taxonID	type	description	language	source
53CC67C870175165968AD28480EFC5C1.taxon	materials_examined	Material examined. Holotype ♂: " VIETNAM: Quang Nam, Tay Giang, Axan Mt, 1,300 meters: July 2017 (Coll RC 17 - 261) ". Deposited in the Montreal Insectarium (IMQC).	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
53CC67C870175165968AD28480EFC5C1.taxon	distribution	Distribution. Currently only known from the type locality in Central Vietnam: Quang Nam Province, Axan Mountain.	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
53CC67C870175165968AD28480EFC5C1.taxon	etymology	Etymology. Noun, Latin for " endowed with spirit ". We felt that these incredible insects are so leaf-like in texture, color, and movement, they simply appear to be a leaf on a tree come to life and walking right out of a fantasy novel and therefore the full binomial translating to " hidden leaf endowed with spirit " was fitting.	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
FD7965C10ABE5CC29D3C500CF043194D.taxon	materials_examined	Material examined. Very few specimens are known. We examined the holotype female in the NHMUK " Holo-type. Ceylon. Athanysus, Westw. Phyllium Athanysus Westw, Ceylon. BMNH (E) # 845230 " (Fig. 11 A) and a male in the OUMNH collection " P. Cryptophyllium athanysus, Westw. Ceylon. E Coll. (1830 - 73), W. W. Saunders., Purchased and, pres. ' 73 by Mrs. F. W. Hope. " Greene (1906: fig. 1 b, plate between page 220 and 221) illustrated a single female specimen as well as an egg (fig. 2 b, page 221). We were unable to trace these specimens for proper examination. Besides these few specimens, we reviewed images of one live female (Fig. 12 A, B) and one live male (Fig. 12 C) from the Sinharaja Forest Reserve, but no other records> are known to us.	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
FD7965C10ABE5CC29D3C500CF043194D.taxon	distribution	Distribution. Sri Lanka. All old records> simply state " Ceylon' ' as the location, but the two modern records> we have come across were both from within the Sinharaja Forest Reserve on the southern end of the island (Fig. 12). Even Greene (1906) regarded this species as rarely encountered with most records> belonging to the other phylliid species present on the island.	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
803CD1D08BC65EBB869D05E0B758D8B4.taxon	materials_examined	Material examined. Holotype ♂: " VIETNAM: Quang Nam, Tay Giang, Axan Mt, 1,300 meters: June 2017, Coll RC 17 - 228 ". Deposited in the Montreal Insectarium (IMQC). Paratypes: (4 ♂♂, 1 ♂ nymph, 1 ♀ nymph) • 1 ♂; " Vietnam; Daknong, June, 2017, Coll RC 17 - 338 " (Coll RC) • 1 ♂; " Vietnam; Quang Ngai Province, Bato Mt. 900 m. elv: May 2015, Coll RC 16 - 141 " (Coll RC) • 1 ♂: " Coll. I. R. Sc. N. B., Da Nang prov., Ba Na-Nui Chua Nat. Res., 18 ° 09 ' N 105 ° 55 ' E, 16 - 19. vii. 2017, GTI Project, Leg. J. Constant and J. Bresseel, I. G.: 33.498 " [vomer dissected] (RBINS) • 1 ♂: " Ngoc Linh, Kon Tum prov., Vietnam, 1700 m, VI. 2016, leg. Luong coll. TB- 05 - 134 " (Coll TB) • 1 subadult ♀: " C. Vietnam, Bach Ma N. P., 16 ° 12 ' N 107 ° 52 ' E, 12 - 17. vii. 2011, Leg J. Constant and J. Bresseel, I. G.: 31.933 " [RBINS- Phyllium - DNA sample 0006] (RBINS) • 1 subadult ♂: " Vietnam, Gia Lai prov. Kon Chu Rang N. R., 600 - 1200 m, 13 - 20. vii. 2018, GTI project, 14 ° 28 ' 28 " N 108 ° 32 ' 27 " E, Leg. J. Constant, J. Bresseel and X. Vermeersch, I. G.: 33.769 " [RBINS- Phyllium - DNA sample 0008] (RBINS).	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
803CD1D08BC65EBB869D05E0B758D8B4.taxon	distribution	Distribution. Central and Southern Vietnam. Known from Quang Ngai, Thua Thien Hue, Da Nang, Gia Lai, Quang Nam, and Dak Nong Provinces.	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
803CD1D08BC65EBB869D05E0B758D8B4.taxon	etymology	Etymology. Patronym. This species is dedicated to our friend and colleague Alexandre Banko for his extensive efforts to discover new species and his long collaboration with Team Phyllies to present us with fresh material to sequence and study.	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
74B70D7C21635BE19FE3D1049C04008F.taxon	materials_examined	Material examined. Holotype ♂: " Coll. I. R. Sc. N. B., Ex breeding Tim Bollens, 2018, Coll. I. R. Sc. N. B., Vietnam, Ninh Thuan prov., Phuoc Binh N. P., 12 ° 04 ' N, 108 ° 45 ' E, 26. vii. 2014, night coll., Leg.: J. Constant & J. Bresseel, GTI project, IG: 32.779 ". Deposited in the Royal Belgian Institute of Natural Sciences (RBINS). Paratypes: (16 ♀♀, 25 ♂♂, 46 eggs) • 1 ♀: " Coll. I. R. Sc. N. B., Vietnam, Ninh Thuan prov., Phuoc Binh N. P., 12 ° 04 ' N, 108 ° 45 ' E, 26. vii. 2014, night coll., Leg.: J. Constant and J. Bresseel, GTI project, IG: 32.779 " (RBINS) • 6 ♂♂: same data as HT [3 with vomer dissected] (RBINS) • 2 ♂♂, 3 ♀♀, 1 subadult ♂: " Coll. I. R. Sc. N. B., Ex breeding Tim Bollens, 2019, Vietnam, Ninh Thuan prov., Phuoc Binh N. P., 12 ° 04 ' N, 108 ° 45 ' E, 26. vii. 2014, night coll., Leg.: J. Constant and J. Bresseel, GTI project, IG: 32.779 " (RBINS) • 1 ♂: " Coll. I. R. Sc. N. B., Ex breeding B. Kneubuhler, Vietnam, Ninh Thuan prov., Phuoc Binh N. P., 12 ° 04 ' N, 108 ° 45 ' E, 26. vii. 2014, night coll., Leg.: J. Constant and J. Bresseel, GTI project, IG: 32.779 " (RBINS) • 1 ♂, 1 ♀: " Coll. I. R. Sc. N. B., Ex breeding Bruno Kneubuehler, 2017 " " Coll. I. R. Sc. N. B., Vietnam, Ninh Thuan prov., Phuoc Binh N. P., 12 ° 04 ' N, 108 ° 45 ' E, 26. vii. 2014, night coll., Leg.: J. Constant and J. Bresseel, GTI project, IG: 32.779 " (VNMN) • 1 ♀; " VIETNAM: Ninh Thuan prov. Phuoc Binh N. P., 12 ° 04 ' N 108 ° 45 ' E, Bred from eggs supplied by Bruno Kneubu ̈ hler (Switzerland), May, 2018, Coll RC 18 - 414 " (Coll RC) • 2 subadult ♀♀; " VIETNAM: Ninh Thuan prov. Phuoc Binh N. P., 12 ° 04 ' N 108 ° 45 ' E, Bred from eggs supplied by Bruno Kneubu ̈ hler (Switzerland), May, 2018 " Coll RC 18 - 412 and 18 - 413 (Coll RC) • 1 ♂; " VIETNAM: Ninh Thuan prov. Phuoc Binh N. P., 12 ° 04 ' N 108 ° 45 ' E, Bred from eggs supplied by Bruno Kneubu ̈ hler (Switzerland), April, 2018, Coll RC 18 - 2017 " (Coll RC) • 19 eggs; " VIETNAM: Ninh Thuan prov. Phuoc Binh N. P., 12 ° 04 ' N 108 ° 45 ' E, eggs supplied by Bruno Kneubu ̈ hler (Switzerland), 2018 ", Coll RC 17 - 375, 17 - 376, 18 - 282 - 18 - 298 (Coll RC) • 1 ♀, 3 ♂♂, 27 eggs; " ex Zucht T. Bollens 2018, Herkunft: Vietnam, Prov. Nin Thuan, Bac Ai Distr., Phuoc Binh N. P., leg. Bresseel and Constant 2014 " [coll. FH, No's 1061 - 1 to 4, E], (Coll FH) • 1 ♀, 1 ♂ (nymph n 3); " ex Zucht F. Hennemann 2019, Herkunft: Vietnam, Prov. Nin Thuan, Bac Ai Distr., Phuoc Binh NP, leg. Bresseel and Constant 2014 " [coll. FH, No's 1061 - 5 and 6], (Coll FH) • 2 ♀♀, 3 ♂♂; " VIETNAM: Ninh Thuan prov. Phuoc Binh N. P., 12 ° 04 ' N 108 ° 45 ' E, Bred by Maxime Ortiz, France, circa 2020 " (Coll MO) • 4 ♀♀, 6 ♂♂; " VIETNAM: Ninh Thuan prov. Phuoc Binh N. P., bred by Bruno Kneubu ̈ hler (Switzerland), circa 2017 - 2018 " (Coll OC).	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
74B70D7C21635BE19FE3D1049C04008F.taxon	distribution	Distribution. At present only known from the type locality of Phuoc Binh N. P., Ninh Thuan Province, southern Vietnam.	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
74B70D7C21635BE19FE3D1049C04008F.taxon	description	Description. Female. Coloration. Coloration description is based upon living individuals (Figs 16, 17 A). Overall coloration is pale green throughout, with variable areas highlighted with burnt red or brown coloration. These areas tend to be the margins on the lobes of the legs, some striping on the lobes of the legs, the thorax, abdominal margins, and the venation in the tegmina (Fig. 16). Morphology. Head. Head capsule about as long as wide, vertex relatively smooth with the only notable feature being the posteromedial tubercle which is finely pointed (Fig. 18 E). Frontal convexity broad and blunt, with a slightly granular surface. Compound eyes slightly protruding from the head capsule, and are not particularly large, taking up ca. 1 / 4 of the head capsule lateral margins (Fig. 18 E). Ocelli absent. Antennal fields slightly wider than the width of the first antennomere. Antennae. Antennae consist of nine segments, with the terminal segment about the same length as the preceding 21 / 2 segments' lengths combined (Fig. 6 B). Antennomeres I-VIII sparsely marked with small transparent setae, the terminal antennomere and the anterior margin of antennomere VIII are covered in stout, brown setae (Fig. 18 C). Thorax. Pronotum with gently concave anterior margin and slightly convex lateral margins, which converge to a straight posterior margin that is half the width of the anterior margin (Fig. 18 E). The pronotum surface is smooth, with only a prominent pit in the center, and slight furrows anterior and lateral to the pit (Fig. 18 E). The pronotum has moderately formed anterior and lateral rims and a weakly formed posterior rim, all of which are relatively smooth (Fig. 18 E). Prosternum and the mesosternum are covered with numerous broad nodes, but the metasternum has a somewhat wrinkled surface. Prescutum longer than wide, lateral rims with 9 - 11 small to medium tubercles, similar in size giving the margin a rough appearance (Fig. 18 E). Prescutum anterior rim prominent but not strongly protruding, rim surface is granular, lacking a large sagittal spine (Fig. 18 F). Prescutum surface heavily granular, with those along the sagittal plane slightly larger than the rest (Fig. 18 E). Mesopleura begin ca. 1 / 4 of the way through the prescutum length and evenly diverge; lateral margin with nine or ten small tubercles with about half of those slightly larger than the rest, with the smaller ones interspersed throughout (Fig. 18 E). Face of the mesopleura smooth or slightly wrinkled, with two notable divots, one on the anterior margin and one near the middle (Fig. 18 F). Wings. Tegmina long, reaching 1 / 2 through abdominal segment VII. Tegmina venation; the subcosta (Sc) is the first vein in the forewing, running parallel with the margin for the first half, and then bending and running towards the margin. The radius (R) spans the central portion of the forewing with two subparallel branched veins; the first radius (R 1) branches ca. 1 / 4 of the way through the wing length and terminates slightly proximal to the midline, and the radial sector (Rs) branches ca. ⅖ of the way through the wing length and terminates near the distal ⅓ of the wing length. There is a weak continuation of the radius following the prominent Rs branching which continues on as a short and thin R-M crossvein that weakly connects the two veins. The media (M) is simply bifurcate with both the media anterior (MA) and media posterior (MP) terminating near to the posterior 1 / 4 of the wing. The cubitus (Cu) is also bifurcate, branching near the posterior ⅕ of the wing into the cubitus anterior (CuA) and cubitus posterior (CuP) which both terminate at or very near the wing posterior apex. The first anal vein (1 A) is simple and fuses with the cubitus early on, at the length about midway between the splitting of the R 1 and Rs. Alae short, with their apex only just passing the anterior margin of abdominal segment IV. Abdomen. Abdominal segments II through the anterior half of IV uniformly diverging. The posterior half of segment IV through the anterior of segment VII are parallel, giving the abdomen a boxy appearance. The posterior half of segment VII ends in a slightly rounded lobe. Segments VIII-X are notably narrower than the previous segments, and have converging margins to the broad rounded apex (Fig. 18 G). Genitalia. Subgenital plate starts at the anterior margin of tergum VIII, is moderately broad, and extends halfway onto tergum X with straight margins ending in a fine point (Fig. 18 H). Gonapophyses VIII are long and moderately broad, slightly exceeding the apex of abdominal tergum X; gonapophyses IX are shorter and narrower, hidden below (Figs 17 C, 18 H). Cerci flat, not strongly cupped, with a granular surface and few detectable setae (Fig. 18 H). Legs. Profemoral exterior lobe broad, rounded, and obtusely angled, smoothly arcing from end to end, ca. ⅓ wider than the width of the interior lobe (Fig. 18 D). Edge of the profemoral exterior lobe granular, or with a slightly serrate surface of eight or nine small teeth (Fig. 18 D). Profemoral interior lobe ca. 2 x as wide as the greatest width of the profemoral shaft, obtusely angled, and marked with five teeth arranged in a two-one-two pattern with looping gaps between them (Fig. 18 D). Mesofemoral exterior lobe arcs from end to end but is slightly bent in the center, weighted towards the distal half, and marked with three or four small serrate teeth distributed on the distal half only. Interior lobe is about the same width as the mesofemoral shaft, and the exterior lobe is slightly wider. Mesofemoral interior lobe arcs smoothly end to end with 6 - 8 small serrate teeth only on the distal half of the arc which is slightly wider than the proximal half of the arc. Metafemoral interior lobe arcs end to end, with the distal half slightly wider than the proximal half and marked with 7 - 10 serrate teeth on the distal half of the lobe. Metafemoral exterior lobe is thin and smooth, hugging the metafemoral shaft and lacks dentation. Protibiae lacking an exterior lobe (Fig. 18 D). Protibiae interior lobe spans the entire length of the protibiae and is ca. 2 x the width of the protibiae shaft itself. The lobe is roundly triangular with the widest portion on the distal half. Mesotibiae and metatibiae lacking exterior and interior lobes. Measurements of paratype female [mm] (wild caught). Length of body (including cerci and head, excluding antennae) 75.6, length / width of head 8.0 / 6.1, antennae 4.3, pronotum 4.8, mesonotum 7.2, length of tegmina 45.3, length of alae 23.1, greatest width of abdomen 28.6, profemora 16.7, mesofemora 13.6, metafemora 16.3, protibiae 10.8, mesotibiae 9.4, metatibiae 12.6. Measurements of paratype females [mm] (ex culture). Length of body (including cerci and head, excluding antennae) 74.6 - 80.1, length / width of head 7.6 - 8.5 / 5.9 - 6.2, antennae 3.9 - 4.2, pronotum 4.5 - 5.4, mesonotum 6.7 - 6.8, length of tegmina 43.2 - 45.1, length of alae (not measurable as they were hidden by the tegmina), greatest width of abdomen 29.7, profemora 17.2 - 18.2, mesofemora 13.7 - 14.4, metafemora 16.5 - 16.6, protibiae 10.9 - 11.0, mesotibiae 9.1 - 9.6, metatibiae 12.5 - 12.8. Male. Coloration. Coloration based upon live bred specimens in captivity (Fig. 17 B). Overall coloration pale green throughout with variable patches of tan to reddish coloration (Fig. 17 B). These tan to reddish areas are primarily around the margins of the lobes of the legs, the margins of the thorax, the tips of the antennae, and the margins of the abdomen. In darker colored specimens the mesofemoral lobes can also have coloration, not just along the margins. Abdominal segment V has a pair of slightly transparent eye spots. Morphology. Head. Head capsule about as long as wide, with a vertex that is relatively smooth with only light granulation throughout. Frontal convexity stout with sparse thin setae. The posteromedial tubercle is not broad but is distinctly raised from the head capsule. Compound eyes large and bulbous, taking up ca. ⅖ of the head capsule lateral margins (Fig. 19 E). There are three well-developed ocelli located between and slightly posterior to the compound eyes. Antennae. Antennae (including the scapus and pedicellus) consist of 23 - 26 segments, all segments except the scapus and pedicellus and terminal three segments are covered in dense setae that are as long as or longer than the antennae segment is wide. The terminal three segments are covered in dense short setae and the scapus and pedicellus are nearly completely bare. Thorax. Pronotum with anterior margin slightly concave and lateral margins that are slightly convex and converging to a straight posterior margin that is ca. 1 / 2 the width of the anterior rim (Fig. 19 E). Anterior and lateral margins of the pronotum have moderately formed rims and the posterior margin lacks a rim (Fig. 19 D). Face of the pronotum is marked by a distinct furrow and pit in the center and a relatively smooth lumpy surface with weak granulation (Fig. 19 D). Prosternum surface is weakly granular with small nodes of even size and spacing. Mesosternum surface marked with slightly more prominent nodes, with the largest along the sagittal plane and denser on the anterior margin, posterior margin with less prominent and smaller nodes. Prescutum slightly longer than wide, with lateral margins that are only slightly converging to the posterior (Fig. 19 E). Lateral rims with eight or nine node-like tubercles, giving the lateral margins a rough textured appearance. Prescutum surface with minimal nodes throughout, with those along the sagittal plane slightly larger than the others. Prescutum anterior rim moderately formed but not strongly raised, with a granular surface and lacking a prominent sagittal tubercle. Mesopleura begin on the anterior prescutum margin but are narrow throughout the anterior ⅓ of their length, only diverging gently for the posterior ⅔ (Fig. 19 E). Lateral margin with eight or nine minor tubercles throughout the length except for the posterior ⅓ which is relatively smooth. Face of the mesopleura mostly smooth, with slight wrinkling throughout. Wings. Tegmina moderate length, extending ⅓ of the way onto abdominal segment III. Tegmina wing venation: the subcosta (Sc) is the first vein, is simple, and terminates the earliest ca. ⅓ of the way through the overall tegmina length. The radius (R) spans the entire length of the tegmina with the first radius (R 1) branching just proximal to the midline and terminating just distal to the midline, followed by the branching and termination of the second radius (R 2) near the distal ⅓ of the wing, and then the radial sector runs to the wing apex. The media (M) also spans the entire length of the tegmina with the first media posterior (MP 1) branching off slightly more than ⅓ of the way through the wing length, and then the second media posterior (MP 2) branches just distal to the midline, and the media anterior (MA) runs to the wing apex. The cubitus (Cu) runs along the edge of the wing as the two media posterior veins fuse with it and as the cubitus reaches the apex it fades. The first anal (1 A) vein terminates upon reaching the cubitus ca. ⅓ of the way through the wing length. Alae well developed in an oval fan configuration, long, reaching to the middle or posterior of abdominal segments IX. Alae wing venation: the costa (C) is present along the entire foremargin giving stability to the wing. The subcosta (Sc) is long, spanning ca. ⅔ of the wing length and is mostly fused with the radius in the beginning but terminates when it meets the costa. The radius (R) spans the entire wing and branches slightly proximal to the midline into the first radius (R 1) and radial sector (Rs) which run gently diverging for most of their length and then converge at the apex of the wing where they terminate near each other but not touching. The media (M) branches early, ca. ⅙ of the way through the wing into the media anterior (MA) and the media posterior (MP) which run parallel with each other throughout the wing until the distal ⅕ of the wing where the media posterior fuses with the media anterior which then run fused to the wing apex where they terminate near the radial sector. The cubitus (Cu) runs unbranched and terminates at the wing apex. Of the anterior anal veins, the first anterior anal (1 AA) fuses with the cubitus near the point where the media branches into the media anterior and media posterior and then the first anterior anal branches from the cubitus ⅔ of the way through the wing length where it uniformly diverges from the cubitus until it terminates at the wing margin. The anterior anal veins two-seven (2 AA- 7 AA) have a common origin and run unbranched in a folding fan pattern of relatively uniform spacing to the wing margin. The posterior anal veins (1 PA- 6 PA) share a common origin separate from the anterior anal veins and run unbranched to the wing margin with slightly thinner spacing than the anterior anal veins. Abdomen. Lateral margins of abdominal segment II parallel, III through the anterior ⅔ of segment IV gradually diverging, the remainder of IV and segment V are parallel-sided, segment VI starts parallel-sided but then gently starts to converge and the remaining segments converge uniformly to the rounded apex of the abdomen. Genitalia. Poculum broad and ends in a rounded apex that slightly passes the anterior margin of segment X (Fig. 19 G). Cerci long and slender, extending from under the anal abdominal segment, nearly flat, not strongly cupped, covered in a granulose surface and numerous short setae (Fig. 19 G). Vomer broad and stout with straight sides evenly converging and ending in a thick apical hook with a smaller second hook adjacent to it (Fig. 5 E). Interestingly while examining the vomers of type material we found several aberrant vomers with some bearing only a singular hook (Fig. 20 A) or even three hooks (Fig. 20 B), based on other specimens examined and the trend within the Cryptophyllium gen. nov. males we expect that a typical male of this species has a two hooked vomer. Legs. Profemoral exterior lobe slightly broader than the interior lobe, ca. 21 / 2 x the greatest width of the profemoral shaft, roundly arcing end to end in a broad obtuse angle that is not distinctly bent, with the proximal margin slightly granulose, and the distal margin with four or five small serrate teeth (Fig. 20 C). Profemoral interior lobe roundly triangular and marked with five sharp teeth arranged in a two-one-two pattern with looping gaps between them, and the central tooth slightly larger than the others (Fig. 20 C). Mesofemoral exterior lobe arcs end to end but is slightly wider on the distal ⅓ which is marked with three or four serrate teeth, and a proximal half that is rather thin. Mesofemoral interior lobe is about the same width as the exterior, is broader on the distal end and is marked with 6 - 8 small serrate teeth. Metafemoral exterior lobe lacks dentation and has a straight margin along the metafemoral shaft. Metafemoral interior lobe smoothly arcs end to end with eight or nine small serrate teeth on the distal ⅔, which is slightly wider than the proximal ⅓. Protibiae lacking exterior lobe, interior lobe reaching end to end in a smooth triangle which is slightly weighted to the distal half and at its widest is ca. 21 / 2 as wide as the protibial shaft (Fig. 