietnam, Phuoc Binh C Cryptophyllium celebicum comb. nov., Indonesia, Sulawesi D Cryptophyllium khmer gen. et sp. nov., Cambodia, Siem Reap Province E Cryptophyllium nuichuaense gen. et sp. nov., Vietnam, Nui Chua F Cryptophyllium phami gen. et sp. nov., Vietnam, Cat Tien G Cryptophyllium westwoodii comb. nov., Laos, Bokeo Province H Cryptophyllium westwoodii comb. nov., Thailand, Chiang Mai. Drawn to relative scale.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
9DD319B2D2195D248366348AE8D1C48F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510512	https://doi.org/10.3897/zookeys.1018.61033.figure48	Figure 48. Cryptophyllium nuichuaense gen. et sp. nov. holotype female where she was found in August 2014 in Nui Chua N. P. by Jerome Constant (RBINS) and Joachim Bresseel (RBINS). Photographs by Jerome Constant (RBINS) A dorsal view B dorsolateral view C location where the female was found.	Figure 48. Cryptophyllium nuichuaense gen. et sp. nov. holotype female where she was found in August 2014 in Nui Chua N. P. by Jerome Constant (RBINS) and Joachim Bresseel (RBINS). Photographs by Jerome Constant (RBINS) A dorsal view B dorsolateral view C location where the female was found.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
9DD319B2D2195D248366348AE8D1C48F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510513	https://doi.org/10.3897/zookeys.1018.61033.figure49	Figure 49. Cryptophyllium nuichuaense gen. et sp. nov. holotype female, photographs by Jerome Constant (RBINS) A habitus, dorsal B habitus, ventral C details of the antennae and anterior half of the head capsule D pro- tibial and femoral lobes, dorsal E details of the antennae, head, and thorax, dorsal F details of the base of the antennae, head, and thorax, lateral G terminalia, dorsal H genitalia, ventral.	Figure 49. Cryptophyllium nuichuaense gen. et sp. nov. holotype female, photographs by Jerome Constant (RBINS) A habitus, dorsal B habitus, ventral C details of the antennae and anterior half of the head capsule D pro- tibial and femoral lobes, dorsal E details of the antennae, head, and thorax, dorsal F details of the base of the antennae, head, and thorax, lateral G terminalia, dorsal H genitalia, ventral.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
BD05CDBF570B51D98A8877617AD023EB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510469	https://doi.org/10.3897/zookeys.1018.61033.figure5	Figure 5. Details of the Cryptophyllium gen. nov. male vomer. All from the RBINS collection, prepared and photographed by Jerome Constant (RBINS). Views 1 ventral 2 right lateral of the ventrally oriented vomer 3 left lateral of the ventrally oriented vomer A Cryptophyllium westwoodii comb. nov., Thailand, Na Haeo B Cryptophyllium khmer gen. et sp. nov. Cambodia, Kbal Spean C Cryptophyllium phami gen. et sp. nov., Vietnam, Cat Tien D Cryptophyllium khmer gen. et sp. nov., Cambodia, Tatai E Cryptophyllium bollensi gen. et sp. nov., Vietnam, Phuoc Binh F Cryptophyllium oyae comb. nov., Laos, Mt. Phu Phan G Cryptophyllium bankoi gen. et sp. nov., Vietnam, Ba Na-Nui Chua H Cryptophyllium icarus gen. et sp. nov., Vietnam, Bidoup-Nui Ba I Cryptophyllium rarum comb. nov., Vietnam, Cuc Phuong J Cryptophyllium limogesi gen. et sp. nov., Vietnam, Ta Dung. Scale bars, 1.0 mm.	Figure 5. Details of the Cryptophyllium gen. nov. male vomer. All from the RBINS collection, prepared and photographed by Jerome Constant (RBINS). Views 1 ventral 2 right lateral of the ventrally oriented vomer 3 left lateral of the ventrally oriented vomer A Cryptophyllium westwoodii comb. nov., Thailand, Na Haeo B Cryptophyllium khmer gen. et sp. nov. Cambodia, Kbal Spean C Cryptophyllium phami gen. et sp. nov., Vietnam, Cat Tien D Cryptophyllium khmer gen. et sp. nov., Cambodia, Tatai E Cryptophyllium bollensi gen. et sp. nov., Vietnam, Phuoc Binh F Cryptophyllium oyae comb. nov., Laos, Mt. Phu Phan G Cryptophyllium bankoi gen. et sp. nov., Vietnam, Ba Na-Nui Chua H Cryptophyllium icarus gen. et sp. nov., Vietnam, Bidoup-Nui Ba I Cryptophyllium rarum comb. nov., Vietnam, Cuc Phuong J Cryptophyllium limogesi gen. et sp. nov., Vietnam, Ta Dung. Scale bars, 1.0 mm.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
BD05CDBF570B51D98A8877617AD023EB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510472	https://doi.org/10.3897/zookeys.1018.61033.figure8	Figure 8. Known eggs for the Cryptophyllium gen. nov. set in couplets of lateral and then dorsal views. All images except for K and L were taken by Bruno Kneubu ̈ hler (Switzerland), K and L images taken by the first author, Q and R images taken by Rene ́ Limoges (IMQC) A, B Cryptophyllium westwoodii comb. nov. C, D Cryptophyllium tibetense comb. nov. E, F Cryptophyllium bollensi gen. et sp. nov. G, H Cryptophyllium oyae comb. nov. I, J Cryptophyllium icarus gen. et sp. nov. K, L Cryptophyllium liyananae gen. et sp. nov. M, N Cryptophyllium chrisangi comb. nov. O, P Cryptophyllium celebicum comb. nov. Q, R Cryptophyllium limogesi gen. et sp. nov. Scale bars 2.0 mm long.	Figure 8. Known eggs for the Cryptophyllium gen. nov. set in couplets of lateral and then dorsal views. All images except for K and L were taken by Bruno Kneubu ̈ hler (Switzerland), K and L images taken by the first author, Q and R images taken by Rene ́ Limoges (IMQC) A, B Cryptophyllium westwoodii comb. nov. C, D Cryptophyllium tibetense comb. nov. E, F Cryptophyllium bollensi gen. et sp. nov. G, H Cryptophyllium oyae comb. nov. I, J Cryptophyllium icarus gen. et sp. nov. K, L Cryptophyllium liyananae gen. et sp. nov. M, N Cryptophyllium chrisangi comb. nov. O, P Cryptophyllium celebicum comb. nov. Q, R Cryptophyllium limogesi gen. et sp. nov. Scale bars 2.0 mm long.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
BD05CDBF570B51D98A8877617AD023EB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510473	https://doi.org/10.3897/zookeys.1018.61033.figure9	Figure 9. Freshly hatched nymphs known for the Cryptophyllium gen. nov., all except for C bred and photographed by Bruno Kneubu ̈ hler (Switzerland) A Cryptophyllium celebicum comb. nov. B Cryptophyllium chrisangi comb. nov. C Cryptophyllium khmer gen. et sp. nov. bred and photographed by Tim Bollens (Belgium) D Cryptophyllium tibetense comb. nov. E Cryptophyllium oyae comb. nov. F Cryptophyllium westwoodii comb. nov. G Cryptophyllium phami gen. et sp. nov. H Cryptophyllium bollensi gen. et sp. nov. I Cryptophyllium icarus gen. et sp. nov.	Figure 9. Freshly hatched nymphs known for the Cryptophyllium gen. nov., all except for C bred and photographed by Bruno Kneubu ̈ hler (Switzerland) A Cryptophyllium celebicum comb. nov. B Cryptophyllium chrisangi comb. nov. C Cryptophyllium khmer gen. et sp. nov. bred and photographed by Tim Bollens (Belgium) D Cryptophyllium tibetense comb. nov. E Cryptophyllium oyae comb. nov. F Cryptophyllium westwoodii comb. nov. G Cryptophyllium phami gen. et sp. nov. H Cryptophyllium bollensi gen. et sp. nov. I Cryptophyllium icarus gen. et sp. nov.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
BD05CDBF570B51D98A8877617AD023EB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510514	https://doi.org/10.3897/zookeys.1018.61033.figure50	Figure 50. Live Cryptophyllium oyae comb. nov. bred and photographed by Bruno Kneubu ̈ hler (Switzerland) A female B male C lateral view of female with tegmina held up to expose the alae D female genitalia, ventral E male genitalia, ventral.	Figure 50. Live Cryptophyllium oyae comb. nov. bred and photographed by Bruno Kneubu ̈ hler (Switzerland) A female B male C lateral view of female with tegmina held up to expose the alae D female genitalia, ventral E male genitalia, ventral.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
BD05CDBF570B51D98A8877617AD023EB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510515	https://doi.org/10.3897/zookeys.1018.61033.figure51	Figure 51. Live Cryptophyllium oyae males from additional, non-type localities A from Vietnam: Ha Giang, Dung Ba Commune, observed and photographed by Chien C. Lee (Malaysia) in January 2020 B from Thailand: Nan Province, Bo Kluea Tai District observed and photographed by Tatsatorn Dharithai (Thailand) in September 2019 C from Thailand: Chiang Mai Province, San Ku Ruins, observed and photographed by Rob Thacker (United Kingdom) in January 2021.	Figure 51. Live Cryptophyllium oyae males from additional, non-type localities A from Vietnam: Ha Giang, Dung Ba Commune, observed and photographed by Chien C. Lee (Malaysia) in January 2020 B from Thailand: Nan Province, Bo Kluea Tai District observed and photographed by Tatsatorn Dharithai (Thailand) in September 2019 C from Thailand: Chiang Mai Province, San Ku Ruins, observed and photographed by Rob Thacker (United Kingdom) in January 2021.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
