taxonID	type	description	language	source
7F0D87D3FFA0FFF2FF7D1E1879CDDF80.taxon	type_taxon	Type species: Harpactopus crudelis F. Smith, 1856, by subsequent designation of Patton 1880: 384. Junior subjective synonym of Sphex Linnaeus, 1758 according to Kohl 1890: 106 and Prionyx Vander Linden, 1827 by Bohart & Menke 1976: 39. Resurrected to subgeneric status by Danilov & Odintsev 2023: 430.	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFA0FFF2FF7D1E1879CDDF80.taxon	diagnosis	Diagnosis. The subgenus Harpactopus F. Smith is characterized by the claws with two ventral teeth, male antennae with distinct placoids, female clypeus entire or with U-shaped median notch, metapleuron channel-like, and metasoma wholly black.	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFA0FFF2FF7D1E1879CDDF80.taxon	distribution	Distribution. Arid and semi-arid areas of the Old World and Australasia.	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFA0FFF2FF7D19667AD4DDA5.taxon	discussion	Note. The species group name crudelis is used in this paper after Bohart & Menke 1963, 1976 because Harpactopus crudelis F. Smith is the type species of Harpactopus.	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFA0FFF2FF7D19667AD4DDA5.taxon	diagnosis	Diagnosis. ♂ ♀: labial palpomeres 2 – 4 usual; arolia present; recurrent vein II of forewing received by submarginal cell III. ♂: placoids on flagellomeres narrow, depressed; S 6 entire, without teeth. ♀: clypeus with Ushaped median notch; tarsal rake of foreleg with long, stout, bladelike and some hairlike setae or only with some long, stout, bladelike setae.	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFA0FFF2FF7D1A8A7917DCE1.taxon	diagnosis	Diagnosis. ♂ ♀: labial palpomeres 2 – 4 elongate; arolia present; recurrent vein II of forewing interstitial with 1 st intersubmarginal vein or received by submarginal cell III at its basal angle. ♂: placoids on flagellomeres broad with medial line of setae; S 6 entire, without teeth. ♀: clypeus entire, slightly emarginate anteriorly; tarsal rake of foreleg with long, hairlike setae and short, stout, bladelike setae. Included species: Prionyx globosus (F. Smith, 1856).	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFA7FFF5FF7D1CEF7FB6DA8D.taxon	diagnosis	Diagnosis. ♂ ♀: labial palpomeres 2 – 4 short, not club-shaped; arolia present; recurrent vein II of forewing interstitial with 1 st intersubmarginal vein or received by submarginal cell III at its basal angle. ♂: placoids on F 3 – F 6 broad, flat; S 6 angularly convex posterolaterally, without teeth. ♀: clypeus entire, slightly emarginate anteriorly; tarsal rake of foreleg with long, stout, bladelike and some short, hairlike or bladelike setae. Included species: Prionyx chobauti (Roth, 1925), P. funebris (Berland, 1926), P. macula (Fabricius, 1804).	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFA7FFF5FF7D1DA37EB3D9C9.taxon	diagnosis	Diagnosis. ♂ ♀: labial palpus extremely shortened, inconspicuous, labial palpomeres 2 – 4 club-shaped; maxillar palpomere 6 inconspicuous; arolia absent; recurrent vein II of forewing received by submarginal cell III. ♂: placoids on F 3 – F 6 broad, flat; S 6 with pair teeth posterolaterally. ♀: clypeus entire, slightly emarginate anteriorly; tarsal rake with short, stout, bladelike setae. Included species: Prionyx persicus (Mocsáry, 1883), P. radoszkowskyi (Kohl, 1888).	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFA7FFF5FF7D1F677F10DF15.taxon	diagnosis	Diagnosis. ♂ ♀: labial palpomeres 2 – 4 elongate; arolia present; recurrent vein II of forewing interstitial with 1 st intersubmarginal vein or received by submarginal cell III at its basal angle. ♂: placoids on flagellomeres narrow, depressed; S 6 entire, without teeth. ♀: clypeus entire, slightly emarginate anteriorly; tarsal rake of foreleg with long, bladelike and long, fine, hairlike setae. Included species: Prionyx melanotus (F. Morawitz, 1890), P. stschurowskii (Radoszkowski, 1877).	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFA7FFF4FF7D181B7EE7DAA9.taxon	description	(Figs 2 b, 8 a, 9 a)	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFA7FFF4FF7D181B7EE7DAA9.taxon	diagnosis	Diagnosis. Both sexes of P. chobauti closely resemble P. macula and P. funebris in having the labial palpomeres 2 – 4 short, and the male in having F 3 – F 6 with broad placoids, S 6 angularly convex posterolaterally, but differ in having the vertex, pronotum and the scutum without dense, appressed, silvery setae (P. macula and P. funebris has the scutum with dense, appressed, silvery setae). Both sexes of P. chobauti resemble P. persicus, and P. radoszkowskyi in habitus, and the male in having F 3 – F 6 with broad placoids. They differ from P. persicus, and P. radoszkowskyi in having the labial palpomeres 2 – 4 short, conspicuous, and the arolia present (P. persicus and P. radoszkowskyi have the labial palpomeres 2 – 4 extremely short, club-shaped, and the arolia absent). Unlike P. stschurowskii and P. persicus the female of P. chobauti has the scutum with no appressed, silvery setae.	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFA7FFF4FF7D181B7EE7DAA9.taxon	description	Description. Head. Labrum trapeziform; labial palpomeres 2 – 4 short, conspicuous. Clypeus laterally and posteriorly, paraocular area anteriorly with semiappressed, brownish setae, not concealing integument; frons, vertex, occiput and gena slightly shiny, microsculptured, with black erect setae. Mesosoma. Pronotum and scutum rugose, with sparse, semiappressed, brownish setae, not concealing integument, and with black erect setae. Scutellum longitudinally depressed, punctate and rugose. Metanotum tuberculate and longitudinally depressed dorsally. Mesopleuron, metapleuron, and propodeum coarsely areolate-rugose, with black erect setae. Wings intensely dark; veins black; recurrent vein II of forewing interstitial with 1 st intersubmarginal vein or received by submarginal cell III at its basal angle. Legs. Claws with two ventral teeth; arolia present. Male. Body length 20 – 25 mm. Head. F 3 – F 6 with broad, flat placoids. Metasoma. S 6 angularly convex posterolaterally, without teeth. Female. Body length 24 – 28 mm. Head. Clypeus slightly emarginate anteriorly; clypeal disk moderately convex, with transverse oval depression anteriorly. Legs. Tarsal rake of foreleg with long and some short, stout, bladelike setae.	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFA7FFF4FF7D181B7EE7DAA9.taxon	materials_examined	Material examined. MOROCCO. Gharb-Chrarda-Béni Hssen: Kenitra [34 ° 15 ’ N, 6 ° 35 ’ W], 23. V. 1947, J. de Beaumont (1 ♀, ZISP).	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFA7FFF4FF7D181B7EE7DAA9.taxon	distribution	Distribution. Parts of North African arid and semi-arid areas (Algeria, Libya and Morocco).	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFA6FFF6FF7D1D877BBCDF31.taxon	description	(Figs 1 a, 4 a, 8 d, 9 b, 11 c)	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFA6FFF6FF7D1D877BBCDF31.taxon	diagnosis	Diagnosis. Both sexes of P. crudelis closely resemble P. subfuscatus, P. saevus, P. judaeus, and P. indus. The female of P. crudelis differs from P. subfuscatus and P. saevus in having the head and mesosoma with dark setae, the propodeal enclosure with long, dense, hairlike black setae, the tarsal rake of the foreleg with long, stout, bladelike and some hairlike setae (P. subfuscatus and P. saevus have the head and mesosoma with pale setae, propodeal enclosure without long, dense, black setae, tarsal rake of foreleg only with some long, stout, bladelike setae). The male of P. crudelis differs from P. subfuscatus and P. saevus in larger size: body length more than 18 mm, and in having the propodeal enclosure with long, dense, hairlike black setae (body length up to 18 mm in P. subfuscatus and P. saevus, and the propodeal enclosure without long, dense, black setae). Unlike P. indus, both sexes of P. crudelis have the wings yellow (the wings dark in P. indus). Both sexes of P. crudelis differ from P. judaeus in having the mesopleuron ventrally rugose (the mesopleuron ventrally smooth and shiny with sparse punctures in P. judaeus). The male of P. crudelis differs from P. judaeus in having F 3 – F 5 with placoids (F 3 – F 4 with placoids in P. judaeus).	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFA6FFF6FF7D1D877BBCDF31.taxon	description	Description. Body black with black erect setae. Head. Labial palpomeres 2 – 4 usual. Vertex, occiput and gena microsculptured, punctate. Mesosoma. Pronotum moderately angulate laterodorsally, finely rugose, with black erect setae. Scutum punctatorugose, slightly shiny. Scutellum slightly longitudinally depressed, longitudinlly rugose. Metanotum distinctly tuberculate dorsally. Mesopleuron, metapleuron, and propodeum coarsely areolate-rugose, with black erect setae, not concealing integument, except propodeal enclosure with long, dense, hairlike black setae, moderately concealing integument. Wings yellow in venation zone, smoky apically; veins light brown to yellow; recurrent vein II of forewing received by submarginal cell III. Legs. Claws with 2 ventral teeth; arolia present. Male. Body length 22 – 33 mm. Head. Labrum trapeziform. Clypeus with wide V-shaped median emargination. Clypeus, subantennal sclerite and paraocular area with appressed, silvery setae, moderately concealing integument. Mandible black and dark brown mesally. F 3 – F 5 with narrow, depressed placoids. Metasoma. S 6 entire, without teeth. Female. Body length 24 – 40 mm. Head. Labrum trapeziform, emarginate apically. Clypeus angulate, with Ushaped median notch; anterior margin of clypeal disk slightly beak-like hangs over notch. Clypeus, subantennal sclerite and paraocular area with appressed, silvery setae, concealing integument. Mesosoma. Pronotum with sparse appressed, silvery setae, not concealing integument. Legs. Tarsal rake of foreleg with long, stout, bladelike and some short, hairlike setae.	