taxonID	type	description	language	source
7F0B87C4FFA58035FDE2FE3CFACAFD63.taxon	type_taxon	Type species Guaranita goloboffi Huber, 2000.	en	Huber, Bernhard A., Meng, Guanliang, Král, Jiří, Ávila Herrera, Ivalú M., Izquierdo, Matías A. (2023): Revision of the South American Ninetinae genus Guaranita (Araneae, Pholcidae). European Journal of Taxonomy 900 (1): 32-80, DOI: 10.5852/ejt.2023.900.2301, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2301/9981
7F0B87C4FFA58035FDE2FE3CFACAFD63.taxon	diagnosis	Diagnosis Small (body length ~ 1 mm) short-legged pholcids with globular abdomen (Fig. 2), distinguished from most other genera of Ninetinae by dorsal flap on procursus (e. g., Figs 4 F, 9 F, 12 D); from Galapa, which shares a dorsal process on the procursus (cf. Huber 2000: figs 383, 387), by pair of prominent apophyses on male chelicerae (e. g., Figs 4 A – C, 11 A – B; absent in Galapa) and by unmodified fangs of male chelicerae (with processes in Galapa).	en	Huber, Bernhard A., Meng, Guanliang, Král, Jiří, Ávila Herrera, Ivalú M., Izquierdo, Matías A. (2023): Revision of the South American Ninetinae genus Guaranita (Araneae, Pholcidae). European Journal of Taxonomy 900 (1): 32-80, DOI: 10.5852/ejt.2023.900.2301, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2301/9981
7F0B87C4FFA58035FDE2FE3CFACAFD63.taxon	description	Description Male MEASUREMENTS. Total body length 0.9 – 1.1, carapace width 0.4 – 0.5. Legs relatively short, tibia 1 0.5 – 0.6; tibia 1 L / d 7 – 9; leg formula 4 - 1 - 2 - 3; metatarsus 1 shorter than tibia 1 or same length (metatarsus 1 / tibia 1: 0.95 – 1.00); tibia 2 much shorter than tibia 4 (tibia 2 / tibia 4: 0.65 – 0.75). COLOUR. Live specimens reddish brown (Fig. 2); abdomen without or with very indistinct marks; legs without dark or light bands. Color in ethanol similar but paler, ochre-yellow. BODY. Ocular area barely raised, eight eyes (Figs 6 A, 11 A – D, 17 A, 27 A), AME relatively large (diameter: 25 – 30 µm, i. e., 50 – 60 % of PME diameter). Carapace without thoracic groove (Figs 6 A, 11 A – D, 17 A, 27 A). Clypeus usually unmodified, only in G. dobby with distinct median process (Torres et al. 2016: fig. 9). Sternum slightly wider than long, with pair of rounded anterior processes near leg coxae 1, processes apparently without pores. Abdomen globular; four (rarely five) epiandrous spigots arranged in two pairs (Figs 11 E – F, 17 F, 27 F); ALS with seven spigots each (as in female, cf. Figs 6 C, 11 H, 17 B – D, 27 C – E): one strongly widened spigot, one long pointed spigot, and five cylindrical spigots (one of which is unusually large); PMS with two short, pointed spigots (as in female, cf. Figs 6 D, 27 D); PLS without spigots (Figs 17 B, 27 C – D). CHELICERAE. With pair of long frontal apophyses (e. g., Figs 4 A – C, 11 A – B); with stridulatory files on relatively small lateral patches (Figs 12 A, 18 A – B, 27 G), with ~ 15 – 25 stridulatory ridges each. PALPS. Coxa unmodified; trochanter without or with very indistinct ventral projection; femur cylindrical, slightly widened distally, proximally without or with very low retrolateral hump, with prolateral stridulatory pick (modified hair; Fig. 27 H); patella short; tibia oval to globular, with two trichobothria; palpal tarsal organ raised, capsulate (Figs 13 A, 28 A – B), with small opening (diameter of opening ~ 1.1 – 1.5 µm); procursus with distinctive dorsal flap, large semi-transparent ventral membrane, and complex tip bent towards dorsal (e. g., Figs 4 D – F, 12 C – F); genital bulb with simple proximal sclerite, distinct distal (main) sclerite, and variably complex ‘ embolar division’ consisting of membranous and sclerotized elements (Figs 4 G – I, 29). LEGS. Without spines and curved hairs; with ‘ short vertical hairs’ in 1 – 2 rows on tibia 1 (Figs 13 D, 19 A – C, 30 A – B; length of hairs ~ 10 – 15 µm). Trichobothria in usual arrangement: three on each tibia (except tibia 1: prolateral trichobothrium absent), one on each metatarsus, slightly feathered (as in female, cf. Fig. 28 E – F); length of trichobothria ~ 60 µm; retrolateral trichobothrium of tibia 1 in very distal position (at ~ 55 – 65 % of tibia length). Tibiae and metatarsi with tiny pores with cuticular rim (diameter of opening ~ 0.6 µm; as in female, cf. Figs 6 E – F, 19 F, 31 A). Metatarsi 3 and 4 with ~ 1 – 5 slender hairs ventrally (Figs 13 E, 19 G – H, 31 C), with bases as in regular hairs but shafts reminding of trichobothria (i. e., feathered and small proximal diameter: ~ 2 µm; regular leg hair proximal diameter: 3 – 4 µm). Tarsus 1 with 5 – 6 pseudosegments, poorly visible in dissecting microscope; tarsus 4 distally with one comb-hair on prolateral side (as in female, cf. Fig. 31 D); leg tarsal organs very small, not raised, capsulate (Fig. 13 C), with small opening (diameter of opening ~ 0.8 – 1.0 µm); three claws, superior claws with 8 – 11 tines (as in female, cf. Figs 7 F, 13 F – G, 20 D – H, 31 D – F). Female In general, similar to male but chelicerae without stridulatory files (Figs 12 B, 18 C), sternum without pair of anterior humps, palpal tarsal organ only weakly raised (Figs 19 E, 28 C – D), and tibia 1 with usual low number of short vertical hairs; legs either slightly shorter than in males or of same length [only G. goloboffi Huber, 2000, G. munda (Gertsch, 1982), and G. yaculica Huber, 2000 with reasonable sample sizes: male / female tibia 1 length: 1.00 – 1.08]. Spinnerets, leg pores, leg tarsal organs, and combhairs as in male. Main (anterior) epigynal plate usually trapezoidal, only in G. dobby Torres et al., 2016 rather triangular, weakly protruding (e. g., Figs 5 A, 10 A, 16 A); posterior plate simple, short but wide. Internal genitalia very simple, usually with distinct median structure (poorly developed in G. dobby), sometimes with membranous median sac (receptacle?) (e. g., Figs 5 C – D, 10 C – D, 32); apparently with very small pore plates (arrows in Fig. 32). The “ pair of receptacles ” mentioned and illustrated in Torres et al. (2016: 10, fig. 14) is a misinterpretation either of the book lungs or of a pair of silk glands. Relationships The molecular analysis of Eberle et al. (2018) included only a single species of Guaranita (G. yaculica), which was placed (with moderate support) as sister to the South American Ninetinae genera Pemona and Kambiwa. Preliminary analyses of molecular (UCE) data (G. Meng, B. A. Huber, L. Podsiadlowski, unpubl. data) support the close relationship among these three genera and add Galapa to this clade, a genus not included in Eberle et al. (2018). Our new SEM data confirm the position of Guaranita among Ninetinae (in particular the small opening of the tarsal organs; cf. character 57 in Huber 2000). Within Guaranita, our CO 1 data suggest that the morphologically distinct G. dobby is sister to the other species, a topology that is also supported by preliminary analyses of UCE data. Natural history While Guaranita auadae Huber sp. nov., G. dobby, and G. goloboffi were found in relatively arid environments with cacti and low bushes (Fig. 34 A, D – F), G. munda and G. yaculica were collected in dry to humid forests (Fig. 33 B – C). In arid environments, the specimens were collected by turning stones and rocks; in more humid environments by shaking dead bromeliads lying on the ground and by sifting leaf litter. Guaranita munda was collected by turning stones of a loosely built wall situated in a low forest (Fig. 33 B). When disturbed by turning a rock, the spiders ran rapidly a few centimeters over the rock surface but seemed reluctant to drop to the ground. Webs were not seen in the field but the spiders quickly built flimsy webs in small glass vials. We never found more than one species of Guaranita at one locality. Other pholcid spiders sharing the microhabitats of Guaranita were Gertschiola macrostyla (Mello-Leit „ o, 1941) and Nerudia spp. (Ninetinae), and several small undescribed representatives of Modisiminae Simon, 1893. Eggs sacs were carried under the prosoma and contained 5 – 8 eggs arranged in a single layer (Fig. 2 A – B, D, H); they are thus among the smallest egg-sacs known in pholcids (Huber & Eberle 2021).	en	Huber, Bernhard A., Meng, Guanliang, Král, Jiří, Ávila Herrera, Ivalú M., Izquierdo, Matías A. (2023): Revision of the South American Ninetinae genus Guaranita (Araneae, Pholcidae). European Journal of Taxonomy 900 (1): 32-80, DOI: 10.5852/ejt.2023.900.2301, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2301/9981
7F0B87C4FFA58035FDE2FE3CFACAFD63.taxon	distribution	Distribution Guaranita is widespread in northern Argentina and reaches into Paraguay and southern Brazil (and probably southern Bolivia and Uruguay) (Fig. 33). It does not seem to cross the Andes into Chile.	en	Huber, Bernhard A., Meng, Guanliang, Král, Jiří, Ávila Herrera, Ivalú M., Izquierdo, Matías A. (2023): Revision of the South American Ninetinae genus Guaranita (Araneae, Pholcidae). European Journal of Taxonomy 900 (1): 32-80, DOI: 10.5852/ejt.2023.900.2301, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2301/9981
7F0B87C4FFA58035FDE2FE3CFACAFD63.taxon	description	Composition The genus now includes five described species, all of which are treated below. Our limited genetic data support the species limits (Table 2): intraspecific K 2 P distances (N = 4) range from 0.2 – 3.0 % (mean 0.9 %); interspecific distances within Guaranita (N = 24) range from 13.6 – 21.7 % (mean 17.1 %).	en	Huber, Bernhard A., Meng, Guanliang, Král, Jiří, Ávila Herrera, Ivalú M., Izquierdo, Matías A. (2023): Revision of the South American Ninetinae genus Guaranita (Araneae, Pholcidae). European Journal of Taxonomy 900 (1): 32-80, DOI: 10.5852/ejt.2023.900.2301, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2301/9981
7F0B87C4FFA98032FDC2FD8EFEE8FD89.taxon	description	Figs 2 A – B, 3 – 7, 32 A	en	Huber, Bernhard A., Meng, Guanliang, Král, Jiří, Ávila Herrera, Ivalú M., Izquierdo, Matías A. (2023): Revision of the South American Ninetinae genus Guaranita (Araneae, Pholcidae). European Journal of Taxonomy 900 (1): 32-80, DOI: 10.5852/ejt.2023.900.2301, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2301/9981
7F0B87C4FFA98032FDC2FD8EFEE8FD89.taxon	diagnosis	Diagnosis (amendments; see Torres et al. 2016) Distinguished from known congeners by median process on male clypeus (cf. Torres et al. 2016: fig. 9; unmodified in congeners), by male cheliceral apophyses (Fig. 4 A – C; short and diverging in distal view), by very small (compared with congeners) dorsal flap on procursus (Fig. 4 F), and by roughly triangular (rather than trapezoidal as in congeners) anterior epigynal plate (Fig. 5 A); also by relatively slender male palpal tibia (Fig. 3 C; width / length 0.75; other species 0.85 – 1.00) and by female internal genitalia (Figs 5 C – D, 32 A; median structure poorly developed compared with congeners); from G. auadae sp. nov. and G. goloboffi also by narrow distal bulbal sclerite (Fig. 4 G).	en	Huber, Bernhard A., Meng, Guanliang, Král, Jiří, Ávila Herrera, Ivalú M., Izquierdo, Matías A. (2023): Revision of the South American Ninetinae genus Guaranita (Araneae, Pholcidae). European Journal of Taxonomy 900 (1): 32-80, DOI: 10.5852/ejt.2023.900.2301, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2301/9981
7F0B87C4FFA98032FDC2FD8EFEE8FD89.taxon	materials_examined	Material examined (new record) ARGENTINA – Salta • 2 ♂♂, 3 ♀♀; ~ 55 km NW of Campo Quijano; 24.4716 ° S, 65.9272 ° W; 3040 m a. s. l.; 19 Mar. 2019; B. A. Huber and M. A. Izquierdo leg.; ZFMK Ar 24121 • 11 ♀♀ in pure ethanol (four prosomata used for molecular work; one female and one female abdomen used for SEM); same collection data as for preceding; ZFMK Arg 187 • 2 ♀♀; same collection data as for preceding; LABRE-Ar 876 • 1 ♀, in pure ethanol; same collection data as for preceding; LABRE-Ar 865.	en	Huber, Bernhard A., Meng, Guanliang, Král, Jiří, Ávila Herrera, Ivalú M., Izquierdo, Matías A. (2023): Revision of the South American Ninetinae genus Guaranita (Araneae, Pholcidae). European Journal of Taxonomy 900 (1): 32-80, DOI: 10.5852/ejt.2023.900.2301, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2301/9981