BD05CDBF570B51D98A8877617AD023EB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510516	https://doi.org/10.3897/zookeys.1018.61033.figure52	Figure 52. Live Cryptophyllium oyae pair from China, Yunnan Province, Maguan County, Gulinqing town (古林箐乡) collected by Xiang-Jing Liu (China) in October 2020, photographed by Zhiwei Dong (KIZ) A adult female dorsal B adult male dorsal.	Figure 52. Live Cryptophyllium oyae pair from China, Yunnan Province, Maguan County, Gulinqing town (古林箐乡) collected by Xiang-Jing Liu (China) in October 2020, photographed by Zhiwei Dong (KIZ) A adult female dorsal B adult male dorsal.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
BD05CDBF570B51D98A8877617AD023EB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510466	https://doi.org/10.3897/zookeys.1018.61033.figure2	Figure 2. Distribution map for the 24 Cryptophyllium gen. nov. species presently known (with solid lines from their name pointing to the type locality) as well as additional Cryptophyllium gen. nov. species which we could not herein describe / differentiate (indicated by dashed lines). Note that the line for Cryptophyllium westwoodii comb. nov. is pointing to the neotype locality and the type locality for Cryptophyllium athanysus comb. nov. is simply " Ceylon " therefore the line is pointing to the present-day localities we are aware of. Inset is of southern Vietnam showing the distributions of three additional species which could not fit within the main map. The colors in this map are noted to the left of the names within the phylogenetic tree in Fig. 4. Newly herein described species have names noted in bold. Note that with Cryptophyllium khmer sp. nov. not easily distinguishable from Cryptophyllium westwoodii comb. nov. from photos alone, only the locations for these two species where they were genetically sampled are solid colored, all observational images without genetic sampling have bicolored circles and could represent either of these species. Additionally, one symbol is split three ways for Cryptophyllium gen. nov. specimens from the Thai offshore islands of Ko Phangan and Ko Samui which could not be differentiated from C. chrisangi comb. nov., C. westwoodii comb. nov., and C. khmer sp. nov. from images alone. See Suppl. material 4 for a full list of the specimens / observations utilized to make the distribution map with deposition data for specimens and links to observational records>.	Figure 2. Distribution map for the 24 Cryptophyllium gen. nov. species presently known (with solid lines from their name pointing to the type locality) as well as additional Cryptophyllium gen. nov. species which we could not herein describe / differentiate (indicated by dashed lines). Note that the line for Cryptophyllium westwoodii comb. nov. is pointing to the neotype locality and the type locality for Cryptophyllium athanysus comb. nov. is simply " Ceylon " therefore the line is pointing to the present-day localities we are aware of. Inset is of southern Vietnam showing the distributions of three additional species which could not fit within the main map. The colors in this map are noted to the left of the names within the phylogenetic tree in Fig. 4. Newly herein described species have names noted in bold. Note that with Cryptophyllium khmer sp. nov. not easily distinguishable from Cryptophyllium westwoodii comb. nov. from photos alone, only the locations for these two species where they were genetically sampled are solid colored, all observational images without genetic sampling have bicolored circles and could represent either of these species. Additionally, one symbol is split three ways for Cryptophyllium gen. nov. specimens from the Thai offshore islands of Ko Phangan and Ko Samui which could not be differentiated from C. chrisangi comb. nov., C. westwoodii comb. nov., and C. khmer sp. nov. from images alone. See Suppl. material 4 for a full list of the specimens / observations utilized to make the distribution map with deposition data for specimens and links to observational records>.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
BD05CDBF570B51D98A8877617AD023EB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510527	https://doi.org/10.3897/zookeys.1018.61033.figure63	Figure 63. Cryptophyllium tibetense comb. nov. live female bred and photographed by Bruno Kneubu ̈ hler (Switzerland) A dorsal, habitus B ventral genitalia, note the subgenital plate which reaches all the way to the apex of the abdomen C dorsal head, thorax, and front legs D lateral view with tegmina raised to expose the alae length.	Figure 63. Cryptophyllium tibetense comb. nov. live female bred and photographed by Bruno Kneubu ̈ hler (Switzerland) A dorsal, habitus B ventral genitalia, note the subgenital plate which reaches all the way to the apex of the abdomen C dorsal head, thorax, and front legs D lateral view with tegmina raised to expose the alae length.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
BD05CDBF570B51D98A8877617AD023EB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510541	https://doi.org/10.3897/zookeys.1018.61033.figure77	Figure 77. Female Cryptophyllium yunnanense comb. nov. A, C, D, F are photographs of the female used in our molecular analysis (DZW 04) (the same female in Fig. 76) collected by Zhiwei Dong in China, Yunnan, Daweishan, Pinbian County, Honghe Prefecture, photographs by Zhiwei Dong B, E, G, H of a female from Vietnam, Lao Cai Province, Mt. Fan-si-pan (Coll RC 17 - 270), photographs by RTC A habitus, dorsal B habitus, dorsal C details of antennae, dorsal D details of front leg, dorsal E genitalia, ventral F details of thorax, dorsal G details of thorax, dorsal H details of the thorax, lateral.	Figure 77. Female Cryptophyllium yunnanense comb. nov. A, C, D, F are photographs of the female used in our molecular analysis (DZW 04) (the same female in Fig. 76) collected by Zhiwei Dong in China, Yunnan, Daweishan, Pinbian County, Honghe Prefecture, photographs by Zhiwei Dong B, E, G, H of a female from Vietnam, Lao Cai Province, Mt. Fan-si-pan (Coll RC 17 - 270), photographs by RTC A habitus, dorsal B habitus, dorsal C details of antennae, dorsal D details of front leg, dorsal E genitalia, ventral F details of thorax, dorsal G details of thorax, dorsal H details of the thorax, lateral.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
BD05CDBF570B51D98A8877617AD023EB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510539	https://doi.org/10.3897/zookeys.1018.61033.figure75	Figure 75. Male Cryptophyllium yunnanense comb. nov. (Coll RC 16 - 120), photographed by RTC A habitus, dorsal B left front leg details C head-thorax details D genitalia details, ventral E head-thorax details, lateral.	Figure 75. Male Cryptophyllium yunnanense comb. nov. (Coll RC 16 - 120), photographed by RTC A habitus, dorsal B left front leg details C head-thorax details D genitalia details, ventral E head-thorax details, lateral.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
553D7DA04C9C541383562C995A6AEAD2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510517	https://doi.org/10.3897/zookeys.1018.61033.figure53	Figure 53. Cryptophyllium parum comb. nov. from Hainan A-D male which was used in our molecular analysis (Coll RC 19 - 156) from China: Hainan Island, leg. Yingtong Wang, photographs by RTC A habitus, dorsal B details of the head-thorax, dorsal C front leg details, dorsal D dorsolateral view E presumed female Cryptophyllium parum comb. nov. habitus, dorsal, from Ledong Li Autonomous County, Hainan Province, China, observed in December 2018 and uploaded to iNaturalist. com under the creative commons license (CC BY-NC 4.0) by user chenhanlin.	Figure 53. Cryptophyllium parum comb. nov. from Hainan A-D male which was used in our molecular analysis (Coll RC 19 - 156) from China: Hainan Island, leg. Yingtong Wang, photographs by RTC A habitus, dorsal B details of the head-thorax, dorsal C front leg details, dorsal D dorsolateral view E presumed female Cryptophyllium parum comb. nov. habitus, dorsal, from Ledong Li Autonomous County, Hainan Province, China, observed in December 2018 and uploaded to iNaturalist. com under the creative commons license (CC BY-NC 4.0) by user chenhanlin.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