20 C). Meso- and metatibiae simple, lacking lobes completely. Measurements of holotype male [mm]. Length of body (including cerci and head, excluding antennae) 57.6, length / width of head 4. ⅓. 4, antennae 41.4, pronotum 3.0, mesonotum 4.3, length of tegmina 18.3, length of alae 43.4, greatest width of abdomen 16.0, profemora 12.8, mesofemora 11.2, metafemora 13.0, protibiae 8.9, mesotibiae 7.7, metatibiae 9.5. Measurements of paratype males [mm] (ex culture). Length of body (including cerci and head, excluding antennae) 55.8 - 65.5, length / width of head 4.1 - 4.9 / 3.2 - 3.5, antennae 39.9 - 41.5, pronotum 3.0 - 3.4, mesonotum 4.0 - 4.8, length of tegmina 18.2 - 20.1, length of alae 41.5 - 47.2, greatest width of abdomen 13.6 - 16.3, profemora 12.4 - 13.6, mesofemora 10.4 - 12.3, metafemora 12.7 - 14.4, protibiae 8.7 - 10.2, mesotibiae 7.5 - 8.4, metatibiae 9.0 - 10.2. Eggs. (Fig. 21). The overall color is muted dark brown, with the moss-like pinnae lighter in color, generally tan or light brown. The lateral surfaces are flat or slightly convex, with eggs either the same width anterior to posterior or with the posterior of the egg slightly wider. The lateral surfaces are marked with 40 - 50 small to medium sized pits, unevenly spaced in no detectable pattern, with sparse tufts of moss-like pinnae between these pits (Fig. 21 A). The dorsal surface has the micropylar plate spanning a majority of the length but not quite reaching end to end. On either side of the micropylar plate is variable pitting (generally eight or so pits) with those one the anterior and posterior ends slightly larger than the central pits (Fig. 21 C). The micropylar plate is symmetrical with the anterior and posterior thin and the middle the widest point. The micropylar cup is not located at this widest midpoint but is instead located on the posterior ⅓ of the micropylar plate. The micropylar plate margin is lined with short moss-like pinnae. Operculum slightly ovular, outer margin with a distinct row of short moss-like pinnae and in from the outer margin is a singular semi-circle of small to medium pits on the dorsal and lateral aspects (not fully surrounding the apex of the operculum as the ventral portion lacks these pits; Fig. 21 D). Operculum is roundly raised with the height ca. 1 / 2 the operculum width and the apex of the raised operculum has a tuft of moss-like pinnae. The ventral surface of the egg capsule has a slightly raised sagittal crest marked sparsely with short moss-like pinnae on the anterior ⅔, and the posterior ⅓ has longer moss-like pinnae. On either side of this raised sagittal crest is pitting, near the posterior ⅓ on each side of the longer moss-like pinnae of the sagittal crest is a large circular pit, and anterior to the lowest large pit are around ten small to medium pits arranged in no detectable pattern (Fig. 21 F). Measurements including the extended pinnae [mm]. Length (including operculum): 4.1 - 4.4; maximum width of capsule when viewed from lateral aspect 2.6 - 2.8; length of micropylar plate 3.3 - 3.4 Newly hatched nymphs. (Fig. 9 H). The general color throughout the body is dark brown with slightly lighter brown on the legs. The basitarsi are yellow and remaining tarsal segments are dark brown. All tibiae lack exterior lobes but do have extremely thin smoothly arcing interior lobes which have several tan to brown stripes throughout their length. All femoral lobes are similar in width and have distinct serration on their distal halves. The interior profemoral lobe lacks a white spot, but the exterior lobe has a distinct white patch on the proximal ⅓ with an additional small white patch at the proximal most margin. The meso- and metafemoral interior lobes have two white patches, one on the proximal most edge, and a larger white patch ⅓ of the way through the length. The meso- and metafemoral exterior lobes also have a large white patch on the proximal ⅓, but lack a smaller white patch on the proximal most margin. The distal ends of the meso- and metafemora also have minimal white edges. The abdomen is mostly brown, but abdominal segments II and III have distinct green patches on their lateral surfaces (the centerline of the abdomen is uniform brown throughout). The terminal three abdominal segments also have a little bit of green on their margins. The widest point of the abdomen is abdominal segment IV.	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
74B70D7C21635BE19FE3D1049C04008F.taxon	etymology	Etymology. Patronym. Named after Tim Bollens (Belgium) who has been instrumental in bringing many new phylliid species into the phasmid breeding community over the years. With his expertise in breeding these difficult phasmids he has allowed us to compare the informative sets of male, female, freshly hatched nymph, and egg morphology instead of only comparing singular dead specimens collected in the wild.	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
9E1ECF645E4D5FD6AAC328B2E0E41250.taxon	materials_examined	Material examined. (11 ♀♀, 3 ♂♂, 5 eggs): 6 ♀♀: " Indonesia: Sulawesi " (Coll RC 16 - 069, 16 - 070, 16 - 075, 16 - 238, (nymph) 16 - 074, (nymph) 16 - 072); 2 ♀♀: " Indonesia: Sulawesi, Palolo, Palu, 2.2008 " (Coll RC 16 - 071, (nymph) 16 - 073); 1 ♀: " Indonesia: Peleng, Tattendeng, Sept. 2019 " (Coll RC 19 - 181); 2 ♂♂: " Indonesia: Sulawesi " (Coll RC 16 - 146, 16 - 076); 1 ♂: " Sulawesi, Central Sulawesi Province, Palu Palolo: February, 2008 " (Coll RC 16 - 145); 1 ♀: " Coll. I. R. Sc. N. B., Indonesia, Sulawesi, Puncak BEI, Palopo, VI. 2001 " (RBINS); 1 ♀: " Indonesia: Bugadidi, ex culture T. Bollens " (RBINS); 2 eggs: " Indonesia: S-Sulawesi, Tiulapolu leg. Jasmin III. 2008, F- 1 Generation, Cultured F. Hennemann 2009 Ex. Coll. Frank Hennemann (Germany) " (Coll RC 18 - 250, 18 - 251); 3 eggs: " Indonesia: Sulawesi; removed from specimen Coll RC 16 - 075 " (Coll RC 17 - 345, 17 - 346, 17 - 347).	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
9E1ECF645E4D5FD6AAC328B2E0E41250.taxon	distribution	Distribution. Known from throughout the island of Sulawesi and from the nearby offshore islands of Peleng to the east and Buton to the south.	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
A16BAF5141C05232A40975226EEEBA20.taxon	materials_examined	Material examined. (7 ♀♀, 5 ♂♂, 10 eggs): 2 ♂♂, 2 ♀♀: " Singapore, ex breeding " (RBINS); 1 ♂: " Singapore (Pulau Ubin), Collected at light, by Francis Seow-Choen, January 5 th, 2018 (0 - 75 m elv) " (Coll RC 18 - 279); 1 ♂: " Thailand: Satun Province, Tarutao Island " (UCR); 1 ♀: " Sarawak: Kuching, 12.12, J. M. Bryan., B. M. 1931 - 150. " (NHMUK); 1 ♀: " P. Phyllium siccifolium, 4.93 Tapah, Dr. Yeh " (LKCNHM); 10 eggs: " Singapore " (Coll RC 18 - 332 - 18 - 341); 1 ♀: " Presented by Dr. Brooke, St. John's I. 26.4.09, Coll Freie Universitat Berlin Sammlung Exotische Insekten, DEI Hemimetabola # 100117 ". Photographic records: 2 ♀♀: Thailand: Nakhon Si Thammarat Province, Thung Song District, August 2020 (photographed by Tatsatorn Dharithai, Thailand); 1 ♂: Indonesia, Pulau Weh Island off the coast of Sumatra (photographed by Mathieu MJP Van Goethem, South Africa).	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
A16BAF5141C05232A40975226EEEBA20.taxon	distribution	Distribution. The type locality for Cryptophyllium chrisangi comb. nov. is mainland Singapore and it has additionally been recorded from St. John's Island from a record in the SDEI collections. Additionally, we have observed specimens and photographs from several areas, including several from Thailand: Nakhon Si Thammarat Province, Thung Song District (Fig. 24 D) and Satun Province, Tarutao Island (UCR coll.); one tentative old record of a large female from Malaysia, Sarawak, Kuching, from the NHMUK collection (Fig. 26); an adult female from Tapah, Malaysia (LKCNHM); and a record from Indonesia, Pulau Weh Island off the coast of Sumatra (Fig. 24 C). Hopefully molecular samples from these far-reaching areas can one day be obtained to confirm the identification as Cryptophyllium chrisangi comb. nov. but for now morphologically this is what these records> appear to represent.	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
DB5E76716257505885921E4B977E7CF1.taxon	materials_examined	Material examined. Holotype ♀: " CHINA: Yunnan, Wangtianshu, Mengla County, Xishuangbanna Prefecture, VII- 2016, Legit: Xiao-Yu Zhu ". Deposited in the Kunming Institute of Zoology (KIZ), Yunnan, China.	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
DB5E76716257505885921E4B977E7CF1.taxon	distribution	Distribution. At present only known from the unique holotype collected in China, Yunnan, Wangtianshu, Mengla County, Xishuangbanna Prefecture.	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
DB5E76716257505885921E4B977E7CF1.taxon	description	Description. Female. Coloration. Coloration description is based upon the dried holotype which is somewhat discolored (Fig. 27). Leaf insects are a more vibrant lime green in life and we expect that this specimen was likely originally this color. The holotype female is mostly tan to pale green throughout and appears to not have any natural brown patches of color as are sometimes present on the lobes or thorax. Morphology. Head. Head capsule about as long as wide, vertex with moderately spaced small granulation, and two notable tubercles, one on each side of the sagittal plane near the midline of the head capsule which are larger than the rest, but not as prominent as the posteromedial tubercle (Fig. 27 D). The posteromedial tubercle is about as broad as the two tubercles on the capsule, but the posteromedial tubercle is notably taller and pointed (easily seen from the lateral aspect; Fig. 27 F). Frontal convexity broad but not very large, with an irregularly lumpy surface and sparse transparent setae throughout. Compound eyes only slightly protruding from the head capsule and only occupying ca. ⅕ of the head capsule length (Fig. 27 D). Ocelli absent. Antennae. Antennae consisting of nine segments, with the terminal segment slightly shorter than the length of the preceding two segments' lengths combined (Fig. 27 B). Antennomeres I-VIII sparsely marked with small transparent setae, the terminal antennomere is covered in stout, brown setae. Thorax. Pronotum with a distinctly concave anterior margin and nearly straight lateral margins, which converge to a straight posterior margin that is half the width of the anterior margin (Fig. 27 D). The pronotum surface is marked with slight granulation, with a prominent pit in the center, and distinct furrows anterior and posterior to the pit and slight furrows lateral to the central pit (Fig. 27 D). The pronotum has a prominent anterior rim which is marked throughout by minute setae and moderately formed lateral rims, and a posterior rim which is weakly formed (Fig. 27 D). Prosternum with moderate nodes, irregularly sized and spaced. Mesosternum with similar nodes as on the pronotum, but only along the margins and on the anterior half of the sagittal plane, the remainder of the surface is rather smooth. Metasternum with an irregularly lumpy surface, no strong nodes. Prescutum slightly longer than wide and with nearly parallel lateral margins (Fig. 27 D). Lateral rims with eight or nine irregularly shaped but short tubercles with various small nodes mixed throughout, giving the margins a rough textured appearance (Fig. 27 D). Prescutum anterior rim distinct, but not strongly raised above the prescutum surface, the rim has a granular surface, no distinct sagittal tubercle larger than the rest (Fig. 27 F). Prescutum surface with irregular nodes throughout, with those along the sagittal plane slightly more prominent (Fig. 27 D). Mesopleura beginning slightly posterior to the anterior margin of the prescutum and evenly diverging; lateral margin with three or four larger tubercles throughout the length, and ten or eleven smaller node-like tubercles interspersed (Fig. 27 D). Face of the mesopleura with granulation throughout and slight wrinkling on areas where the nodes are less prominent (Fig. 27 F). Wings. Tegmina long, extending three quarters of the way through abdominal segment VII. The subcosta (Sc) is the first vein in the forewing and runs parallel with the wing for the first half and then distinctly bends towards the distal margin where it terminates ca. 1 / 4 of the way through the wing length. The radius (R) fills approximately the anterior ⅓ of the wing as two subparallel branched veins; radius 1 (R 1) terminates ca. ⅓ of the way through the wing length, and the radial sector (Rs) terminates posterior to the widest portion of the tegmina, just past the midline. There is a thinner continuation of the radius following the prominent Rs branching which continues on as a short R-M crossvein that does not appear to solidly connect the two veins. The media (M) is bifurcate with both the media anterior (MA) and media posterior (MP) terminating close to the posterior ⅓ of the wing. The cubitus (Cu) is also bifurcate, branching near the posterior 1 / 4 of the wing into the cubitus anterior (CuA) and cubitus posterior (CuP) which both terminate at or very near the wing posterior apex. The first anal vein (1 A) is simple and fuses with the cubitus early on, only slightly past the branching distance of the R 1 from the R. Alae well developed, 58.5 mm long, only a little shorter than the tegmina. Abdomen. Segments II through the anterior ⅓ of IV gradually diverging, with the posterior ⅔ of segments IV and V parallel-sided. Segments VI-X are gradually converging to the broadly rounded apex, giving the abdomen a smooth spade-shaped appearance (Fig. 27 A). Genitalia. Subgenital plate starts at the anterior margin of segment VIII, is broad, and extends ca. ⅔ of the way onto segment X, ending in a fine point (Fig. 27 G). Gonapophyses VIII are long and moderately broad, slightly exceeding the apex of abdominal segment X (Fig. 27 G); gonapophyses IX are thinner and shorter and are concealed below the larger gonapophyses VIII. Cerci flat, not strongly cupped, with a granular surface and rough granular lateral margins (Fig. 27 E). Legs. Profemoral exterior lobes slightly broader than the interior lobe, roundly arcing from end to end in a broad obtuse angle (Fig. 27 C). Edge of the profemoral exterior lobe with a highly granular surface on the proximal margin, and the distal margin has less granulation, but does have five or six small but notable teeth (Fig. 27 C). Profemoral interior lobe ca. 3 x as wide as the greatest width of the profemoral shaft, and with a distinct obtuse angle and a distal margin marked by four or five prominent serrate teeth, with a larger gap between the middle teeth (Fig. 27 C). Mesofemoral exterior lobe arcs from end to end with a distinct bend near the center slightly weighted towards the distal half and marked with two or three dulled serrate teeth on the distal half only. Interior mesofemoral lobe is slightly narrower than the exterior lobe. Mesofemoral interior lobe arcs end to end with five or six serrate teeth on the distal half of the arc only. Metafemoral interior lobe narrow, arcing end to end, and marked with five or six serrate teeth and slight granulation on the distal half of the lobe only. Metafemoral exterior lobe is thin and smooth, hugging the metafemoral shaft and lacks teeth. Pro-, meso-, and meta- tibiae lacking exterior lobes. Protibial interior lobe spans the entire length, is ca. 2 x the width of the shaft, and is roundly triangular with the widest portion on the distal half (Fig. 27 A). Meso- and metatibiae lacking interior lobes. Measurements of holotype female [mm]. Length of body (including cerci and head, excluding antennae) 107.0, length / width of head 8.5 / 7.0, antennae 5.0, pronotum 6.3, mesonotum 7.0, length of tegmina 67.5, length of alae 58.5, greatest width of abdomen 32.0, profemora 27.0, mesofemora 19.2, metafemora 20.0, protibiae 17.4, mesotibiae 18.0, metatibiae 18.5.	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
DB5E76716257505885921E4B977E7CF1.taxon	etymology	Etymology. Noun, named for the artistic company " DAPARO ", owned by Daparo-Yeung which is well-known for their beautiful natural history themed brooches. Several years ago, DAPARO even produced a beautifully crafted leaf insect themed brooch which helped to shed light on these beautiful creatures and bring them into the public eye.	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
37ECB48AD86050F7AB814CAADB108F15.taxon	materials_examined	Material examined. We examined the holotype female from within the Beijing Agricultural University from detailed photographs taken by Yu-Chen Zheng (China Agricultural University, China). Additionally, we examined a tentatively identified male from " Yunnan China: Qinglangdang, Dulongjiang Township, Gongshan County, Nujiang Prefecture, II- 2016, Local " (Coll ZD). A tentative male specimen collected very near the type locality was used in our molecular analysis, which cannot be confidently confirmed as Cryptophyllium drunganum comb. nov. due to extreme sexual dimorphism of the phylliids and lack of a fresh tissue sample from a true Cryptophyllium drunganum comb. nov. female (Fig. 29).	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
37ECB48AD86050F7AB814CAADB108F15.taxon	distribution	Distribution. At present only known from northern Yunnan Province, from the type locality of Nujian Lisu Autonomous Prefecture, Gongshan County (Drung-Nu), and our tentative male Cryptophyllium drunganum comb. nov. from Du Longjiang Township, Qing Lang Dang in the same prefecture.	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
B7FA223E233555279DF4863A17B08329.taxon	materials_examined	Material examined. Holotype ♀: INDONESIA: Wangi-wangi Island. Collected prior to 2020, but no exact date given. Deposited in the Montreal Insectarium (IMQC).	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
B7FA223E233555279DF4863A17B08329.taxon	distribution	Distribution. At present only known from the small Indonesian island of Wangi-wangi off the east coast of Buton Island.	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
B7FA223E233555279DF4863A17B08329.taxon	description	Description. Female. Coloration. At present we only have the dried holotype female to describe the color from which has a bit of rot through the legs, head, thorax, and the central area of the abdomen. The rotten areas are brown but are assumed to have been green in life. The remainder of the female is lime-green in color throughout, with no indication of natural brown patches (which even on somewhat rotten specimens can generally be identified) but this female appears to have been uniform green in life. Morphology. Head. Head capsule slightly longer than wide, vertex with minimal granulation throughout the surface, all relatively well-spaced with no areas on the head tightly packed. The posteromedial tubercle is the most prominent feature on the vertex of the head capsule. Frontal convexity broad and stout, shorter than the length of the first antennomere, and with a lumpy surface marked by few short transparent setae. Compound eyes not particularly large, only slightly protruding from the head capsule, taking up ca. 1 / 4 of the length of the lateral head capsule margins (Fig. 30 A). Ocelli absent. Antennal fields slightly wider than and about as long as the length of the first antennomere. Antennae. Antennae consist of nine segments, with the terminal segment approximately the same length as the preceding two segments' lengths combined (Fig. 30 B). Antennomeres I-III sparsely marked with small transparent setae (with the longest on the first segment), the terminal two antennomeres are densely covered in stout, brown setae. Thorax. Pronotum with a gently concave anterior margin and slightly convex lateral margins, which converge to a straight posterior margin that is half the width of the anterior margin (Fig. 30 D). The pronotum surface lacks granulation but is slightly lumpy, with only a prominent pit in the center, and slight furrows anterior and lateral to the pit (Fig. 30 D). The pronotum has a prominent anterior rim and moderate lateral and posterior rims (Fig. 30 D). Prosternum and mesosternum with numerous nodes throughout the surface, all about the same size and spacing throughout. Metasternum with slightly less granulation but they are slightly larger than those on the pro- and mesosternum. Prescutum longer than wide, with a slightly broader anterior margin (Fig. 30 D). Lateral rims with 6 - 8 prominent tubercles with various small, lumpy granules interspersed throughout the margins (Fig. 30 D). Prescutum anterior rim prominent but not strongly protruding, surface is granular and lacks a prominent sagittal spine (Fig. 30 D). Prescutum surface without a strongly raised sagittal crest, instead the surface is only slightly raised along the sagittal plane. The prescutum surface has moderate granulation throughout ranging in size from small to medium with irregular spacing (Fig. 30 D). Mesopleura not spanning the entire length, instead with the anterior ⅓ narrow and only starting to fan out near the midline of the prescutum length. Mesopleura lateral margins with five or six larger, sharp tipped tubercles with an additional five or six smaller nodes interspersed throughout (Fig. 30 D). Face of the mesopleura lacking granulation, but instead highly wrinkled and with two notable pits, one on the anterior ⅓ and one nearer the posterior ⅓ (Fig. 30 D). Wings. Tegmina long, reaching nearly to the posterior margin of abdominal segment VII. The subcosta (Sc) is the first vein in the forewing and runs subparallel with the wing for the first half of its length, and then bends towards the wing margin for the second half. The radius (R) spans the central portion of the tegmina with two subparallel branched veins. The first radius (R 1) branches ca. ⅗ of the way through the radius length and terminates ca. ⅓ of the way through the wing length. The radial sector (Rs) branches from the end of the radius and runs angled to the wing margin where it terminates just posterior to the wing midline length. There is a weak continuation of the radius following the prominent radial sector branching which continues on as a short and thin radius to media crossvein (R-M). The media (M) is simply bifurcate with both the media anterior (MA) and media posterior (MP) terminating close to the posterior 1 / 4 of the wing. The cubitus (Cu) runs throughout the entire wing length simply, and then near the posterior ⅕ of the wing becomes bifurcate into the cubitus anterior (CuA) and cubitus posterior (CuP) which both terminate at or very near the wing posterior apex. The first anal vein (1 A) is simple and fuses with the cubitus early on, near where the radial sector branches from the radial. Alae of moderate length, reaching abdominal segment IV. Abdomen. Abdominal segments II through the anterior 1 / 2 of IV diverging, with the middle of segment IV the widest segment. Segments V-VII gently converging, with segment VII ending with a distinct lobe which bends inward to a notably narrower segment VIII. Segments VIII-X converging to a broad apex. Genitalia. Subgenital plate starts at the anterior margin of segment VIII, is broad, and only extends ⅓ of the way under segment X, ending in a fine point (Fig. 30 E). Gonapophyses VIII are long (exceeding the tip of the abdomen but not as long as the tips of the cerci) and moderately broad (together side by side are about as broad as the subgenital plate; Fig. 30 E). Gonapophyses IX are smaller and shorter, and mostly covered from view by the notably larger gonapophyses VIII. Cerci flat, with a granular surface throughout and few setae on the ventral surface, the dorsal surface has thin, transparent setae throughout the surface (Fig. 30 E). Legs. Profemoral exterior lobes broad (ca. 2 x width of the interior lobe) and approximately right angled (Fig. 30 A). Proximal edge of the exterior profemoral lobe gently undulates giving this margin a slightly wavy appearance, whereas the distal margin is nearly straight, both margins have notable serration throughout their lengths (Fig. 30 A). Profemoral interior lobe with a slightly obtuse angle and marked with five large, serrate teeth with looping gaps between them. These teeth are arranged somewhat into a two-one-two pattern with the gaps between these sets larger. Mesofemoral exterior lobe arcs from end to end and is almost evenly weighted on both sides, but with the broadest point just off center to the distal side of the midline, and on the distal half only marked with three small serrate teeth. Interior mesofemoral lobe arcs end to end, is ca. 1 / 4 narrower than the exterior lobe, not as strongly angled, and marked with six or seven teeth on the distal half. Metafemoral interior lobe arcs end to end but is notably wider on the distal ⅔ of the lobe and this wider portion is marked by 11 or 12 serrate teeth. Metafemoral exterior lobe is thin and smooth, hugging the metafemoral shaft and marked with one or two small, rounded teeth on the distal edge. Protibiae lacking an exterior lobe. Protibiae interior lobe spans the entire length of the protibiae as a rounded scalene triangle with the widest portion on the distal ⅓. Mesotibiae and metatibiae simple, lacking exterior and interior lobes. Measurements of holotype female [mm]. Length of body (including cerci and head, excluding antennae) 94.5, length / width of head 7.8 / 7.5, antennae 5.0, pronotum 6.6, mesonotum 7.9, length of tegmina 59.1, length of alae 31.6, greatest width of abdomen 41.3, profemora 20.7, mesofemora 16.5, metafemora 21.2, protibiae 14.0, mesotibiae 12.1, metatibiae 16.4.	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