7F0B87C4FFA98032FDC2FD8EFEE8FD89.taxon	description	Redescription of male (amendments; see Torres et al. 2016) Measurements of male from 55 km NW of Campo Quijano: total body length 1.1 (1.2 with clypeus process), carapace width 0.48; distance PME – PME 45 µm; diameter PME 40 µm; distance PME – ALE 20 µm; distance AME – AME 20 µm; diameter AME 25 µm. Leg 1: 2.33 (0.66 + 0.16 + 0.62 + 0.55 + 0.34), tibia 2: 0.54, tibia 3: 0.48, tibia 4: 0.72; tibia 1 L / d: 9; diameters of leg femora 0.10 – 0.11, of leg tibiae: 0.065. Tibia 1 of second newly collected male: 0.58. Tip of clypeus process straight but at tip with hairs pointing upwards and backwards. Sternum slightly wider than long (0.34 / 0.30). Chelicerae as in Fig. 4 A – C. Pedipalp as in Fig. 3 A – C; tibia with two trichobothria; procursus as in Fig. 4 D – F, with large transparent ventral membrane, distinctive dorsal flap, and tip bent towards dorsal; genital bulb as in Fig. 4 G – I, with simple proximal sclerite and band-like distal sclerite (same width over most of its length). Legs without spines and curved hairs; vertical hairs not seen; retrolateral trichobothrium of tibia 1 at 60 %; prolateral trichobothrium absent on tibia 1, present on other leg tibiae; tarsus 1 with 5 pseudosegments, poorly visible in dissecting microscope. Description of female In general similar to male (Fig. 2 A – B) but clypeus without process, sternum without pair of anterior humps, and chelicerae without stridulatory files. Tibia 1 in seven females: 0.58 – 0.64 (mean 0.62). Epigynum (Figs 5 A, 6 B) with simple triangular anterior plate weakly bulging; posterior plate short and simple. Internal genitalia (Figs 5 C – D, 32 A) very simple, with median sclerotized structure (receptacle?), apparently with small pore plates. Each ALS with one strongly widened spigot, one long pointed spigot, and five cylindrical spigots (of which one is much wider than the others; Fig. 6 C); each PMS with two conical spigots (Fig. 6 D); PLS without spigots. Leg tibiae and metatarsi with tiny pores with cuticular rim (pore diameter 0.6 µm; Fig. 6 E – F) and with small round cuticular ‘ plates’ (diameter 4 – 5 µm; Fig. 6 E). Tarsal organs with very small openings (diameters of openings 0.8 – 0.9 µm; Fig. 7 D – E). Metatarsi 3 and 4 with one long slender hair each on retrolateral side (Fig. 7 C). Remarks (notes on type locality) This species was previously known from two specimens supposedly from two localities in Salta province: the holotype locality, 9 km E of Cabra Corral dam; and a second locality, 1 km N of “ Charrillos ” (should be Chorrillos). Our newly collected specimens of G. dobby are from close to the second locality, in the same river valley, ~ 37 km NW of Chorrillos. However, we failed to find G. dobby at the holotype locality and at several nearby sites E of Cabra Corral dam we visited. Instead, we found G. goloboffi at two sites in that area. Previous collectors also found numerous specimens of G. goloboffi E of Cabra Corral dam (Torres et al. 2015). This sheds doubt on the origin of the G. dobby holotype. We suspect that the holotype specimen is mislabeled but according to José Corronca (pers. com. Jan. 2022) this is unlikely to be the case. Natural history The newly collected specimens were found under rocks in a very arid environment (Fig. 34 A). Egg sacs (N = 4) contained 6 – 8 eggs and were carried in a single layer under the prosoma (Fig. 2 B); egg diameter: 0.46 – 0.48.	en	Huber, Bernhard A., Meng, Guanliang, Král, Jiří, Ávila Herrera, Ivalú M., Izquierdo, Matías A. (2023): Revision of the South American Ninetinae genus Guaranita (Araneae, Pholcidae). European Journal of Taxonomy 900 (1): 32-80, DOI: 10.5852/ejt.2023.900.2301, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2301/9981
7F0B87C4FFA98032FDC2FD8EFEE8FD89.taxon	distribution	Distribution Known from three localities in Argentina, Salta Province (Fig. 33 A); but see Notes on type locality above.	en	Huber, Bernhard A., Meng, Guanliang, Král, Jiří, Ávila Herrera, Ivalú M., Izquierdo, Matías A. (2023): Revision of the South American Ninetinae genus Guaranita (Araneae, Pholcidae). European Journal of Taxonomy 900 (1): 32-80, DOI: 10.5852/ejt.2023.900.2301, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2301/9981
7F0B87C4FFAE8028FDCAFD34FCFDFADA.taxon	description	Figs 2 C – D, 8 – 13, 32 B	en	Huber, Bernhard A., Meng, Guanliang, Král, Jiří, Ávila Herrera, Ivalú M., Izquierdo, Matías A. (2023): Revision of the South American Ninetinae genus Guaranita (Araneae, Pholcidae). European Journal of Taxonomy 900 (1): 32-80, DOI: 10.5852/ejt.2023.900.2301, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2301/9981
7F0B87C4FFAE8028FDCAFD34FCFDFADA.taxon	diagnosis	Diagnosis (amendments; see Huber 2000) Distinguished from known congeners by size and shape of dorsal flap on procursus (Fig. 9 F; larger than in congeners; distally widened) and by female internal genitalia (Fig. 10 C – D; large membranous median sac; lateral elements medially curved, creating median posterior indentation also sometimes visible in uncleared epigyna); from G. auadae sp. nov. and G. goloboffi also by narrow distal bulbal sclerite (Fig. 9 G); from most congeners (except G. dobby) also by relatively long male palpal femur (Fig. 8 C; length / width 2.50 – 2.55, most other species 1.85 – 2.25, G. dobby 2.50).	en	Huber, Bernhard A., Meng, Guanliang, Král, Jiří, Ávila Herrera, Ivalú M., Izquierdo, Matías A. (2023): Revision of the South American Ninetinae genus Guaranita (Araneae, Pholcidae). European Journal of Taxonomy 900 (1): 32-80, DOI: 10.5852/ejt.2023.900.2301, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2301/9981