19CD5EF761A957D89DA569CC8CCB7D94.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510469	https://doi.org/10.3897/zookeys.1018.61033.figure5	Figure 5. Details of the Cryptophyllium gen. nov. male vomer. All from the RBINS collection, prepared and photographed by Jerome Constant (RBINS). Views 1 ventral 2 right lateral of the ventrally oriented vomer 3 left lateral of the ventrally oriented vomer A Cryptophyllium westwoodii comb. nov., Thailand, Na Haeo B Cryptophyllium khmer gen. et sp. nov. Cambodia, Kbal Spean C Cryptophyllium phami gen. et sp. nov., Vietnam, Cat Tien D Cryptophyllium khmer gen. et sp. nov., Cambodia, Tatai E Cryptophyllium bollensi gen. et sp. nov., Vietnam, Phuoc Binh F Cryptophyllium oyae comb. nov., Laos, Mt. Phu Phan G Cryptophyllium bankoi gen. et sp. nov., Vietnam, Ba Na-Nui Chua H Cryptophyllium icarus gen. et sp. nov., Vietnam, Bidoup-Nui Ba I Cryptophyllium rarum comb. nov., Vietnam, Cuc Phuong J Cryptophyllium limogesi gen. et sp. nov., Vietnam, Ta Dung. Scale bars, 1.0 mm.	Figure 5. Details of the Cryptophyllium gen. nov. male vomer. All from the RBINS collection, prepared and photographed by Jerome Constant (RBINS). Views 1 ventral 2 right lateral of the ventrally oriented vomer 3 left lateral of the ventrally oriented vomer A Cryptophyllium westwoodii comb. nov., Thailand, Na Haeo B Cryptophyllium khmer gen. et sp. nov. Cambodia, Kbal Spean C Cryptophyllium phami gen. et sp. nov., Vietnam, Cat Tien D Cryptophyllium khmer gen. et sp. nov., Cambodia, Tatai E Cryptophyllium bollensi gen. et sp. nov., Vietnam, Phuoc Binh F Cryptophyllium oyae comb. nov., Laos, Mt. Phu Phan G Cryptophyllium bankoi gen. et sp. nov., Vietnam, Ba Na-Nui Chua H Cryptophyllium icarus gen. et sp. nov., Vietnam, Bidoup-Nui Ba I Cryptophyllium rarum comb. nov., Vietnam, Cuc Phuong J Cryptophyllium limogesi gen. et sp. nov., Vietnam, Ta Dung. Scale bars, 1.0 mm.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
19CD5EF761A957D89DA569CC8CCB7D94.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510470	https://doi.org/10.3897/zookeys.1018.61033.figure6	Figure 6. Details of female phylliid antennae illustrated from specimens within the RBINS collection, illustrations by Julien Caudron (RBINS). Note that the figures start with the scapus, therefore the diagnostic fourth antennomere is the third one shown; to ensure clarity A has the segment number listed to the right of the image. Note that the fourth antennomere of Phyllium (A) is longer than the fifth antennomere in comparison to the Cryptophyllium gen. nov. representatives (B-H), which have the fourth antennomere short and disk-like A Phyllium ericoriai, Philippines B Cryptophyllium bollensi gen. et sp. nov., Vietnam, Phuoc Binh C Cryptophyllium celebicum comb. nov., Indonesia, Sulawesi D Cryptophyllium khmer gen. et sp. nov., Cambodia, Siem Reap Province E Cryptophyllium nuichuaense gen. et sp. nov., Vietnam, Nui Chua F Cryptophyllium phami gen. et sp. nov., Vietnam, Cat Tien G Cryptophyllium westwoodii comb. nov., Laos, Bokeo Province H Cryptophyllium westwoodii comb. nov., Thailand, Chiang Mai. Drawn to relative scale.	Figure 6. Details of female phylliid antennae illustrated from specimens within the RBINS collection, illustrations by Julien Caudron (RBINS). Note that the figures start with the scapus, therefore the diagnostic fourth antennomere is the third one shown; to ensure clarity A has the segment number listed to the right of the image. Note that the fourth antennomere of Phyllium (A) is longer than the fifth antennomere in comparison to the Cryptophyllium gen. nov. representatives (B-H), which have the fourth antennomere short and disk-like A Phyllium ericoriai, Philippines B Cryptophyllium bollensi gen. et sp. nov., Vietnam, Phuoc Binh C Cryptophyllium celebicum comb. nov., Indonesia, Sulawesi D Cryptophyllium khmer gen. et sp. nov., Cambodia, Siem Reap Province E Cryptophyllium nuichuaense gen. et sp. nov., Vietnam, Nui Chua F Cryptophyllium phami gen. et sp. nov., Vietnam, Cat Tien G Cryptophyllium westwoodii comb. nov., Laos, Bokeo Province H Cryptophyllium westwoodii comb. nov., Thailand, Chiang Mai. Drawn to relative scale.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
19CD5EF761A957D89DA569CC8CCB7D94.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510473	https://doi.org/10.3897/zookeys.1018.61033.figure9	Figure 9. Freshly hatched nymphs known for the Cryptophyllium gen. nov., all except for C bred and photographed by Bruno Kneubu ̈ hler (Switzerland) A Cryptophyllium celebicum comb. nov. B Cryptophyllium chrisangi comb. nov. C Cryptophyllium khmer gen. et sp. nov. bred and photographed by Tim Bollens (Belgium) D Cryptophyllium tibetense comb. nov. E Cryptophyllium oyae comb. nov. F Cryptophyllium westwoodii comb. nov. G Cryptophyllium phami gen. et sp. nov. H Cryptophyllium bollensi gen. et sp. nov. I Cryptophyllium icarus gen. et sp. nov.	Figure 9. Freshly hatched nymphs known for the Cryptophyllium gen. nov., all except for C bred and photographed by Bruno Kneubu ̈ hler (Switzerland) A Cryptophyllium celebicum comb. nov. B Cryptophyllium chrisangi comb. nov. C Cryptophyllium khmer gen. et sp. nov. bred and photographed by Tim Bollens (Belgium) D Cryptophyllium tibetense comb. nov. E Cryptophyllium oyae comb. nov. F Cryptophyllium westwoodii comb. nov. G Cryptophyllium phami gen. et sp. nov. H Cryptophyllium bollensi gen. et sp. nov. I Cryptophyllium icarus gen. et sp. nov.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
19CD5EF761A957D89DA569CC8CCB7D94.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510518	https://doi.org/10.3897/zookeys.1018.61033.figure54	Figure 54. Paratype female Cryptophyllium phami gen. et sp. nov. where she was found in July 2012 by Joachim Bresseel (RBINS) and Jerome Constant (RBINS) in the Cat Tien N. P., Vietnam. This is the same individual as in Figure 1, but these photos were taken with flash. Photographs by Jerome Constant (RBINS) A branch the female was found on, note the leaves which show signs of the damage which led to her discovery B same individual, zoomed in to show details.	Figure 54. Paratype female Cryptophyllium phami gen. et sp. nov. where she was found in July 2012 by Joachim Bresseel (RBINS) and Jerome Constant (RBINS) in the Cat Tien N. P., Vietnam. This is the same individual as in Figure 1, but these photos were taken with flash. Photographs by Jerome Constant (RBINS) A branch the female was found on, note the leaves which show signs of the damage which led to her discovery B same individual, zoomed in to show details.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
19CD5EF761A957D89DA569CC8CCB7D94.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510519	https://doi.org/10.3897/zookeys.1018.61033.figure55	Figure 55. Live Cryptophyllium phami gen. et sp. nov. bred and photographed by Bruno Kneubu ̈ hler (Switzerland) A adult female, dorsal B nymph, dorsal C adult male, doorsal.	Figure 55. Live Cryptophyllium phami gen. et sp. nov. bred and photographed by Bruno Kneubu ̈ hler (Switzerland) A adult female, dorsal B nymph, dorsal C adult male, doorsal.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
19CD5EF761A957D89DA569CC8CCB7D94.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510520	https://doi.org/10.3897/zookeys.1018.61033.figure56	Figure 56. Cryptophyllium phami gen. et sp. nov. paratype female (RBINS), photographs by Jerome Constant (RBINS) A habitus, dorsal B habitus, ventral C details of antennae and anterior of head capsule D profemoral and tibial lobes, dorsal E details of antennae, head, and thorax, dorsal F details of head and thorax, lateral G terminalia, dorsal H genitalia, ventral.	Figure 56. Cryptophyllium phami gen. et sp. nov. paratype female (RBINS), photographs by Jerome Constant (RBINS) A habitus, dorsal B habitus, ventral C details of antennae and anterior of head capsule D profemoral and tibial lobes, dorsal E details of antennae, head, and thorax, dorsal F details of head and thorax, lateral G terminalia, dorsal H genitalia, ventral.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