B7FA223E233555279DF4863A17B08329.taxon	etymology	Etymology. Noun, Greek in origin. Relating to the tenth labor of Heracles (apparently a favorite story of Gray (1843) as four of his therein described species came from this myth) in which Heracles was tasked with capturing the red cattle from the monster Geryon (for which Gray named Phyllium geryon Gray, 1843). Before Heracles sailed out to the island Erythia where Geryon and his red cattle lived, he rested on the mainland and forgot to tie up his horses. When he eventually found them, they were in a cave with an Echidna (a monster which is described as women from the waist up and snake on her lower half). She refused to give back his horses unless he lay with her, which being a classic Greek hero, he did. Before he left the Echidna, she told him that she was pregnant with three of his sons and asked him which of the three should rule her lands one day. Heracles then left her with his bow and a girdle, and told her that whichever of his three sons could draw the bow and wear the girdle best would inherit her land and the other two should be banished. Those three sons were Agathyrsus, Gelonus, and Scythes (for which Gray named Phyllium agathyrsus Gray, 1843; Phyllium gelonus Gray, 1843; and Phyllium scythe Gray, 1843). We felt that this species could help to finish telling the story which Gray was so fond of. With Phyllium geryon being a species from the Philippines, one biogeographical bridge for species to the Philippines is from Sulawesi through the Sangihe Islands (Evans et al. 2003), therefore as a steppingstone to the Philippines we felt there should be an Echidna along the route to the Philippines where Phyllium geryon can be found.	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
C8E17D58FF8F562DAD931973898C355E.taxon	materials_examined	Material examined. Holotype ♂: " VIETNAM: Quang Ngai Province, Bato Mt. 900 m. elv: May 2015 (Coll RC 16 - 114) ". Molecularly sampled within our analysis. Deposited in the Montreal Insectarium (IMQC). Paratypes (2 ♂): 1 ♂: " Ngoc Linh, Kon Tum Prov. Vietnam, 1700 m, VI. 2016, leg. Luong coll. TB- 05 - 134 ' (Coll TB) • ♂ nymph: " VIETNAM: Lam Dong, Bao Lam, Da Tom: March 2016 (Coll RC 16 - 236) ", molecularly sampled within our analysis (Coll RC).	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
C8E17D58FF8F562DAD931973898C355E.taxon	distribution	Distribution. Presently only known from central and southern Vietnam, no other specimens are presently known. The two Vietnamese provinces that this species is known from are Quang Ngai and Lam Dong Provinces.	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
C8E17D58FF8F562DAD931973898C355E.taxon	etymology	Etymology. Patronym. Dedicated to David Faulkner, California, United States. Forensic entomology mentor to RTC and dear friend.	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
875D19DBF6005EC786793DEA37E816C1.taxon	materials_examined	Material examined. Holotype ♂: " Coll. I. R. Sc. N. B., Ex breeding Tim Bollens, 2018, Coll. I. R. Sc. N. B., Vietnam, Lam Dong prov., Bidoup-Nui Ba N. P., 12 ° 26 ' N, 108 ° 30 ' E, 21 - 25. vii. 2014, Leg. J. Constant and J. Bresseel, GTI Project, I. G.: 32.779 ". Deposited in the Royal Belgian Institute of Natural Sciences (RBINS). Paratypes: (26 ♀♀, 49 ♂♂, 78 eggs) • 1 ♂: " Coll. I. R. Sc. N. B., Ex breeding Tim Bollens, 2018, Coll. I. R. Sc. N. B., Vietnam, Lam Dong prov., Bidoup-Nui Ba N. P., 12 ° 26 ' N, 108 ° 30 ' E, 21 - 25. vii. 2014, Leg. J. Constant and J. Bresseel, GTI Project, I. G.: 32.779 " [vomer dissected] (RBINS) • 5 ♀♀, 4 ♂♂: " Coll. I. R. Sc. N. B., Ex breeding Tim Bollens, 2018, Coll. I. R. Sc. N. B., Vietnam, Lam Dong prov., Bidoup-Nui Ba N. P., 12 ° 26 ' N, 108 ° 30 ' E, 21 - 25. vii. 2014, Leg. J. Constant and J. Bresseel, GTI Project, I. G.: 32.779 " (RBINS) • 1 ♀, 1 ♂: " Coll. I. R. Sc. N. B., Ex breeding Tim Bollens, 2018, Coll. I. R. Sc. N. B., Vietnam, Lam Dong prov., Bidoup-Nui Ba N. P., 12 ° 26 ' N, 108 ° 30 ' E, 21 - 25. vii. 2014, Leg. J. Constant and J. Bresseel, GTI Project, I. G.: 32.779 " (VNMN) • 1 ♀, 5 ♂♂: " Coll. I. R. Sc. N. B., Ex breeding Tim Bollens, 2019, Coll. I. R. Sc. N. B., Vietnam, Lam Dong prov., Bidoup-Nui Ba N. P., 12 ° 26 ' N, 108 ° 30 ' E, 21 - 25. vii. 2014, Leg. J. Constant and J. Bresseel, GTI Project, I. G.: 32.779 " (RBINS) • 1 ♀, 1 ♂: " Coll. I. R. Sc. N. B., Ex breeding Bruno Kneubuehler, 2017, Coll. I. R. Sc. N. B., Vietnam, Lam Dong prov., Bidoup-Nui Ba N. P., 12 ° 26 ' N, 108 ° 30 ' E, 21 - 25. vii. 2014, Leg. J. Constant and J. Bresseel, GTI Project, I. G.: 32.779 " (RBINS) • 6 ♀♀, subadult ♀, ♂: " Coll. I. R. Sc. N. B., Vietnam, Lam Dong prov., Bidoup-Nui Ba N. P., 12 ° 26 ' N, 108 ° 30 ' E, 21 - 25. vii. 2014, Leg. J. Constant and J. Bresseel, GTI Project, I. G.: 32.779 " (RBINS) • 1 ♂; " Vietnam, Lam Dong, Bao Loc " (molecular sample SLT 005 within our analysis) (Coll SLT) • 3 ♂♂; " ex Zucht F. Hennemann 2019, Herkunft: Z-Vietnam, Lam Dong Prov., Dam Rong, Bidoup Nui Ba NP, leg. Bresseel and Constant VII. 2014 [ex coll. FH] " (Coll SLT) • 1 ♀; " VIETNAM, Lam Dong Prov., Dam Rong, Bidoup-Nui Ba NP, Leg. Bresseel and Constant VII. 2014. Ex breeding F. Hennemann 2015 - 2017. Ex coll. F. H. Hennemann (Germany); Coll RC 18 - 226 " (Coll RC) • 4 ♂♂; " ex Zucht F. Hennemann 2019, Herkunft: Z-Vietnam, Lam Dong Prov., Dam Rong, Bidoup Nui Ba NP, leg. Bresseel and Constant VII. 2014 [ex coll. FH] ", Coll RC 20 - 099 - 20 - 102 (Coll RC) • 1 ♂; " VIETNAM, Dak Lak Prov., May 2018. Coll RC 18 - 408 " (Coll RC) • 1 ♂; " VIETNAM, Lam Dong Prov., Dam Rong, Bidoup-Nui Ba NP, Leg. Bresseel and Constant VII. 2014. Ex breeding F. Hennemann 2015 - 2017. Ex coll. F. H. Hennemann, Germany; Coll RC 18 - 225 " (Coll RC) • 1 ♂; " VIETNAM, Lam Dong Prov., Bao Loc, October 2016, Coll RC 16 - 248 " (Coll RC) • 1 ♂; " VIETNAM, Lam Dong Prov., Bao Loc, May 2016, Coll RC 17 - 264 " (Coll RC) • 3 ♂♂; " VIETNAM, Lam Dong Prov., Bao Loc, July 2017 ", Coll RC 17 - 266, 17 - 267, 17 - 268 (Coll RC) • 1 ♂; " VIETNAM, Lam Dong Prov., Bao Loc, June 2017. Coll RC 17 - 265 " (Coll RC) • 2 eggs: " VIETNAM, Lam Dong Prov., Dam Rong, Bidoup-Nui Ba NP, Leg. Bresseel and Constant VII. 2014. Ex breeding F. Hennemann 2015 - 2017. Ex coll. F. H. Hennemann, Germany "; Coll RC 18 - 238 and 18 - 239 (Coll RC) • 20 eggs: " VIETNAM: Lam Dong Prov., Bidoup Nui Ba N. P., culture from Maxime Ortiz (France), 2018 ". Coll RC 18 - 343 - 18 - 362 (Coll RC) • 7 ♀♀, 1 ♂; " VIETNAM: Lam Dong Prov., Bidoup Nui Ba N. P., bred by Bruno Kneubu ̈ hler (Switzerland), circa 2016 " (Coll OC) • 3 ♀♀, 7 ♂♂, 28 eggs: " ex Zucht F. Hennemann 2018, Herkunft: Z-Vietnam, Lam Dong Prov., Dam Rong, Bidoup Nui Ba NP, leg. Bresseel and Constant VII. 2014 " [coll. FH, No's 0896 - 1 to 10, E 1] (Coll FH) • 1 ♀, 7 ♂♂, 1 ♂ nymph n 5, 28 eggs; " ex Zucht F. Hennemann 2019, Herkunft: Z-Vietnam, Lam Dong Prov., Dam Rong, Bidoup Nui Ba NP, leg. Bresseel and Constant VII. 2014 " [coll. FH, No's 0896 - 11 to 19, E 2], (Coll FH) • 1 ♀: " ex Zucht F. Hennemann 2020, Herkunft: Z-Vietnam, Lam Dong Prov., Dam Rong, Bidoup Nui Ba NP, leg. Bresseel and Constant VII. 2014 " [coll. FH, No's 0896 - 20], (Coll FH) • 3 ♀♀, 4 ♂♂; " Vietnam, Lam Dong Prov., Dam Rong, Bidoup Nui Ba, bred by Maxime Ortiz, France, circa 2020 " (Coll MO).	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
875D19DBF6005EC786793DEA37E816C1.taxon	distribution	Distribution. Only presently known from southern Vietnam, from the provinces of Lam Dong and Dak Lak.	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
875D19DBF6005EC786793DEA37E816C1.taxon	description	Description. Female. Coloration. Coloration is variable, as when it was bred in captivity females were almost uniformly pale green, with only slightly orange / tan areas along the profemoral lobes, antennae, eyes, thorax, and terminal abdominal margins (Fig. 33 A). In contrast, the paratype female when she was found in the wild was quite colorful with a vibrant green body with highlights of red throughout the antennae, legs, thorax, tegmina, and abdomen (Fig. 32 A), much darker and more plentiful than on captive bred specimens. Morphology. Head. Head capsule longer than wide, vertex relatively smooth, with the only notable feature the posteromedial tubercle which is not notably broad but is significantly raised above the head capsule (Fig. 34 E). Frontal convexity broad and stout, with a lumpy surface, and with several setae throughout (Fig. 34 E). Compound eyes slightly protruding from the head capsule, not notably large, only taking up slightly <1 / 4 of the length of the lateral head capsule margins (Fig. 34 E). Ocelli absent. Antennal fields slightly wider than the first antennomere (Fig. 34 C). Antennae. Antennae consisting of nine segments, with the terminal segment about as long as 21 / 2 x the preceding segments lengths (Fig. 34 C). Antennomeres I-VIII sparsely marked with small transparent setae, the terminal antennomere and the distal margin of antennomere VIII has darker, shorter, and denser setae than the other segments (Fig. 34 C). Thorax. Pronotum with a slightly concave anterior margin and slightly convex lateral margins, which converge to a straight posterior margin that is half the width of the anterior margin (Fig. 34 E). The pronotum surface and moderately formed pronotum rims are only slightly lumpy, lacking significant granulation, with only a prominent pit in the center, and slight furrows anterior, posterior, and lateral to the pit (Fig. 34 E). Prosternum with moderate granulation, mesosternum anterior half and lateral margins with moderate granulation (Fig. 34 B). Metasternum relatively smooth, lacking notable nodes. Prescutum slightly longer than wide, lateral rims with four larger nodes and four or five nodes interspersed throughout (Fig. 34 E). Prescutum anterior rim prominent but not strongly protruding, surface marked throughout with irregular granulation, no prominent singular sagittal spine present (Fig. 34 F). Prescutum surface covered densely by small tubercles and numerous nodes, with those along the sagittal plane the largest (Fig. 34 E). Mesopleura beginning slightly posterior to the anterior margin of the prescutum and evenly diverging; lateral margin with five larger tubercles, and five or six smaller tubercles interspersed unevenly throughout (Fig. 34 E). Face of the mesopleura slightly wrinkled, with a distinct pit near the anterior margin and one near the center (Fig. 34 E). Wings. Tegmina long, reaching 1 / 2 way through abdominal segment VII. The subcosta (Sc) is the first vein in the forewing and runs parallel with the tegmina lateral margin for the first half of the vein, then bends gently and runs to the to the lateral margin of the wing where it terminates ca. ⅓ through the length. The radius (R) spans the central portion of the forewing with two subparallel branched veins; radius 1 (R 1) terminates slightly proximal to the midline, and the radial sector (Rs) terminates ca. ⅔ of the way through the wing length. There is a weak continuation of the radius following the prominent Rs branching which continues on as a short and thinner R-M crossvein that does not solidly connect the two veins as it reaches the media. The media (M) is simply bifurcate with both the media anterior (MA) and media posterior (MP) terminating on the posterior 1 / 4 of the wing. The cubitus (Cu) is also bifurcate, branching near the posterior ⅕ of the wing into the cubitus anterior (CuA) and cubitus posterior (CuP) which both terminate at or very near the wing posterior apex. The first anal vein (1 A) is simple and fuses with the cubitus early on, halfway between the branching of the R 1 and the Rs. Alae are highly reduced, not reaching the anterior margin of abdominal segment II. Abdomen. Abdominal segments II through the anterior half of IV diverge uniformly towards the posterior. The posterior half of IV and segment V are parallel-sided and are the widest portion of the abdomen. Segment VI and VII are variable depending on the environmental conditions, with wild collected females (such as in the paratype; Fig. 32 A) having segments VI and VII strongly converging and VII ending with a small bulge on the posterior margin with a final width about the same as the anterior margin of segment VIII, vs. captive bred individuals generally having VI and VII subparallel and ending with VII having a distinct lobe which is notably wider than the anterior margin of segment VIII (Fig. 33 A). Segments VIII-X converge uniformly to the rounded apex. Genitalia. Subgenital plate starts at the anterior margin of segment VIII, is long and narrow reaching ca. 1 / 2 onto segment X (Fig. 34 H). Gonapophyses VIII are long and significantly broad, each about as wide as the subgenital plate projection, with their tips reaching the apex of segment X, gonapophyses IX are smaller and slender, hidden below the gonapophyses VIII (Fig. 34 H). Cerci flat, not strongly cupped, with a heavily granular surface and a setae throughout (Fig. 34 H). Legs. Profemoral exterior lobes notably wider than the interior lobe with a rounded obtuse angle (Fig. 34 D). Exterior lobe margin is not marked by teeth and is instead rather smooth or at most slightly granular (Fig. 34 D). Profemoral interior lobe ca. 2 x as wide as the greatest width of the profemoral shaft, with an obtuse angle, and marked with five prominent teeth arranged in a two-one-two pattern with large looping gaps between the teeth (Fig. 34 D). Mesofemoral exterior lobe arcs from end to end in a slightly bent lobe slightly weighted to the distal 1 / 2 and marked with two or three small serrate teeth on the distal 1 / 2 only. Interior and exterior lobes are of a similar width, or the exterior is slightly wider. Mesofemoral interior lobe arcs smoothly end to end, is marked with six or seven serrate teeth only on the distal half of the arc, and is about as wide as the mesofemoral shaft. Metafemoral interior lobe arcs end to end, but is slightly wider on the distal half, and has six or seven serrate teeth on the distal half of the lobe only. Metafemoral exterior lobe is thin and smooth, hugging the metafemoral shaft and lacks dentation. Protibiae interior lobe spans the entire length of the protibiae and is at ca. 2 x as wide as the protibial shaft. The lobe is distinctly triangular with the broadest point distal to the midline (Fig. 34 D). Pro-, meso-, and meta- tibiae lacking exterior lobes; meso-, and meta- tibiae lack interior lobes as well. Measurements of paratype females [mm]. Length of body (including cerci and head, excluding antennae) 81.8 - 94.1, length / width of head 8.0 - 9.7 / 6.4 - 7.4, antennae 3.3 - 4.2, pronotum 4.6 - 5.7, mesonotum 6.7 - 9.1, length of tegmina 45.4 - 57.5, length of alae 6.0 - 7.2, greatest width of abdomen 30.7 - 34.4, profemora 17.9 - 21.6, mesofemora 13.8 - 15.9, metafemora 16.2 - 18.6, protibiae 11.1 - 11.8, mesotibiae 10.1 - 10.7, metatibiae 13.6 - 13.7. Male. Coloration. Coloration description based on live captive reared males (Fig. 33 B) and the male nymphs found in the wild (Fig. 32 B). Captive males are generally mostly pale green throughout with the margins of the legs, thorax, and abdomen a reddish brown, and slightly transparent eye spots on abdominal segment V. The wild caught nymphs had these areas typically darker, more dark red than reddish brown and dark eyespots on abdominal segment V. Morphology. Head. Head capsule about as long as wide, with a vertex that is weakly granular; posteromedial tubercle small but notable and slightly raised above the head capsule (Fig. 35 D). Frontal convexity stout with a few short setae near the apex. Compound eyes large and bulbous, occupying ca. ⅖ of the head capsule lateral margins and significantly protruding from the head capsule (Fig. 35 D). There are three well-developed ocelli between and slightly posterior to the compound eyes. Antennae (including the scapus and pedicellus) consists of 23 segments. The scapus and pedicellus are bare, all other segments are covered in dense, thin, pale setae that are as long as or longer than the antennae segment is wide. The terminal three segments have shorter darker setae. Thorax. Pronotum anterior margin is distinctly concave and lateral margins are slightly convex and converge to a straight posterior margin that is slightly> 1 / 2 width of the anterior rim (Fig. 35 D). Anterior and lateral margins of the pronotum have moderate rims and the posterior margin lacks a rim (Fig. 35 D). Face of the pronotum is slightly lumpy, has a distinct sagittal furrow, a pit just posterior to the center, a moderate perpendicular furrow just anterior to the central pit, and has a distinct pit on each side near the anterior margin (Fig. 35 D). The prosternum surface is slightly granular. The mesosternum surface is marked densely with prominent nodes, with the largest along the sagittal plane and more prominent on the anterior margin, posterior margin with less prominent and slightly smaller nodes. Prescutum slightly longer than wide, with lateral margins slightly converging to the posterior which is ca. 3 / 4 the width of the anterior rim (Fig. 35 D). Lateral rims with nine or ten nodes of slightly varying size, none very large or prominent, but each marked with a single stiff seta protruding from the tip. The surface of the prescutum is notably granulose along the sagittal plane with lateral surfaces rather smooth (Fig. 35 D). Prescutum anterior rim slightly granulose with no distinct central tubercle (Fig. 35 E). Mesopleura narrow, only gradually diverging from the anterior to the posterior (Fig. 35 D). Lateral margin granulose throughout, with only four or five slightly larger than the rest, but not significantly larger. The largest nodes along the mesopleura have a singular seta protruding from them like those on the prescutum margins. Face of the mesopleura smooth but slightly wrinkled and with two faint pits, one on the anterior margin and one near the middle of the mesopleura. Wings. Tegmina short, only reaching the anterior margin of abdominal segment III. Tegmina wing venation: the subcosta (Sc) is the first vein and terminates the earliest, ca. ⅖ of the way through the overall tegmina length. The radius (R) spans the entire length of the tegmina with the first radius (R 1) branching ca. ⅓ of the way through the wing length and terminating just posterior to the middle of the wing, the second radius (R 2) branches near the distal ⅓ of the wing, and then the radial sector (Rs) runs straight to the tegmina apex and terminates. The media (M) spans the entire length of the tegmina, terminating at the wing apex as the media anterior (MA) with the first media posterior (MP 1) beginning and terminating near the tegmina mid length followed by the second media posterior (MP 2) which begins ca. ⅔ of the way through the tegmina length and terminates near the posterior quarter of the wing. The cubitus (Cu) runs through the wing surface angled until it meets the margin ca. ⅓ of the way through the tegmina length and then runs along the margin as the two media posterior veins then meet it and fuse and the cubitus continues to run nearly to the wing apex. The first anal (1 A) vein runs subparallel to the cubitus until it meets it slightly> ⅓ of the way through the tegmina length and fuses with it. Alae well-developed in an oval fan configuration, reaching to the anterior margin of abdominal segment IX or halfway through it. Alae wing venation: the costa (C) is present along the entire foremargin giving stability to the wing. The subcosta (Sc) spans ca. ⅔ of the wing length and is mostly fused with the radius in the beginning but terminates when it meets the costa. The radius (R) spans the