taxonID	type	description	language	source
DC82C0062D235FA08F17B1D83A9DD9EB.taxon	materials_examined	Material examined. Holotype: ♂, Cambodia, Kampong Chhnang Province, Khom Domnatpopol, Tonle Sap Lake, 12 ° 14 ' 14 " N, 104 ° 41 ' 15 " E, (light trap), 21. V. 2018, Rossi, Bernardi and Kong leg. (CAl). Paratypes: (1 ♂ and 3 ♀♀); 1 ♀ same data as holotype; 1 ♂ Cambodia, Kampong Chhnang, Rolea B'ier District, Toulkrolanh Village, 12 ° 13 ' 31 " N, 104 ° 39 ' 50 " E, light trap, 7. XI. 2018, W. Rossi and V. Kong leg.; 2 ♀♀, Cambodia, Kampong Chhnang, banks of Tonle Sap Lake, light trap, 17. V. 2019, W. Rossi and V. Kong leg. (CAl, CGi).	en	Allegro, Gianni, Giachino, Pier Mauro (2021): The genus Diplocheila Brulle, 1834 in Cambodia, with descriptions of two new species (Coleoptera, Carabidae, Licinini). ZooKeys 1044: 427-448, DOI: http://dx.doi.org/10.3897/zookeys.1044.60072, URL: http://dx.doi.org/10.3897/zookeys.1044.60072
DC82C0062D235FA08F17B1D83A9DD9EB.taxon	diagnosis	Diagnosis. A medium-sized to small Diplocheila (ABL: 12 - 14 mm) of the Diplocheila polita group in the subgenus Diplocheila Diplocheila (sensu Ball 1959). It is easily distinguished from the other species of the group with a sexsetose labrum (D. indus, D. laevigata, D. laevigotoides, and D. walterrossii sp. nov.) by the smaller body size (≤ 14 mm), the narrower and almost quadrangular pronotum (transverse in the four other species), with hind angles not protruding (protruding in D. indus), and by the morphology of the aedeagus.	en	Allegro, Gianni, Giachino, Pier Mauro (2021): The genus Diplocheila Brulle, 1834 in Cambodia, with descriptions of two new species (Coleoptera, Carabidae, Licinini). ZooKeys 1044: 427-448, DOI: http://dx.doi.org/10.3897/zookeys.1044.60072, URL: http://dx.doi.org/10.3897/zookeys.1044.60072
DC82C0062D235FA08F17B1D83A9DD9EB.taxon	description	Description. Habitus: ABL: 12 - 14 mm (HT ♂ 13.5 mm). Body parallel-sided, moderately shiny, black with antennae and palpi piceous-brown (Fig. 5). Head: almost quadrangular, glabrous except for the supraorbital setae, narrow in comparison with pronotum. Eyes markedly convex; a single supraorbital seta on each side. Dorsum with microsculpture not evident, only with scattered punctures visible at> 100 x magnification; frontal impressions short and superficial. Labrum symmetrically and deeply (LR = 0.78) emarginate, with six setigerous punctures on anterior margin (4 medial equidistant + 2 lateral on lobes). Clypeus trapezoid, distinctly concave anteriorly, with 1 seta on each side at anterolateral corner. Antennae moderately long, densely pubescent from segment 4, with terminal 2 articles surpassing base of pronotum; segments elongate, the second one short, as long as a half of first. Mandibles elongate, broad, approximately similar each another (the left with apical cutting edge more concave), with scrobe well-defined and glabrous and apex blunt; terebral tooth triangular and prominent. Labial and maxillary palps fusiform, with apices narrowly truncate. Thorax: pronotum smooth, with very faint microsculpture evident at> 200 x magnification and with scattered punctures, subquadrate (PW / PL = 1.18), widest at middle (Fig. 11). Disk moderately convex. Sides from rectilinear to hardly rounded in anterior half; rectilinear or very slightly sinuate backwards. Hind angles rounded obtuse, provided with a postero-lateral