19CD5EF761A957D89DA569CC8CCB7D94.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510521	https://doi.org/10.3897/zookeys.1018.61033.figure57	Figure 57. Cryptophyllium phami gen. et sp. nov. holotype male (RBINS), photographs by Jerome Constant (RBINS) A habitus, dorsal B habitus, ventral C front leg, dorsal D details of base of antennae and head-thorax, dorsal E details of head and thorax, lateral F terminalia, dorsal G genitalia, ventral.	Figure 57. Cryptophyllium phami gen. et sp. nov. holotype male (RBINS), photographs by Jerome Constant (RBINS) A habitus, dorsal B habitus, ventral C front leg, dorsal D details of base of antennae and head-thorax, dorsal E details of head and thorax, lateral F terminalia, dorsal G genitalia, ventral.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
19CD5EF761A957D89DA569CC8CCB7D94.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510522	https://doi.org/10.3897/zookeys.1018.61033.figure58	Figure 58. Cryptophyllium phami gen. et sp. nov. egg, RBINS collection, photographs by Jerome Constant A lateral view B dorso-lateral view C dorsal view D opercular (anterior) view E posterior view F ventral view.	Figure 58. Cryptophyllium phami gen. et sp. nov. egg, RBINS collection, photographs by Jerome Constant A lateral view B dorso-lateral view C dorsal view D opercular (anterior) view E posterior view F ventral view.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
19CD5EF761A957D89DA569CC8CCB7D94.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510465	https://doi.org/10.3897/zookeys.1018.61033.figure1	Figure 1. Paratype Cryptophyllium phami sp. nov. where it was found in Cat Tien N. P. (Vietnam) feeding on a Guava tree (Psidium guajava) in July 2012 by Jerome Constant (RBINS) and Joachim Bresseel (RBINS). The photograph was taken without flash to show how they appear naturally from underneath with the light passing through their thin bodies like the light passing through a leaf.	Figure 1. Paratype Cryptophyllium phami sp. nov. where it was found in Cat Tien N. P. (Vietnam) feeding on a Guava tree (Psidium guajava) in July 2012 by Jerome Constant (RBINS) and Joachim Bresseel (RBINS). The photograph was taken without flash to show how they appear naturally from underneath with the light passing through their thin bodies like the light passing through a leaf.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
64EBAECAF9BB521EAAB400E22E3CB59E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510469	https://doi.org/10.3897/zookeys.1018.61033.figure5	Figure 5. Details of the Cryptophyllium gen. nov. male vomer. All from the RBINS collection, prepared and photographed by Jerome Constant (RBINS). Views 1 ventral 2 right lateral of the ventrally oriented vomer 3 left lateral of the ventrally oriented vomer A Cryptophyllium westwoodii comb. nov., Thailand, Na Haeo B Cryptophyllium khmer gen. et sp. nov. Cambodia, Kbal Spean C Cryptophyllium phami gen. et sp. nov., Vietnam, Cat Tien D Cryptophyllium khmer gen. et sp. nov., Cambodia, Tatai E Cryptophyllium bollensi gen. et sp. nov., Vietnam, Phuoc Binh F Cryptophyllium oyae comb. nov., Laos, Mt. Phu Phan G Cryptophyllium bankoi gen. et sp. nov., Vietnam, Ba Na-Nui Chua H Cryptophyllium icarus gen. et sp. nov., Vietnam, Bidoup-Nui Ba I Cryptophyllium rarum comb. nov., Vietnam, Cuc Phuong J Cryptophyllium limogesi gen. et sp. nov., Vietnam, Ta Dung. Scale bars, 1.0 mm.	Figure 5. Details of the Cryptophyllium gen. nov. male vomer. All from the RBINS collection, prepared and photographed by Jerome Constant (RBINS). Views 1 ventral 2 right lateral of the ventrally oriented vomer 3 left lateral of the ventrally oriented vomer A Cryptophyllium westwoodii comb. nov., Thailand, Na Haeo B Cryptophyllium khmer gen. et sp. nov. Cambodia, Kbal Spean C Cryptophyllium phami gen. et sp. nov., Vietnam, Cat Tien D Cryptophyllium khmer gen. et sp. nov., Cambodia, Tatai E Cryptophyllium bollensi gen. et sp. nov., Vietnam, Phuoc Binh F Cryptophyllium oyae comb. nov., Laos, Mt. Phu Phan G Cryptophyllium bankoi gen. et sp. nov., Vietnam, Ba Na-Nui Chua H Cryptophyllium icarus gen. et sp. nov., Vietnam, Bidoup-Nui Ba I Cryptophyllium rarum comb. nov., Vietnam, Cuc Phuong J Cryptophyllium limogesi gen. et sp. nov., Vietnam, Ta Dung. Scale bars, 1.0 mm.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
64EBAECAF9BB521EAAB400E22E3CB59E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510523	https://doi.org/10.3897/zookeys.1018.61033.figure59	Figure 59. Cryptophyllium rarum comb. nov. male and female A male, habitus, dorsal, molecular sample within this study Coll RC 16 - 115 from Vietnam, Da Nang Province, photograph by RTC B details of the head and thorax, dorsal, Coll RC 16 - 115, photograph by RTC C female, habitus, dorsal, from China: Guangxi Province, Nanning city, Shuangding town, molecular sample DZW 08 within this study, Photographs by Zhiwei Dong (KIZ) D female, genitalia, ventral, photographs by Zhiwei Dong (KIZ).	Figure 59. Cryptophyllium rarum comb. nov. male and female A male, habitus, dorsal, molecular sample within this study Coll RC 16 - 115 from Vietnam, Da Nang Province, photograph by RTC B details of the head and thorax, dorsal, Coll RC 16 - 115, photograph by RTC C female, habitus, dorsal, from China: Guangxi Province, Nanning city, Shuangding town, molecular sample DZW 08 within this study, Photographs by Zhiwei Dong (KIZ) D female, genitalia, ventral, photographs by Zhiwei Dong (KIZ).	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
64EBAECAF9BB521EAAB400E22E3CB59E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510524	https://doi.org/10.3897/zookeys.1018.61033.figure60	Figure 60. Cryptophyllium rarum comb. nov. male collected by Jerome Constant (RBINS) and Joachim Bresseel (RBINS) in Cuc Phuong N. P., Vietnam in August 2016. Photographs by Jerome Constant (RBINS). This male is molecular sample RBINS 04 in our analysis.	Figure 60. Cryptophyllium rarum comb. nov. male collected by Jerome Constant (RBINS) and Joachim Bresseel (RBINS) in Cuc Phuong N. P., Vietnam in August 2016. Photographs by Jerome Constant (RBINS). This male is molecular sample RBINS 04 in our analysis.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
64EBAECAF9BB521EAAB400E22E3CB59E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510525	https://doi.org/10.3897/zookeys.1018.61033.figure61	Figure 61. Cryptophyllium rarum comb. nov. male from Cuc Phuong N. P., Ninh Binh Province, Vietnam collected by Jerome Constant and Joachim Bresseel (RBINS), photographs by Jerome Constant (RBINS) A habitus, dorsal B habitus, ventral C details of the base of antennae, head, and thorax, dorsal D details of the base of antennae, head, and thorax, lateral E pro- femoral and tibial lobes, dorsal F terminalia, dorsal G genitalia, ventral.	Figure 61. Cryptophyllium rarum comb. nov. male from Cuc Phuong N. P., Ninh Binh Province, Vietnam collected by Jerome Constant and Joachim Bresseel (RBINS), photographs by Jerome Constant (RBINS) A habitus, dorsal B habitus, ventral C details of the base of antennae, head, and thorax, dorsal D details of the base of antennae, head, and thorax, lateral E pro- femoral and tibial lobes, dorsal F terminalia, dorsal G genitalia, ventral.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
64EBAECAF9BB521EAAB400E22E3CB59E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510526	https://doi.org/10.3897/zookeys.1018.61033.figure62	Figure 62. Presumed Cryptophyllium rarum comb. nov. a historic female from Hoa Binh, Vietnam, loaned to the RBINS from the MNHN collection, photographs by Jerome Constant (RBINS) A habitus, dorsal B habitus, ventral C details of antennae and anterior of head capsule D profemoral lobe, dorsal E details of the head and thorax, dorsal F details of the head-thorax, lateral G terminalia, dorsal H genitalia, ventral I data label.	Figure 62. Presumed Cryptophyllium rarum comb. nov. a historic female from Hoa Binh, Vietnam, loaned to the RBINS from the MNHN collection, photographs by Jerome Constant (RBINS) A habitus, dorsal B habitus, ventral C details of antennae and anterior of head capsule D profemoral lobe, dorsal E details of the head and thorax, dorsal F details of the head-thorax, lateral G terminalia, dorsal H genitalia, ventral I data label.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
64EBAECAF9BB521EAAB400E22E3CB59E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510468	https://doi.org/10.3897/zookeys.1018.61033.figure4	Figure 4. Phylogenetic relationships among 35 phylliid species (67 specimens). The species of the Cryptophyllium gen. nov. are highlighted with colors and symbols corresponding to the distribution map (Fig. 2). Support values based on maximum likelihood (UFBoot, IQ-TREE) are indicated at each node.	Figure 4. Phylogenetic relationships among 35 phylliid species (67 specimens). The species of the Cryptophyllium gen. nov. are highlighted with colors and symbols corresponding to the distribution map (Fig. 2). Support values based on maximum likelihood (UFBoot, IQ-TREE) are indicated at each node.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