entire wing and branches ca. ⅖ of the way through into the radius 1 (R 1) and radial sector (Rs) which run slightly diverging for the first ⅓ of their length, parallel for the central portion until the terminal quarter where they converge and terminate on the wing margin near each other but not touching. The media (M) branches early, ca. ⅙ of the way through the wing into the media anterior (MA) and the media posterior (MP) which run parallel with each other until the distal ⅙ of the wing where the media posterior fuses with the media anterior which then run fused together to the wing margin. The cubitus (Cu) runs unbranched and terminates at the wing apex. Of the anterior anal veins, the first anterior anal (1 AA) fuses with the cubitus near the point where the media branches into the media anterior and media posterior and then the first anterior anal branches from the cubitus ⅔ of the way through the wing length where it uniformly diverges from the cubitus until it terminates at the wing margin. The anterior anal veins two-seven (2 AA- 7 AA) have a common origin and run unbranched in a folding fan pattern of relatively uniform spacing to the wing margin. The posterior anal veins (1 PA- 7 PA) share a common origin separate from the anterior anal veins and run unbranched to the wing margin with slightly thinner spacing than the anterior anal veins. Abdomen. Segment II, parallel-sided, segment III and the anterior half of IV diverging, the posterior 1 / 2 of IV through segment V either parallel-sided or slightly diverging, VI through the apex converging to a blunt rounded apex. The margins of segments VIII-X have a line of setae along them (Fig. 35 F). Genitalia. Poculum broad and rounded, ending in a rounded apex that passes beyond the anterior margin of segment X (Fig. 35 G). Cerci long and slender, with ca. 1 / 2 of their length extending out from under the anal abdominal segment. The cerci are slightly cupped, with a granulose surface and numerous short setae throughout (Fig. 35 F). Vomer broad and stout with rounded sides converging to the apex which is armed with two upwards turning hooks, one at the apex which is larger and one lateral to it which is slightly smaller (Fig. 5 H). Legs. Profemoral exterior lobe a rounded arc without a distinct angle, slightly thinner than the interior lobe (at its widest slightly> 2 x the greatest width of the profemoral shaft), and with the distal half marked by three or four small but sharp anteriorly pointing teeth (Fig. 35). Profemoral interior lobe roundly triangular, at its widest ca. 21 / 2 x as wide as the profemoral shaft at its widest. The profemoral interior lobe is generally marked with five, serrate, anteriorly pointing teeth arranged in a two-one-two pattern, with shallow looping gaps between them, and occasionally marked with an extra tooth within the set (like as can be seen in Fig. 35 C). Mesofemoral exterior lobe arcs end to end but with a more prominent bend near the distal ⅓ of the lobe which is marked with two or three small serrate teeth on the distal ⅓ only, with the proximal portion of the lobe smooth. The mesofemoral interior lobe at its widest is approximately the same width as the exterior lobe, but the proximal half is slightly wider than the proximal half of the exterior lobe. The distal half of the mesofemoral interior lobe is marked with six or seven small serrate teeth and the proximal half is rather smooth. Metafemoral exterior lobe lacks dentation and has a straight margin along the metafemoral shaft. Metafemoral interior lobe smoothly arcs end to end with six or seven small serrate teeth on the distal 1 / 2 only. Protibiae lacking exterior lobe, interior lobe reaching end to end in a smooth triangle ca. 3 x as wide as the protibial shaft, with the widest point just distal to the midline, and all margins notably marked with short setae throughout (Fig. 35 C). Mesotibiae and metatibiae simple, lacking lobes completely. Measurements of holotype male [mm]. Length of body (including cerci and head, excluding antennae) 58.6, length / width of head 3.7 / 3.3, antennae 32.5, pronotum 2.8, mesonotum 4.5, length of tegmina 16.1, length of alae 42.0, greatest width of abdomen 15.8, profemora 12.0, mesofemora 9.5, metafemora 11.4, protibiae 7.5, mesotibiae 6.3, metatibiae 8.2. Measurements of paratype males [mm]. Length of body (including cerci and head, excluding antennae) 56.0 - 69.4, length / width of head 2.9 - 4.6 / 2.9 - 4.0, antennae 34.5 - 43.9, pronotum 2.5 - 3.0, mesonotum 3.8 - 4.8, length of tegmina 16.4 - 19.8, length of alae 40.5 - 49.9, greatest width of abdomen 14.4 - 18.3, profemora 11.7 - 14.8, mesofemora 9.5 - 12.3, metafemora 11.0 - 13.6, protibiae 7.7 - 9.9, mesotibiae 6.5 - 8.0, metatibiae 8.5 - 10.4. Description of egg (Fig. 36). The lateral surfaces are flat but with the posterior half slightly wider than the anterior half. The dorsal surface is slightly convex, which gives the margin a slight undulating appearance when viewed from the lateral aspect as the middle is thinner than either end of the egg. When viewed from the lateral aspect; the ventral margin is also not straight but is instead with the posterior slightly protruding more than the anterior. All surfaces have numerous small sized pits throughout with short moss-like pinnae interspersed throughout the capsules surfaces with those on the margins and those on the dorsal surface slightly more prominent. Dorsal surface with irregular medium sized pitting and moss-like pinnae around the micropylar plate. Micropylar plate long, ca. 7 / 8 of the overall dorsal surface length, with the widest portion around the micropylar cup. Micropylar plate nearly symmetrical with the anterior and posterior thin and the area around the micropylar cup the widest point. Micropylar cup of moderate size and placed just slightly posterior to the micropylar plate midline. Operculum slightly ovular, with the outer margin with a distinct row of moss-like pinnae surrounding the operculum. Operculum is roundly raised with the height ca. 1 / 2 the operculum width. The overall color is light brown, with the moss-like pinnae sometimes slightly lighter in color.	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
875D19DBF6005EC786793DEA37E816C1.taxon	description	Measurements including the extended pinnae [mm]. Length (including operculum) 2.2, maximum width of capsule when viewed from lateral aspect 1.3 mm, length of micropylar plate 1.1 mm. Newly hatched nymphs. (Fig. 9 I). The body is made up of two general blocks of color, the legs, head, pronotum, and mesonotum are primarily chocolate brown and the mesonotum and abdomen are burnt red in color. Basitarsi are white and the remaining tarsal segments are burnt reddish. The tibiae only have interior lobes which smoothly span the full length and lack exterior lobes. The tibiae interior lobes are brown with two white patches on the proximal half only, the proximal most is notably larger. Interior and exterior femoral lobes are all about the same width, are smoothly arcing, and all have minimal serration. On the profemoral interior lobe there is a notable whitish patch, the exterior lobe and profemoral shaft itself are devoid of prominent white markings. The meso- and metafemoral interior lobes are similar in that they have a small white patch right at the proximal end, and then another white marking ca. ⅖ of the way through the length. The exterior meso- and metafemoral lobes only have one white patch located on the proximal ⅓, but notably wider than the interior lobe white patch. The meso- and metafemoral shafts lack white coloration. The abdomen is slender with segment II through the anterior 1 / 2 of IV diverging and the posterior 1 / 2 of IV-X converging.	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
875D19DBF6005EC786793DEA37E816C1.taxon	etymology	Etymology. Noun, from Greek mythology. Named for the tragic story of Icarus, son of Daedalus. During their escape from the island of Crete, Icarus flew too close to the sun and melted the wax wings his father built. We felt it was fitting that this mythological name is shared with this species that lacks the hindwings within the former Cryptophyllium celebicum species group (characterized by females with well-developed alae).	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
7F2F01FA305959E39E1EC7B005EBDF4C.taxon	materials_examined	Material examined. Holotype ♂: " Coll. I. R. Sc. N. B., collected as nymph, Coll. I. R. Sc. N. B., Cambodia, Koh Kong prov., Tatai, 11 ° 35 ' 13 " N 103 ° 05 ' 50 " E, 9 - 19. x. 2016, day collecting, GTI Project, Leg. J. Constant and J. Bresseel, I. G.: 33.345 (RBINS-PHYLLIUM DNA sample 0002) " [vomer dissected], deposited in RBINS. Paratypes (9 ♀♀, 2 ♂♂): • " Coll. I. R. Sc. N. B., collected as nymph, Coll. I. R. Sc. N. B., Cambodia, Koh Kong prov., Tatai, 11 ° 35 ' 13 " N 103 ° 05 ' 50 " E, 9 - 19. x. 2016, day collecting, GTI Project, Leg. J. Constant and J. Bresseel, I. G.: 33.345, RBINS-PHYLLIUM DNA sample 0001 " (RBINS) • 3 ♀♀, 1 ♂: " Coll. I. R. Sc. N. B., Ex breeding Tim Bollens, 2018, Coll. I. R. Sc. N. B., Cambodia, Koh Kong prov., Tatai, 11 ° 35 ' 13 " N 103 ° 05 ' 50 " E, 9 - 19. x. 2016, day collecting, GTI Project, Leg. J. Constant and J. Bresseel, I. G.: 33.345 " • 2 ♂♂: " Coll. I. R. Sc. N. B., Ex breeding Tim Bollens, 2018, Coll. I. R. Sc. N. B., Cambodia, Koh Kong prov., Tatai, 11 ° 35 ' 13 " N 103 ° 05 ' 50 " E, 9 - 19. x. 2016, day collecting, GTI Project, Leg. J. Constant and J. Bresseel, I. G.: 33.345; ex breeding Tim Bollens, 2018 " • 2 ♀♀: " Coll. I. R. Sc. N. B., CAMBODIA, Siem Reap Prov., Phnom Kulen N. P., Forest near Preah Thom, 26 - 27 - VII- 2006, Leg K. Smets, Y. Oul and D. Jump. " (1 ♀: RBINS; 1 ♀: RUPP) • 1 ♂: " Cambodia, Siem Reap; Kbal Spean, 13 ° 40.858 ' N 104 ° 01.111 ' E, 122 m, 6 - jul- 2015, Hap, Sour, Phauk, Khearn, Chhum, Ly, Lom, Heang, Hok, CA 0028, Lighttrap in the forest with canopy cover. " (RUPP) • 3 ♀, 1 ♂: " Coll. I. R. Sc. N. B., Ex breeding Tim Bollens, 2019, Coll. I. R. Sc. N. B., Cambodia, Koh Kong prov., Tatai, 11 ° 35 ' 13 " N 103 ° 05 ' 50 " E, 9 - 19. x. 2016, day collecting, GTI Project, Leg. J. Constant and J. Bresseel, I. G.: 33.345; ex breeding Tim Bollens, 2019 " (RBINS) • 1 ♀, 1 ♂: " Coll. I. R. Sc. N. B., Ex breeding Tim Bollens, 2019, Coll. I. R. Sc. N. B., Cambodia, Koh Kong prov., Tatai, 11 ° 35 ' 13 " N 103 ° 05 ' 50 " E, 9 - 19. x. 2016, day collecting, GTI Project, Leg. J. Constant and J. Bresseel, I. G.: 33.345; ex breeding Tim Bollens, 2018 " (Coll RC).	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
7F2F01FA305959E39E1EC7B005EBDF4C.taxon	distribution	Distribution. At present only confirmed from two Cambodian provinces, Koh Kong Province (Tatai) and Siem Reap Province (Kbai Spean and Phnom Kulen N. P., Forest Near Prean Thom). It is likely that other nearby localities may also represent this species, but due to a lack of molecular data we cannot at this time confirm them.	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
7F2F01FA305959E39E1EC7B005EBDF4C.taxon	description	Description. Female. Coloration. Coloration description is based upon photographs of living individuals (Fig. 38 A, B) reared by Tim Bollens (Belgium). Overall coloration pale mint green with variable slight highlighting of orange or tan coloration throughout. Compound eyes are slightly more yellow with tan highlights. Antennae are tan. The prescutum and mesopleura are reddish tan with pale cream granulation throughout. Throughout the head, legs, and body there is slight speckling as granulation is slightly paler in color than the surface it is found on. In lighter individuals, the venation of the tegmina is pale yellow to pale mint green (Fig. 38 A) and in darker individuals the venation is yellow with highlights of orange interspersed throughout (Fig. 38 B). Darker individuals also have variable reddish patches throughout the lobes of the legs and slightly darker coloration on the abdomen. Morphology. Head. Head capsule slightly longer than wide, vertex with granulation throughout the surface, none as prominent as the posteromedial tubercle which is not notably wide but is distinctly taller than any other nodes on the head (Fig. 39 E). Frontal convexity stout, marked throughout with slight granulation and several short setae. Compound eyes slightly protruding from the head capsule, but are significantly large, taking up slightly <⅓ of the head capsule margins (Fig. 39 E). Ocelli absent. Antennal fields slightly wider than the first antennomere. Antennae. Antennae consisting of nine segments, with the terminal segment slightly longer than the preceding two segments' lengths combined (Fig. 39 C). Antennomeres I-VIII sparsely marked with small transparent setae, the terminal antennomere is covered densely in slightly shorter setae. Thorax. Pronotum slightly wider than long, with gently concave anterior margin and slightly convex lateral margins, which converge to a slightly convex posterior margin that is half the width of the anterior margin (Fig. 39 E). The pronotum surface is marked with granulation throughout, a prominent pit in the center, and slight furrows anterior and lateral to the pit (Fig. 39 E). The pronotum has a prominent anterior rim and weakly formed lateral and posterior rims (Fig. 39 E). Prosternum and the anterior half of the mesosternum are marked with stout and numerous nodes, with the remainder of the mesosternum and the metasternum lacking prominent nodes (Fig. 39 B). Prescutum about as long as wide with lateral rims with 11 or 12 lumpy tubercles ranging in size from small to medium with granulation present throughout the length giving the margins a tough textured appearance (Fig. 39 E). Prescutum anterior rim not strongly protruding and marked with a granular surface (Fig. 39 F). Prescutum surface with 14 or 15 distinct nodes predominantly along the sagittal plane, with those on the anterior half slightly larger than the rest (Fig. 39 E). Mesopleura are narrow and parallel on the anterior ⅓, and then bend distinctly and diverge uniformly throughout their length; lateral margin with 13 - 16 small to medium lumpy tubercles, of which three or four are slightly larger than the rest, but most are small and variable in shape, giving the margin a rough textured appearance (Fig. 39 E). Face of the mesopleura with granulation along the margin, with the remainder of the surface relatively smooth or with slight wrinkles. The surface of the mesopleura also has two distinct pits, one near the anterior ⅓ where the mesopleura bend, and one near the posterior ⅓ (Fig. 39 E). Wings. Tegmina long, reaching onto abdominal segment VIII. The subcosta (Sc) is the first vein in the forewing and runs parallel with the wing for the first half of its length, and then bends towards the wing margin for the second half, terminating ca. ⅓ of the way through the wing length. The radius (R) spans the central portion of the tegmina with two subparallel branched veins. The first radius (R 1) branches ca. 1 / 2 through the radius length and terminates ca. ⅖ of the way through the wing length. The radial sector (Rs) branches from the end of the radius and runs angled to the wing margin where it terminates near the posterior ⅓ of the wing length. There is a weak continuation of the radius following the prominent radial sector branching which continues on as a short and thin radius to media crossvein (R-M). The media (M) is simply bifurcate with both the media anterior (MA) and media posterior (MP) terminating close to the posterior ⅕ of the wing. The cubitus (Cu) runs throughout the entire wing length simply, and then near the posterior ⅕ of the wing splits into the cubitus anterior (CuA) and cubitus posterior (CuP) which both terminate at or very near the wing posterior apex. The first anal vein (1 A) is simple and fuses with the cubitus early on, at around the midline between the first radial branching and the radial sector branching. Alae well-developed, reaching abdominal segment VI. Abdomen. Abdominal segments II through the anterior half of IV diverging, the posterior half of IV through the anterior half of VII parallel-sided (giving the abdomen a boxy appearance), the remainder of VII smoothly rounded and converging to the apex with segments VIII-X. Genitalia. Subgenital plate starts at the anterior margin of segment VIII, is moderately broad, and extends 1 / 2 to 3 / 4 of the way onto segment X, ending in a fine point (Fig. 39 H). Gonapophyses VIII are long and moderately broad, exceeding the apex of the abdomen with the tips slightly longer than the cerci, gonapophyses IX are thinner and shorter, hidden below gonapophyses VIII (Fig. 39 H). Cerci flat, not strongly cupped, with a finely granular surface and moderately marked with a few short setae. Legs. Profemoral exterior lobe broad and smoothly rounded, ca. 11 / 2 to ca. 2 x wider than the interior lobe (Fig. 39 D). Margin of the profemoral exterior lobe with 10 - 12 small weakly formed teeth throughout the length (Fig. 39 D). Profemoral interior lobe obtusely angled and typically marked with five teeth arranged in a two-one-two pattern with looping gaps between them, but occasionally individuals can have doubly serrate teeth or an extra small tooth between sets (Fig. 39 D). Mesofemoral exterior lobe arcs from end to end but is weighted towards the distal half with a detectable bend and marked with four or five rounded teeth distributed on the distal half only. Interior and exterior lobes of a similar width. Mesofemoral interior lobe arcs end to end smoothly with five or six small serrate teeth only on the distal half of the arc which is slightly wider than the proximal half of the arc. Metafemoral interior lobe arcs end to end and has five or six serrate teeth on the distal half of the lobe which is slightly wider than the proximal half. Metafemoral exterior lobe is thin and smooth, hugging the metafemoral shaft and lacks notable teeth but the distal ⅓ can be slightly granular. Protibial interior lobe spans the entire length of the protibiae and is ca. 21 / 2 the width of the protibiae shaft itself. The lobe is roundly triangular and is slightly wider on the distal half. Protibiae lacking a distinct exterior lobe. Mesotibiae and metatibiae lacking exterior and interior lobes. Measurements of paratype females [mm] (from Tatai, Cambodia). Length of body (including cerci and head, excluding antennae) 83.3 - 90.0, length / width of head 8.4 - 8.7 / 6.6 - 7.1, antennae 4.1 - 4.6, pronotum 5.6 - 6.0, mesonotum 7.6 - 7.8, length of tegmina 52.8 - 53.6, length of alae 42.6 (only measured on one specimen, the others have the alae covered by the tegmina), greatest width of abdomen 31.3 - 36.2, profemora 19.1 - 21.4, mesofemora 15.1 - 15.4, metafemora 18.7 - 19.6, protibiae 12.5 - 12.6, mesotibiae 11.4 - 11.6, metatibiae 14.7 - 15.0. Male. Coloration. Coloration description based on images of live males bred by Tim Bollens (Belgium). Overall coloration pale mint green throughout with highlighting of tan to orange (Fig. 39 C). The areas most often with the orange highlighting are the tips of the antennae, margins of the lobes of the legs, the thorax, and the margins of the abdomen. Additionally, on more prominently colored individuals the base of the antennae and the posteromedial tubercle of the head capsule can also be colored. Compound eyes are a muddled tan to reddish. Morphology. Head. Head capsule about as long as wide, with a vertex that has moderate granulation throughout and a prominent but not broad posteromedial tubercle which is larger than any of the granules on the head capsule (Fig. 40 E). Frontal convexity not particularly long but ending in a fine point and covered with sparse thin setae. Compound eyes large and bulbous, taking up ca. ⅖ of the head capsule lateral margins (Fig. 40 E). There are three moderately developed ocelli located between and slightly posterior to the compound eyes. Antennal fields about as wide as the scapus. Antennae. Antennae (including the scapus and pedicellus) consists of 25 segments, all segments except the scapus and pedicellus and terminal three segments are covered in dense setae that are as long as or longer than the antennae segment is wide. The terminal three segments are covered in dense short setae and the scapus and pedicellus are nearly completely bare. Thorax. Pronotum with anterior margin slightly concave and lateral margins that are straight or slightly convex and converging to a straight posterior margin that is half the width of the anterior rim (Fig. 40 E). Anterior margin of the pronotum has a distinct rim, lateral margins have moderate rims, and the posterior margin lacks a rim (Fig. 40 E). Face of the pronotum is marked by a distinct sagittal furrow and pit in the center, a granular surface, and a slight perpendicular furrow from the central pit. Prosternum is granulose throughout with small nodes of nearly even size. Mesosternum anterior half with nodes of a similar size to the prosternum and those on the posterior half slightly less prominent. The metasternum has a slightly wrinkled surface and sparse granulation. Prescutum longer than wide, with lateral margins slightly converging to the posterior (Fig. 40 E). Lateral rims with small granulation throughout giving them a rough textured appearance, only three or four are slightly larger than the rest. Prescutum surface with granulation throughout with those along the sagittal plane slightly larger than the others. Prescutum anterior rim weakly formed but marked with a surface which is granular. Mesopleura narrow, almost parallel for the anterior quarter, and then only gradually diverge for the remainder of the length (Fig. 40 E). Lateral margin lacking prominent tubercles, instead marked with sharp granulation throughout with only two or three slightly larger than the rest, giving the margins a rough textured appearance. Face of the mesopleura slightly wrinkled and with two faint divots, one near the anterior margin and one half-way through the length (Fig. 40 D). Wings. Tegmina moderate length, reaching ⅓ to 1 / 2 onto abdominal segment III. Tegmina wing venation: the subcosta (Sc) is the first vein, is simple, and terminates ca. 1 / 2 through the overall tegmina length. The radius (R) spans the entire length of the tegmina with the first radius (R 1) branching <ca. 1 / 2 through the wing length and terminating just distal to the midline, followed by the branching and termination of the second radius (R 2) near the distal ⅓ of the wing, and then the radial sector runs to the wing apex. The media (M) also spans the entire length of the tegmina with the first media posterior (MP 1) branching off ca. ⅖ of the way through the wing