seta. Posterior margin rectilinear between basal impressions, which are linear and markedly impressed; anterior margin with front angles nearly obsolete. A single lateral seta on each side just above middle. Lateral bead continuous, separated from the discal area by a narrow groove, only scarcely dilated before hind angles. Medial longitudinal impression fine, nearly reaching anterior and posterior margins; anterior transversal impression absent. Elytra: moderately long (EL / EW = 1.59), parallel-sided, slightly convex and flattened on disk, widest at middle, with rounded shoulders and sides delicately sinuate before apex. Surface moderately shiny; microsculpture evident only at high magnification (> 100 x), consisting of fine, slightly transverse meshes. Epipleura without any distinct external plicae (" uncrossed epipleura "). Intervals moderately convex, smooth; striae deeply impressed on the whole length, distinctly punctate. Parascutellar stria present; scutellar setigerous pore present at base of stria 1, just before conjunction with stria 2. Basal margin complete. Discal setigerous punctures absent; umbilicate series of setigerous punctures continuous, not interrupted at middle. Hind wings fully developed. Ventral surface (thorax and abdomen): prosternum and proepisterna glabrous and impunctate (only with very fine punctures). Metepisterna as long as twice the width of anterior side; metepimera large, broadly rounded. Prosternal intercoxal process widely rounded and bordered at apex. Abdominal ventrites IV-VI shiny but shagreened at sides, glabrous except one pair subapical central setae; males with 2, females with 4 marginal setae at apex of ventrite VII. Legs: moderately slender. Posterior face of femora with 1 seta in profemora, 2 in mesofemora and metafemora. Metatrochanters glabrous and slightly shorter than half length of metafemora. Protibial antennal cleaning organ well developed, with 2 clip setae. Protibiae robust, with 4 or 5 outer apical spines; mesotibiae with a group of setae at middle of inner face; metatibiae longitudinally furrowed at inner face. Dorsal face of tarsomeres smooth. Male protarsomeres 1 - 3 distinctly dilated, slightly asymmetrical; meso- and metatarsomeres not dilated in both sexes; tarsomere 5 ventrally glabrous, dorsally with 2 apical setae; claws smooth. Male genitalia: median lobe of aedeagus short and markedly swollen before apex in lateral view (Fig. 23); the apical lamella shortly triangular in dorsal view, with blunt tip (Fig. 30), apex very slightly bent downwards in lateral view. Ostium long, in dorsal position. Right paramere oval; left paramere conchoid.	en	Allegro, Gianni, Giachino, Pier Mauro (2021): The genus Diplocheila Brulle, 1834 in Cambodia, with descriptions of two new species (Coleoptera, Carabidae, Licinini). ZooKeys 1044: 427-448, DOI: http://dx.doi.org/10.3897/zookeys.1044.60072, URL: http://dx.doi.org/10.3897/zookeys.1044.60072
DC82C0062D235FA08F17B1D83A9DD9EB.taxon	etymology	Etymology. This species is named, as a token of our esteem, after our late colleague Terry Erwin, a world-renowned specialist in world and tropical Carabidae.	en	Allegro, Gianni, Giachino, Pier Mauro (2021): The genus Diplocheila Brulle, 1834 in Cambodia, with descriptions of two new species (Coleoptera, Carabidae, Licinini). ZooKeys 1044: 427-448, DOI: http://dx.doi.org/10.3897/zookeys.1044.60072, URL: http://dx.doi.org/10.3897/zookeys.1044.60072