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFA6FFF6FF7D1D877BBCDF31.taxon	materials_examined	Material examined. AFGHANISTAN. Uruzgan Province: Gizab [33 ° 22 ’ N, 66 ° 15 ’ E], 8 – 15. VI. 1970, O. Kabakov (2 ♀, ZISP). IRAN. Sistan and Baluchestan Province: Mirjaveh County, Siah Jangal [28 ° 47 ’ N, 61 ° 14 ’ E], 11 – 12. V. 1901, N. Zarudny (1 ♀, ZISP); Iranshahr County, Bazman District, Bazman [27 ° 51 ’ N, 60 ° 10 ’ E], 3 – 5. VIII. 1898, N. Zarudny (2 ♂, ZISP); Zahedan County, Harmak [29 ° 59 ’ N, 60 ° 51 ’ E], 11. VI. 1898, N. Zarudny (2 ♂, ZISP); Zahedan [formerly Duzzap or Duzdab] [29 ° 30 ’ N, 60 ° 51 ’ E], 28. V. 1955, D. Steinberg (2 ♂, 1 ♀, ZISP); 14 – 18. V. 1901, N. Zarudny (1 ♂, ZISP); South Khorasan Province: Darmian County, Gazik [33 ° 00 ’ N, 60 ° 13 ’ E], 12 – 13. VII. 1901, N. Zarudny (1 ♂, 1 ♀, ZISP); TAJIKISTAN. Khatlon Province: Ayvadj [36 ° 59 ’ N, 68 ° 2 ’ E], 3. VIII. 1934, V. Gussakovskij (1 ♀, ZISP); Qabodiyon [37 ° 24 ’ N, 68 ° 11 ’ E], 27. VI. 1934, V. Gussakovskij (1 ♀, ZISP); Bokhtar [formerly Qurghonteppa or Kurgan-Tyube] [37 ° 49 ’ N, 68 ° 46 ’ E], 31. V. 1931, N. Fursov (1 ♀, ZISP); Dusti District, Jilikul [37 ° 29 ’ N, 68 ° 31 ’ E], 12. VI. 1934, V. Gussakovskij (1 ♂, ZISP); 30. IX. 1931, N. Fursov (1 ♂, ZISP); Panj District, Panj [37 ° 14 ’ N, 69 ° 06 ’ E], 16 – 17. VII. 1943, D. Stackelberg (2 ♂, ZISP); TURKMENISTAN. Ashgabat [37 ° 57 ’ N, 58 ° 23 ’ E], 21. V. 1916, collector unknown (1 ♂, ZISP); 7. VII. 1915, collector unknown (1 ♂, ZISP); 28. VI. 1928, V. Gussakovskij (1 ♂, ZISP); Ahal Province: Tejen [= Tedzhen] [37 ° 22 ’ N, 60 ° 30 ’ E], 4. VII. 1929, Predtetschensky (1 ♀, ZISP); Komarovsky near Ashgabat [37 ° 45 ’ N, 58 ° 33 ’ E], 22. V. 1928, V. Gussakovskij (1 ♂, ZISP); Balkan Province: Eldere gorge near Magtymguly [formerly Kara-Kala] [38 ° 30 ’ N, 56 ° 23 ’ E], 29. VI. 1953, E. Arens (1 ♀, ZISP); Sumbar river valley near Magtymguly [formerly Kara-Kala] [38 ° 25 ’ N, 56 ° 24 ’ E], 18. VI. 1955, A. Ponomareva (1 ♀, ZISP); Garabogaz near Gyzylarbat [formerly Serdar or Kyzyl-Arvat] [39 ° 17 ’ N, 56 ° 27 ’ E], 31. V. 1953, D. Steinberg (1 ♂, ZISP); Lebap Province: Repetek Biosphere State Reserve [38 ° 34 ’ N, 63 ° 10 ’ E], 14. V. 1914, K. Bil’kewitch (1 ♀, ZISP); Çärjew District, Farap [= Farab] [39 ° 10 ’ N, 63 ° 36 ’ E], 16 – 20. V. 1911, Kuleshov (1 ♂, ZISP); Mary Province: Imeni Chapayeva [formely Ymam Baba] [36 ° 42 ’ N, 62 ° 27 ’ E], 18 – 20. VI. 1925, V. Gussakovskij (1 ♂, 3 ♀, ZISP); 28 – 31. V. 1912, V. Kozhanchikov (3 ♂, 3 ♀, ZISP); 8. VI. 1912, V. Kozhanchikov (1 ♂, ZISP); 25. V. 1912, V. Kozhanchikov (1 ♂, ZISP); 1932, A. Shestakov (2 ♂, ZISP); Baýramaly [37 ° 37 ’ N, 62 ° 09 ’ E], N. Simonov (1 ♀, ZISP); 17. VI. 1910, B. Uvarov (1 ♀, ZISP); 11. IX. 1916, M. Ryabov (1 ♀, ZISP); 11. V. 1917, M. Ryabov (2 ♂, ZISP); 21. V. 1917, collector unknown (1 ♀, ZISP); Serhetabat [formerly Gushgy or Kushka] [35 ° 17 ’ N, 62 ° 21 ’ E], 13 – 18. VI. 1925, V. Gussakovskij (1 ♂, 1 ♀, ZISP); Sary-Yazi [36 ° 27 ’ N, 62 ° 37 ’ E], V. 1904, K. Arris (1 ♀, ZISP); Ýolöten [37 ° 18 ’ N, 62 ° 21 ’ E], 18. VI. 1927, V. Kizeritskij (1 ♀, ZISP); 1. X. 1926, V. Kizeritskij (1 ♂, ZISP); 12. VI. 1898, K. Angher (1 ♂, ZISP); UZBEKISTAN. Jizzakh Province: Jizzakh [40 ° 07 ’ N, 67 ° 51 ’ E], VI. 1919, Chernevsky expedition (1 ♀, ZISP); Khorazm Province: Khiva [41 ° 22 ’ N, 60 ° 21 ’ E], 25. VI. 1927, V. Gussakovskij (1 ♂, ZISP); YEMEN. Sanaa [15 ° 20 ’ N, 44 ° 12 ’ E], 16 – 18. VIII. 1930, collector unknown (1 ♂, 2 ♀, ZISP).	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFA6FFF6FF7D1D877BBCDF31.taxon	distribution	Distribution. Arid and semi-arid to tropical wet and dry areas from Central and South Asia to Southern Balkans and from Middle East to East and Equatorial Africa and Western Indian Ocean Islands (* Afghanistan, Bulgaria, Comoros, Cyprus, Democratic Republic of the Congo, Egypt, Eritrea, Ethiopia, Greece, Guinea, Guinea-Bissau, India, Iran, Iraq, Israel, Kazakhstan, Kenya, Kuwait, Madagascar, Mauritania, Mauritius, Oman, Saudi Arabia, Seychelles, Somalia, South Sudan, Sudan, Syria, Tajikistan, Tanzania, Turkey, Turkmenistan, United Arab Emirates, * Uzbekistan, Yemen).	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFA4FFF9FF7D18FF78E0DA8D.taxon	description	(Figs 2 c, 4 d, 5 d, 6 d, 7 c, 8 b, 9 c, 12 a)	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFA4FFF9FF7D18FF78E0DA8D.taxon	diagnosis	Diagnosis. Both sexes of P. funebris closely resemble P. macula in having the scutum with dense, appressed, silvery setae, concealing the integument, in having the labial palpomeres 2 – 4 short, and the male in having F 3 – F 6 with broad placoids, and S 6 angularly convex posterolaterally, but differ in having the vertex and the pronotum without dense, appressed, silvery setae and in having the wings intensely dark with purple reflexes (P. macula has the vertex and pronotum with dense, appressed, silvery setae, and the wings dark to smoky).	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFA4FFF9FF7D18FF78E0DA8D.taxon	description	Description. Head. Labrum trapeziform; labial palpomeres 2 – 4 short, conspicuous. Clypeus laterally and posteriorly, paraocular area anteriorly with appressed, silvery setae; frons with transverse band of dense appressed, silvery setae; vertex, occiput and gena slightly shiny, microsculptured, with black erect setae. Mesosoma. Pronotum moderately angulate laterodorsally, transversely rugose laterally and laterodorsally, finely rugose dorsally, with black erect setae. Scutum with dense, appressed, silvery setae, concealing integument, except anterolateral angles and narrow area anteriorly distinctly transversely rugose. Scutellum slightly longitudinally depressed, punctate and rugose. Metanotum tuberculate dorsally. Mesopleuron, metapleuron, and propodeum coarsely areolate-rugose with black erect setae. Wings intensely dark with purple reflexes; veins black; recurrent vein II of forewing interstitial with 1 st intersubmarginal vein or received by submarginal cell III at its basal angle. Legs. Claws with two ventral teeth; arolia present. Male. Body length 25 mm. Head. Clypeus with wide V-shaped median emargination, glabrous and shiny anteriorly. F 3 – F 6 with broad, flat placoids. Metasoma. S 6 angularly convex posterolaterally, without teeth. Female. Body length 29 mm. Head. Clypeus slightly emarginate anteriorly; clypeal disk moderately convex, with transverse oval depression anteriorly. Legs. Tarsal rake of foreleg with long, stout, bladelike and some short, hairlike setae.	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFA4FFF9FF7D18FF78E0DA8D.taxon	materials_examined	Material examined. KENYA. Coast Province: Tsavo River, no specific locality [3 ° 00 ’ S, 38 ° 26 ’ E], 27. XII. 1911, Svatosh (1 ♀, ZISP). YEMEN. Sanaa [15 ° 20 ’ N, 44 ° 12 ’ E], 30. VIII. 1914, collector unknown (1 ♂, ZISP).	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFA4FFF9FF7D18FF78E0DA8D.taxon	distribution	Distribution. Southern and eastern African and southern Arabian semi-arid and arid areas of Afrotropical region (Ethiopia, Kenya, Namibia, South Africa, * Yemen).	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFABFFF8FF7D194479AADE09.taxon	description	(Figs 2 a, 4 f, 5 i, 6 i, 9 d)	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFABFFF8FF7D194479AADE09.taxon	diagnosis	Diagnosis. Both sexes of P. globosus closely resemble P. saevus, P. subfuscatus, P. reymondi, and P. zarudnyi. The female of P. globosus differs in having the clypeus entire, slightly emarginate anteriorly (P. saevus, P. subfuscatus, P. reymondi, and P. zarudnyi have the clypeus with U-shaped median notch). The male of P. globosus differs in having the placoids on flagellomeres broad with medial line of setae (the placoids on flagellomeres narrow, depressed in P. saevus, P. subfuscatus, P. reymondi, and P. zarudnyi).	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFABFFF8FF7D194479AADE09.taxon	description	Description. Body black with pale erect setae. Head. Labial palpomeres 2 – 4 elongate. Palpi, labrum, mandible in basal ⅔, and often clypeus anteriorly dark reddish. Clypeus entire, slightly emarginate anteriorly with appressed, silvery setae, distinctly concealing integument, and with glabrous triangle area medially. Subantennal sclerite and paraocular area with appressed, silvery setae, distinctly concealing integument. Gena with appressed, silvery setae, moderately concealing integument. Mesosoma. Pronotum with appressed, silvery setae, moderately concealing integument dorsally. Scutum punctatorugose, slightly shiny with appressed, silvery setae laterally, slightly concealing integument. Scutellum shiny. Mesopleuron, metapleuron, and propodeum finely rugose with patchs of appressed, silvery setae, slightly concealing integument. Wings smoky; veins light brown; recurrent vein II of forewing interstitial with 1 st intersubmarginal vein or received by submarginal cell III at its basal angle. Legs. Claws with 2 ventral teeth; arolia present. Male. Body length 10 – 12 mm. Head. Placoids on F 3 – F 6 broad with medial line of setae. Metasoma. S 6 entire, without teeth. Female. Body length 11 – 22 mm. Legs. Tarsal rake of foreleg with long, hairlike setae and short, stout, bladelike setae. Justification of synonymy. Harpactopus australis was described by de Saussure (1867) from materials collected during the voyage of the frigate Novara along the coast of Australia (Nova Hollandia). Kohl (1890), discussing this taxon, noted the similarity with P. globosus, but concluded that a number of characters prevented him from synonymy, although he did not completely reject it. R. Turner (1910) disagreed with Kohl’s opinion that these taxa are different and placed australis in synonymy of globosus. The research of these authors was based on material from NHMW or conspecific. Having studied the material in MHNG in 1971, van der Vecht (1973) came to the conclusion that Harpactopus australis is a Palmodes, probably P. occitanicus, as he indicated on the label of the specimen. In addition, he is indicates that the type material is stored in MHNG, not in NHMW. A little later, Bohart and Menke (1976) listed australis as a possible subspecies of Palmodes occitanicus. Danilov and Byvaltsev (2020) raised the question of the status of this taxon again, and later australis was placed in the subgenus Harpactopus of Prionyx (Danilov & Odintsev 2023). The specimen from MHNG, designated as the holotype is a male of the Palaearctic species Palmodes occitanicus. The characters of the taxon in de Saussure’s description, based on a female, match those of the typelabeled specimen in NHMW. At the same time, the drawing in the appendix to de Saussure’s description shows a male with the bicolor metasoma, which, as Kohl noted, was due to the draftsman’s error. In any case, the original description has an undoubted advantage over the drawing. The designation of the holotype in MHNG was made after de Saussure’s publication and is therefore invalid according to Article 73.1.3 of the Code (ICZN 1999), which states that the holotype can only be fixed in the original publication and by the original author.	