length, and then the second media posterior (MP 2) branches near the midline, and the media anterior (MA) runs to the wing apex. The cubitus (Cu) runs along the edge of the wing as the two media posterior veins fuse with it and as the cubitus reaches the apex it fades. The first anal (1 A) vein terminates upon reaching the cubitus slightly <⅓ of the way through the wing length. Alae well-developed in an oval fan configuration, long, reaching onto abdominal segments IX or X. Alae wing venation: the costa (C) is present along the entire foremargin giving stability to the wing. The subcosta (Sc) is long, spanning slightly> ⅔ of the wing length and is mostly fused with the radius in the beginning but terminates when it meets the costa. The radius (R) spans the entire wing and branches ca. ⅓ of the way through into the first radius (R 1) and radial sector (Rs) which run gently diverging for most of their length and then converge at the apex of the wing where they terminate near each other but not touching. The media (M) branches early, ca. ⅙ of the way through the wing into the media anterior (MA) and the media posterior (MP) which run parallel with each other throughout the wing until the distal ⅙ of the wing where the media posterior fuses with the media anterior which then run fused together to the wing apex where they terminate near the radial sector. The cubitus (Cu) runs unbranched and terminates at the wing apex. Of the anterior anal veins, the first anterior anal (1 AA) fuses with the cubitus near the point where the media branches into the media anterior and media posterior and then the first anterior anal branches from the cubitus ⅔ of the way through the wing length where it uniformly diverges from the cubitus until it terminates at the wing margin. The anterior anal veins two-seven (2 AA- 7 AA) have a common origin and run unbranched in a folding fan pattern of relatively uniform spacing to the wing margin. The posterior anal veins (1 PA- 6 PA) share a common origin separate from the anterior anal veins and run unbranched to the wing margin with slightly thinner spacing than the anterior anal veins. Abdomen. Margins of abdominal segment II either slightly converging or parallel-sided. Abdominal segments III through the anterior ⅔ of IV diverging. Segment V with parallel margins and VI-X converging slowly at first then more prominently for the terminal three segments, giving the abdomen a spade-shaped appearance. Genitalia. Poculum broad, posteriorly rounded and with a shallow notch medioapically; slightly passes the anterior margin of segment X (Fig. 40 G). Cerci long and slender, extending from under the anal abdominal segment, slightly cupped with a granular surface and numerous short setae throughout (Fig. 40 F). Vomer broad and stout with sides evenly converging and terminating in an upward hooking apical spine with a smaller hook next to the base of the primary spine (Fig. 5 D). Legs. Profemoral exterior lobe about the same width as the interior lobe or slightly wider, smoothly arcing from end to end and marked with a granular margin and five or six small serrate teeth on the distal half only (Fig. 40 C). Profemoral interior lobe roundly triangular and marked with five teeth arranged in a two-one-two pattern with prominent looping gaps between the sets and the middle tooth larger than the others (Fig. 40 C). Mesofemoral exterior lobe arcs end to end, but is slightly more bent than the interior lobe and is broader on the distal half which can either be lacking dentation or have three or four dulled teeth, and the proximal half that is rather thin and lacking teeth. Mesofemoral interior lobe of a similar width to the exterior lobe, is broader on the distal end and is marked with five or six serrate teeth mostly situated on the distal ⅓ to 1 / 2 of the lobe. Metafemoral exterior lobe lacks dentation, and has a straight margin along the metafemoral shaft. Metafemoral interior lobe smoothly arcs end to end with eight or nine serrate teeth on the slightly wider distal half. Protibiae lacking exterior lobe, interior lobe reaching end to end in a smoothly rounded triangle with the widest portion ca. 3 - 31 / 2 x as wide as the protibial shaft and situated just distal to the midline. Meso- and metatibiae simple, lacking lobes completely. Measurements of holotype male [mm]. Length of body (including cerci and head, excluding antennae) 61.9, length / width of head 4.5 / 3.9, antennae 37.31, pronotum 3.6, mesonotum 4.3, length of tegmina 19.0, length of alae 49.3, greatest width of abdomen 17.3, profemora 13.5, mesofemora 11.3, metafemora 13.4, protibiae 9.7, mesotibiae 7.8, metatibiae 10.1. Measurements of paratype males [mm]. Length of body (including cerci and head, excluding antennae) 63.8 - 70.2, length / width of head 5.0 - 5.5 / 4.1 - 4.3, antennae 38.8 - 39.5, pronotum 3.6 - 4.1, mesonotum 5.0 - 6.2, length of tegmina 20.0 - 20.4, length of alae 50.0 - 52.1, greatest width of abdomen 17.1 - 17.9, profemora 15.72, mesofemora 12.1, metafemora 13.7 - 14.1, protibiae 9.8 **, mesotibiae 8.7 - 8.9, metatibiae 11.2 - 11.4. Eggs. (Fig. 41). The lateral surfaces are flat with a length ca. 11 / 2 x the width with parallel margins, giving the capsule a rectangular appearance. All surfaces have numerous small to medium sized pits throughout, the lateral surface has around 35 pits (mostly on the smaller end of the spectrum) arranged in no detectable order, some more closely spaced than others. In addition, between the pits the surfaces are covered with short moss-like pinnae with the pinnae along the margins slightly longer than the pinnae on the other surfaces. The dorsal surface is marked with six or seven slightly irregular medium sized pits on each half running the length of the capsule with short moss-like pinnae around the micropylar plate and between the pits. The micropylar plate is not overly long, occupying ca. 1 / 2 of the dorsal surface length but not perfectly centered, with ca. ⅓ of the unoccupied space below and ⅔ above the micropylar plate. The micropylar cup is the widest portion of the micropylar plate and is located ca. ⅓ of the dorsal surface length from the posterior. The micropylar plate is approximately teardrop-shaped with the anterior portion longer and thinner than the posterior after the micropylar cup. Operculum slightly ovular, with the outer margin encircled with short moss-like pinnae surrounding the operculum and four or five medium pits surrounding the dorsal and lateral margins. The operculum is roundly raised with a height slightly> 1 / 2 operculum width. This rounded raised cap is marked with a sagittal raised row of pinnae similar in length to those along the capsule margins. The rounded raised cap is not perfectly centered and instead the rounded projection is shifted slightly towards the ventral surface. The overall egg color is tan to light brown, with the moss-like pinnae sometimes slightly lighter in color. Measurements including the extended pinnae [mm]. Length (including operculum): 5.6; maximum width of capsule when viewed from lateral aspect 3.2; length of micropylar plate 3.0.	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
7F2F01FA305959E39E1EC7B005EBDF4C.taxon	etymology	Etymology. Noun. The species epithet is the Hindi word Cryptophyllium khmer, meaning Cambodia, referring to the country of origin for this species.	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
3634D0DD98725A759E2C550D8B673343.taxon	materials_examined	Material examined. Holotype ♀: " VIETNAM, Lam Dong Province, Bao Lam, Dambri, V. 2018 ". Deposited within the Montreal Insectarium (IMQC). Paratypes: (13 ♀♀, 2 ♂, 7 eggs) • 1 ♂: " Coll. I. R. Sc. N. B., VIETNAM, Dak Nong prov., Ta Dung N. P., 11 ° 52 ' 22 " N 107 ° 58 ' 40 " E, 5 - 8. viii. 2019, GTI Project, Leg. J. Constant and J. Bresseel, I. G.: 34.048 (Coll. I. R. Sc. N. B.) ", vomer dissected (RBINS) (SB 0531 molecular sample within our analysis) • 1 ♂: " Vietnam: Dak Lak Province, Local collector, September 2020 " (IMQC) • 5 ♀♀; " Vietnam: Dak Lak Province, Local collector, September 2020 " (Coll RC 20 - 127, 20 - 128, 20 - 129) • 5 ♀♀; " Vietnam: Dak Lak Province, Local collector, September 2020 " (Coll SLT) • 3 ♀♀; " Vietnam: Dak Lak Province, Local collector, September 2020 " (IMQC) • 1 egg: " Vietnam: Dak Lak Province, Local collector, September 2020 " (Coll SLT) • 1 egg: " Vietnam: Dak Lak Province, Local collector, September 2020 " (Coll FH) • 3 egg: " Vietnam: Dak Lak Province, Local collector, September 2020 " (IMQC) • 2 egg: " Vietnam: Dak Lak Province, Local collector, September 2020 " (Coll RC).	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
3634D0DD98725A759E2C550D8B673343.taxon	distribution	Distribution. Southern Vietnam: presently known from three provinces: the type locality of Lam Dong Province, Bao Lam, Dambri; the male paratype record from Dak Nong Province, Ta Dung N. P.; an observational record from Dak Lak Province, Chu Yang Sin N. P.; and paratype records> from Dak Lak Province.	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
3634D0DD98725A759E2C550D8B673343.taxon	description	Description. Female. Coloration. Coloration description is based on photos of the holotype female shortly after being preserved. Nearly the entire body was of a uniform lime-green without differing colored markings (all legs and even wing venation a similar color to the rest of the body). Only the compound eyes were slightly yellow and not the same shade of green as the rest of the body. Morphology. Head. Head capsule longer than wide, vertex with a moderately granular surface, and the posteromedial tubercle which is three or four times larger than the most prominent granules on the capsule. Frontal convexity broad and stout, notably shorter than the length of the first antennomere, and with several long, thin, clear setae across the surface. Compound eyes are not large, only slightly protruding from the head capsule and with a width of ca. 1 / 4 the head capsule length (Fig. 42 C). Ocelli absent. Antennal fields wider than the first antennomere but not protruding back farther than the frontal suture. Antennae. Antennae consist of nine segments, with the terminal segment approximately the same length as the previously three segments combined (Fig. 42 D). The eighth antennal segment has a distinct furrow around the middle which makes the segment appear to be two separate segments (giving the antennae a ten segmented appearance), but this furrow appears to only be superficial (Fig. 42 D). Antennomeres I-III sparsely marked with thin transparent setae, similar to those found on the frontal convexity, but slightly shorter in length. Antennomere IV is short, disk-like, and wider than the following segments, and interestingly has a base which is narrow and somewhat longer than other congeneric antennal segments IV, giving it a raised appearance (Fig. 42 D). The terminal antennomere and the distal half of segment VIII (distal to the midline furrow) are covered in dense, stout, setae. Thorax. Pronotum with anterior margin slightly concave and lateral margins that are relatively straight, converging to a narrow, straight posterior margin that is ca. 1 / 2 the width of the anterior rim (Fig. 42 E). The pronotum surface is smooth, with only a prominent pit in the center, and slight furrows anterior and lateral to the pit, no prominent wrinkles or granulation. The pronotum has a prominent anterior rim and moderate lateral rims, the posterior is lacking a rim. Prosternum with notable nodes throughout the surface, relatively evenly spaced. Mesosternum with prominent nodes on the anterior margin, followed by moderate nodes on the anterior ⅓ of the surface, with the remainder with dispersed, weak granulation which continues onto the metasternum. Prescutum notably longer than wide, with lateral margins running parallel to the posterior margin giving it a distinctly rectangular appearance. Lateral rims with five or six medium sized tubercles situated on the anterior ⅔, with only small granulation on the remainder. Prescutum anterior rim prominent but not strongly protruding, with the surface granular and lacking a prominent sagittal tubercle. Prescutum surface with granulation throughout with those along the sagittal plane slightly larger (Fig. 42 E). Mesopleura start ca. ⅓ down the prescutum and evenly diverge with straight lateral margins. Lateral margin with six or seven major and distinctly pointed tubercles and six or seven smaller tubercles intermixed amongst them. This mix of tubercles is only prominent along the anterior ⅔ of the length with the remaining ⅓ lacking notable tubercles and instead marked with consistent granulation (Fig. 42 E). Face of the mesopleura with slight wrinkles throughout most of the surface and a few irregular nodes along the lateral margin, as well as two faint divots, one on the anterior margin and one closer to the center. Wings. Tegmina reaching slightly past the anterior margin of abdominal segment VII. Tegmina venation is rather typical for the Cryptophyllium gen. nov. The subcosta (Sc) is the first vein in the forewing and runs parallel with the wing for the first half of its length, and then bends towards the wing margin for the second half. The radius (R) spans the central portion of the tegmina with two subparallel branched veins. The first radius (R 1) branches ca. 1 / 2 through the radius length and terminates ca. ⅓ of the way through the wing length. The radial sector (Rs) branches from the end of the radius and runs angled to the wing margin where it terminates near the wing midline length. There is a weak continuation of the radius following the prominent radial sector branching which continues on as a short and thin radius to media crossvein (R-M). The media (M) is simply bifurcate with both the media anterior (MA) and media posterior (MP) terminating close to the posterior 1 / 4 of the wing. The cubitus (Cu) runs throughout the entire wing length simply, and then near the posterior ⅕ of the wing becomes bifurcate into the cubitus anterior (CuA) and cubitus posterior (CuP) which both terminate at or very near the wing posterior apex. The first anal vein (1 A) is simple and fuses with the cubitus early on, at around the midline between the first radial branching and the radial sector branching. Alae short, only 21.8 mm long. Abdomen. Abdominal segments II through the anterior half of IV diverging, posterior half of IV through the anterior ⅓ of VII parallel. Abdominal segment VII with a distinct looping lobe which meets abdominal segment VIII which is notably narrower. Segments VIII-X uniformly converge to a broad rounded apex. Genitalia. Projecting portion of the subgenital plate stout, beginning at the anterior margin of abdominal segment IX and projecting with nearly straight sides to just under the anterior margin of the terminal abdominal segment (Fig. 42 H). Gonapophyses VIII are long and broad with a dagger-like shape (parallel-sided at first and then after ca. 1 / 2 of the length uniformly converging to the point) with the points just projecting from under the terminal abdominal segment (Fig. 42 H). Gonapophyses IX are smaller and not visible from under the large gonapophyses VIII. Cerci strongly pointed and relatively flat (not strongly cupped) with weakly crenate margins, and the dorsal surface heavily granular and marked by thin transparent setae throughout (Fig. 42 G, H). Legs. Profemoral exterior lobes broad, at its broadest ca. 2 x wider than the interior lobe, and distinctly serrate throughout the entire length (with 13 - 17 small, pointed teeth). The proximal edge is slightly concave and the distal edge is smoothly convex, therefore giving the lobe a distinct recurved appearance and an acute exterior angle (Fig. 42 F). Interior profemoral lobe ca. 21 / 2 x the width of the profemoral shaft at its widest and with doubly serrate, large, triangular teeth. The largest teeth are grouped into a two-one-two pattern with large looping gaps between these groupings, with these large gaps also finely serrate, not smooth (Fig. 42 F). Interior mesofemoral lobe arcs evenly weighted from end to end, and at its widest is approximately the same width as the mesofemoral shaft itself. The interior mesofemoral lobe is finely serrate for ca. 3 / 4 of its distal length with seven or eight teeth. Mesofemoral exterior lobe is also approximately at its widest as wide as the mesofemoral shaft, but the exterior lobe is distinctly bent in the center with straight margins, not smoothly arcing from end to end. On the distal half of the lobe only there are six or seven small serrate teeth. Metafemoral interior lobe arcs end to end and has five or six dull teeth pointing distally. Metafemoral exterior lobe is thin and smooth, hugging the metafemoral shaft and lacks dentation. Protibiae with a thin but notable exterior lobe on the distal ⅕ only. Protibial interior lobe spans the entire length as a broad scalene triangle with the broad end on the distal half of the protibiae. Mesotibiae simple, completely lacking lobes. Metatibiae lacking interior lobes, exterior is marked by a very slender lobe which only occupies the distal 1 / 4 of the shaft. Measurements of holotype female [mm]. Length of body (including cerci and head, excluding antennae) 99.0, length / width of head 8.9 / 6.8, antennae 5.1, pronotum 5.9, mesonotum 6.5, length of tegmina 55.7, length of alae 21.8, greatest width of abdomen 41.5, profemora 22.8, mesofemora 16.0, metafemora 19.1, protibiae 11.2, mesotibiae 11.8, metatibiae 15.0. Male. Coloration. Coloration description based on the captive reared paratype male when it was alive (Fig. 43). Overall coloration mint-green throughout with highlights of tan coloration on the distal half of the protibiae, along all femoral lobe margins, the frontal convexity, the margins of the thorax, the base of the antennae and the distal tips of each longer antennomere, and intermittently along the tegmina and alae sclerotized veins. Abdominal segment V has a slightly darker patch and a transparent eye spot on each side of the midline. Compound eyes are pale yellow with slight orange marbling throughout. Morphology. Head. Head capsule approximately as long as wide, with a vertex that is only slightly granular with no discernable pattern (Fig. 44 C). The posteromedial tubercle is not broad and is only weakly raised from the head capsule. Compound eyes are large and bulbous, taking up ca. ⅖ of the head capsule lateral margins (Fig. 44 C). There are three well-developed ocelli between and slightly posterior to the compound eyes (Fig. 44 C). Antennal fields are about as wide as the scapus. Antennae. Antennae (including the scapus and pedicellus) consist of 29 segments. The scapus and pedicellus are nearly completely bare, lacking long setae. The following segments except the terminal three are covered in dense setae that are as long as or longer than the antennae segment is wide, and the terminal three segments are covered in dense short setae. Thorax. Pronotum with anterior margin slightly concave and lateral margins that are straight and converging to a slightly curved posterior margin that is ca. 1 / 2 the width of the anterior rim (Fig. 44 C). Anterior and lateral margins have moderate rims and the posterior margin lacks a rim (Fig. 44 C). Face of the pronotum is marked by a distinct pit in the center with a furrow anterior to the pit along the sagittal plane, weakly formed furrows lateral to the central pit, and a smooth surface with only slight granulation in no detectable pattern (Fig. 44 C). The prosternum is slightly granulose throughout and the anterior ⅓ of the mesosternum surface is marked with more prominent nodes, with the remainder of the surface with fewer and smaller nodes (Fig. 44 B). Prescutum longer than wide, with lateral margins slightly converging to the posterior (Fig. 44 C). Lateral rims with eight or nine small tubercles (Fig. 44 C). Prescutum surface slightly raised along the sagittal plane with six or seven nodes of varying size, with the remainder of the surface with only slight granulation throughout. Prescutum anterior margin weakly formed and with a granular surface, lacking a prominent central tubercle. Mesopleura narrow on the anterior quarter of the length but then gently diverging with nearly straight margins to the posterior (Fig. 44 C). Mesopleura lateral margin with three large conical tubercles, three or five small tubercles, and four or five nodes throughout the length (Fig. 44 C). Face of the mesopleura slightly wrinkled, with slight granulation throughout, and with two faint divots, one on the anterior margin and one near the midline. Wings. Tegmina moderate length, extending halfway through abdominal segment III. Tegmina wing venation: the subcosta (Sc) is the first vein, is simple, and terminates the earliest slightly <1 / 2 through the overall tegmina length. The radius (R) spans the entire length of the tegmina with the first radius (R 1) branching ca. ⅓ of the way through the wing length and terminating near the midline, followed by the branching and termination of the second radius (R 2) slightly distal to the midline, and then the radial sector runs to the wing apex. The media (M) also spans the entire length of the tegmina with the first media posterior (MP 1) branching off ca. ⅓ of the way through the wing length, then the second media posterior (MP 2) branching near the midline, and the media anterior (MA) runs to the wing apex. The cubitus (Cu) runs along the edge of the wing as the two media posterior veins fuse with it and as the cubitus reaches the apex it fades. The first anal (1 A) vein terminates upon reaching the cubitus ca. ⅓ of the way through the wing length. Alae well-developed in an oval fan configuration, long, reaching the middle of abdominal segment VIII. Abdomen. Abdominal segment II gently diverging, III through the anterior half of segment IV diverging to the widest portion of the abdomen. The posterior of IV-V parallel-sided giving the abdomen a broad spade-shaped appearance. Segments VI-X uniformly converging with slightly undulating margins (Fig. 44 B). Genitalia. Poculum broad, and ends in a broadly rounded apex that slightly passes the anterior margin of segment X (Fig. 44 G). Cerci long and slender, with> 1 / 2 their length extending from underneath the terminal abdominal segment (Fig. 44 F), relatively flat, covered in a heavily granulose surface and with equally granular margins, and surface with numerous short setae (Fig. 44 G). Vomer stout with slightly rounded sides converging to an apex with two side by side thick apical hooks which are the same size and hook upwards into the paraproct (Fig. 43 C). Legs. Profemoral