64EBAECAF9BB521EAAB400E22E3CB59E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510498	https://doi.org/10.3897/zookeys.1018.61033.figure34	Figure 34. Cryptophyllium icarus gen. et sp. nov. female paratype, photographs by Jerome Constant (RBINS) A habitus, dorsal B habitus, ventral C antennae and anterior of head capsule D profemoral and protibial lobes E head and thorax details dorsal F head and thorax details lateral G genitalia dorsal H genitalia ventral.	Figure 34. Cryptophyllium icarus gen. et sp. nov. female paratype, photographs by Jerome Constant (RBINS) A habitus, dorsal B habitus, ventral C antennae and anterior of head capsule D profemoral and protibial lobes E head and thorax details dorsal F head and thorax details lateral G genitalia dorsal H genitalia ventral.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
64EBAECAF9BB521EAAB400E22E3CB59E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510497	https://doi.org/10.3897/zookeys.1018.61033.figure33	Figure 33. Live adult Cryptophyllium icarus bred and photographed by Bruno Kneubu ̈ hler (Switzerland) A female B male C lateral view of female with tegmina held up to expose the dwarfed alae underneath.	Figure 33. Live adult Cryptophyllium icarus bred and photographed by Bruno Kneubu ̈ hler (Switzerland) A female B male C lateral view of female with tegmina held up to expose the dwarfed alae underneath.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
64EBAECAF9BB521EAAB400E22E3CB59E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510478	https://doi.org/10.3897/zookeys.1018.61033.figure14	Figure 14. Cryptophyllium bankoi gen. et sp. nov. holotype male, photographs by RTC A front leg details, dorsal B details of the base of antennae, head, thorax C habitus, dorsal D details of genitalia, ventral.	Figure 14. Cryptophyllium bankoi gen. et sp. nov. holotype male, photographs by RTC A front leg details, dorsal B details of the base of antennae, head, thorax C habitus, dorsal D details of genitalia, ventral.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
64EBAECAF9BB521EAAB400E22E3CB59E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510479	https://doi.org/10.3897/zookeys.1018.61033.figure15	Figure 15. Cryptophyllium bankoi gen. et sp. nov. paratype male from the Ba Na-Nui Chua Nature Reserve (RBINS), photographs by Jerome Constant (RBINS) A habitus, dorsal B habitus, ventral C pro- femoral and tibial, dorsal D details of the base of antennae, head, and thorax, dorsal E details of the base of antennae, head, and thorax, lateral F terminalia, dorsal G genitalia, ventral.	Figure 15. Cryptophyllium bankoi gen. et sp. nov. paratype male from the Ba Na-Nui Chua Nature Reserve (RBINS), photographs by Jerome Constant (RBINS) A habitus, dorsal B habitus, ventral C pro- femoral and tibial, dorsal D details of the base of antennae, head, and thorax, dorsal E details of the base of antennae, head, and thorax, lateral F terminalia, dorsal G genitalia, ventral.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
A26EBE1EFDCC5CDB863F9069FF5C0F01.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510472	https://doi.org/10.3897/zookeys.1018.61033.figure8	Figure 8. Known eggs for the Cryptophyllium gen. nov. set in couplets of lateral and then dorsal views. All images except for K and L were taken by Bruno Kneubu ̈ hler (Switzerland), K and L images taken by the first author, Q and R images taken by Rene ́ Limoges (IMQC) A, B Cryptophyllium westwoodii comb. nov. C, D Cryptophyllium tibetense comb. nov. E, F Cryptophyllium bollensi gen. et sp. nov. G, H Cryptophyllium oyae comb. nov. I, J Cryptophyllium icarus gen. et sp. nov. K, L Cryptophyllium liyananae gen. et sp. nov. M, N Cryptophyllium chrisangi comb. nov. O, P Cryptophyllium celebicum comb. nov. Q, R Cryptophyllium limogesi gen. et sp. nov. Scale bars 2.0 mm long.	Figure 8. Known eggs for the Cryptophyllium gen. nov. set in couplets of lateral and then dorsal views. All images except for K and L were taken by Bruno Kneubu ̈ hler (Switzerland), K and L images taken by the first author, Q and R images taken by Rene ́ Limoges (IMQC) A, B Cryptophyllium westwoodii comb. nov. C, D Cryptophyllium tibetense comb. nov. E, F Cryptophyllium bollensi gen. et sp. nov. G, H Cryptophyllium oyae comb. nov. I, J Cryptophyllium icarus gen. et sp. nov. K, L Cryptophyllium liyananae gen. et sp. nov. M, N Cryptophyllium chrisangi comb. nov. O, P Cryptophyllium celebicum comb. nov. Q, R Cryptophyllium limogesi gen. et sp. nov. Scale bars 2.0 mm long.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
A26EBE1EFDCC5CDB863F9069FF5C0F01.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510473	https://doi.org/10.3897/zookeys.1018.61033.figure9	Figure 9. Freshly hatched nymphs known for the Cryptophyllium gen. nov., all except for C bred and photographed by Bruno Kneubu ̈ hler (Switzerland) A Cryptophyllium celebicum comb. nov. B Cryptophyllium chrisangi comb. nov. C Cryptophyllium khmer gen. et sp. nov. bred and photographed by Tim Bollens (Belgium) D Cryptophyllium tibetense comb. nov. E Cryptophyllium oyae comb. nov. F Cryptophyllium westwoodii comb. nov. G Cryptophyllium phami gen. et sp. nov. H Cryptophyllium bollensi gen. et sp. nov. I Cryptophyllium icarus gen. et sp. nov.	Figure 9. Freshly hatched nymphs known for the Cryptophyllium gen. nov., all except for C bred and photographed by Bruno Kneubu ̈ hler (Switzerland) A Cryptophyllium celebicum comb. nov. B Cryptophyllium chrisangi comb. nov. C Cryptophyllium khmer gen. et sp. nov. bred and photographed by Tim Bollens (Belgium) D Cryptophyllium tibetense comb. nov. E Cryptophyllium oyae comb. nov. F Cryptophyllium westwoodii comb. nov. G Cryptophyllium phami gen. et sp. nov. H Cryptophyllium bollensi gen. et sp. nov. I Cryptophyllium icarus gen. et sp. nov.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
A26EBE1EFDCC5CDB863F9069FF5C0F01.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510527	https://doi.org/10.3897/zookeys.1018.61033.figure63	Figure 63. Cryptophyllium tibetense comb. nov. live female bred and photographed by Bruno Kneubu ̈ hler (Switzerland) A dorsal, habitus B ventral genitalia, note the subgenital plate which reaches all the way to the apex of the abdomen C dorsal head, thorax, and front legs D lateral view with tegmina raised to expose the alae length.	Figure 63. Cryptophyllium tibetense comb. nov. live female bred and photographed by Bruno Kneubu ̈ hler (Switzerland) A dorsal, habitus B ventral genitalia, note the subgenital plate which reaches all the way to the apex of the abdomen C dorsal head, thorax, and front legs D lateral view with tegmina raised to expose the alae length.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
A26EBE1EFDCC5CDB863F9069FF5C0F01.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510528	https://doi.org/10.3897/zookeys.1018.61033.figure64	Figure 64. Cryptophyllium tibetense comb. nov. live pair from Tibet and bred and photographed by Zhiwei Dong A female, habitus, dorsal B male, habitus, dorsal C male, details of the antennae, front legs, head, thorax, and tegmina, dorsal D male, details of the antennae, front legs, head, thorax, and tegmina, lateral view.	Figure 64. Cryptophyllium tibetense comb. nov. live pair from Tibet and bred and photographed by Zhiwei Dong A female, habitus, dorsal B male, habitus, dorsal C male, details of the antennae, front legs, head, thorax, and tegmina, dorsal D male, details of the antennae, front legs, head, thorax, and tegmina, lateral view.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
A26EBE1EFDCC5CDB863F9069FF5C0F01.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510529	https://doi.org/10.3897/zookeys.1018.61033.figure65	Figure 65. Additional Cryptophyllium tibetense comb. nov. distribution records> A live adult female from Kalimpong, West Bengal, India, observed and photographed by Vandana Wadwa Sood (West Bengal, India) in September 2019 B female nymph, NHMUK from Sikkim, India, photographs by RTC C female nymph, NHMUK: " Burma: Mishmi Hills. Delei River. 1,700 ft. 28. i. 1935, M. Steele. ", photographs by RTC D male nymph, NHMUK: " Burma: Mishmi Hills. Lohit River. 22. iii. 1935, M. Steele. ", photographs by RTC E female nymph observed and photographed in Digboi, Assam by Rajib Rudra Tariang (India).	Figure 65. Additional Cryptophyllium tibetense comb. nov. distribution records> A live adult female from Kalimpong, West Bengal, India, observed and photographed by Vandana Wadwa Sood (West Bengal, India) in September 2019 B female nymph, NHMUK from Sikkim, India, photographs by RTC C female nymph, NHMUK: " Burma: Mishmi Hills. Delei River. 1,700 ft. 28. i. 1935, M. Steele. ", photographs by RTC D male nymph, NHMUK: " Burma: Mishmi Hills. Lohit River. 22. iii. 1935, M. Steele. ", photographs by RTC E female nymph observed and photographed in Digboi, Assam by Rajib Rudra Tariang (India).	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