DC82C0062D235FA08F17B1D83A9DD9EB.taxon	distribution	Distribution and ecology. Geographical distribution: this species is recorded only from the extreme south banks of the Tonle Sap Lake, Kampong Chhnang Province, Cambodia. It seems to have a more restricted Geographical distribution than D. walterrossii sp. nov., which has been recorded from the same site as well as from other two localities in north-western Cambodia (Fig. 33). Life habits: the specimens of the type series were collected on lake banks by light trapping. No other data are available.	en	Allegro, Gianni, Giachino, Pier Mauro (2021): The genus Diplocheila Brulle, 1834 in Cambodia, with descriptions of two new species (Coleoptera, Carabidae, Licinini). ZooKeys 1044: 427-448, DOI: http://dx.doi.org/10.3897/zookeys.1044.60072, URL: http://dx.doi.org/10.3897/zookeys.1044.60072
382C4F214C4D5D4F801E642E5B814422.taxon	materials_examined	Material examined. Myanmar: HT ♂, Kawkareet in Tenasserim, Gen. Febbr. 1887, Fea legit (MCSNG) (figs 2, 6, 12, 26). Cambodia: 3 ♂♂ 2 ♀♀ Kampong Chhnang, banks of Tonle Sap Lake, 17. V. 2019, W. Rossi and V. Kong leg. (CAl, CGi); 1 ♂ Kampong Chhnang, Rolea B'ier District, Toekchenh Village, 18. V. 2019, W. Rossi and V. Kong leg. (CAl); 1 ♂, Khsam, Kampong Chhnang, 12 ° 16 ' 47 " N, 104 ° 39 ' 28.6 " E, 29. XI- 3. XII. 2019, light trap, W. Rossi and V. Kong leg. (CAl).	en	Allegro, Gianni, Giachino, Pier Mauro (2021): The genus Diplocheila Brulle, 1834 in Cambodia, with descriptions of two new species (Coleoptera, Carabidae, Licinini). ZooKeys 1044: 427-448, DOI: http://dx.doi.org/10.3897/zookeys.1044.60072, URL: http://dx.doi.org/10.3897/zookeys.1044.60072
382C4F214C4D5D4F801E642E5B814422.taxon	diagnosis	Diagnosis. Diplocheila laevigata may be distinguished by the combination of the following characters. ABL = 14 - 16 mm; head with 1 supraorbital setiferous pore on each side; labrum with 6 setae (4 medial + 2 lateral), symmetrical and moderately emarginate (LR = 0.63 - 0.65) (Figs 12, 14); anterior margin of clypeus hardly concave (Fig. 12); pronotum transverse (PW / PL = 1.29 - 1.31), with sides delicately sinuate backwards (Fig. 6); elytral striae distinctly punctate; apical lamella of the median lobe of aedeagus in dorsal view shortly triangular, with blunt apex (Figs 20, 26).	en	Allegro, Gianni, Giachino, Pier Mauro (2021): The genus Diplocheila Brulle, 1834 in Cambodia, with descriptions of two new species (Coleoptera, Carabidae, Licinini). ZooKeys 1044: 427-448, DOI: http://dx.doi.org/10.3897/zookeys.1044.60072, URL: http://dx.doi.org/10.3897/zookeys.1044.60072
900ED10AF38053AAAD0A9152CA9D3958.taxon	materials_examined	Material examined. Philippine Islands, ♀ HT, Manila, 4. II. 1914, Coll. Bottcher (BMNH) (Fig. 3).	en	Allegro, Gianni, Giachino, Pier Mauro (2021): The genus Diplocheila Brulle, 1834 in Cambodia, with descriptions of two new species (Coleoptera, Carabidae, Licinini). ZooKeys 1044: 427-448, DOI: http://dx.doi.org/10.3897/zookeys.1044.60072, URL: http://dx.doi.org/10.3897/zookeys.1044.60072