exterior lobe broader than the interior lobe, arcing end to end with a distinct rounded bend in the center, with the proximal half margin with a distinctly granular surface, and the distal half with six small serrate teeth (Fig. 44 E). Profemoral interior lobe roundly triangular, at least 3 x wider than the profemoral shaft, and marked with five large, serrate teeth arranged in a two-one-two pattern with looping gaps between them (Fig. 44 E). Mesofemoral exterior lobe arcs end to end, is slightly wider than the mesofemoral shaft, but with the widest portion on the distal ⅓ which is marked with granulation or with one or two weakly formed teeth. Mesofemoral interior lobe is slightly thinner than the exterior lobe and is slightly broader on the distal end which is marked with six serrate teeth. Metafemoral exterior lobe lacks dentation, and has a straight margin hugging the metafemoral shaft. Metafemoral interior lobe smoothly arcs end to end, with the distal half wider than the proximal half, and the distal half is marked with seven or eight serrate teeth on the distal half. Protibiae with a small but notable exterior lobe on the anterior ⅕ which is no wider than the width of the protibial shaft (Fig. 44 E). Protibial interior lobe reaching end to end in a smoothly rounded triangle with the widest portion on the distal third ca. 3 x as wide as the protibial shaft (Fig. 44 E). Meso- and metatibiae simple, lacking lobes. Measurements of reared paratype male [mm]. Length of body (including cerci and head, excluding antennae) 79.7, length / width of head 4.9 / 3.9, antennae 41.8, pronotum 3.7, mesonotum 5.7, length of tegmina 24.4, length of alae 57.5, greatest width of abdomen 26.5, profemora 16.4, mesofemora 11.4, metafemora 13.7, protibiae 9.1, mesotibiae 7.7, metatibiae 10.9. Eggs. (Fig. 45). When viewed from the anterior the egg capsule cross section is rounded pentagonal, therefore the lateral surfaces are raised into dorsolateral and ventrolateral surfaces. All surfaces and margins slightly undulate giving the egg an overall lumpy appearance. The ventrolateral surface is marked by four large, evenly spaced pits from the anterior to the posterior in a singular line. The dorsolateral surface has five large pits, arranged with one on the anterior, one on the posterior, and three in the middle spaced out in a two-one pattern. Both lateral surfaces are primarily bare but do have sparse and small moss-like pinnae between the pits. The dorsal surface is marked with seven medium sized pits total (one on the anterior end along the sagittal plane at the apex of the micropylar plate, followed by three on each slide of the plate with broad spacing between them ending with the posterior most near the base of the capsule). The dorsal surface is also marked throughout with short moss-like pinnae around the micropylar plate, with the area immediately around each pit bare. The micropylar plate is long, ca. 5 / 7 of the overall dorsal surface length with the micropylar cup situated on the posterior ⅓ of the length. The micropylar plate is thin with the widest portion the area around the micropylar cup. The operculum is slightly ovular with a surface that is roundly raised and a height slightly <1 / 2 the operculum width. The operculum is marked intermittently with moss-like pinnae similar in shape but slightly smaller than those found on the rest of the capsule, as well as marked with two medium sized pits (one on each side of the sagittal plane). The overall egg color is dark brown, with the moss-like pinnae light brown in color so they stand out clearly on the surface. Measurements including the extended pinnae [mm]. Length (including operculum): 5.2; maximum width of capsule when viewed from lateral aspect 3.8; length of micropylar plate 3.0.	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
3634D0DD98725A759E2C550D8B673343.taxon	etymology	Etymology. Patronym. Named after Rene Limoges (Canada) from the Montreal Insectarium to thank him for his many years of assisting Team Phyllies with countless publication worthy photographs.	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
DC10D79283405E2C8DC092214AC2E7B9.taxon	materials_examined	Material examined. Holotype ♀: " CHINA: Guangxi Prov., Jinxiu County, Liuzhou City, Dayaoshan Mountain, 875 - 1,500 m., 18 - 19. IX. 2019. (Coll RC 20 - 002) ". Deposited in the Montreal Insectarium (IMQC). Paratypes: (6 ♀♀, 4 eggs) • 1 ♀: " CHINA: Guangxi Prov., Jinxiu County, Liuzhou City, Dayaoshan Mountain, 875 - 1,500 m., 19. IX. 2019. Coll RC 19 - 182 " (Coll RC) • 1 ♀: " CHINA: Guangxi Prov., Jinxiu County, Liuzhou City, Dayaoshan Mountain, 875 - 1,500 m., 18 - 19. IX. 2019. Coll RC 20 - 003 " (Coll RC) • 3 ♀♀: Same data as the holotype (Coll SLT) • 1 ♀: Same data as the holotype (Coll FH) • 4 eggs: Removed from the abdomen of paratype female 19 - 182. " CHINA: Guangxi Prov., Jinxiu County, Liuzhou City, Dayaoshan Mountain, 875 - 1,500 m., 19. IX. 2019 "; Coll RC 20 - 069 - 20 - 072: 20 - 069, 20 - 072 (Coll RC); 20 - 070, 20 - 071 (IMQC). Photographic record: (1 ♀) In addition to the type material examined, images of a live female observed by Dr. Lu Qiu in Guangxi, Huaping Nature Reserve, 900 - 1000 m, in August 2020 (Fig. 46 B) were shared with us and compared to type material.	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
DC10D79283405E2C8DC092214AC2E7B9.taxon	distribution	Distribution. Presently only confirmed from two localities in Guangxi Province, Liuzhou prefecture-level city (Fig. 46 A, C) and Guilin prefecture-level city (Fig. 46 B). The other specimens within Liu (1993) with varying locality data which may represent additional localities have not been reviewed so it is possible that these may represent other species or range expansions.	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
DC10D79283405E2C8DC092214AC2E7B9.taxon	description	Description. Female. Coloration. Coloration descriptions are based upon photos of the live individuals which became the type material herein described (Fig. 46 A, C). The overall coloration is pale mint green, with highlights of yellow, orange, and red. The antennae, interior profemoral lobe margins, and the margins of the terminal abdominal segments are red. The mesopleura margins, veins of the tegmina, and exterior profemoral lobe margins are orange to yellow. Morphology. Head. Head capsule about as long as wide, vertex smooth (Fig. 47 D). The posteromedial tubercle is the only notable feature on the head capsule. Frontal convexity broad for the posterior half then narrowing on the anterior half, about as long as the first antennomere, and with slight granulation on the dorsal surface and several setae present which are longer than any setae on the antennae. Compound eyes slightly protruding from the head capsule, taking up ca. 1 / 4 of the length of the lateral head capsule margins (Fig. 47 D). Ocelli absent. Antennal fields slightly wider than the first antennomere and slightly shallower than the first antennomere is tall (Fig. 47 B). Antennae. Antennae consisting of nine segments, with the terminal segment slightly shorter than the preceding two segment lengths combined (Fig. 47 B). Antennomeres I-VIII sparsely marked with small transparent setae, the terminal antennomere is more densely covered in stout, brown setae. The pars stridens of antennomere III has 46 or 47 teeth. Thorax. Pronotum with gently concave anterior margin and straight lateral margins, which converge to a convex posterior margin that is 1 / 2 the width of the anterior margin (Fig. 47 D). The pronotum surface is smooth, with only a prominent pit in the center, and slight furrows anterior, posterior, and lateral to the pit (Fig. 47 D). The pronotum has a prominent anterior rim and weakly formed lateral and posterior rims, all of which are relatively smooth (Fig. 47 D). Prosternum and the mesosternum with stout and numerous nodes, with the central area of the mesosternum with less nodes and relatively smooth. Metasternum with granulation reduced and only minimal. Prescutum anterior margin as wide as the presuctum is long, with margins slightly narrowing on the anterior ⅓ and then running parallel to the posterior margin which is slightly narrower than the anterior margin (Fig. 47 D). Lateral rims with 7 - 9 short tubercles which are all about the same size (Fig. 47 D). Prescutum anterior rim prominent but not strongly protruding, with a surface that is granular, lacking a singular prominent sagittal tubercle (Fig. 47 D). Prescutum surface with minimal granulation throughout, with those along the sagittal plane only slightly larger (Fig. 47 D). Mesopleura beginning near the anterior margin of the prescutum and evenly diverging; lateral margin with 9 - 13 tubercles which are variable in size, with three larger than the rest (Fig. 47 D). Face of the mesopleura relatively smooth, and with two notable divots, one near the anterior margin and another ca. ⅗ of the way through the length (Fig. 47 D). Wings. Tegmina reaching ca. 1 / 2 through abdominal segment VII. The subcosta (Sc) is the first vein in the forewing, is distinctly bent in the center, and terminates ca. 1 / 4 of the way through the wing length. The radius (R) spans the anterior half of the forewing with two subparallel branched veins; radius 1 (R 1) terminates ca. ⅓ of the way through the wing length, and the radial sector (Rs) terminates in the center of the wing at the widest portion. There is a weak continuation of the radius following the prominent Rs branching which continues on as a short and thinner R-M crossvein that does not appear to solidly connect the two veins fading as it reaches the media. The media (M) is bifurcate with both the media anterior (MA) and media posterior (MP) terminating on the posterior ⅓ of the wing. In some individuals there is a weak continuation of the media as a thin crossvein to the cubitus, but this was not present in all individuals observed. The cubitus (Cu) is also bifurcate, branching near the posterior ⅓ to 1 / 4 of the wing into the cubitus anterior (CuA) and cubitus posterior (CuP) which both terminate at or very near the wing posterior apex. The first anal vein (1 A) is simple and fuses with the cubitus early on, only slightly past the branching distance of the first radius from the radius. Alae well-developed, reaching abdominal segment VI (41.0 - 43.0 mm long). Abdomen. Abdominal shape relatively stable between all females observed. Segments II through the anterior ⅓ of IV evenly diverging, with the posterior ⅔ of segment IV the widest segment. Segments V-VI are slightly subparallel, converging gently to the posterior, giving the abdomen a slight narrowing appearance. Segment VII is distinctly rounded to the terminal three segments which are notably narrower than the previous segments (segment VIII on the anterior is slightly> 1 / 2 width of the widest portion of the abdomen). Segments VIII-X converge to the apex which is broad and rounded. Genitalia. Subgenital plate starts at the anterior margin of segment VIII, is moderately broad, and extends ca. ⅔ of the way onto segment X, ending in a fine point (Fig. 47 E). Gonapophyses VIII are long and moderately broad, with their tips notably exceeding the apex of the abdomen, and slightly shorter than the tips of the cerci; gonapophyses IX are slender and long, extending ca. 3 / 4 of the way onto segment X (Fig. 47 E). Cerci flat, not strongly cupped, with a heavily granular surface (Fig. 47 E). Legs. Profemoral exterior lobe broad (broader than the interior lobe), approximately right angled, and with a rounded exterior angle. The margin is marked by 7 - 9 small serrate teeth throughout the length, none prominent (Fig. 47 C). Profemoral interior lobe narrower than the exterior lobe (only ca. 3 x the greatest width of the profemoral shaft) and with a slightly obtuse angle giving the interior lobe a triangular appearance marked by five teeth on the distal margin (Fig. 47 C). These teeth are arranged in a two-one-two pattern with the exterior pairs closer together and with a shallow gap between them, and the gap to the center tooth is deeper and wider than these exterior pairs (Fig. 47 C). Mesofemoral exterior lobe arcs from end to end as a rounded triangle, but is slightly weighted towards the distal 1 / 2 and marked with two or three small serrate teeth distributed on the distal half only. Mesofemoral exterior lobe is slightly wider than the interior lobe. Mesofemoral interior lobe arcs end to end more evenly than the exterior lobe, is marked with five or six small serrate teeth only on the distal half of the arc. Metafemoral interior lobe arcs end to end but is significantly more heavily weighted on the distal half and has six or seven serrate teeth on the broader distal half of the lobe. Metafemoral exterior lobe is thin and smooth, hugging the metafemoral shaft. Protibiae interior lobe spans the entire length of the protibiae and is at its widest ca. 2 x the width of the protibiae shaft itself. The lobe is distinctly triangular and widest on the distal half. There is a small but notable exterior protibial lobe on the distal quarter of the length, but this is only ca. 1 / 2 as wide as the protibial shaft itself. Mesotibiae simple, lacking lobes completely. Metatibiae lacks an interior lobe, but does have a small but notable anteriorly situated exterior lobe. Measurements of holotype female [mm]. Length of body (including cerci and head, excluding antennae) 88.7, length / width of head 7.2 / 7.1, antennae 4.6, pronotum 5.3, mesonotum 7.5, length of tegmina 52.3, length of alae 41.5, greatest width of abdomen 37.9, profemora 19.7, mesofemora 15.0, metafemora 17.2, protibiae 12.6, mesotibiae 11.4, metatibiae 15.2. Measurements of paratype females [mm]. Length of body (including cerci and head, excluding antennae) 90.8 - 92.0, length / width of head 7.5 - 7.7 / 6.6 - 7.3, antennae 4.9 - 5.3, pronotum 5.5 - 5.6, mesonotum 8.0 - 8.4, length of tegmina 50.5 - 53.8, length of alae 41.0 - 43.5, greatest width of abdomen 35.9 - 39.5, profemora 19.9 - 21.1, mesofemora 15.1 - 16.0, metafemora 17.6 - 18.0, protibiae 12.0 - 12.6, mesotibiae 10.8 - 11.0, metatibiae 15.6 - 15.9. Description of egg (Fig. 8 K, L). The entire capsule is covered in short moss-like pinnae and pitting of various size, depth, and spacing. The dorsal, ventral, and lateral surfaces are flattened, giving the egg a rectangular appearance. When viewed from the lateral aspect, the egg has an almost uniform width throughout. The lateral margins have slightly longer pinnae than the faces, but not drastically. Lateral surfaces slightly raised along the center of the length of the egg, and the entire surface has various shallow pitting in no detectable pattern. Micropylar plate spans the entire dorsal surface, with the thickest portion near the posterior ⅓ around the micropylar cup. The remainder of the micropylar plate is narrower, but not thin, at the thinnest still ca. 1 / 2 of the width of the widest portion. Operculum ovular and shallowly raised in the center and a singular circle of eight or nine shallow pits around the margin. Overall color light tan to brown. Measurements including the extended pinnae [mm]. Length (including operculum) 4.8 - 5.0, maximum width of capsule when viewed from lateral aspect 3.5 - 3.6 mm, length of micropylar plate 3.9 - 4.0 mm.	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
DC10D79283405E2C8DC092214AC2E7B9.taxon	etymology	Etymology. Patronym. The type specimens for this species were discovered by Chengzhi Bian (China) who recognized the scientific importance of the specimens and shared them with the authors to review. Chengzhi Bian has decided to name this species after his mother Liyanan to thank her for her amazing support of his passion for entomology.	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
9DD319B2D2195D248366348AE8D1C48F.taxon	materials_examined	Material examined. Holotype ♀: " Coll. I. R. Sc. N. B., Vietnam, Ninh Thuan prov., Nui Chua N. P., 11 ° 42 ' N 109 ° 09 ' E, 3 - 9. VII. 2014, night coll. Leg. J. Constant and J. Bresseel, GTI project I. G.: 32.779, DNA PH 006 ". Deposited in the Royal Belgian Institute of Natural Sciences (RBINS). Paratype: 1 ♀, " Coll. I. R. Sc. N. B., Vietnam, Ninh Thuan prov., Nui Chua N. P., 11 ° 42 ' N 109 ° 09 ' E, 3 - 9. VII. 2014, night coll. Leg. J. Constant and J. Bresseel, GTI project I. G.: 32.779 " (VNMN).	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
9DD319B2D2195D248366348AE8D1C48F.taxon	distribution	Distribution. At present only known from Nui Chua N. P., in Ninh Thuan Province, Vietnam.	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
9DD319B2D2195D248366348AE8D1C48F.taxon	description	Description. Female. Coloration. Coloration description is based upon the living type material (Fig. 48). Overall coloration is pale green throughout. The antennae, compound eyes, interior profemoral lobe margin, small patches along the protibial interior lobe, and the anterior of the prescutum margins are orange to red, but these areas are only sparsely marked. Morphology. Head. Head capsule about as long as wide, vertex relatively smooth with the only notable feature being the posteromedial tubercle which is finely pointed (Fig. 49 E). Frontal convexity broad and blunt, with a slightly granular surface. Compound eyes slightly protruding from the head capsule, and are not particularly large, taking up slightly <1 / 4 of the head capsule lateral margins (Fig. 49 E). Ocelli absent. Antennal fields slightly wider than the width of the first antennomere. Antennae. Antennae consisting of nine segments, with the terminal segment about the same length as the preceding two segments' lengths combined (Fig. 49 C). Antennomeres I-VII sparsely marked with small transparent setae, the terminal antennomere and antennomere VIII are covered in stout, brown setae (Fig. 49 C). Antennomeres V-VII ventral margins project farther than the ventral margin of segment VIII, therefore giving the antennae a slight lamelatte appearance (Fig. 6 E). Thorax. Pronotum with gently concave anterior margin and slightly convex lateral margins, which converge to a straight posterior margin that is slightly <half the width of the anterior margin (Fig. 49 E). The pronotum surface is smooth, with only a prominent pit in the center, and slight furrows anterior and lateral to the pit (Fig. 49 E). The pronotum has moderately formed anterior and lateral rims and a weakly formed posterior rim, all of which are relatively smooth (Fig. 49 E). Prosternum and the anterior half of the mesosternum are covered with numerous nodes, the metasternum has lateral margins which are slightly granular, and the central area is relatively smooth. Prescutum longer than wide, lateral rims with four or five small tubercles on the anterior ⅓, the remainder only has nodes throughout, giving the margin a rough textured appearance (Fig. 49 E). Prescutum anterior rim not strongly protruding, rim surface is granular, lacking a large sagittal spine (Fig. 49 E). Prescutum surface granular, with those along the sagittal plane slightly larger than the rest (Fig. 49 E). Mesopleura begin ca. ⅓ of the way through the prescutum length and evenly diverge; lateral margin with seven or eight small tubercles and several nodes interspersed (Fig. 49 E). Face of the mesopleura smooth or slightly wrinkled, with two notable divots, one on the anterior margin and one near the middle (Fig. 49 E). Wings. Tegmina long, reaching 1 / 2 through abdominal segment VII. Tegmina venation; the subcosta (Sc) is the first vein in the forewing, running parallel with the margin for the first half, and then bending and running towards the margin. The radius (R) spans the central portion of the forewing with two subparallel branched veins; the first radius (R 1) branches ca. 1 / 4 of the way through the wing length, terminates ca. ⅖ of the way through the wing length, and the radial sector (Rs) branches ca. ⅖ of the way through the wing length and terminates near the distal ⅓ of the wing length. There is a weak continuation of the radius following the prominent Rs branching which continues on as a short and thin R-M crossvein that weakly connects the two veins. The media (M) is simply bifurcate with both the media anterior (MA) and media posterior (MP) terminating near to the posterior 1 / 4 of the wing. The cubitus (Cu) is also bifurcate, branching near the posterior ⅕ of the wing into the cubitus anterior (CuA) and cubitus posterior (CuP) which both terminate at or very near the wing posterior apex. The first anal vein (1 A) is simple and fuses with the cubitus early on, at the length about midway between the splitting of the R 1 and Rs. Alae short, with their apex only just passing the posterior margin of abdominal segment III or slightly passing onto the anterior margin of abdominal segment IV. Abdomen. Abdominal segments II through the anterior half of IV diverging. The posterior half of segment IV through segment VI are parallel, giving the abdomen a boxy appearance. Abdominal segment VII has a slightly rounded margin, no notable protruding lobe present. Segments VIII-X are notably narrower than the previous segments, and have converging margins to the broad rounded apex (Fig. 49 G). Genitalia. Subgenital plate starts at the anterior margin of segment VIII, is moderately broad, and extends ca. 3 / 4 of the way onto segment X with straight margins ending in a fine point (Fig. 49 H). Gonapophyses VIII are long and moderately broad, exceeding the apex of abdominal segment X; gonapophyses IX are shorter and narrower, hidden below (Fig. 49 H). Cerci only slightly cupped, with a granular surface and margins, and few detectable setae (Fig. 49 G). Legs. Profemoral exterior lobe broad, rounded, and obtusely angled, smoothly arcing from end to end, ca. ⅓ again wider than the width of the interior lobe (Fig. 49 D). Edge of the profemoral exterior lobe granular, or with a slightly serrate surface of four or five very small teeth (Fig. 49 D). Profemoral interior lobe ca. 21 / 2 x as wide as the greatest width of the profemoral shaft, obtusely angled, and marked with five teeth arranged in a two-one-two pattern with shallow gaps between them (Fig. 49 D). Mesofemoral exterior lobe arcs from end to end but is slightly bent in the center, weighted towards the distal 1 / 2, with a smooth proximal margin and a slightly lumpy distal half appearing to be weakly formed teeth. Interior and exterior mesofemoral lobes about the same width. Mesofemoral interior lobe arcs smoothly end to end with six or seven small serrate teeth only on the distal half of the arc which is slightly wider than the proximal half of the arc. Metafemoral interior lobe arcs end to end, with the distal half slightly wider than the proximal half and marked with seven or eight serrate teeth on the distal half of the lobe only. Metafemoral exterior lobe is thin and smooth, hugging the metafemoral shaft and lacks dentation. Protibiae lacking an exterior lobe (Fig. 49 D). Protibiae interior lobe spans the entire length of the protibiae and is ca. 21 / 2 x the width of the protibiae shaft itself. The lobe is roundly triangular with the widest portion just slightly distal to the midline. Mesotibiae and metatibiae lacking exterior and interior lobes. Measurements of holotype female [mm]. Length of body (including cerci and head, excluding antennae) 64.9, length / width of head 6.7 / 5.6, antennae 3.2, pronotum 4.5, mesonotum 5.9, length of tegmina 38.2, length of alae 15.2, greatest width of abdomen 22.0 (abdomen not perfectly flat), profemora 15.7, mesofemora 11.6, metafemora 14.2, protibiae 10.0, mesotibiae 8.2, metatibiae 11.4. Measurements of paratype female [mm]. Length of body (including cerci and head, excluding antennae) 67.6, length / width of head 7.3 / 6.2, antennae 3.3, pronotum 4.9, mesonotum 6.2, length of tegmina 40.3, length of alae 16.4, greatest width of abdomen 24.6 (abdomen not perfectly flat), profemora 15.5, mesofemora 11.5, metafemora 15.9, protibiae 10.0, mesotibiae 8.6, metatibiae 11.8.	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