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFABFFF8FF7D194479AADE09.taxon	materials_examined	Material examined. AUSTRALIA. Queensland: Brisbane [27 ° 28 ’ S, 153 ° 01 ’ E], 30. X. 1938, C. F. Ashby (1 ♂, ZISP).	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFABFFF8FF7D194479AADE09.taxon	distribution	Distribution. Semi-arid areas of Australasian region (Australia).	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFAAFFFBFF7D1927798CD83D.taxon	description	(Figs 1 b, 4 a, 9 e)	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFAAFFFBFF7D1927798CD83D.taxon	diagnosis	Diagnosis. Both sexes of P. indus closely resemble P. crudelis and P. judaeus, but differ in having the wings dark (the wings are yellow in P. crudelis and P. judaeus).	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFAAFFFBFF7D1927798CD83D.taxon	description	Description. Body black with black erect setae. Head. Labial palpomeres 2 – 4 usual. Vertex, occiput and gena microsculptured, punctate. Mesosoma. Pronotum moderately angulate laterodorsally, finely rugose, with black erect setae. Scutum punctatorugose, slightly shiny. Scutellum slightly longitudinally depressed, longitudinlly rugose. Metanotum distinctly tuberculate dorsally. Mesopleuron, metapleuron, and propodeum coarsely areolate-rugose with black erect setae, not concealing integument, except propodeal enclosure with long, dense, hairlike black setae, moderately concealing integument. Wings dark in venation zone, smoky apically; veins dark brown; recurrent vein II of forewing received by submarginal cell III. Legs. Claws with 2 ventral teeth; arolia present. Male. Body length 21 – 32 mm. Head. Labrum trapeziform. Clypeus with wide V-shaped median emargination. Clypeus, subantennal sclerite and paraocular area with appressed, silvery setae, moderately concealing integument. Mandible black. F 3 – F 5 with narrow, depressed placoids. Metasoma. S 6 entire, without teeth. Female. Body length 28 – 35 mm. Head. Labrum trapeziform, emarginate apically. Clypeus with U-shaped median notch. Clypeus, subantennal sclerite and paraocular area with appressed, silvery setae, concealing integument. Mesosoma. Pronotum with sparse appressed, silvery setae, not concealing integument. Legs. Tarsal rake of foreleg with long, stout, bladelike and some short, hairlike setae.	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFAAFFFBFF7D1927798CD83D.taxon	materials_examined	Material examined. None.	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFAAFFFBFF7D1927798CD83D.taxon	distribution	Distribution. Arid and semi-arid areas of Southern Africa (Angola, Botswana, Democratic Republic of the Congo, Malawi, Namibia, South Africa, Tanzania, Zambia, Zimbabwe).	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFA9FFFAFF7D1B687E4CDFA5.taxon	description	(Figs 1 c, 4 b, 9 f)	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFA9FFFAFF7D1B687E4CDFA5.taxon	diagnosis	Diagnosis. Both sexes of P. judaeus closely resemble P. subfuscatus, P. saevus, P. crudelis, and P. indus. The female of P. judaeus differs from P. subfuscatus and P. saevus in having the head and mesosoma with dark setae, the propodeal enclosure with long, dense, hairlike black setae, the tarsal rake of the foreleg with long, stout, bladelike and some hairlike setae (P. subfuscatus and P. saevus have the head and mesosoma with pale setae, propodeal enclosure without long, dense, black setae, tarsal rake of foreleg only with some long, stout, bladelike setae). The male of P. judaeus differs from P. subfuscatus and P. saevus in having the propodeal enclosure with long, dense, hairlike black setae (the propodeal enclosure without long, dense, black setae in P. subfuscatus and P. saevus). Unlike P. indus, both sexes of P. judaeus have the wings yellow (the wings dark in P. indus). Both sexes of P. judaeus differ from P. crudelis in having the mesopleuron ventrally smooth and shiny with sparse punctures (the mesopleuron is ventrally rugose in P. crudelis). The male of P. judaeus differs from P. crudelis in having F 3 – F 4 with placoids (F 3 – F 5 with placoids in P. crudelis).	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFA9FFFAFF7D1B687E4CDFA5.taxon	description	Description. Body black with black erect setae. Head. Labial palpomeres 2 – 4 usual. Vertex, occiput and gena microsculptured, punctate. Mesosoma. Pronotum moderately angulate laterodorsally, finely rugose, with black erect setae. Scutum punctatorugose, slightly shiny. Scutellum slightly longitudinally depressed, longitudinlly rugose. Metanotum distinctly tuberculate dorsally. Mesopleuron laterally, metapleuron, and propodeum coarsely areolate-rugose, with black erect setae. Mesopleuron ventrally smooth and shiny, with sparse punctures. Metapleuron and propodeum coarsely areolate-rugose with black erect setae, not concealing integument, except propodeal enclosure with long, dense, hairlike black setae, moderately concealing integument. Wings yellow in venation zone, smoky apically; veins light brown to yellow; recurrent vein II of forewing received by submarginal cell III. Legs. Claws with 2 ventral teeth; arolia present. Male. Body length 18 – 21 mm. Head. Labrum trapeziform. Clypeus with wide V-shaped median emargination. Clypeus, subantennal sclerite and paraocular area with appressed, silvery setae, moderately concealing integument. Mandible black and dark brown mesally. F 3 – F 4 with narrow, depressed placoids. Metasoma. S 6 entire, without teeth. Female. Body length 24 mm. Head. Labrum trapeziform, emarginate apically. Clypeus angulate, with Ushaped median notch. Clypeus, subantennal sclerite and paraocular area with appressed, silvery setae, concealing integument. Legs. Tarsal rake of foreleg with long, stout, bladelike and some short, hairlike setae.	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFA9FFFAFF7D1B687E4CDFA5.taxon	materials_examined	Material examined. PALESTINE. Jericho [31 ° 51 ’ N, 35 ° 27 ’ E], 27. VIII. 1941, H. Bytinski-Salz (♀, holotype, MCZL). ISRAEL. Ein Gedi [31 ° 27 ’ N, 35 ° 23 ’ E], 2. VI. 1965, W. Schlaefle (1 ♂, paratype, MCZL)	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFA9FFFAFF7D1B687E4CDFA5.taxon	distribution	Distribution. Arid and semi-arid areas of the Middle East (Egypt, Israel, Palestine).	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFA8FFFDFF7D188B7920DE8E.taxon	description	(Figs 2 d, 4 d, 5 c, 6 c, 7 d, 8 c, 9 g, 12 b)	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFA8FFFDFF7D188B7920DE8E.taxon	diagnosis	Diagnosis. Both sexes of P. macula closely resemble P. funebris in having the scutum with dense, appressed, silvery setae, concealing the integument, in having the labial palpomeres 2 – 4 short, and the male in having F 3 – F 6 with broad placoids, and S 6 angularly convex posterolaterally, but differ in having the vertex and pronotum with dense, appressed, silvery setae, and the wings dark to smoky (P. funebris has the vertex and the pronotum without dense, appressed, silvery setae and in having the wings intensely dark with purple reflexes).	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFA8FFFDFF7D188B7920DE8E.taxon	description	Description. Head. Labrum trapeziform; labial palpomeres 2 – 4 short, conspicuous. Mandible in basal ⅔ dark reddish. Clypeus except apical depression, paraocular area, frons and vertex with appressed, silvery setae; gena shiny, with black erect setae. Mesosoma. Pronotum moderately angulate laterodorsally with dense, appressed, silvery setae dorsally. Scutum with dense, appressed, silvery setae, concealing integument. Scutellum slightly longitudinally depressed, punctate and rugose. Metanotum tuberculate dorsally. Mesopleuron, metapleuron, and propodeum coarsely areolate-rugose. Wings dark to smoky; veins black; recurrent vein II of forewing interstitial with 1 st intersubmarginal vein or received by submarginal cell III at its basal angle. Legs. Claws with two ventral teeth; arolia present. Male. Body length 19 – 24 mm. Head. Clypeus with wide V-shaped median emargination, glabrous and shiny anteriorly. F 3 – F 6 with broad, flat placoids. Metasoma. S 6 angularly convex posterolaterally, without teeth. Female. Body length 26 – 36 mm. Head. Clypeus slightly emarginate anteriorly; clypeal disk moderately convex, with transverse oval depression anteriorly. Legs. Tarsal rake of foreleg with long, stout, bladelike and some short, hairlike setae.	