taxonID	type	description	language	source
7F4A8799FFF1FF80FF44FC14FABCFBC5.taxon	type_taxon	Type genus: Eodysagrion Rust, Petrulevičius & Nel, 2008 (earliest Eocene, Fur Formation, Denmark).	en	Nel, André (2024): Two new damselflies from the Eocene Green River Formation (Odonata, Zygoptera, Dysagrionidae, Thaumatoneuridae). Zootaxa 5446 (4): 588-594, DOI: 10.11646/zootaxa.5446.4.11, URL: http://dx.doi.org/10.11646/zootaxa.5446.4.11
7F4A8799FFF1FF80FF44FC14FABCFBC5.taxon	description	Other genera: Tynskysagrion Bechly, Garrouste, Aase, Karr, Grande & Nel, 2021, Oreodysagrion Bechly, Garrouste, Aase, Karr, Grande & Nel, 2021, Gusagrion gen. nov. (Eocene, Green River Formation, USA).	en	Nel, André (2024): Two new damselflies from the Eocene Green River Formation (Odonata, Zygoptera, Dysagrionidae, Thaumatoneuridae). Zootaxa 5446 (4): 588-594, DOI: 10.11646/zootaxa.5446.4.11, URL: http://dx.doi.org/10.11646/zootaxa.5446.4.11
7F4A8799FFF1FF80FF44FB08FB2DFA58.taxon	type_taxon	Type species: Gusagrion coloratum sp. nov.	en	Nel, André (2024): Two new damselflies from the Eocene Green River Formation (Odonata, Zygoptera, Dysagrionidae, Thaumatoneuridae). Zootaxa 5446 (4): 588-594, DOI: 10.11646/zootaxa.5446.4.11, URL: http://dx.doi.org/10.11646/zootaxa.5446.4.11
7F4A8799FFF1FF80FF44FB08FB2DFA58.taxon	diagnosis	Diagnosis. Wing venation characters only. Elongate and narrow discoidal cell; long petiole; nodus situated in a very basal position, at 10 % of distance between Ax 2 and wing apex; distal third of wing very broad; narrow anal area below subdiscoidal cell, at most as broad as subdiscoidal cell; only one crossvein below subdiscoidal cell.	en	Nel, André (2024): Two new damselflies from the Eocene Green River Formation (Odonata, Zygoptera, Dysagrionidae, Thaumatoneuridae). Zootaxa 5446 (4): 588-594, DOI: 10.11646/zootaxa.5446.4.11, URL: http://dx.doi.org/10.11646/zootaxa.5446.4.11
7F4A8799FFF1FF80FF44FB08FB2DFA58.taxon	etymology	Etymology. Named after the type locality Gus’ pit and the suffix Agrion. Gender neutral.	en	Nel, André (2024): Two new damselflies from the Eocene Green River Formation (Odonata, Zygoptera, Dysagrionidae, Thaumatoneuridae). Zootaxa 5446 (4): 588-594, DOI: 10.11646/zootaxa.5446.4.11, URL: http://dx.doi.org/10.11646/zootaxa.5446.4.11
7F4A8799FFF1FF82FF44F9F5FED1FC55.taxon	description	(Figure 1) urn: lsid: zoobank. org: act: 7 A 54 A 54 E- 4503 - 4 BD 6 - B 206 - 1 FDF 3 EA 84 AA 0	en	Nel, André (2024): Two new damselflies from the Eocene Green River Formation (Odonata, Zygoptera, Dysagrionidae, Thaumatoneuridae). Zootaxa 5446 (4): 588-594, DOI: 10.11646/zootaxa.5446.4.11, URL: http://dx.doi.org/10.11646/zootaxa.5446.4.11
7F4A8799FFF1FF82FF44F9F5FED1FC55.taxon	materials_examined	Material. Holotype IP. 358925 (part and counterpart of a complete wing), stored at Invertebrate Paleontology Collection, Yale Peabody Museum, USA. Age and outcrop. Eocene (Ypresian), Green River Formation, Parachute Creek member, Gus’ pit, two miles west of Rio Blanco store, Colorado, USA.	en	Nel, André (2024): Two new damselflies from the Eocene Green River Formation (Odonata, Zygoptera, Dysagrionidae, Thaumatoneuridae). Zootaxa 5446 (4): 588-594, DOI: 10.11646/zootaxa.5446.4.11, URL: http://dx.doi.org/10.11646/zootaxa.5446.4.11
7F4A8799FFF1FF82FF44F9F5FED1FC55.taxon	etymology	Etymology. Named after the colored distal third of wing.	en	Nel, André (2024): Two new damselflies from the Eocene Green River Formation (Odonata, Zygoptera, Dysagrionidae, Thaumatoneuridae). Zootaxa 5446 (4): 588-594, DOI: 10.11646/zootaxa.5446.4.11, URL: http://dx.doi.org/10.11646/zootaxa.5446.4.11