A26EBE1EFDCC5CDB863F9069FF5C0F01.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510530	https://doi.org/10.3897/zookeys.1018.61033.figure66	Figure 66. A Cryptophyllium tibetense comb. nov. from Pasighat in the East Siang district, observed and photographed by Oken Tayeng (India) next to a visually similar leaf from the area the female was found B habitat near where the female was found.	Figure 66. A Cryptophyllium tibetense comb. nov. from Pasighat in the East Siang district, observed and photographed by Oken Tayeng (India) next to a visually similar leaf from the area the female was found B habitat near where the female was found.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
A26EBE1EFDCC5CDB863F9069FF5C0F01.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510511	https://doi.org/10.3897/zookeys.1018.61033.figure47	Figure 47. Holotype Cryptophyllium liyananae gen. et sp. nov., photographs by Rene ́ Limoges (IMQC) A habitus, dorsal B details of the antennae, dorsal C profemoral lobes, dorsal D details of the antennae, head, and thorax, dorsal E genitalia, ventral.	Figure 47. Holotype Cryptophyllium liyananae gen. et sp. nov., photographs by Rene ́ Limoges (IMQC) A habitus, dorsal B details of the antennae, dorsal C profemoral lobes, dorsal D details of the antennae, head, and thorax, dorsal E genitalia, ventral.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
5B088CA6B89C59CD808B3FABF59F3EC1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510531	https://doi.org/10.3897/zookeys.1018.61033.figure67	Figure 67. Holotype female Cryptophyllium wennae gen. et sp. nov., photographs by Zhiwei Dong (KIZ) A habitus, dorsal B details of antennae, dorsal C details of profemoral lobes D details of the head and thorax, dorsal E terminalia, dorsal F details of the head and thorax, lateral G genitalia, ventral.	Figure 67. Holotype female Cryptophyllium wennae gen. et sp. nov., photographs by Zhiwei Dong (KIZ) A habitus, dorsal B details of antennae, dorsal C details of profemoral lobes D details of the head and thorax, dorsal E terminalia, dorsal F details of the head and thorax, lateral G genitalia, ventral.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
602EFA0F6D165D669074356F7C502A98.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510469	https://doi.org/10.3897/zookeys.1018.61033.figure5	Figure 5. Details of the Cryptophyllium gen. nov. male vomer. All from the RBINS collection, prepared and photographed by Jerome Constant (RBINS). Views 1 ventral 2 right lateral of the ventrally oriented vomer 3 left lateral of the ventrally oriented vomer A Cryptophyllium westwoodii comb. nov., Thailand, Na Haeo B Cryptophyllium khmer gen. et sp. nov. Cambodia, Kbal Spean C Cryptophyllium phami gen. et sp. nov., Vietnam, Cat Tien D Cryptophyllium khmer gen. et sp. nov., Cambodia, Tatai E Cryptophyllium bollensi gen. et sp. nov., Vietnam, Phuoc Binh F Cryptophyllium oyae comb. nov., Laos, Mt. Phu Phan G Cryptophyllium bankoi gen. et sp. nov., Vietnam, Ba Na-Nui Chua H Cryptophyllium icarus gen. et sp. nov., Vietnam, Bidoup-Nui Ba I Cryptophyllium rarum comb. nov., Vietnam, Cuc Phuong J Cryptophyllium limogesi gen. et sp. nov., Vietnam, Ta Dung. Scale bars, 1.0 mm.	Figure 5. Details of the Cryptophyllium gen. nov. male vomer. All from the RBINS collection, prepared and photographed by Jerome Constant (RBINS). Views 1 ventral 2 right lateral of the ventrally oriented vomer 3 left lateral of the ventrally oriented vomer A Cryptophyllium westwoodii comb. nov., Thailand, Na Haeo B Cryptophyllium khmer gen. et sp. nov. Cambodia, Kbal Spean C Cryptophyllium phami gen. et sp. nov., Vietnam, Cat Tien D Cryptophyllium khmer gen. et sp. nov., Cambodia, Tatai E Cryptophyllium bollensi gen. et sp. nov., Vietnam, Phuoc Binh F Cryptophyllium oyae comb. nov., Laos, Mt. Phu Phan G Cryptophyllium bankoi gen. et sp. nov., Vietnam, Ba Na-Nui Chua H Cryptophyllium icarus gen. et sp. nov., Vietnam, Bidoup-Nui Ba I Cryptophyllium rarum comb. nov., Vietnam, Cuc Phuong J Cryptophyllium limogesi gen. et sp. nov., Vietnam, Ta Dung. Scale bars, 1.0 mm.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
602EFA0F6D165D669074356F7C502A98.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510470	https://doi.org/10.3897/zookeys.1018.61033.figure6	Figure 6. Details of female phylliid antennae illustrated from specimens within the RBINS collection, illustrations by Julien Caudron (RBINS). Note that the figures start with the scapus, therefore the diagnostic fourth antennomere is the third one shown; to ensure clarity A has the segment number listed to the right of the image. Note that the fourth antennomere of Phyllium (A) is longer than the fifth antennomere in comparison to the Cryptophyllium gen. nov. representatives (B-H), which have the fourth antennomere short and disk-like A Phyllium ericoriai, Philippines B Cryptophyllium bollensi gen. et sp. nov., Vietnam, Phuoc Binh C Cryptophyllium celebicum comb. nov., Indonesia, Sulawesi D Cryptophyllium khmer gen. et sp. nov., Cambodia, Siem Reap Province E Cryptophyllium nuichuaense gen. et sp. nov., Vietnam, Nui Chua F Cryptophyllium phami gen. et sp. nov., Vietnam, Cat Tien G Cryptophyllium westwoodii comb. nov., Laos, Bokeo Province H Cryptophyllium westwoodii comb. nov., Thailand, Chiang Mai. Drawn to relative scale.	Figure 6. Details of female phylliid antennae illustrated from specimens within the RBINS collection, illustrations by Julien Caudron (RBINS). Note that the figures start with the scapus, therefore the diagnostic fourth antennomere is the third one shown; to ensure clarity A has the segment number listed to the right of the image. Note that the fourth antennomere of Phyllium (A) is longer than the fifth antennomere in comparison to the Cryptophyllium gen. nov. representatives (B-H), which have the fourth antennomere short and disk-like A Phyllium ericoriai, Philippines B Cryptophyllium bollensi gen. et sp. nov., Vietnam, Phuoc Binh C Cryptophyllium celebicum comb. nov., Indonesia, Sulawesi D Cryptophyllium khmer gen. et sp. nov., Cambodia, Siem Reap Province E Cryptophyllium nuichuaense gen. et sp. nov., Vietnam, Nui Chua F Cryptophyllium phami gen. et sp. nov., Vietnam, Cat Tien G Cryptophyllium westwoodii comb. nov., Laos, Bokeo Province H Cryptophyllium westwoodii comb. nov., Thailand, Chiang Mai. Drawn to relative scale.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
602EFA0F6D165D669074356F7C502A98.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510472	https://doi.org/10.3897/zookeys.1018.61033.figure8	Figure 8. Known eggs for the Cryptophyllium gen. nov. set in couplets of lateral and then dorsal views. All images except for K and L were taken by Bruno Kneubu ̈ hler (Switzerland), K and L images taken by the first author, Q and R images taken by Rene ́ Limoges (IMQC) A, B Cryptophyllium westwoodii comb. nov. C, D Cryptophyllium tibetense comb. nov. E, F Cryptophyllium bollensi gen. et sp. nov. G, H Cryptophyllium oyae comb. nov. I, J Cryptophyllium icarus gen. et sp. nov. K, L Cryptophyllium liyananae gen. et sp. nov. M, N Cryptophyllium chrisangi comb. nov. O, P Cryptophyllium celebicum comb. nov. Q, R Cryptophyllium limogesi gen. et sp. nov. Scale bars 2.0 mm long.	Figure 8. Known eggs for the Cryptophyllium gen. nov. set in couplets of lateral and then dorsal views. All images except for K and L were taken by Bruno Kneubu ̈ hler (Switzerland), K and L images taken by the first author, Q and R images taken by Rene ́ Limoges (IMQC) A, B Cryptophyllium westwoodii comb. nov. C, D Cryptophyllium tibetense comb. nov. E, F Cryptophyllium bollensi gen. et sp. nov. G, H Cryptophyllium oyae comb. nov. I, J Cryptophyllium icarus gen. et sp. nov. K, L Cryptophyllium liyananae gen. et sp. nov. M, N Cryptophyllium chrisangi comb. nov. O, P Cryptophyllium celebicum comb. nov. Q, R Cryptophyllium limogesi gen. et sp. nov. Scale bars 2.0 mm long.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