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFA8FFFDFF7D188B7920DE8E.taxon	materials_examined	Material examined. AZERBAIJAN. Baku [40 ° 27 ’ N, 49 ° 48 ’ E], 1911, collector unknown (1 ♀, ZISP); 7. VII. 1960, L. Rzaeva (1 ♀, ZISP); 23. VII. 1988, S. Storozhenko (1 ♀, SZMN); Nakhchivan [39 ° 12 ’ N, 45 ° 24 ’ E], collector unknown (1 ♀, ZISP); Ganja [formerly Yelisavetpol] [40 ° 40 ’ N, 46 ° 21 ’ E], 11 – 28. VI. [year?], I. Babadjanidi (2 ♀, ZISP). IRAN. Ardabil Province: Parsabad County, Owltan [39 ° 36 ’ N, 47 ° 46 ’ E], 30. VI. 1927, Zapolsky (1 ♀, ZISP); 21. VI. 1927, Bocharnikov (1 ♂, ZISP); Sistan and Baluchestan Province: Khash County, Karvandar [27 ° 50 ’ N, 60 ° 46 ’ E], 25. IV. 1901, N. Zarudny (1 ♀, ZISP); Nik Shahr County, Shisheh Posht [26 ° 49 ’ N, 60 ° 23 ’ E], 8 – 9. IV. 1901, N. Zarudny (1 ♂, ZISP). KAZAKHSTAN. Kyzylorda Province: Aral District, Sapak [46 ° 38 ’ N, 61 ° 46 ’ E], 19. VI. 1927, collector unknown (1 ♀, ZISP); Shieli District, near Baigekum [Djulek] [44 ° 17 ’ N, 66 ° 24 ’ E], 25. VI. 1915, E. Pavlovsky (5 ♂, ZISP). TAJIKISTAN. Khatlon Province: Qabodiyon [37 ° 24 ’ N, 68 ° 11 ’ E], 3. VII. 1934, V. Gussakovskij (1 ♀, ZISP); Dusti District, Jilikul [37 ° 29 ’ N, 68 ° 31 ’ E], 13. VI. 1934, V. Gussakovskij (1 ♀, ZISP); Dusti District, Jilikul [37 ° 29 ’ N, 68 ° 31 ’ E], 25. V. 1931, N. Fursov (1 ♂, 1 ♀, ZISP). TURKEY. Kars Province, no specific locality [40 ° 30 ’ N, 43 ° 10 ’ E], VIII. 1911, K. Satunin (1 ♀, ZISP). TURKMENISTAN. Ashgabat [37 ° 57 ’ N, 58 ° 23 ’ E], 6. VI. 1924, collector unknown (1 ♀, ZISP); Ahal Province: Komarovsky near Ashgabat [37 ° 45 ’ N, 58 ° 33 ’ E], 25. VI. 1928, V. Gussakovskij (1 ♂, ZISP); Mary Province: Imeni Chapayeva [formely Ymam Baba] [36 ° 42 ’ N, 62 ° 27 ’ E], Kuzjakin (1 ♂, ZISP); 1932, A. Shestakov (1 ♂, ZISP); 5. VI. 1931, Frolova (2 ♂, ZISP); 18 – 20. VI. 1925, V. Gussakovskij (1 ♂, ZISP); Sary-Yazi [36 ° 27 ’ N, 62 ° 37 ’ E], V. 1913, K. Bil’kewitch (1 ♂, ZISP). UZBEKISTAN. Kyzylkum Desert, no specific locality [42 ° 30 ’ N, 63 ° 30 ’ E], 1912, collector unknown (1 ♀, ZISP); Qashqadaryo Province: Qamashi District, Qamashi [38 ° 49 ’ N, 66 ° 27 ’ E], 29. VI. 1931, A. Rodd (1 ♀, ZISP).	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFA8FFFDFF7D188B7920DE8E.taxon	distribution	Distribution. Arid and semi-arid areas from Central Asia to North Africa (Afghanistan, Algeria, Armenia, Azerbaijan, Egypt, Iran, Iraq, Israel, Kazakhstan, Kuwait, Libya, Palestine, Saudi Arabia, Tajikistan, Tunisia, * Turkey, Turkmenistan, Uzbekistan). Distribution note. The specimens from Kenya (Makindu) listed as Sphex eatoni by R. Turner 1912: 413, belongs to Prionyx funebris Berland according to his description.	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFAFFFFCFF7D19A37EA6DEC5.taxon	description	(Figs 3 c, 4 e, 5 g, 6 g, 8 e, 9 h, 12 c)	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFAFFFFCFF7D19A37EA6DEC5.taxon	diagnosis	Diagnosis. The male of P. melanotus closely resembles P. stschurowskii in having the labial palpomeres 2 – 4 elongate, the placoids on flagellomeres narrow, depressed, and S 6 entire, without teeth, but differs in having F 3 – F 8 with the placoids (P. stschurowskii has F 3 – F 6 with the placoids). It is similar to P. persicus and P. radoszkowskyi in habitus, but differs in having the labial palpomeres 2 – 4 elongate, the placoids on flagellomeres narrow, depressed, S 6 entire, without teeth, and the arolia present (P. persicus and P. radoszkowskyi have the labial palpomeres 2 – 4 extremely short, club-shaped, the placoids on flagellomeres broad, flat, S 6 with pair teeth posterolaterally, and the arolia absent). The female of P. melanotus resembles P. stschurowskii, P. persicus, and P. radoszkowskyi in habitus. It differs from P. persicus, and P. radoszkowskyi in having the labial palpomeres 2 – 4 elongate, and the arolia present (P. persicus and P. radoszkowskyi have the labial palpomeres 2 – 4 extremely short, club-shaped, and the arolia absent). Unlike P. stschurowskii and P. persicus it has the scutum with no appressed, silvery setae.	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFAFFFFCFF7D19A37EA6DEC5.taxon	description	Description. Head with black erect setae. Labrum trapeziform, slightly emarginate apically; labial palpomeres 2 – 4 elongate; head without appressed, silvery setae; frons microsculptured, punctate, slightly shiny; vertex, occiput and gena punctate, shiny. Mesosoma with black erect setae. Pronotum finely rugose and punctate, slightly shiny. Scutum shiny, punctate (punctures about 0.5 – 1.5 diameters apart). Scutellum slightly longitudinally depressed, punctate and rugose, shiny. Metanotum tuberculate dorsally. Metapleuron and propodeum areolate-rugose and punctate, slightly shiny. Wings dark; veins black; recurrent vein II of forewing interstitial with 1 st intersubmarginal vein or received by submarginal cell III at its basal angle. Legs. Claws with 2 ventral teeth; arolia present. Male. Body length 15 – 19 mm. Head. Clypeus slightly emarginate anteriorly. F 3 – F 8 or F 3 – F 7 with narrow, depressed placoids. Mesosoma. Mesopleuron shiny, punctate (punctures about 0.5 – 1.5 diameters apart). Metasoma. S 6 entire, without teeth. Female. Body length 16 – 23 mm. Head. Anterior margin of clypeus rounded; clypeal disk moderately convex, slightly depressed anteriorly. Legs. Tarsal rake of foreleg with long, bladelike and long, fine, hairlike setae.	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFAFFFFCFF7D19A37EA6DEC5.taxon	materials_examined	Material examined. KAZAKHSTAN. Jambyl Province: Bayzak District, Sarykemer [formely Michailovka] near Taraz [43 ° 00 ’ N, 71 ° 30 ’ E], VI. 1912 (1 ♂, 1 ♀, ZISP); Kyzylorda Province: Kazaly District, Kazaly [formely Kazalinsk] [45 ° 46 ’ N, 62 ° 06 ’ E], 07. VI. 1928, Rukavishnikov (1 ♂, ZISP); Shieli District, Balamurun Mount near Shieli [44 ° 23 ’ N, 66 ° 50 ’ E], 20. V. 1913, Koshantschikow (1 ♀, ZISP); 21. V. 1913 (1 ♀, ZISP); 23. V. 1913 (2 ♀, ZISP); 30. V. 1913 (1 ♂, ZISP). TURKMENISTAN. no specific locality, Komarov (1 ♂, paralectotype, 1 ♀, lectotype, ZISP); Ahal Province: Kirpili, 100 km NE Serdar [formerly Kyzyl-Arvat] [39 ° 35 ’ N, 57 ° 13 ’ E], 07. V. 1953, E. Arens (1 ♂, ZISP); 10. V. 1953 (3 ♂, 1 ♀, ZISP); 11. V. 1953 (4 ♂, ZISP); 15. V. 1953 (4 ♂, 1 ♀, ZISP); 11 – 12. V. 1953, Kirjakova (1 ♂, ZISP); Tejen [37 ° 23 ’ N, 60 ° 30 ’ E], 20. IV. 1893, Glasunov (1 ♂, ZISP); Balkan Province: Garabogaz, 40 km N Serdar [formerly Kyzyl-Arvat] [39 ° 13 ’ N, 56 ° 23 ’ E], 17. V. 1953, D. Steinberg (3 ♀, ZISP); 18. V. 1953 (2 ♀, ZISP); 18. V. 1953, Odintsova (1 ♂, 1 ♀, ZISP); 19. V. 1953 (2 ♂, 1 ♀, ZISP); Jebel [39 ° 38 ’ N, 54 ° 13 ’ E], 03 – 05. VI. 1934, V. Popov (2 ♂, ZISP); Mollagara near Jebel [39 ° 36 ’ N, 54 ° 10 ’ E], 06. VI. 1934, V. Popov (1 ♀, ZISP); Daşoguz Province: Ortakui well, 240 km SW Daşoguz [40 ° 41 ’ N, 57 ° 29 ’ E], 29. V. 1953, Kirjakova (1 ♀, ZISP); 26. V. 1953, E. Arens (1 ♂, ZISP); 22. V. 1953 (1 ♂, ZISP); 29. V. 1953 (2 ♂, ZISP); 19. V. 1953 (2 ♂, ZISP); ruins of Schahsenem, 100 km SWW Daşoguz [41 ° 34 ’ N, 58 ° 42 ’ E], 30. V. 1953, E. Arens (1 ♀, ZISP); Lebap Province: Repetek Biosphere State Reserve [38 ° 34 ’ N, 63 ° 10 ’ E], 23. V. 1913, N. Androssov (1 ♀, ZISP); Mary Province: Imeni Chapayeva [formely Ymam Baba] [36 ° 42 ’ N, 62 ° 27 ’ E], 05. VI. 1931, Frolova (1 ♂, ZISP); 03 – 08. V. 1912, Koshantschikow (2 ♀, ZISP); 12 – 14. V. 1912 (1 ♀, ZISP); Utsch-Adshi [38 ° 05 ’ N, 62 ° 47 ’ E], 30. IV. 1929, A. Shestakov (1 ♂, 5 ♀, ZISP); 01 – 03. V. 1929 (3 ♂, 10 ♀, ZISP); 01. V. 1929 (2 ♂, 2 ♀, ZISP); 10. V. 1932, E. Kuznetsova (1 ♀, ZISP); 12. V. 1932 (1 ♀, ZISP); 11. VI. 1921 (1 ♀, ZISP). UZBEKISTAN. Karakalpakstan: Qo’ng’irot District, Qiziljar [43 ° 08 ’ N, 59 ° 07 ’ E], 02. VI. 1939, Veltischev (1 ♀, ZISP).	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFAFFFFCFF7D19A37EA6DEC5.taxon	distribution	Distribution. Desert areas of Central Asia (* Kazakhstan, Turkmenistan, * Uzbekistan).	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFAEFFFFFF7D1A6B7EB7DEE1.taxon	description	(Figs 3 a, 4 d, 5 a, 6 a, 7 a, 8 f, 10 a, 11 a)	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFAEFFFFFF7D1A6B7EB7DEE1.taxon	diagnosis	Diagnosis. Both sexes of P. persicus closely resemble P. radoszkowskyi in having the labial palpus extremely shortened, inconspicuous, with the club-shaped labial palpomeres 2 – 4, the arolia absent, the male F 3 – F 6 with broad placoids, and the female tarsal rake with short, stout, bladelike setae, but differ in having the labial palpomere 1 elongate: 3 – 4 × as long as the combined length of club-shaped labial palpomeres 2 – 4 (P. radoszkowskyi has the labial palpomere 1 short: 1 – 1.5 × as long as the combined length of club-shaped labial palpomeres 2 – 4). The male of P. persicus differs from P. radoszkowskyi in having S 6 with distinctly prominent teeth posterolaterally (P. radoszkowskyi has S 6 with slightly prominent teeth). It is similar to P. melanotus and P. stschurowskii in habitus, but differs in having labial palpomeres 2 – 4 extremely short, the placoids on flagellomeres broad, flat, S 6 with pair teeth posterolaterally, and the arolia absent (P. melanotus and P. stschurowskii have labial palpomeres 2 – 4 elongate, the placoids on flagellomeres narrow, depressed, and S 6 entire, without teeth). The female of P. persicus differs from P. radoszkowskyi in having the scutum with appressed, silvery setae and thick, black, erect setae (P. radoszkowskyi has the scutum with black erect setae). It is similar to P. stschurowskii in having the scutum with appressed, silvery setae, but differs in having the labial palpus extremely shortened, inconspicuous, the arolia absent, and the tarsal rake of foreleg with short, stout, bladelike setae (P. stschurowskii has labial palpomeres 2 – 4 elongate, the arolia present, and the tarsal rake with long, bladelike and long, fine, hairlike setae).	