900ED10AF38053AAAD0A9152CA9D3958.taxon	diagnosis	Diagnosis. Diplocheila laevigotoides may be distinguished by the combination of the following characters. ABL = 14 mm; head with 1 supraorbital setiferous pore on each side; labrum with 6 setae (4 medial + 2 lateral), symmetrical and deeply emarginate (LR = 0.80) (Fig. 13); anterior margin of clypeus markedly concave (Fig. 13); pronotum transverse (PW / PL = 1.31), with sides markedly sinuate backwards (Fig. 7); elytral striae nearly impunctate.	en	Allegro, Gianni, Giachino, Pier Mauro (2021): The genus Diplocheila Brulle, 1834 in Cambodia, with descriptions of two new species (Coleoptera, Carabidae, Licinini). ZooKeys 1044: 427-448, DOI: http://dx.doi.org/10.3897/zookeys.1044.60072, URL: http://dx.doi.org/10.3897/zookeys.1044.60072
582C63BBE91F5B488E6973704D0409DA.taxon	materials_examined	Material examined. Holotype: ♂, Cambodia, Siem Reap Province, N Siem Reap City, 13 ° 26 ' 29 " N, 103 ° 52 ' 25 " E, light trap, 13. XI. 2018, W. Rossi and V. Kong leg. (CGi). Paratypes: (7 ♂♂ and 3 ♀♀); 2 ♂♂, same data as HT; 2 ♂♂ Cambodia, Kampong Chhnang Province, Khom Domnatpopol, Tonle Sap Lake, 12 ° 14 ' 14 " N, 104 ° 41 ' 15 " E, light trap, 21. V. 2018, Rossi, Bernardi and Kong leg.; 1 ♂ 1 ♀, Cambodia, Kampong Chhnang, banks of Tonle Sap Lake, light trap, 17. V. 2019, W. Rossi and V. Kong leg.; 1 ♂, Cambodia, Kampong Chhnang, Rolea B'ier District, Toulkrolanh Village, 12 ° 13 ' 31 " N, 104 ° 39 ' 50 " E, light trap, 7. XI. 2018, W. Rossi and V. Kong leg.; 2 ♀♀, Cambodia, Banteay Meanchey Province, near Sisophon, Campus of the Mean Chey University, light trap, 22. X- 23. XI. 2019, P. Bun and W. Rossi leg.; 1 ♂, Cambodia, Khsam, Kampong Chhnang, 12 ° 16 ' 47 " N, 104 ° 39 ' 28.6 " E, light trap, 29. XI- 3. XII. 2019, W. Rossi and V. Kong leg. (CAl, CCa, CGi, BMNH, MCSNG).	en	Allegro, Gianni, Giachino, Pier Mauro (2021): The genus Diplocheila Brulle, 1834 in Cambodia, with descriptions of two new species (Coleoptera, Carabidae, Licinini). ZooKeys 1044: 427-448, DOI: http://dx.doi.org/10.3897/zookeys.1044.60072, URL: http://dx.doi.org/10.3897/zookeys.1044.60072
582C63BBE91F5B488E6973704D0409DA.taxon	diagnosis	Diagnosis. A medium-sized (ABL: 15 - 18 mm) Diplocheila of the Diplocheila polita group in the subgenus Diplocheila Diplocheila (sensu Ball 1959). Among the species of this group having a sexsetose labrum, it is easily distinguished from D. erwini sp. nov. by the larger body size (15 - 18 mm vs 12 - 14 mm), from D. laevigata and D. laevigotoides by the more transverse pronotum (PW / PL = 1.38 vs 1.28 - 1.32), from D. indus by the hind angles of pronotum not protruding (externally protruding in D. indus) and from all these species by the morphology of the aedeagus.	en	Allegro, Gianni, Giachino, Pier Mauro (2021): The genus Diplocheila Brulle, 1834 in Cambodia, with descriptions of two new species (Coleoptera, Carabidae, Licinini). ZooKeys 1044: 427-448, DOI: http://dx.doi.org/10.3897/zookeys.1044.60072, URL: http://dx.doi.org/10.3897/zookeys.1044.60072
582C63BBE91F5B488E6973704D0409DA.taxon	description	Description. Habitus: ABL: 15 - 18 mm (HT ♂ 15.6 mm). Body parallel-sided, moderately shiny, black with antennae and palpi piceous-brown (Fig. 4). Head: almost quadrangular, robust, glabrous except for the supraorbital setae. Eyes markedly convex; a single supraorbital seta on each side. Dorsum with microsculpture not evident, only with scattered punctures visible at> 100 x magnification; frontal impressions short and superficial. Labrum symmetrically and deeply (LR = 0.72) emarginate, with six setigerous punctures on anterior margin (4 medial equidistant + 2 lateral on lobes). Clypeus trapezoid, distinctly concave