7F0B87C4FFAE8028FDCAFD34FCFDFADA.taxon	materials_examined	Material examined (new records) ARGENTINA – Córdoba • 6 ♂♂, 5 ♀♀ (one male and one female used for SEM); ~ 2.5 km E of Nono; 31.8025 ° S, 64.9762 ° W; 915 m a. s. l.; 2 Mar. 2019; B. A. Huber and M. A. Izquierdo leg.; ZFMK Ar 24122 • 7 ♀♀, in pure ethanol (three prosomata used for molecular work; one female used for SEM); same collection data as for preceding; ZFMK Arg 127 • 3 ♂♂, 2 ♀♀; same collection data as for preceding; LABRE-Ar 877 • 3 ♂♂; same collection data as for preceding; LABRE-Ar 878 • 3 ♀♀, in pure ethanol; same collection data as for preceding; LABRE-Ar 882, 883, 856 • 1 ♀, in pure ethanol; ~ 1.5 km E of Nono; 31.7980 ° S, 64.9877 ° W; 895 m a. s. l.; 2 Mar. 2019; B. A. Huber and M. A. Izquierdo leg.; ZFMK Arg 126 • 2 ♂♂, 1 ♀, 2 juvs; Villa La Merced; 31.8397 ° S, 64.5249 ° W; 765 m a. s. l.; 17 Dec. 2019; Izquierdo and Palen Pietri leg.; litter and bark of Eucalyptus plantation; LABRE-Ar 873 • 1 ♂; Villa La Merced; 31.8419 ° S, 64.5240 ° W; 775 m a. s. l.; 27 Jan. 2020; Izquierdo, Abregú, and Palen Pietri leg.; LABRE-Ar 874 • 4 ♀♀, some juvs; same collection data as for preceding; LABRE-Ar 626. – Entre Ríos • 5 ♂♂, 3 ♀♀, 2 juvs (one male used for SEM); Dept. Colón, Parque Nacional El Palmar; 31.8653 ° S, 58.2375 ° W; 20 m a. s. l.; 6 – 8 Aug. 2011; M. J. Ramírez et al. leg.; MACN Ar 32745 • 2 ♂♂, 4 ♀♀, 2 juvs; same collection data as for preceding; MACN Ar 32741 • 1 ♂, 1 ♀; same collection data as for preceding; MACN Ar 32744 • 1 ♂; same collection data as for preceding, with label “ muestra de tejido prep. CJG- 3350 ”; MACN Ar 32743 • 1 ♀; Dept. Colón, Parque Nacional El Palmar, Arroyo El Palmar; 31.8931 ° S, 58.2385 ° W; 10 m a. s. l.; 7 Aug. 2011; M. J. Ramírez et al. leg.; MACN Ar 32742 • 1 ♂; Dept. Colón, Parque Nacional El Palmar, Sector Sur; 31.8877 ° S, 58.3119 ° W; 30 m a. s. l.; 7 Aug. 2011; M. J. Ramírez et al. leg.; MACN Ar 32740 • 1 ♀; Parque Nacional El Palmar (no precise locality information); 22 – 23 Nov. 2003; C. Grismado, A. Ojanguren and F. Labarque leg.; MACN Ar 25453 • 1 ♀; Villa Urquiza; ~ 31.65 ° S, 60.38 ° W (no precise locality information); 17 Feb. 1988; P. Goloboff and C. Szumik leg.; MACN Ar 20030.	en	Huber, Bernhard A., Meng, Guanliang, Král, Jiří, Ávila Herrera, Ivalú M., Izquierdo, Matías A. (2023): Revision of the South American Ninetinae genus Guaranita (Araneae, Pholcidae). European Journal of Taxonomy 900 (1): 32-80, DOI: 10.5852/ejt.2023.900.2301, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2301/9981
7F0B87C4FFAE8028FDCAFD34FCFDFADA.taxon	description	Redescription (amendments; see Huber 2000) Measurements of male from E of Nono: total body length 1.06, carapace width 0.42; distance PME – PME 40 µm; diameter PME 50 µm; distance PME – ALE 15 µm; distance AME – AME 15 µm; diameter AME 30 µm. Leg 1: 1.98 (0.52 + 0.14 + 0.50 + 0.50 + 0.32), tibia 2: 0.42, tibia 3: 0.38, tibia 4: 0.60; tibia 1 L / d: 8; diameters of leg femora 0.09; of leg tibiae: 0.06. Tibia 1 in 25 males (incl. holotype): 0.49 – 0.58 (mean 0.53). Sternum slightly wider than long (0.33 / 0.29). Chelicerae as in Fig. 9 A – C; stridulatory files (Fig. 12 A) with ~ 15 – 17 ridges each; distances between ridges proximally ~ 1.0 µm, distally ~ 2.1 µm. Pedipalp as in Fig. 8 A – C; tibia with two trichobothria; palpal tarsal organ capsulate (Fig. 13 A) with small opening (diameter of opening 1.15 µm); procursus as in Fig. 9 D – F, with large transparent ventral membrane, distinctive dorsal flap, and tip bent towards dorsal; genital bulb as in Fig. 9 G – I, with simple proximal sclerite, distal sclerite short and simple, not widened in mid-section. Legs without spines and curved hairs; vertical hairs not seen in dissecting microscope but present on tibia 1 (Fig. 13 D), apparently only one row; prolateral trichobothrium absent on tibia 1, present on other leg tibiae; metatarsi 3 and 4 with few (3 – 5) slender hairs proximally on retrolateral-ventral side (Fig. 13 E). Gonopore with 4 – 5 epiandrous spigots (Fig. 11 E – F); spinnerets as in female (see below). Tibia 1 in 22 females: 0.48 – 0.58 (mean 0.54). Female chelicerae without stridulatory ridges (Fig. 12 B). Female internal genitalia with strong median structure and membranous sac (receptacle?) (Fig. 10 C – D); apparently with small pore plates (Fig. 32 B). Each ALS (Fig. 11 H) with one strongly widened spigot, one long pointed spigot, and five cylindrical spigots (of which one is much wider than the others); each PMS with two conical spigots; PLS without spigots. Leg tarsal organs with very small openings (diameters of openings 0.8 – 0.9 µm; Fig. 13 B). Metatarsi 3 and 4 with long slender hairs as in male; tarsus 4 with single prolateral comb-hair as in male.	en	Huber, Bernhard A., Meng, Guanliang, Král, Jiří, Ávila Herrera, Ivalú M., Izquierdo, Matías A. (2023): Revision of the South American Ninetinae genus Guaranita (Araneae, Pholcidae). European Journal of Taxonomy 900 (1): 32-80, DOI: 10.5852/ejt.2023.900.2301, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2301/9981
7F0B87C4FFAE8028FDCAFD34FCFDFADA.taxon	discussion	Remarks (notes on type locality) The type locality of this species has been confused twice. Gertsch (1982) interpreted the label information as referring to Cerro Colorado in Nuevo León, Mexico. Later, Huber (2000), read the handwritten label as “ Crro Colorado, Cta., 14. X- 61, Col: O. de Ferrariis ”, and suggested that this referred to Cerro Colorado in the province of Catamarca (“ Cta. ”), Argentina, i. e. ~ 28.46 ° S, 65.85 ° W. Another interpretation for a label accompanying a specimen of the linyphiid Scolecura propinqua Millidge, 1991 collected by O. de Ferrariis on the same day, was offered by Miller (2007): Cerro Colorado in the province of Córdoba, i. e. ~ 30.10 ° S, 63.93 ° W. A new look at both labels confirms Miller’s (2007) interpretation: the label in the type vial of Guaranita munda quite clearly reads “ Cba. ” rather than “ Cta. ”, and the machinewritten label accompanying the Scolecura propinqua specimen explicitly says “ Prov. Cordoba ”.	en	Huber, Bernhard A., Meng, Guanliang, Král, Jiří, Ávila Herrera, Ivalú M., Izquierdo, Matías A. (2023): Revision of the South American Ninetinae genus Guaranita (Araneae, Pholcidae). European Journal of Taxonomy 900 (1): 32-80, DOI: 10.5852/ejt.2023.900.2301, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2301/9981
7F0B87C4FFAE8028FDCAFD34FCFDFADA.taxon	biology_ecology	Natural history Near Nono, the spiders were collected by turning the uppermost rocks of a stone wall in a low forest (Fig. 34 B). The spiders started to run rapidly but did not drop from the rocks. A label accompanying specimens from Parque Nacional El Palmar suggests a very similar habitat: “ piedras palmeras con pastizal y bosque bajo ”. Two egg-sacs contained 6 and 7 eggs, respectively, and were carried under the prosoma; egg diameter: 0.44.	en	Huber, Bernhard A., Meng, Guanliang, Král, Jiří, Ávila Herrera, Ivalú M., Izquierdo, Matías A. (2023): Revision of the South American Ninetinae genus Guaranita (Araneae, Pholcidae). European Journal of Taxonomy 900 (1): 32-80, DOI: 10.5852/ejt.2023.900.2301, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2301/9981