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFAEFFFFFF7D1A6B7EB7DEE1.taxon	description	Description. Head. Labial palpomeres 2 – 4 extremely shortened, club-shaped, palpomere 1 elongate: 3 – 4 × as long as combined length of club-shaped labial palpomeres 2 – 4. Maxillar palpomere 6 inconspicuous. Vertex and occiput slightly shiny, microsculptured and punctate, with black erect setae. Gena shiny, punctate. Mesosoma. Scutellum slightly longitudinally depressed, rugose. Metanotum slightly tuberculate dorsally. Mesopleuron, metapleuron, and propodeum coarsely areolate-rugose with black erect setae. Wings dark; veins black; recurrent vein II of forewing received by submarginal cell III. Legs. Claws with 2 ventral teeth; arolia absent. Male. Body length 23 – 25 mm. Head. Labrum trapeziform; Clypeus with wide V-shaped median emargination, glabrous and shiny along apical margin. Face dull, densely punctate and microsculptured with long, black, erect setae. F 3 – F 6 with broad, flat placoids. Mesosoma. Pronotum slightly angulate laterodorsally. Scutum and propodeal enclosure dull, finely, densely rugose, with black erect setae. Metasoma. S 6 with distinctly prominent teeth posterolaterally. Female. Body length 26 – 32 mm. Head. Clypeus entire, nearly rectangular; clypeal disk moderately convex, with rounded depression anteriorly; clypeus laterally and posteriorly, subantennal sclerite, paraocular area, and frons anteriorly with appressed, silvery setae, moderately concealing integument. Mesosoma. Pronotum moderately triangularly prominent dorsally, with black erect setae. Scutum with appressed, silvery setae, moderately concealing integument. Legs. Tarsal rake of foreleg with short, stout, bladelike setae.	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFAEFFFFFF7D1A6B7EB7DEE1.taxon	materials_examined	Material examined. AFGHANISTAN. North Afghanistan, no specific locality [37 ° 00 ’ N, 65 ° 40 ’ E, approximately], 25. VI. 1930, Umnov (1 ♀, ZISP). IRAN. No specific locality (1 ♀, ZISP); Sistan and Baluchestan Province: near Taftan Mount [28 ° 37 ’ N, 61 ° 07 ’ E], 25. V. 1955, D. Steinberg (1 ♀, ZISP); South Khorasan Province: Nehbandan [31 ° 32 ’ N, 60 ° 02 ’ E], 12 – 13. VII. 1901, N. Zarudny (1 ♀, ZISP); Semnan Province: Shahrud [36 ° 23 ’ N, 55 ° 02 ’ E] (1 ♀, ZISP). TURKMENISTAN. Sumbar River, no specific locality, Herz (1 ♂, ZISP); Ahal Province: Archman [38 ° 35 ’ N, 57 ° 07 ’ E], Christoph (1 ♀, lectotype, ZISP); Pulikhatum [35 ° 58 ’ N, 61 ° 08 ’ E] (1 ♀, ZISP); Balkan Province: Sunt-Hasardag Nature Reserve, Sunt mount [38 ° 31 ’ N, 56 ° 21 ’ E], 21. VI. 1952, Odintsova (1 ♂, ZISP); 03. VI. 1956, G. Medvedev (1 ♂, ZISP).	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFAEFFFFFF7D1A6B7EB7DEE1.taxon	distribution	Distribution. Desert areas of the Iranian Plateau (* Afghanistan, Iran, Turkmenistan).	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFADFFE1FF7D1A4F7B10D8CD.taxon	description	(Figs 3 b, 4 d, 5 b, 6 b, 7 b, 8 g, 10 b, 11 b)	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFADFFE1FF7D1A4F7B10D8CD.taxon	diagnosis	Diagnosis. Both sexes of P. radoszkowskyi closely resemble P. persicus in having the labial palpus extremely shortened, inconspicuous, with the club-shaped labial palpomeres 2 – 4, the arolia absent, the male F 3 – F 6 with broad placoids, and the female tarsal rake with short, stout, bladelike setae, but differ in having the labial palpomere 1 short: 1 – 1.5 × as long as the combined length of club-shaped palpomeres 2 – 4 (P. persicus has the labial palpomere 1 elongate: 3 – 4 × as long as the combined length of club-shaped palpomeres 2 – 4). The male of P. radoszkowskyi differs from P. persicus in having S 6 with slightly prominent teeth posterolaterally (P. persicus has S 6 with distinctly prominent teeth). It is similar to P. melanotus and P. stschurowskii in habitus, but differs in having labial palpomeres 2 – 4 extremely short, the placoids on flagellomeres broad, flat, S 6 with pair of teeth posterolaterally, and the arolia absent (P. melanotus and P. stschurowskii have the labial palpomeres 2 – 4 elongate, the placoids on flagellomeres narrow, depressed, the arolia present, and S 6 entire, without teeth). The female of P. radoszkowskyi differs from P. persicus in having the scutum with black erect setae (P. persicus has the scutum with appressed, silvery setae and thick, black, erect setae). It is similar to P. melanotus in habitus, but differs in having the labial palpus extremely shortened, inconspicuous, the arolia absent, and the tarsal rake of foreleg with short, stout, bladelike setae (P. melanotus has the labial palpomeres 2 – 4 elongate, the arolia present, and the tarsal rake with long, bladelike and long, fine, hairlike setae).	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFADFFE1FF7D1A4F7B10D8CD.taxon	description	Description. Head. Labial palpomeres 2 – 4 extremely shortened, club-shaped, palpomere 1 short: 1 – 1.5 × as long as combined length of club-shaped palpomeres 2 – 4. Maxillar palpomere 6 inconspicuous. Vertex and occiput slightly shiny, microsculptured and punctate, with black erect setae. Gena shiny, punctate. Mesosoma with black erect setae. Pronotum slightly angulate laterodorsally. Scutum punctate and slightly obliquely rugose on disk. Scutellum moderately longitudinally depressed, rugose. Metanotum moderately tuberculate dorsally. Mesopleuron, metapleuron, and propodeum areolate-rugose. Wings dark; veins black; recurrent vein II of forewing received by submarginal cell III. Legs. Claws with 2 ventral teeth; arolia absent. Male. Body length 15 – 24 mm. Head. Labrum trapeziform; Clypeus slightly emarginate anteriorly, glabrous and shiny along apical margin. Face densely punctate and microsculptured with long, black, erect setae. F 3 – F 6 with broad, flat placoids. Metasoma. S 6 with slightly prominent teeth posterolaterally. Female. Body length 17 – 25 mm. Head. Clypeus entire, trapeziform; clypeal disk moderately convex, with fine rounded depression anteriorly; clypeus laterally and posteriorly, subantennal sclerite, and paraocular area with sparse appressed, silvery setae, not concealing integument. Legs. Tarsal rake of foreleg with short, stout, bladelike setae. Justification of new synonymy. Having studied the photographs of the paratypes of Sphex guichardi from MZL (two female specimens), as well as the Beaumont’s description, we did not find any differences from radoszkowskyi.	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFADFFE1FF7D1A4F7B10D8CD.taxon	materials_examined	Material examined. KAZAKHSTAN. Aqmola Province: Zharkain District, near Shoindykol Lake [50 ° 27 ’ N, 67 ° 22 ’ E], 24. VI. 1957, V. Rudolf (1 ♂, ZISP); Kyzylorda Province: Shieli District, Baygekum [44 ° 18 ’ N, 66 ° 28 ’ E], 15. V. 1908, L. Wollman (1 ♂, ZISP). TURKMENISTAN. Ahal Province: Kirpili, 100 km NE Serdar [formerly Kyzyl-Arvat] [39 ° 35 ’ N, 57 ° 13 ’ E], 11 – 12. V. 1953, Kirjakova (1 ♂, ZISP); Daşoguz Province: Ortakui well, 240 km SW Daşoguz [40 ° 41 ’ N, 57 ° 29 ’ E], 26. V. 1953, E. Arens (1 ♂, ZISP); 22. V. 1953 (1 ♂, ZISP); 29. V. 1953 (2 ♂, ZISP); 19. V. 1953 (2 ♂, ZISP); Mary Province: Imeni Chapayeva [formely Ymam Baba] [36 ° 42 ’ N, 62 ° 27 ’ E], 03 – 08. V. 1912, Koshantschikow (2 ♀, ZISP); 12 – 14. V. 1912 (1 ♀, ZISP). UZBEKISTAN. Jizzakh Province: Gallaorol District, Lalmikor [39 ° 56 ’ N, 67 ° 27 ’ E], 26. VII. 1930 (1 ♀, ZISP); Qashqadaryo Province: Kamashi District, Kamashi [38 ° 49 ’ N, 66 ° 27 ’ E], 28. VI. 1931, V. Gussakovskij (4 ♀, ZISP).	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFADFFE1FF7D1A4F7B10D8CD.taxon	distribution	Distribution. Arid and semi-arid areas from Central Asia to Anatolia (Kazakhstan, Turkey, Turkmenistan, Uzbekistan).	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFB3FFE1FF7D1863790CDC65.taxon	description	(Fig. 10 c)	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFB3FFE1FF7D1863790CDC65.taxon	diagnosis	Diagnosis. Both sexes of P. reymondi resemble P. subfuscatus, P. judaeus, and P. crudelis, but differ in having the wings smoky and the body brownish (P. subfuscatus, P. judaeus, and P. crudelis have the wings yellow and the body black).	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFB3FFE1FF7D1863790CDC65.taxon	description	Description. Body brownish with fine dark erect setae. Head. Clypeus, subantennal sclerite, paraocular area and frons with fine, sparse, appressed, silvery setae. Mesosoma. Scutum slightly shiny, finely micropunctate and sparsely punctate. Metanotum tuberculate dorsally. Mesopleuron, metapleuron and propodeum areolate-rugose, shiny; propodeal enclosure dull. Wings smoky; veins light brown. Legs. Claws with 2 ventral teeth; arolia present. Male. Body length 22 mm. Head. Clypeus with wide V-shaped median emargination. Metasoma. S 6 entire, without teeth. Female. Body length 22 mm. Head. Clypeus angulate, with U-shaped median notch. Legs. Tarsal rake of foreleg with long, stout, bladelike and some short, hairlike setae.	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFB3FFE1FF7D1863790CDC65.taxon	materials_examined	Material examined. None.	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFB3FFE1FF7D1863790CDC65.taxon	distribution	Distribution. Desert areas of the Saoura area (Algeria).	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFB2FFE3FF7D18EC7F16D965.taxon	description	(Figs 1 d, 4 b, 5 f, 6 f, 10 d)	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFB2FFE3FF7D18EC7F16D965.taxon	diagnosis	Diagnosis. Both sexes of P. saevus closely resemble P. subfuscatus, P. crudelis, and P. judaeus. The female of P. saevus differs from P. crudelis and P. judaeus in having the head and mesosoma with pale setae, propodeal enclosure without long, dense, black setae, tarsal rake of foreleg only with some long, stout, bladelike setae (P. crudelis and P. judaeus have the head and mesosoma with dark setae, the propodeal enclosure with long, dense, hairlike black setae, the tarsal rake of the foreleg with long, stout, bladelike and some hairlike setae). The male of P. saevus differs from P. crudelis and P. judaeus in having the propodeal enclosure without long, dense, black setae (propodeal enclosure with long, dense, hairlike black setae in P. crudelis and P. judaeus). The female of P. saevus differs from P. subfuscatus in having the mandible reddish mesally (mandible black in P. subfuscatus). The male of P. saevus differs from P. subfuscatus in having F 3 – F 4 with placoids and S 4 – S 6 with long black setae apically (F 3 – F 6 with placoids and S 4 – S 6 without long black setae in P. subfuscatus).	