602EFA0F6D165D669074356F7C502A98.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510473	https://doi.org/10.3897/zookeys.1018.61033.figure9	Figure 9. Freshly hatched nymphs known for the Cryptophyllium gen. nov., all except for C bred and photographed by Bruno Kneubu ̈ hler (Switzerland) A Cryptophyllium celebicum comb. nov. B Cryptophyllium chrisangi comb. nov. C Cryptophyllium khmer gen. et sp. nov. bred and photographed by Tim Bollens (Belgium) D Cryptophyllium tibetense comb. nov. E Cryptophyllium oyae comb. nov. F Cryptophyllium westwoodii comb. nov. G Cryptophyllium phami gen. et sp. nov. H Cryptophyllium bollensi gen. et sp. nov. I Cryptophyllium icarus gen. et sp. nov.	Figure 9. Freshly hatched nymphs known for the Cryptophyllium gen. nov., all except for C bred and photographed by Bruno Kneubu ̈ hler (Switzerland) A Cryptophyllium celebicum comb. nov. B Cryptophyllium chrisangi comb. nov. C Cryptophyllium khmer gen. et sp. nov. bred and photographed by Tim Bollens (Belgium) D Cryptophyllium tibetense comb. nov. E Cryptophyllium oyae comb. nov. F Cryptophyllium westwoodii comb. nov. G Cryptophyllium phami gen. et sp. nov. H Cryptophyllium bollensi gen. et sp. nov. I Cryptophyllium icarus gen. et sp. nov.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
602EFA0F6D165D669074356F7C502A98.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510532	https://doi.org/10.3897/zookeys.1018.61033.figure68	Figure 68. Live adult Cryptophyllium westwoodii comb. nov. bred and photographed by Bruno Kneubu ̈ hler (Switzerland) from stock collected in Tha Pla Duk subdistrict, Mae Tha district, Lamphun, Thailand, collected in May / June 2014 by Suttah Ek-Amnuay (Thailand) A female habitus B male habitus C female details of the thorax, dorsal D lateral view female with tegmina lifted to show long alae E female genitalia details, ventral.	Figure 68. Live adult Cryptophyllium westwoodii comb. nov. bred and photographed by Bruno Kneubu ̈ hler (Switzerland) from stock collected in Tha Pla Duk subdistrict, Mae Tha district, Lamphun, Thailand, collected in May / June 2014 by Suttah Ek-Amnuay (Thailand) A female habitus B male habitus C female details of the thorax, dorsal D lateral view female with tegmina lifted to show long alae E female genitalia details, ventral.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
602EFA0F6D165D669074356F7C502A98.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510533	https://doi.org/10.3897/zookeys.1018.61033.figure69	Figure 69. Strikingly colored live adult female Cryptophyllium westwoodii comb. nov. observed in Thailand, Nan Province, Bo Kluea Tai A habitus, dorsal and B habitus, ventral, photographed by Lek Karton (Thailand) in November 2019 C habitus, dorsal, photographed by Tatsatorn Dharithai (Thailand) in October 2020.	Figure 69. Strikingly colored live adult female Cryptophyllium westwoodii comb. nov. observed in Thailand, Nan Province, Bo Kluea Tai A habitus, dorsal and B habitus, ventral, photographed by Lek Karton (Thailand) in November 2019 C habitus, dorsal, photographed by Tatsatorn Dharithai (Thailand) in October 2020.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
602EFA0F6D165D669074356F7C502A98.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510534	https://doi.org/10.3897/zookeys.1018.61033.figure70	Figure 70. Female Cryptophyllium westwoodii comb. nov. from Laos, Bokeo Province, Ban Muang Kan (RBINS), photographs by Jerome Constant (RBINS) A habitus, dorsal B habitus, ventral C details of the antennae D pro- tibial and femoral lobes, dorsal E details of the antennae, head, and thorax, dorsal F details of the antennae, head, and thorax, lateral G terminalia, dorsal H genitalia, ventral.	Figure 70. Female Cryptophyllium westwoodii comb. nov. from Laos, Bokeo Province, Ban Muang Kan (RBINS), photographs by Jerome Constant (RBINS) A habitus, dorsal B habitus, ventral C details of the antennae D pro- tibial and femoral lobes, dorsal E details of the antennae, head, and thorax, dorsal F details of the antennae, head, and thorax, lateral G terminalia, dorsal H genitalia, ventral.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
602EFA0F6D165D669074356F7C502A98.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510535	https://doi.org/10.3897/zookeys.1018.61033.figure71	Figure 71. Male Cryptophyllium westwoodii comb. nov. from Laos, Bokeo Province, Ban Muang Kan (RBINS), photographs by Jerome Constant (RBINS) A habitus, dorsal B habitus, ventral C front leg, dorsal D details of the antennae, head, and thorax, lateral E details of the antennae, head, and thorax, dorsal F terminalia, dorsal H genitalia, ventral.	Figure 71. Male Cryptophyllium westwoodii comb. nov. from Laos, Bokeo Province, Ban Muang Kan (RBINS), photographs by Jerome Constant (RBINS) A habitus, dorsal B habitus, ventral C front leg, dorsal D details of the antennae, head, and thorax, lateral E details of the antennae, head, and thorax, dorsal F terminalia, dorsal H genitalia, ventral.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
602EFA0F6D165D669074356F7C502A98.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510536	https://doi.org/10.3897/zookeys.1018.61033.figure72	Figure 72. Neotype male Cryptophyllium westwoodii comb. nov. from Chiang Mai Province, Thailand (Coll RC 16 - 148), photographed by RTC A habitus, dorsal B details of the base of the antennae, head, and thorax, dorsal C front leg details, dorsal D lateral view of the anterior portion of the specimen.	Figure 72. Neotype male Cryptophyllium westwoodii comb. nov. from Chiang Mai Province, Thailand (Coll RC 16 - 148), photographed by RTC A habitus, dorsal B details of the base of the antennae, head, and thorax, dorsal C front leg details, dorsal D lateral view of the anterior portion of the specimen.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
602EFA0F6D165D669074356F7C502A98.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510466	https://doi.org/10.3897/zookeys.1018.61033.figure2	Figure 2. Distribution map for the 24 Cryptophyllium gen. nov. species presently known (with solid lines from their name pointing to the type locality) as well as additional Cryptophyllium gen. nov. species which we could not herein describe / differentiate (indicated by dashed lines). Note that the line for Cryptophyllium westwoodii comb. nov. is pointing to the neotype locality and the type locality for Cryptophyllium athanysus comb. nov. is simply " Ceylon " therefore the line is pointing to the present-day localities we are aware of. Inset is of southern Vietnam showing the distributions of three additional species which could not fit within the main map. The colors in this map are noted to the left of the names within the phylogenetic tree in Fig. 4. Newly herein described species have names noted in bold. Note that with Cryptophyllium khmer sp. nov. not easily distinguishable from Cryptophyllium westwoodii comb. nov. from photos alone, only the locations for these two species where they were genetically sampled are solid colored, all observational images without genetic sampling have bicolored circles and could represent either of these species. Additionally, one symbol is split three ways for Cryptophyllium gen. nov. specimens from the Thai offshore islands of Ko Phangan and Ko Samui which could not be differentiated from C. chrisangi comb. nov., C. westwoodii comb. nov., and C. khmer sp. nov. from images alone. See Suppl. material 4 for a full list of the specimens / observations utilized to make the distribution map with deposition data for specimens and links to observational records>.	Figure 2. Distribution map for the 24 Cryptophyllium gen. nov. species presently known (with solid lines from their name pointing to the type locality) as well as additional Cryptophyllium gen. nov. species which we could not herein describe / differentiate (indicated by dashed lines). Note that the line for Cryptophyllium westwoodii comb. nov. is pointing to the neotype locality and the type locality for Cryptophyllium athanysus comb. nov. is simply " Ceylon " therefore the line is pointing to the present-day localities we are aware of. Inset is of southern Vietnam showing the distributions of three additional species which could not fit within the main map. The colors in this map are noted to the left of the names within the phylogenetic tree in Fig. 4. Newly herein described species have names noted in bold. Note that with Cryptophyllium khmer sp. nov. not easily distinguishable from Cryptophyllium westwoodii comb. nov. from photos alone, only the locations for these two species where they were genetically sampled are solid colored, all observational images without genetic sampling have bicolored circles and could represent either of these species. Additionally, one symbol is split three ways for Cryptophyllium gen. nov. specimens from the Thai offshore islands of Ko Phangan and Ko Samui which could not be differentiated from C. chrisangi comb. nov., C. westwoodii comb. nov., and C. khmer sp. nov. from images alone. See Suppl. material 4 for a full list of the specimens / observations utilized to make the distribution map with deposition data for specimens and links to observational records>.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