7F0B87C4FFAE8028FDCAFD34FCFDFADA.taxon	distribution	Distribution Widely distributed in north-eastern Argentina, reaching Rio Grande do Sul (Brazil) (Fig. 33 A). Presumably also present in Uruguay and southern Paraguay. The single record from Jujuy (Torres et al. 2016) appears dubious (misidentified G. yaculica?).	en	Huber, Bernhard A., Meng, Guanliang, Král, Jiří, Ávila Herrera, Ivalú M., Izquierdo, Matías A. (2023): Revision of the South American Ninetinae genus Guaranita (Araneae, Pholcidae). European Journal of Taxonomy 900 (1): 32-80, DOI: 10.5852/ejt.2023.900.2301, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2301/9981
7F0B87C4FFB48025FDDFFA79FB77FE36.taxon	description	Figs 2 E – F, 14 – 20, 32 C	en	Huber, Bernhard A., Meng, Guanliang, Král, Jiří, Ávila Herrera, Ivalú M., Izquierdo, Matías A. (2023): Revision of the South American Ninetinae genus Guaranita (Araneae, Pholcidae). European Journal of Taxonomy 900 (1): 32-80, DOI: 10.5852/ejt.2023.900.2301, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2301/9981
7F0B87C4FFB48025FDDFFA79FB77FE36.taxon	diagnosis	Diagnosis (amendments; see Huber 2000; Torres et al. 2016) Distinguished from known congeners by size and shape of dorsal flap on procursus (Fig. 15 F; rounded, larger than in the similar G. goloboffi) and by female internal genitalia (Fig. 16 C – D; membranous median sac, similar to G. munda but smaller; lateral elements straight, not curved as in G. munda); from G. auadae sp. nov. and G. goloboffi also by narrower distal bulbal sclerite (Fig. 15 G).	en	Huber, Bernhard A., Meng, Guanliang, Král, Jiří, Ávila Herrera, Ivalú M., Izquierdo, Matías A. (2023): Revision of the South American Ninetinae genus Guaranita (Araneae, Pholcidae). European Journal of Taxonomy 900 (1): 32-80, DOI: 10.5852/ejt.2023.900.2301, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2301/9981
7F0B87C4FFB48025FDDFFA79FB77FE36.taxon	materials_examined	Material examined (new records) ARGENTINA – Jujuy • 2 ♂♂, 2 ♀♀, 1 juv.; Calilegua National Park, Guaraní trail, near camping area; 23.7612 ° S, 64.8517 ° W; 620 m a. s. l.; 15 Mar. 2019; B. A. Huber and M. A. Izquierdo leg.; ZFMK Ar 24123 • 2 ♀♀, in pure ethanol; same collection data as for preceding; LABRE-Ar 515 • 6 ♂♂, 3 ♀♀ (one male used for SEM); Calilegua National Park, ~ 1 km NW of headquarters; 23.7540 ° S, 64.8537 ° W; 710 m a. s. l.; 15 Mar. 2019; B. A. Huber and M. A. Izquierdo leg.; ZFMK Ar 24124 • 14 ♀♀, in pure ethanol (four prosomata used for molecular work; two females used for SEM); same collection data as for preceding; ZFMK Arg 175 • 4 ♂♂, 7 ♀♀, 1 juv.; same collection data as for preceding; LABRE-Ar 514 • 7 ♀♀, 1 juv.; same collection data as for preceding; LABRE-Ar 520 • 1 ♂, 2 ♀♀; Calilegua National Park, Seccional Aguas Negras; 23.7619 ° S, 64.8514 ° W; 605 m a. s. l.; 6 – 11 Dec. 2008; C. Grismado et al. leg.; MACN Ar 22134 • 1 ♀, in pure ethanol; same collection data as for preceding; MACN Ar 34688 • 1 ♂, 1 ♀; Calilegua National Park, entrance area; ~ 23.76 ° S, 64.85 ° W; ~ 620 m a. s. l.; 23 – 24 Sep. 1995; M. Ramírez, P. Goloboff and C. Szumik leg.; MACN Ar 19977 • 1 ♂, 2 juvs; Calilegua National Park, no precise locality information; 22 Dec. 1994; C. Grismado leg.; MACN Ar 19976 • 3 ♀♀; Calilegua National Park, Aguas Negras at ~ 1100 m, no precise locality information; 5 – 7 Aug. 1997; M. Ramírez and L. Compagnucci leg.; MACN Ar 19978, 19981. PARAGUAY – Boquerón • 1 ♂; Enciso, “ T 88.09.0 r 1 ”; 21.2061 ° S, 61.6575 ° W; 255 m a. s. l.; 3 Nov. 2001; M. Leponce leg.; IRSNB • 1 ♂ prosoma; Enciso, “ T 90.09.0 r 1 ”; 21.1998 ° S, 61.6608 ° W; 255 m a. s. l.; 4 Nov. 2001; M. Leponce leg.; IRSNB • 2 ♂♂; same collection data as for preceding, “ T 90.14.0 r 1 ”; IRSNB • 1 ♀; same collection data as for preceding, “ T 90.12.0 r 1 ”; IRSNB.	en	Huber, Bernhard A., Meng, Guanliang, Král, Jiří, Ávila Herrera, Ivalú M., Izquierdo, Matías A. (2023): Revision of the South American Ninetinae genus Guaranita (Araneae, Pholcidae). European Journal of Taxonomy 900 (1): 32-80, DOI: 10.5852/ejt.2023.900.2301, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2301/9981
7F0B87C4FFB48025FDDFFA79FB77FE36.taxon	description	Redescription (amendments; see Huber 2000, Torres et al. 2016) Measurements of male from Calilegua National Park: total body length 0.98, carapace width 0.45; distance PME – PME 45 µm; diameter PME 45 µm; distance PME – ALE 20 µm; distance AME – AME 20 µm; diameter AME 25 µm. Leg 1: 2.20 (0.62 + 0.14 + 0.56 + 0.54 + 0.34), tibia 2: 0.46, tibia 3: 0.40, tibia 4: 0.68; tibia 1 L / d: 9; diameters of leg femora 0.10; of leg tibiae: 0.06. Tibia 1 in 16 males (incl. holotype): 0.50 – 0.62 (mean 0.57). Sternum slightly wider than long (0.33 / 0.31). Chelicerae as in Fig. 15 A – C, 18 A; stridulatory files with ~ 17 – 23 ridges; distances between ridges proximally ~ 0.6 µm, distally ~ 2.3 µm (Fig. 18 B). Pedipalp as in Fig. 14 A – C; tibia with two trichobothria; palpal tarsal organ capsulate, with small opening; procursus as in Fig. 15 D – F and 18 D – F, with large transparent ventral membrane, distinctive dorsal flap, and tip bent towards dorsal; genital bulb as in Figs 15 G – I and 18 D – F, with simple proximal sclerite, distal sclerite not widened in mid-section. Legs without spines and curved hairs; vertical hairs not seen in dissecting microscope but present on tibia 1 (Fig. 19 A – C), apparently in two rows (one prolateral and one retrolateral); prolateral trichobothrium absent on tibia 1, present on other leg tibiae; metatarsi 3 and 4 with few (1 – 3) slender hairs proximally on retrolateral-ventral side (Fig. 19 H). Gonopore with four epiandrous spigots (Fig. 17 F); spinnerets as in female (Fig. 17 D; see below). Tibia 1 in 33 females: 0.48 – 0.64 (mean 0.55). Female chelicerae without stridulatory ridges (Fig. 18 C). Female internal genitalia with median membranous sac (receptacle?) (Fig. 16 C – D); apparently with small pore plates (Fig. 32 C). Each ALS (Fig. 17 B – C) with one strongly widened spigot, one long pointed spigot, and five cylindrical spigots (of which one is much wider than the others); each PMS with two conical spigots; PLS without spigots. Palpal tarsal organ capsulate with small opening (diameter of opening 1.1 µm); leg tarsal organs with very small openings (diameters 0.7 – 0.9 µm; Fig. 20 A – C). Metatarsi 3 and 4 with long slender hairs as in male (Fig. 19 G).	en	Huber, Bernhard A., Meng, Guanliang, Král, Jiří, Ávila Herrera, Ivalú M., Izquierdo, Matías A. (2023): Revision of the South American Ninetinae genus Guaranita (Araneae, Pholcidae). European Journal of Taxonomy 900 (1): 32-80, DOI: 10.5852/ejt.2023.900.2301, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2301/9981