7F4A8799FFF1FF82FF44F9F5FED1FC55.taxon	diagnosis	Diagnosis. As for the genus. A dark brown area covering apical third of wing.	en	Nel, André (2024): Two new damselflies from the Eocene Green River Formation (Odonata, Zygoptera, Dysagrionidae, Thaumatoneuridae). Zootaxa 5446 (4): 588-594, DOI: 10.11646/zootaxa.5446.4.11, URL: http://dx.doi.org/10.11646/zootaxa.5446.4.11
7F4A8799FFF1FF82FF44F9F5FED1FC55.taxon	description	Description. A complete wing, with a dark brown area covering apical third, wing 28.8 mm long, 8.4 mm wide, broad and with numerous small cells; distance from base to arculus 3.6 mm, from arculus to nodus 3.2 mm; primary antenodal crossveins Ax 1 and Ax 2 well-defined; Ax 1 2.2 mm from wing base, distance between Ax 1 and Ax 2 1.4 mm; Ax 2 just distad arculus; no secondary antenodal crossveins; ca. 34 postnodal crossveins, not well aligned with postsubnodal crossveins; pterostigma rather short but broad, 2.4 mm long and 0.8 mm wide, covering four cells; distal edge of pterostigma more oblique than proximal one; pterostigmal brace absent; one row of crossveins between C and RA distal of pterostigma; posterior part of arculus (= basal discoidal crossvein) present, therefore discoidal cell basally closed; discoidal cell rectangular, elongate, narrow (1.6 mm long, 0.4 mm wide), and free of crossveins; origins of RP and MA separated in arculus; bend of ScP at nodus very abrupt and Z-like; nodal vein and subnodus oblique; subnodus aligned with nodal crossvein; no crossveins in antesubnodal area; median space free of crossveins; submedian space free of crossveins except for CuP-crossing; subdiscoidal cells free; anal area narrow (0.3 mm wide in middle), not wider than subdiscoidal cell, with one crossvein below subdiscoidal cell; cubito-anal area max. 2.0 mm wide in its broadest part, with rows of cells; CuA well-defined and curved; CuA not forked; area between MP and CuA progressively broadened; MP regularly curved, not distally zigzagged, and ending between nodus and pterostigma; basal half of postdiscoidal area between MA and MP narrow with only one row of cells, but greatly widened distally, and with two long secondary longitudinal veins; MA ending well basad pterostigma level; MA, RP 3 / 4, and IR 2 rather straight, but distally curved towards hind margin; no antefurcal crossveins between basal parts of RP and MA; area between RP 3 / 4 and MA narrow with one row of cells between them in basal half, but area distally widened with two long secondary longitudinal veins; area between RP 3 / 4 and IR 2 progressively broadened, with two long secondary longitudinal veins; RP 3 / 4 ending beneath pterostigma; base of RP 3 / 4 1.8 mm basad subnodus; base of IR 2 below nodus; RP 1 / 2 straight at subnodus; base of RP 2 very far from subnodus, at 13 cells distal from it; IR 1 well-defined, long, and curved, originating four cells distad RP 2; no lestine oblique vein ‘ O’ between RP 2 and IR 2; RP 2 and IR 2 with only one row of cells between them until level of base of pterostigma; RP 1 without distinct bend at pterostigmal brace; wings well petiolated, petiole 2.8 mm long.	en	Nel, André (2024): Two new damselflies from the Eocene Green River Formation (Odonata, Zygoptera, Dysagrionidae, Thaumatoneuridae). Zootaxa 5446 (4): 588-594, DOI: 10.11646/zootaxa.5446.4.11, URL: http://dx.doi.org/10.11646/zootaxa.5446.4.11