602EFA0F6D165D669074356F7C502A98.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510468	https://doi.org/10.3897/zookeys.1018.61033.figure4	Figure 4. Phylogenetic relationships among 35 phylliid species (67 specimens). The species of the Cryptophyllium gen. nov. are highlighted with colors and symbols corresponding to the distribution map (Fig. 2). Support values based on maximum likelihood (UFBoot, IQ-TREE) are indicated at each node.	Figure 4. Phylogenetic relationships among 35 phylliid species (67 specimens). The species of the Cryptophyllium gen. nov. are highlighted with colors and symbols corresponding to the distribution map (Fig. 2). Support values based on maximum likelihood (UFBoot, IQ-TREE) are indicated at each node.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
602EFA0F6D165D669074356F7C502A98.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510504	https://doi.org/10.3897/zookeys.1018.61033.figure40	Figure 40. Cryptophyllium khmer gen. et sp. nov. holotype male, from Tatai, Cambodia, photographs by Jerome Constant (RBINS) A habitus, dorsal B habitus, ventral C pro- tibial and femoral lobes, dorsal D details of the antennae, head, and thorax, lateral E details of the antennae, head, and thorax, dorsal F terminalia, dorsal G genitalia, ventral.	Figure 40. Cryptophyllium khmer gen. et sp. nov. holotype male, from Tatai, Cambodia, photographs by Jerome Constant (RBINS) A habitus, dorsal B habitus, ventral C pro- tibial and femoral lobes, dorsal D details of the antennae, head, and thorax, lateral E details of the antennae, head, and thorax, dorsal F terminalia, dorsal G genitalia, ventral.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
2547FE3A5146576C863ABAB416DE4B82.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510537	https://doi.org/10.3897/zookeys.1018.61033.figure73	Figure 73. Holotype female Cryptophyllium yapicum comb. nov. photographed by RTC A habitus, dorsal B ventrolateral view of the protibiae showing the unique lobe shape C details of the thorax and head, dorsal D lateral view of the thorax-head E terminalia, ventral.	Figure 73. Holotype female Cryptophyllium yapicum comb. nov. photographed by RTC A habitus, dorsal B ventrolateral view of the protibiae showing the unique lobe shape C details of the thorax and head, dorsal D lateral view of the thorax-head E terminalia, ventral.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
2547FE3A5146576C863ABAB416DE4B82.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510538	https://doi.org/10.3897/zookeys.1018.61033.figure74	Figure 74. Three additional non-type specimens of Cryptophyllium yapicum comb. nov., photographs by Miho Maeda, Jerilynn Chun, and James Boone, 2020 (BPBM). Corresponding data labels below the specimen A female habitus, dorsal B associated data labels C female habitus, dorsal D associated data labels E male habitus, dorsal F associated data labels.	Figure 74. Three additional non-type specimens of Cryptophyllium yapicum comb. nov., photographs by Miho Maeda, Jerilynn Chun, and James Boone, 2020 (BPBM). Corresponding data labels below the specimen A female habitus, dorsal B associated data labels C female habitus, dorsal D associated data labels E male habitus, dorsal F associated data labels.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
933CCA9F6C54533EBA59DD0BC6428B28.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510539	https://doi.org/10.3897/zookeys.1018.61033.figure75	Figure 75. Male Cryptophyllium yunnanense comb. nov. (Coll RC 16 - 120), photographed by RTC A habitus, dorsal B left front leg details C head-thorax details D genitalia details, ventral E head-thorax details, lateral.	Figure 75. Male Cryptophyllium yunnanense comb. nov. (Coll RC 16 - 120), photographed by RTC A habitus, dorsal B left front leg details C head-thorax details D genitalia details, ventral E head-thorax details, lateral.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
933CCA9F6C54533EBA59DD0BC6428B28.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510540	https://doi.org/10.3897/zookeys.1018.61033.figure76	Figure 76. Live female Cryptophyllium yunnanense comb. nov. collected and photographed in China, Yunnan, Daweishan, Pinbian County, Honghe Prefecture by Zhiwei Dong (tissue sample DZW 04 within our molecular analysis).	Figure 76. Live female Cryptophyllium yunnanense comb. nov. collected and photographed in China, Yunnan, Daweishan, Pinbian County, Honghe Prefecture by Zhiwei Dong (tissue sample DZW 04 within our molecular analysis).	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
933CCA9F6C54533EBA59DD0BC6428B28.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510541	https://doi.org/10.3897/zookeys.1018.61033.figure77	Figure 77. Female Cryptophyllium yunnanense comb. nov. A, C, D, F are photographs of the female used in our molecular analysis (DZW 04) (the same female in Fig. 76) collected by Zhiwei Dong in China, Yunnan, Daweishan, Pinbian County, Honghe Prefecture, photographs by Zhiwei Dong B, E, G, H of a female from Vietnam, Lao Cai Province, Mt. Fan-si-pan (Coll RC 17 - 270), photographs by RTC A habitus, dorsal B habitus, dorsal C details of antennae, dorsal D details of front leg, dorsal E genitalia, ventral F details of thorax, dorsal G details of thorax, dorsal H details of the thorax, lateral.	Figure 77. Female Cryptophyllium yunnanense comb. nov. A, C, D, F are photographs of the female used in our molecular analysis (DZW 04) (the same female in Fig. 76) collected by Zhiwei Dong in China, Yunnan, Daweishan, Pinbian County, Honghe Prefecture, photographs by Zhiwei Dong B, E, G, H of a female from Vietnam, Lao Cai Province, Mt. Fan-si-pan (Coll RC 17 - 270), photographs by RTC A habitus, dorsal B habitus, dorsal C details of antennae, dorsal D details of front leg, dorsal E genitalia, ventral F details of thorax, dorsal G details of thorax, dorsal H details of the thorax, lateral.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
933CCA9F6C54533EBA59DD0BC6428B28.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510506	https://doi.org/10.3897/zookeys.1018.61033.figure42	Figure 42. Holotype female Cryptophyllium limogesi, photographs by Rene Limoges (IMQC) A habitus, dorsal B habitus, ventral C head and antennae dorsal D antennae detail dorsal E thorax dorsal F front leg, dorsal G terminalia, dorsal H genitalia, ventral.	Figure 42. Holotype female Cryptophyllium limogesi, photographs by Rene Limoges (IMQC) A habitus, dorsal B habitus, ventral C head and antennae dorsal D antennae detail dorsal E thorax dorsal F front leg, dorsal G terminalia, dorsal H genitalia, ventral.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
933CCA9F6C54533EBA59DD0BC6428B28.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/510527	https://doi.org/10.3897/zookeys.1018.61033.figure63	Figure 63. Cryptophyllium tibetense comb. nov. live female bred and photographed by Bruno Kneubu ̈ hler (Switzerland) A dorsal, habitus B ventral genitalia, note the subgenital plate which reaches all the way to the apex of the abdomen C dorsal head, thorax, and front legs D lateral view with tegmina raised to expose the alae length.	Figure 63. Cryptophyllium tibetense comb. nov. live female bred and photographed by Bruno Kneubu ̈ hler (Switzerland) A dorsal, habitus B ventral genitalia, note the subgenital plate which reaches all the way to the apex of the abdomen C dorsal head, thorax, and front legs D lateral view with tegmina raised to expose the alae length.	2021-02-18	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven		Zenodo	biologists	Cumming, Royce T.;Bank, Sarah;Bresseel, Joachim;Constant, Je ́ ro ̂ me;Tirant, Stephane Le;Dong, Zhiwei;Sonet, Gontran;Bradler, Sven			