7F0B87C4FFB48025FDDFFA79FB77FE36.taxon	biology_ecology	Natural history At Calilegua National Park, the spiders were collected in forest leaf litter (Fig. 34 C). Two egg-sacs contained five and six eggs, respectively.	en	Huber, Bernhard A., Meng, Guanliang, Král, Jiří, Ávila Herrera, Ivalú M., Izquierdo, Matías A. (2023): Revision of the South American Ninetinae genus Guaranita (Araneae, Pholcidae). European Journal of Taxonomy 900 (1): 32-80, DOI: 10.5852/ejt.2023.900.2301, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2301/9981
7F0B87C4FFB48025FDDFFA79FB77FE36.taxon	distribution	Distribution Most known records are from northern Argentina and north-eastern Paraguay (Fig. 33 B). The single record from Corrientes in Torres et al. (2015) is dubious (misidentified G. munda?).	en	Huber, Bernhard A., Meng, Guanliang, Král, Jiří, Ávila Herrera, Ivalú M., Izquierdo, Matías A. (2023): Revision of the South American Ninetinae genus Guaranita (Araneae, Pholcidae). European Journal of Taxonomy 900 (1): 32-80, DOI: 10.5852/ejt.2023.900.2301, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2301/9981
7F0B87C4FFBC8023FDD5FECFFC7DFC8B.taxon	description	urn: lsid: zoobank. org: act: 8409459 A-BDA 1 - 4650 - 9765 - 44 EA 30 ABDECF Figs 2 G – H, 21 – 23, 32 D	en	Huber, Bernhard A., Meng, Guanliang, Král, Jiří, Ávila Herrera, Ivalú M., Izquierdo, Matías A. (2023): Revision of the South American Ninetinae genus Guaranita (Araneae, Pholcidae). European Journal of Taxonomy 900 (1): 32-80, DOI: 10.5852/ejt.2023.900.2301, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2301/9981
7F0B87C4FFBC8023FDD5FECFFC7DFC8B.taxon	diagnosis	Diagnosis Distinguished from known congeners by shape of dorsal flap on procursus (Fig. 22 F; distally narrow and curved); also by wider distal bulbal sclerite (Fig. 22 G; similar only in G. goloboffi), by relatively short male palpal femur (Fig. 21 C; length / width 1.9; other species 2.1 – 2.6) and by female internal genitalia (Fig. 23 C – D; median structure rectangular, similar to G. goloboffi but smaller).	en	Huber, Bernhard A., Meng, Guanliang, Král, Jiří, Ávila Herrera, Ivalú M., Izquierdo, Matías A. (2023): Revision of the South American Ninetinae genus Guaranita (Araneae, Pholcidae). European Journal of Taxonomy 900 (1): 32-80, DOI: 10.5852/ejt.2023.900.2301, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2301/9981
7F0B87C4FFBC8023FDD5FECFFC7DFC8B.taxon	etymology	Etymology The species name honors Ángela Auad (1945 – 1977), an Argentine social activist who worked with the Mothers of the Plaza de Mayo until she was kidnapped, tortured and murdered.	en	Huber, Bernhard A., Meng, Guanliang, Král, Jiří, Ávila Herrera, Ivalú M., Izquierdo, Matías A. (2023): Revision of the South American Ninetinae genus Guaranita (Araneae, Pholcidae). European Journal of Taxonomy 900 (1): 32-80, DOI: 10.5852/ejt.2023.900.2301, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2301/9981
7F0B87C4FFBC8023FDD5FECFFC7DFC8B.taxon	materials_examined	Type material Holotype ARGENTINA – Jujuy • ♂; between San Salvador and Purmamarca, ‘ site 2 ’; 23.8849 ° S, 65.4613 ° W; 2150 m a. s. l.; 16 – 17 Mar. 2019; B. A. Huber and M. A. Izquierdo leg.; LABRE-Ar 1016. Paratypes ARGENTINA • 1 ♂, 2 ♀♀; same collection data as for holotype; ZFMK Ar 24125 • 2 ♂♂, 7 ♀♀ (together with 11 juvs); same collection data as for holotype; LABRE-Ar 880. Other material examined ARGENTINA – Jujuy • 6 ♀♀, 1 juv., in pure ethanol (two female prosomata used for molecular work, two cleared female genitalia transferred to ZFMK Ar 24125); same collection data as for holotype; ZFMK Arg 179 • 3 ♀♀, in pure ethanol; same collection data as for holotype; LABRE-Ar 867 • 1 ♀, with 6 eggs, in pure ethanol; same collection data as for holotype; LABRE-Ar 866.	en	Huber, Bernhard A., Meng, Guanliang, Král, Jiří, Ávila Herrera, Ivalú M., Izquierdo, Matías A. (2023): Revision of the South American Ninetinae genus Guaranita (Araneae, Pholcidae). European Journal of Taxonomy 900 (1): 32-80, DOI: 10.5852/ejt.2023.900.2301, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2301/9981
7F0B87C4FFBC8023FDD5FECFFC7DFC8B.taxon	description	Description Male (holotype) MEASUREMENTS. Total body length 0.97, carapace width 0.42. Distance PME – PME 40 µm; diameter PME 40 µm; distance PME – ALE 20 µm; distance AME – AME 25 µm; diameter AME 25 µm. Leg 1: 2.02 (0.58 + 0.14 + 0.50 + 0.48 + 0.32), tibia 2: 0.40, tibia 3: 0.36, tibia 4: 0.60; tibia 1 L / d: 7; diameters of leg femora 0.095, of leg tibiae: 0.07. COLOUR (in ethanol). Prosoma and legs ochre-yellow, legs without darker rings; abdomen ochre-grey with indistinct internal marks. BODY (Fig. 2 G). Ocular area barely raised. Carapace without thoracic groove. Clypeus unmodified. Sternum slightly wider than long (0.34 / 0.31), with pair of rounded anterior processes near coxae 1. Abdomen globular. CHELICERAE (Fig. 22 A – C). With pair of long frontal apophyses; with stridulatory files poorly visible in dissecting microscope. PALPS (Fig. 21 A – C). Coxa unmodified; trochanter without process; femur proximally with prolateral stridulatory pick, distally widened but simple; femur-patella joints slightly shifted towards prolateral side; tibia globular, with two trichobothria; tibia-tarsus joints not shifted to one side; procursus as in Fig. 22 D – F, with dorsal flap curved towards distal, large transparent ventral membrane, tip of procursus bent towards dorsal; genital bulb as in Fig. 22 G – I, with simple proximal sclerite, distal sclerite wide, narrowing distally. LEGS. Without spines and curved hairs; vertical hairs not seen; trichobothria of tibia 1 not seen; tarsus 1 with 5 – 6 pseudosegments, poorly visible in dissecting microscope. VARIATION (male). Tibia 1 in three other males: 0.51, 0.52, 0.55. Female In general similar to male (Fig. 2 H) but sternum without pair of anterior humps, and chelicerae apparently without stridulatory files. Tibia 1 in 16 females: 0.50 – 0.60 (mean 0.56). Epigynum (Fig. 23 A) with simple trapezoidal anterior plate; posterior plate short and simple. Internal genitalia (Fig. 23 C – D) very simple, with median sclerotized structure (receptacle?), apparently with small pore plates (Fig. 32 D).	