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFB2FFE3FF7D18EC7F16D965.taxon	description	Description. Head. Labial palpomeres 2 – 4 usual. Vertex, occiput and gena microsculptured, punctate. Mesosoma. Pronotum moderately angulate laterodorsally, finely rugose. Scutum, mesopleuron, metapleuron, and propodeum areolate-rugose, slightly shiny. Scutellum slightly longitudinally depressed, longitudinlly rugose and punctate. Metanotum slightly tuberculate dorsally. Wings yellow in venation zone, smoky apically; veins light brown to yellow; recurrent vein II of forewing received by submarginal cell III. Legs. Claws with 2 ventral teeth; arolia present. Male. Body length 15 – 22 mm. Body black with black erect setae. Head. Labrum trapeziform. Clypeus with wide V-shaped median emargination. Clypeus, subantennal sclerite and paraocular area with appressed, silvery setae, moderately concealing integument. F 3 – F 6 with narrow, depressed placoids. Mesosoma. Propodeal enclosure with long, sparse black setae. Metasoma. S 4 – S 6 with long black setae; S 6 entire, without teeth. Female. Body length 20 – 28 mm. Body black with light erect setae. Head. Labrum trapeziform, emarginate apically. Mandible reddish mesally. Clypeus angulate, with U-shaped median notch. Clypeus, subantennal sclerite, paraocular area and frons with appressed, goldish or silvery setae, concealing integument. Mesosoma. Pronotum and scutum with sparse semiappressed, goldish or silvery setae, not concealing integument. Propodeal enclosure with sparse light setae. Legs. Tarsal rake of foreleg with long, stout, bladelike and some short, hairlike setae.	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFB2FFE3FF7D18EC7F16D965.taxon	materials_examined	Material examined. None.	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFB2FFE3FF7D18EC7F16D965.taxon	distribution	Distribution. Semi-arid areas of Australasian region (Australia, Indonesia (Lesser Sunda Islands)).	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFB1FFE5FF7D1AE27EA8DA61.taxon	description	(Figs 3 d, 4 c, 5 h, 6 h, 8 h, 10 e, 12 d)	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFB1FFE5FF7D1AE27EA8DA61.taxon	diagnosis	Diagnosis. The male of P. stschurowskii closely resembles P. melanotus in having the labial palpomeres 2 – 4 elongate, the placoids on flagellomeres narrow, depressed, and S 6 entire, without teeth, but differs in having F 3 – F 6 with placoids (P. melanotus has F 3 – F 8 with the placoids). It is similar to P. persicus and P. radoszkowskyi in habitus, but differs in having the labial palpomeres 2 – 4 elongate, the placoids on flagellomeres narrow, depressed, S 6 entire, without teeth, and the arolia present (P. persicus and P. radoszkowskyi have the labial palpomeres 2 – 4 extremely short, the placoids on flagellomeres broad, flat, S 6 with pair teeth posterolaterally, and the arolia absent). The female of P. stschurowskii resembles P. melanotus, P. persicus, and P. radoszkowskyi in habitus. It differs from P. persicus, and P. radoszkowskyi in having the labial palpomeres 2 – 4 elongate, and the arolia present (P. persicus and P. radoszkowskyi have the labial palpomeres 2 – 4 extremely short, and the arolia absent). Unlike P. melanotus and P. radoszkowskyi it has the scutum with appressed, silvery setae.	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFB1FFE5FF7D1AE27EA8DA61.taxon	description	Description. Head with black erect setae. Labrum trapeziform, distinctly emarginate apically; labial palpomeres 2 – 4 elongate; clypeus laterally and posteriorly, subantennal sclerite, paraocular area, and frons with appressed, silvery setae; vertex, occiput and gena microsculptured, punctate, slightly shiny. Mesosoma with black erect setae. Scutellum slightly longitudinally depressed, punctate and rugose, shiny. Metanotum tuberculate dorsally. Mesopleuron areolate-rugose and punctate, shiny. Metapleuron and propodeum areolate-rugose, shiny; propodeal enclosure slightly shiny or dull. Wings dark or smoky (in var. hyalipennis); veins black; recurrent vein II of forewing interstitial with 1 st intersubmarginal vein or received by submarginal cell III at its basal angle. Legs. Claws with 2 ventral teeth; arolia present. Male. Body length 20 – 23 mm. Head. Clypeus slightly emarginate anteriorly; clypeal disk moderately convex, with fine depression anteriorly. F 3 – F 6 with narrow, depressed placoids. Mesosoma. Pronotum and scutum finely rugose and punctate, slightly shiny, with black erect setae only. Metasoma. S 6 entire, without teeth. Female. Body length 23 – 30 mm. Head. Anterior margin of clypeus slightly rounded; clypeal disk moderately convex, slightly depressed anteriorly. Mesosoma. Pronotum and scutum with appressed, silvery setae, moderately concealing integument. Legs. Tarsal rake of foreleg with long, bladelike and long, fine, hairlike setae. Justification of new synonymy. By all key features (labial palpomeres 2 – 4 elongate; arolia present; recurrent vein II of forewing interstitial with 1 st intersubmarginal vein or received by submarginal cell III at its basal angle; placoids on flagellomeres narrow, depressed; S 6 entire, without teeth), Sphex sirdariensis Radoszkowski belongs in the stschurowskii group. Both the lectotype female of stschurowskii and the lectotype male of sirdariensis come from the same locality. In addition, the description of male of stschurowskii by Morice (1897) well agrees with sirdariensis.	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFB1FFE5FF7D1AE27EA8DA61.taxon	materials_examined	Material examined. IRAN. Hormozgan Province: 30 km S of Hajjiabad [28 ° 01 ’ N, 55 ° 53 ’ E], 27. VI. 1955, D. Steinberg (1 ♀, ZISP); 7. V. 1955 (1 ♀, ZISP); Sistan and Baluchestan Province: Bampur [27 ° 11 ’ N, 60 ° 27 ’ E], 26 – 27. IV. 1901, N. Zarudny (1 ♀, ZISP). KAZAKHSTAN. Turkistan Province: Shardara District, Kyzylkum Desert on the left bank of Syr Darya River [42 ° 00 ’ N, 68 ° 00 ’ E approximately], 13. V. 1871, A. Fedchenko (1 ♀, lectotype of stschurowskii, ZMMU); 19. V. 1871 (1 ♂, lectotype of sirdariensis, ZMMU). MOROCCO. Midelt [32 ° 40 ’ N, 4 ° 43 ’ W], 30. V. 1947, J. de Beaumont (1 ♀, ZISP). TAJIKISTAN. Khatlon Province: Jilikul District, Jilikul [37 ° 29 ’ N, 68 ° 31 ’ E], 25. V. 1931, N. Fursov (1 ♀, ZISP). TURKMENISTAN. No specific locality [Transcaspia] (1 ♀, ZISP); Mary Province: Imeni Chapayeva [formely Ymam Baba] [36 ° 42 ’ N, 62 ° 27 ’ E], 1932, A. Shestakov (1 ♂, 1 ♀, ZISP).	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFB1FFE5FF7D1AE27EA8DA61.taxon	distribution	Distribution. Arid and semi-arid areas from Central Asia to North Africa (Afghanistan, Algeria, Egypt, Iran, Iraq, Israel, Kazakhstan, Libya, Morocco, Saudi Arabia, Tajikistan, Tunisia, Turkmenistan).	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFB7FFEBFF7D19527B71DCD1.taxon	description	(Figs 1 e, 4 c, 8 i, 10 f, 11 d)	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFB7FFEBFF7D19527B71DCD1.taxon	diagnosis	Diagnosis. Both sexes of P. subfuscatus closely resemble P. saevus, P. crudelis, and P. judaeus. The female of P. subfuscatus differs from P. crudelis and P. judaeus in having the head and mesosoma with pale setae, propodeal enclosure without long, dense, black setae, tarsal rake of foreleg only with some long, stout, bladelike setae (P. crudelis and P. judaeus have the head and mesosoma with dark setae, the propodeal enclosure with long, dense, hairlike black setae, the tarsal rake of the foreleg with long, stout, bladelike and some hairlike setae). The male of P. subfuscatus differs from P. crudelis and P. judaeus in having the propodeal enclosure without long, dense, black setae (propodeal enclosure with long, dense, hairlike black setae in P. crudelis and P. judaeus). The female of P. subfuscatus differs from P. saevus in having the mandible black (mandible reddish mesally in P. saevus). The male of P. subfuscatus differs from P. saevus in having F 3 – F 6 with placoids and S 4 – S 6 without long black setae (F 3 – F 4 with placoids and S 4 – S 6 with long black setae apically in P. saevus).	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFB7FFEBFF7D19527B71DCD1.taxon	description	Description. Head. Labial palpomeres 2 – 4 usual. Mandible black. Vertex, occiput and gena microsculptured, punctate. Mesosoma. Pronotum moderately angulate laterodorsally, finely rugose. Scutum, mesopleuron, metapleuron, and propodeum areolate-rugose, slightly shiny. Scutellum slightly longitudinally depressed, longitudinlly rugose and punctate. Metanotum slightly tuberculate dorsally. Wings yellow in venation zone, smoky apically; veins light brown to yellow; recurrent vein II of forewing received by submarginal cell III. Legs. Claws with 2 ventral teeth; arolia present. Male. Body length 10 – 17 mm. Body black with black erect setae. Head. Labrum trapeziform. Clypeus with wide V-shaped median emargination. Clypeus, subantennal sclerite and paraocular area with appressed, silvery setae, moderately concealing integument. F 3 – F 6 with narrow, depressed placoids. Mesosoma. Propodeal enclosure with long, sparse black setae. Metasoma. S 4 – S 6 without long black setae; S 6 entire, without teeth. Female. Body length 14 – 22 mm. Body black with light erect setae. Head. Labrum trapeziform, emarginate apically. Clypeus angulate, with U-shaped median notch. Clypeus, subantennal sclerite, paraocular area and frons with appressed, goldish or silvery setae, concealing integument. Mesosoma. Pronotum and scutum with sparse semiappressed, goldish or silvery setae, not concealing integument. Propodeal enclosure with sparse light setae. Legs. Tarsal rake of foreleg with long, stout, bladelike and some short, hairlike setae. Justification of new synonymy. Having studied the photographs of the paratypes of Prionyx binghami from NPC, as well as the original description, we did not find any clear differences from subfuscatus. Questionable synonymy. Sphex rukwaensis described by G. Arnold (1959) in the subgenus Calosphex was tentatively treated as Prionyx subfuscatus rukwaensis by R. Bohart and A. Menke (1976). Arnold in diagnosis of rukwaensis pointed out the differences between his new species and the other two African species of the subgenus Calosphex: P. niveatus (Dufour, 1854) and P. nigropectinatus (Taschenberg, 1869). However, he did not mention subfuscatus, which was well known to him. Unfortunately, we were unable to study the holotype of rukwaensis, and the description does not provide sufficient evidence to resurrect this taxon to full species status.	