9DD319B2D2195D248366348AE8D1C48F.taxon	etymology	Etymology. Toponym, named for the type locality, Nui Chua N. P. where this species was first discovered in Ninh Thuan Province, Vietnam.	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
BD05CDBF570B51D98A8877617AD023EB.taxon	materials_examined	Material examined. The extensive paratype series within Coll RC and Coll SLT were examined to review the intraspecific variation within this species. See Cumming and Le Tirant (2020) for a detailed list of material examined. Additionally, 1 ♂: " Coll. I. R. Sc. N. B., Laos, NE, Mt Phu Phan, vi. 2019, local collectors, I. G.: 34.159 " [vomer dissected] (paratype male from original description) (RBINS). Photographic records: 1 ♂, 1 ♀: " China, Yunnan Province, Maguan County, Gulinqing town (古林箐乡) " (observed by Xiang-Jing Liu, China); 1 ♂: " Vietnam, Ha Giang Province, Dung Ba Commune " (observed by Chien C. Lee, Malaysia); 1 ♂: " Thailand, Nan Province, Bo Kluea Tai District " (observed by Tatsatorn Dharithai, Thailand); 1 ♂: " Thailand: Phetchabun Province, Phetchabun Research Station " (Photograph shared by Tatsatorn Dharithai, Thailand); 1 ♀: " San Ku Ruins, Chiang Mai, Thailand " (observed by Rob Thacker, United Kingdom).	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
BD05CDBF570B51D98A8877617AD023EB.taxon	distribution	Distribution. Widely ranging through several countries with only inland records> to date, no coastal records> yet known. This species has been located in Vietnam, Ha Giang Province, Dung Ba Commune (observed by Chien C. Lee, Malaysia; Fig. 51 A); the type locality of Laos, Houaphanh Province, Xam Neua District, Mount Phu Phan; China, Yunnan Province, Maguan County, Gulinqing town (古林箐乡) (observed by Xiang-Jing Liu, China; Fig. 52); and three localities from Thailand, one from Nan Province, Bo Kluea Tai District (observed by Tatsatorn Dharithai, Thailand, Fig. 51 B), from San Ku Ruins, Chiang Mai (observed by Rob Thacker, United Kingdom; Fig. 51 C), and from Phetchabun Province, Phetchabun Research Station.	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
553D7DA04C9C541383562C995A6AEAD2.taxon	materials_examined	Material examined. (1 ♀, 1 ♂): 1 ♂: " China: Hainan Island, Jianfengling Park: Tropical forest with saturated moisture at night, nymph found on 2 m tall shrub: 13 th July 2019 leg. Yingtong Wang. " (Coll RC 19 - 156). Photographic records: 1 ♀: " 中国海南省乐东黎族自治县 " (Ledong Li Autonomous County, Hainan Province, China) (iNaturalist user @ chenhanlin) (https: // www. inaturalist. org / observations / 34767801)	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
553D7DA04C9C541383562C995A6AEAD2.taxon	distribution	Distribution. At present only known from a few localities on Hainan Island, China (Baisha Li Autonomous County and Ledong Li Autonomous County).	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
19CD5EF761A957D89DA569CC8CCB7D94.taxon	materials_examined	Material examined. Holotype ♂: " Coll. I. R. Sc. N. B., Vietnam, Cat Tien N. P., 11 ° 26 ' N 107 ° 26 ' E, 6 - 16. vii. 2012, Leg. J. Constant and J. Bresseel, I. G.: 32.161 ". Deposited in RBINS. Paratypes (5 ♀♀, 4 ♂♂): • 1 ♀: same data as HT (RBINS) • 1 ♂, 3 ♀♀: same data as HT, " Ex Breeding: Bruno Kneubuehler " (2 ♀♀: RBINS ♂; 1, • 1 ♀: VNMN) • 1 ♂: same data as HT, " Ex Breeding Tim Bollens " [damaged; vomer dissected] (RBINS) • 1 ♀: same data as HT, " Ex Breeding Tim Bollens " (RBINS) • 1 ♂: " Vietnam, Dong Nai Province, Cat Tien N. P., bred by Bruno Kneubu ̈ hler (Switzerland), circa 2012 " (Coll OC) • 1 ♂: " Vietnam, Binh Thuan Province, Dong Tien, IV. 2019 ", molecular sample SLT 03 in our analysis (Coll SLT).	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
19CD5EF761A957D89DA569CC8CCB7D94.taxon	distribution	Distribution. At present only known from two southern Vietnamese provinces, Dong Nai and Binh Thuan Provinces.	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
19CD5EF761A957D89DA569CC8CCB7D94.taxon	description	Description. Female. Coloration. Coloration description is based upon both living wild caught (Fig. 54 B) and captive reared individuals (Fig. 55 A) as their coloration appears to be rather similar. Overall coloration is pale green, with many areas of the body highlighted with cream or very pale green. These areas tend to be the margins on the lobes of the legs, some striping on the lobes of the legs, the pronotum, abdominal margins, and the venation in the tegmina (Fig. 55 A). Morphology. Head. Head capsule about as long as wide, vertex relatively smooth with the only notable feature being the posteromedial tubercle which is finely pointed (Fig. 56 E). Frontal convexity broad and blunt, with a slightly granular surface. Compound eyes slightly protruding from the head capsule, and are not particularly large, taking up slightly <⅓ of the head capsule lateral margins (Fig. 56 E). Ocelli absent. Antennal fields slightly wider than the width of the first antennomere. Antennae. Antennae consist of nine segments, with the terminal segment about the same length as the preceding two and a half segments' lengths combined (Fig. 56 C). Antennomeres I-VII sparsely marked with small transparent setae, the terminal two antennomeres are covered in stout, brown setae. Thorax. Pronotum with gently concave anterior margin and slightly convex lateral margins, which converge to a straight posterior margin that is half the width of the anterior margin (Fig. 56 E). The pronotum surface is smooth, with only a prominent pit in the center, and slight furrows anterior and lateral to the pit (Fig. 56 E). The pronotum has moderately formed anterior and lateral rims and a weakly formed posterior rim, all of which are relatively smooth (Fig. 56 E). Prosternum and the anterior of the mesosternum are covered with numerous nodes, but the remainder of the mesosternum and the metasternum are relatively smooth. Prescutum longer than wide, lateral rims with nine to eleven small to medium tubercles, not ranging dramatically in size giving the margin a rough textured appearance (Fig. 56 E). Prescutum anterior rim prominent but not strongly protruding, rim surface is granular, lacking a large sagittal spine (Fig. 56 F). Prescutum surface heavily granular, with all about the same size. Mesopleura begin ca. ⅓ of the way through the prescutum length and evenly diverge; lateral margin with seven or eight small tubercles with ca. 1 / 2 of those slightly larger than the rest, with the smaller ones interspersed throughout (Fig. 56 E). Face of the mesopleura smooth, with two notable divots, one on the anterior margin and one near the middle (Fig. 56 F). Wings. Tegmina long, reaching 1 / 2 through abdominal segment VII. Tegmina venation; the subcosta (Sc) is the first vein in the forewing, running parallel with the margin for the first half, and then bending and running towards the margin. The radius (R) spans the central portion of the forewing with two subparallel branched veins; the first radius (R 1) branches ca. 1 / 4 of the way through the wing length and terminates slightly proximal to the midline, the radial sector (Rs) branches ca. ⅖ of the way through the wing length and terminates near the distal ⅓ of the wing length. There is a weak continuation of the radius following the prominent Rs branching which continues on as a short and thin R-M crossvein that connects the two veins. The media (M) is simply bifurcate with both the media anterior (MA) and media posterior (MP) terminating near to the posterior 1 / 4 of the wing. The cubitus (Cu) is also bifurcate, branching near the posterior ⅕ of the wing into the cubitus anterior (CuA) and cubitus posterior (CuP) which both terminate at or very near the wing posterior apex. The first anal vein (1 A) is simple and fuses with the cubitus early on, at the length about midway between the splitting of the R 1 and Rs. Alae reduced, with their apex only just passing the anterior margin of abdominal segment III. Abdomen. Abdominal segments II through the anterior half of IV uniformly diverging. The posterior half of segment IV through segment VII are subparallel, gradually converging, and segment VII is ending in a slightly rounded lobe. Segments VIII-X are notably narrower than the previous segments, and have converging margins to the broad rounded apex (Fig. 56 G). Genitalia. Subgenital plate starts at the anterior margin of segment VIII, is moderately broad, and extends 1 / 2 onto segment X with straight margins ending in a fine point (Fig. 56 H). Gonapophyses VIII are long and moderately broad, reaching the apex of abdominal segment X; gonapophyses IX are shorter and narrower, hidden below (Fig. 56 H). Cerci flat, not strongly cupped, with a granular surface and few detectable setae (Fig. 56 H). Legs. Profemoral exterior lobe broad, rounded, and obtusely angled, smoothly arcing from end to end, ca. ⅓ again wider than the width of the interior lobe (Fig. 56 D). Edge of the profemoral exterior lobe granular, with a few slightly larger than the rest, but none very large to resemble teeth (Fig. 56 D). Profemoral interior lobe ca. 2 x as wide as the greatest width of the profemoral shaft, obtusely angled, and marked with five teeth arranged in a two-one-two pattern with looping gaps between them (Fig. 56 D). Mesofemoral exterior lobe arcs from end to end but is slightly bent in the center, weighted towards the distal 1 / 2, and marked with three or four small serrate teeth distributed on the distal half only. Interior lobe is about the same width as the mesofemoral shaft, and the exterior lobe is slightly wider. Mesofemoral interior lobe arcs smoothly end to end with 6 - 8 small serrate teeth only on the distal half of the arc which is slightly wider than the proximal half of the arc. Metafemoral interior lobe arcs end to end, with the distal half slightly wider than the proximal half and marked with 7 - 10 serrate teeth on the distal half of the lobe. Metafemoral exterior lobe is thin and smooth, hugging the metafemoral shaft and lacks dentation. Protibiae lacking an exterior lobe. Protibiae interior lobe spans the entire length of the protibiae and is slightly> 2 x the width of the protibiae shaft itself. The lobe is roundly triangular with the widest portion on the distal half. Mesotibiae and metatibiae lacking exterior and interior lobes. Measurements of paratype females [mm] (wild caught). Length of body (including cerci and head, excluding antennae) 76.7, length / width of head 7.9 / 6.1, antennae 3.8, pronotum 5.1, mesonotum 6.8, length of tegmina 45.8, length of alae 17.9, greatest width of abdomen 30.0, profemora 17.6, mesofemora 13.8, metafemora 17.1, protibiae 11.6, mesotibiae 9.8, metatibiae 12.7. Measurements of paratype females [mm] (ex culture) 3. Length of body (including cerci and head, excluding antennae) 79.4 - 90.3, length / width of head 8.2 - 9.5 / 6.4 - 7.4, antennae 3.8 - 4.3, pronotum 5.1 - 5.8, mesonotum 7.0 - 7.5, length of tegmina 45.3 - 53.8, length of alae 17.2 - 22.9, greatest width of abdomen 28.7 - 36.3, profemora 17.8 - 20.7, mesofemora 14.7 - 15.6, metafemora 16.6 - 19.5, protibiae 11.5 - 12.0, mesotibiae 10.3 - 11.0, metatibiae 13.4 - 14.8. Male. Coloration. Coloration based upon live bred specimens in captivity (Fig. 55 C). Overall coloration pale green throughout with variable patches of tan to reddish coloration (Fig. 55 C). These tan to reddish areas are primarily around the margins of the lobes of the legs, the margins of the thorax, the tips of the antennae, and the margins of the abdomen. In darker colored specimens the mesofemoral lobes can also have coloration, not just along the margins. Abdominal segment V has a pair of slightly transparent eye spots. Morphology. Head. Head capsule about as long as wide, with a vertex that is relatively smooth with only sparse granulation throughout. Frontal convexity stout with sparse thin setae. The posteromedial tubercle is not broad but is distinctly raised from the head capsule. Compound eyes large and bulbous, taking up slightly <1 / 2 head capsule lateral margins (Fig. 57 D). There are three well-developed ocelli located between and slightly posterior to the compound eyes. Antennae. Antennae (including the scapus and pedicellus) consists of 24 segments, all segments except the scapus and pedicellus and terminal three segments are covered in dense setae that are as long as or longer than the antennae segment is wide. The terminal three segments are covered in dense short setae and the scapus and pedicellus are nearly completely bare. Thorax. Pronotum with anterior margin distinctly concave and lateral margins that are slightly convex and converging to a straight posterior margin that is ca. 1 / 2 the width of the anterior rim. Anterior and lateral margins of the pronotum have moderately formed rims and the posterior margin lacks a rim. Face of the pronotum is marked by a distinct furrow and pit in the center and a relatively smooth surface with weak granulation. Prosternum surface is weakly granular with small nodes of even size and spacing. Mesosternum surface marked with slightly more prominent nodes, with the largest along the sagittal plane and more strongly on the anterior margin, posterior margin with less prominent and smaller nodes. Prescutum slightly longer than wide, with lateral margins that are only slightly converging to the posterior (Fig. 57 D). Lateral rims with nine or ten node-like tubercles, giving the lateral margins a rough textured appearance. Prescutum surface with minimal nodes throughout, with those along the sagittal plane slightly larger than the others. Prescutum anterior rim prominent but not strongly raised, with a granular surface and lacking a prominent sagittal tubercle. Mesopleura begin on the anterior prescutum margin but are narrow throughout most of their length, only diverging gently for the posterior ⅔. Lateral margin with nine or ten minor tubercles throughout the length except for the posterior ⅓ which is relatively smooth. Face of the mesopleura mostly smooth, with slight wrinkling throughout. Wings. Tegmina moderate length, extending ⅓ of the way through abdominal segment III. Tegmina wing venation: the subcosta (Sc) is the first vein, is simple, and terminates the earliest ca. ⅓ of the way through the overall tegmina length. The radius (R) spans the entire length of the tegmina with the first radius (R 1) branching just proximal to the midline and terminating just distal to the midline, followed by the branching and termination of the second radius (R 2) near the distal ⅓ of the wing, and then the radial sector runs to the wing apex. The media (M) also spans the entire length of the tegmina with the first media posterior (MP 1) branching off slightly> ⅓ of the way through the wing length, and then the second media posterior (MP 2) branches just distal to the midline, and the media anterior (MA) runs to the wing apex. The cubitus (Cu) runs along the edge of the wing as the two media posterior veins fuse with it and as the cubitus reaches the apex it fades. The first anal (1 A) vein terminates upon reaching the cubitus ca. ⅓ of the way through the wing length. Alae well-developed in an oval fan configuration, long, reaching to the middle or posterior of abdominal segments IX. Alae wing venation: the costa (C) is present along the entire foremargin giving stability to the wing. The subcosta (Sc) is long, spanning ca. ⅔ of the wing length and is mostly fused with the radius in the beginning but terminates when it meets the costa. The radius (R) spans the entire wing and branches slightly proximal to the midline into the first radius (R 1) and radial sector (Rs) which run gently diverging for most of their length and then converge at the apex of the wing where they terminate near each other but not touching. The media (M) branches early, ca. ⅙ of the way through the wing into the media anterior (MA) and the media posterior (MP) which run parallel with each other throughout the wing until the distal ⅕ of the wing where the media posterior fuses with the media anterior which then run fused together to the wing apex where they terminate near the radial sector. The cubitus (Cu) runs unbranched and terminates at the wing apex. Of the anterior anal veins, the first anterior anal (1 AA) fuses with the cubitus near the point where the media branches into the media anterior and media posterior and then the first anterior anal branches from the cubitus ⅔ of the way through the wing length where it uniformly diverges from the cubitus until it terminates at the wing margin. The anterior anal veins two-seven (2 AA- 7 AA) have a common origin and run unbranched in a folding fan pattern of relatively uniform spacing to the wing margin. The posterior anal veins (1 PA- 6 PA) share a common origin separate from the anterior anal veins and run unbranched to the wing margin with slightly thinner spacing than the anterior anal veins. Abdomen. Lateral margins of abdominal segment II are parallel, III through the anterior ⅔ of segment IV gradually diverging, the remainder of IV and segment V are parallel-sided, segment VI starts parallel-sided but almost immediately starts to converge and the remaining segments converge uniformly to the rounded apex of the abdomen. Genitalia. Poculum broad and ends in a rounded apex that slightly passes the anterior margin of segment X (Fig. 57 G). Cerci long and slender, extending from under the anal abdominal segment, nearly flat, not strongly cupped, covered in a granulose surface and numerous short setae (Fig. 57 F). Vomer broad and stout with straight sides evenly converging and ending in a thick apical hook with a smaller second hook adjacent to it (Fig. 5 C). Legs. Profemoral exterior lobe about the same width as the interior lobe, ca. 21 / 2 x the greatest width of the profemoral shaft, roundly arcing end to end in a broad obtuse angle that is not distinctly bent, with the proximal margin slightly granulose, and the distal margin with four or five small serrate teeth (Fig. 57 C). Profemoral interior lobe roundly triangular and marked with five sharp teeth arranged in a two-one-two pattern with looping gaps between them (Fig. 57 C). Mesofemoral exterior lobe arcs end to end but is slightly wider on the distal ⅓ which is marked with four or five serrate teeth, and the proximal half that is rather thin. Mesofemoral interior lobe is about the same width as the exterior, is broader on the distal end and is marked with seven or eight small serrate teeth. Metafemoral exterior lobe lacks dentation and has a straight margin along the metafemoral shaft. Metafemoral interior lobe smoothly arcs end to end with eight or nine small serrate teeth on the distal ⅔, which is slightly wider than the proximal ⅓. Protibiae lacking exterior lobe, interior lobe reaching end to end in a smooth triangle which is slightly weighted to the distal 1 / 2 and at its widest is ca. 21 / 2 x as wide as the protibial shaft (Fig. 57 C). Meso- and metatibiae simple, lacking lobes completely. Measurements of holotype male [mm]. Length of body (including cerci and head, excluding antennae) 58.7, length / width of head 3.8 / 3.5, antennae 38.2, pronotum 2.9, mesonotum 3.8, length of tegmina 17.3, length of alae 42.9, greatest width of abdomen 13.4, profemora 12.8, mesofemora 10.6, metafemora 13.0, protibiae 10.1, mesotibiae 7.6, metatibiae 10.0. Measurements of paratype male [mm] (ex culture). 4 Length of body (including cerci and head, excluding antennae) 54.3, length / width of head 4.0 / 3.1, antennae 36.1, pronotum 2.7, mesonotum 4.3, length of tegmina 15.5, length of alae 41.1, greatest width of abdomen 13.7, profemora 11.6, mesofemora 10.1, metafemora 12.2, protibiae 8.05, mesotibiae 7.4, metatibiae 8.9. Eggs. (Fig. 58). The lateral surfaces are flat but with the posterior half slightly wider than the anterior half. The center of the dorsal surface is slightly convex, which gives the margin a slight undulating appearance when viewed from the lateral aspect as the middle is thinner than either end of the egg. When viewed from the lateral aspect; the ventral margin has the posterior slightly protruding more than the anterior, adding to the overall undulating shape of the egg. All surfaces have numerous small to medium sized pits throughout, the lateral surface has around 20 pits arranged in no particular order. The surfaces are also covered with short moss-like pinnae interspersed throughout the capsules with those along the margins slightly longer than the other surfaces. The dorsal surface is marked with six slightly irregular medium sized pits on each half and short moss-like pinnae around the micropylar plate. The micropylar plate is long, ca. 6 / 7 of the overall dorsal surface length and the shape is nearly symmetrical with the anterior and posterior thin and the area around the micropylar cup the widest point. Micropylar cup of moderate size and placed slightly posterior to the micropylar plate midline. Operculum slightly ovular, with the outer margin with a distinct row of moss-like pinnae surrounding the operculum and four or five medium pits surrounding the dorsal and lateral margins. The operculum is roundly raised with a height slightly> 1 / 2 operculum width, this rounded raised cap is marked with a sagittal raised row of pinnae similar in length to those along the capsule margins. The overall color is tan to light brown, with the moss-like pinnae sometimes slightly lighter in color. Measurements including the extended pinnae [mm]. Length (including operculum): 5.1; maximum width of capsule when viewed from lateral aspect 3.2 - 3.3; length of micropylar plate 2.9 - 3.2 Newly hatched nymphs. (Fig. 9 G). The general color throughout the body is dark brown with slightly lighter brown on the legs. The basitarsi are yellow and remaining tarsal segments are dark brown. All tibiae lack exterior lobes, and all have smoothly arcing interior lobes which have several tan to brown stripes throughout their length. All femoral lobes are similar in width and have distinct serration on their distal halves. The interior profemoral lobe lacks a white spot, but the exterior lobe has a narrow white crescent on the proximal ⅓ with an additional small white patch at the proximal most margin. The meso- and metafemoral interior lobes have two white patches, one on the proximal most edge, and a larger white patch ⅓ of the way through the length. The meso- and metafemoral exterior lobes also have the large white patch on the proximal ⅓, but lack a smaller white patch on the proximal most margin. The distal ends of the meso- and metafemora also have minimal white edges. The abdomen is mostly brown, but abdominal segments II and III have distinct green patches on their lateral surfaces (the centerline of the abdomen is uniform brown throughout). The terminal three abdominal segments also have a little bit of green on their margins. The widest point of the abdomen is abdominal segment IV.	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