en	Huber, Bernhard A., Meng, Guanliang, Král, Jiří, Ávila Herrera, Ivalú M., Izquierdo, Matías A. (2023): Revision of the South American Ninetinae genus Guaranita (Araneae, Pholcidae). European Journal of Taxonomy 900 (1): 32-80, DOI: 10.5852/ejt.2023.900.2301, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2301/9981
7F0B87C4FFBC8023FDD5FECFFC7DFC8B.taxon	biology_ecology	Natural history The spiders were found under rocks on an arid slope (Fig. 34 D). The habitat was shared with another species of Ninetinae, Nerudia colina Huber, 2023. Two egg-sacs contained six and eight eggs, respectively; egg diameter: 0.36.	en	Huber, Bernhard A., Meng, Guanliang, Král, Jiří, Ávila Herrera, Ivalú M., Izquierdo, Matías A. (2023): Revision of the South American Ninetinae genus Guaranita (Araneae, Pholcidae). European Journal of Taxonomy 900 (1): 32-80, DOI: 10.5852/ejt.2023.900.2301, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2301/9981
7F0B87C4FFBC8023FDD5FECFFC7DFC8B.taxon	distribution	Distribution Known from type locality only, in Argentina, Jujuy (Fig. 33 B).	en	Huber, Bernhard A., Meng, Guanliang, Král, Jiří, Ávila Herrera, Ivalú M., Izquierdo, Matías A. (2023): Revision of the South American Ninetinae genus Guaranita (Araneae, Pholcidae). European Journal of Taxonomy 900 (1): 32-80, DOI: 10.5852/ejt.2023.900.2301, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2301/9981
7F0B87C4FFBF8015FDD2FC29FB89FE91.taxon	description	Figs 2 I – J, 24 – 31, 32 E	en	Huber, Bernhard A., Meng, Guanliang, Král, Jiří, Ávila Herrera, Ivalú M., Izquierdo, Matías A. (2023): Revision of the South American Ninetinae genus Guaranita (Araneae, Pholcidae). European Journal of Taxonomy 900 (1): 32-80, DOI: 10.5852/ejt.2023.900.2301, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2301/9981
7F0B87C4FFBF8015FDD2FC29FB89FE91.taxon	diagnosis	Diagnosis (amendments; see Huber 2000) Distinguished from known congeners by shape of dorsal flap on procursus (Fig. 25 F; rounded, smaller than in the similar G. yaculica); also by wide distal bulbal sclerite (Fig. 25 G; similar only in G. auadae sp. nov.), by relatively wide male palpal tibia (Fig. 24 C; width / length 1.00; other species 0.85 – 0.95; tibia width / femur width: 1.75 – 1.80; other species 1.40 – 1.70) and by female internal genitalia (Fig. 26 C – D; median structure rectangular, similar to G. auadae but larger).	en	Huber, Bernhard A., Meng, Guanliang, Král, Jiří, Ávila Herrera, Ivalú M., Izquierdo, Matías A. (2023): Revision of the South American Ninetinae genus Guaranita (Araneae, Pholcidae). European Journal of Taxonomy 900 (1): 32-80, DOI: 10.5852/ejt.2023.900.2301, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2301/9981
7F0B87C4FFBF8015FDD2FC29FB89FE91.taxon	materials_examined	Material examined (new records) ARGENTINA – Salta • 1 ♂, 1 ♀; ~ 1 km SW of Alemanía; 25.6300 ° S, 65.6180 ° W; 1210 m a. s. l.; 23 Mar. 2019; B. A. Huber and M. A. Izquierdo leg.; ZFMK Ar 24126 • 1 ♀, 3 juvs, in pure ethanol; same collection data as for preceding; ZFMK Arg 203 • 4 ♀♀, 4 juvs, in pure ethanol; same collection data as for preceding; LABRE-Ar 860 • 1 ♀; same collection data as for preceding; LABRE-Ar 861 • 1 ♀; ~ 5 km W of Cafayate, ‘ site 1 ’; 26.0641 ° S, 66.0294 ° W; 2060 m a. s. l.; 24 Mar. 2019; B. A. Huber and M. A. Izquierdo leg.; ZFMK Ar 24127 • 2 ♀♀, in pure ethanol; same collection data as for preceding; LABRE-Ar 857 • 1 ♀, in pure ethanol; same collection data as for preceding; LABRE-Ar 858 • 1 ♀, 1 juv.; 6 km NW of Cafayate, Chuscha; ~ 26.04 ° S, 66.02 ° W; ~ 1980 m a. s. l.; 17 Jul. 1995; M. Ramírez and P. Goloboff leg; MACN Ar 20094 • 1 ♂, 2 ♀♀; Cabra Corral, ‘ site 1 ’, ~ 5 km E of Coronel Moldes; 25.2870 ° S, 65.4238 ° W; 1080 m a. s. l.; 20 Mar. 2019; B. A. Huber and M. A. Izquierdo leg.; ZFMK Ar 24128 • 2 ♀♀, 3 juvs, in pure ethanol; same collection data as for preceding; ZFMK Arg 190 • 1 ♀, same collection data as for preceding; LABRE-Ar 881 • 6 ♀♀, 1 juv., in pure ethanol; same collection data as for preceding; LABRE-Ar 864 • 5 ♂♂, 3 ♀♀ (one male and two females used for µ-CT study; one male used for karyotype study); Cabra Corral, ‘ site 3 ’, ~ 3.5 km SE of dam; 25.2907 ° S, 65.3057 ° W; 1000 m a. s. l.; 21 Mar. 2019; B. A. Huber and M. A. Izquierdo leg.; ZFMK Ar 24129 • 4 ♀♀, 15 juvs, in pure ethanol; same collection data as for preceding; ZFMK Arg 196 • 1 ♂, 1 ♀; same collection data as for preceding; LABRE-Ar 855 • 3 ♀♀, 4 juvs, in pure ethanol; same collection data as for preceding; LABRE-Ar 863. – Catamarca • 8 ♂♂, 4 ♀♀ (two males and two females used for µ-CT study; two males used for karyotype study, one male used for SEM); ~ 5 km NW of Chumbicha, near Balneario El Caolín, ‘ site 1 ’; 28.8152 ° S, 66.2478 ° W; 610 m a. s. l.; 28 – 29 Mar. 2019; B. A. Huber and M. A. Izquierdo leg.; ZFMK Ar 24130 • 1 ♂, 17 ♀♀, 5 juvs, in pure ethanol (two females used for SEM); same collection data as for preceding; ZFMK Arg 220 • 8 ♂♂, 2 juvs; same collection data as for preceding; LABRE-Ar 875 • 11 ♀♀, 18 juvs, in pure ethanol; same collection data as for preceding; LABRE Ar 859. Assigned tentatively (no males available) ARGENTINA – Tucumán • 2 ♀♀, 1 juv., in pure ethanol; San Miguel de Tucumán, Parque 9 de Julio; 26.828 ° S, 65.186 ° W; 430 m a. s. l.; 1 Apr. 2015; A. Porta leg.; MACN Ar 34678. – Salta • 3 ♀♀; between Alemanía and Cafayate; 25.7023 ° S, 65.7022 ° W; 1340 m a. s. l.; 23 Mar. 2019; B. A. Huber and M. A. Izquierdo leg.; LABRE-Ar 862.	en	Huber, Bernhard A., Meng, Guanliang, Král, Jiří, Ávila Herrera, Ivalú M., Izquierdo, Matías A. (2023): Revision of the South American Ninetinae genus Guaranita (Araneae, Pholcidae). European Journal of Taxonomy 900 (1): 32-80, DOI: 10.5852/ejt.2023.900.2301, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2301/9981