anteriorly, with 1 seta at each anterolateral corner. Antennae moderately long, densely pubescent from segment 4, with terminal two articles surpassing base of pronotum; segments elongate, the second one short, as long as a half of first. Mandibles elongate, broad, approximately similar to one another (the left with apical cutting edge more concave), with scrobe well-defined and glabrous and apex blunt; terebral tooth triangular and prominent. Labial and maxillary palps fusiform, with apices narrowly truncate. Thorax: pronotum smooth, with very faint isodiametric microsculpture evident at> 200 x magnification and with scattered punctures, transverse (PW / PL = 1.38), widest just above middle (Fig. 10). Disk moderately convex. Sides moderately rounded in anterior half, delicately sinuate backwards. Hind angles rounded obtuse, with a postero-lateral seta. Posterior margin rectilinear between basal impressions, which are linear and markedly impressed; anterior margin with front angles nearly obsolete. A single lateral seta on each side at anterior third. Lateral bead continuous, separated from the discal area by a narrow groove, only scarcely dilated before hind angles. Medial longitudinal impression fine, nearly reaching anterior and posterior margins; anterior transversal impression absent. Elytra: moderately long (EL / EW = 1.65), parallel-sided, slightly convex and flattened on disk, widest at middle, with rounded shoulders and sides delicately sinuate before apex. Surface moderately shiny; microsculpture evident only at high magnification (> 100 x), consisting of fine, slightly transverse meshes. Epipleura without any distinct external plicae (" uncrossed epipleura "). Intervals moderately convex, smooth; striae deeply impressed on the whole length, delicately punctate. Parascutellar stria present; scutellar setigerous pore present at base of stria 1, just before conjunction with stria 2. Basal margin complete. Discal setigerous punctures absent; umbilicate series of setigerous punctures continuous, but punctures more widely spaced at middle. Hind wings fully developed. Ventral surface (thorax and abdomen): prosternum and proepisterna glabrous and impunctate (only with very fine punctures). Metepisterna twice as long as their width at anterior side; metepimera narrow, nearly rectangular. Prosternal intercoxal process parallel-sided with blunt apex, delicately bordered. Abdominal ventrites IV-VI shiny but shagreened at sides, glabrous except one pair of subapical central setae; males with 2, females with 4 setae at apex of ventrite VII. Legs: moderately slender. Posterior face of femora with 1 seta in profemora, 2 setae in mesofemora and metafemora. Metatrochanters glabrous and as long as one-third of metafemora. Protibial antennal cleaning organ well developed, with 2 clip setae. Protibiae robust, with 6 or 7 outer apical spines; mesotibiae with a group of setae at middle of inner face; metatibiae longitudinally furrowed at inner face. Dorsal face of tarsomeres smooth. Protarsomeres 1 - 3 of males moderately dilated, slightly asymmetrical; meso- and metatarsomeres not dilated in both sexes; tarsomere 5 ventrally glabrous, dorsally with 2 apical setae; claws smooth. Male genitalia: median lobe of aedeagus short and moderately swollen before apex in lateral view (Fig. 22); apical lamella very short and apically rounded in dorsal view (Fig. 29), apex thick and very slightly bent downwards in lateral view. Ostium long, in dorsal position. Right paramere elongate and subtruncate at apex; left paramere conchoid.	en	Allegro, Gianni, Giachino, Pier Mauro (2021): The genus Diplocheila Brulle, 1834 in Cambodia, with descriptions of two new species (Coleoptera, Carabidae, Licinini). ZooKeys 1044: 427-448, DOI: http://dx.doi.org/10.3897/zookeys.1044.60072, URL: http://dx.doi.org/10.3897/zookeys.1044.60072