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFB7FFEBFF7D19527B71DCD1.taxon	materials_examined	Material examined. AZERBAIJAN. Ganja [formerly Yelisavetpol] [40 ° 40 ’ N, 46 ° 21 ’ E], 14 – 19. V. 1909, I. Babadjanidi (1 ♂, ZISP); Nakhchivan Autonomous Republic: Shahbuz District, Türkeş [39 ° 25 ’ N, 45 ° 28 ’ E], 18 – 26. VII. 1940, M. Ryabov (2 ♂, ZISP). CHINA. Xinjiang Uyghur Autonomous Region: Ili Kazakh Autonomous Prefecture, Künes River, no specific locality [43 ° 35 ’ N, 82 ° 33 ’ E], VIII. 1877, N. Przhevalsky (1 ♀, ZISP). IRAN. South Khorasan Province: Birjand [32 ° 52 ’ N, 59 ° 12 ’ E], 21 – 26. VI. 1896 [the dates are possibly in error], N. Zarudny, (5 ♀, ZISP); Khusf County, Chahkand-e Gol [32 ° 41 ’ N, 59 ° 13 ’ E], 17. VI. 1896 [the dates are possibly in error], N. Zarudny (1 ♂, ZISP); Zirkuh County, Dakkepatargun salt-marsh near Shahrakht [33 ° 27 ’ N, 60 ° 37 ’ E], 24. IV. 1898, N. Zarudny (7 ♂, 2 ♀, ZISP); Zirkuh County, Makhunik [32 ° 27 ’ N, 60 ° 25 ’ E], 1. V. 1898, N. Zarudny (1 ♀, ZISP); Zirkuh County, Ardakul near Mohammadabad [33 ° 29 ’ N, 60 ° 03 ’ E], 22. IV. 1898, N. Zarudny (1 ♂, ZISP); Ardabil Province: Parsabad County, Owltan [39 ° 36 ’ N, 47 ° 46 ’ E], 30. VI. 1927, Zapolsky (2 ♀, ZISP). KAZAKHSTAN. Abai Province: Beskaragay District, near Kanonerka [50 ° 41 ’ N, 79 ° 41 ’ E], 18. VI. 1955, P. Rafes (1 ♀, ZISP); Akmola Province: Kokshetau Mountains [53 ° 05 ’ N, 70 ° 16 ’ E], 23. VI. 1957, V. Rudolf (1 ♂, ZISP); 12. VII. 1957, V. Tobias (2 ♂, ZISP); Aktobe Province: Mugalzhar District, Zhaksytau in Mugodzhar Hills [48 ° 10 ’ N, 58 ° 23 ’ E], 26. VII. 1904, V. Dubiansky (1 ♀, ZISP); Yrgyz District, Shet-Irgiz River [48 ° 45 ’ N, 60 ° 08 ’ E], 10. VII. 1928, V. Popov (3 ♀, ZISP); Lesser Barsuki desert [47 ° 12 ’ N, 60 ° 44 ’ E], 28. VI – 1. VII. 1910, N. Androssow (1 ♂, 1 ♀, ZISP); 21. VII. 1930, E. Luppova (1 ♀, ZISP); Greater Barsuki desert near Shalkar [47 ° 35 ’ N, 59 ° 32 ’ E], 14. VI. 1907, N. Androssow (1 ♂, 2 ♀, ZISP); 24. VI. 1930, A. Gozhev (1 ♀, ZISP); 1. VI. 1908, N. Androssow (1 ♀, ZISP); 3. VI. 2011, A. Ivanov (1 ♀, ZISP); Mugalzhar District, Kokzhide Sands [48 ° 24 ’ N, 57 ° 13 ’ E], 19 – 23. VI. 1908, D. Borodin, B. Uvarov (9 ♂, 1 ♀, ZISP); Shalkar District, Berchogur [48 ° 25 ’ N, 58 ° 43 ’ E], 26. VI. 1910, L. Wollman (1 ♂, ZISP); Shalkar District, near Shalkar [47 ° 47 ’ N, 59 ° 41 ’ E], 2. VII. 1930, A. Gozhev (1 ♂, ZISP); Almaty Province: Ile District, Usharal [44 ° 14 ’ N, 76 ° 51 ’ E], 4. VII. 1903, L. Bergh (2 ♂, ZISP); Mountains near Almaty, no specific locality [43 ° 05 ’ N, 76 ° 50 ’ E], 17. VI. 1903, L. Bergh (1 ♂, ZISP); Balkhash District, Bokkary natural boundary near Akkol [45 ° 08 ’ N, 75 ° 20 ’ E], 8. VII. 1903, L. Bergh (1 ♀, ZISP); Balkhash District, near Akzhar [44 ° 52 ’ N, 75 ° 53 ’ E], 26. V. 2015, K. Fadeev (1 ♀, ZISP); Balkhash District, 17 km NW Bakanas [44 ° 52 ’ N, 76 ° 01 ’ E], 23. VI – 15. VII. 1970, V. Kazenas (2 ♂, SZMN); 17 – 19. VII. 1970, V. Kazenas (2 ♀, FSCV); Atyrau Province: Inder District, Lake Inder [48 ° 28 ’ N, 51 ° 54 ’ E], 1. VI. 1907, B. Uvarov (2 ♂, ZISP); 31. V – 9. VI. 1907, D. Borodin (15 ♂, 1 ♀, ZISP); 27. VI. 1951, D. Steinberg (1 ♂, ZISP); 23. VI. 1951, V. Tobias (1 ♂, ZISP); East Kazakhstan Province: Katonkaragay District, Akkaynar [formerly Chernovoye] [49 ° 13 ’ N, 85 ° 53 ’ E], 8 – 10. VIII. 1897, Silantiev (2 ♂, 2 ♀, ZISP); Jambyl Province: Moiynkum District, Mynaral [45 ° 26 ’ N, 73 ° 41 ’ E], 26. VII. – 16. VIII. 1903, L. Bergh (2 ♀, ZISP); Moiynkum District, Basaral island in Lake Balkhash [45 ° 28 ’ N, 73 ° 44 ’ E], 1. VIII. 1903, L. Bergh (2 ♀, ZISP); Moiynkum District, near Moiynkum [45 ° 26 ’ N, 73 ° 38 ’ E], 10. VI. 1952, M. Baitenov (1 ♂, SZMN); Turar Ryskulov District, Akyrtobe [42 ° 59 ’ N, 72 ° 06 ’ E], 7. VII. 1931, Veltischev (1 ♀, ZISP); Turar Ryskulov District, Kargaly lake in Moiynkum Desert [43 ° 19 ’ N, 72 ° 01 ’ E], 23. V. 1910, A. Kirichenko (1 ♂, ZISP); Taraz [formerly Auliye-Ata or Dzhambul] [42 ° 53 ’ N, 71 ° 22 ’ E], 21. VIII. 1931, Veltischev (2 ♂, ZISP); Kostanay Province: Auliekol District, Timofeevka [52 ° 17 ’ N, 63 ° 31 ’ E], 27. VII. 1928, Raevsky (1 ♀, ZISP); Terisaqqan river near Arkalyk [50 ° 15 ’ N, 67 ° 39 ’ E], 11 – 15. VI. 1957, V. Tobias (1 ♂, 1 ♀, ZISP); Kyzylorda Province: Kyzylorda [formerly Ak-Mechet or Perovsk] [44 ° 51 ’ N, 65 ° 31 ’ E], 7. VI. 1925, A. Ushinsky (1 ♀, ZISP); 16. V. 1916, N. Pulikovskaja (1 ♀, ZISP); near Kyzylorda [44 ° 47 ’ N, 65 ° 45 ’ E], 8. VI. 2015, K. Fadeev (4 ♂, ZISP); Kazaly District, Syr Darya River mouth [45 ° 40 ’ N, 61 ° 44 ’ E], VI. 1898, I. Heyer (1 ♀, ZISP); Shieli district, Syr Darya River near Baigekum [Djulek] [44 ° 14 ’ N, 66 ° 20 ’ E], 29. V. 1898, I. Heyer (1 ♂, ZISP); Shieli district, Meshugul natural boundary near Shieli [44 ° 01 ’ N, 66 ° 31 ’ E], 28. V. 1898, I. Heyer (2 ♂, ZISP); Zhanakorgan District, Zhanakorgan [formerly Yany-Kurgan] [43 ° 54 ’ N, 67 ° 15 ’ E], 27. V. 1898, I. Heyer (1 ♂, ZISP); Aral District, Barsa-Kelmes [45 ° 38 ’ N, 59 ° 52 ’ E], 4. VI. 1983, D. Piryulin (1 ♀, ZISP); Aral District, Barsa-Kelmes Lake [46 ° 20 ’ N, 59 ° 41 ’ E], 30. V. 2011, A. Ivanov (1 ♂, ZISP); Mangystau Province: Mangystau District, Sengyrkum sands [43 ° 48 ’ N, 53 ° 31 ’ E], no date, A. Ivanov (1 ♂, ZISP); Qarağandy Province: Konyrat [formerly Kounradskiy] near Balkhash [46 ° 57 ’ N, 74 ° 59 ’ E], 3. VII. 2014, K. Fadeev (1 ♂, ZISP); Turkistan Province: Shardara District, the border of the Kyzylkum desert and the Golodnaya Steppe [Mirzachoʻl] on Syr Darya River [approximately 41 ° 11 ’ N, 67 ° 54 ’ E], 10 – 12. V. 1903, G. Jakobson (9 ♂, 4 ♀, ZISP); Ushkaik [43 ° 09 ’ N, 67 ° 54 ’ E], 24. V. 1898, I. Heyer (1 ♀, ZISP); Ulytau Province: Zhanaarka District, Koksengir mount [48 ° 22 ’ N, 71 ° 31 ’ E], 28. VI. 1959, M. Kozlov (1 ♀, ZISP); West Kazakhstan Province: Akzhaik District, Kharkino [48 ° 45 ’ N, 51 ° 48 ’ E], 2 – 19. VI. 1951, V. Rudolf (7 ♂, ZISP); 2. VI. 1951, V. Tobias (2 ♂, ZISP); Oral [= Uralsk] [51 ° 14 ’ N, 51 ° 23 ’ E], 1. VII. 1906, B. Uvarov (1 ♀, ZISP); 16. VII. 1908, B. Uvarov (2 ♀, ZISP); 10. VII. 1908, D. Borodin, B. Uvarov (1 ♂, ZISP). KYRGYZSTAN. Talas Province: Talas [formerly Dmitrievka] [42 ° 31 ’ N, 72 ° 13 ’ E], 24. V. 1909, Minkwitz (1 ♂, ZISP); Chüy Province: Moskva District, Sretenka [42 ° 55 ’ N, 74 ° 07 ’ E], 17. VI. 1931, L. Zimin (1 ♀, ZISP); Moskva District, Belovodskoye [42 ° 49 ’ N, 74 ° 09 ’ E], 5. VIII. 1931, L. Zimin (1 ♂, ZISP); Moskva District, near Telek [43 ° 07 ’ N, 74 ° 05 ’ E], 5. VIII. 1999, D. Milko (1 ♀, SZMN); Kemin District, near Cholok [42 ° 41 ’ N, 75 ° 53 ’ E], 31. VII. 2000, V. Dubatolov (2 ♂, SZMN); Ysyk-Ata District, near Sary-Dzhon [42 ° 56 ’ N, 74 ° 50 ’ E], 6. VIII. 1999 (1 ♀, SZMN). MONGOLIA. Govi-Altai Province: Bugat [45 ° 33 ’ N, 94 ° 22 ’ E], 20. VII. 1970, E. Nartshuk (1 ♂, ZISP). RUSSIA. Altai Territory: near Kulunda [52 ° 33 ’ N, 78 ° 56 ’ E], 6 – 13. VII. 1922, A. Reichardt (2 ♂, 1 ♀, ZISP); Klyuchevsky District, near Klyuchi [52 ° 13 ’ N, 79 ° 06 ’ E], 22. VIII. 1952, G. Zolotarenko (1 ♀, SZMN); Astrakhan Province: Astrakhan [46 ° 19 ’ N, 48 ° 01 ’ E], A. Gebel’ (1 ♀, ZISP); 1868, A. Bekker (1 ♂, 1 ♀, ZISP); Limansky District, 13 km S of Liman [45 ° 39 ’ N, 47 ° 14 ’ E], 25. VII. 2015, S. Belokobylskij, V. Loktionov, M. Mokrousov, M. Proshchalykin (1 ♂, PCMM); Enotayevsky District, near Zamjany [46 ° 49 ’ N, 47 ° 37 ’ E], 22. VIII. 2015, S. Belokobylskij, V. Loktionov, M. Mokrousov, M. Proshchalykin (2 ♂, 3 ♀, PCMM); 22. VIII. 2015, M. Mokrousov (1 ♀, SZMN); 22. VII. 2017, M. Mokrousov (1 ♀, SZMN); Enotayevsky District, near Nikolskoye [47 ° 46 ’ N, 46 ° 23 ’ E], 8. VIII. 1989, P. Lehr (1 ♂, SZMN); Narimanovsky District, Volzhskoye [46 ° 39 ’ N, 47 ° 52 ’ E], 26. VII. 2015, S. Belokobylskij, V. Loktionov, M. Mokrousov, M. Proshchalykin (2 ♂, PCMM); 8. VII – 18. VIII. 2017, M. Mokrousov (13 ♂, 3 ♀, SZMN); Akhtubinsky District, near Kapustin Yar [48 ° 35 ’ N, 45 ° 49 ’ E], 16 – 21. VIII. 2003, V. Sapronov (1 ♀, ZISP); Akhtubinsky District, near Baskunchak Lake [48 ° 12 ’ N, 46 ° 49 ’ E], 30. V. 1996, V. Dubatolov (1 ♂, SZMN); Buryat Republic: Selenginsky District, near Baraty [51 ° 15 ’ N, 106 ° 21 ’ E], 26. VII. 2007, A. Lelej, M. Proshchalykin, V. Loktionov (3 ♂, FSCV; 1 ♀, SZMN); Dzhidinsky District, near Dzhida [50 ° 41 ’ N, 106 ° 13 ’ E], A. Lelej, M. Proshchalykin, V. Loktionov (1 ♂, FSCV); Crimea Republic: Kerch [45 ° 22 ’ N, 36 ° 25 ’ E], no date, A. Kirichenko (1 ♂, 1 ♀, ZISP); Simferopol [44 ° 57 ’ N, 34 ° 05 ’ E], 15. VI. 1924, collector unknown (1 ♀, ZISP); Dagestan Republic: Tarumovsky District, Kuma river [44 ° 56 ’ N, 46 ° 27 ’ E], 19. VII. 2015, S. Belokobylskij, V. Loktionov, M. Mokrousov, M. Proshchalykin (6 ♂, PCMM); Tarumovsky District, Alexandro-Nevskoye [43 ° 55 ’ N, 46 ° 33 ’ E], 9. VI. – 9. VIII. 1927, N. Olsufjev (3 ♀, ZISP); Kalmyk Republic: Chernozemelsky District, near Komsomolsky [45 ° 17 ’ N, 45 ° 56 ’ E], 2. VI. 2016, S. Belokobylskij, V. Loktionov, M. Mokrousov, M. Proshchalykin (17 ♂, PCMM); Chernozemelsky District, near Mekleta [45 ° 42 ’ N, 45 ° 43 ’ E], 19. VI. 2017, M. Mokrousov (4 ♂, 2 ♀, SZMN); Chernozemelsky District, 26 km NWW Artesian [45 ° 00 ’ N, 46 ° 19 ’ E], 14. VIII. 2015, M. Mokrousov (1 ♀, SZMN); Ketchenerovsky District, near Ketchenery [47 ° 17 ’ N, 44 ° 39 ’ E], 14. VII. 2015, S. Belokobylskij, V. Loktionov, M. Mokrousov, M. Proshchalykin (1 ♂, PCMM); Ketchenerovsky District, near Tsagan-Nur [47 ° 20 ’ N, 45 ° 14 ’ E], 13. VII. 