7F0B87C4FFBF8015FDD2FC29FB89FE91.taxon	description	Redescription (amendments; see Huber 2000) Measurements of male from Cabra Corral, ‘ site 3 ’: total body length 1.08, carapace width 0.40; distance PME – PME 40 µm; diameter PME 45 µm; distance PME – ALE 20 µm; distance AME – AME 20 µm; diameter AME 25 µm. Leg 1: 2.02 (0.56 + 0.14 + 0.50 + 0.48 + 0.34), tibia 2: 0.42, tibia 3: 0.38, tibia 4: 0.63; tibia 1 L / d: 8; diameters of leg femora 0.090 – 0.095; of leg tibiae: 0.060. Tibia 1 in 19 males (incl. males in Huber 2000): 0.49 – 0.59 (mean 0.53). Sternum slightly wider than long (0.32 / 0.30). Chelicerae as in Fig. 25 A – C; stridulatory files (Fig. 27 G) with ~ 17 – 19 ridges each; distances between ridges proximally ~ 0.6 µm, distally ~ 2.7 µm. Pedipalp as in Fig. 24 A – C; tibia with two trichobothria; palpal tarsal organ capsulate (Fig. 28 A – B), raised, with small opening (diameter of opening 1.45 µm); procursus as in Figs 25 D – F and 29 A – C, with large transparent ventral membrane, distinctive dorsal flap, and tip bent towards dorsal; genital bulb as in Figs 25 G – I and 29 A – F, with simple proximal sclerite, distal sclerite widened in mid-section. Legs without spines and curved hairs; vertical hairs not seen in dissecting microscope but present in two retrolateral rows on tibia 1 (Fig. 30 A – B); prolateral trichobothrium absent on tibia 1, present on other leg tibiae; metatarsus 4 with a few slender hairs on retrolateral-ventral side (as in female, cf. Fig. 30 C); tarsus 4 with single prolateral comb-hair (as in female, cf. Fig. 31 D). Gonopore with four epiandrous spigots (Fig. 27 F). Tibia 1 in 55 newly collected females 0.48 – 0.58 (mean 0.52). Female internal genitalia (Fig. 26 C – D) with strong median structure; apparently with small pore plates (Fig. 32 E). Each ALS (Fig. 27 C – E) with one strongly widened spigot, one long pointed spigot, and five cylindrical spigots (of which one is much wider than the others); each PMS with two conical spigots (Fig. 27 D); PLS without spigots. Leg tibiae and metatarsi with tiny pores with cuticular rim (pore diameter 0.5 µm; Fig. 31 A) and with small round cuticular ‘ plates’ (diameter 4 – 5 µm; Fig. 31 A). Tarsal organs with very small openings (palp: 1.2 µm; legs: ~ 0.8 µm; Figs 28 C – D, 31 B). Metatarsi 3 and 4 with long slender hairs as in male (Fig. 30 C); tarsus 4 with single prolateral comb-hair as in male (Fig. 31 D).	en	Huber, Bernhard A., Meng, Guanliang, Král, Jiří, Ávila Herrera, Ivalú M., Izquierdo, Matías A. (2023): Revision of the South American Ninetinae genus Guaranita (Araneae, Pholcidae). European Journal of Taxonomy 900 (1): 32-80, DOI: 10.5852/ejt.2023.900.2301, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2301/9981
7F0B87C4FFBF8015FDD2FC29FB89FE91.taxon	biology_ecology	Natural history The newly collected specimens were found in relatively arid environments (Fig. 34 E – F), under rocks, in leaf litter, and in the dry leaves of dead bromeliads lying on the ground. Three egg-sacs contained 6 – 7 eggs, respectively, and were carried under the prosoma.	en	Huber, Bernhard A., Meng, Guanliang, Král, Jiří, Ávila Herrera, Ivalú M., Izquierdo, Matías A. (2023): Revision of the South American Ninetinae genus Guaranita (Araneae, Pholcidae). European Journal of Taxonomy 900 (1): 32-80, DOI: 10.5852/ejt.2023.900.2301, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2301/9981
7F0B87C4FFBF8015FDD2FC29FB89FE91.taxon	distribution	Distribution Known from several localities in Salta, Tucumán, and Catamarca provinces, Argentina (Fig. 33 B).	en	Huber, Bernhard A., Meng, Guanliang, Král, Jiří, Ávila Herrera, Ivalú M., Izquierdo, Matías A. (2023): Revision of the South American Ninetinae genus Guaranita (Araneae, Pholcidae). European Journal of Taxonomy 900 (1): 32-80, DOI: 10.5852/ejt.2023.900.2301, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2301/9981
7F0B87C4FFBF8015FDD2FC29FB89FE91.taxon	description	Karyology While the preparation of the G. goloboffi specimen from Cabra Corral contained rare mitoses, prophases and metaphases I, preparations of the males from Chumbicha contained only a few premeiotic interphases and prophases of the second meiotic division. The male karyotype of the G. goloboffi specimen from Cabra Corral consisted of 11 exclusively metacentric chromosomes, namely five chromosome pairs that decreased gradually in length and a single large X chromosome (Fig. 35 E). Chromosome pairs decreased gradually in length, except for the prominent first pair. The X chromosome was twice as long as the chromosomes of the first pair. Fused sister prophases II of specimens from Chumbicha also comprised 11 chromosomes (Fig. 35 C), which confirms the diploid number and sex chromosome system. For the specimen from Cabra Corral we also obtained data on the NOR pattern. Two bivalents included a terminal NOR. Another NOR was possibly placed in the middle of an X chromosome arm. However, this was visible in only one of three metaphase I plates suitable for the detection of NORs. The sex chromosome did not differ in its intensity of condensation and staining from the other chromosomes at the mitotic prophase and metaphase (Fig. 35 E). The male prophase of the first meiotic division included a diffuse stage (Fig. 35 B). The X chromosome was positively heteropycnotic (i. e., more intensively stained than the other chromosomes) during the premeiotic interphase (Fig. 35 A) and the diffuse stage (Fig. 35 B). During the prophase of the second meiotic division (Fig. 35 C), however, it exhibited the same behavior and intensity of staining as the other chromosomes.	en	Huber, Bernhard A., Meng, Guanliang, Král, Jiří, Ávila Herrera, Ivalú M., Izquierdo, Matías A. (2023): Revision of the South American Ninetinae genus Guaranita (Araneae, Pholcidae). European Journal of Taxonomy 900 (1): 32-80, DOI: 10.5852/ejt.2023.900.2301, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2301/9981