7F4A8799FFF1FF82FF44F9F5FED1FC55.taxon	discussion	Remark. This damselfly wing fits quite well with those of the two eodysagrionine genera and species Tynskysagrion brookeae Bechly, Garrouste, Aase, Karr, Grande & Nel, 2021 and Oreodysagrion tenebris Bechly, Garrouste, Aase, Karr, Grande & Nel, 2021 because of the elongate and narrow discoidal cell, long petiole, nodus situated in a very basal position, distal third of wing very broad. Eodysagrion has a shorter discoidal cell, the nodus situated in a more distal position, and much less postnodal crossveins (Rust et al. 2008). The new fossil differs from Tynskysagrion in the much narrower anal area below the subdiscoidal cell, at most as broad as subdiscoidal cell, as in Oreodysagrion vs. as broad as subdiscoidal plus discoidal cells in Tynskysagrion. The new fossil also has only one crossvein below subdiscoidal cell, vs. two in Oreodysagrion and 2 – 3 in Tynskysagrion. In Oreodysagrion, the nodus is situated in a very basal position, 14 % of the distance between Ax 2 and wing apex vs. 19 % in the species of Tynskysagrion. In the new fossil, the nodus is situated at 10 % of the distance between Ax 2 and wing apex, which better fits with Oreodysagrion. But the wing of Oreodysagrion is more slender than in the new fossil and in Tynskysagrion. Thus the new fossil does not fit in these two genera. Lastly, the color pattern of the new wing is strongly different from those of Oreodysagrion tenebris and Tynskysagrion brookeae. Oreodysagrion tenebris has a dark brown band between the base of RP 2 and the basal side of pterostigma. Tynskysagrion brookeae has a completely darkened wings or with a narrow hyaline band in distal half of the wing.	en	Nel, André (2024): Two new damselflies from the Eocene Green River Formation (Odonata, Zygoptera, Dysagrionidae, Thaumatoneuridae). Zootaxa 5446 (4): 588-594, DOI: 10.11646/zootaxa.5446.4.11, URL: http://dx.doi.org/10.11646/zootaxa.5446.4.11
7F4A8799FFF3FF82FF44FB41FC43FA89.taxon	type_taxon	Type species: Petrolestes hendersoni Cockerell, 1927 (Eocene, Green River Formation, Colorado, USA). Other species. Petrolestes messelensis Garrouste & Nel, 2015 (Eocene, Messel, Germany), Petrolestes inexpectatus sp. nov. (Eocene, Green River Formation, Colorado, USA).	en	Nel, André (2024): Two new damselflies from the Eocene Green River Formation (Odonata, Zygoptera, Dysagrionidae, Thaumatoneuridae). Zootaxa 5446 (4): 588-594, DOI: 10.11646/zootaxa.5446.4.11, URL: http://dx.doi.org/10.11646/zootaxa.5446.4.11
7F4A8799FFF3FF85FF44FABDFAEEF849.taxon	description	(Figure 2) urn: lsid: zoobank. org: act: 64 CA 7115 - 25 F 1 - 4 FAD- 87 E 4 - 200896 B 07 A 63	en	Nel, André (2024): Two new damselflies from the Eocene Green River Formation (Odonata, Zygoptera, Dysagrionidae, Thaumatoneuridae). Zootaxa 5446 (4): 588-594, DOI: 10.11646/zootaxa.5446.4.11, URL: http://dx.doi.org/10.11646/zootaxa.5446.4.11
7F4A8799FFF3FF85FF44FABDFAEEF849.taxon	materials_examined	Material. Holotype IP. 359116 (part and counterpart of a nearly complete wing), stored at Invertebrate Paleontology Collection, Yale Peabody Museum, USA. Age and outcrop. Eocene (Ypresian), Green River Formation, Parachute Creek member, Gus’ pit, two miles west of Rio Blanco store, Colorado, USA.	en	Nel, André (2024): Two new damselflies from the Eocene Green River Formation (Odonata, Zygoptera, Dysagrionidae, Thaumatoneuridae). Zootaxa 5446 (4): 588-594, DOI: 10.11646/zootaxa.5446.4.11, URL: http://dx.doi.org/10.11646/zootaxa.5446.4.11
7F4A8799FFF3FF85FF44FABDFAEEF849.taxon	etymology	Etymology. Named after the unexpected presence of a second species of Petrolestes in the Green River Formation.	en	Nel, André (2024): Two new damselflies from the Eocene Green River Formation (Odonata, Zygoptera, Dysagrionidae, Thaumatoneuridae). Zootaxa 5446 (4): 588-594, DOI: 10.11646/zootaxa.5446.4.11, URL: http://dx.doi.org/10.11646/zootaxa.5446.4.11