19CD5EF761A957D89DA569CC8CCB7D94.taxon	etymology	Etymology. Patronym. This species is dedicated to Pham Hong Thai (VNMN), a good friend and colleague who co-organized the GTI entomological expeditions to Vietnam with the RBINS team since 2010.	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
64EBAECAF9BB521EAAB400E22E3CB59E.taxon	materials_examined	Material examined. (5 ♀♀, 7 ♂♂): 1 ♀: " China: Guangxi Province, Nanning city, Shuangding town （ 双定镇 ） N 22 ° 59 ' 41.39 " E 108 ° 6 ' 53.59 ", VI- 2016, Xiao-Yu Zhu. Molecular sample: DZW 08 " (Coll ZD); 1 ♀: " China, Guangxi, Liuzhou, Dayaoshan Mts, IX. 2019 " (IMQC); 1 ♀: " China, Guangxi, Jinxiu County, Dayao Mountain, July-Oct 2020 " (Coll SLT); 1 ♀: " China, Guangxi, Jinxiu County, Dayao Mountain, Oct 2020 " (Coll SLT); 1 ♀: " Museum Paris, Tonkin, reg. de Hoa-Binh, A. De Cooman 1927 " (MNHN); 1 ♂: " Kon Tum Province, Ngoc Linh Mt. 1,700 m. elv. May 2015 " (Coll RC 16 - 116); 1 ♂: " Vietnam: Da Nang Prov., Ba Na Mt.: May, 2015 1,450 m. " (Coll RC 16 - 115); 1 ♂ " Da Nang Province, Ba Na Mt. 1,450 m. elv. May 2015 " (Coll RC 16 - 119); 1 ♂: " Coll. I. R. Sc. N. B., Vietnam, Ninh Binh prov., Cuc Phuong Nat. Park, 20 ° 20 ' 53 " N 105 ° 35 ' 52 " E, 31. vii- 3. viii. 2016, GTI Project, Leg. J. Constant & J. Bresseel, I. G.: 33.282, RBINS-PHYLLIUM DNA sample 0004 " [vomer dissected] (RBINS); 1 ♂: " Coll. I. R. Sc. N. B., Vietnam, Ninh Binh prov., Cuc Phuong Nat. Park, 20 ° 20 ' 53 " N 105 ° 35 ' 52 " E, 2 - 8. vii. 2019, Leg. J. Constant, I. G.: 34.032 " (RBINS); 1 ♂: " Coll. I. R. Sc. N. B., Vietnam, Tay Yen Tu Nat. Res., 21 ° 11 ' 10 '' N 106 ° 43 ' 25 " E, 7 - 11. vii. 2013, night collecting, Leg J. Constant & J. Bresseel, I. G. 32.454 " (VNMN); 1 ♂: " Coll. I. R. Sc. N. B., Vietnam, Tay Yen Tu Nat. Res., 21 ° 11 ' 10 '' N 106 ° 43 ' 25 " E, 7 - 11. vii. 2013, night collecting, Leg J. Constant & J. Bresseel, I. G. 32.454, RBINS-PHYLLIUM DNA sample 0005 " (RBINS).	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
64EBAECAF9BB521EAAB400E22E3CB59E.taxon	distribution	Distribution. China, Guangxi Province (recorded from Hexian, Shuangding town, and Dayaoshan Mountains) and distributed south through Vietnam. At present we have records> for five Vietnamese provinces: Vinh Phuc, Quang Ninh, Ninh Binh, Da Nang, and Kon Tum Provinces.	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
A26EBE1EFDCC5CDB863F9069FF5C0F01.taxon	materials_examined	Material examined. (7 ♀♀, 4 ♂♂, 10 eggs): 1 ♂: " Tibet, Nyingchi Area, De'Ergong Village, Motutown 2020 " (Coll SLT); 1 ♂: " China, Tibet, Motuo, Beibeng. 2019. 6 - 8. " (Coll RC 20 - 001); 1 ♀ nymph: " Crowley Bequest. 1901 - 78. Sikkim " (NHMUK); 1 ♀ nymph: " Arunachal Pradesh from the Mishmi Hills. Delei River. 1,700 ft. 28. i. 1935, M. Steele. " (NHMUK); 1 ♂ nymph: " Arunachal Pradesh from the Mishmi Hills. Lohit River. 22. iii. 1935, M. Steele. " (NHMUK); 1 ♂, 1 ♀: " bred from material collected in: Tibet China: Beibeng Township, Medog County, IX- 2016, Jin Chen. " (Coll ZD); 10 eggs: " China: Tibet: Medong Region: Bred by Bruno Kneubu ̈ hler, 2018 " (Coll RC 18 - 396 - 18 - 405). Photographic records: 1 ♀: " Kalimpong, West Bengal, India, September, 2019, photographed by Vandana Wadwa Sood (West Bengal, India) "; 1 ♀ nymph: " Digboi, Assam 786171, India, iNaturalist user @ rajib, by Rajib Rudra Tariang " (https: // www. inaturalist. org / observations / 61945900); 1 ♀: " Samthar, iNaturalist user @ ripbumlepcha " (https: // www. inaturalist. org / observations / 35911522); 1 ♀: " Pasighat in the East Siang district, observed and photographed by Oken Tayeng "	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
A26EBE1EFDCC5CDB863F9069FF5C0F01.taxon	distribution	Distribution. The type locality for Cryptophyllium tibetense comb. nov. stated by Liu (1993) is Tibet (Xizang Autonomous Region), Medog County (Motuo). There do appear to be additional distribution records> for this high elevation species however. Unfortunately, we have only been presented with photographs of individuals or antique subadults / nymphs, and no fresh adult specimens to examine or include in our molecular phylogeny at this time. Therefore, these additional distribution records> are only presumed to be Cryptophyllium tibetense comb. nov. due to their morphological similarity to bred nymphs of this species and are here presented to give as thorough a view into the Cryptophyllium gen. nov. distribution as possible. From Southeast Tibet, Tenga Valley, we have been presented with an image of an adult male which looks to be the right size and has morphological features which suggest it could represent the Cryptophyllium tibetense comb. nov. male. Additionally, there is a pair of nymphs in the NHMUK from " Arunachal Pradesh from the Mishmi Hills " from 1935, a female from Anjaw District " Delei River. 1,700 ft. 28. i. 1935, M. Steele. " (Fig. 65 C) and a male from the Lohit District " Lohit River. 22. iii. 1935, M. Steele. " (Fig. 65 D). Recently we also were sent photos of a female from Pasighat in the East Siang district, which matches well with the Cryptophyllium tibetense comb. nov. morphology (Fig. 66 A). From Nepal we are aware of two records>, both female subadults, one found in Gandaki Pradesh, Tanahun District and the other found in Province No. 1, Ilam. Both of these subadult females appear to have Cryptophyllium tibetense comb. nov. shaped abdomens, profemoral lobes, and importantly small exterior tibial lobes which help to characterize Cryptophyllium tibetense comb. nov. females. From India we have located a nymph from within the NHMUK collection from Sikkim (Fig. 65 B), and we have been lucky enough to be presented with photographs of an adult female from West Bengal, Kalimpong (Fig. 65 A) both of which have Cryptophyllium tibetense comb. nov. like features. From the state of Assam we have been sent an image of a female nymph from the town of Digboi which can clearly be seen as having distinct but small exterior tibial lobes (Fig. 65 E). Interestingly, we have yet to be presented with records> from Bhutan, but as it lies between areas where Cryptophyllium gen. nov. species have been confirmed we expect that there likely is at least one species present, just not yet officially recorded.	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
5B088CA6B89C59CD808B3FABF59F3EC1.taxon	materials_examined	Material examined. Holotype ♀: " CHINA: Yunnan Province, Chashan Park, Simao District, Puer City, VI- 2017, leg. Xiao-Yu Zhu ". deposited in the Kunming Institute of Zoology (KIZ), Yunnan, China.	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
5B088CA6B89C59CD808B3FABF59F3EC1.taxon	distribution	Distribution. At present only known from the type locality of Puer City in Yunnan Province, China.	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
5B088CA6B89C59CD808B3FABF59F3EC1.taxon	description	Description. Female. Coloration. Based upon the singular dried holotype specimen (Fig. 67). Living individuals are always a more vibrant green and this specimen appears to have been fully green, without patches of variable brown. The dried holotype female is pale green throughout with discoloration of tan along the shafts of the legs and head, thorax, and abdomen likely due to the drying process. Morphology. Head. Head capsule slightly longer than wide, vertex relatively smooth with only slight granulation throughout the surface, all relatively well-spaced (Fig. 67 D). The posteromedial tubercle is broader and taller than any other nodes on the head capsule (Fig. 67 F). Frontal convexity broad and about as long as the first antennomere, with a slightly lumpy surface, and with several setae present throughout (Fig. 67 D). Compound eyes slightly protruding from the head capsule, not notably large, taking up slightly <1 / 4 of the length of the lateral head capsule margins (Fig. 67 D). Ocelli absent. Antennal fields wider than the first antennomere (Fig. 67 D). Antennae. Antennae consisting of nine segments, with the terminal segment about the same length as the preceding two segments' lengths combined (Fig. 67 B). Antennomeres I-VIII sparsely marked with small transparent setae, the terminal antennomere has darker, shorter, and denser setae than the other segments (Fig. 67 B). Thorax. Pronotum with gently concave anterior margin and straight lateral margins, which converge to a convex posterior margin that is half the width of the anterior margin (Fig. 67 D). The pronotum surface and moderately formed pronotum rims are smooth, lacking significant granulation, with only a prominent pit in the center, and slight furrows anterior, posterior, and lateral to the pit (Fig. 67 D). Prosternum with moderate granulation, mesosternum anterior margin and the anterior half of the sagittal plane with moderate granulation. Metasternum relatively smooth, lacking notable nodes. Prescutum as long as wide, lateral rims with nine or ten irregularly shaped but not large tubercles with those on the anterior slightly larger than those on the posterior (Fig. 67 D). Prescutum anterior rim prominent but not strongly protruding, surface marked throughout with irregular granulation, no prominent singular sagittal spine present (Fig. 67 D). Prescutum surface covered irregularly by moderate nodes, with slightly larger nodes along the sagittal plane (Fig. 67 D). Mesopleura beginning on the anterior margin of the prescutum and evenly diverging; lateral margin with five larger tubercles, and eight or nine smaller tubercles interspersed unevenly throughout with some tubercles touching side by side or with a slight gap between them (Fig. 67 D). Face of the mesopleura is mostly smooth but with lateral margins that have a slightly granular surface (Fig. 67 D). Wings. Tegmina long, reaching 1 / 2 through abdominal segment VII. The subcosta (Sc) is the first vein in the forewing and runs parallel with the tegmina lateral margin for the first 1 / 2 of the vein, then bends gently and runs to the lateral margin of the wing where it terminates ca. 1 / 4 of the way through the length. The radius (R) spans the central portion of the forewing with two subparallel branched veins; radius 1 (R 1) terminates ca. ⅓ of the way through the wing length, and the radial sector (Rs) terminates ca. ⅔ of the way through the wing length. There is a weak continuation of the radius following the prominent Rs branching which continues on as a short and thinner R-M crossvein that does not solidly connect the two veins as it reaches the media. The media (M) is simply bifurcate with both the media anterior (MA) and media posterior (MP) terminating near the posterior 1 / 4 of the wing. The cubitus (Cu) is also bifurcate, branching near the posterior ⅕ of the wing into the cubitus anterior (CuA) and cubitus posterior (CuP) which both terminate at or very near the wing posterior apex. The first anal vein (1 A) is simple and fuses with the cubitus early on, near the branching distance of the R 1 from R. Alae in the holotype are unfortunately deformed, so we do not yet know what length alae this species has. Abdomen. Abdominal segments II through the anterior ⅓ of IV diverging slowly at first and then more strongly towards the posterior, with the posterior ⅔ of segment IV the widest segment. Segments V-VI are subparallel, and segment VII converges slightly more prominently and ends in a slight lobe. Segments VIII-X are notably narrower than the previous segments and converge uniformly to the rounded apex. Genitalia. Subgenital plate starts at the anterior margin of segment VIII, is long and narrow reaching significantly onto the terminal segment (Fig. 67 G). Gonapophyses VIII are long and moderately broad, exceeding the apex of abdominal segment X, gonapophyses IX are smaller and slender, hidden below the gonapophyses VIII (Fig. 67 G). Cerci flat, not strongly cupped, with a heavily granular surface and few detectable setae (Fig. 67 E). Legs. Profemoral exterior lobes notably wider than the interior lobe with an acute angle due to a slight recurve of the lobe (Fig. 67 C). Proximal edge of the profemoral exterior lobe slightly granular, and the bend and distal edge is marked by six or seven small serrate teeth (Fig. 67 C). Profemoral interior lobe ca. 3 x as wide as the greatest width of the profemoral shaft, with an obtuse angle, and marked with five prominent teeth arranged in a two-one-two pattern with large looping gaps between the teeth (Fig. 67 C). Mesofemoral exterior lobe arcs from end to end in a slightly bent lobe weighted on the distal half and marked with two serrate teeth on the distal half only. Interior and exterior lobes are of a similar width. Mesofemoral interior lobe arcs smoothly end to end, is marked with four or five serrate teeth only on the distal half of the arc, and is about as wide as the mesofemoral shaft. Metafemoral interior lobe arcs end to end, but is wider on the distal half, and has four or five serrate teeth on the distal half of the lobe only. Metafemoral exterior lobe is thin and smooth, hugging the metafemoral shaft. Protibiae interior lobe spans the entire length of the protibiae and is at least 2 x as wide as the protibial shaft. The lobe is distinctly triangular with the broadest point distal to the midline. Pro-, meso-, and meta- tibiae with small anteriorly situated exterior lobes, meso-, and meta- tibiae lack interior lobes. Measurements of holotype female [mm]. Length of body (including cerci and head, excluding antennae) 78.5, length / width of head 7.9 / 7.5, antennae 4.6, pronotum 5.5, mesonotum 6.2, length of tegmina 54.5, length of alae (unknown, deformed in the holotype), greatest width of abdomen 31.8, profemora 17.5, mesofemora 13.0, metafemora 14.2, protibiae 9.7, mesotibiae 9.0, metatibiae 12.5.	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
5B088CA6B89C59CD808B3FABF59F3EC1.taxon	etymology	Etymology. Patronym. Named in honor of Zhiwei Dong's wife, Ms. Wen-Na Chen, for her support and love over the years.	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
602EFA0F6D165D669074356F7C502A98.taxon	materials_examined	Material examined. Neotype ♂: " THAILAND: Chiang Mai Province: October 2010. Coll RC 16 - 148 ". Deposited within the Montreal Insectarium (IMQC). Molecular sample 16 - 148 within this study.	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
602EFA0F6D165D669074356F7C502A98.taxon	materials_examined	Additional material examined. (15 ♀♀, 21 ♂♂, 4 eggs): 3 ♀♀: " Thailand: Chiang Mai, July 2017. " (Coll RC 18 - 145, 18 - 146, 18 - 147); 1 ♀: " Chiang Mai, Fang: February, 2011 " (Coll RC 16 - 211); 1 ♀: " Thailand, Fang, II- 2011 " (Coll RC 16 - 212); 1 ♀: " Lamphun Province, Maetha: September, 2011 " (Coll RC 16 - 080); 1 ♀: " Thailand, Lamphun Province, 2009 November " (Coll RC 16 - 078); 1 ♀: " Northern Thailand, Chiang Mai Province, 2010, October " (Coll RC 16 - 079); 1 ♀: " Laos: Luang Prabang Province, Kiew Mak Nao Village, 900 m.: June, 2014 " (Coll RC 16 - 077); 1 ♂: " Thailand: Chiangmai, Doi Pui, 25 May 1985 " (Coll RC 16 - 082); 1 ♂: " Thailand: Lampon, Mae Tha, 09 / 2011 " (Coll RC 16 - 083); 1 ♂: " Thailand: Chiangmai, Doi Pui, 19 May 1985 " (Coll RC 16 - 214); 1 ♂: " Thailand: Chiangmai, Doi Pui, 28 May 1985 " (Coll RC 16 - 215); 1 ♂: " Thailand: Chiangmai, Doi Pui, 24 May 1985 " (Coll RC 16 - 216); 1 ♂: " Thailand: Chiangmai, Doi Pui, 25 May 1985 " (Coll RC 16 - 217); 1 ♂: " Thailand: Lampang, May 2001 " (Coll RC 16 - 218); 1 ♂: " North Laos: Kiew Mak Nao, VII. 2015, 900 m. S. Collard leg " (Coll RC 18 - 030); 2 ♂♂: " Thailand: Lampoon, Mae Tha, 09 - 2011 " (Coll RC 16 - 147, 16 - 213); 1 ♂: " Burma: 4 km E. Karathuri, Top of Hill, 350 to 400 m., VI. 2011., Coll. A. Banko / Collected by beating tree in Forest " (Coll RC 18 - 029); 1 ♂: " Coll. I. R. Sc. N. B., Thailande (Loei), Na Haeo (bio station), 05 - 12. V. 2001, Light trap, Leg. J. Constant & P. Grootaert " [vomer dissected] (RBINS); 2 ♂♂: " Coll. I. R. Sc. N. B., Thailande (Loei), Na Haeo (field res stat )), 15 - 19. V. 2003, Light trap, Leg. J. Constant, K. Smets & P. Grootaert " (RBINS); 1 ♂: " Coll. I. R. Sc. N. B., Thailande (Loei), Na Haeo, light trap, 15 - 19. V. 2003, Light trap, Leg. J. Constant & K. Smets " (RBINS); 1 ♂: " Coll. I. R. Sc. N. B., Thailande (Loei), Na Haeo, light trap, 15 - 19. V. 2003, Light trap, Leg. J. Constant & K. Smets, RBINS-PHYLLIUM DNA sample 0002 " (RBINS); 1 ♂: " Coll. I. R. Sc. N. B., Thailande (Loei), Na Haeo, forest clearing, light trap, 16. V. 2003, Light trap, Leg. J. Constant & K. Smets " (RBINS); 1 ♂: " Coll. I. R. Sc. N. B., Thailand, Loei, Na Haeo, 22. V. 2000, Station 20007, Leg P. Grootaert " (RBINS); 1 ♂: " Coll. I. R. Sc. N. B., Laos, Bokeo prov., Ban Muang Kan, 1 - 15. vi. 2012, local collectors, I. G.: 32.213, RBINS-PHYLLIUM DNA sample 0012 " (RBINS); 1 ♀: " Coll. I. R. Sc. N. B., Laos, Bokeo prov., Ban Muang Kan, 1 - 15. vi. 2012, local collectors, I. G.: 32.213, RBINS-PHYLLIUM DNA sample 0011 " (RBINS); 2 ♀♀: " Coll. I. R. Sc. N. B., Laos, Bokeo prov., Ban Muang Kan, 1 - 15. vi. 2012, local collectors, I. G.: 32.213 " (RBINS); 1 ♀: " Coll. I. R. Sc. N. B., Thailand, SE Chiang Mai, Salok, Wang Chin, Near Lamphang, ex breeding A. & C. Bauduin, 2015 " (RBINS); 1 ♀, 1 ♂: " Coll. I. R. Sc. N. B., Ex breeding Bruno Kneubuehler, 2017, Thailand, Lamphun prov., Tha Pla Duk " (RBINS); 1 ♀, 1 ♂: " Thailand, Ex Culture Kristien Rabaey (RBINS); 4 eggs: N-Thailand, Cultured F. Hennemann 1995 - 2001. Ex. Coll. Frank Hennemann (Germany) " (Coll RC 18 - 242 - 18 - 245).	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
602EFA0F6D165D669074356F7C502A98.taxon	distribution	Distribution. Cryptophyllium westwoodii comb. nov. has only been confirmed through genetic analysis from northern Thailand, northern Laos, and southern Myanmar. With the description of Cryptophyllium khmer sp. nov. which morphologically cannot be differentiated from photographs of nymphs or females, we are unsure where these two species biogeographically are separated, but at this time we only know of Cryptophyllium khmer sp. nov. from Cambodia and are unsure if Cryptophyllium westwoodii comb. nov. also occurs in this country. Until additional Cryptophyllium westwoodii comb. nov. samples from throughout the range are also sequenced, the true distribution must remain somewhat vague at this point (as indicated by the bi-colored symbols in our distribution map; Fig. 2).	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
2547FE3A5146576C863ABAB416DE4B82.taxon	materials_examined	Material examined. (3 ♀♀, 1 ♂): At present only the holotype within the CAS collection, and three additional specimens within the BPBM collection are known to us. 1 ♀: " M. R. Lundgren, Collector. Kaday, Yap, Xi. 10.1980. CASTYPE # 19438, Holotype " (Fig. 73); 1 ♀: " Yap Islands, Yap I., Kolonia. R. P. Owen, Collector. 6 - xii- ' 63, unknown tree. BPBMENT, 0000080399 " (Fig. 74 A); 1 ♀: " Yap, 25. XI. 1940, H. Fujishima. Micronesia coll., Entomology Lab., Kyushu univ., Fukuoka, Japan. BPBMENT, 0000080401. " (Fig. 74 C); 1 ♂: " Yap, S. Ikuta. Micronesia coll., Entomology Lab., Kyushu univ., Fukuoka, Japan. Yap. The specimens was at, first preserved in Formalin, and therefore the coloration, was faded. BPBMENT, 0000080400. " (Fig. 74 E).	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
2547FE3A5146576C863ABAB416DE4B82.taxon	distribution	Distribution. Only known from the areas of Colonia and Kaday on Yap Island, Micronesia.	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
933CCA9F6C54533EBA59DD0BC6428B28.taxon	materials_examined	Material examined. (3 ♀♀, 6 ♂♂): 1 ♀: " Vietnam: Lao Cai Prov. Mt. Fan-si-pan, North Side, 1,600 m., 22 17 ' N 103 44 ' E, Primary Forest, 28. October- 3. November 1994. " (Coll RC 17 - 270); 1 ♀ nymph: " Lao Cai Province, Sapa mt. 1,600 m.: May 2015 " (Coll RC 16 - 081); 1 ♀: " Yunnan China: Daweishan, Pinbian County, Honghe Prefecture, 3 - VI- 2017, Zhiwei Dong. (Coll ZD). Molecular sample: DZW 04. " (Coll ZD); 3 ♂♂: " Vietnam: Lao Cai Prov. Mt. Fan-si-pan, North Side, 1,600 m., 22 17 ' N 103 44 ' E, Primary Forest, 20 - 30. October, 1995 " (Coll RC 17 - 271, 17 - 272, 17 - 273); 1 ♂: " Vietnam: Lao Cai Prov. Mt. Fan-si-pan, North Side, 1,600 m., 22 17 ' N 103 44 ' E, Primary Forest, 1 - 7, November, 1995 " (Coll RC 17 - 274); 1 ♂: " Yunnan Province, Xinping Country, Mt. Ailao: June 2015 " (Coll RC 16 - 120); 1 ♂: " Vietnam: Yen Bai, Nghia Lo: June, 2017 " (Coll RC 17 - 240).	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
933CCA9F6C54533EBA59DD0BC6428B28.taxon	distribution	Distribution. Presently only known from southern China (Yunnan Province) and adjacent northern Vietnam (Lai Chau and Yen Bai Provinces).	en	Cumming, Royce T., Bank, Sarah, Bresseel, Joachim, Constant, Je ́ ro ̂ me, Tirant, Stephane Le, Dong, Zhiwei, Sonet, Gontran, Bradler, Sven (2021): Cryptophyllium, the hidden leaf insects - descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae). ZooKeys 1018: 1-179, DOI: http://dx.doi.org/10.3897/zookeys.1018.61033, URL: http://dx.doi.org/10.3897/zookeys.1018.61033