2015, S. Belokobylskij, V. Loktionov, M. Mokrousov, M. Proshchalykin (2 ♀, PCMM); Yashkulsky District, near Yashkul [46 ° 12 ’ N, 45 ° 22 ’ E], 16. VII. 2015, S. Belokobylskij, V. Loktionov, M. Mokrousov, M. Proshchalykin (3 ♂, PCMM); Kherson Province: Aleshky near Kherson [46 ° 37 ’ N, 32 ° 43 ’ E], 20 VIII. 1925, A. Kirichenko (1 ♀, ZISP); 22. VI – 14. VII. 1924, S. Fedorov (10 ♂, 1 ♀, ZISP); Krasnodar Territory: Apsheronsky District, Kamyshanov [44 ° 10 ’ N, 40 ° 02 ’ E], 27. VIII. 2016, V. Zintshenko (1 ♀, SZMN); 25. VIII. 2016, K. Yalinchuk (1 ♀, SZMN); Orenburg Province: Svetlinsky District, Shalkar-Yega-Kara Lake [50 ° 46 ’ N, 60 ° 49 ’ E], 15 – 25. VIII. 2003, T. Tunyova (1 ♀, ZISP); Svetlinsky District, Aschisayskaya steppe Nature Reserve [50 ° 59 ’ N, 61 ° 13 ’ E], 7 – 8. VIII. 2015, K. Fadeev (5 ♂, 9 ♀, ZISP); Primorskii Territory: Lazovsky District, Lazovski Nature Reserve, near Glazkovka [43 ° 01 ’ N, 134 ° 07 ’ E], 16. VII. 2006, P. Nemkov (1 ♂, 1 ♀, FSCV; 1 ♂, SZMN); Rostov Province: Salsky District, no specific locality [46 ° 30 ’ N, 41 ° 30 ’ E], 3. VIII. 1931, B. Roddendorf (1 ♂, ZISP); Saratov Province: Saratov [51 ° 32 ’ N, 45 ° 55 ’ E], 20. VI – 5. VII. 1896, collector unknown (1 ♀, ZISP); Pugachyovsky District, no specific locality [51 ° 56 ’ N, 49 ° 03 ’ E], 1913, V. Bostanzhoglo (5 ♂, ZISP); Stavropol Territory: Shpakovsky District, Pelagiada [45 ° 13 ’ N, 42 ° 01 ’ E], 3. VII. 1925, D. Tratnikov (1 ♀, ZISP); Neftekumsky District, Achikulak [44 ° 32 ’ N, 44 ° 49 ’ E], 11. IX. 1953, P. Rafes (1 ♂, 1 ♀, ZISP); Tuva Republic: Kyzyl [51 ° 42 ’ N, 94 ° 22 ’ E], 16. VII. 2009, S. Belokobylskij (1 ♂, 2 ♀, ZISP); 18. VII. 2000, O. Kosterin, N. Priidak (1 ♀, SZMN); Chaa-Kholsky District, Chaa-Khol river near Ak-Durug [51 ° 21 ’ N, 92 ° 13 ’ E], 6. VIII. 2009, S. Belokobylskij (1 ♀, ZISP); 14. VII. 1962, collector unknown (1 ♂, SZMN); Erzinsky District, Tore-Khol Lake [50 ° 05 ’ N, 95 ° 10 ’ E], 6. VII. 1997, Yu. Mikhailov (1 ♀, ZISP); 12. VII. 1993, S. Tshernyshev (1 ♂, SZMN); Tandinsky District, Sosnovka [51 ° 08 ’ N, 94 ° 31 ’ E], 5. VII. 1949, Dyatlova (1 ♂, SZMN); Kyzylsky District, Elegest River [51 ° 28 ’ N, 94 ° 11 ’ E], 22. VII. 2014, A. Lelej, M. Proshchalykin, V. Loktionov (1 ♂, SZMN); Piy-Khemsky District, Piy-Khem River [51 ° 54 ’ N, 94 ° 25 ’ E], 4. VII. 2010, A. Erst (1 ♀, SZMN); Ulug-Khemsky District, near Shagonar [51 ° 29 ’ N, 92 ° 54 ’ E], 20 – 26. VI. 2017, A. Byvaltsev, N. Moskalenko (3 ♂, SZMN); Volgograd Province: Volgograd [48 ° 31 ’ N, 44 ° 32 ’ E], 1866, A. Bekker (4 ♂, ZISP); 1867, A. Bekker (1 ♀, ZISP); 1872, A. Bekker (3 ♂, 1 ♀, ZISP); 11. VI. 1907, M. Koch (1 ♂, ZISP); 28. VI. 1993, V. Krivokhatsky (1 ♂, ZISP); Kamyshinsky District, Yelshanka [50 ° 10 ’ N, 45 ° 23 ’ E], 21. VII – 6. VIII. 1933, L. Zimin (4 ♂, ZISP); Pallasovsky District, Lake Elton [49 ° 10 ’ N, 46 ° 31 ’ E], 20. VI. 2003, V. Krivokhatsky, O. Ovtshinnikova (1 ♂, ZISP); Zabaikalskii Territory: Onon river, no specific locality [50 ° 37 ’ N, 115 ° 32 ’ E], 12. VII. 1926, Naumov (1 ♀, ZISP); Peschanka near Chita [51 ° 59 ’ N, 113 ° 36 ’ E], 15 – 20. VIII. 1927, Mikhno (1 ♂, ZISP). SOUTH KOREA. Busan [35 ° 11 ’ N, 129 ° 04 ’ E], 6 – 7. IX. 1900, A. Semenov (1 ♀, ZISP). TAJIKISTAN. Gorno-Badakhshan Autonomous Region: Vanj [38 ° 21 ’ N, 71 ° 26 ’ E], 18. IX. 1943, D. Stackelberg (1 ♂, 1 ♀, ZISP); Khatlon Province: Bokhtar [formerly Qurghonteppa or Kurgan-Tyube] [37 ° 49 ’ N, 68 ° 46 ’ E], 31. V. 1931, N. Fursov (1 ♂, ZISP); Region of Republican Subordination: Varzob District, Varzob [38 ° 46 ’ N, 68 ° 49 ’ E], 14. VI. 2012, Sh. Nazarova (1 ♂, SZMN). TURKMENISTAN. Ashgabat: Büzmeýin District, Archabil [= Firjuza] [37 ° 55 ’ N, 58 ° 05 ’ E], 8. VII. 1906, K. Angher (1 ♀, ZISP); Ashgabat [37 ° 57 ’ N, 58 ° 23 ’ E] 31. V – 6. VI. 1925, V. Gussakovskij (4 ♂, 1 ♀, ZISP); 26. V. [year?], K. Angher (1 ♀, ZISP); 20. V. 1916, A. Ushinsky (1 ♂, ZISP); 8. VII. 1924, A. Ushinsky (2 ♂, ZISP); 19. V. 1932, A. Ushinsky (1 ♂, ZISP); Balkan Province: Magtymguly [formerly Kara-Kala] [38 ° 26 ’ N, 56 ° 18 ’ E], 6 – 8. VIII. 1934, V. Popov (2 ♂, 2 ♀, ZISP); Gyzylarbat [formerly Serdar or Kyzyl-Arvat] [38 ° 59 ’ N, 56 ° 16 ’ E], 2 – 4. V. 1912, Gedevanov (1 ♀, ZISP); Daşoguz Province: Daşoguz [formerly Tashauz] [41 ° 50 ’ N, 59 ° 58 ’ E], 21. VII. 1931, A. Ushinsky (1 ♀, ZISP); Lebap Province: Repetek Biosphere State Reserve [38 ° 34 ’ N, 63 ° 10 ’ E], 30. VI. 1982, V. Krivokhatsky (1 ♀, ZISP); Mary Province: Imeni Chapayeva [formely Ymam Baba] [36 ° 42 ’ N, 62 ° 27 ’ E], 18 – 20. VI. 1925, V. Gussakovskij (1 ♀, ZISP); Baýramaly [37 ° 37 ’ N, 62 ° 09 ’ E], 14. VII. 1928, V. Gussakovskij (1 ♀, ZISP); 26. IV. 1917, M. Ryabov (2 ♂, ZISP); 17. VII. 1918, M. Ryabov (1 ♂, ZISP). UKRAINE. Kyiv [50 ° 25 ’ N, 30 ° 26 ’ E], 5. IX. 1928, A. Ushinsky (1 ♀, ZISP); Odesa [46 ° 27 ’ N, 30 ° 41 ’ E], 5. VIII. 1926, Aksinin (1 ♀, ZISP); Poltava Province: Shyshaky District, Jaresky [49 ° 50 ’ N, 33 ° 54 ’ E], 8. VIII. 1925, Fabri (1 ♂, ZISP); 3. IX. 1925, Fabri (1 ♀, ZISP); 21. VII. 1923, M. Belchevsky (1 ♂, 1 ♀, ZISP). UZBEKISTAN. Bukhara Province: near Bukhara [39 ° 49 ’ N, 64 ° 21 ’ E], 6. VII. 1928, V. Gussakovskij (1 ♀, ZISP); Jizzakh Province: Golodnaya Steppe [Mirzachoʻl] [40 ° 30 ’ N, 68 ° 00 ’ E], 16 – 25. V. 1903, G. Jakobson (40 ♂, ZISP); 27 – 29. V. 1903, Grebner (1 ♂, ZISP); Gʻallaorol District, Lalmikor [formerly Kuropatkino] [39 ° 56 ’ N, 67 ° 27 ’ E], 1. VIII. 1930, V. Gussakovskij (1 ♂, ZISP); Karakalpakstan: Nukus [42 ° 28 ’ N, 59 ° 35 ’ E], VI – VII. 1898, K. Rossikov (1 ♀, ZISP); Khorazm Province: Khiva [41 ° 22 ’ N, 60 ° 21 ’ E], 16. VI – 24. VII. 1927, V. Gussakovskij (5 ♂, ZISP); Tashkent Province: Boʻstonliq District, Karankul’ [41 ° 34 ’ N, 69 ° 46 ’ E], 17. V. 1898, I. Heyer (2 ♂, 6 ♀, ZISP); Yangiyoʻl District, near Yangiyoʻl [41 ° 08 ’ N, 69 ° 02 ’ E], 17. VI. 1930, L. Zimin (1 ♂, ZISP).	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFB7FFEBFF7D19527B71DCD1.taxon	distribution	Distribution. Arid and semi-arid areas of southern Palaearctic (Portugal, Spain, Italy, Malta, Croatia, Albania, Greece, Cyprus, Czech Republic, Germany, Poland, Slovakia, Hungary, Ukraine, Romania, Morocco, Algeria, Libya, Egypt, Georgia, Abkhazia, * Azerbaijan, Russia (Belgorod Province, Voronezh Province, * Kherson Province, Ulyanovsk Province, Saratov Province, Rostov Province, Volgograd Province, Kalmykia, Astrakhan Province, Krasnodar Territory, * Stavropol Territory, Dagestan, Crimea, Orenburg Province, Altai Territory, Tuva, Irkutsk Province, Buryatia, Zabaykalsky Territory, Amurskaya Province, Primorsky Territory), Turkey, Israel, Palestine, Iraq, Iran, Turkmenistan, Uzbekistan, Tajikistan, Kyrgyzstan, Kazakhstan, Mongolia, China (Liaoning, Gansu, Beijing, Hebei, Shandong, * Xinjiang, Jiangsu, Zhejiang), North Korea, South Korea), Afrotropic (Angola, Ethiopia, Namibia, Senegal, Somalia, South Africa, Tanzania, Zimbabwe, Oman, Saudi Arabia) and western Indomalayan regions (India).	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFB8FFEDFF7D1ACF790DD885.taxon	description	(Figs 1 f, 4 a, 5 e, 6 e, 10 g, 11 e)	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFB8FFEDFF7D1ACF790DD885.taxon	diagnosis	Diagnosis. Both sexes of P. zarudnyi resemble P. subfuscatus, P. judaeus, and P. crudelis, but differ in having the wings smoky and the body brownish (P. subfuscatus, P. judaeus, and P. crudelis have the wings yellow and the body black).	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFB8FFEDFF7D1ACF790DD885.taxon	description	Description. Body brownish with fine brown erect setae. Head. Labial palpomeres 2 – 4 usual. Vertex, occiput and gena microsculptured, with fine, sparse, inconspicuous, appressed, goldish or brownish setae. Mesosoma. Metanotum tuberculate dorsally. Mesopleuron, metapleuron and propodeum areolate-rugose, shiny; propodeal enclosure dull. Wings smoky; veins light brown; recurrent vein II of forewing received by submarginal cell III. Legs. Claws with 2 ventral teeth; arolia present. Male. Body length 14 – 19 mm. Head. Labrum trapeziform. Clypeus with wide V-shaped median emargination. Clypeus, subantennal sclerite, paraocular area and frons with appressed, silvery setae. Mandible reddish mesally. F 3 – F 5 with narrow, depressed placoids. Mesosoma. Pronotum microsculptured, slightly shiny. Scutum finely transversely rugose anteriorly, longitudinally rugose posteriorly and smooth posterolaterally, slightly shiny. Pronotum and scutum anteriorly with fine, sparse, inconspicuous, appressed, goldish or brownish setae. Scutellum finely longitudinally rugose, dull. Metasoma. S 6 entire, without teeth. Female. Body length 19 – 24 mm. Head. Labrum trapeziform, emarginate apically. Clypeus angulate, with U-shaped median notch. Clypeus, subantennal sclerite, paraocular area and frons with appressed, goldish setae. Mesosoma. Pronotum microsculptured, distinctly longitudinally emarginate dorsally, slightly shiny. Scutum micropunctate, slightly shiny. Pronotum and scutum with fine, sparse, inconspicuous, appressed, goldish or brownish setae. Scutellum micropunctate, finely longitudinally rugose posteriorly, shiny. Legs. Tarsal rake of foreleg with long, stout, bladelike and some short, hairlike setae.	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFB8FFEDFF7D1ACF790DD885.taxon	materials_examined	Material examined. IRAN. Sistan and Baluchestan Province: Iranshahr County, Bazman District, Bazman [27 ° 51 ’ N, 60 ° 10 ’ E], 10. VIII. 1898, N. Zarudny (3 ♂, 1 ♀ paralectotypes, 1 ♀, lectotype, ZISP); 30. VIII. 1898 (1 ♂, paralectotype, ZISP).	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
7F0D87D3FFB8FFEDFF7D1ACF790DD885.taxon	distribution	Distribution. Desert areas of the Iranian Plateau (Iran).	en	Odintsev, Yuriy N. Danilov Oleg A., Kobets, Anastasia S. (2024): A taxonomic study on the genus Prionyx Vander Linden, 1827 (Hymenoptera: Sphecidae: Prionychini). Subgenus Harpactopus F. Smith, 1856. Zootaxa 5551 (2): 263-298, DOI: 10.11646/zootaxa.5551.2.3, URL: https://doi.org/10.11646/zootaxa.5551.2.3