7F4A8799FFF3FF85FF44FABDFAEEF849.taxon	diagnosis	Diagnosis. Wing characters only. Base of RP 2 four cells distad subnodus; pterostigma covering two cells; presence of very long cells in cubito-anal area.	en	Nel, André (2024): Two new damselflies from the Eocene Green River Formation (Odonata, Zygoptera, Dysagrionidae, Thaumatoneuridae). Zootaxa 5446 (4): 588-594, DOI: 10.11646/zootaxa.5446.4.11, URL: http://dx.doi.org/10.11646/zootaxa.5446.4.11
7F4A8799FFF3FF85FF44FABDFAEEF849.taxon	description	Description. Wings hyaline, distal third of wing deformed, thus its exact length is difficult to determine, wing ca. 27.8 mm long; max. width 6.8 mm, wing 6.2 mm wide at nodus level; a very short petiole, 2.3 mm long; anal area with one row of cells between AA and AP; subdiscoidal cell broad elongate, 1.7 mm long, 0.5 mm wide; distance from base to arculus 5.0 mm, from arculus to nodus 7.8 mm, from nodus to pterostigma ca. 10.0 mm; nodus in a basal position; pterostigma rather short but broad, 2.1 mm long, 6.2 mm wide, covering two cells; pterostigmal brace perpendicular to RA and RP 1; distance from wing base to Ax 1 3.4 mm, from Ax 1 to Ax 2 1.4 mm, Ax 2 nearly aligned with arculus; no supplementary antenodal crossvein; discoidal cell unicellular, 1.1 mm long, 1.1 mm wide, distinctly broadened in its distal part, with basal side 0.3 mm, anterior side 0.9 mm, posterior side 1.1 mm, distal side MAb 1.1 mm; ScP not crossing through nodus as in Stenolestes Scudder, 1895; nodal crossvein very oblique; subnodus perpendicular to RA and RP with corresponding crossvein below (between RP 1 / 2 and IR 2) of inverted obliquity; ca. 14 postnodal crossveins preserved, not aligned with 14 postsubnodal crossveins; bases of RP 3 / 4 and IR 2 between arculus and nodus, closer to nodus than to arculus, base of RP 3 / 4 4.6 mm from arculus and 3.2 mm from nodus; base of RP 2 four cells, 3.0 mm distal of subnodus; base of IR 1 two cells distally; oblique crossvein ‘ O’ absent; cubito-anal area with three-four rows of cells between CuA and posterior wing margin, some of these being very long; CuA reaching posterior wing margin well distal of nodus level; CuA weakly curved; postdiscoidal area with one row of cells; area between RP 3 / 4 and MA with two secondary longitudinal veins in between; three rows of cells between RP 3 / 4 and IR 2 and one row between RP 1 and IR 1; three rows of cells between IR 1 and RP 2 opposite pterostigma, but only one basally.	en	Nel, André (2024): Two new damselflies from the Eocene Green River Formation (Odonata, Zygoptera, Dysagrionidae, Thaumatoneuridae). Zootaxa 5446 (4): 588-594, DOI: 10.11646/zootaxa.5446.4.11, URL: http://dx.doi.org/10.11646/zootaxa.5446.4.11
7F4A8799FFF3FF85FF44FABDFAEEF849.taxon	discussion	Remark. This wing is strongly similar to those of the two species of the genus Petrolestes, especially in the shape of the petiole, discoidal cell, subnodus, cubital veins, patterns of veins between main longitudinal veins, etc. (Cockerell 1927; Garrouste & Nel 2015). It can be attributed to this genus, but differs from the two described species in the base of RP 2 situated four cells distad subnodus vs. only two, in the pterostigma covering only two cells vs. four or more, and in the presence of very long cells in the cubito-anal area. Thus I consider it as a new species.	en	Nel, André (2024): Two new damselflies from the Eocene Green River Formation (Odonata, Zygoptera, Dysagrionidae, Thaumatoneuridae). Zootaxa 5446 (4): 588-594, DOI: 10.11646/zootaxa.5446.4.11, URL: http://dx.doi.org/10.11646/zootaxa.5446.4.11