582C63BBE91F5B488E6973704D0409DA.taxon	etymology	Etymology. This species is named after its collector, Walter Rossi, a world-renowned specialist in entomoparasitic fungi, as a token of our esteem and as a sign of gratitude for the gift to the authors of the specimens of the new species.	en	Allegro, Gianni, Giachino, Pier Mauro (2021): The genus Diplocheila Brulle, 1834 in Cambodia, with descriptions of two new species (Coleoptera, Carabidae, Licinini). ZooKeys 1044: 427-448, DOI: http://dx.doi.org/10.3897/zookeys.1044.60072, URL: http://dx.doi.org/10.3897/zookeys.1044.60072
582C63BBE91F5B488E6973704D0409DA.taxon	distribution	Distribution and ecology. Geographical distribution: this species is recorded from Central and North-Western Cambodia (Fig. 33). Life habits: the specimens of the type series were collected by light trapping. No other data are available.	en	Allegro, Gianni, Giachino, Pier Mauro (2021): The genus Diplocheila Brulle, 1834 in Cambodia, with descriptions of two new species (Coleoptera, Carabidae, Licinini). ZooKeys 1044: 427-448, DOI: http://dx.doi.org/10.3897/zookeys.1044.60072, URL: http://dx.doi.org/10.3897/zookeys.1044.60072
662CE56DE2A759D8894AC18070578185.taxon	materials_examined	Material examined. Cambodia: 1 ♂ Kampong Chhnang Province, Khom Domnatpopol, Tonle Sap Lake, 21. V. 2018, Rossi, Bernardi and Kong leg. (CAl); 1 ♀ Lamphun, Mueang Lamphun District, near Umong, 24. II. 2017, W. Rossi and V. Kong leg. (CAl); 1 ♂ Banteay Meanchey, near Sisophon, campus of the Mean Chey University, 1. XI. 2018, W. Rossi and V. Kong leg. (CAl); 1 ♂ Banteay Meanchey, near Sisophon, campus of the Mean Chey University, 20. V. 2019, W. Rossi and V. Kong leg. (CGi); 1 ♀ Mean Chey, 20. V. 2019, W. Rossi and V. Kong leg. (CAl); 1 ♀ Kampong Chhnang, Rolea B'ier District, Ourung Village, 20 - 23. V. 2018, Rossi, Bernardi and Kong leg. (CGi). Thailand: 1 ♂ 1 ♀ Chiang Mai, 6. V. 1988 (CGi). India: 1 ♀ Uttar Pradesh, Jhansi District, Babina, VIII. 1987 (CGi).	en	Allegro, Gianni, Giachino, Pier Mauro (2021): The genus Diplocheila Brulle, 1834 in Cambodia, with descriptions of two new species (Coleoptera, Carabidae, Licinini). ZooKeys 1044: 427-448, DOI: http://dx.doi.org/10.3897/zookeys.1044.60072, URL: http://dx.doi.org/10.3897/zookeys.1044.60072
3C1B7F5006C75DE790EAECA39CF1CC62.taxon	materials_examined	Material examined. Cambodia: 1 ♂ Kampong Chhnang, banks of Tonle Sap Lake, 17. V. 2019, W. Rossi and V. Kong leg. (CAl); 1 ♂ Kampong Chhnang Province, Sankat Kampong Chhnang, Phum Toul Ompel, banks of a branch of Tonle Sap Lake, 12 ° 14 ' N, 104 ° 41 ' E, 4. XI. 2018, W. Rossi and V. Kong leg. (CGi).	en	Allegro, Gianni, Giachino, Pier Mauro (2021): The genus Diplocheila Brulle, 1834 in Cambodia, with descriptions of two new species (Coleoptera, Carabidae, Licinini). ZooKeys 1044: 427-448, DOI: http://dx.doi.org/10.3897/zookeys.1044.60072, URL: http://dx.doi.org/10.3897/zookeys.1044.60072
